Bollettino della Societd Paleontologica ltaliana, 44 (l), 2005, 47-53. Mode, Anew speciesof Gibbula(Gastropoda:Trochidae) fi of Killini (north-westernPeloponnesus, Greece) Vittorio GnruLLI,Magda Cnlsct & RosaMpssne V. Garilli, Dipartimento di Geologia e Geodesia,Universitir degli Studi di Palermo, Corso Tr [email protected] M. Crisci, Via TommasoAversa 70, 90145, Palermo,Italy. R. Messina, Via Messina Marina 435c, 90100, Palermo,Italy. KEY WORDS - Gastropoda, Trochidae, Gibbula olympica n. sp., Mediterranean Plet ABSTRACT - A nev,species of the gastropod Gibbula, G. olympica n. sp., is described western Peloponnesus, Greece). Material comes from two sandy-muddy beds (N2 and H6, mollusc assemblages (mainlv trochids, rissoids and ceriths) related to the modern Posidonia (HP).Gibbula olympica n. sp. appears particularly similar to G. spratti (Forbes, 1844) at Aegean Sea and the Island of Malta respectively. G. olympica n. sp. is also compared witl the Mediterranean erea. RIASSUI,{TO - [Una nuova specie di Gibbula (Gastropoda: Trochidae) dal Pleistocer Grecia)] - Gibbula olympica n. sp. d descritta sulla base di circa 160 esemplari, in bt l'originaria colorazione. Il materiale proviene da due orizzonti stratigrafici prevalentem lungo il tratto di costa distante circa 500 m dal piccolo villaggio di Killini (Elea, Pelopont giacciono circa 70 e 20 m sopra un livello pelitico contenente i nannofossili Gephyroca^'sa Igg2) e, pertanto, essi potrebbero essere attribuiti alla parte superiore del sottopiano Si medio-superiore. G. olympica n. sp. mostra notevoli ffinitd con G. spratti (Forbes, , endemiche del Mar Egeo e dell'lsola di Malta. G. olympica n. sp. vien rispettivamente '15 mediterranee ,o, quali presenta una certa ffinitd: le viventi G. racketti (Payraudeau, pliocenica G. distefanoi Crema, 1903. Sulla base del contenuto paleontologico associato alr spp. (tra cui la specie termofita J. ? bullula (Fischer 1877), Alvania spp., Bittium spp. e J specie vivesse in una biocenosi riferibite a quella attuale della Posidonia oceanica (Linnaea calde. INTRODUCTION MATERIAL EXAMINED In the MediterraneanSea the trochid gastropod One hundred sixty specimens,covering complete genus Gibbula Risso, 1826 ex Leach ms. is a common ontogenetic series, of Gibbula olympica n. sp. (type component of littoral environmentsespecially in algal material included) come from two sandy-muddylayers, or phanerogambeds. It is also well representedin the outcropping near the village of Killini. They were uppermostTertiary and Quaternaryof the Mediterranean obtainedby hand-pickingor washing six bulk samples, area. The richest assemblagesare recorded for the each one of about l5 lt., on a sievesbattery (diameter Recent time: at least 30 Mediterraneanliving well- 0.5, l, 2 mm). As a whole, specimensare very well establishedtaxa are recorded(Sabelli et al., 1990);little preservedusually showing the original coloration. more than 20 species are recorded for the The following comparative material was studied: malacologicallyrich Plioceneof Tuscany(Chirli,2004). Gibbula distefanoi Crema, 1903, one specimen,ex In this article, which presentsthe secondresult of a coll. Palazzi,Pliocene of Pietrafitta(Siena, North Italy). wider researchdealing with the study of the mollusc G. nivosaAdams A., 185l, five specimens,ex coll. assemblagescoming from the Plio-Pleistoceneof the Lugli, Recent,Isle of Malta. central Mediteffaneanarea (Garilli,2004), we describe G. racketti (Payraude&u, 1826), forty-three a new speciesof Gibbula, G. olympica n. SP.,based specimens,ex coll. Garilli, Recent,Capo Gallo, Palermo, on specimenscollected from the northern part of the NW Sicily; one specimen,same coll., Recent,Marina sedimentary sequencecropping out along the cliffed di Cinisi, Palerrno. coast between Killini (also transliteratedfrom the G. spratti (Forbes, 1844),eleven specimens,ex coll. modern Greek as Kyllene) and Cape Troupito (Elea, Tenekidis, Recent, Aegean Sea, Goulandris National NW Peloponnesus,Greece, Fig. I ). History Museum (GNHM), Kifissia, Athens, Greece; /SSN 0375-7633 48 Bollettino della SocietaPaleontologica ltaliana,44 (l), 2005 Fig. I - Site,type materialand H6 and N2 layers locations;KI : Killini; TR : CapeTroupito. two specimeJls(GNHM-A 11.62),ex coll. Tenekidis, trochids (mainly Jujubinus spp.), rissoids (mainly Isle of Dilos, Aegean Sea (specimenfigured by Alvania spp.) and ceriths (mainly Bittium spp.). With "Gibbula Tenekidis, 1989, fig. 6 as spratti var. regardto the genusGibbula,, the following specieswere alveolata") and Anavyssos, Saronikos gulf, Aegean collectedfrom N2 and H6 layers:G. ardens(Von Salis, Greece (specimenfigured by Tenekidis, 1989, fig. 6 1793),G. magus(Linnaeus, 17 58), G. fanulum (Gmelin, "Gibbula as spratti typus"; two specimens,eX coll. 179l), and G. guttadauri (Philippi, 1836).Posidonia Delamotte,GNHN, Marpissa,Isle of Paros,Cyclades, remains (both leaves and rhizomes) also abundantly Aegean Greece. occur, mainly at the base of N2 bed. From a G. turbinoides(Deshayes, 1835), ten specimens, paleoclimaticpoint of view, it is noteworthy to remark ex coll. Garilli, Recent,Capo Gallo, Palermo. that both levels contain the trochid Juiubinus ? bullula (Fischer, 1877) (see Ruggieri & Unti, 1988 for the interesting information about its stratigraphical STRATIGRAPHIC AND PALEOECOLOGICAL distribution)which can be considereda thermophilic SETTING speciesrelated to warrn-temperateconditions (Garilli, 1998). LayersN2 and H6 respectivelycrop out at about The sequenceKillini-Troupito is a partof an anticline 70 and 20 m abovea turritellid-richblue-greyish muddy structureoriginated by a diapiric intrusion of Triassic level containing the nannofossils Gephyrocapsasp. 3 halite and gypsum (Christodoulou,1969,l97 l; and Crenalithusasanoi (Sato & Takayama, 1992) (Fig. Hageman, 1976; Underhill, 1988). It was referredby 2), whose Mediteffaneandistributions are respectively Hageman (1976) to the upper part of the Vounargon recordedbetween 0.990 Ma B.P. (uppermostpart of formation. In particular, material comes from two Early Pleistocene)and 0.584 Ma B.P. (Middle yellow-grey sandy-muddybeds, here named N2 and Pleistocene)(Castradori , 1993; Sprovieri, 1993; Di H6 (Fig. 2), laterally changingin a yellow sand with Stefano,1998) and between1.122 Ma B.P.(lower part Lucinomaboreale (Linnaeus , 1767) in life position(H6) of the Early PleistoceneSicilian Substage,sensu Ruggieri or in a yellow, almost sterile calcarenite(N2). Both et al., 1984)and 0.781 Ma B.P.(lower part of Middle layers, inclined along the cliff plane of about 8-9o Pleistocene)(De Kaenel et al., 1999).Consequently an -Late northward, are characterrzedby having a rich shallow upper Sicilian Substageor, more likely, an Middle marine mollusc assemblagerelated to the modern Pleistoceneage could be likely assignedto N2 and H6 Posidoniaoceanica (Linnaeus) Delile, l8l3 biocoenosis layers. (HP, sensu Pdrds& Picard, 1964), being dominated by V.Garilli, M. Crisci, R. Messina- Gibbula olympican. sp. from the MediterraneanPleistt SYSTEMATICS t-rr., il Sandstone Phylum MolluscA Linnaeus,1758 - Mudstone ClassisGnsrnoPoDA Cuvier, 1797 FamiliaTnocutDAE Rafinesque, l8l5 Genus Gibbula Risso. 1826 ex Leach ms. Type speciesTrochus magus Linnaeus, 1758 by subsequentdesignation, Herrmannsen, 1847 . Gibbula olympica n. sp. Gibbula olympica n sp. (Figs. 3a-c; Pl. I , figs. I a-b, 4a-b,5a-b) Paratypes Tvp, material - Holotype (GNHM-52/18), paratypes I (GNHM-5 3l19) and 2 (GNHM-54 120)are housedin uJ the GoulandrisNatural History Museum (GNHM), Kifissia, Athens, Greece.Paratypes 3 (KIGR003) and 4 (KIGR004) arehoused in the Dipartimentodi Geologia z z e Geodesiaof the Universityof Palermo,Italy. Fifteen o unnumberedparatypes are in Palazzi coll. (Modena). - Holotype, height (H) 8.2 ffiffi, IU up to 3o-40m Measurements F ) diameterof last whorl (D) 7.95 mm; paratypel, H 7.5 ffiffi, D 7.4 mm; paratype 2, H 7.3 ffiffi, D 6.85 mm; paratype3, H 7.1 mm, D 6 mm; paratype4, H 8.5 O ffiffi, D 7.8 mm. t Locus typicus Pleistoceneof Killini (Elea, NW Peloponnesus, Greece), Vounargon formation o o Gibbulaolympica n. sp. II ' (accordingto Hageman, 1976).Holotype is from the Holotype yellow-greyish sandy-muddybed (H6) outcropping at F about0.5 Km Eastfrom the villageof Killini (Elea,NW z Peloponnesus,Greece, UTM 34S 5 I I 7938 4199216N, o Fig. I ). All paratypesare from layer N2, similar for lithologyand paleontological contents to the underlying (, a H6 (Fie. 2). t Derivatio nominis G. olvmpica n. sp. is named after the ancient town of Olympia (Peloponnesus, z Greece)where the first Olympic Games took place. uJ f Diagnosis Sturdy, small, conical shell reaching o about 9 mm in height. Protoconchpaucispiral, J consistingof about one whorl. Protoconch/teleoconch demarcationis markedand slightly sinuated.Teleoconch consistsof more or less convex whorls, bearing tr irregular and fine spiral cords separatedby very naffow interspaceswhich are sculpturedby microscopical dots. Body whorl well developed.Aperture is subquadrangular.Outer lip rounded.Inner lip straight. Basemoderately convex crossed by very fine concentric threadsand with a deep, wide umbilicus. Patternof colorationusually consisting of rectangular-rhomboidal Gephyrocapsasp. 3 white spiralling mottles in a reddish background. Crenalifhusasanoi Description - Sturdy,conical shell reaching8.9 mm in height (8.2 mm in holotype). Height/maximum diameterratio (H/D) rangesfrom 0.95 to I . l5 ( 1.03in fir holotype).Protoconch paucispiral, consisting of about l.l smooth whorls (countedaccording to