) Cyprinus Carpio (

) Cyprinus Carpio (

(%&' $#) !" (Cyprinus carpio) * 00 , /. (* +,- )%%&' ($ ) ! "#!$ (77/<4/4; :89' 3!' 77/6/4 :2! 3!') ( ! ) ! * ! #' ! . & !$% ! "# . ! !9: !; & ! &24 : 6-7 89 54 123 & -$ /0 & , + $ +9! B!; !-) A9 ! !@ ! < !, * ? & 9 + , < = > + $ , , 6< $ &# #' < * ? , $ +> .: (EFG :) < + @ 0 => (EFGH %) +9C gonado-) !2 ! 4 &!24 !: * ? & 23 C ( & 24 & 2 $ 7; >, . > . !4 89! ,$ > %3 C0$, *#' .4 ,$ (samatic indicex = GSI !# . ! ! EFFG *!C7 > !, + ! A ! .: 6< C "4 C0 & 23 C ( &24 4 , % 2 4 4 ": .: ",0 / " : 2 4 ! & 3 0 +> & 0 &24 : ,& 2 4 > 6? .: ",0 64 .& 6< , (; & &-$ /0 &, 54123 >, 2 4 (- (Cyprinus carpio) &-$ /0 : Archive of SID E-mail: [email protected] : : : * www.SID.ir ... 4 5 & > C ' * &'$ U! '$ B` &>a b( . 5 _7 * & c$ . @' O & ($ )* & ' . & $% ! "# d>K BC ' $ <K? FU\ & ' -/ (Cyprinus carpio) &($ )* T#G '$ B` BC 8!# .2 & !# 42 3( 2 ' 01 & ' . '$ B` =E . (D B%! %/ 01 ' : ' 0 9 4' &8 67 '$!5 O' (Oncorhynchus mykiss) @$ab W1=5 =>? @A !* 401 <; .$& ! 2 &' &' @ 4>C (Syngnathus scovelli) Naderi ) 8*& B$? C &(1B 4>C "2 ( &8M( Garra Labeo capesis) .(Jolodar and Abdoli, 2004 U' 4(Bardacki et al., 2002) >C e87 (rufa) 82 '& ' BE ( &82! (D (Mojazi et al., 1996) >C e87 (K!) &'^ 8 % 1 & & : F5 >C "2 ( 5 =Ra Mullus surmuletus) & 8 BH &82G .(Sparre et al., 1988) . 2 !# ( ) N’da and Deniel, 1993 Crook and ) 6 %/ K% IJ & G;/ ' g J & ($ )* &' f> M L B5C 4(Robertson, 1999 VW$ ' . <RK !8/ / <RK L J8$ (Appleford et al., 1998) .( ) 2& h( ' Tempero et al., 2006 - 4(McDowall, 1996) @$' * & b( j i & ($ )* @ ' * .2 *& L 48& f> & b( i k ' @ ' &>E ( NO BC &H f> & b( m l & ' & 6K F\8 Q #G PJ #G .2& &K / n @ .( ) & McCrimmon, 1968 Agrawal , ) -# $/ * &$#K! VW $ ' & ($ )* & ' f > & RK & K @ ' .(1996 Horvath, ) 2& ljooo C ' loooo H$ BE ( BSG . & >SG ( 82& BE ( AK 5 & ($ )* &' .(1985 B#K Q & ' 01 T 2 > F\8 . $ &D ? T 2 / 4$ 1 BU5 .(Gougnard et al., 1987) 2 i m H$ * X ) " G VW$ @$ @ T 2 / .(McCrimmon, 1968) OH =\ NO %/ @$ @ 3>Y Z( 4& "'* = 8 : &Archive 4& ($ )* & ' &$>5 of SID Nikolsky , ) 2& RK &>E ( : @* gJ & b K ! <G . $ (GSI) & ' B * @ 8 & U! (D .(1963 >LG .(Tempero et al., 2006) & G;/ ) [D N2 @8J & n lpoo B 5 C / n mooo VW$ & 2 ' $ &82G & .(Biswass , 1993 .( ) 2& / Linhart et al., 1995 FU\ .(Booth and Weyl, 2000) 2& $KJ B5 )* &' BE( 2 PM <##? E* ^ '$ & >C West (1990) ] 1 7 @1 *K q7 rU &($ / @ 2& KGO 7 BC * &'$ * &(D s ^ .(Szabo et al., 2000) U? * * @O\ . 4%1 www.SID.ir ( " !"#" $% &' ()* +, (#- <L L 7 ' ' <L &($ )* * 2 NO 42 PM &U\ T? P$ @ G lj @ H8 / GZ7 Fida ) <RK &K! b( 4d>K 'T? G & * / @H 2& 7 L et al., 1988; Swee and McCrimmon, 1966; 4&G 48@ tS% .( K2 ] & W\ $/ Shikhshabekov, 1972 f> 1 '42 4 =\ L F \ ' $ 8 42 48K!' d>K ' * d>K '@H &' D5 lkl y$M .2 KG &##? .( ) 2 & <RK B* . G 5 & d>K & (\ ' z* Alikunhi, 1966 &82! ' K * G & 8 N2 2 &M8! & '$ Peyghan et al. (2002 & \ ( & 2 7 ) & ( $ ) * ' 8 &82 G ' z * & ' J {p 8_$' . 2 %1 .$ & q7 BSG %1 & !8/ f> BC 2 d>K * &##? 5 K K @ ' =\ . G & Smith and walker (2004) BH 41 B$J (K 08/ P o/l 5 =KM @ K$> 'q RK &($ )* &' @$ =KM RK 8 @ . 2 .$ & &7HH &7H* .2 P o/ol 5 % ' &H &($ )* &' H8 / Presnell ) 6 n>!*$' q F#? T2 &G\ &K @' '$ 1} %/ (and Schreibman, 1997 d>K &G;/ 'gJ &' &>E( p >* ^ & !8/ f> BC &82 . 2 RK 4$& ; 4d>K '= #D8 &KC .G <L (Biswas, 1993) >C : 2 PM u' =SC %/ (D GSI= WG/Wt × looD 8 N2 =\ &G 48@ n & ' B* @ = WT 48 @ = WG =G .1 L) w208/ &($ )* &' .(Biswas, 1993) ((H F>D 1' # r ' B >? M rU ( ) 8 N2 < & n (1 RK d>K 8 @' &U! GSI w208/ &($ )* &' @1 H @1 F \ 'b ! # GDH X ' B >? M %/ SPSS15 G P . 2 PM &' @$ $ BH d>K BC xO n 5 R K ' $ % / Excel GP w208/ (& A) &($ )* Archive .G of SID 7 L & 4&($ )* & ' $ B? #$!" . &52 08/ BC #K! (H B C 4 1 B $ J &82 G '& / BC & K @ ' L Y1 @$' [\ @ 7 * Y1 @K!> @K & Y &52 Cyprinus ) & ( $ ) * & ' & !8/ & d>K & ' $ .G & K @ ' L :2 8 !# [2 >C "2 (carpio www.SID.ir ... z7K G P K$* ' (Peri-nucleolar) # &'( % K!' ({p m) $ . * =;2 (l{o m) ]8 BH2 '$ 2C 2 PY ' :* ' .G "G K!' &> 2C K!' K!' .(l BH2) BHO K!' ' .(l BH2) ;K 'K!' BH2 . (Tertiary yolk) +/+ 0 % =;2 z7K G P B5C ' (Yolk vesicle) /+ -.,+ % E*C K!' ({ m) '$ . * OY &> 2C K :* '=H .(l BH2) # .2 8*7 z7K P$ X) => BH2 .G "G eK 6 K!' !C (Migratory nucleus) '( 1 % .(l BH2) 2 ]8 e 'K!' K!' $ 2C $ K!' *C @W y2 '$ $ BH >C .2& NO (Primary yolk) +/+ - % .O 'O BG86 V2 &87 ' &>L >C Y1 >C .2 BHO '=H + -2 K$* >C ' . & ($ )* 2 & ' $ @ =\ D . * =;2 z7K G P ! K$> jo & ' =\ # KO * @O "G @1 ]8 BH2 K!' $ @ # K$* . K$> mjo @ 1 K$* .(l BH2) G "G (ip m) $ UQ P koo @ 1 KO P mpj 2 & .(m BH2) (Secondary yolk) +/+ 0/ % Archive of SID (*+, (') %& $# "! X, ) ($7 $6 *) <5 * ;(89X, HandE) (/57 $6 *) 45 * :0 0 1. 2 3//*. <5 *) ;(9X, HandE) ($2(2 45 *) <=> * ;(9X, HandE) ((2 $ 4#25 *) < * ;(89HandE (AX, HandE) (*# ?/ *) ?@ * ;(9X, HandE) ($2(2 < *) ;(9X, HandE) ($2(2 www.SID.ir ( " !"#" $% &' ()* +, (#- 4000 W = 0.0112*L3.0587 و 2 2000 R = 0.9457 زن (م ( 0 020406080 ل () (*+, (') %& $# "! ( D25 5 4E F. *C. B + VM 8R .G "'* X) &M 2 L @ ' @ = \ & D @ O%U 2 b$O: "G N2 4 & ]Cz m BH2 * D$' . U? Y1 & ( O%U . # E*C @ =\ (%o " ) & W ! &bK!U$' .(k BH2) G %2 @7 "'* 2 . / @' (GSI) /# 34$ )* & ' (GSI) 8 N2 < ; & * @O $ =\ &($ " G 1 4@1 ' N2 20 10 17 a 13 15 14 15 13 15 ab 12 7 ab ab ab 8 ab ab ab 4 10 3 ab b b b د( (% 5Archive of SID 0 د ذ داد دا ا ر ن ( ر د 5 اردورد ( 38 (1 (1386 (*+, (') %& $# "! ( $ I& FH, G .. ( *,, / H,#, F/1 L5 $1. ( K J www.SID.ir ... 100% 80% 60% %40 اوا ( (% 20% 0% ا د د رد و ذر اد رد ا ن ( ر داد ) (1385 (1386 ر ا زرد ز زرد ز وز زرد ه ر (*+, (') %& $# "! $= ( 0, + 76 # 5" d>K 8 &U! F>D 1' b @ 1 E*C = m 42 L @ ' B5C - F>D 1 ' E*C B5C . 2 U? i & 8 ' r & 8 & G KO . = 1 ' b . p{ mjmok 3 .(j BH2) = p .(n= {i) 2 J likkom/ {{jjk 3 & U! 1 ' E*C B5C 1 @ @ P> * ' mkilo .(l =/) 20 16 Archive12 of SID 8 اوا 4 0 23456789 (*+, (') %& $# "! ( (/) ,5 + www.SID.ir ( " !"#" $% &' ()* +, (#- %& $# "! F , 5 C $5 ? D% 45 /± / (a) /± / (a) /± / (a) /± / (a) /± / (a) /± / (ab) /± / (a) /± / (ab) /± / (a) /± / (b) /± / (a) /± /(b) .' $( %)* % %&% !$"#" ! / / lk/m) @1 @ K$* (bK 87 / . UQ (bK B H 2 $ d>K ' 01 @ / m{/) @ KO . 1 { 30 25 20 د 15 10 ر اد 5 0 ا رد و د ذر داد ر ا ن ( ر داد د (1385 ) 86 (13 (*+, (') %& $# "! $( ! $ + @$' y &($ )* &' @$ y .(Tempero et al., 2006 4 Smith and Walker, 2004) &' @$ BH F#? .8$& BH y2 '$ ' Archive of w2 08/SID (Cyprinus carpio) &($ )* @ ( P$ * x 8 5 & &82G <(D RK '$ @& &' @$ . ' . KG 'K &( 4$ 'O & d>K BC @& $ BH >C "2 &($ )* P$ . 5 & >C '$ M FD $ BH BC xO .$ 'O - &$ K K '$ Hunter and Macwicz (1985) T * _1 8: =\ H$ M 8 .2 Rahman et al. (2000) Matsuyama et al. (1988) J! T2 8 .(Horvath, 1985) ' KR' . 2 @ / lp W J " ) www.SID.ir ... 5 K * &'$ P$ VW$ .(1990 & &($ )* &' 8 BH 4(&K Bishai et al., ) 2& K @ 4KG 7 2 PM <(D & .2 PM K$ \ H$ L mo C &( 4(1974 lm B5C &( 4* $' ( ' .(Smith and Walker, 2004) 2 K Swee and ) K! W J VW$KC 8* " H$ McCrimmon, 1966; Brzuska, 1989; Guha and ]8 8: ' .( .82& B5 '$ M K Mukherjee, 1991 ) "# &($ )* &' '$ 2& K h( '$ * & Sivakumaran ) % "# &( 4( *& 8& &5 BH &K >C Davies and et al., 2003 @H J! & T2 .( . 0Z/ K h( '$E* Hanyu, 1986 $K! \ &' 42 1 @ &>E ( .(Horvath, 1985; Crivelli, 1981) * ' (GSI) 8 ' N2 RK '&' &($ )* &' * G MK @& 8 .(Biswass, 1993) <UQ Vz* 5 = =\ P$ ! F\8 & G #K! rU 8' @O "'A7 eK .2& * (GSI) 8 N2 & 8?8 & ' f> BC &($ )* &' >LC eK / 8 N2 # .'& @O BSG .& <L = B5C KO = ' .

View Full Text

Details

  • File Type
    pdf
  • Upload Time
    -
  • Content Languages
    English
  • Upload User
    Anonymous/Not logged-in
  • File Pages
    14 Page
  • File Size
    -

Download

Channel Download Status
Express Download Enable

Copyright

We respect the copyrights and intellectual property rights of all users. All uploaded documents are either original works of the uploader or authorized works of the rightful owners.

  • Not to be reproduced or distributed without explicit permission.
  • Not used for commercial purposes outside of approved use cases.
  • Not used to infringe on the rights of the original creators.
  • If you believe any content infringes your copyright, please contact us immediately.

Support

For help with questions, suggestions, or problems, please contact us