KALMIOPSIS Journal of the Native Plant Society of Oregon Castilleja chambersii Egger and Meinke A, habit; B, flowering stem to show leaf and size; C, bracts; D, calyx with its corolla; E, variability in calyces; F, seed with closeup of wall. ISSN 1055-419X Volume 8, 2001 EDITORIAL KALMIOPSIS Secrets can be difficult to keep, especially for as long as two Journal of the Native Plant Society of Oregon, ©2001 years, but we hope that this issue of Kalmiopsis surprises the individual that we as NPSO members are all proud to EDITORS claim as one of our own, Ken Chambers. From front cover Cindy Roché, PhD through the appendices, every page is dedicated to his honor. Linda Ann Vorobik, PhD Conceived at the time of Ken’s seventieth birthday, this Festschrift is the work of many individuals, whom Aaron EDITORIAL BOARD Liston acknowledges in his introduction. However, Ken’s stature as a botanist and as a person is no secret, as noted on Stuart Garrett, MD the Certificate of Merit presented to him by the Botanical Susan Kephart, PhD Society of America in August 1990: Rhoda M. Love, PhD “Eminent biosystematist, internationally recognized for his studies of various genera of Asteraceae; a pioneer in the NPSO Web PAG E development of plant conservation in Oregon; and stimu- http://www.NPSOregon.org lating teacher who has inspired many students to become botanists.” MEMBERSHIP Membership in the Native Plant Society of Oregon is open to all. Membership applications, renewals and changes of address (include old address and zip code) should be sent to the NPSO Membership Chair, Jan Dobak, 2584 NW Savier St., Portland, OR 97210-2412 Student $12; Regular $18; Family $24; Sustaining $50; Patron $100; Life Member $500 Subscription Only (Bulletin and Kalmiopsis) $18 CONTACT INFORMATION KALMIOPSIS EDITORS: Cindy Roché, 109 Meadow View Dr., Medford, OR 97504, [email protected] & Linda Ann Vorobik, P.O. Box 866, Lopez Island, WA 98261, [email protected] BULLETIN EDITOR: Tanya Harvey, 40237 Reuben Leigh Rd., Lowell, OR 97452-9712 [email protected] MEMBERSHIP CHAIR: Jan Dobak, 2584 NW Savier St., Portland, OR 97210-2412, (503) 248-9242 WEBMASTER: Michael McKeag, 7461 SW 184th Pl., Aloha, OR 97007-5740, [email protected] EMAIL DISCUSSION LIST LOGIN: See NPSO Web Page, select subcribe to list under membership. WEB page: http://www.NPSOregon.org COVER ILLUSTRATION Castilleja chambersii Egger and Meinke, as published in “Castilleja chambersii (Scrophulariaceae), a new rare species from the northern Coast Range of Oregon” (Egger, M. and R. J. Meinke. 1999. Brittonia 51(4):445-451). An Indian Paintbrush en- demic to the Coast Range of southwestern Clatsop County, Oregon, first recognized as distinct by Ken Chambers, and named for him in 1999 in honor of his 70th birthday. Kalmiopsis: ISSN 1055-419X. Volume 8, 2001. Published annually. Subscription $18 per year. Native Plant Society of Oregon General business address: PO Box 902, Eugene OR 97440 NATIVE PLANT SOCIETY OF OREGON STATE OFFICERS CHAPTER PRESIDENTS DIRECTORS (TO 6/02) .................Karl Anderson, Dan Luoma, BLUE MOUNTAIN (PEN D LETON ) ..............................Jerry Baker ..........................................................................Danna Lytjen CHEAHMILL .......................................................Dave Hanson DIRECTORS (TO 6/03) .................... Jim Duncan, Rhoda Love, COR V ALLIS ...................................................... Esther McEvoy ..................................................................... Kareen Sturgeon EMERAL D (EUGENE ) .............................................Nick Otting PRESI D ENT ....................................................Bruce Newhouse HIGH DESERT (BEN D ) ............................................Stu Garrett IMME D IATE PAST PRESI D ENT ................................. Michael Igo KLAMATH BASIN ....................................................David Lebo VICE PRESI D ENT ...........................................Michael McKeag MI D -COLUMBIA .........................................................Jerry Igo SECRETARY ..................................................Kelli Van Norman NORTH COAST .............................................. Vivian Starbuck TREASURER ............................................................. Eric Wold PORTLAN D .............................................................Dee White SIS K IYOU .......................................................... Molly Sullivan STATE COMMITTEE CHAIRS UMPQUA VALLEY (ROSEBURG ) ........................... Jack Hausotter ED UCATION ...............................................................Jerry Igo WILLAMETTE VALLEY (SALEM ) ........................... Karl Anderson CONSER V ATION , EAST SI D E .....................................Susan Geer WMCUSIC K (LAGRAN D E ) ..................................Frazier Nichol CONSER V ATION , WEST SI D E ............................ Steven L. Jessup LEGISLATI V E ................................................................. vacant PUBLICATIONS MEMBERSHIP ......................................................... Jan Dobak BULLETIN ED ITOR .............................................. Tanya Harvey BU D GETS AN D GRANTS ......................................... Dan Luoma KALMOPSIS ED ITORS ............ Cindy Roché, Linda Ann Vorobik NOTICE TO CONTRIBUTORS Members of the Native Plant Society of Oregon and others Contributions without accompanying manuscripts, such are invited to submit articles, book reviews, artwork, poetry, car- as art, cartoons, or photos, also are welcomed. toons, and photographs for publication in KALMIOPSIS. All Send all submissions to KALMIOPSIS EDITORS, or materials submitted should pertain to Oregon’s vegetation and e-mail KALMIOPSIS EDITORS to request an electronic sub- flora. Acceptance will be based on suitability (articles dealing mission. (Specific instructions for submission of photographs, with formal nomenclatural proposals or of a highly technical figures, and tables will be sent to contributor upon request or nature are not acceptable). after article has been accepted for publication.) Contributors of articles should submit one hard copy of a Please feel free to contact KALMIOPSIS EDITORS for double-spaced manuscript accompanied by an electronic copy further information, or to inquire if an article is suitable for of the file (PC or MAC; Word, Word-Perfect, or aT ext or Rich publication in this journal. Text Format File; indicate name of Program and Version in the accompanying cover letter). Computer facilities are available for use at libraries and printing or word-processing businesses. ACKNOWLEDGEMENTS Kalmiopsis Logo: Linda Ann Vorobik (VorobikBotanicalArt.com) Pagesetting: Cindy Roché and Linda Ann Vorobik Art work: © Linda Ann Vorobik Printing: Mustard Press, (MustardPress.com) TABLE OF CONTENTS A Festschrift Honoring Kenton L. Chambers ....................................................................................... 1 By Aaron Liston Ken Chambers, Oregon Botanist .......................................................................................................... 2 By Henrietta Laing Chambers Ken Chambers: Taxonomic Rigor and Rare Plant Protection in Oregon ............................................. 10 By Rhoda M. Love Flora of Walker Flat, Yamhill County, Oregon.................................................................................... 17 By Richard R. Halse and Judith B. Glad The Eunanus Monkey Flowers of Genus Mimulus (Scrophulariaceae) in Oregon ................................ 25 By Wayland L. Ezell Non-morphological Evidence in Biosystematics: Kenton Chambers and the Annual Species of Microseris ................................................................................................. 30 By Konrad Bachmann Appendix 1: Graduate students directed by Kenton L. Chambers at OSU, and their thesis titles ........................................................................................................................... 39 Appendix 2: Publications by Kenton L. Chambers ............................................................................. 40 PURPOSE ~ Native Plant Society of Oregon ~ Dedicated to the enjoyment, conservation, and study of Oregon’s native vegetation. Kalmiopsis NONPROFIT ORG Native Plant Society of Oregon US POSTAGE Jan Dobak, Membership Chair PAID 2584 NW Savier Street MEDFORD OR Portland, OR 97210-2412 Permit No.113 A Festschrift Honoring Kenton L. Chambers Aaron Liston Herbarium Director, Oregon State University, Corvallis, OR 97331 estschrift is the German word for “celebratory writing,” a them properly ... has been the decisive ability of great taxonomists. unique publication devoted to the research and influences Kenton Chambers has this gift to perfection.” Microseris was the Fof an exceptional scholar. In this issue of Kalmiopsis, we cel- subject of Ken’s PhD thesis, and its taxonomy and evolution remain ebrate the career and accomplishments of Kenton Lee Chambers one of his central interests. Together, these five articles celebrate the on the occasion of his 70th birthday on September 27, 1999. Ken breadth and influence of Ken’s botanical career. has made important contributions to plant systematics through his I wish to acknowledge the role of Nancy Fredricks in the initial monographic and biosystematic studies of Microseris (Asteraceae) development and organization of this festschrift. Nancy conceived and related genera. He published the new genus, Stebbinsoseris,