Checklist of Beetles of the British Isles 2nd Edition Edited by A.G. Duff with an Introduction by D.A. Lott and a chapter on Fossil Beetles by P.I. Buckland & P.C. Buckland Checklist of Beetles of the British Isles Copyright © A.G. Duff, 2012. First edition 2008 published by A.G. Duff Revised edition 2012 ISBN: 978-0-9573357-0-7 Published by Pemberley Books (Publishing), United Kingdom. All rights reserved. No part of this book may be reproduced, stored in a retrieval system, or transmitted by any means (electronic, mechanical, photocopying, recording, or otherwise) without written permission from the publisher. Although every precaution has been taken in the preparation of this book, the publisher and authors assume no responsibility for errors or omissions. Neither is any liability assumed for damages resulting from the use of the information contained herein. Within the meaning of Article 8.2 of the International Code of Zoological Nomenclature (4th Edition, 1999), this work is not issued for the purposes of zoological nomenclature. Pemberley Books 18 Bathurst Walk, Iver SL0 9AZ United Kingdom Tel.: +44(0)1753 631114 Fax: +44(0)1753 631115 E-mail: [email protected] WWW: www.pemberleybooks.com Cover: Platyrhinus resinosus (Scopoli, 1763) (Anthribidae) © John Walters Summary of changes in the 2012 edition This section details the main changes in the current edition of the checklist, compared to the previous edition (Duff, 2008). For further details of these changes, please refer to the endnotes. Family-group names: . Nomenclature of family-group names follows Bouchard et al. (2011) . Family PAELOBIIDAE is renamed to HYGROBIIDAE . Family SCYDMAENIDAE becomes a subfamily of STAPHYLINIDAE . Family BOLBOCERATIDAE becomes a subfamily of GEOTRUPIDAE . Order of families in SCARABAEOIDEA changed . Family ANOBIIDAE is renamed to PTINIDAE . Family LANGURIIDAE deleted and included species moved to EROTYLIDAE Species added to the main list (47): . Suphrodytes figuratus . Isorhipis melasoides . Somotrichus unifasciatus . Gastrallus laevigatus . Hololepta plana . Mesocoelopus collaris . Ochthebius nilssoni . Mirosternomorphus heali . Catops borealis . Colotes punctatus . Brachygluta klimschi . Meligethes matronalis . Mycetoporus ambiguus . Meligethes symphyti . Mycetoporus reichei . Typhaea haagi . Acrotona convergens . Berginus tamarisci . Gyrophaena transversalis . Semanotus russicus . Anotylus hammondi . Poecilium lividum . Bledius lohsei . Leiopus linnei . Carpelimus erichsoni . Bruchus brachialis . Carpelimus zealandicus . Aphthona pallida . Carpelimus alutaceus . Longitarsus symphyti . Edaphus beszedesi . Pseudoperapion . Paederidus ruficollis brevirostre . Philonthus alpinus . Bradybatus fallax . Quedius lyszkowskii . Cotaster uncipes . Quedius lucidulus . Allopentarthrum elumbe . Gyrohypnus wagneri . Conarthrus praeustus . Agrilus cuprescens . Otiorhynchus cribricollis . Agrilus cyanescens . Sitona gressorius . Trachys subglaber . Polygraphus grandiclava Species deleted from the main list (9): . Hydaticus continentalis . Abax parallelus . Stenolophus comma . Aglyptinus agathidioides . Alevonota aurantiaca . Bledius praetermissus . Heterocerus hispidulus (duplication error) . Protapion ryei . Cathormiocerus maritimus 1 Acknowledgements I would like to thank everyone who has helped to create and maintain the Checklist of Beetles of the British Isles. During the early part of this project I was greatly assisted by draft checklist manuscripts written by Roger Booth and Eric Philp. For keeping track of new and deleted species, published updates to the British list were invaluable (Lott, 1995, 1998; Mann, 2000b, 2002; Owen, 1993a, 1994c). Recent published checklists were also very helpful covering the Irish fauna (Anderson, Nash & O'Connor, 1997; Nash, Anderson & O'Connor, 1997), water beetles (Foster, 2004, 2005; Foster & Friday, 2011), Carabidae (Luff, 2007), Leiodidae (Cooter, 1996b), Staphylinidae part (Lott, 2009a; Lott & Anderson, 2011), Dermestidae (Peacock, 1993), Nitiduloidea (Cooter, 1995), Kateretidae and Nitidulidae (Kirk-Spriggs, 1996), Ciidae (Orledge & Booth, 2006; Orledge, 2009), Anthicidae (Telnov, 2010), Scraptiidae (Levey, 2009), seed and leaf beetles (Cox, 2007) and weevils (Morris, 2003, 2011). The late Derek Lott deserves a special mention, not only as the original section author for the very large family Staphylinidae, but also for writing his stimulating introduction to the project, and for his unstinting encouragement. With his encyclopaedic knowledge of the British fauna, and nomenclatural issues in particular, a great debt is owed to Roger Booth for undertaking to author many individual families and for answering numerous questions. All the family authors are to be warmly thanked: Keith Alexander (Cantharidae), Max Barclay (Tetratomidae and Tenebrionidae), Roger Booth (numerous families), Jon Cooter (Leiodidae), Mike Cox (Megalopodidae, Orsodacnidae and Chrysomelidae), Garth Foster (water beetles), Colin Johnson (Ptiliidae, Clambidae, Monotomidae, Cryptophagidae, Corylophidae and Latridiidae), Brian Levey (Buprestidae, Mordellidae and Scraptiidae), the late Derek Lott (Staphylinidae), Martin Luff (Carabidae), Darren Mann (Scarabaeoidea), Howard Mendel (Elateroidea), Mike Morris (Curculionoidea), Glenda Orledge (Ciidae) and Martin Rejzek (Cerambycidae). Many thanks also to Phil and Paul Buckland for their chapter on beetles known only as fossils from Quaternary sediments. Several coleopterists have reviewed all or part of the checklist. Tony (A.J.) Allen helped with literature, reviewed numerous drafts of the complete checklist and suggested several formatting improvements. Max Barclay reviewed the arrangement of Otiorhynchus. Martin Collier spotted several errors and suggested some format changes. Scotty Dodd spotted an important error in Altica. Peter Hammond is thanked for correcting several errors, for some formatting suggestions and for thoroughly reviewing the entire checklist. Mark Hill corrected some errors during the course of extracting data for the Biological Records Centre species dictionary. Peter Hodge assisted with literature and drew my attention to a change in Globicornis. Eric Philp spotted a duplication in Heteroceridae. Mark Telfer is to be thanked for his comments on various drafts of the checklist, and for maintaining the MapMate® Coleoptera species dictionary. Finally, the Board of Governors of The Coleopterist journal are to be thanked for allowing the checklist to be hosted on the journal's website at www.coleopterist.org.uk. Andrew Duff Contents Introduction ................................................................................................................ 3 Extant or recently extinct native and naturalised species ...................................... 7 Species found as fossils in Quaternary sediments ..............................................127 Non-established introductions ...............................................................................131 Bibliography ............................................................................................................136 Index to family-group and genus-group names ...................................................157 2 Introduction Introduction D.A. Lott ince publication of the Coleoptera volume of the second edition of Kloet & Hincks' S checklist of British insects (Pope, 1977), a large number of additions and deletions to the list have been published, as well as numerous nomenclatural changes. In addition, recent advances in our understanding of the phylogeny of Coleoptera have led to major modifications of higher classification in checklists and catalogues that have been published more recently in Europe and North America. In the absence of any widely accepted, modern checklist of the British fauna, British workers have been using binomial names taken from a variety of often conflicting sources. It is intended that this checklist should have a wider currency and be suitable for use in future conservation reviews, survey reports and taxonomic dictionaries for biological recording. It is, therefore, vital that the nomenclature used should be widely accepted by coleopterists working on the British fauna. It is also important that the checklist should be as useful as possible to those working at larger scales ranging from European through Palaearctic to the world fauna. It is envisaged that the checklist will be updated regularly to keep abreast of published changes and suggested amendments from correspondents. To this end, comments on the checklist are welcome and should be addressed to the Editor by e-mail to [email protected]. Scope The checklist aims to include all species that have been reliably recorded from the British Isles as possible residents. Exotic species which are only known from casual importation, and have never formed established populations, are listed in an appendix. The British Isles is here taken to include Great Britain and Ireland, including the Isle of Man but not the Channel Islands. The checklist aims to be comprehensive in its listing of subfamilies, genera and species. Tribes and subtribes are only listed when supported by a recent, authoritative review. Contemporary checklists vary in the extent to which they include subgenera and many subgeneric classifications are old and in need of review. Consequently, subgenera have only been listed when they enjoy current and widespread usage. Rationale It is considered that there are two main uses for a checklist: .