Tineidae I Microlepidoptera of Europe Edited by M. Nuß (Senckenberg Museum of Zoology, Dresden) O. Karsholt (Zoologisk Museum, Copenhagen) P. Huemer (Tiroler Landesmuseen Betriebsgesellschaft m.b.H., Innsbruck) volume 7 The titles published in this series are listed at brill.com/mle Tineidae I (Dryadaulinae, Hapsiferinae, Euplocaminae, Scardiinae, Nemapogoninae and Meessiinae) By Reinhard Gaedike LEIDEN | BOSTON Cover illustration: Euplocamus anthracinalis (Scopoli, 1763), Germany, Buch, 8.iv.2007, leg. et photograph P. Lichtmannecker. This publication has been typeset in the multilingual ‘Brill’ typeface. With over 5,100 characters covering Latin, IPA, Greek, and Cyrillic, this typeface is especially suitable for use in the humanities. For more information, please see www.brill.com/brill-typeface. ISSN 1395-9506 ISBN 978-90-04-25641-5 (hardback) ISBN 978-90-04-28916-1 (e-book) Copyright 2015 by Koninklijke Brill NV, Leiden, The Netherlands. 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Dedicated to my wife Hannelore and in memory of Günther Petersen (1924 – 2012) ∵ Contents Acknowledgements IX Abstract xi Introduction XII Material and Methods xii The Body Structure of the Tineoidea xii Systematics of the Tineoidea and their Family Groups xvii Collection and Preparation Techniques xxiii Abbreviations xxvi 1 Key to the European subfamilies of Tineidae 1 2 Check-list of European and Macaronesian Dryadaulinae, Hapsiferinae, Euplocaminae, Scardiinae, Nemapogoninae and Meessiinae 3 3 Systematic Treatment of the Genera and Species of European Tineidae 10 Dryadaulinae 10 Hapsiferinae 18 Euplocaminae 23 Scardiinae 25 Nemapogoninae 32 Meessiinae 78 4 Distribution Catalogue 165 5 Colour Plates 176 6 Drawings, Male Genitalia 191 7 Drawings, Female Genitalia 237 References 277 Index to Entomological Genus Names 301 Index to Entomological Species-Group Names 303 Index to Plant and Other Hosts Names 307 Acknowledgements First of all it is my desire to thank the late Austria); Christian Gibeaux (Avon, France); Günther Petersen (1924–2012), who was Stanislav Gomboc (Kranj, Slovenija); Laszlo my mentor, colleague and friend during Gozmány (†) (HNHM); K. Gregersen (Soro, my working time at the Deutsches Ento- Denmark) Theo Grünewald (Landshut, mologisches Institut (DEI), now part of Germany); Richard Heindel (Günzburg, the Senckenberg Gesellschaft für Natur- Germany); Marcel Hellers (Bissen, forschung, and therefore abbreviated as Luxembourg); Peter Huemer (TLMF); SDEI. He initiated my interest in tineids, Tomasz Jaworski (Raszyn, Poland); Jan Å. supported my studies at all times and pro- Jonasson (Gothenburg, Sweden); Jari vided generous access to his unpublished Junnilainen (Vantaa, Finland); Lauri Kaila results on tineid taxonomy, distribution (FMNH); Tim Karisch (MNVD); Ole and biology, compiled on card files. Karsholt (ZMUC); Friedrich Kasy (†) I would like to express my special thanks (NMW); Rudolf Keller (Sulzemoos, to my colleagues at the DEI / SDEI. Holger Germany); András Kun (HNHM); Bernard Dathe, formerly director of the institute, Landry (Genève, Switzerland); Mojmir kindly agreed on a long-time loan of the Lasan (Ljubljana, Slovenija); Martin Lödl tineid collection after my retirement, (NMW); Wolfram Mey (ZMHB); Joel which served as base for the continuation Minet (MNHN); Timo Muus (Steenwijk, of my scientific work. The librarians Holger The Netherlands); Erik van Nieukerken Framke, Ute Kaczinski, Gabriele Mirschel (RMNH); Jacques Nel (La Ciotat, France); and Renate Riedelsheimer supplied me Eivind Palm (Højer, Denmark); Pietro with copies of scientific literature when- Passerin d’Entrèves (Torino, Italy); Lukasz ever needed. Last but not least, I thank Przybylowicz (ISZP); Emili Requena Christian Kutzscher for taking some colour (Igualada, Spain); Gaden S. Robinson (†) images of several taxa. (BMNH); Hartmut Roweck (Kiel, Ger many); Without the help of many people it Willy Sauter (Illnau, Switzerland); Nikolajs would have been impossible to complete Savenkovs (Kiel, Germany); Willibald the present volume. I am grateful to all who Schmitz (Bergisch-Gladbach, Germany); supported me by the loan of material under Andreas Segerer (ZSM); Rudi Seliger their care or from their own collections, as (Schwalmtal, Germany); Sergej Sinjev well as by giving practical advice and help- (ZIN); Thomas Sobszyk, Bautzen, ful comments. These are Helen Alipanah Germany); Andreas Stübner (Peitz, (IRIPP); Ernst Arenberger (NMW); Joaquin Germany); Dieter Stüning (ZFMK); Jan Baixeras (Burjassot, Valencia, Spain); Šumpich (Česka Bela, Czech Republic); Giorgio Baldizzone (Asti, Italy); Ian Barton Reinhard Sutter (Bitterfeld, Germany); (Stretham, United Kingdom); Bengt Å. Franz Theimer (Berlin, Germany); Zdeno Bengtsson (Färjestaden, Sweden); Erich Tokar (Sala, Slovakia); Paolo Triberti Bettag (Dudenhofen, Germany); Don R. (Verona, Italy); Robert Trusch (SMNK); Davis (USNM); Helmut Deutsch (Lienz, Kevin Tuck (BMNH); Harry van der Wolf x acknowledgements (Nuenen, Belgium); Thierry Varenne (Nice, cover photo. The courtesy of Kevin Tuck France); Andreas Werno (Nunkirchen, (BMNH) enabled me to use some copy- Germany); Wolfgang Wittland (Wegberg- right images made with the permission Dalheim, Germany). of the Trustees of the Natural History My colleague, the late Gaden Robinson Museum, London. (1949–2009) at the BMNH, was in so many I thank Bernhard Oertel (Bonn, cases my “court of last resort” on taxonomic Germany) for his help with the authorship questions. Our correspondence and face- of the fungus names. to-face encounter at the BMNH will never I would like to express my thank to be forgotten. the editors (Peter Huemer (Innsbruck, Special thanks to Igor Kostjuk (Kiev, Austria), Ole Karsholt (Copenhagen, Ukraine) for making the colour pictures of Denmark) and Matthias Nuss (Dresden, most of the adults and for the compilation Germany) for their intensive support of all images onto plates. Some images during preparation of the manuscript and were made by Giorgio Baldizzone (Asti, especially the invaluable discussions with Italy), Christian Kutzscher from SDEI Matthias Nuss (MTD). My special thank (Müncheberg, Germany), Tomasz Jaworski go to Martin Corley (Faringdon, U.K.) for (Roszyn, Poland). A special thank to Peter the linguistic revision and many useful Lichtmannecker (Adlkofen, Germany) for comments on the manuscript. My col- the permission to use some of his colour leagues Sergey Sinev and Zdenek Laštůvka images for the text figures and for the eventually reviewed the manuscript. Abstract The first volume of Tineidae treats the N. similella Gaedike, 2007, syn. n. is syn- subfamilies Dryadaulinae, Hapsiferinae, onymised with N. sardicus Gaedike, 1983, Euplocaminae, Scardiinae, Nemapogoninae N. teberdellus Zagulajev, 1963, syn. n. and its and Meessiinae from Europe. A total of 180 misspelling N. teberdensis Zagulajev, 1964 species are diagnosed and figures of the as well as N. georgiellus Zagulajev, 1963, moths, male and female genitalia are given. syn. n. and its misspelling N. georgicus Information is added on the life history and Zagulajev, 1964 are synonymised with N. distribution of the species. The distribution lagodechiellus Zagulajev, 1962. Lichenotinea data are summarised in a table showing the maculata Petersen, 1957, syn. n. is syn- records for each European country. onymised with L. pustulatella (Zeller, 1852). The following nomenclatural innova- Novotinea ochripennella Nel & Varenne, 2011 tions are introduced: Nemapogon nevellus syn. n. is synonymised with Xystrologa gren- Zagulajev, 1963, syn. n. is synonymised with adella (Walsingham, 1897). Infurcitinea N. cloacella (Haworth, 1828), N. wolffiella nedae Gaedike, 1983, comb. n. is transferred Karsholt & Nielsen, 1976, syn. n. is syn- to Dryadaula. onymised with N. koenigi Capuşe, 1967, Introduction Material and Methods one square centimetre divided into four hundred squares of half a square millime- Sources for the studied material were tre. The magnification used is adapted to mainly European collections of museums the size of the genitalia. The most com- and private entomologists (see list of mon magnification is 160x (16x ocular and abbreviations and acknowledgments). 10x objective). For larger objects a magnifi- Beside literature cited, an important source cation of 100.8x (16x ocular and 6.3x objec- of information are the card files on the tax- tive) was used and for a few very large onomy, faunistics and biology of tineids, objects a magnification of 51.2x (16x ocular compiled by my predecessor Günther and 3.2x objective). An analogous net with Petersen, and continued by myself. These squares of 7.5 mm was drawn on paper to card files are now deposited at the SDEI have a scale for the drawing. (Senckenberg Deutsches Entomologisches Institut, Müncheberg/Germany). Since the late 1990s, bibliographical and faunistic The Body Structure information