Plant Reproduc/on Sexual - Selfing Gavin Douglas and Young Wha Lee Capsella rubella, selfing species (l); and Capsella grandiflora, outcrossing species (r) Sexual - Selfing chasmogamous flower cleistogamous flowers Sexual - Outcrossing selF-incompability Sexual - Outcrossing early protandry late male flowers not yet mature Female flowers recep/ve protogyny dichogamy (temporal separaon oF sexes) Sexual - Outcrossing monoecy dioecy heterostyly unisexual flowers herkogamy (spaal separaon oF sexes) Asterids - Lamiids Asterids Core asterids Lamiids Campanulids Asterids - Lamiids Asterids Lamiales Scrophulariaceae, Lamiaceae Lamiales Scrophulariaceae, Lamiaceae, Verbenaceae (verbena), Oleaceae (olive) Lamiales corolla oLen zygomorphic, bilabiate; stamens didynamous (2 long + 2 short) Lamiaceae / Labiatae Scutellaria lateriflora Monardella odora:ssima Salvia splendens skullcap mountain monardella scarlet sage Salvia dorii Monarda didyma Plectranthus scutellarioides gray-ball sage scarlet beebalm coleus Lamiaceae / Labiatae Salvia officinalis Rosmarinus officinalis Thymus vulgaris sage rosemary thyme Origanum vulgare Ocimum basilicum Mentha spicata Lavandula angusfolia oregano basil spearmint lavender Lamiaceae / Labiatae leaves opposite and decussate, stems sQuare Lamiaceae / Labiatae inflorescence: oen ver/cillate Lamiaceae / Labiatae corolla zygomorphic, oLen bilabiate Lamiaceae / Labiatae stamens didynamous (2 long + 2 short) Lamiaceae / Labiatae ovary 2-carpellate, with 4 lobes and 4 ovules Fruit: schizocarp Forming 4 nutlets “Scrophulariaceae” Penstemon davidsonii Davidson’s beardtongue Caslleja hispida An:rrhinum majus indian paintbrush snapdragon Mimulus guCatus Verbascum blaaria Digitalis purpurea seep monkeyflower mullein foxglove “Scrophulariaceae” “Scrophulariaceae sensu lato” Olmstead et al., 2001 “Scrophulariaceae” corolla zygomorphic, oLen bilabiate “Scrophulariaceae” some, with 4 petals and 2 stamens Veronica “Scrophulariaceae” stamens didynamous (2 long + 2 short) “Scrophulariaceae” stamens didynamous (2 long + 2 short) “Scrophulariaceae” many ovules / carpel Fruit: capsule “Scrophulariaceae” Scrophulariaceae sensu stricto Scrophularia Verbascum Selago figwort mullein Plantaginaceae Angelonia Linaria Penstemon toadflax beardtongue Phrymaceae Mimulus monkeyflower Orobanchaceae Caslleja Pedicularis Orobanche indian paintbrush lousewort broomrape Biogeography Biogeography Biogeography http://www.gbif.org/species/3033663 Distribution of Aconitum Biogeography C A B C A B Dispersal Speciation Biogeography NOAA Satellite and Information service, 1961 - 1990 Mean annual precipitation Biogeography NOAA Satellite and Information service, 1961 - 1990 Minimum temperature Biogeography Biogeography WA DNR Biogeography http://www.nwhi.org/index/gisdata Biogeography Lowland and montane forest Biogeography Prairie Biogeography Temperate rainforest Biogeography Alpine and subalpine Biogeography Ponderosa pine woodland Biogeography Sagebrush steppe .