Naturschutz und Biologische Vielfalt Heft 28

Bat Migrations in Europe A Review of Banding Data and Literature

Rainer Hutterer TeodoraIvanova Christine Meyer-Cords Luisa Rodrigues

Federal Agency for Nature Conservation Bonn 2005 Contents

Acknowledgements 8 Preface 9 1 Introduction 11 2 Methods and surveys 13 3 Composition and nomenclature of the European fauna (P. Benda & R. Hutterer) 17 3.1 Faunal content 17 3.2 Taxonomic and nomenclatorial changes 19 4 History and practice of bat banding in Europe 23 4.1 Aims and techniques of banding 23 4.2 Risk assessment 26 4.3 A key to European bat band signatures 28 4.4 Context and legislation 29 4.5 National reports on the history and practice of bat banding 32 4.5.1 Austria (K. Bauer) 32 4.5.2 Belgium (J. Fairon) 34 4.5.3 Bosnia and Herzegovina (B. Karapandza & M. Paunovic) 34 4.5.4 Bulgaria (T. Ivanova) 35 4.5.5 Czech Republic (J. Gaisler) 37 4.5.6 Denmark (H. J. Baagoe) 38 4.5.7 Estonia (M. Masing) 39 4.5.8 Finland (E.-M. Kyheroinen & T. Stjernberg) 41 4.5.9 France (S. Aulagnier) 41 4.5.10 Germany (R. Hutterer) 44 4.5.11 Hungary (P. Paulovics) 46 4.5.12 Italy (GIRC) 46 4.5.13 Latvia (G. Petersons & J. Kazubiernis) 47 4.5.14 Lithuania (J. Jusys, R. Patapavicius, N. Pauzien & D.H. Pauza) 49 4.5.15 Malta (J. Borg) 51 4.5.16 Portugal (L. Rodrigues & J. Palmeirim) 52 4.5.17 Romania (D. Murariu) 53 4.5.18 Serbia and Montenegro (M. Paunovic & B. Karapandza) 53 4.5.19 Slovakia (P. Kanuch) 54 4.5.20 Slovenia (B. Krystufek) 55 4.5.21 Spain (J. Benzal) 55 4.5.22 Switzerland (P. Moeschler) 58 4.5.23 Ukraine (A.-T. Bashta) 60 4.5.24 Other countries 62 5 Results of bat banding in Europe 69 5.1 Egyptian fruit-bat (Rousettus aegyptiacus) 69 5.2 Blasius' horseshoe bat {Rhinolophus blasii) 69 5.3 Mediterranean horseshoe bat {Rhinolophus euryale) 69 5.4 Greater horseshoe bat {Rhinolophus ferrumequinum) 70 5.5 Lesser horseshoe bat {Rhinolophus hipposideros) 71 5.6 Mehely's horseshoe bat {Rhinolophus mehelyi) 73 5.7 Barbastelle { barbastellus) 73 5.8 Northern bat { nilssonii) 74 5.9 Serotine {Eptesicus serotinus) 75 5.10 Savi's pipistrelle { savii) 77 5.11 Bechstein's bat {Myotis bechsteinii) 77 5.12 Lesser mouse-eared bat {Myotis blythii) 78 5.13 Brandt's bat {Myotis brandtii) 80 5.14 Long-fingered bat {Myotis capaccinii) 81 5.15 Pond bat {Myotis dasycneme) 81 5.16 Daubenton's bat {Myotis daubentonii) 83 5.17 Geoffroy's bat {Myotis emarginatus) 84 5.18 Greater mouse-eared bat {Myotis myotis) 86 5.19 Whiskered bat {Myotis mystacinus) 88 5.20 Natterer's bat {Myotis nattereri) 90 5.21 Maghreb mouse-eared bat {Myotispunicus) 91 5.22 Azorean bat { azoreum) 91 5.23 Greater noctule {Nyctalus lasiopterus) 92 5.24 Leisler's bat {Nyctalus leisleri) 92 5.25 Noctule {Nyctalus noctula) 94 5.26 Kuhl's pipistrelle { kuhlii) 96 5.27 Madeira pipistrelle {Pipistrellus maderensis) 97 5.28 Nathusius' bat {Pipistrellus nathusii) 97 5.29 Common pipistrelle {Pipistrelluspipistrellus) 100 5.30 Soprano pipistrelle {Pipistrelluspygmaeus) 101 5.31 Brown long-eared bat ( auritus) 102

4 5.32 Grey long-eared bat {Plecotus austriacus) 103 5.33 Tenerife long-eared bat {Plecotus teneriffae) 104 5.34 Parti-coloured bat { murinus) 104 5.35 Schreibers' bat {Miniopterus schreibersii) 106 5.36 European free-tailed bat {Tadarida teniotis) 108 6 Discussion and conclusion 109 References 119 Appendix 1. EUROBATS Resolution 4.6: Guidelines for the Issue of Permits for the Capture and Study of Captured Wild 163 Appendix 2. List of contributors 171 Appendix 3. Bat banding centres in Europe 175