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Taxonomie Goniatitové Fauny, Biostratigrafie a Paleoekologie Jesenického a Drahanského Kulmu

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Taxonomie Goniatitové Fauny, Biostratigrafie a Paleoekologie Jesenického a Drahanského Kulmu MASARYKOVA UNIVERZITA Přírodovědecká fakulta Tomáš LEHOTSKÝ TAXONOMIE GONIATITOVÉ FAUNY, BIOSTRATIGRAFIE A PALEOEKOLOGIE JESENICKÉHO A DRAHANSKÉHO KULMU Disertační práce Školitel: prof. RNDr. Jan Zapletal, CSc. Brno, 2008 Bibliografická identifikace Jméno a příjmení autora: Tomáš Lehotský Název disertační práce: Taxonomie goniatitové fauny, biostratigrafie a paleoekologie drahanského a jesenického kulmu Název disertační práce anglicky: Taxonomy of the Goniatite Fauna, Biostratigraphy and Palaeoecology of the Drahany and Nízký Jeseník Culm Studijní program: 1201V Geologie, doktorský studijní program Studijní obor (směr), kombinace oborů: 1201V003 Geologické vědy se zaměřeními Školitel: prof. RNDr. Jan Zapletal, CSc. Rok obhajoby: 2008 Klíčová slova v češtině: Taxonomie, goniatitová fauna, biostratigrafie, paleoekologie, spodní karbon, kulmská facie, Nízký Jeseník, Drahanská vrchovina Klíčová slova v angličtině: Taxonomy, Goniatite Fauna, Biostratigraphy, Palaeoecology, Lower Carboniferous, Culm Facies, Nízký Jeseník Mts., Drahany Upland 2 © Tomáš Lehotský, Masarykova univerzita, 2008 3 Na tomto místě bych chtěl poděkovat všem, kteří byli nápomocni sepsání této disertační práce. Především děkuji školiteli prof. RNDr. Janu Zapletalovi CSc. za cenné rady a připomínky a dále všem pracovníkům muzejních institucí a soukromým sběratelům nejen v České republice za ochotu, se kterou zpřístupnili své sbírkové fondy. 4 Prohlašuji, že jsem disertační práci vypracoval sám a že jsem uvedl veškeré zdroje literatury. V Brně 20. 8. 2008 ...................................................... 5 Abstrakt: Taxonomie goniatitové fauny, biostratigrafie a paleoekologie drahanského a jesenického kulmu Kulmské sedimenty na Moravě a ve Slezsku jsou vyvinuty v podélné struktuře jz. – sv. směru na východním okraji Českého masivu. V zásadě se rozlišují dvě hlavní sedimentační pánve: drahanská a nízkojesenická. Sedimenty Drahanské vrchoviny se litostratigraficky dělí na tři souvrství – protivanovské (spodní – střední visé), rozstáňské (svrchní visé, zóny Peδ – Goα) a myslejovické (svrchní visé, zóny Goα – Goγ). Nízký Jeseník je členěn do čtyř souvrství: andělskohorské (spodní až střední visé), hornobenešovské (střední – svrchní visé), moravické (svrchní visé, subzóny Goα2–3 - Goβmu) a hradecko-kyjovické (svrchní visé až namur, subzóny Goβspi až po zónu E1). V disertační práci je provedena revize goniatitové fauny. V drahanském a jesenickém kulmu bylo zjištěno a popsáno celkem 16 rodů goniatitů sdružených do osmi čeledí: Eoglyphioceras, Sudeticeras, Nomismoceras, Girtyoceras, Calygirtyoceras, Sulcogirtyoceras, Edmooroceras, Paradimorphoceras, Goniatites, Arnsbergites, Hibernicoceras, Paraglyphioceras, Lusitanoceras, Emstites, Neoglyphioceras a Lusitanites. Na základě studia společenstev fauny a ichnofauny jsou diskutovány biostratigrafické a paleoekologické závěry. 6 Dissertation Abstract: Taxonomy of the Goniatite Fauna, Biostratigraphy and Palaeoecology of the Drahany and Nízký Jeseník Culm The Culm sediments of the Moravian-Silesian region are preserved in an elongated, SW-NE trending structure on the eastern margin of the Bohemian Massif. Two Culm basins are generally recognized in the Moravian-Silesian Zone: the Drahany Basin and the Nízký Jeseník Basin. The fill of the Drahany Basin has been lithostratigraphically subdivided into three formations: the Protivanov Formation of early to late Viséan age (Peγ to Peδ Zone), the Rozstání Formation of late Viséan age (Peδ to Goα Zone), and the Myslejovice Formation of late Viséan age (Goα to Goγ Zone), each consisting of several members. The lithostratigraphic succession of the Nízký Jeseník Basin has been subdivided into four formations: the Andělská Hora Formation of early Viséan to middle Viséan age, the Horní Benešov Formation of middle to late Viséan age, the Moravice Formation of late Viséan age (Goα2–3 to Goβmu Subzone), and the Hradec-Kyjovice Formation ranging from the late Viséan to the lowermost part of the Namurian (Goβspi Subzone to E1 Zone). The Goniatite Fauna of the Lower Carboniferous strata of the Nízký Jeseník Mts. and Drahany Upland is revisited in the present Dissertation Thesis. The sixteen genera of goniatites: Eoglyphioceras, Sudeticeras, Nomismoceras, Girtyoceras, Calygirtyoceras, Sulcogirtyoceras, Edmooroceras, Paradimorphoceras, Goniatites, Arnsbergites, Hibernicoceras, Paraglyphioceras, Lusitanoceras, Emstites, Neoglyphioceras a Lusitanites in eight families were described in detail. On the basis of fauna/ichnofauna assemblages study are biostratigraphical and palaeoecological consequences discussed. 7 Obsah: ÚVOD.......................................................................................................................................................................11 VYMEZENÍ ZKOUMANÉHO ÚZEMÍ..................................................................................................................12 PŘEHLED DOSAVADNÍCH VÝZKUMŮ GONIATITOVÉ FAUNY V KULMU NÍZKÉHO JESENÍKU........17 PŘEHLED DOSAVADNÍCH VÝZKUMŮ GONIATITOVÉ FAUNY V KULMU DRAHANSKÉ VRCHOVINY ..................................................................................................................................................................................24 STAV ZACHOVÁNÍ FAUNY.................................................................................................................................26 POZNÁMKY K MORFOLOGII A TERMINOLOGII GONIATITŮ...................................................................27 BIOSTRATIGRAFIE SPODNÍHO KARBONU NA ZÁKLADĚ GONIATITOVÉ FAUNY................................ 28 METODIKA ............................................................................................................................................................ 32 SYSTEMATICKÁ ČÁST........................................................................................................................................34 ŘÁD: GONIATITIDA HYATT, 1884 ......................................................................................................................... 35 Podřád: Goniatitina HYATT, 1884....................................................................................................................35 Nadčeleď: Pericycloidea HYATT, 1900.........................................................................................................................35 Čeleď: Muensteroceratidae LIBROVICH, 1957 ........................................................................................................35 Rod: Eoglyphioceras BRÜNING 1923.................................................................................................................35 Eoglyphioceras truncatum (PHILLIPS 1836).................................................................................................35 Eoglyphioceras sp..........................................................................................................................................36 Čeleď: Anthracoceratidae PLUMMER & SCOTT, 1937............................................................................................37 Rod: Sudeticeras PATTEISKY, 1930...................................................................................................................37 Sudeticeras crenistriatum (BISAT 1928)........................................................................................................37 Sudeticeras cf. crenistriatum (BISAT 1928)...................................................................................................39 Sudeticeras subtile MOORE 1950..................................................................................................................40 Sudeticeras cf. subtile MOORE 1950 .............................................................................................................41 Sudeticeras wilczeki - hoeferi PATTEISKY 1929...........................................................................................41 Sudeticeras cf. wilczeki-hoeferi PATTEISKY 1929........................................................................................44 Sudeticeras turneri MOORE&HODSON 1958...............................................................................................44 Sudeticeras cf. turneri MOORE&HODSON 1958 ..........................................................................................45 Sudeticeras stolbergi PATTEISKY 1929 .......................................................................................................45 Sudeticeras cf. newtonense MOORE 1950 .....................................................................................................47 Sudeticeras ostraviense PATTEISKY 1929 ...................................................................................................48 Nadčeleď: Nomismoceratoidea LIBROVICH, 1957 .......................................................................................................50 Čeleď: Nomismoceratidae LIBROVICH, 1957..........................................................................................................50 Rod: Nomismoceras HYATT, 1884 .....................................................................................................................50 Nomismoceras vittiger (PHILLIPS, 1836).......................................................................................................50 Nomismoceras sp...........................................................................................................................................52
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