IUB No. Enzyme Name Source Approved in EC 1,4-Alpha-Glucan Bacillus Subtilis JECFA, Denmark, France, Brazil, 2.4.1.18 Branching Enzyme USA (GRAS Notice No

<ul style="display: flex;"><li style="flex:1">IUB No. </li><li style="flex:1">Enzyme name </li><li style="flex:1">Source </li><li style="flex:1">Approved in </li></ul>EC 2.4.1.18 1,4-alpha-glucan branching enzyme Bacillus subtilis JECFA, Denmark, France, Brazil, USA (GRAS Notice No. GRN 00274) <ul style="display: flex;"><li style="flex:1">Alpha-acetolactate Bacillus subtilis expressed in </li><li style="flex:1">USA 21 CFR 173.115 </li></ul>decarboxylase Bacillus brevis EC 3.2.1.1 Alpha amylase Aspergillus niger Australia/New Zealand, Canada, France, Brazil, Mexico, USA (GRN 89) Aspergillus oryzae Australia/New Zealand, Canada, France, Brazil, Mexico, USA (GRN 90) Bacillus amyloliquefaciens Bacillus licheniformis Australia and New Zealand, Canada, Mexico, Brazil, USA 21 CFR 184.1148 Belgium, France, China, Japan, Denmark, Australia and New Zealand, Canada, Mexico, Brazil Bacillus licheniformis expressed Australia/New Zealand, Canada, in Bacillus licheniformis France, Brazil, Denmark, Mexico, USA GRAS Notice No. GRN 000079, JECFA Bacillus licheniformis and Bacillus amyloliquefaciens expressed in Bacillus licheniformis GRAS Notice No. GRN 00022, Brazil, Mexico Bacillus amyloliquefaciens expressed in Bacillus licheniformis Brazil, Denmark, France Bacillus megaterium expressed JECFA, Canada, Brazil, Mexico in Bacillus subtilis Bacillus stearothermophilus JECFA, Canada, Brazil, Mexico, USA 21 CFR 184.1012 Bacillus stearothermophilus expressed in Bacillus licheniformis Australia/New Zealand, Canada, France, Brazil, Denmark, Mexico, Japan, GRAS Notice No. GRN 000024 Bacillus stearothermophilus expressed in Bacillus subtilis Australia/New Zealand, Canada, France, Brazil, Denmark, Mexico Bacillus stearothermophilus (Geobacillus stearothermophilus) Australia/New Zealand, Canada, France, Europa, Brazil, Mexico, USA (GRN 594) Bacillus subtilis Australia and New Zealand, Canada, Mexico, Brazil, USA 21 CFR 184.1148 Rhizopus delemar Rhizopus oryzae Brazil Brazil, Mexico, USA GRAS Notice No. GRN 000090 Thermoccocales expressed in Brazil, USA (GRN126) Pseudomonas fluorecens Rhizomucor pusillus expressed in Aspergillus niger Brazil, Denmark, France, Mexico, Australia and New Zealand. Thermoactinomyces vulgaris expressed in B. subtilis Alicyclobacillus pohliae expressed in Bacillus subtillis Canada, Denmark, Mexico USA (GRAS Notice No. GRN 604) France Cytophaga sp. expressed in Bacillus licheniformis Aspergillus kawachii expressed in Trichoderma reesei Barley malt, malted cereals Pseudomonas fluorecens Trichoderma reesei USA 21 CFR 184.1443a Canada, France Denmark, France EC 4.1.1.5 Alfa-acetolactate decarboxylase Bacillus brevis expressed in Bacillus subtilis Australia/New Zealand, Canada, France, Europe, JECFA, Brazil, Denmark Bacillus brevis expressed in Bacillus licheniformis Brazil, Denmark, France, Mexico, Canada, GRAS Notice No. GRN 00587, Bacillus subtilis Australia, Denmark Bacillus amyloliquefaciens Australia Denmark Bacillus licheniformis Aspergillus niger EC 3.2.1.1 Alphagalactosidase Brazil, Mexico, New Zealand, Australia Mortierella vinacea Brazil, Mexico, USA 21 CFR 173.145 Saccharomyces carlsbergensis Brazil, Mexico EC 3.2.1.133 Maltogenic amylase Bacillus stearothermophilus expressed in Bacillus licheniformis Brazil, Australia, Canada, China, France, JECFA, Japan, Denmark, Mexico, USA (GRN 751) Bacillus stearothermophilus expressed in Bacillus subtilis Brazil, Mexico, Canada, China, France, JECFA, USA (GRN 746 y GRN 751) Pseudomonas stutzeri expressed in Bacillus licheniformis Australia and New Zealand, Canada, Mexico, Brazil Geobacillus stearothermophilus Canada, China, France, JECFA, expressed in Bacillus subtilis Aspergillus niger var. awamori Aspergillus niger Japan, Australia, Denmark, Brazil, Mexico, USA (GRN 751) <ul style="display: flex;"><li style="flex:1">Canada, Brazil </li><li style="flex:1">EC 3.2.1.3 Amyloglucosidase </li></ul>or Glucoamylase Australia and New Zealand, Canada, France, Brazil, Mexico, Japan, Denmark, USA (GRN 89) Aspergillus niger expressed in Aspergillus niger Brazil, Mexico, Australia and New Zealand, Canada, Mexico, USA (GRN 751) Aspergillus oryzae Rhizopus arrhizus JECFA, Australia and New Zealand, Canada, France, Brazil, Mexico, USA (GRN 90) Australia and New Zealand, Canada, Brazil, Mexico Rhizopus delemar Rhizopus niveus Australia and New Zealand, Brazil Mexico, Australia and New Zealand, Brazil, USA 21 CFR 173.110 Rhizopus oryzae Mexico, Australia and New Zealand, Brazil, USA (GRN 90) Talaromyces emersonii expressed in Aspergillus niger Brazil, Canada, China, Denmark, France, Mexico Trichoderma reesei Brazil Trametes cingulata expressed in Denmark, France Aspergillus niger Trichoderma reesei expressed Brazil, USA (GRN 372), JECFA in Trichoderma reesei Gloeophyllum trebeum Brazil, Denmark, France, Mexico expressed in Aspergillus niger Rhizopus delemar var. multiplicisporus Aspergillus fumigatus expressed Mexico in Trichoderma reesei Fusarium verticilloides expressed in Trichoderma reesei Mexico Penicillium oxalicum expressed in Aspergillus niger Denmark, Mexico, Canada Glucoamylase de Aspergillus niger 41SAM2-54 EC 2.4.1.25 Amylomaltase Thermus thermophilus expressed in Bacillus amyloliquefaciens Australia/New Zealand, Canada, USA (GRN 507) Aspergillus oryzae USA (GRAS Notice GRN No. 274) USA (GRN 274) Aspergillus oryzae Bacillus amyloliquefaciens Australia Aminoglycoside 3′- Eschericha coli expressed in phosphotransferas Eschericha coli e II USA (21 CFR 173.170) Eschericha coli expressed in Eschericha coli USA 21 CFR 173.170 <ul style="display: flex;"><li style="flex:1">EC </li><li style="flex:1">Aminopeptidase </li></ul>Lactococcus lactis USA GRAS 3445, Brazil, <ul style="display: flex;"><li style="flex:1">3.4.11.1 </li><li style="flex:1">Australia/New Zealand, Canada </li></ul><ul style="display: flex;"><li style="flex:1">Aspergillus oryzae </li><li style="flex:1">Australia/New Zealand, China, </li></ul>France, Denmark, USA (GRN 90) Lactococcus lactis (Streptococcus lactis) Aspergillus melleus USA 21 CFR 184.1985 China, Denmark, France, Japan Japan EC 3.5.4.6 AMP deaminase Streptomyces murinus EC 3.2.1.55 Arabinofuranosidas Talaromyces pinophilus Denmark, Mexico, USA (GRN <ul style="display: flex;"><li style="flex:1">000680) </li><li style="flex:1">e</li></ul>expressed in Trichoderma reesei Aspergillus niger Australia, China, France, JECFA EC 3.5.1.1 Asparaginase Aspergillus niger expressed in Aspergillus niger Australia/New Zealand, Brazil, Canada, China, Denmark, France, JECFA, Mexico, USA (GRN 214) Aspergillus oryzae expressed in Australia/New Zealand Brazil, Aspergillus oryzae Mexico, Europe, Denmark, France, Canada, China, JECFA, Mexico, USA (GRN 201) Pyrococcus furiosus expressed in Bacillus subtilis Australia/New Zealand Brazil, Mexico, Europe, Denmark, Canada, USA (GRN 476), France Aspergillus niger Australia Aspergillus oryzae JECFA, Denmark, France, Brazil Bacillus subtilis Denmark, France, Brazil <ul style="display: flex;"><li style="flex:1">EC 3.4.23 </li><li style="flex:1">Aspartic protease </li></ul>Cynara cardunculus L. USA (GRAS Notice GRN No. 551) Cynara cardunculus L. Bacillus licheniformis USA (GRN 551) <ul style="display: flex;"><li style="flex:1">Denmark </li><li style="flex:1">EC 3.2.1.2 Beta-amylase </li></ul>Bacillus cereus Brazil Bacillus megaterium Bacillus subtilis Brazil Brazil Bacillus flexus expressed in Bacillus licheniformis Brazil, Mexico, Denmark, France, USA (GRAS) Cebeda 21 CFR 184.1443a EC 3.2.1.6 Beta-glucanase Aspergillus aculeatus Australia/New Zealand, Brazil, Canada, China, Denmark, France, Mexico Aspergillus niger Bacillus subtilis Brazil, Canada, France Brazil, Canada, France Disporotrichum Brazil, Canada, USA (GRN 482) dimorphosphorum Humicola insolens Brazil, Canada, France. USA (GRN 195) Penicillium emersonii Brazil Talaromyces emersonii Brazil, Canada, France, USA (GRN 479), Australia Trichoderma harzianum Trichoderma longibrachiatum Trichoderma reesei Brazil, Canada, France Brazil Brazil, Canada, France, Australia, JECFA Penicillium funiculosum USA (GRN 584) USA (GRN 592) Bacillus subtilis expressed in Bacillus subtilis Rasamsonia composticola Aspergillus niger Beta-glucosidase or cellobiase Brazil Brazil Trichoderma harzianum Trichoderma reesei Apergillus niger Brazil EC 3.2.1.21 E.C. Beta-glucosidase Bromelain Brazil, USA (GRN 750) Ananas comosus USA 21 CFR 184.1024 3.4.22.32 A. bracteatus L. Bacillus licheniformis USA 21 CFR 184.1024 <ul style="display: flex;"><li style="flex:1">Mexico </li><li style="flex:1">EC 3.2.1.3 Carbohydrase </li></ul>Rhizopus oryzae USA (GRAS Notice GRN No. 90, 21 CFR 173.130) Aspergillus niger Rhizopus oryzae USA 21 CFR 173.120 JECFA, Mexico, Brazil, USA (GRAS Notice GRN No. 90) EC 3.2.1.3 Diastase <ul style="display: flex;"><li style="flex:1">3.4.16.5 </li><li style="flex:1">Carboxypeptidase </li><li style="flex:1">Aspergillus niger expressed in Canada, Denmark, France, Mexico, </li></ul>CAspergillus niger Aspergillus niger Russia, USA (GRN 435) Australia/New Zealand, Brazil, Canada, China, France, USA (GRN 89), Mexico EC 1.11.1.6 Catalase Micrococcus lysodeicticus (micrococcus juteus) USA 21 CFR 173.135, Canada, JECFA, Brazil, Mexico Aspergillus niger expressed in Aspergillus niger Australia/New Zealand, USA (GRN 89) Bovine liver Aspergillus niger USA 21 CFR 184.1034 Brazil, Mexico, Australia, New Zeland, Canada, JECFA, USA (GRN 89) EC 3.2.1.4 Cellulase Aspergillus oryzae Brazil, USA (GRN 90) Penicillium funiculosum Rhizopus delemar Brazil, JECFA, USA (GRN 584) Brazil Brazil Brazil Brazil Rhizopus oryzae Sporotrichum dimorphosporum Thielavia terrestris Trichoderma longibrachiatum Brazil, Australia/New Zealand, Canada, China, Denmark, France, Mexico Trichoderma reesei Brazil, Canada, USA 21 CFR 184.1250 Rasamsonia (Talaromyces) emersonii JECFA, USA (GRAS GRN 479) Trichoderma viride Australia <ul style="display: flex;"><li style="flex:1">EC </li><li style="flex:1">Cyclodextrin </li></ul>Bacillus licheniformis Denmark, France <ul style="display: flex;"><li style="flex:1">2.4.1.19 </li><li style="flex:1">glucanotransferase </li></ul>Paenibacillus macerans Australia Thermoanaerobacter sp. expressed in Bacillus licheniformis Arthrobacter globiformis expressed in E. coli K-12 W3110 China, Denmark, France <ul style="display: flex;"><li style="flex:1">USA (GRN 624) </li><li style="flex:1">EC </li></ul>5.1.3.30 D-Allulose 3- epimerase <ul style="display: flex;"><li style="flex:1">EC </li><li style="flex:1">Dextranase </li></ul>Bacillus subtilis Brazil 3.2.1.11 Chaetomium erraticum Brazil, Australia, New Zeland, Brazil, Denmark, USA GRAS GRN 000027 Chaetomium gracile Klebsiella aerogenes Brazil Brazil Penicillium funiculosum Brazil Penicillium lilacinum Brazil Branching enzyme Rhodothermus obamensis Brazil Milk clotting enzymes Endothia parasitica USA 21 CFR 173.150 Bacillus cereus USA 21 CFR 173.150 USA 21 CFR 173.150 USA 21 CFR 173.150 Mucor pusillus Lindt Mucor miehei Cooney et Emerson Aspergillus oryzae Aspergillus niger USA 21 CFR 173.150 <ul style="display: flex;"><li style="flex:1">Brazil </li><li style="flex:1">Esterase </li></ul>Mucor miehei Australia, New Zeland, Brazil, Denmark, USA 21 CFR 173.140 <ul style="display: flex;"><li style="flex:1">Brazil </li><li style="flex:1">EC 3.1.3.8 Phytase </li></ul>Aspergillus niger Aspergillus niger expressed in Aspergillus niger Ficus Brazil, Australia, France, JECFA <ul style="display: flex;"><li style="flex:1">USA 21 CFR 184.1316 </li><li style="flex:1">Ficin </li></ul>EC 2.3.1.43 Phosfatidylcoline sterol O- acyltransferase Aeromonas salmonicida expressed in Bacillus licheniformis Brazil, Australia, GRAS GRN 265 <ul style="display: flex;"><li style="flex:1">EC 3.1.4.1 Phosphodiesterase Leptographium procerum </li><li style="flex:1">Denmark, France, USA (GRN 505) </li></ul>USA (GRAS GRN 574) I EC 3.1.4.11 Phosphoinositide phospholipase C Pseudomonas fluorescens Pseudomonas fluorescens gene extracted from soil sample Pseudomonas sp-62186 expressed in Bacillus licheniformis Denmark, USA (GRAS GRN 728), Canada, Mexico <ul style="display: flex;"><li style="flex:1">EC </li><li style="flex:1">Phospholipase A1 </li><li style="flex:1">Fusarium venenatum expressed Brazil, USA (GRN 142), Australia, </li></ul>3.1.1.32 in Aspergillus oryzae Canada, China, Denmark, France, Mexico Talaromyces leycettanus expressed in Aspergillus niger Streptomyces violaceoruber Denmark, France, USA (GRN 651) Brazil, USA (GRN 145) USA (GRN 524) EC 3.1.1.4 Phospholipase A2 A. nishimurae (ex fumigatus) expressed in T. reesei Aspergillus niger Australia, Canada, China, Denmark, France, Japan S. violaceoruber expressed in Streptomyces violaceoruber USA (GRN 212) USA (GRN 183) Porcine pancreas expressed in Aspergillus niger A. nishimurae (ex fumigatus) expressed in France, USA GRAS #524, Mexico EC 3.1.4.3 Phospholipase C Pichia pastoris Brazil, Argentina, Canada, China, France, JECFA, Mexico, USA (GRAS GRN 204) Bacillus thuringiensis expressed Denmark, USA GRN (689), Canada in Bacillus licheniformis Pseudomonas sp. 62186 expressed in Bacillus licheniformis Canada, Denmark, USA (GRN 728) Pichia pastoris gene extracted from soil sample EC 3.2.1.6 Glucanase Aspergillus niger Canada Canada Bacillus amyloliquefaciens Bacillus subtillis Rasamsonia emersonii (Talaromyces emersonii) Canada Canada Humicola insolens Australia/New Zealand, Brazil, China, Denmark, France, Mexico, Canada, USA (GRN 195) Trichoderma reesei JECFA, Australia e Brazil (longibrachiatum) expressed in Trichoderma reesei (longibrachiatum) Bacillus amyloliquefaciens expressed in Bacillus amyloliquefaciens Australia, China, France Australia, Brazil Geosmithia emersonii Trichoderma reesei <ul style="display: flex;"><li style="flex:1">Australia/New </li><li style="flex:1">Zealand, </li><li style="flex:1">Brazil, </li></ul>Canada, France, JECFA, China, Singapore, South Korea Trichoderma harzianum Brazil, China, JECFA, USA (GRN 149), EC 3.2.1.39 Glucanase Streptomyces violaceoruber expressed in Streptomyces violaceoruber JECFA, USA (GRAS GRN 728), Brazil, Japan <ul style="display: flex;"><li style="flex:1">EC </li><li style="flex:1">Glucanase (beta) </li></ul>Disporotrichum Canada 3.2.1.58 dimorphosporum EC 3.2.1.6 endo-1,3(4)-β- T. reesei expressed in T. reesei France, Denmark, Australia, USA <ul style="display: flex;"><li style="flex:1">glucanase </li><li style="flex:1">21 CFR </li></ul><ul style="display: flex;"><li style="flex:1">EC 5.3.1.5 Glucose isomerase Actinoplanes missourienses </li><li style="flex:1">Autralia, New Zeland, Canada, </li></ul>JECFA, France, Brazil, Mexico Bacillus coagulans Autralia, New Zeland, Canada, JECFA, France, Brazil, Mexico, USA 21 CFR 184.1372 Brazil, Mexico Microbacterium arborensens Streptomyces albus Brazil Streptomyces murinus Brazil, Australia, New Zealand, Canada, Denmark, France, Mexico Streptomyces olivaceus Brazil, USA 21 CFR 184.1372 Brazil, USA 21 CFR 184.1372 Streptomyces olivochromogenes Streptomyces rubiginosus Streptomyces violaceoniger Brazil, USA 21 CFR 184.1372 Brazil Penicillium chrysogenum Denmark, France expressed in Aspergillus niger Actinoplanes missouriensis Aspergillus niger USA 21 CFR 184.1372 Brazil, China, Australia and New Zealand, USA (GRN 89) Aspergillus oryzae Brazil, Mexico, Denmark, USA GRAS GRN 000106 Aspergillus niger expressed in Aspergillus niger Brazil Brazil Aspergillus niger expressed in Aspergillus oryzae Penicillium amagasakiense Brazil Streptomyces rubiginosus expressed in Streptomyces rubiginosus Mexico <ul style="display: flex;"><li style="flex:1">EC 1.1.3.4 Glucose oxidase </li><li style="flex:1">Australia/New Zealand, Brazil, </li></ul>Canada, China, Denmark, France, Mexico, USA (GRN 89) Aspergillus niger USA (GRN 106) Aspergillus oryzae Aspergillus niger expressed in Aspergillus niger Aspergillus niger expressed in Australia/New Zealand, Brazil, Canada, China, Denmark, France, Mexico, USA (GRN 106) Aspergillus oryzae Penicillium amagasakiense Penicillium chrysogenum Brazil, Denmark, France expressed in Aspergillus niger Penicillium expressed in T. reesei amagasakiense France, Denmark, Canada, USA GRAS #707 <ul style="display: flex;"><li style="flex:1">EC </li><li style="flex:1">Glutaminase </li></ul>Chryseobacterium proteolyticum Brazil, USA (GRAS GRN 000267), <ul style="display: flex;"><li style="flex:1">3.5.1.44 </li><li style="flex:1">France </li></ul>Bacillus licheniformis expressed USA (GRN 774) in Bacillus licheniformis Protein-glutaminase from Chryseobacterium proteolyticum Canada AE-PG EC 3.2.1.32 Hemicellulase endo-1,3-β- xylanase Humicola insolens Australia, Nueva Zelanda Hemicellulase Aspergillus niger Aspergillus oryzae Bacillus subtilis Australia, Nueva Zelanda, Brazil Brazil Australia, Nueva Zelanda, Canada, Brazil Rhizopus delemar Rhizopus oryzae Brazil Brazil Sporotrichum dimorphosporum Trichoderma reesei Brazil <ul style="display: flex;"><li style="flex:1">EC 3.2.1.8 Hemicellulase </li><li style="flex:1">Australia, Nueva Zelanda, Brazil </li></ul>EC 3.2.1.78 Bacillus amyloliquefaciens Bacillus amyloliquefaciens Australia, Nueva Zelanda Australia EC 1.1.3.5 Hexose oxidase EC 3.2.1.7 Inulinase Chondrus crispus expressed in Hansenula polymorpha Aspergillus niger Brazil, JECFA, Mexico Australia, Nueva Zelanda, France, Brazil, China, Mexico, USA (GRN 89) Kluyveromyces fragilis Brazil, Mexico Sporotrichum dimorphosporum Aspergillus niger Brazil, Mexico <ul style="display: flex;"><li style="flex:1">Brazil </li><li style="flex:1">EC </li><li style="flex:1">Invertase </li></ul>3.2.1.26 Bacillus subtilis Brazil Kluyveromyces fragilis Brazil, USA (GRAS Notice GRN No. 88) Saccharomyces carlsbergensis Saccharomyces cerevisiae Brazil Mexico, Brazil, Australia, New Zealand, Canada, Denmark, France, USA (GRAS Notice GRN No. 88), JECFA Protaminobacter rubrum Brazil EC 3.2.1.68 EC 1.10.3.2 EC Isoamylase Laccase Lactase Pseudomonas amyloderamosa GRN No. 85 Myceliphthora thermophila expressed in Aspergillus oryzae No.122) Brazil, USA (GRAS Notice GRN Bacillus subtilis Aspergillus niger Aspergillus oryzae

IUB No. Enzyme Name Source Approved in EC 1,4-Alpha-Glucan Bacillus Subtilis JECFA, Denmark, France, Brazil, 2.4.1.18 Branching Enzyme USA (GRAS Notice No

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