School of Medicine Vanderbilt University 2011/2012 Archived 2011/2012 Medical School Catalog Containing general information and courses of study for the 2011/2012 session corrected to August 2011 Nashville The university reserves the right, through its established procedures, to modify the requirements for admission and graduation and to change other rules, regulations, and provisions, including those stated in this bulletin and other publications, and to refuse admission to any student, or to require the with- drawal of a student if it is determined to be in the interest of the student or the university. All students, full- or part-time, who are enrolled in Vanderbilt courses are subject to the same policies. Policies concerning noncurricular matters and concerning withdrawal for medical or emotional reasons can be found in the Student Handbook, which is on the Vanderbilt website at www.vanderbilt.edu/student_handbook.Archived 2011/2012 NONDISCRIMINATION STATEMENTMedical School Catalog In compliance with federal law, including the provisions of Title VII of the Civil Rights Act of 1964, Title IX of the Education Amendment of 1972, Sections 503 and 504 of the Rehabilitation Act of 1973, the Americans with Disabilities Act (ADA) of 1990, the ADA Amendments Act of 2008, Executive Order 11246, the Uniformed Services Employment and Reemployment Rights Act, as amended, and the Genetic Information Nondiscrimination Act of 2008, Vanderbilt University does not discriminate against individuals on the basis of their race, sex, religion, color, national or ethnic origin, age, disability, military service, or genetic information in its administration of educational policies, programs, or activities; admissions policies; scholarship and loan programs; athletic or other university-administered programs; or employment. In addition, the university does not discriminate against individuals on the basis of their sexual orientation, gender identity, or gender expression consistent with the university’s nondiscrimination policy. Inquiries or complaints should be directed to the Equal Opportunity, Affirmative Action, and Disability Services Department, Baker Building, PMB 401809, Nashville, TN 37240-1809. Tele- phone (615) 322-4705 (V/TDD); Fax (615) 343-4969. The text of this publication is printed on paper with 30% post-consumer recycled content with ink made from renewable resources. Copyright © 2011 Vanderbilt University Printed in the United States of America Contents Calendar 4 Administration 5 Medical Center Overview 12 Life at Vanderbilt 19 Medical Education at Vanderbilt 26 Admission 34 Academic Program and Policies 41 Honors and Awards 50 Financial Information 52 Courses of Study 59 Research in Medical Sciences 88 Faculty 94 Index 174 Archived 2011/2012 Medical School Catalog School of Medicine Calendar 2011/2012 FALL SEMESTER 2011 Registration and classes begin for 3rd and 4th year MD students / Tuesday 5 July Registration for 1st and 2nd year MD students / Thursday 28 July to Friday 29 July Classes begin for 1st and 2nd year MD students / Monday 1 August Fall semester begins for medical master's and other doctoral programs / Wednesday 24 August Labor Day— No MD class or clinical activities / Monday 5 September Falll break for medical master's and other doctoral programs / Thursday 6 October–Friday 7 October Fall break for 1st and 2nd year MD students / Saturday 15 October to Tuesday 18 October Thanksgiving holiday for all MD classes / Thursday 24 November to Sunday 27 November Thanksgiving holiday for medical master's and other doctoral programs / Saturday 19 November to Sunday 27 November Fall semester ends for all MD students / Friday 16 December Fall semester ends for medical master's and other doctoral programs / Saturday 17 December Holiday break for all MD classes / Saturday 17 December to Monday 2 January Holiday break for medical master's and other doctoral programs / Saturday 18 December to Sunday 8 January SPRING SEMESTER 2012 Spring semester begins for all MD classes / Tuesday 3 January Spring semester begins for medical master's and other doctoral programs / Monday 9 January Martin Luther King, Jr., Day—No MD class or clinical activities / Monday 16 January Spring holidays for 1st and 2nd year MD students / Saturday 3 March to Sunday 11 March Spring holidays for medical master's and other doctoral program students / Saturday 3 March to Sunday 11 March Spring holidays for 3rd year MD students / Saturday 24 March to Sunday 1 April Spring holidays for 4th year MD students / Friday 16 March to Sunday 18 March Instruction ends for 4th year MD students / Sunday 22 April Spring semester ends for medical master's and other doctoral programs / Friday 4 May Instruction ends required courses for 1st year MD students / Monday 7 May Instruction ends required courses for 2nd year MD students / Friday 11 May Commencement / Friday 11 May Emphasis for 1st year MD students / Monday 14 May to Sunday 20 July Memorial Day—No MD class or clinical activities / Monday 28 May Instruction ends for 3rd year MD students / Friday 22 June SUMMER SESSION 2012 Maymester begins for Master of Education of the Deaf program / Monday 7 May Summer session begins for medical master's and other doctoral programs / Tuesday 5 June Summer session ends for medical master's and other doctoral programs / Friday 10 August Archived 2011/2012 Medical School Catalog Vanderbilt University Board of Trust MARK F. DALTON, Chairman of the Board, Scarsdale, NY JACKSON W. MOORE, Vice Chairman, Memphis, TN NANCY PEROT MULFORD, Vice Chairman, Dallas, TX JOANNE F. HAYES, Secretary, Nashville, TN NICHOLAS S. ZEPPOS, Chancellor of the University, Nashville, TN MARY BETH ADDERLEY-WRIGHT E JOHN R. HALL E RICHARD H. SINKFIELD La Jolla, CA Lexington, KY Atlanta, GA MICHAEL L. AINSLIE L. HALL HARDAWAY, JR. E WYATT H. SMITH Palm Beach, FL Nashville, TN Reform, AL JOHN D. ARNOLD H. RODES HART E CAL TURNER Houston, TX Brentwood, TN Franklin, TN WILLIAM W. BAIN, JR. JOHN R. INGRAM EUGENE H. VAUGHAN E Boston, MA Nashville, TN Houston, TX LEE M. BASS MARTHA R. INGRAM E THOMAS B. WALKER, JR. E Fort Worth, TX Nashville, TN Dallas, TX ELIZABETH S. BENNETT ORRIN H. INGRAM LEVI WATKINS, JR., M.D. Pittsboro, NC Nashville, TN Baltimore, MD DARRYL D. BERGER EDITH CARELL JOHNSON DUDLEY BROWN WHITE E New Orleans, LA Nashville, TN Nashville, TN CAMILLA DIETZ BERGERON LESLIE C. LABRUTO W. RIDLEY WILLS II E New York, NY Spring Lake, NJ Nashville, TN DENNIS C. BOTTORFF J. HICKS LANIER J. LAWRENCE WILSON E Nashville, TN Atlanta, GA Bonita Springs, FL LEWIS M. BRANSCOMB E EDWARD A. MALLOY, C.S.C. E REBECCA WEBB WILSON La Jolla, CA Notre Dame, IN Memphis, TN BILLY RAY CALDWELL ALYNE QUEENER MASSEY E WILLIAM M. WILSON Nashville, TN Nashville, TN Nashville, TN SHERYLL D. CASHIN EDWARD G. NELSON E Washington, DC Nashville, TN E Emerita/Emeritus Trustee THOMAS F. CONE E AYO OSITELU MARIBETH GERACIOTI, Assistant Secretary Nashville, TN ArchivedWestfield, IN 2011/2012 of the University CECIL D. CONLEE E COURTNEY C. PASTRICK Atlanta, GA Bethesda, MD BROWNLEE O. CURREY, JR. E MedicalHENRY ROSS PEROT, School JR. Catalog Nashville, TN Plano, TX CLAIBORNE P. DEMING JUDSON G. RANDOLPH, M.D. E El Dorado, AR Nashville, TN BRUCE R. EVANS JOHN W. RICH E Boston, MA Nashville, TN WILLIAM W. FEATHERINGILL KENNETH L. ROBERTS E Birmingham, AL Nashville, TN KAREN T. FESMIRE JOE L. ROBY Germantown, TN New York, NY FRANK A. GODCHAUX III E EUGENE B. SHANKS, JR. Houston, TX Greenwich, CT Vanderbilt University Administration NICHOLAS S. ZEPPOS, J.D., Chancellor RICHARD C. MCCARTY, Ph.D., Provost and Vice Chancellor for Academic Affairs JEFFREY R. BALSER, M.D., Ph.D., Vice Chancellor for Health Affairs and Dean of the School of Medicine JERRY G. FIFE, B.S., Vice Chancellor for Administration BETH A. FORTUNE, M.A., Vice Chancellor for Public Affairs SUSIE S. STALCUP, B.B.A., C.F.P., Vice Chancellor for Development and Alumni Relations BRETT SWEET, M.B.A., Vice Chancellor for Finance and Chief Financial Officer DAVID WILLIAMS II, J.D., LL.M., M.B.A., Vice Chancellor for University Affairs and Athletics; General Counsel; Secretary of the University MATTHEW WRIGHT, M.B.A., Vice Chancellor for Investments Academic Deans JEFFREY R. BALSER, M.D., Ph.D., Vice Chancellor for Health Affairs and Dean of the School of Medicine MARK D. BANDAS, Ph.D., Associate Provost and Dean of Students CAMILLA PERSSON BENBOW, Ed.D., Dean of Peabody College JAMES W. BRADFORD, JR., J.D., Dean of Owen Graduate School of Management DOUGLAS L. CHRISTIANSEN, Ph.D., Vice Provost for Enrollment Management and Dean of Admissions COLLEEN CONWAY-WELCH, Ph.D., Dean of the School of Nursing CAROLYN DEVER, Ph.D., Dean of the College of Arts and Science CONNIE VINITA DOWELL, M.L.S., Dean of Libraries KENNETH F. GALLOWAY, Ph.D., Dean of the School of Engineering CHRIS GUTHRIE, J.D., Dean of the Law School DENNIS G. HALL, Ph.D., Vice Provost for Research and Dean of the Graduate School JAMES HUDNUT-BEUMLER, Ph.D., Dean of the Divinity School MARK WAIT, D.M.A., Dean of Blair School of Music FRANCIS W. WCISLO, Ph.D., Dean of The Ingram Commons Archived 2011/2012 Medical School Catalog Vanderbilt University Board of Trust Medical Center Affairs Committee ORRIN H. INGRAM, Chair JOE L. ROBY Nashville, TN New York, NY LEVI WATKINS, JR., M.D., Vice Chair EUGENE B. SHANKS, JR. Baltimore, MD Greenwich, CT CAL TURNER Franklin, TN ELIZABETH S. BENNETT Pittsboro, NC REBECCA WEBB WILSON Memphis, TN DENNIS C. BOTTORFF Nashville, TN WILLIAM M. WILSON Nashville, TN MARK F. DALTON Scarsdale, NY NICHOLAS S. ZEPPOS Nashville, TN KAREN T. FESMIRE Germantown, TN Community Members JOHN R. INGRAM Nashville, TN MICHAEL J. BURRY, M.D. Cupertino, CA EDITH CARELL JOHNSON Nashville, TN RICHARD B. JOHNSTON, JR., M.D. Aurora, CO JACKSON W.