Arthropoda Selecta 5 (1/2): 75-137 © ARTHROPODA SELECTA, 1996 A checklist of the spiders of Russia and other territories of the former USSR Ðååñòð ïàóêîâ Ðîññèè è äðóãèõ òåððèòîðèé áûâøåãî ÑÑÑÐ K.G. Mikhailov Ê.Ã. Ìèõàéëîâ Zoological Museum of the Moscow State University, Bolshaya Nikitskaya Str. 6, K-9 Moscow 103009 Russia. Çîîëîãè÷åñêèé ìóçåé ÌÃÓ, óë. Á. Íèêèòñêàÿ 6, Ê-9 Ìîñêâà 103009 Ðîññèÿ. KEY WORDS: spiders, Russia, ex-USSR, faunistics, checklist. ÊËÞ×ÅÂÛÅ ÑËÎÂÀ: ïàóêè, Ðîññèÿ, áûâøèé ÑÑÑÐ, ôàóíèñòèêà, êîíòðîëüíûé ñïèñîê. ABSTRACT: A checklist of the spiders of Russia Ç+È mountainous Middle Asia, and other ex-USSR territories is given for the first Ê deserts of Middle Asia, time since Kharitonov [1932, 1936]. All valid Ë Kazakhstan hills, names, the main synonyms as well as nomina dubia Ì West Siberia, and nomina nuda are indicated. Í+Î Middle Siberia, Ï mountains of South Siberia, ÐÅÇÞÌÅ: Âïåðâûå ïîñëå ðàáîò Õàðèòîíîâà Ð Northeastern Siberia, [1932, 1936] äàí êîíòðîëüíûé ñïèñîê ïàóêîâ Ñ1 Far North-East, Ðîññèè è äðóãèõ òåððèòîðèé áûâøåãî ÑÑÑÐ. Ñ2 Kamchatka, Óêàçàíû âñå âàëèäíûå íàçâàíèÿ, îñíîâíûå ñèíî- Ñ3 Northern Kurile Islands, íèìû, ñîìíèòåëüíûå è ãîëûå íàçâàíèÿ. Ñ4 Commander Islands, Ò1 continental Southern Far East (Amur- Introduction Maritime area), Ò2 Sakhalin and Moneron islands, The checklist below is actually an extraction Ò3 Southern Kurile Islands. from a complete card catalogue of the spiders of the Republics: USSR started by the author in 1981. It is compiled ÐÔ Russia, almost exclusively as based on the literature data, Ýñò Estonia, with new materials mentioned only occasionally. Ëàòâ Latvia, Some calculations of the number of species between Ëèò Lithuania, different regions have already been published else- Áåë Byelorussia, where [Mikhailov, 1996ab]. Óêð Ukraine, Each spider species included in this checklist is Ìîëä Moldova (Moldavia), supplied with an attribution both to main physio- Ãð Georgia, graphical areas delimited after Gvozdetskiy [1968] Àç Azerbaijan, and to post-Soviet republics. Àðì Armenia, The physiographical areas and republics are Êàç Kazakhstan, coded by Russian litters and abbreviations, respec- Óçá Uzbekistan, tively. The sequence of references is as follows: Òóðê Turkmenistan (Turkmenia), Physiographical areas (see also Map 1): Êèðã Kyrgyzstan (Kirghizia), À Atlantic-Arctic insular area, Òàäæ Tajikistan. Á Fennoscandia (Karelian-Kola area), Synonyms are marked with a = sign, only names  Russian Plain, cited in the Russian/Soviet arachnological literature Ã1 Novaya Zemlya and Vaigach islands, are reported. Some old combinations not accepted Ã2 Urals, here are also incorporated without =. Valid subspe- Ä Carpathians, cies are mentioned with a + sign. For nomina dubia Å1 Crimea, and nomina nuda, all available sources are reported. Å2 Caucasus, Nomina dubia are given in the main list, whereas Æ1 Armenian Upland, nomina nuda are removed to the end of each family Æ2 Kopetdagh Mts., list. Doubtful identifications are marked with a ?, 76 K.G. Mikhailov Ñ1 Ã1 Ð Ñ4 Á Ñ2  Ñ1 Ä Ì Ñ3  Ò2 Ã2 Í+Î Ò1 Å1 Ï Ò3 Â Ï Å2 Ë Æ1 Ê Ç+È Æ2 Ç+È Map. Physiographical regions of the ex-USSR territory (after Gvozdetskiy, 1968). Explanations see in text. Êàðòà. Ôèçèêî-ãåîãðàôè÷åñêèå ðàéîíû òåððèòîðèè áûâøåãî ÑÑÑÐ (ïî Ãâîçäåöêîìó, 1968). Îáúÿñíåíèÿ ñì. â òåêñòå. misidentifications begin a separate paragraph and are Verzhbitskiy, Zawadsky instead of Zavadsky. Yet in marked with a ?? symbol. Species names recorded the list of references all such names are anglified as in the ex-USSR territory with doubts or definitely opposed to their latinized names used after their taxa erroneously are cited in figure brackets ({}) and in in the main list. regular brackets (), respectively. If necessary, new materials are supplied with Correct spellings of most Latin generic and indications of the museum in which they are kept: BI specific names have been checked after catalogues Zoological Museum, Biological Institute, Siberi- and the original sources, if necessary. Dates of an Branch of the Russian Academy of Sciences, publication of some papers by Savigny & Audouin, Novosibirsk. C.W. Hahn and C.L.Koch are established with the help of special literature sources [Brignoli, 1985; ACKNOWLEDGMENTS. I am deeply obliged Sacher, 1988; Sherborn, 1914; Tollitt, 1986]. to my colleagues and friends Drs. Yu.M. Marusik According to ICZN [1985], Article 33d, original (Magadan), A.V. Tanasevitch (Moscow), and D.V. species endings cannot be changed from -ii to -i. So, in Logunov (Novosibirsk) for valuable and numerous theory a lot of traditionally accepted names of Europe- comments on this checklist. I am also grateful to Drs. an spiders are to be changed: Maso sundevallii instead K.Yu. Eskov (Moscow), A.A. Zyuzin (Alma-Ata) of M. sundevalli, Pachygnatha degeerii instead of P. and V.I. Ovtsharenko (St.-Petersburg) for their degeeri, etc. I am not prepared yet to applying this rule remarks on earlier versions of my card catalogue. I directly, so I have simply marked all such cases in the greatly appreciate the bibliographical help by Dr. T. text of the current checklist by a double asterisk (**). Kronestedt (Stockholm, Sweden). The work is Only for synonyms some replacements have been made supported in part through the INTAS grant 94-3708. without special reference. Please note that the spelling of some Russian Checklist names has traditionally been non-English, mostly German: Bachwalow instead of Bakhvalvov, Chari- Atypidae tonov instead of Kharitonov, Mcheidze instead of Mkheidze, Michailov instead of Mikhailov, Pono- Atypus affinis Eichwald, 1830 marjov instead of Ponomarev, Saveljeva instead of Â. Áåë, Óêð, Ìîëä. Savelyeva, Tystshenko instead of Tyshchenko, Atypus magnus Namkung, 1986 Wagner instead of Vagner, Wierzbitzki instead of Ò1. ÐÔ. A checklist of spiders of Russia and other ex-USSR territories 77 Atypus muralis Bertkau, 1890 Zaitunia martynovae (Andreeva et Tystshenko, 1969) = A. anachoreta L.Koch in Ausserer, 1871 Ç+È. Òàäæ. Â, Ä, Å1, Å2, Æ2. ÐÔ, Óêð, Ìîëä, Ãð, Àç, Òóðê. Zaitunia monticola (Spassky, 1941) Atypus piceus (Sulzer, 1776) Ç+È. Òàäæ. Â. Ìîëä. Sicariidae Ctenizidae Loxosceles rufescens (Dufour, 1820) Sterrhochrotus ferghanensis (Kroneberg, 1875) Æ2, Ç+È, Ê. Êàç, Óçá, Òóðê. Ç+È, Ê. Êàç, Óçá. Òóðê?? Scytodidae Sterrhochrotus kitabensis (Charitonov, 1946) Ç+È, Óçá, Êèðã. Scyloxes asiatica Dunin, 1992 Ummidia gandjinoi (Andreeva, 1968) Ç+È. Òàäæ. Ç+È. Òàäæ. Scytodes bertheloti Lucas, 1839 Ê. Òóðê. Dipluridae Scytodes strandi Spassky, 1941 Æ2, Ç+È, Ê. Óçá, Òóðê, Êèðã, Òàäæ. Phyxioschaema raddei Simon, 1889 Scytodes thoracica Latreille, 1804 = Ischnotrhele strandi Spassky, 1937 Â, Ä, Å1, Å2, Æ1. ÐÔ, Óêð, Ìîëä, Ãð, Àç, Àðì. Æ2, Ç+È, Ê. Óçá, Òóðê, Òàäæ. Scytodes univittatus Simon, 1882 Æ2, Ç+È. Òóðê, Êèðã. Nemesiidae Leptonetidae (Amblyocarenum walckenaerii (Lucas, 1846)) Ç+È?? Óçá?? Leptonetela caucasica Dunin, 1990 Anemesia birulai (Spassky, 1937) Å2. Ãð. Æ2, Ê. Òóðê. Anemesia karatauvi (Andreeva, 1968) Pholcidae Ç+È. Òàäæ. Raveniola fedotovi (Charitonov, 1946) Artema transcaspica Spassky, 1934 Æ2?, Ç+È. Óçá, Òóðê? Æ2, Ç+È, Ê. Êàç, Óçá, Òóðê, Òàäæ. Raveniola ferghanensis (Zonshtein, 1984) Ceratopholcus maculipes Spassky, 1934 Ç+È. Óçá. Êèðã. Ç+È, Ê. Óçá, Òóðê, Òàäæ. Raveniola hyrcanica Dunin, 1988 Hoplopholcus forskali (Thorell, 1871)** Å2. Àç. Ê. Òóðê. Raveniola kopetdaghensis (Fet, 1984) Hoplopholcus longipes (Spassky, 1934) Æ2. Òóðê. Å2. ÐÔ, Ãð. Raveniola pontica (Spassky, 1937) Pholcus alticeps Spassky, 1932 Å2. ÐÔ, Ãð. Â, Å2, Ç+È. ÐÔ, Ãð, Àç, Òàäæ. Raveniola recki (Mcheidze, 1983) Pholcus crassipalpis Spassky, 1937 Å2. Ãð. Â. ÐÔ, Óêð. Raveniola redikorzevi (Spassky, 1937) Pholcus crypticolens Bösenberg et Strand, 1906 Ê. Òóðê. Ò1. ÐÔ. Raveniola virgata (Simon, 1891) Pholcus nenjukovi Spassky, 1936 = Brachythele arnoldii Zonshtein, 1984 Ç+È.- Óçá, Êèðã. Æ2, Ç+È. Òóðê, Òàäæ. Raveniola zaitzevi (Charitonov, 1948) Pholcus opilionoides (Schrank, 1781) Å2. Ãð, Àç. Â, Ä, Å1, Å2, Ç+È, Ê, Ò1. ÐÔ, Óêð, Ìîëä, Àç, Êàç. Á?? Mygale caucasicus Wagner, nomen nudum Pholcus phalangioides (Fuesslin, 1775) Å2. Ãð [Vagner, 1896]. Á, Â, Ä, Å1, Å2, Ê. ÐÔ, Áåë, Óêð, Ãð, Àç, Êàç, Òóðê. Pholcus phungiformes Oliger, 1983 Filistatidae Ò1. ÐÔ. Pholcus ponticus Thorell, 1875 Filistata insidiatrix (Forskal, 1775) Â, Å2, Ç+È, Ê. ÐÔ, Óêð, Àç, Êàç, Óçá. Å2, Æ2. Ãð, Àç, Òóðê. Pholcus sidorenkoi Dunin, 1994 Microfilistata tyshchenkoi Zonshtein, 1990 Â. ÐÔ. Ç+È. Òàäæ. Pholcus sogdianae Brignoli, 1983 Pritha crosbyi (Spassky, 1938) = P. fagei Spassky, 1940, praeocc. Æ2, Ç+È. Êàç, Óçá, Òóðê, Òàäæ. Â, Ç+È. ÐÔ, Êàç. Zaitunia beschkentica (Andreeva et Tystshenko, 1969) Pholcus velitchkovskyi Kulczyñski, 1913 Ç+È. Òàäæ. Â, Å1. ÐÔ, Óêð. Zaitunia maracandica (Charitonov, 1946) Spermophora senoculata (Dugès, 1836) Ç+È. Óçá. Å1, Å2. ÐÔ, Óêð, Àç. 78 K.G. Mikhailov Segestriidae Dysdera gmelini Dunin, 1991 Å2. Ãð. Segestria bavarica (C.L.Koch, 1843) Dysdera hungarica Kulczyñski in Chyzer et Kulczyñski, Ä, Å2. Óêð, Ãð, Àç. 1897 Ç+È?? Òàäæ?? + D. h. atra Mcheidze, 1979, Segestria florentina (Rossi, 1790) + D. h. subalpina Dunin, 1992 Å1, Å2. Óêð, Ãð. Â, Å1, Å2. Óêð, Ìîëä, Ãð, Àç. Segestria senoculata (Linnaeus, 1758) Dysdera imeretiensis Mcheidze, 1979 Á, Â, Ä, Å1, Å2. ÐÔ, Áåë, Óêð, Ìîëä, Ãð. Å2. Ãð. Segestria turkestanica Dunin, 1986 Dysdera incognita Dunin, 1991 Ç+È. Êèðã, Òàäæ. Å2. ÐÔ. Dysdera inopinata Dunin, 1991 Dysderidae Å2. Ãð. Dysdera karabachica Dunin, 1990