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University Morifilms International 300N.Zeeb Road Ann Arbor, Ml 48106 8305402 Urbanek, Richard Paul ARTHROPOD COMMUNITY STRUCTURE ON STRIP-MINED LANDS IN OHIO The Ohio Stale University PH.D. 1982 University Microfilms International300 N. Zeeb Road, Ann Arbor, MI 48106 PLEASE NOTE: In all cases this material has been filmed in the best possible way from the available copy. Problems encountered with this document have been identified here with a check mark V . 1. Glossy photographs or pages______ 2. Colored illustrations, paper or print______ 3. Photographs with dark background ______ 4. Illustrations are poor copy______ 5. Pages with black marks, not original copy ______ 6. Print shows through as there is text on both sides of page ______ 7. Indistinct, broken or small print on several pages 8. Print exceeds margin requirements ______ 9. Tightly bound copy with print lost in spine______ 10. Computer printout pages with indistinct print i-— 11. P a g e (s)____________ lacking when material received, and not available from school or author. 12. Page(s)____________ seem to be missing in numbering only as text follows. 13. Two p ag es n u m b e re d ____________ . Text follows. 14. Curling and wrinkled p a g e s ______ 15. O ther_____________________________________________________________________________ University Microfilms International ARTHROPOD COMMUNITY STRUCTURE ON STRIP-MINED LANDS IN OHIO DISSERTATION Presented in Partial Fulfillment of the Requirements for the Degree Doctor of Philosophy in the Graduate School of The Ohio State University By Richard Paul Urbanek, B.S., M.A. The Ohio State University 1982 Reading Committee: Approved By Theodore A. Bookhout Tony J. Peterle Gordon R. Stairs Advisor Charles A. Triplehorn Department of Zoology ACKNOWLEDGMENTS I thank my advisor, Dr. T. A. Bookhout, for his constant patience and support throughout the study. I also thank my other committee members, Drs. C. A. Triplehorn, D. J. Horn, T. J. Peterle, and G. R. Stairs, for their valuable comments and suggestions. I wish to thank Consolidation Coal Company, Cadiz, Ohio, for allowing me to work on company property. G. Cybulski, J. Smith, A. Wallace, N. P. Neumann, T. Michael, H. Myers, and C. Cullen were especially helpful. I am grateful to the late C. V. Riley, Kent State University, for his assistance in locating suitable study areas. I thank W. Sagrilla for allowing me to use his old field as a control area. I am grateful to M. Piccin for allowing me to use laboratory facilities at Belmont Technical College, St. Clairsville, Ohio. I also thank J. George of the same institution for his assistance and advice. This project would not have been possible without the many hours of tedious labor performed by numerous assistants. Special thanks are due to M. J. Young and C. M. Lackney. The help of the following persons is also appreciated: J. Angerer, M. E. Bankiewicz, D. Bevill, B. Boehm, J. Bolitho, S. F. Borgwald, J. L. Brown, B. Butterfield, J. Chittenden, D. Coffman, G. Coffman, C. Colvin, M. S. Crowley, G. A. Dahlem, J. Dowdle, J. Eickleberry, K. S. Gilliland, ii C. J. Hafke, N. A. Heaslip, S. M. Hof, N. J. Houston, S. Howes, E. Karr, H. King, J. A. Kinnee, D. Lallathin, M. Lanchman, K. J. McGowan, S. Moreland, D. Morawetz, S. W. Nichols, B. Niemczura, R. Phillips, A. Pollack, D. Reed, D. Romsliak, R. L. Smith, J. R. Stenzel, D. M. Urbanek, D. P. Urbanek, Margaret M. Urbanek, Mary M. Urbanek, M. P. Urbanek, P. M. Urbanek, D. A. Wagner, and L. A. Young. Thanks are due the Ohio Young Adult Conservation Corps for supplying many of the field and laboratory assistants. J. D. Bittner and R. Ward deserve specific recognition. I wish to thank J. R. Stenzel, S. Teraguchi, S. B. White, L. E. Watrous, 0. Wheeler, R. Mitchell, G. Hardy, L. E. Rogers, J. W. Leetham, and the late M. J. Glorioso for their suggestions dealing with methodology. I thank J. R. Schrock and B. A. Hawkins for their comments concerning strip-mine arthropods. I am grateful to D. Fairley, F. S. Ruland, L. D. Vangilder, J. R. Bart, N. Reichenbach, and S. Dynin for statistical consultation. Thanks are due to the Instructional Research and Computer Center, The Ohio State University, for providing computer time and services, and to the Statistics Laboratory, OSU, for supplying consultation. I thank R. L. Stuckey and W. Carr for identification of plant specimens. Most importantly, I wish to thank the specialists who provided arthropod identifications and feeding habits information. Without the contributions of these authorities, whose names follow, this study could not have been accurately conducted: R. T. Allen, D. M. Anderson, F. G. Andrews, G. E. Ball, J. A. Beatty, E. C. Becker, M. W. Boesel, G. W. Byers, D. S. Chandler, J. B. Chapin, K. A. Christiansen, W. E. Clark, G. A. Coovert, J. F. Cornell, Jr., R. E. Crabill, Jr., G. A. Dahlem, C. E. Dasch, D. M. DeLong, D. L. Deonier, the late H. S. Dybas, G. Ekis, T. L. Erwin, F. D. Fee, R. J. Gagne, R. C. Graves, R. D. Hall, T. J. Henry, R. L. Hoffman, P. M. Holeski, D. J. Horn, M. A. Ivie, N. F. Johnson, P. J. Johnson, P. B. Kannowski, K. C. Kim, J. M. Kingsolver, J. E. McPherson, R. M. Miller, R. S. Miller, E. 0. Moore, W. B. Muchmore, L. B. O'Brien, S. B. Peck, A. J. Penniman, R. W. Rings, W. H. Robinson, J. A. Sargent, D. Shpeley, W. E. Steiner, Jr., J. B. Stribling, M. C. Thomas, G. A. Triplehorn, B. D. Valentine, the late P. Vaurie, K. Valley, L. E. Watrous, A. A. Weaver, Q. Wheeler, J. A. Wilcox, W. W. Wirth, C. P. Withrow, S. L. Wood, and R. E. Woodruff. I also wish to acknowledge the assistance of the Insect Identification and Beneficial Insect Introduction Institute, L. Knutson, Chairman, SEA, AR, USDA, Beltsville, Maryland. I am solely responsible for any missortings or inaccurate identifications that might have been made during processing. I thank the Ohio Division of Wildlife, which, through the Ohio Cooperative Wildlife Research Unit, provided most of the funding for this study. I also acknowledge The Ohio State University, which provided me with the following sources of personal support while I was working on the project: University Fellowship, Teaching Associateship (Department of Zoology), Research Associateship (Department of Zoology), Mining and Mineral Resources Research Fellowship, and Presidential Fellowship. Finally, I thank all of the people whom I have forgotten to mention. Their support has been appreciated. This project was a contribution of the Ohio Cooperative Wildlife Research Unit, jointly sponsored by The Ohio State University, Ohio Division of Wildlife, U.S. Fish and Wildlife Service, and Wildlife Management Institute. v VITA 25 November 1953 ................. Born - Belleville, Illinois 1974 ............................ B.S., Southern Illinois Univer­ sity, Carbondale, Illinois 1974-76.......................... Research Assistant, Graduate Fellow, Southern Illinois Uni­ versity, Carbondale, Illinois 1976 ............................ M.A., Southern Illinois University, Carbondale, Illinois 1976-82.......................... Graduate Fellow, Teaching Assoc­ iate, Research Associate, The Ohio State University, Columbus, Ohio PAPERS Urbanek, R. P. 1976. Vertebrate and floral diversity on strip-mined land in Williamson and Saline counties, Illinois. M.A. Thesis, Southern Illinois Univ., Carbondale. 146pp. FIELDS OF STUDY Major Field: Wildlife Ecology Wildlife and Strip-mine Ecology ............. Dr. W. D. Klimstra Wildlife and Strip-mine Ecology ............. Dr. T. A. Bookhout Insect Systematics Dr. C. A. Triplehorn Population Ecology........................... Dr. T.