4)L ©Zoologische Staatssammlung München;download: http://www.biodiversitylibrary.org/; www.biologiezentrum.at \ 57Cls ^üniilTifiiiiiiii] miffit^iilllllliiiiülr SPIXIANA. |H, Zeitschrift für Zoologie ,ni|^S SPIXIANA • Supplement 24 • München, 15. Oktober 1997 • ISSN 0177-7424 • ISBN 3-931516-24-5 ©Zoologische Staatssammlung München;download: http://www.biodiversitylibrary.org/; www.biologiezentrum.at ©Zoologische Staatssammlung München;download: http://www.biodiversitylibrary.org/; www.biologiezentrum.at The Flesh-Flies of Central Europe (Insecta, Diptera, Sarcophagidae) Dalibor Povolny & Yuriy Verves ©Zoologische Staatssammlung München;download:SPIXIAMA http://www.biodiversitylibrary.org/; www.biologiezentrum.at ZEITSCHRIFT FÜR ZOOLOGIE herausgegeben von der ZOOLOGISCHEN STAATSSAMMLUNG MÜNCHEN SPIXIANA bringt Originalarbeiten aus dem Gesamtgebiet der Zoologischen Systematik mit Schwerpunkten in Morphologie, Phylogenie, Tiergeographie und Ökologie. Manuskripte werden in Deutsch, Englisch oder Französisch angenommen. Pro Jahr erscheint ein Band zu drei Heften. Umfangreiche Beiträge können in Supplementbänden herausgegeben werden. SPIXIANA publishes original papers on Zoological Systematics, with emphasis on Morphology, Phylogeny, Zoogeography and Ecology. Manuscripts will be accepted in German, English or French. A volume of three issues will be published annually. Extensive contributions may be edited in Supplement volumes. Redaktion - Editor-in-chief M. BAEHR Fotoarbeiten: M. MÜLLER Manuskripte, Korrekturen und Besprechungsexemplare Manuscripts, galley proofs, commentaries and review sind zu senden an die copies of books should be addressed to Redaktion SPIXIANA ZOOLOGISCHE STAATSSAMMLUNG MÜNCHEN Münchhausenstraße 21, D-81247 München Tel. (089) 8107-0 - Fax (089) 8107-300 SPIXIANA - Journal of Zoology published by The State Zoological Collections München Die Deutsche Bibliothek - CIP-Einheitsaufnahme [Spixiana / Supplement] Spixiana : Zeitschrift für Zoologie / hrsg. von der Zoologischen Staatssammlung, München. - Supplement. München : Pfeil Früher Schriftenreihe Reihe Supplement zu: Spixiana 24. Povolny, Dalibor: The flesh flies of central Europe (Insecta, Diptera, Sarcophagidae). - 1997 Povolny, Dalibor: The flesh flies of central Europe (Insecta, Diptera, Sarcophagidae) - / Dalibor Povolny & Yuriy Verves. München : Pfeil, 1997 (Spixiana : Supplement ; 24) ISBN 3-931516-24-5 Gedruckt mit Unterstützung des Förderungs- und Beihilfefonds Wissenschaft der VG WORT Copyright © 1997 by Verlag Dr. Friedrich Pfeil, München Alle Rechte vorbehalten - All rights reserved. No part of this publication may be reproduced, stored in a retrieval System, or transmitted in any form or by any means, electronic, mechanical, photocopying or othera/ise, without the prior permission of the Copyright owner. Applications for such permission, with a Statement of the purpose and extent of the reproduction, should be addressed to the Publisher, Verlag Dr. Friedrich Pfeil, P.O. Box 65 00 86, D-81214 München, Germany. Satz und Litho: Verlag Dr. Friedrich Pfeil, München Druck: grafik -i- druck GmbH Peter Pöllinger, München Buchbinder: Thomas, Augsburg ISSN 0177-7424 - ISBN 3-931516-24-5 Printed in Germany - Gedruckt auf alterungsbeständigem Papier - Verlag Dr. Friedrich Pfeil, P.O. Box 65 00 86, D-81214 München, Germany Tel. (089) 74 28 27 - Fax (089) 72 42 772 - E-Mail [email protected] ©Zoologische Staatssammlung München;download: http://www.biodiversitylibrary.org/; www.biologiezentrum.at SPIXIANA Zeitschrift für Zoologie Supplement 24 The Flesh-Flies of Central Europe (Insecta, Diptera, Sarcophagidae) Dalibor Povolny & Yuriy Verves Gedruckt mit Unterstützung des Förderungs- und Beihilfefonds Wissenschaft der VG WORT Verlag Dr. Friedrich Pfeil • München, 15. Oktober 1997 • ISSN 0177-7424 • ISBN 3-931516-24-5 ©Zoologische Staatssammlung München;download: http://www.biodiversitylibrary.org/; www.biologiezentrum.at iRi» ©Zoologische Staatssammlung München;download: http://www.biodiversitylibrary.org/; www.biologiezentrum.at SPIXIANA Supplement 24 1-260 München, 15. 10. 1997 ISSN 0341-8391 ISBN 3-931516-24-5 The Flesh-Flies of Central Europe (Insecta, Diptera, Sarcophagidae) Dalibor Povolny & Yuriy Verves Povolny, D. & Y. Verves (1997): The Flesh-Hies of Central Europe (Insecta, Diptera, Sarcophagidae). - Spixiana Suppl. 24: 1-260 The present publication is the first attempt to review the central European taxa of the family Sarcophagidae or flesh-flies. More than 150 species of the family are known to occur in this area, a number of which is representative for central Europe, and no essential changes in number of taxa are to be expected in future. The paper comprises Information on nomenclature and synonymy, Classification and phylog- eny, morphology, relation to hosts, ethology of adults, flesh-fly taxocenoses, foren- sic importance of flesh-flies and, finally, a review of all taxa distributed in the individual countries of central Europe. Chapters on the flesh-fly taxocenoses and on the forensic importance of the flesh-flies are published for the first time in Sarcophagidae literature. Most important morphological characters of adults in- cluding male and female genitalia are illustrated. Prof. Dr. Dalibor Povolny, Department of Zoology and Apiculture, Mendel University of Agriculture and Forestry, 639 00 Brno, Czech Republic. Prof. Dr. Yuriy G. Verves, Department of Zoology, Shevtshenko University, Volodimirskaya ul. 64, 252017 Kiev, Ukrainian Republic. Contents Introduction 8 Nomenclature and Synonymy 9 Classification and Phylogeny 9 Morphology 9 6 terminalia 16 9 terminalia 32 Relations of Sarcophagidae to their hosts 33 Appendix I 37 The hilltopping in flesh-flies 38 The most important flesh-fly taxocenoses of central Europe 42 ©Zoologische Staatssammlung München;download: http://www.biodiversitylibrary.org/; www.biologiezentrum.at Extrazonal flesh-fly taxocenoses of central Europe 46 Forensic importance 47 Review of Central European Sarcophagidae 49 Acknowledgements 52 Key to the subfamilies of Sarcophagidae 53 Subfamily Macronychiinae 53 Genus Mncroin/chia Rondani, 1859 53 Subfamily Miltogrammatinae Brauer & Bergenstamm, 1889 58 Tribe Miltogrammatini B. B 60 Subtribe Senotainiina Rohdendorf, 1930 60 Genus Soiotaiiüa Macquart 1844 60 Subtribe Pterellina Rohdendorf, 1967 64 Genus Pwtomiltogramma Townsend, 1916 65 Genus Pteniln Robineau-Desvoidy, 1963 66 Subtribe Miltogrammatina Brauer & Bergenstamm, 1889 70 Genus Anacanihoihecum Rohdendorf, 1930 70 Genus Cyliiidrothecum Rohdendorf, 1930 71 Genus MiUognvnma Meigen, 1803 72 Genus Miltogmiumatidium Rohdendorf, 1930 78 Subtribe Apodacrina Rohdendorf, 1967 80 Genus Apodacra Macquart, 1854 80 Tribe Amobiini Townsend, 1918 81 Genus Aniobia Robineau-Desvoidy, 1830 82 Tribe Phyllotelini Rohdendorf, 1935 85 Subtribe Metopodiina Rohdendorf, 1967 85 Genus Metopodia Brauer & Bergenstamm, 1891 85 Subtribe Phyllotelina Rohdendorf, 1935 86 Genus PJn/lloteks Loew, 1844 87 Tribe Oebaliini Rohdendorf, 1967 88 Subtribe Oebaliina Rohdendorf, 1967 88 Genus Oebalia Robineau-Desvoidy, 1863 88 Genus Ptychoneura Brauer & Bergenstamm, 1889 91 Tribe Metopiini Townsend, 1908 93 Subtribe Taxigrammatina Rohdendorf, 1967 93 Genus Hilnrella Rondani, 1856 93 Genus Parngiisia Schiner, 1861 96 Genus Taxigiwiima Perris, 1852 97 Subtribe Metopiina Townsend, 1908 98 Genus Mctopia Meigen, 1803 98 Subtribe Mesomelaenina Verves, 1989 106 Genus Mesomelaena Rondani, 1859 106 Subtribe Phrosinellina Verves, 1989 107 Genus PhrosiiicUa Robineau-Desvoidy, 1863 107 Subtribe Sphenometopiina Verves, 1989 109 Genus Sphenomdopa Townsend, 1902 109 Subfamily Paramacronychiinae 111 Tribe Helicoboscini Verves, 1980 112 Genus Eurychaeta Brauer & Bergenstamm, 1891 112 ©Zoologische Staatssammlung München;download: http://www.biodiversitylibrary.org/; www.biologiezentrum.at Tribe Paramacronychiini B. B 115 Subtribe Agriina Enderlein, 1928 116 Genus Agria Robineau-Desvoidy, 1830 116 Genus Angiometopa Brauer & Bergenstamm, 1889 120 Genus Brachicoma Rondani, 1856 120 Subtribe Nyctiina Enderlein, 1928 122 Genus Nydia Robineau-Desvoidy, 1830 123 Subtribe Paramacronychiina Brauer & Bergenstamm, 1889 124 Genus Paramacromjchin Brauer & Bergenstamm, 1889 124 Subtribe Wohlfahrtiina Rohdendorf, 1928 127 Genus Sarcophila Rondani, 1856 127 Genus Wohlfahrtin Brauer & Bergenstamm, 1889 130 Subfamily Sarcophaginae Macquart, 1835 135 Tribe Protodexiini Townsend, 1912Key to the genera of Protodexiini 136 Genus Blaesoxipha Low, 1861 137 Genus Servaisia Robineau-Desvoidy, 1863 143 Genus Tephroim/ia Brauer & Bergenstamm, 1891 145 Tribe Raviniini Rohdendorf, 1937 146 Genus Ravinia Robineau-Desvoidy, 1863 146 Tribe Johnsoniini Rohdendorf, 1967 147 Subtribe Sarcotachinellina Verves, 1988 148 Genus Sarcohichiuella Townsend, 1892 148 Tribe Sarcophagini Macquart, 1835 149 Subtribe Helicophagellina 151 Genus Helicophagella Enderlein, 1928 151 Subtribe Heteronychiina 158 Genus Discachaeta Enderlein, 1928 158 Genus Heterom/clüa Brauer & Begrenstanim, 1889 161 Subtribe Phallanthina Rohdendorf, 1965 185 Genus Arachnidomyia Townsend, 1934 185 Genus Ascelotella Enderlein, 1928 186 Genus Bellieriomima Rohdendorf, 1937 188 Genus Krameromyia Verves, 1982 189 Genus PandeUeana Rohdendorf, 1937 190 Genus Pierretia Robineau-Desvoidy, 1863 192 Genus Thyrsocuemn Enderlein, 1928 199 Subtribe Parasarcophagina Rohdendorf, 1965 201 Genus Bercaea Robineau-Desvoidy, 1863 201 Genus Liosarcophaga Enderlein, 1928 204 Genus Liopygia Enderlein, 1928 214 Genus Parasarcophaga Johnston & Tiegs, 1921