c 1 DFO - L bra y MPO B blto hèque Canadian Special Publication of Fisheries 11111 12038893 and Aquatic Sciences 61 Catalogue of the Living Bivalvia of the Eastern Pacific Ocean: Bering Strait to Cape Horn FR.Bernard Fisheries & Oceans LIBRARY , ratt 16 Igor BIBLIOTHÈQUE Pêc:las & Océans CANADIAN SPECIAL PUBLICATION OF FISHERIES AND AQUATIC SCIENCES 61 Catalogue of the Living Bivalvia of the Eastern Pacific Ocean: Bering Strait to Cape Horn F. R. BERNARD Department of Fisheries and Oceans Pacific Biological Station Nanaimo, British Columbia Canada V9R 5K6 DEPARTMENT OF FISHERIES AND OCEANS Ottawa 1983 Published by Publié par Government of Canada Gouvernement du Canada 1+ Fisheries and Oceans Pêches et Océans Scientific Information Direction de l'information and Publications Branch et des publications scientifiques Ottawa Ki A 0E6 ©Minister of Supply and Services Canada 1983 Available from authorized bookstore agents and other bookstores, or you may send your prepaid order to the Canadian Government Publishing Centre, Supply and Services Canada, Hull, Que. K1A 0S9. Make cheques or money orders payable in Canadian funds to the Receiver General for Canada. A deposit copy of this publication is also available for reference in public libraries across Canada. Canada: $8.00 Catalog No. Fs 41-31/61E Other countries: $9.60 ISBN 0-660-11270-1 ISSN 0706-6481 Price subject to change without notice Ottawa Printed in Canada by Love Printing Services Ltd. Ottawa, Ontario Correct citation for this publication: BERNARD, F.R. 1983. Catalogue of the living Bivalvia of the eastern Map on page viii is reproduced with permission of the Pacific Ocean: Bering Strait to Cape Horn. Can. Spec. Publ. Department of Energy, Mines and Resources. Fish. Aquat. Sci. 61:102 p. To D.B. Quayle '71 Fhariv.s & Oceans LIBRARY. j:Li ' BERLIOTHÈQUE P9i/Cill-,9S OCEiien Contents ABSTRACT/RÉSUMÉ Vii INTRODUCTION Format 1 Origin and classification of Bivalvia 2 Acknowledgments 5 Collections consulted 5 Abbreviations 5 OUTLINE OF CLASSIFICATION • 7 SYSTEMATIC CATALOGUE 9 Notes 67 References cited 71 TAXONOMIC BIBLIOGFtAPHY 73 INDEX 86 Abstract BERNARD, F.R. 1983. Catalogue of the living Bivalvia of the eastern Pacific Ocean: Bering Strait to Cape Horn. Can. Spec. Publ. Fish. Aquat. Sci. 61:102 p. This work consists of a systematic catalogue and primary bibliography of the living Bivalvia of the eastern Pacific Ocean from Bering Strait, Alaska (66°N) to Cape Horn, Tierra del Fuego (60°S). 1308 species from the high intertidal zone to deep waters extending approximately two thousand kilometers offshore are included, representing the total de- scribed fauna. Synonymies for each species and its distribution updated from the recent literature and museum and private collections, are given. Also presented are the thermal range and fossil occurrence in the region, abstracted from the literature and collections. The bibliography lists sources of primary descriptions and replacement names at the specific level only. New species are not proposed, though six replacement names and various changes of suprageneric categories are suggested. The following nomenclatural changes are proposed: Anadara auricula nom. nov. pro Arca auriculata Sowerby, 1833 not Lamarck; Modiolus kurilensis nom. nov. pro Volsella difficilis Kuroda and Habe 1950 not Deshayes; Pecten berryi nom. nov. pro Pecten lunaris Berry 1963 not Renner; Petricola olssoni nom. nov. pro Petricola peruviana Olsson, 1961 not Jay; Semele clydosa nom. nov. pro Amphidesma punctatum Sowerby 1833 not Say; Astartacea superfamily nom. transi.; Axinopsidinae subfamily nov.; Borniinae subfamily nov.; Crassatellacea superfamily emend.; Eryciniinae subfamily nom. transi.; Glycymer- idacea superfamily nov.; Kelliinae subfamily nov.; Lasaeinae subfamily nom. transi.; Montacutinae subfamily nom. transi.; Mysellinae subfamily nov.; Orobitellinae subfamily nov.; Thecodontinae subfamily nov.; Thraciacea superfamily nom. transi.; Thyasirinae subfamily nov. Résumé BERNARD, F.R. 1983. Catalogue of the living Bivalvia of the eastern Pacific Ocean: Bering Strait to Cape Horn. Can. Spec. Publ. Fish. Agnat. Sci. 61:102 p. La présente publication contient un catalogue systématique et une bibliographie de base des Bivalvia existants du Pacifique oriental, depuis le détroit de Béring, en Alaska (66°N), jusqu'au cap Horn, en Terre de Feu (60°S). Elle comprend 1 308 espèces réparties de la zone intertidale supérieure aux eaux profondes jusqu'à environ 2 000 km au large et représentant le total de la faune décrite à ce jour. On y donne, pour chaque espèce, les synonymes et la répartition mise à jour d'après les récentes publications et les collections de musées ou privées. Sont inclus également les extrêmes de température où se trouvent ces espèces, ainsi que l'incidence de fossiles dans la région, ici encore extraits de la littérature et des collections. Dans la bibliographie, on indique les sources de descriptions originelles et les noms de remplacement seulement au niveau spécifique. Aucune nouvelle espèce n'est proposée, bien six noms de remplacement et divers changements de catégories supragénéri- ques soient suggérés. Nous proposons les changements suivants dans la nomenclature : Anadara auricula nom. nov. pro Arca auriculata Sowerby, 1833, non pas Lamarck; Modiolus kurilensis nom. nov. pro Volsella difficilis Kuroda et Habe 1950, non Deshayes; Pecten berryi nom. nov. pro Pecten lunaris Berry 1963, non Römer; Petricola olssoni nom. nov. pro Petricola peruviana Olsson, 1961, non Jay; Semele clydosa nom. nov. pro Amphidesma punctatum Sowerby 1833, non Say; super-famille Astartacea nom, transi.; sous-famille Axinopsidi- nae nov.; sous-famille Borniinae nov.; super-famille Crassatellacea emend.; sous-famille Eryciniinae nom. transi.; super-famille Glycymeridacea nov.; sous-famille Kelliinae nov.; sous-famille Lasaeinae nom, transi.; sous-famille Montacutinae nom, transi.; sous-famille Mysellinae nov.; sous-famille Orobitellinae nov.; sous-famille Thecodontinae nov.; super- famille Thraciacea nom. transi.; sous-famille Thyasirinae nov. vii Os Wand KN. Ha. 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' RePe, ' eUENOS A — r ) NORTH THE WORLD Compiled, Drawn and PrMted by the SURVEYS AND MAPPING BRANCH DEPARTMENT OF ENERGY, MINES AND RESOURCES CANADA 1967 eon der Grinfen Projection Equatorial Seale 1:33,000.000 2000 500 1000 7500 SFATUle 1-s t Commonwealth Countries and Dependencies INTRODUCTION No comprehensive inventory of the bivalve fauna of the changes in concepts or status are given, but do not appear in entire eastern Pacific has appeared, and it is more than 44 the bibliography. bibliography of the years since A. M. Keen's Check list and Genera and subgenera are arranged alphabetically. I have northwestern American marine Mollusca (1937). The avoided listing the nominate genus directly following the present catalogue was started over fifteen years ago to pro- family or subfamily entry as done in the 'Treatise' , as this geography using vide data for a re-interpretation of bivalve merely reflects chronology and is often disruptive of the segregated by depth to resolve new distributional data, current view of natural relationships, which are constantly latitude, effects of submergence of northern species at lower changing as more information accumulates. and submitted to computer techniques for analysis. This work is now in final preparation (Bernard & McKinnel MS), Each species comprises a separate entry, with the current but it appeared useful for taxonomic workers to issue the binomen in heavy type. The original combination and full data, and setting down in juxtaposition the northern and bibliographic reference follows for the species and its junior southern representatives of various groups may stimulate synonyms, arranged chronologically, the status indicated, further study leading to monographie reviews. where necessary, with abbreviated latin terrns. Distribution outside the area of concern is briefly indicated. Occurrence in attempted to include all binomina used regionally, I have the Clipperton, Cocos, or Galapagos Islands fauna is noted, the early literature is replete with extralimital an- though as there appear to be features distinct from the continental alogues. I have built on the important faunistic works of shelf biota, notably the presence of a few endemic taxa and Carpenter (1872), Dall (1921), Keen (1937), and Burch those species with Indo-Pacific affinities. The final line of for the northern fauna; Olsson (1961) and Keen (1944-46) each entry includes the geographic range to nearest degree of (1971) for tropical America; and Carcelles & Williamson latitude and the mid-point of geographical distribution. The and Osorio & Bahamonde (1970) for the southern (1951) bathymetric range is in meters,