Bulletin of the Native Society of Oregon Dedicated to the enjoyment, conservation and study of Oregon’s native and habitats

Volume 49, No. 6 July 2016 Whites in the Minority

Story and photos by Peter Lesica, Clark plants? And why is it that for some dominate in different populations. For Fork Chapter species the white-flowered forms are example, Rocky Mountain columbine This article originally appeared always rare, but in other cases they can (Aquilegia coerulea) is commonly blue in the Fall 2015 issue of Kelseya, the become quite common or even domi- in the central Rocky Mountains, where newsletter of the Native Plant nant? The answer to the first question it is most often visited by . Society. Color photos are available in the is pretty straightforward but not so for However, white-flowered forms, like online Bulletin at npsoregon.org/publica- the second one. those found in Montana, are thought tions.html The blue, red, pink or purple color to be pollinated more by hawk moths, Seems like almost every year some- of flowers is the result of pigments which are attracted to the lighter- one I know will excitedly tell me that called anthocyanins, which are manu- colored flowers. Mathew Streisfeld they saw a white shooting star or a factured by plants from flavonoids. and collaborators studied the red and white lupine. Of course I’m too jaded When one of the genes that contributes whitish forms of orange monkeyflower to get as excited as they are, but the to the production of these anthocya- (Mimulus aurantiacus [now Diplacus occurrence of albino flowers does raise nins becomes mutated, the flowers are aurantiacus]) in . They found some interesting questions. Why is it white instead of colored. Sometimes that a single gene was responsible for that albino morphs usually occur in the occurrence of white-flowered the difference in flower color, and that blue- or pink- or red-flowered species, forms relates to pollination. In this the two forms had different pollinators. but rarely occur in yellow-flowered case, white-or blue/red-colored flowers They suggest that the two forms are on their way to becoming two different species because interbreeding is unlike- ly when different insects are attracted to the different forms. Pollinator preference is not al- ways the explanation when differ- ent flower colors are both common. continued on page 7

In This Issue State Secretary Candidate.....2 New Corvallis President...... 2 Rare Plant Petitions...... 2 NPSO Calendar...... 3 New Members...... 6 Share Meeting Photos...... 6

L e sic a NPSO Contacts...... 9

Pe t e r Friends of OFP...... 10 Many populations of shooting star (Dodecatheon conjugens) include albino plants.

July 2016 1 Candidate for NPSO State Secretary OCIE T S TY N O A F L Steven Yeager (Emerald) is originally from Philadelphia. Steven moved to the P O R E in 1994 and has been rooted in Eugene ever since. He began E V G I exploring Oregon’s wild places and studying the Pacific Northwest’s native flora in

O T

A N 1995. Steven is co-owner of the Columbines School of Botanical Studies in Eu- N gene. Columbines offers a unique educational program focusing on community herbalism, including a heavy emphasis on botany that covers plant terminology F ou 61 and using taxonomic keys, ecology, and ethnobotany. He also is the QA/QC direc- nded 19 tor at Mountain Rose Herbs and is responsible for the proper identity of their bo- Native Plant tanical products. Steven currently serves as NPSO Emerald Chapter President and

is on the Boards of United Plant Savers and American Herbal Products Association Society of Oregon Foundation for Education and Research on Botanicals.

Voting for this position will take place at the Annual Meeting banquet. World Wide Web http://www.NPSOregon.org s E-mail Discussion List To join, send a message to [email protected], with the following in the body of the ­ ­message: subscribe npso General Business Address New Corvallis Chapter President P.O. Box 902 Eugene, OR 97440 or correspond directly with Officers Please welcome Judi Sanders as the president of the Corvallis Chapter. Judi has and Committee Chairs listed on the been an active NPSO member for several years, from serving as state president to inside of the back page. coordinating Native Plant Appreciation Week to working on Arundo donax legisla- tion, and more. Thank you to Jordan Brown for serving as chapter president the Membership Inquiries Only past couple of years. Matt Morales P.O. Box 80714 Portland, OR 97280 Bulletin of the Native Plant Society of Oregon s Editor: Erika Wedenoja Layout Artist: Cat Mead ISSN 0884-5999. Published 10 times NPSO Petitions for Rare Plants per year. Subscription price speci- fied in Membership Form printed on by Olivia Duren inside back cover page of this issue. Date and issue number on page 1. Any concerned Oregon resident can petition to list (or delist) native plant spe- cies on the state’s threatened and endangered plant list. NPSO members from the The submission date is the 10th Portland, Corvallis, High Desert, and Siskiyou Chapters recently took advantage of of each month for the following this opportunity and submitted petitions to increase protection for four of our rare month’s issue except for the Aug/ plant species. Sept issue which is July 24 and the The threatened and endangered plant list, maintained by the Native Plant Dec/Jan issue which is November 24. Send submissions by e-mail to Conservation Program of the Oregon Department of Agriculture (ODA), is an [email protected]. important tool for protection and management of listed plants on Oregon’s non- federal public lands. Citizens are welcome to submit scientific evidence of increas- Opinions expressed in this publica- ing or decreasing threats to native species that justify amendments to this list. tion are those of the authors of the Species that are most appropriate for listing are native to Oregon and rare through- articles. They do not represent the out their range, face threats to natural reproduction potential or habitat, and are opinions of the Native Plant Society of Oregon, unless so stated. inadequately protected by existing state or federal programs or regulations. While a plant must meet certain criteria to be considered for listing (as described in Oregon continued on page 6

2 NPSO Bulletin NPSO Calendar Service will host an open house. Come State Notes Corvallis see 350 different types of blueberries. NPSO Annual Meeting 2016 For questions, information on upcoming You can taste more than 100 different events, or to be on the Corvallis chapter e- blueberry varieties and see the field July 15–17, Friday–Sunday mail list, contact co_president@NPSOre- collection of northern and southern Emerald Chapter will host the 2016 gon.org. highbush blueberry, lowbush blueberry, NPSO Annual Meeting on a private half-highs, and the rabbiteye cultivars, forested 46 acre property near the ma- July 7, Thursday, 1–5 pm among other species from around the jestic McKenzie River, a fifty-minute Open House: National Germplasm world. Location: 33447 Peoria Road, drive from Eugene. McKenzie River Repository. The U.S. Department Corvallis. Conference Center, Rainbow, Oregon. of Agriculture, Agricultural Research July 9, Saturday, 8 am A state board meeting will be held on Sunday. Visit the Annual Meeting web- Field Trip: Iron Mountain and Cone site, emerald.npsoregon.org/2016mtg, IMPORTANT NOTE TO Peak. Hike the Iron Mountain Cone for the latest information. FIELD TRIP PARTICIPANTS Peak loop, identifying wildflowers in Field trips take place rain or this botanical hotspot. We will return Blue Mountain shine, so proper dress and in the late afternoon. Bring a lunch For information on Blue ­Mountain footwear are ­essential. BRING and water, dress for the weather, and Chapter call Jerry Baker,541-566- WATER AND LUNCH. Trips may wear sturdy shoes. Location: meet at 2244. To get on our email list, contact be strenuous and/or hazard- the OSU Wilkinson Hall parking lot [email protected]. ous. Participation is at your southwest of the Beanery at 26th and own risk. Be prepared to sign Monroe. For more information and to Cheahmill a release form indicating this. RSVP, contact Dan: 541-752-8860 or A sample waiver form is avail- [email protected]. For questions, or to be added to the able at http://npsoregon.org/ Cheahmill Chapter's email list for re- documents/NPSO_waiver.pdf. July 10, Sunday, 10 am–noon minders of upcoming programs and Please contact the trip leader Work Party: Avery House. Come events, please contact the Cheahmill or chapter representative about and help with summer clean-up at the Chapter President at ch_president@ difficulty, distance, and terrain Avery House native plant garden. Bring NPSOregon.org. to be expected on field trips. gloves, clippers, trowel, and a bucket. Meeting and work party location: Dogs are not allowed. All NPSO field trips are open to the pub- Contact Esther for more information at Carnegie Room, McMinnville Public lic at no charge (other than 541-754-0893 or [email protected]. Library, 225 NW Adams St. (corner of contribution to carpool driver) July 16, Saturday, 9 am–noon Highway 99W and SW Second St.), and newcomers and visitors are Field Trip: Backcountry of Bald Hill McMinnville, Ore. 97128. Parking lot always welcome. You must pre- entrances on Adams and Second streets. register for all events. National Farm. Matt Blakeley-Smith will take Meetings start at 7 pm. Doors open at Forests require a Northwest us to parts of Bald Hill Farm that are 6:45 pm. for socializing. Forest Pass for some field trip not open to the public. We will explore locations. Permits can be ac- the hidden corners of the property for July 23 and August 27, Satur- quired at forest headquarters milkweed patches and monarch but- days, 10 am–noon and ranger districts. terflies. This is a rugged off trail hike Work Party: Native Plant Garden. of about 3 miles that may involve Join NPSO volunteers tending the na- NOTICE TO FIELD TRIP some bush whacking and jumping tive plant garden at McMinnville Public CHAIRS AND LEADERS over creeks. Location: meet at the Library. Dress for the weather and bring The Forest Service and other coffee stand at the Benton County gloves and gardening tools if possible. agencies have set policies limit- Fairgrounds, 53rd and Reservoir Ave. No experience is necessary; the garden ing group size in many wilder- For more information, contact Matt is a great place to get more familiar with ness areas to 12. The reason Blakeley-Smith, [email protected] local native species. Location: McMin- is to limit human impacts on or 541-231-9861. nville Public Library, see above for these fragile areas. Groups details. For more information, contact using wilderness areas should be no larger than 12. A trip Emerald Susan Williams at helgesusan@comcast. leader checklist and sample Visit the Emerald Chapter Web site at net or 503-538-1865. waiver form are available at: http://emerald.npsoregon.org to learn more http://npsoregon.org/calendar. about the latest chapter events, plant lists html & botanical information about Lane

July 2016 3 County plants & the people who love them. Clatsop County. Coniferous forests org) for details and registration. Meeting Location: Conference Room with Abies amabilis, A. grandis, A. proc- at Lane County Mental Health, 2411 era, and Taxus brevifolia; riparian zones, Klamath Basin Martin Luther King Blvd. Turn off seeps, meadows, talus slopes, and a For information on the Klamath Basin MLK Blvd. onto Scout Access Rd. summit rock garden. Saddle Mountain Chapter, visit http://klamathbasinnps.com across from Autzen and take the first supports high species diversity, endem- or contact the chapter president, Kim El- left into the parking lot. The meeting ics (e.g., Micranthes hitchcockiana), and liott, [email protected]. room is on the right side as you face the species at their northern (e.g., Carda- 252. building. mine californica), southern (e.g., Geum triflorumvar. ciliatum) and western July 9, Saturday, 9 am–4 pm (e.g., Anemone multifida) range limits. Mid-Columbia Field Trip: Upper Elk Meadows. Alan The trail climbs 1600 ft. in 2.5 miles For information on the Mid-Columbia Curtis leads this hike, located 30 miles for a strenuous 5 mile round trip, but Chapter, contact Sara Wu at mc_presi- from Cottage Grove. Elevation is 4,000 at a botanist's pace we won't cover [email protected]. ft. Over 200 species of vascular plants the whole trail. Location: meet at the The Mid-Columbia Chapter will be in have been identified here. We expect to Saddle Mountain trailhead off High- the field for the summer, so no meet- see Frasera umpquaensis and several or- way 26 at 9 am. Contact Carla Cole at ings will be held until our September chid species. Surrounding the meadow [email protected] to register and if program. Happy botanizing until then. is an old-growth forest of Abies amabi- interested in carpooling. The hike will lis, Abies grandis, and Pseudotsuga men- be limited to 12 participants. Portland ziesii. We will walk 1.5 miles through If you would like to receive the chapter’s moist meadow. Wear boots and bring High Desert semi-official email newsletter,The lunch and water. Location: meet at For information, visit the High Desert Calochortus, email Don Jacobson at South Eugene High School at 19th and Chapter website: www.highdesertnpsore- [email protected]. Patterson. For more information call gon.org and Facebook page: Native Plant 541-345-5531. Society of Oregon: High Desert Chapter, Facebook: Take a look at our Face- July 23, Saturday, 8:30 am or email [email protected]. book page. Native Plant Society of Or- August 10, Wednesday egon-Portland-Chapter. It will have the Field Trip: South Jetty Dunes. Ed most up-to-date information on hike Alverson will lead an exploration of Field Trip: Fen Exploration with Rick and other events as well as great plant coastal dune habitats just south of Dewey. We will explore a fen on the pictures. Florence. In the morning we will visit Soda Creek drainage with Rick Dewey, South Beach Park, a 300 acre undevel- the Forest Service's bryophyte (moss) Meeting Location: The Portland oped Lane County Park with extensive expert and wetland enthusiast for the Chapter meets the second Thursday of dune and deflation plain communities, entire Pacific Northwest. Fens are rich most months. Since January 2016 we to develop a plant list. An optional af- in biodiversity and Rick is loaded with have been meeting at TaborSpace in ternoon hike will visit a nearby site on information. Come appreciate the Mt. Tabor Presbyterian Church, 5441 U.S. Forest Service land to look for an minutia and learn about this unique SE Belmont St., Portland, 97215. Enter adder's-tongue, Ophioglossum pusillum, habitat. A hand lens and rubber boots the church through the main door off population that was last observed in the are recommended. Check our website Belmont and turn right to enter Cope- 1990s. Each hike will be about 2 miles (www.highdesertnpsoregon.org) for land Commons. Parking is available in round trip. Location: meet at South Eu- details. the church lot on the NW corner of SE 54th and Belmont. Meetings and pro- gene High School at 19th and Patterson August 20–21, Saturday–Sunday, 9 grams begin at 7:00 pm. to carpool. For more information, call am–4:30 pm Ed Alverson at 541-461-1958. August 6, Saturday, 8 am Class: Botanical Illustration. Learn Filipendula botanical illustration with local artist Field Trip: Silver Star Mountain. The Rebecca Brown-Thompson in her stu- rocky soil and open meadows of Silver This is a newly formed chapter for the Star Mountain offer subalpine and northern Oregon coast area. For more dio. You can take one or both classes. Saturday: field sketching class. Learn alpine wildflowers not usually seen at information, visit our Facebook page: this low (4,390 ft.) elevation. Climbing NPSO-Filipendula Chapter, or contact drawing techniques with an emphasis on plant illustration. $25 material fee. higher takes us back in time to earlier Field Trip Chair Carla Cole, colecarla@ spring flowers, includingXerophyllum yahoo.com. Sunday: color class. Watercolor painting and colored pencil techniques and color tenax, Calochortus subalpinus, Erythro- July 9, Saturday, 9 am mixing. $35 material fee. Some drawing nium montanum, Lewisia columbiana, Hike: Saddle Mountain. Join us for experience (i.e. the field sketching class) Ligusticum grayi, and five penstemon an exploration of the diverse habi- is required for the color class. Check species. The summit view offers five- tats found on the tallest mountain in our website (www.highdesertnpsoregon. volcanoes, the Columbia River, and

4 NPSO Bulletin Vancouver and Portland. This moder- July 30, Saturday, 10 am ate hike is 5.6 miles round trip with a Field Trip: Crater Lake National Willamette Valley 1240 ft. gain along a high ridge. Hiking Park. We will visit Poison Meadows on For program information, contact John boots and poles recommended, bring the western boundary of the Park. Join Savage at 503-399-8615 or Alexandra lunch, water, binoculars, etc. For more Jennifer Beck, Crater Lake National Ninneman at 503-391-4145. information and to sign up, contact Park botanist, for a moderate off trail July 3, Sunday, 7 am–5 pm Ginny at [email protected] and hike on rolling terrain through forest Field Trip: Iron Mountain. Enjoy one include a cell phone number for the stands and meadows. We will see a va- of the best wildflower hikes in Oregon. hike day. riety of plants, including Mt. Mazama Iron Mountain (near Sweet Home) has September 8, Thursday, 7 pm (Collomia mazama), leather grape-fern over 300 different species amid stands Program: Conservation and Johnson (Sceptridium multifidum), and moun- of trees and meadows. John Savage of Creek. Noah Jenkins, riparian pro- tain owl clover (Orthocarpus imbrica- the Willamette Valley Chapter will lead gram manager of the Johnson Creek tus). Bring lunch, hat, and sunscreen. the all-day hike over a seven-mile loop Watershed Council, will discuss native Location: meet at the junction of Deer with elevation gain. Bring food and wetland plants, removal of invasive Creek Road (NF 6260) and Highway water. Free and open to the public. For vegetation, and the conservation efforts 62 (just east of the junction of High- more information and to register, call at Johnson Creek Watershed. Location: ways 62 & 230). Dirt road drive to John Savage at 503-399-8615 after 7 Copeland Commons at TaborSpace, hiking spot is 30-45 minutes. For more pm. Mt. Tabor Presbyterian Church, 5441 info, contact [email protected]. SE Belmont in Portland. Contact: William Cusick Mary at [email protected] or South Coast Visit our Web site at http://williamcusick. 503-288-4340. This is a newly formed chapter. For npsoregon.org or contact Susan Geer at more information, contact the chapter 541-963-0477 or Emelie Montgomery- Siskiyou president, Alexis Brickner, sc_president@ Jones at 541-963-3339 for updates and To join the Siskiyou Chapter email list: NPSOregon.org. general information. Chapter notices Send an email, from the address at which and communications are done primarily you want to receive announcements, to Umpqua Valley through a Google group. Members are re- [email protected]. No Visit umpquavalleynativeplants.com for minded to contact Susan or Emelie if they subject or message is required. You will get more information on chapter activities or want to be added to the Google group, a confirmation email. If you would like to to be added to our email list. Check out or if they do not have internet access and volunteer to lead a hike in 2016, make a our Facebook page, www.facebook.com/ want to be contacted by phone for events. suggestion, or provide feedback regarding UmpquaValleyNPSO. Not online - call the field trip program, please contact Julie Donna Rawson 541-459-2821. Spelletich at: [email protected]. Join us August 11, Thursday, 7 pm on Facebook at: facebook.com/SiskiyouChap- Presentation: Restoring the Monarch terNativePlantSocietyOfOregon. Butterfly with Native Plants. The July 10, Sunday, 8:30 am monarch butterfly, which migrates Field Trip: Cook and Green Pass Bo- thousands of miles and has traditionally tanical Area. We will visit Elk Meadow flourished throughout much of North Basin in the California portion of the America, including here in Oregon, is Applegate watershed. Focus of the in decline. Tom Landis, retired U.S. trip is the flora and a unique wetland. Forest Service nursery specialist, will Brewer’s spruce, Baker’s cypress, azaleas, share the current situation of this but- Wiggins’ lily, California ladyslipper terfly and discuss its primary host plant, orchids, Newberry’s gentian, Klamath the milkweed, and the other important Mtn. serpentine endemics, etc. Cross- nectar plants (ones you can grow in your yard) that help support the but- h country hiking with lots of elevation gain and loss but not too brushy. Plan terfly. Find out what we can do to help on a long day. Location: rendezvous reverse downward population trends and carpool in the large public park- for this impressive species. Our chapter ing lot at the west end of West D St. in has Asclepias speciosa seed to give away. Jacksonville, Oregon. Call trip leader Refreshments will be served. Location: Wayne Rolle, 541-482-0093, for more Mercy Community Education Center, information. Folks coming from other 2459 Stewart Parkway, Roseburg. locations may call the trip leader to ar- range additional rendezvous points.

July 2016 5 Rare Plants, cont'd New NPSO Members Administrative Rule 603-073), the pe- tition process is meant to be accessible Please welcome these new members who joined NPSO between January 11, 2016 to any knowledgeable plant enthusiast. and June 12, 2016 NPSO volunteers each chose their favorite rare plant from a list whittled Cheahmill High Desert down to the most deserving candidates Alexander Petit The Garden Specialist by botanists at the Oregon Biodiversity Ruth Bucklin Information Center (ORBIC). After Corvallis Sarah Canham gathering the required information and Gail Arbenbright Judith Foster making our best case, we traded peti- Kimberly Conley Lisa Foster tions for peer review and finally sub- Ann Geyer Irv Givot mitted petitions for Acker Rock wild Michael Gough Patricia Green buckwheat (Eriogonum villosissimum), Richard Little Virginia Harmon a recently described narrow endemic Gary Radice Suzanne & Greg Johannsen found on rock outcrops near Crater Cindy Sagers Emilie Marlinghaus Lake; fragrant kalmiopsis (Kalmiopsis Isaac Sandlin Lori L. Neal fragrans), a shrub associated with steep Zak Weinstein Jake Ringold rocky slopes in the southern Cascades; Tina Smith Ochoco lomatium (Lomatium ocho- Emerald William & Frances Tweed cense), a newly described endemic to Heron Brae Botanical Studies Gerhard Weber lava tablelands in the Ochoco Moun- Anna Bradley Krista Winchell tains; and sessile mousetail (Myosurus Keiko Bryan Cindy Zalunardo sessilis), a tiny annual obligate of vernal Joshua Caraco pools mostly known from Oregon’s Emily Dinan Klamath Basin Columbia Basin as well as from Patty Driscoll Jan Goodyear California. Barbara Dumesnil ODA staff has 90 days from sub- Chuck Gibson Members At Large mittal to assess petitions and decide Robert Ivey Judy Bevington whether or not there is enough sci- Theodora Jaster Alexa DiNicola entific evidence to proceed. Petitions Richard Lemon Forrest Pierce without enough evidence are dismissed, Brianna McTeague while those that are substantiated go on Johanna Schmitt Mid-Columbia to a detailed two-year review to deter- Bud Taylor Izzy Baptista mine listing of the plant as threatened, Louise Sargent endangered, or unlisted. We are happy Filipendula to report that all four of our petitions Vanessa Garner Portland were successful in clearing the first Claudia Rosenthal Robin Albrecht hurdle, and are now undergoing in- Brooke Stanley Amber Ayers depth review! James Thompson Deborah Baker Catherine Varing Reid Bramble Staff at ODA’s Native Plant Con- continued on page 7 servation Program, together with many collaborators, recently began a major review of Oregon’s threatened and endangered plant list that includes consideration of new species to be Share Your Annual Meeting Photos and added, as well as species to be dropped. While new petitions for amending the Stories list are welcome any time, those sub- Are you ready for this year’s Annual Meeting in the McKenzie River area? Per- mitted before the end of 2016 will be haps you will see all eight of the area’s Castilleja species on field trips, catch up with particularly helpful to this effort. If you old friends, or volunteer to help the event run smoothly. After the Annual Meet- are interested in developing a petition, ing, you are invited to gather your memories into text, photos, or illustrations and please contact Olivia Duren at olivia- share them in the Bulletin. The October issue will feature an Annual Meeting recap [email protected] for ideas of the best from the Emerald Chapter and members like you. Please submit your articles and/ candidate species, as well as a petition or high resolution images (minimum of 120 dpi) to [email protected] by template and information resources. September 10.

6 NPSO Bulletin New Members, cont'd Whites, cont'd eye color in humans. Elizabeth Brewster Sandblossoms (Linanthus parryae) is White and colored flowers may Dennis Brown an annual of the Mojave Desert with both be common in some species, but Kim Brown blue and white color forms. Some loca- more often than not albino forms are Heather Buley tions have a preponderance of white- rare throughout a plant’s range. So Carolyn Caldwell flowered plants, but just a little ways what’s with that? People often assume Wing S. Chan away the blue-flowered form predomi- that albinos are rare because they lose Darlene Chirman nates. Doug Schemske and Paulette their pollinators. For example, a com- Thomas James Dee Bierzychudek examined one such site mon species of larkspur (Delphinium Marci Degman and found strong evidence that differ- nelsonii) is ordinarily blue, but rare Sheila DeLaquil ent plant communities with different white-flowered plants occur near Rocky Weina Dinata soils supported different color forms in Mountain Biological Lab in Colorado. Matt Erickson spite of the fact that they were in close Experiments conducted by Nick Waser Verleen Fletcher spatial proximity. They were unable to and Mary Price showed that white- Zeph Friedman-Sowder pinpoint the exact mechanism, but it flowered plants produced very few Jerilyn Gauthier must have been some trait(s) that was seeds because the primary pollinators, Sara Geonczy connected to flower color rather than hummingbirds and bumblebees, dis- Toni Gonzalez the flower color itself. criminated against them. Christina Gullion Missoula phlox (Phlox kelseyi var. However, pollination is often not Andrea & Ted Heid missoulensis) occurs on the hills imme- involved in albino rarity because an- Melissa Jackson diately north of Missoula. Plants with thocyanins do more than just color the Lillian & Jericho Knight deep blue or white or intermediate flowers. They occur throughout plant Wendy & Robert Lee flowers occur together. It was assumed tissue and have been shown to protect Jan Lopez that flower color affected pollination, the leaves and stems from damage by Daniel Mathews but Lisa Campbell, a graduate student too much solar radiation. Anthocya- Angie Neff at the University of Montana, spent nins and especially their flavonoid pre- Antonya Pickard many hours observing all the different cursors also help protect plants against Kevin & Emily Schoenfelder color forms and found that the same diseases and insect pests. So anthocya- Karen Schwartz insects in equal numbers visited all nins and the genetic machinery that Mike & Laurie Skinner the forms, and there was no difference make them have a number of functions Elizabeth Stiles in seed set. In this case it appears that besides just coloring the flowers and Margie Willis flower color has little bearing on sur- helping attract pollinators, although Byerly Woodward vival or reproductive success, just like this is probably what is most noticeable continued on page 8 Siskiyou Tony Alexander Kim Chignell Wendy J. Gere Bonnie Johnson Peggy Martin Goly Ostovar Joan Rowe

Umpqua Valley Michael & Madalyn Dixon Laurel Fisher

Willamette Valley Mahonia Nursery Zena Learning Center LLC Richard Jorgensen Sharon & John Krimmel Matthew Palmquist L e sic a Pe t e r Rare albino Lewis monkeyflower.

July 2016 7 Whites, cont'd antioxidants. She’ll thank you for it. to us. This situation, when a single Further reading flower color in the desert annual gene or gene complex influences more Linanthus parryae: was Wright right? Dick, C. A., J. Buenrostro, T. Butler, Evolution 61: 2528–2543. than one trait, is called pleiotropy, and M. L. Carlson, D. J. Kliebenstein it is the overarching reason that albino Streisfeld, M. A., W. N. Young and J. and J. B. Whittall. 2011. Arctic M. Sobel. 2013. Divergent selec- plants are rare in most populations, al- mustard flower color polymorphism though the details vary among species. tion drives genetic differentiation in controlled by petal-specific down an R2R3-MYB transcription factor Drummond’s phlox (Phlox drum- regulation at the threshold of the mondii) is a pink- or red-flowered spe- that contributes to incipient specia- anthocyanin biosynthetic pathway. tion in Mimulus aurantiacus. PLoS cies endemic to Texas with rare white- PLoS One e18230. flowered forms occurring sporadically. One 9 e1003385. Levin, D. A. and E. T. Brack. 1995. Warren, J. and S. Mackenzie. 2001. Donald Levin and his collaborators Natural selection against white found that albino plants were visited Why are all colour combinations petals in Phlox. Evolution 49: not equally represented as flower- as often by their insect pollinators as 1017-1022. were their pink-or red-flowered neigh- colour polymorphisms? New Phy- Miller, R. B. 1981. Hawkmoths and tologist 151: 237–241. bors. However, white-flowered plants the geographic patterns of floral produced fewer flowers and had much Waser, N. M. and M. V. Price. 1981. variation in Aquilegia caerulea. Evo- Pollinator choice and stabilizing se- lower survivorship, and the research- lution 35: 763-774. ers found that the loss of the gene lection for flower color in Delphini- Schemske, D. W. and P. Bierzychudek. um nelsonii. Evolution 35: 376-390. that produced pink or red flowers also 2007. Spatial differentiation for made the plants less able to cope with dry conditions that can be common in Texas. Northern wallflower (Parrya nudi- caulis) is a mustard that is common in Alaska. The flowers are usually purple. White-flowered plants do occur, but they are more common in the south- ern part of the state and virtually non-existent in the north. Researchers found that the same biochemical path- way that produces the purple pigment also enhances cold tolerance. For this reason white-flowered plants can persist in the warmer parts of the state (if any parts can be said to be warm), but do not occur where it is colder. John Warren and Sally Mackenzie studied five species of British plants that have both bluish and whitish color forms. They found that blue- flowered plants performed better under drought-induced stressful conditions. They surmised that the flavonoids and anthocyanins, which are anti-oxidants and were lower in white-flowered plants, helped the blue-flowered plants overcome stress. It’s a complex world out there. That little albino flower you found could be suffering from lack of pol- lination or cold or drought or some disease. Perhaps you should take your little white buddy to the natural food L e sic a

store nearest you and get her some Pe t e r A completely albino population of lupine (Lupinus argenteus) near Hot Springs, MT

8 NPSO Bulletin Native Plant Society of Oregon Membership Form Name______DUES include monthly Bulletin and Kalmiopsis when published. Membership is for the calendar year. New memberships enrolled after Address______July 1 include the following year.

City______State______Zip+4______New Renewal Student – $12 Regular – $25 Family – $35 Phone ______E-Mail ______Sustaining – $60 Patron – $125 Life Membership – $500 Chapter (if known)______Subscription Only (Bulletin and Kalmiopsis) – $25 Only for those who wish to subscribe, but do not want to have full membership status. Is this a change of address? If so, please write your old address here: CONTRIBUTIONS to NPSO are tax deductible. Address______Jean Davis Memorial Scholarship Fund $ ______Leighton Ho Memorial Field Botany Fund $ ______City______State______Zip+4______Augusta Rockafellar Memorial Scholarship $ ______ Check to receive electronic Bulletin, sent by email in lieu of paper. Rare and Endangered Plant Conservation Fund $ ______Please make checks for dues and contributions payable to: Friends of the Oregon Flora Project $ ______native plant society of oregon Membership in the Native Plant Society of Oregon is open to all. Send completed form and full remittance to: Membership applications, renewals and changes of address (include Native Plant Society of Oregon old address) should be sent to the Membership Chair. Changes of P.O. Box 80714 address may also be e-mailed to [email protected]. Portland, OR 97280 Officers & Committee Chairs State officers Directors...... Sam Friedman, Linda Hardison, Kevin Weitemier (to 6/2016); Mary Beth Averill, Susan Berger, Billy Don Robinson (to 6/2017) President...... Kelli Van Norman...... [email protected] Immediate Past President...... Judi Sanders...... [email protected] Vice President...... Lisa Blackburn ...... [email protected]; 503-538-3976 Secretary...... Rosalie Crowe...... [email protected] Treasurer...... Cyndi Dion ...... 897 Hillview Dr., Ashland, OR 97520-3517; [email protected]; 541-482-8603 Chapter Presidents Cheahmill ...... Susan Aldrich-Markham ...... [email protected]; 503-537-0587 Corvallis...... Judi Sanders...... [email protected] Emerald (Eugene)...... Steven Yeager...... [email protected]; 541-521-3964 Filipendula...... Kathleen Sayce...... [email protected]; 360-665-5292 High Desert (Bend)...... Desiree Johnson...... [email protected] Klamath Basin...... Kim Elliott...... [email protected]; 541-331-5704 Mid-Columbia...... Sara Wu ...... [email protected] Portland...... Linda Hardie...... [email protected]; 503-309-8810 Siskiyou...... Kristi Mergenthaler...... [email protected]; 541-941-3744 South Coast...... Alexis Brickner ...... [email protected] Umpqua Valley (Roseburg)...... Neal Hadley ...... [email protected]; 541-679-8889 Willamette Valley (Salem)...... John Savage ...... [email protected]; 503-399-8615 Wm Cusick (La Grande)...... Susan Geer...... [email protected] State Committee Chairs Budgets and Grants...... Dan Luoma...... 5 NW Edgewood Dr., Corvallis, OR 97330; 541-752-8860 Conservation, East Side...... Billy Don Robinson...... [email protected]; 503-515-1708 Conservation, West Side...... vacant...... Friends of OR Flora Project. Linda Hardison...... 2082 Cordley Hall, Corvallis, OR 97331-2902; [email protected]; 541-737-4338 Legislative...... Billy Don Robinson...... [email protected]; 503-515-1708 Membership...... Matt Morales...... P.O. Box 80714, Portland, OR 97280; [email protected] NPSO Fellows...... Cindy Roché and Kareen Sturgeon...... [email protected] Rare & Endangered Plants..... Jason Clinch...... [email protected] Publications x Bulletin Editor...... Erika Wedenoja...... [email protected] Bulletin Production...... Cat Mead...... [email protected] Publications Committee Chair. Cindy Roché...... [email protected] Webmaster...... Sunia Yang...... [email protected] July 2016 9 Send change of address notices and membership requests to: NonProfit ORG Native Plant Society of Oregon US Postage Matt Morales, Membership Chair Paid P.O. Box 80714 Eugene, OR Portland, OR 97280 Permit No. 204 RETURN SERVICE REQUESTED

TIME DATED MAIL

Friends of the Oregon Flora Project Version 1.6 of the Oregon Vascular Plant Checklist has been recently published on our website. Some highlights from this important milepost for plant nomenclature: • There are 4,620 species, subspecies and varieties of vascular plants in Oregon • 231 changes were reflected in the Checklist www.oregonflora.org • 77 plants are now referred to by a different name • The manuscripts from a completed portion of Volume 2 of the Flora of Oregon (plant families Cabombaceae through Euphorbiaceae) are reflected in the Checklist This information is now reflected in the online Atlas and Photo Gallery. Details can be found in the table “Summary of Recent Checklist Updates” on the checklist page at: http://oregonflora.org/checklist.php Work on Volume 2 is progressing—as of mid-May we have 62% of the manuscripts for the ~1670 taxa in hand! If you find the OFP website helpful in your enjoyment and study of plants, please contribute to insure we can continue to provide this valuable tool! Contributions can be made online (www.oregonflora.org/supportOFP. php) or mailed directly to: Agricultural Research Foundation, 1600 SW Western Blvd. Ste. 320, Corvallis, OR 97333. Please make checks payable to the Agricultural Research Foundation and include “OFP” on the memo line.

10 NPSO Bulletin