World Catalogue of Coccinellidae World Catalogue of Coccinellidae Part I - Epilachninae Part I - Epilachninae Andrzej S. Jadwiszczak Piotr W^grzynowicz Part II - Sticholotidinae, Chilocorinae, Coccidulinae KOPIE B 15 Part III - Scymninae dei-Bibliolh.kd.. Dsul.ch8n Entomologiichan Instituts MOnchsbsrg. ^_ - 2ALF e.V. - A h. M53, H Part IV - Coccinellinae Olsztyn 2003 © 2003 MANTIS, Olsztyn Cover design: © Piotr Wejjrzynowicz Epilachna sp. from Colombia, phot. Piotr We_grzynowicz All rights reserved. Apart from any fair dealing for the purposes of private study, research, criticism or review, no part of this publication may be reproduced or trans- mitted, in any form or by any means without written permission from the publisher. Authors' addresses: Andrzej S. Jadwiszczak INTRODUCTION ul. Slowicza 11 11-044 Olsztyn Poland The last comprehensive world-catalogue of the Coccinellldae was published by e-mail: [email protected] Korschefsky — as the respective part of the Coleopterorum Catalogus — 72 years ago. Intensive systematic and faunistic studies on this populär group of beetles, Piotr We_grzynowicz pursued by three generations of entomologists, have extended our knowledge so Muzeum i Instytut Zoologii PAN much, that a new, updated catalogue summarizing the data accumulated since the ul. Wilcza 64 time of Linnaeus became urgently needed. The present publication — including 00-679 Warszawa Poland the subfamily Epilachninae: altogether 1051 species in 22 genera — is the first of e-mail: [email protected] planned four parts. Tribes, genera and species have been an-anged alphabetically within their respective higher taxa. The catalogue has been primarily based on the Publisher's address: data from original publications; unfortunately, however, some papers (marked as MANTIS "not seen") had remained unattainable for us, what made quotations from other ul. Slowicza 11 sources necessary. 11-044 Olsztyn Poland For each species the valid name (in bold), füll list of synonyms, and general e-mail: [email protected] A 2830 geographical distribution is given. Synonyms are listed (retaining original combi- www.mantis.mgt.pl nation and orthography) in chronological order, with type-locality [TL] (if the type-series contained more than a single specimen, all the localities mentioned in the original description), Status (holotype, lectotype or neotype if— and by whom Printed in Poland by Gutgraf, Olsztyn — designated) and (as far as possible) sex of the type-specimen, and sources of these informations. Depositories of the type-material [TD] are given according to the original publications or after the later authors who have studied it. Acronyms Date of publication: 5 III 2003 of the respective institutions follow Amett and Samuelson (1986), or — if not found there — have been created by ourselves. Data of this catalogue were actualized until the preparation of the final manu- ISBN 83-918125-3-7 ,!fn 7'li script, i.e. January 2003. 5 ACRONYMS OF COLLECTIONS IPPG - Institute of Plant Protection, Guangxi Academy of Agricultural Sciences, China ISNB - Institut Royal des Sciences Naturelles de Belgique, Brüssels, Bel- AMNH - Department of Entomology Collection, American Museum of Na- gium tural History, New York, USA IZAS - Insect Collection, Institute of Zoology, Beijing, China ANIC - Australian National Insect Collection, Division of Entomology, KEI - Korean Entomological Institute, Korea. CSIRO, Canberra, Australia KIZAS - Kunming Institute of Zoology, Academia Sinica, Kunming, China ApC _ A. Pütz's collection, Eisenhüttenstadt, Germany KSUC - Department of Entomology Collection, Kansas State University, BM - Brno Museum, Brno, Czech Republic Manhattan, USA BMNH - British Museum (Natural History) (now The Natural History Mu- KUEC - Entomological Laboratory in Faculty of Agriculture, Kyushu Uni- seum), London, United Kingdom versity, Fukuoka, Japan LSUK - Linnean Society, London, United Kingdom BPBM - Department of Entomology Collection, Bernice P. Bishop Museum, LMP - Lisboa Museum, Lisboa, Portugal (precise localization unknown) Honolulu, Hawaii, USA MBBJ - Museum Zoologicum Bogoriense, Bogor, Indonesia CASC - Department of Entomology, California Academy of Sciences, San MCSN - Museo Civico di Storia Naturale "Giacomo Doria", Genoa, Italy Francisco, USA MCZC - Entomology Department, Museum of Comparative Zoology, Ha- CMF - Caen Museum, Caen, France (precise localization unknown) rvard University, Cambridge, USA CNCI - Canadian National Insect Collection, Biosystematics Research MEAU - Missäo de Estudos Agronömicos do Ultramar, Brazil Institute, Ottawa, Canada MEPM - Museo Entomologico del Pontificio Instituto Missioni in Monza, CUMZ - University Museum of Zoology Insect Collection, Cambridge, Uni- Italy ted Kingdom MGFT - Entomologisches Institut, Museum G. Frey, Tutzing, Germany (now DEIC - Deutsches Entomologisches Institut, Eberswalde, Germany in NHMB) DPPN - Department of Plant Protection, Nanjing Agricultural University, MHNG - Museum d'Histoire Naturelle, Geneva, Switzerland Nanjing, China MHNL - Musee Guimee d'Histoire Naturelle de Lyon, Lyon, France DZUP - Departamento de Zoologia, Universidade Federal do Paranä, Cu- MIZPAN - Muzeum i Instytut Zoologii PAN, Warszawa, Poland ritiba, Brazil MLPA - Museo de La Plata, Universidad Nacional de La Plata, La Plata, Argentina DZPU - Department of Zoology, Punjabi University, Patiala, India MMNHO - Municipal Museum of Natural History, Osaka, Japan EIHU - Entomological Institute, Faculty of Agriculture, Hokkaido Univer- MNHN - Museum National d'Histoire Naturelle, Paris, France sity, Sapporo, Japan MRAC - Musee Royal de l'Afrique Centrale, Tervuren, Belgium ELSFC - Entomological Laboratory of the Southwestern Forestry College, MZLU - Museum of Zoology, Lund University, Lund, Sweden Kunming, China MZSP - Museu de Zoologia da Uversidade de Säo Paulo, Säo Paulo, Brazil FDYP - Forest Department, Yunnan Province, China MZUB - Museum of Zoology, University of Bologna, Bologna, Italy (pre- HFC - Coll. H. Fürsch (now in ZSMC — Zoologische Sammlung des Bay- cise localization unknown) erischen Staates, München, Germany) NHMB - Entomology Department, Naturhistorisches Museum, Basel, Swit- HNHM - Zoological Department, Hungarian Natural History Museum, Bu- zerland dapest, Hungary NHMW - Naturhistorisches Museum Wien, Austria ICCM - Section of Insects and Spiders, Carnegie Museum of Natural Hi- NHRS - Naturhistoriska Riksmuseet, Sektionen för entomologi, Stoclcholm, story, Pittsburg, USA Sweden IMLA - Fundacion e Instituto Miguel Lillo, Universidad Nacional de Tu- NMNS - National Museum of Natural Science, Taichung, Taiwan cuman, Tucuman, Argentina NMPC - Narodni Muzeum, Entomologicke odd., Praha, Czech Republic INPC - Division of Entomology, National Pusa Collection of Insects, In- dian Agricultural Research Institute, New Delhi, India 6 7 NSMT - National Science Museum (Natural History), Tokyo, Japan NOMENCLATORIAL ACTS NTUT - Department of Plant Pathology and Entomology, National Taiwan University, Taipei, Taiwan Type-species designation NZSI - National Zoological Collection, Zoological Siirvey of India, Cal- cutta, India Epilachna eckloni Mulsant, 1850b: 866 — designation of type species for subge- OXUM - Hope Entomological Collections, University Museum, Oxford, nus Cleta Mulsant, 1850b: 866. United Kingdom PAS - Academy of Natural Science, Philadelphia, USA New synonym PPID - Plant Protection Research Institute, Chinese Academy of Tropical Epilachna cucurbitae Richards, 1983: 37 is a new synonim of Epilachna vigin- Agricultural Sciences, Danzhou, China tioctopunctata sumbana Bielawski, 1959: 147. QCAZ - Zoological Museum, Pontificia Universidad Catolica del Ecuador, Quito, Ecuador Coccinella vigintioctopunctata Montrouzier, 1857: 75 is a new synonim of Cocci- RMNH - Rijksmuseum van Natuurlijke Historie, Leiden, Netherlands nella vigintioctopunctata Fabricius, 1775: 84 (now in Henosepilachnd). SAMA - South Australian Museum, Adelaide, Australia Henosepilachna beniensis Fürsch, 1991: 246 is a new synonym of Epilachna SAMC - Insect Collection, South African Museum, Cape Town, Republic alternata Sicard, 1930a: 59 (now in Henosepilachna). of South Africa SMFD - Forschunginstitut und Naturmuseum Senckenberg, Frankfurt am New statuses Main, Germany SMG - Stralsund Museum, Germany (precise localization unknown) Henosepilachna vigintioctopunctata sumbana (Bielawski, 1959: 147) — upgra- SMNS - Staatliches Museum für Naturkunde, Stuttgart, Germany ded to the species-rank. SMTD - Staatliches Museum für Tierkunde, Dresden, Germany Epilachna grayi var. zebra Sicard, 1925: 448 — upgraded to the species-rank. TMSA - Transvaal Museum, Pretoria, Republic of South Africa UMBB - Übersee-Museum, Bremen, Germany Replacement names USNM - United States National Museum of Natural History, Entomologi- Epilachna aulisoides ruandica is a replacement name for Epilachna aulisoides cal Collection, Washington, USA UZIU - University ofUppsala, Uppsala, Sweden libera Fürsch, 1963b: 270, nee Epilachna libera Dieke, 1947: 85. UZMH - Zoologiska Muset, Universitets Helsinki, Helsinki, Finland Epilachna boymi is replacement name for Epilachna hausen (Mader, 1930: 182), ZFMK - Zoologisches Forschungsinstitut und Museum "Alexander Koenig", nee Epilachna hausen Weise, 1904a: 57. Bonn, Germany Epilachna chelys is replacement name for Epilachna irritans Fürsch, 1985: 205, ZMAS - Zoological Museum, Academy of Science, Petersburg, Russia nee Epilachna irritans (Mader, 1958c: 5). ZMUB - Museum für Naturkunde, Zentralinstitut