ISSN 0031-0204 Transactions and Proceedings of the Palaeontological Society of Japan New Series No. 103 ./ Palaeontological Society of Japan·, Oct. 15, 1976 Editor Takashi HAMADA Associate Editor Iku wo OBAT A Officers for 1975 - 1976 Honorary President: Teiichi KOBAYASH I President: Tatsuro MATSUMOTO Councillors (*Executive): *Kazuo ASAMA, Kiyoshi ASANO, *Kiyotaka CHINZEI, *Takashi HAMADA, *Tetsuro HANA I, *Itaru HAYAMI, Tadao KAMEl, *Kametoshi KANMERA, *Tamio KOTAKA, *Tatsuro MATSUMOTO, Tokio SHIKAMA, Tsugio SHUTO, *yokichi T AKA YANA GI, Toshimasa T ANAl, *Hiroshi UJIIE Executive Committee: General Affairs: Tetsuro HANAI, Itaru HAY AM I, Kiyotaka CHINZEI, Saburo KAN NO Membership: Kazuo ASAMA, Kazuhiko UEMURA Finance: Hiroshi UJIIE Planning: Tamio KOTAKA, Jun'ichi TAZAWA Publications Transactions: Takashi HAMADA, Ikuwo OBATA Special Papers: Kametoshi KANMERA, Ienori FUJIYAMA; Tomowo OZAWA, Juichi YANAGIDA " Fossils" : Yokichi TAKA YANAGI, Kunihiro ISI-IIZAKI Fossil on the cover is the six leaves in a whorl of Trizygia oblongifolia (GERM. & KAULF.) ASAMA from the Maiya Formation (Parafusulina zone), Maiya, N. E. Japan. All communications relating to this journal should be addressed to the P ALAEONTOLOGICAL SOCIETY OF JAPAN c/ o Business Center for Academic Societies, Japan Yayoi 2-4-16, Bunkyo-ku, Tokyo 113, Japan Sole agent: University of Tokyo Press, Hongo, Tokyo Trans. Proc. Palaeont. Soc. Japan, N. S., No. 103, pp. 343-378, pIs. 36-39, Oct. 1:;, 1976 663. MESOZOIC PLANTS FROM THE AKAIW A FORMATION (UPPER NEOCOMIAN), THE ITOSHIRO GROUP, CENTRAL HONSHU, JAPAN* TATSUAKI KIMURA Tokyo Gakugei University, Koganei, Tokyo 184 and SHIN]I SEKIDO Science Education Centre, Ishikawa Prefecture, Kanazawa 920 **::z ~ 7 :/ J:fif,iI'r-!6~ C;fiii{n3~llf.J:fif,) O)flHf0fl:;fi: ;Jf;!5~O)fjM0fl:;fill-tn:.i*fj: /J;!e <, [,'t*a:;;t:;j;;y (1975) O)liif5eHh'<~,·(lH·0)t1;jlJlfu"f:R'Jliif1l:11ITt.l:bn -n,t.l:~'o q-Ji¥l:, ;fi)IIiJf<;fiJ1lIi~SiIJ·f.;j:'t7JiJ~Mn CS0Jq:.)Jrl) :to J: tr*t~1:i1'.J!Jli:/J' i?~~i*ftO) J: ~ ,tlt'!S:/J~~:!h:. %J.i?nt.:O)"'C;: ;: I;:.**,-t 7.:>0 ih~!6~O)fll'i¥!.1fl:;fillri:i'i:O)~ I::J !WO)fll'i¥ollf. t mRlt/J;~t.l: 7.:> 0)'(::, u.r:il\~!5;j1li¥!.1llft J: tr, ~ ~1;:.:;;t:MI;:' J: -:> -C)ic!li:~nt.:m*iJf<*!j!fll15%o;iRM=F!Jli::tt!!~O)FEa1:i C1fBJ1:i) i'Illlrr,q:.ifil)~O)fll'i¥!.1fl:;fi~~6i')7.:>o :iI\~!5fll'i¥!.11tY:11P'J*j1li¥!.1:l1l!~1R C:;;t:H, 1961; 1975) 0) fll'i!['mlr(:""(', t.:/J'bi?trf4, IjGCiSf4:toJ:trJi!TJ,9',/f'IV]O)';"-,;(, 'JT"~~, 17-3t/~:t~J: tr-t.-O)~~O)to), :toJ:tr:E'J(*~ C$~O)to)/J;f!I~) /J'Gt.l:7.:>M¥ollf.""(';57.:>o iI'r-!61iilfuW,u, t.:/J'b G trf4~ll C6i'), ,;,.- ?1"~/J~ $~""(';5 7.:> ;: t, 'J T -;; ~~/J~$~/J'''?/N~HI:-t 7.:> ;: t, it.:~~/J~}J.iiiii:i\~~-t7.:>to)/J;f!I~I;:.t.l:7.:>;:t, :toJ:tr, Ginkgoidium H~~, 17-~ t/ ~O)~/J~·H~IHI:-t 7.:>;: t t.l: t· O)ilf.B"'C~ p fll'i¥!.1llf.t ~d.l: 7.:>0 i t.: J:i:i'i: 0) FE Oi;1:iM¥oM t UIIRlt /J~~t.l: 7.:> 0 ;JJ~!5fll'i¥ollf.~tf.JRlt-t 7.:>~fiO):;i(fif,7t, -tt.l:bi?, Coniopteris, Birisia, Asplenium, Raphaelia, Adiantites, Dictyozamites cfr. cordatus, Nilssonia lobatidentata, Gillkgoites, Leptostrobus, Pseudotorellia, Podozamites t.l: nl, [i'lJ1i!ff(;0),;,.-"":: ~ 7fll'i¥o:tt!!~1R1ill¥olrr. t ;tt<ij t L <11 ~b6i')-CJ!I~O) t 0)""(';5 f), [i'lJ1i!ff(;0) rffil¥ofif,J1I~I1Ii¥ollf. C:;;t:t,t· fFE, 1975) tIH'i? C7.:> L <~t.l:7.:>o tJJ:ll, :;;t:HO)~' Ij B*O)P'J*1tl'!¥o:tt!!~IRO)t1J.Jl'im/J~';"-"":: ~ 7Mi¥o :tt!!~IRO)-t.-ni;:.J!i~""(';5 I), B *O)7f.*fl!l¥o:tt!!~IRO)-t.-h Ul~t.l: -:>t.: t 0)""(';57.:>;: t ~JHt vt -C~' 7.:> 0 ~\"!5)fi/J' G 11, ~:!lO)itfl:;fi~ A'f- L -C ~, 7.:> /J;, ;: h i? I;:. ,,?~, -C 11JjiJKgcII!lt*l'r-t 7.:>0 :;;t:H~ryj ·1)gJpiB"iJ.: Introduction Inner Zone of Central Japan. Table 1 summarises the stratigraphy of this The Tetori Supergroup is widely supergroup. Vile now mention work on distributed over Nagano, Gifu, Toyama, the floras of the Tetori Supergroup. Ishikawa and Fukui Prefectures in the The flora of the Kuzuryu Group(Kuzuryu * Received April 5, 1976: read Jan. 11, 197c1 Flora) was described by KIMURA (1958b. at Fukuoka and June 14, 1975 at Morioka. 1959b). The flora of the Oguchi For. 343 344 Tatsuaki KIMURA and Shinji SEKIDO mati on (also called the Oguchi Flora and Mt. Hakusan (2702 m) and Irahara along by earlier authors, the "Tetori Flora") the Osugidani, a branch of the upper has been described by GEYLER (1877), course of the Tetori River, Shiramine­ YOkOYAMA (1889), Y ABE(1905, 1922,1927a), mura, Ishikawa-gun, Ishikawa Prefecture, OISHI (1936, 1940,1941), MATSUO & OMURA with the cooperation of Mr. K. Y AMA­ (1968), KIMURA (1961) and KIMURA & ZAKI, a staff of the Komatsu City Museum SEKIDO (1965, '66, '67, '71, '72, '74, '75). and several students of the Tokyo Ga­ The monographical study of this flora is kugei University. now in progress by us. Thus, this paper is the second palaeobo­ The Akaiwa Flora is dealt with in this tanical study of the Akaiwa Flora, the paper but we now realize that the lower first being in the paper by KIMURA and middle parts and the upper part of (1975a). the Tamodani Formation along the Tamo­ dani, studied by KIMURA & HAY ASH! Material may correspond to the Akaiwa Formation and the lower part of the Kitadani For­ The plant remains described in this mation respectively. The uppermost paper have been graphitised or removed member of the Tamodani Formation have from the plant substance and so, while yielded a rich and peculiar flora (called showing their form and venation clearly, the Tamodani Flora by KIMURA, 1975a) they are not suitable for the preparation which includes Arctopterisand Jacutopteris of spores or cuticles. and is of a character previously known The letters, BK and OS used for only in Siberia. registered number show the abbreviation Now under a new heading, the flora of fossil localities, Bettokuzure and Osu­ of the Akaiwa Formation, we collected gidani respectively. Specimens here in 1973-1975 many fossil plants and non­ described are all deposited at the Koma­ marine shells from the Akaiwa Formation tsu City Museum, Komatsu City, Ishika­ at Bettokuzure on the southern slope of wa Prefecture. Table 1. Brief stratigraphy and subdivision of the Tetori Supergroup. Upper Cretaceous Asuwa Group (MATSUO, 1962) and its equivalents --- unconformity --- Kitadani Formation (alternation of sandstone, shale and tuff) (adapted from MAEDA, 1958b) ; Late Neo­ comian below and Aptian above -Akaiwa Formation and its equivalents Tetori Supergroup - -Itoshiro Group _ I (KAWAI, 1961) ; Late Neocomian -Oguchi Formation (KAWAI, 1961); Early Neocomian --- mostly unconformity --- -Kuzuryu Group; Late Jurassic 663. Mesozoic plants from the Akaiwa Formation 345 exilijormis, C. hukuiensis, C. ishikawaensis, Composition of the Akaiwa Flora C. lobifolia, C. triangularis, C. sp., Nils· Among the collection from Bettokuzure sonia koloi, N. nipponensis?, N. orientalis, Otozamites klipsteinii, Ginkgodium na­ and Osugidani (Irahara), the determined thorsti, Czekanowskia rigida, Elalocladus genera and species are shown in Table sp., Taxodium sp., Sequoia? sp., Podoza­ 2 together with those from the middle mites griesbachi, P. lanceolalus, P. reinii, member of the Tamodani Formation Taeniopteris richthofeni?, T.? sp. and (after KIMURA, 1975a). Besides the above, Xenoxylon latiporosum. MAEDA, MAEDA & T AKENAMI and KA­ w AI listed the following species from various localities of the Akaiwa Forma­ Characteristics of the Akaiwa Flora tion and its equivalents: The following characteristics of the Kumanogawa Formation (alternation of Akaiwa Flora seem noteworthy. sandstone and shale) along the Jintsu 1) The Equisetales are only represented and the Kumanogawa Rivers, Toyama by several tubers and a fragment of Prefecture (adapted from MAEDA & stem with a node. These tubers resemble TAKENA~Il, 1957); closely in general outline those from the Adiantites sewardi, Onychiopsis elongata, Oguchi Formation, the Nagdong Group Sphenopteris goepperti, Cladophlebis argu­ tula, C. dentieulata, C. distans, C. exili­ of Korea and the Lower Cretaceous of fonnis, C. hukuiensis, C. triangularis, C. the Siberian Palaeofioristic Area proposed lobifolia, Nilssonia orientalis, Ptero­ by VAKHRAMEEV (1964, 1966, 1970, 1971). phyllum ? sp., Ginkgoites digitata, Czeka­ 2) Ferns are fairly numerous and varied, nowskia rigida, Podozamites lanceolatus, though not so predominant as in the P. reinii and Taeniopteris sp. Oguchi Flora. The specimens named Nochino Formation (conglomerate and sand­ Gleichenites aff. porsildi agree well with stone) along the Uchinami and the Ito­ those in the Tamodani Flora. Coniopteris shiro Rivers, branches of the Kuzuryu which is diverse and abundant in the River, Fukui Prefecture (adapted from Oguchi Flora is rather rare. Only MAEDA, 1957); doubtfully determined sterile leaf and Onychiopsis elongata, Cladophlebis exili­ formis, Podozamiles lanceolalus, P. reinii several detached fertile pinnules were and Xenoxylon laliporosum. obtained. Birisia onychioides formerly Tochio Formation (alternation of sandstone called Coniopteris onychioides agrees with and shale) along the Gamata River, Gifu material described from the various Prefecture (adapted from MAEDA, 1958a) ; localities of the Lower Cretaceous in the Onychiopsis elongala, Cladophlebis den. Siberian Palaeofioristic Area. Cladophlebis ticulata, C. exiliformis, Podozamites shinshuensis originally described by T A­ lanceolatus and Xenoxylon latiporosum. TEIW A from the Shinshu (Chinju in Akaiwa Formation around Mt. Hakusan in Korea) Formation, Korea and that followed Fukui, Ishika \Va and Gifu Prefectures by KI:YIURA (1958a) from the uppermost (K-\'\vAI, 1961) ; member of the Tamodani Formation is lv! archant ites yabei, Equiset ites ushi· marensis, E.