"#$% / ! / / / Uncinula necator (Schw.)Burr. !&' () * )+() ,- . !.0 (! * 1! 23 456 @ *!>.6? < :=0! *2 5 9:!.+ 6;. < 9:*! *!7 9" . 7 8 52 5AB 23 456 5E! * H;! * GI ! J-+ G; 5I / !&' () * D )+() ,- . F G ,D >E C;! * O) /A G Uncinula necator (Schw.)Burr.!&' () 5 M K* G N; ( < ! . .K.' H! !.0 (! * 1! 1(O* SO+ U. necator DRAM .5 5;* !.O0 (! * GOI * O. Q; * O1! P!0 8* DGO!7 XW#V O GO 7 DG 0 1! 23 456 D C()U 95 M 7 DG!7 5 ! 5 TT.'R! C;.( .5*. () * DG0 D[. * 9 D[ . 5 /A+ D)+() 5?* #TWZT ! , .55 .D Y .( ( .5O* (! !* GO .'R! 1(* * .( Q; ! , '5 G * 9D* /' D G!7 5 \ * * RAM !] ! D G * ^ _ * 5 !*G1 D[ . J3 D )+() ! K;*! + . ! ! * =? G DRAM * K ; ,D N;5+ G DRAM G!7 I 95* G!7 G * ^ EA .A D` J3 * + . 5?*"a G K;*! C;* . D * K ; ,;!]! N;5+ G 5 6! /;! ! ! * .6? 58* 5 6! /;! ! + * _ \ * * D c AM C()A !. * K ; ,D * C;! * b!* G DRAM PM /) (E .5 .5 /;! ! * .' :/a () * .' # G ]; E * .' T * 58 ! DRAM ;! .* .' "/e !] G G * (! d 6+! () * 1()A .!. * Dc AM \!;* K - ,D !] C;.( G 5 hU 5D* GI ! J-+ G; ! G ! M ,- G 5 = g! I \5;. D[ . fI DRAM .5 !.0 (! D () + * U. necator !&' () ?! /A D)+() 1! 923 456 9!&' () 9RAM 9)+() : 5 Di! '(" ) !" #$$%& . 0 ) (& ,/". ) !" ,'*". )*+ ,- . "#$% / ! / / .(WC WR )R )C) '9. '. 3/. %5 6'7 A S* G 5- 9' n+ 4= V$G& 4 ! 39+ 8 U. necator 2 3& 4 )3/. %5 6'7 '9& S:N U = KF. oI" 3/. %5 6'7 9. @9A& 39/. 9 A ?& >( : ; <%= S*"& Z 4+ c; ? @3F&3 FG ! 9 U. necator 9@ 3F&3 FG 3H .(D C) '+ 99$. 99 399p 499 99 ?99 ? 99 )'.399+99 =J K ,'+ L - MN ! ? FI. 49 ,' '=3& ^/. F! S*+ 3T 4 @3F&3 FG .(R) '9. 3 ,'9+ ,'9 9 93/. O9$. P9GQ 4 39/. % 59 6'7 A ' 8 4= V$G& 3T 4 '.3+ S*"& 4.3- ? 2 3& 4 @3F&3 FG .KN- @8. 9A=U )* )= A 4= V$G& 4 ?& >( S*9"& /.3/X .(W ) )D)K ,'+ L - = Dk *! 99Z 99 ! 3Y99 3* U !99 99@399F&3 FG D ( . .@ ?9$$% 29 3& ,/9"!U 3/. #Q ! ,'+ UAH ) 9X3; S9+ 3/. >( (+ F& [99 . .( RD) R )RW )RR) K99 ,'99+ 99 997G I 39H (+ A+ )'(" ) 38 )cN ) + 9$N (1 1) 9Q )(W\) .7= #$$%& K9F. & >e F! 4 'H ?; ) - S+ @3G9F S*+ 4 U. necator4 K , #] (WW) 3H 9 4. 3H c; $ '. =A+ (+ 4 . ^9- 9F! 39/. _ 39 `9J , 3=U 4. 3H 9 Flag Shoot >9keT ,'9" 9 39 SA % 3/. , 3=U 4 . 3H a S*+ 4 4.3- ? . ^- F! ' '9 ! U. necator 4 3/. - 3=U / ' 3+ ?9 . ^- F! Gb S*+ 3T 4 3/. _ 3 `J ,!9& 39/. 39H 4+ 4 , & , ? N ) 399 ,'99+ 499 N^ 99. 993/. J 399. P99GQ c99; 9; 3 RWD RWDR ` '.3+ S*"& 49+ U 9@ 39F&3 FG 39H 49? 9 >QGT @9; 9q 9 ?9 ,'9+ 4 9+ 39/. ,'9AT @; .' N- )(W) .9 )(WD) 4F. N )(W1 Wd) A=U 3/. Z Vitis vinifera L. 49.3- 9"A" ) 9IN )* ) /FT . 39 ,'9+ L 9- (W) = 9 (R)9* )() .7= .' F . ^9- F! L 4 '. 3 '$ ( g) ? e- f 9$ T 9(.U `9J ?9T = KF. @3G h$; U. necator DRAM D m =!.n+ . .l @39F&3 FG 3Y9 3* U S*9+ 49 c9; ?9 49 ' + *9FT >9; , 3=U 3/. Z RWDR ,( ?9 i=9 3Y9 3* U 9.! 49. 3H .' . ^- F! !9" #$$%& ,/ F 4 rG . F& ! @3F&3 FG 9* V9$G& 3 ,'+ ,' (R1) RDg ` S c; U9AH 9X3; s'(9" , H d 3G ; *AG- j37 3 (R1) RDg 3=- 2 3& 3Y 3* U 3/. ,'9+ ,' . #5; 4 Z ? .'.'+ 3Y99 3* U 99$. (W) RgCd `399kU (WD) RgW !9" #$$%& 3/. q f SAG 4F . ^99- 99F! he99; 499 998.U! . 99 99A '99=3& ! 9 f9 ! Rd 9 .'.'9+ U ('(") 39 ,'+ #] 3/. _ 3 `J 4 . 3H U . necator !99 99 RWD ,99 499A. 99& RWDR ,99( c9; ? l3* U $. 4 '. 3 '$ ! ?$$% 39 L M9 @3F&3 FG ? ! 3Y 3* U 9A( 9$. KF. m )> U . ^- F! 1 ... Uncinula necator(Schw.)Burr. !&' () * )+() ,- . d ! t9 '.'9+ 4 9/. 9- .9 4H U 999AH 999 .'999+ 999-, ! '999. () 999- 99@v V599 3Y99 3* U 99.! 499. 3H 99 KT99 9- 495; , A ST c; @3F&3 FG 3G ?& s SN3 v `3G% U x. ! t 3* * 9*& 95$ _U 3 % * . R\\ ?= 4"+ f .'.'+ -,! '. _U tY9 .'.39+ !U _U 9@39F&3 FG 9& '.'+ , ! tY9 9 9G ^9N 5; f= ! 4"+ S 1()A .!. * )+() !;* K - .'!5! .Z 9@ 3F&3 FG UAH * D ^N 5; f= 9 f9 ,9 RWD , K"( ' & RWDR ,( ! U AH f = V5 ! l3* U )'+ , 3T M9 / 9F*+ l 3* U , 3 K$ 'T W\ '& 4 . / 5; ^Q fF V5 L ? M .KN- @8. (Rd) 3 - L f f "& L3 S . '.'+ !3& l3* U 49 / 9F*+ 9 @39F&3 FG '9T W\ , 3 K$ 9. f "& 4 S '+ 4& ^Q fF f9 @3F&3 FG 'T f KFI. .'+ - ,! '. f*7& 5$ _U =G 4 '+ , ; Nb ^Q Z @v f9 V5 \ ?k3& 'b \/\ J 5$ _U ,5; Nb ^Q 3 ,'+ t n+ K3 8 .'9+ , 9; U S9v f '+ 4 + ^- 3* * 9 K9 _39X `e9 '9T 49 f "& S ! 4 @-\/\\R K; ` 8 ! & f @v ? _3X `e ? 3* * @v f '+ 4 + ^- 4.' A `3 7 f '+ 4 + ^- K+ ; =33 f V59 ,'+ ! 3Y 3* U & '+ , ; K v9 U y39m >9p& j KJ KF. 3& 4 =33 9 9 9 f "& L3 4 tY )'.3+ U AH (.U , 39 ! ?9 9] & "N Sv 4 N ? f9A 9 9 f 9"& 9 w '.'+ , ; > ! 9& 29 3& "N ? P:. 3+ 4 F*+ l3* U ! t 9 3Y9 3* U ! ' - 'F >GN ?9 .'9+ K9] -, ! '. @3F&3 FG W\ V59 9U x9. ! ' ) - . 4H KT d z9&) 9A& f9 9 N 9:& he9 y9 P=; !U .'+ -,! '. 3G ?& sSN3 v `3G% @v .'+ @8. (@3F&3 FG ) *& W\ ( -,! '. DRAM G!7 m =!.n+ . .# DRAM oE( PM /) (E .'!5! .T 9- L M9 9 9 f 4 7 .! Sb 3N 9*A. 9` 39G% 3Y9 3* U =3G @3& -,! '. '9T W\ )RWD 9 2 & RWDR ,( ! (RW) 3 L ?99 M99 .'99 + @998. ( d) `'99 L M99 9 .9 95; 9 ^9Q 9 fF @3F&3 FG 99 ! 3Y99 3* U f99 99 3Y99 3* U 99 # 99{& .'.'999+ t999 9995$ _U 999 tY999 '.'999+ 4 999+ ^- K9FI. .'+ - ,! '. l3* U W\ '& 4 l3* U 3999 L '999.A !999. 3999 999@3999F&3 FG 99 995$ _U 3Y99 3* U =3A99 e n99+ 4995$. !999 S(999F& 999 .'.'999+ 4999(& () 999- >'9 '9G v39 f9 \/D )\/1 )\/d )\/ 9 `3G% 49 & SY=* f fA l3* U , 3 3Y 3* U S9b 3N 9-, ! '9. 9@ 3F&3 FG UAH ! f 9"& '9T L3 f F ? '.'+ 4 F*+ U 9 3Y9 3* U =3G9 # {& . f , .! @v f9 S9; L '9.A '.'+ 4& *& 3T 4 )79b >'9 'G `3G% 3Y 3* U W\ -,! '. 9 f "& .'+ , ; f "& 4 3* * C "#$% / ! / / DRAM ;! . .$ 9 3Y 3* U |T `3 -,! '. v3 f \/ 9A S9T c9; 3Y9 3* U 9 ! A #] 99 R\\\ 99A.Z99 399. l3*99 * ! , 7 99 9 f "& 4 4 *FT >; 3/. Z KFI. .'+ 4 % `39G% 9 49$; #'9 4 _I . '. 3 . g 5; ~39N 9 Z .'.'+ .37T'm 'b f >' KG3Y D ( * DRAM D !] .'!5! .a 9 =G 3 % f "& f S Z@ S% ! .'R! 1(* ! * 6(! 39G KA9F; R\\9 , 9A 'b R/ -U s _U 2% f9RWD ,9 29 & , '7 4A. ! RWDR ` K9 _39X `e9 9(X U V59 he; 4 )`! 'A 9; *AG- #$$%& ,/ F 3 -3Y ,/ K9" 29% V59 Z ? 4GbN 8 3p 4 S 9 9a9 ?9 9X3; –'(9" , H d 3G 9. 9 f 9"& 9 _ .'+ , N 3 ,'+ 4& '9. 3 ,'9+ 4 + Q X #3b 4 4 *FT >; 3/. 5$ _U P* . 4& S Nb ^Q Z 1\ n97& U 49.! w*9+ 49 39 49 '9 - P:. f9 4 ^Q fF (X tY '+ t S >9p& 39 ,/ 9 . K9+ ; ?9! V5 ! . 39 ,'9+ , 9; 9A S9T c; @3F&3 FG 'T \ )49 7 .'+ ,/ 4$; 3 = , 4 ' - '9+ 4 + ^- f "& _ 3H3 Nb ^Q (9A8J %R\) ?N 9 , A ?=! U V5 4 3. f ' + 4 + ^- U _ 4. #3b 4 f "& 4 ~39N 39. .' - 4& ,/ KT 3 ,'+ 4 "QU 'F K3 €7J3p 4 >GN f "& 4= w @v ,'9+ ,'9 49. ! S9b 3N 49 •3& 4 7 )9A& N :& he y f P=; !U ? .'+ S9N3 v s3G ?& `3G% '+ , ; 3* * A A& (@3F&3 FG ' ^Q f F ) '+ ! 9 ,'9+ !U 3Y9 3* U '9& Ux. 9*& 9 f9 (@3F&3 FG ^Q f F ) ! .'+ LA+ d\\ A.Z .(RR)KN- @8. ( f "&) /m * )+() - K ^ .'!5! .X N; ( , 39=U 9 49+ )9 Z V5 3H3 l3* U 949+ %59 6'79 39•J 49 O39 >keT ! , 7 9 RWD , ? N & RWDR ,( ! 6 V5 '9. 3 ,'9+! S9:N ` 49 39/. 9- 94. 3H U 99AH 99 f99 ,99 (D) 99*A 99F&3 L ,'9 K%& (+ 3 @ v 97 #39b 49 l3*9 U 'T \ .'.'+ 9 9G #39b 4 3(‚ @/ ) A ? >keT .'". `39G% 9(.U '. ,'.! KG; ,'+ S$ l3* * 9Z9 49+J P9GQ 9 Z V5 8 499 $ ; ! '99 _U # 99 ?99 3k3GN '99b \/\R )' 9+ `3 . CsW\ 'J 4 3H .H 4+ 99- L M99 99 t.99 3k3GN l3*99 * K99%& 3Y 3* U ! . '+ ,' , K"( _39F% ,'9.! 9 l3*9 U .'9+ (RC) *A ,9( 9- .9 49H n+ K3 ?99+ 99 x99. U 3Y99 3* U %\ 499 '.'99+99 l3*99 * @v 99 99 .99 '99T 1 '99J RWDR .' ".