Epiphytes of Las Cuevas Research Station, Cayo District, BELIZE 1 Bruce K. Holst, Sally Chambers, Antone Jones1 & David Amaya, Ella Baron, Marvin Paredes, Sayuri Tzul2 1Marie Selby Botanical Gardens, 2Ian Anderson’s Caves Branch Botanical Garden © Marie Selby Botanical Gardens ([email protected]), Ian Anderson’s Caves Branch Botanical Garden ([email protected]). Photos by D. Amaya (DA), E. Baron (EB), S. Chambers (SC), W. Collier (WC), B. Holst (BH), E. Kay (EK), A. Jones (AJ), P. Nelson (PN), M. Paredes (MP). Support from Marie Selby Botanical Gardens, Ian Anderson’s Caves Branch Jungle Lodge.

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Fruit Flower and Flower seed-bearing

Anthurium pentaphyllum Anthurium pentaphyllum Anthurium scandens Anthurium scandens (Araceae; summit; BH) (Araceae; summit; BH) (Araceae; EB) (Araceae; EB)

Fruit Flower

Anthurium schlechtendalii Anthurium schlechtendalii Anthurium schlechtendalii Monstera deliciosa Monstera deliciosa (Araceae; BH) (Araceae; EB) (Araceae; BH) (Araceae; AJ) (Araceae; PN)

Monstera sp. Philodendron hederaceum Philodendron hederaceum Philodendron radiatum Philodendron radiatum (Araceae; BH) (Araceae; AJ) (Araceae; EK) (Araceae; BH) (Araceae; BH)

Philodendron smithii Philodendron smithii Philodendron tripartitum Philodendron tripartitum Syngonium podophyllum (Araceae; BH) (Araceae; BH) (Araceae; EB) (Araceae; EB) (Araceae; BH) Epiphytes of Las Cuevas Research Station, Cayo District, BELIZE 2 Bruce K. Holst, Sally Chambers, Antone Jones, David Amaya, Ella Baron, Marvin Paredes, Sayuri Tzul

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Fruit Flower

Syngonium podophyllum Syngonium podophyllum Oreopanax guatemalensis Oreopanax guatemalensis Begonia glabra Female flowers (Araceae; BH) (Araceae; BH) (Araliaceae; EB) (Araliaceae; EB) (Begoniaceae; EB)

Begonia glabra Androlepis skinneri Androlepis skinneri (inset, fruit) berteroniana (Begoniaceae; EB) (; EB) (Bromeliaceae; EB) (Bromeliaceae; BH) (Bromeliaceae; EB)

Catopsis juncifolia Catopsis juncifolia Tillandsia brachycaulos Tillandsia brachycaulos Tillandsia bulbosa (Bromeliaceae; PN) (Bromeliaceae; PN) (Bromeliaceae; EB) (Bromeliaceae; EB) (Bromeliaceae; BH)

Tillandsia bulbosa Tillandsia festucoides Tillandsia festucoides Tillandsia filifolia Tillandsia filifolia (Bromeliaceae; EB) (Bromeliaceae, BH) (Bromeliaceae, EB) (Bromeliaceae; EB) (Bromeliaceae; EB) Epiphytes of Las Cuevas Research Station, Cayo District, BELIZE 3 Bruce K. Holst, Sally Chambers, Antone Jones, David Amaya, Ella Baron, Marvin Paredes, Sayuri Tzul

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Tillandsia juncea Tillandsia juncea Tillandsia pruinosa Tillandsia pruinosa Tillandsia schiedeana (Bromeliaceae, EB) (Bromeliaceae, EB) (Bromeliaceae; PN) (Bromeliaceae; PN) (Bromeliaceae; EB)

Tillandsia schiedeana Tillandsia variabilis Tillandsia variabilis heliconioides Vriesea heliconioides (Bromeliaceae; PN) (Bromeliaceae; EB) (Bromeliaceae; EB) (Bromeliaceae; EB) (Bromeliaceae; EB)

Epiphyllum oxypetalum Epiphyllum oxypetalum Pseudorhipsalis ramulosa Pseudorhipsalis ramulosa Rhipsalis baccifera (Cactaceae; EB) (Cactaceae; EB) (Cactaceae; BH) (Cactaceae; EB) (Cactaceae; EB)

Rhipsalis baccifera Anathallis sertularioides Anathallis sertularioides Camaridium densum Camaridium densum (Cactaceae; EB) (Orchidaceae; BH) (Orchidaceae; MP) (Orchidaceae; BH) (Orchidaceae; BH) Epiphytes of Las Cuevas Research Station, Cayo District, BELIZE 4 Bruce K. Holst, Sally Chambers, Antone Jones, David Amaya, Ella Baron, Marvin Paredes, Sayuri Tzul

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Christensonella uncata Christensonella uncata Cohniella ascendens Cohniella ascendens Cycnoches ventricosum (Orchidaceae; EB) (Orchidaceae; EB) (Orchidaceae; BH) (Orchidaceae; DA) (Orchidaceae; EB)

Cycnoches ventricosum Epidendrum cardiophorum Epidendrum cardiophorum Epidendrum isomerum Epidendrum isomerum (Orchidaceae; EB) (Orchidaceae; EB) (Orchidaceae; WC) (Orchidaceae; EB) (Orchidaceae; MP)

Epidendrum martinezii Epidendrum martinezii Gongora cassidea Gongora cassidea Guarianthe bowringiana (Orchidaceae; EB) (Orchidaceae; WC) (Orchidaceae; BH) (Orchidaceae; BH) (Orchidaceae; BH)

Guarianthe bowringiana Isochilus carnosiflorus Isochilus carnosiflorus Masdevallia floribunda Masdevallia floribunda (Orchidaceae; BH) (Orchdiaceae; EB) (Orchdiaceae; EB) (Orchidaceae; EB) (Orchidaceae; EB) Epiphytes of Las Cuevas Research Station, Cayo District, BELIZE 5 Bruce K. Holst, Sally Chambers, Antone Jones, David Amaya, Ella Baron, Marvin Paredes, Sayuri Tzul

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Mormolyca hedwigiae Mormolyca hedwigiae Mormolyca ringens Mormolyca ringens Muscarella fimbriata (Orchidaceae; EB) (Orchidaceae; WC) (Orchidaceae; BH) (Orchidaceae; EB) (Orchidaceae; EB)

Muscarella fimbriata Nemaconia striata Nemaconia striata Nidema boothii Nidema boothii (Orchidaceae; EB) (Orchidaceae; BH) (Orchidaceae; EB) (Orchidaceae; EB) (Orchidaceae; EB) Note white spider

Notylia barkeri Notylia barkeri Oncidium sphacelatum Oncidium sphacelatum Ornithocephalus inflexus (Orchidaceae; EB) (Orchidaceae; BH) (Orchidaceae; EK) (Orchidaceae; EB) (Orchidaceae; EB)

Ornithocephalus inflexus Pleurothallis sp. Prosthechea cochleata Prosthechea cochleata Prosthechea cochleata (Orchidaceae; EB) (Orchidaceae; BH) National Flower of Belize National Flower of Belize National Flower of Belize (Orchidaceae; EB) (Orchidaceae; EB) (Orchidaceae; EB) Epiphytes of Las Cuevas Research Station, Cayo District, BELIZE 6 Bruce K. Holst, Sally Chambers, Antone Jones, David Amaya, Ella Baron, Marvin Paredes, Sayuri Tzul [fieldguides.fieldmuseum.org] [1149] version 1 5/2019

Rhetinantha aciantha Rhetinantha aciantha Specklinia brighamii Specklinia brighamii Specklinia tribuloides (Orchidaceae; DA) (Orchidaceae; EB) (Orchidceae; BH) (Orchidceae; WC) (Orchidaceae; EB)

Specklinia tribuloides Stelis sp. Stanhopea sp. Trichocentrum sp. Trigonidium egertonianum (Orchidaceae; EB) (Orchidaceae; AJ) (Orchidaceae; AJ) (Orchidaceae; AJ) (Orchidaceae; EB)

Trigonidium egertonianum Peperomia alata Peperomia alata Peperomia deppeana Peperomia deppeana (Orchidaceae; EB) (Piperaceae; EB) (Piperaceae; EB) (Piperaceae; BH) (Piperaceae; BH)

Peperomia obtusifolia Peperomia obtusifolia Peperomia rhombea Peperomia tetraphylla Peperomia tetraphylla (Piperaceae; BH) (Piperaceae; EB) (Piperaceae; AJ) (Piperaceae; BH) (Piperaceae; AJ) Epiphytes of Las Cuevas Research Station, Cayo District, BELIZE 7 Bruce K. Holst, Sally Chambers, Antone Jones, David Amaya, Ella Baron, Marvin Paredes, Sayuri Tzul

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Spore-bearing Plants

Coussapoa oligocephala “strangler” Coussapoa oligocephala “strangler” Asplenium auritum Asplenium auritum (Urticaceae; BH) (Urticaceae; BH) (Aspleniaceae; SC) (Aspleniaceae; SC)

Asplenium cristatum Asplenium cristatum Asplenium formosum Asplenium formosum Asplenium serratum (Aspleniaceae; EB)) (Aspleniaceae; EB)) (Aspleniaceae; DA) (Aspleniaceae; EB) (Aspleniaceae; base; BH)

Asplenium serratum Campyloneurum brevifolium Campyloneurum brevifolium Didymoglossum krausii Didymoglossum krausii (Aspleniaceae; base; BH) (Polypodiaceae; PN; Nicaragua) (Polypodiaceae; PN; Nicaragua) (Hymenophyllaceae; EB) (Hymenophyllaceae; EB)

Phlegmariurus linifolius Phlegmariurus linifolius Microgramma lycopodioides Microgramma lycopodioides Microgramma percussa (Lycopodiaceae; BH) (Lycopodiaceae; EB) (Polypodiaceae; slopes; BH) (Polypodiaceae; slopes; EB) (Polypodiaceae; BH) Sterile leaves. Fertile leaves Sterile leaves Epiphytes of Las Cuevas Research Station, Cayo District, BELIZE 8 Bruce K. Holst, Sally Chambers, Antone Jones, David Amaya, Ella Baron, Marvin Paredes, Sayuri Tzul

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Microgramma percussa Niphidium crassifolium Niphidium crassifolium Pecluma dispersa Pecluma dispersa (Polypodiaceae; BH) (Polypodiaceae; summit; BH (Polypodiaceae; summit; BH) (Polypodiaceae; WC) (Polypodiaceae; WC) Fertile leaves

Pecluma plumula Pecluma plumula Pleopeltis marginata Pleopeltis marginata Polypodium fallax (Polypodiaceae; SC) (Polypodiaceae; BH) (Polypodiaceae; BH) (Polypodiaceae; BH) (Polypodiaceae; SC)

Polypodium fallax Polypodium lindenianum Polypodium lindenianum (Polypodiaceae; AJ) (Polypodiaceae; SC) (Polypodiaceae; SC)

Las Cuevas Research Station (LCRS) is located at the northern portion of Belize's Maya Mountains and in the heart of Chi- quibul Forest Reserve. Since 1994 LCRS has pro- moted biological, botanical, wildlife, climatic, and archaeological research and education. This unique setting provides a range of opportunities for researchers, naturalists, environmentalists, and students to enjoy nature and conduct biodiversity research. LCRS is co-managed by Friends for Conservation and Development and the Government of Belize.

The Chiquibul Forest represents nearly 8% of the Belize's terrestrial surface and is part of the tri- national Maya Forest bioregion forming the largest remaining contiguous block of tropical forest north of the Amazon.

The following epiphytic species are reported from Las Cuevas, but photos are not available: Campyloneurum fasciale, Lycaste Serpocaulon dissimile Serpocaulon dissimile sp., Nephrolepis pendula, Polypodium hispidulum, Rhetinantha sp., Scaphyglottis sp. (Polypodiaceae; SC) (Polypodiaceae; AJ) The authors appreciate the help of the Las Cuevas Research Station staff and the Friends of Conservation and Development, Belize, for logistical support.