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The Marine Arthropods from the Solnhofen Lithographic Limestones (Late Jurassic, Germany) in the Collections of the Muséum National D’Histoire Naturelle, Paris

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The Marine Arthropods from the Solnhofen Lithographic Limestones (Late Jurassic, Germany) in the Collections of the Muséum National D’Histoire Naturelle, Paris The marine arthropods from the Solnhofen Lithographic Limestones (Late Jurassic, Germany) in the collections of the Muséum national d’Histoire naturelle, Paris Sylvain CHARBONNIER Muséum national d’Histoire naturelle, Département Histoire de la Terre, UMR 7207 CNRS, Centre de Recherche sur la Paléobiodiversité et les Paléoenvironnements, case postale 38, 57 rue Cuvier, F-75231 Paris cedex 05 (France) [email protected] Alessandro GARASSINO Museo di Storia Naturale di Milano, Sezione di Paleontologia, Corso Venezia 55, I-20121 Milano (Italia) [email protected] Charbonnier S. & Garassino A. 2012. — The marine arthropods from the Solnhofen Litho- graphic Limestones (Late Jurassic, Germany) in the collections of the Muséum national d’Histoire naturelle, Paris. Geodiversitas 34 (4): 857-871. http://dx.doi.org/10.5252/g2012n4a8 KEY WORDS Crustacea, Decapoda, ABSTRACT Jurassic, The palaeontological collections of the Muséum national d’Histoire naturelle, Tithonian, Kimmeridgian, Paris, France, house a broad range of marine arthropods from the Solnhofen Solnhofen, Lithographic Limestones (Late Jurassic, Germany). These arthropods include Eichstätt, decapod crustaceans, stomatopods, supposed mysidaceans, and limulids. Franconia, Germany, A commented systematic list is proposed in order to give to the specialists an paleontological collections. instrument of comparison and reference. RÉSUMÉ MOTS CLÉS Les arthropodes marins des Calcaires lithographiques de Solnhofen (Jurassique supé- Crustacea, rieur, Allemagne) dans les collections du Muséum national d’Histoire naturelle, Paris. Decapoda, Jurassique, Les collections paléontologiques du Muséum national d’Histoire naturelle, Tithonien, Paris, France, renferment un ensemble d’arthropodes marins provenant des Kimméridgien, Solnhofen, Calcaires lithographiques de Solnhofen (Jurassique supérieur, Allemagne). Eichstätt, Ces arthropodes incluent des crustacés décapodes, des stomatopodes, des Franconie, mysidacés présumés et des limules. Une liste systématique commentée est Allemagne, collections proposée dans le but de fournir aux spécialistes un outil de comparaison et paléontologiques. de référence. GEODIVERSITAS • 2012 • 34 (4) © Publications Scientifiques du Muséum national d’Histoire naturelle, Paris. www.geodiversitas.com 857 Charbonnier S. & Garassino A. INTRODUCTION to date. After their taxonomic identification, we have decided to illustrate some of these specimens The palaeontological collections of the Muséum na- in order to give to the specialists an instrument of tional d’Histoire naturelle, Paris, house a number of comparison and reference. specimens of marine arthropods from the Solnhofen Lithographic Limestones (Bavaria, Germany) includ- ing decapod crustaceans, stomatopods, mysidaceans, MATERIAL AND METHODS and limulids. The Solnhofen collection is composed of several generations of material added over the COLLECTION ARRANGEMENT last 250 years. This collection was mainly derived The history of the palaeontological collections from specimens accumulated by scientific changes housed in the Paris Museum is complicated be- between the Paris Muséum and different German cause these collections are the heritage of dif- museums and institutions. Other material was also ferent arrangements from the Royal Cabinet of collected by the French Revolutionary armies and Natural History to the public collections of the during the Napoleonic wars. Moreover, a group of new Muséum of Natural History in 1793. In specimens was traded in the late 19th century to this time, the department of palaeontology did different fossil sellers. The scientific changes were not exist at the Paris Muséum and fossils were supported by some very important French professors distributed into different collections: geology, such as Barthélemy Faujas de Saint-Fond (1741- botany, mineralogy. In 1853, when the chair of 1819), Anselme-Gaétan Desmarest (1781-1838), palaeontology was especially created for Alcide Alexandre Brongniart (1770-1847) and above all d’Orbigny, numerous professors of the Museum Alphonse Milne Edwards (1835-1900). Other refused to give him their fossils to constitute a specimens came from changes with other muse- new palaeontological collection (Brygoo 2002; ums (e.g., Manchester Museum) and donations Vénec-Peyré 2002). This conflicting situation by fortunate collectors (e.g., Boué, de Roissy, de explains the dispersion of the fossils into the Drée, du Mouy, Stuer) that sometimes collaborated Paris Museum. For instance, the type specimens with the Paris Muséum professors. Sometimes, of Eryon cuvieri Desmarest, 1817, were forgotten fossil specimens were traded to some famous sell- in a drawer of the geological collection during ers such as Émile Deyrolle (Paris, 46 rue du Bac, almost two centuries and recently rediscovered since 1831), Louis Saemann (Paris, between 1850 by one of the authors (S. Charbonnier). Today, and 1866), Meyrat (Basel, Comptoir Géologique à this situation is quite solved but there are always Birsfelden), Grebel, Wendler & Company (Geneva, numerous fossils housed in the geological collec- Comptoir Minéralogique et Géologique Suisse, tion. Consequently, this study includes specimens between 1905 and c. 1917), Jean Danhauser (Paris, housed into different buildings having at the same Comptoir Minéralogique et Géologique, 51 rue de time different catalogue numbers. Seine, between 1832 and 1836) and Dr. F. Krantz (Bonn, Rheinisches Mineralien-Kontor GmbH & FOSSIL LOCALITIES Co. KG, since 1833). As reported by Garassino & Schweigert (2006), The “Konservat-Lagerstätte of Solnhofen” is well the lithographic limestones of the Late Jurassic in known since the beginning of the 18th century and southern Germany (Bavaria) comprise a large set it is still subject of interest by the specialists of the of localities usually summarized under the name world. Therefore, the aim of the present study is to “Solnhofen”. The study of the Paris collections propose a commented list of the marine arthropods has allowed identifying mainly three localities: from the Solnhofen Lithographic Limestones housed Solnhofen, Eichstätt and Pappenheim. However, in the Paris Muséum. Indeed, several specimens the specimens reported as discovered in Pappen- housed in old collections in the Paris Muséum, heim are probably from other locality. Indeed, the are rather well preserved and totally unknown town of Pappenheim is located in the valley of the 858 GEODIVERSITAS • 2012 • 34 (4) Marine arthropods from Solnhofen Altmühl River, 5 km north of the outcrop area Palaeastacus fuciformis (von Schlotheim, 1822) of the Solnhofen Lithographic Limestones, and (3 specimens). probably these specimens were purchased there from some local persons (Günter Schweigert pers. GLYPHEIDAE comm. 2011). Following the last geological setting Glyphea pseudoscyllarus (von Schlotheim, 1822) by Schweigert (2007), the Solnhofen Lithographic (4 specimens). Limestones range from the late Kimmeridgian to the early Tithonian (Late Jurassic). The “classical” MECOCHIRIDAE lithographic limestones of Solnhofen have an early Mecochirus longimanatus (von Schlotheim, 1820) Tithonian age clearly younger than those of the (27 specimens). nearby Eichstätt district. PALINURIDAE ABBREVIATIONS Palinurina longipes Münster, 1839 (8 specimens). MNHN.F Muséum national d’Histoire na- turelle, Paris (France), Collection ERYONIDAE de Paléontologie. MNHN.GG Muséum national d’Histoire na- Cycleryon propinquus (von Schlotheim, 1822) turelle, Paris (France), Collection (5 specimens); de Géologie. Cycleryon orbiculatus (Münster, 1839) (1 specimen); Eryon cuvieri Desmarest, 1817 (16 specimens). SYstematics SCULDIDAE Sculda pennata Münster, 1840 (1 specimen). The following listing of crustaceans (including Decapoda, Stomatopoda, Mysidacea) and limulids MYSIDACEA (Xiphosurida) housed in the Paris Museum is ar- Elder ungulatus Münster, 1839 (3 specimens). ranged taxonomically. This commented systematic list includes 106 specimens identified as follows: LIMULIDAE Mesolimulus walchii (Desmarest, 1817) (4 specimens). AEGERIDAE Aeger spinipes (Desmarest, 1817) (1 specimen); REMARKS Aeger tipularius (von Schlotheim, 1822) (7 specimen); Some specimens belonging to the penaeids (10 speci- Acanthochira longipes (Oppel, 1862) (1 specimen). mens), erymids (1 specimen), mecochirids (1 speci- men) and eryonids (2 specimens) do not show the PENAEIDAE main diagnostic characters to identify them at spe- Antrimpos speciosus Münster, 1839 (6 specimens); cific level. Most of them correspond to fragmentary Rauna angusta Münster, 1839 (1 specimen); and/or incomplete remains. Therefore, we do not Bylgia hexadon Münster, 1839 (1 specimen); consider them in the commented list. Moreover, Albertoppelia kuempeli Schweigert & Garassino, three specimens corresponding to larval stage of 2004 (1 specimen); decapod crustaceans are tentatively attributed to Drobna deformis Münster, 1839 (1 specimen). Phalangites Münster, 1836. Their poorly preserva- tion does not allow supplementary dissertation (see CARIDEA Polz 1970-1973). Hefriga serrata Münster, 1839 (2 specimens). In conclusion, each identified species is listed, as follows: the specimen number, collecting locality, ERYMIDAE collector (when known), and comment. The type Eryma modestiforme (von Schlotheim, 1822) (8 speci- species is given also for each taxon. The systematic mens); arrangement follows De Grave et al. (2009). GEODIVERSITAS • 2012 • 34 (4) 859 Charbonnier S. & Garassino A. COMMENTED SYSTEMATIC LIST Genus Acanthochirana Strand,
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