22 3 267 306 Tanasevitch for Inet.P65

22 3 267 306 Tanasevitch for Inet.P65

Arthropoda Selecta 22(3): 267306 © ARTHROPODA SELECTA, 2013 The linyphiid spiders of the Altais, southern Siberia (Aranei: Linyphiidae) Ïàóêè-ëèíèôèèäû Àëòàÿ (Aranei: Linyphiidae) A.V. Tanasevitch À.Â. Òàíàñåâè÷ Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospect, 33, Moscow 119071, Russia. E-mail: [email protected] Èíñòèòóò ïðîáëåì ýêîëîãèè è ýâîëþöèè ÐÀÍ, Ëåíèíñêèé ïðîñïåêò, 33, Ìîñêâà 119071, Ðîññèÿ. KEY WORDS: Spiders, southern Siberia, Altai, taxonomy, faunistics, new species, distribution. ÊËÞ×ÅÂÛÅ ÑËÎÂÀ: Ïàóêè, Þæíàÿ Ñèáèðü, Àëòàé, òàêñîíîìèÿ, ôàóíèñòèêà, íîâûå âèäû, ðàñïðîñò- ðàíåíèå. ABSTRACT. New data on linyphiid spiders from 1996. C ó÷åòîì íîâûõ äàííûõ ôàóíà ëèíèôèèä ðå- the Altais are presented: 52 species are recorded in ãèîíà íàñ÷èòûâàåò íå ìåíåå 210 âèäîâ, èìååò ÿâ- the area for the first time, including four described íûé ïåðåõîäíûé õàðàêòåð ìåæäó åâðîïåéñêîé è as new: Halorates altaicus sp.n., Hilaira meridiona- þæíîñèáèðñêîé ôàóíàìè, õàðàêòåðèçóåòñÿ âûñîêèì lis sp.n., Incestophantes brevilamellus sp.n. and ïðîöåíòîì øèðîêîàðåàëüíûõ âèäîâ (58.1%), íåâû- Mecynargus minutus sp.n. An unknown female of ñîêîé äîëåé ñèáèðñêèõ ýëåìåíòîâ (23.8%), à òàêæå Anguliphantes sibiricus (Tanasevitch, 1996) and the îòñóòñòâèåì ìíîãèõ òèïè÷íûõ «ñèáèðÿêîâ» è þæ- males of Mughiphantes sobrioides Tanasevitch, 2000 íîñèáèðñêèõ âèäîâ. Ïðèâåäåí ïîëíûé ñïèñîê ïàó- and Walckenaeria katanda Marusik, Hippa et Ko- êîâ-ëèíèôèèä Àëòàÿ è èõ ðàñïðåäåëåíèå ïî âûñîò- ponen, 1996 are described for the first time. Based íûì ïîÿñàì. Øèðîêîàðåàëüíûå âèäû ëó÷øå îñâàè- on the new data, the linyphiid fauna of the Altais âàþò âåñü âåðòèêàëüíûé ïðîôèëü è äîìèíèðóþò âî currently contains 210 species, obviously being tran- âñåõ âûñîòíûõ ïîÿñàõ, ñóùåñòâåííî ïðåîáëàäàÿ (â sitional in character between the European and South 4 ðàçà) â íèâàëüíîé çîíå. Âåðîÿòíûå ýíäåìèêè Siberian faunas. It is dominated by widespread spe- (3.8%) ÿâíî ñèáèðñêîãî ãåíåçèñà, îáèòàþò (çà åäèí- cies (58.1%), the proportion of Siberian elements ñòâåííûì èñêëþ÷åíèåì) â âûñîêîãîðüÿõ. Àðêòî- being low (23.8 %), while many typical Siberians àëüïèéöû íàéäåíû èñêëþ÷èòåëüíî â âûñîêîãîðüÿõ and South Siberian species absent. A checklist of the è ïîäíèìàþòñÿ âïëîòü äî ìàêñèìàëüíûõ (èçó÷åí- Altai linyphiids and their vertical distribution are íûõ) âûñîò (3100 ì). Scotinotylus eutypus (Cham- given. The widespread species dominate in all altitu- berlin, 1948) â ñìûñëå Saito & Yasuda [1988], îòìå- dinal belts, essentially prevailing (4 times) in the ÷åííûé â ßïîíèè, âåðîÿòíåå âñåãî ÿâëÿåòñÿ àëü- nival zone. Presumed endemics (3.8 %) are obvious- ïèéñêî-àëòàéñêèì S. antennatus (O. Pickard- ly of Siberian origins and live (with one exception) Cambridge, 1875). in highlands. Arcto-alpine elements occur exclusive- ly at high altitudes and all of them reach the maxi- Introduction mum elevations explored (3100 m a.s.l.). Scotinoty- lus eutypus (Chamberlin, 1948) in the sense of Saito The Altais, altyn meaning gold in Turkic lan- & Yasuda [1988], recorded from Japan, most likely guages, is accepted here, according to Gvozdetsky, belongs to the Alps-Altai S. antennatus (O. Pickard- Mikhailov [1987], as the westernmost mountain prov- Cambridge, 1875). ince of the South Siberian Mountain system. The Altais extend over the territory of Russia, Kazakhstan, China ÐÅÇÞÌÅ. Ïðèâåäåíû íîâûå ñâåäåíèÿ ïî ïàó- and Mongolia, the latter two parts being termed the êàì-ëèíèôèèäàì Àëòàÿ: 52 âèäà âïåðâûå îòìå÷åíû Mongolian Altais. This paper deals with the Russian â ðåãèîíå, èç íèõ 4 âèäà îïèñàíû êàê íîâûå äëÿ and Kazakhstan territories of the Altais, concluded by íàóêè: Halorates altaicus sp.n., Hilaira meridionalis a few words about the Mongolian Altais. sp.n., Incestophantes brevilamellus sp.n. è Mecynargus The Altai territory is considered here in a little minutus sp.n. Âïåðâûå îïèñàíà ñàìêà Anguliphantes more restricted sense than by some authors. Thus, spe- sibiricus (Tanasevitch, 1996), à òàêæå íåèçâåñòíûå cies recorded from the localities like Kosikha, Talmen- ñàìöû Mughiphantes sobrioides Tanasevitch, 2000 è ka [Eskov, 1992] or Akutikha [Marusik et al., 1996] Walckenaeria katanda Marusik, Hippa et Koponen, are omitted from the Altai list. 268 A.V. Tanasevitch Since the publication of the first larger paper on Notioscopus jamalensis Grese, 1909 [Marusik et the spiders of the Altais [Marusik et al., 1996], the al., 1996; Levina, Mikhailov, 2004] = Notioscopus Altai list which contained 95 linyphiids has been en- sibiricus Tanasevitch, 2007 riched by 65 species, 11 of which described directly Oreoneta uralensis Saaristo et Marusik, 2004 from this mountain country [Azarkina, Trilikauskas, [Saaristo, Marusik, 2004] = Oreoneta eskovi Saaristo 2013; Ermolajev, 1937; Fomichev, Marusik, 2011; et Marusik, 2004 Hippa, Oksala, 1985; Levina, Mikhailov, 2004; Panamomops mengei Simon, 1926 [Marusik et al., Marusik, Logunov, 2009; Tanasevitch, 2000a, b, 1996] = Panamomops tauricornis (Simon, 1881) 2005a, 2007, 2008, 2010, 2011b, 2012, 2013; Ta- Porrhomma montanum Jackson, 1913 [Marusik et nasevitch et al., 2012; Trilikauskas, 2012; Volynkin al., 1996; Levina, Mikhailov, 2004] = Porrhomma pyg- et al., 2011]. The information contained in Savelyeva maeum (Blackwall, 1834) [1970, 1976, 1979] on the spiders from East Kaza- Saloca diceros (O. Pickard-Cambridge, 1871) [Lev- khstan lacks specific localities and is thus mostly ina, Mikhailov, 2004] = Horcotes strandi (Sytschev- omitted. Only Savelyevas records found and checked skaya, 1935) in the collections have been considered. The species Thyreosthenius biovatus (O. Pickard-Cambridge, referred to by Volkovskij [2006] and Volkovskij, Ro- 1875) [Marusik et al., 1996; Levina, Mikhailov, 2004] manenko [2005a, b, c; 2010] for the Altais have been = Thyreosthenius parasiticus (Westring, 1851) omitted as well, because they seem to be dubious and Trichopterna mengei (Simon, 1884) [Levina, requiring confirmation. Mikhailov, 2004] = Pelecopsis palmgreni Marusik et The new data in this paper add another 52 linyphiid Esyunin, 1998 species to the Altai list. Four of them are described as new: Halorates altaicus sp.n., Hilaira meridionalis Material and methods sp.n., Incestophantes brevilamellus sp.n. and Mecy- nargus minutus sp.n. Besides the spider material collected by Sergei Golo- The paper first lists the most interesting records. vatch and Andrei Tanasevitch (Moscow) from the Al- Some of them are supplied with figures of the genitalia. tais in 1997, this paper is also based on extensive At the end, a checklist of the Altai Linyphiidae and spider collections kindly offered for study mostly by their vertical distribution patterns are given. Galina Azarkina (Novosibirsk). Spiders were also tak- The following 16 species are eliminated from the en by Andrei & Roman Dudko, Andrei Fedotov, Na- Altai list, representing either synonyms or misidentifi- dezhda Levina, Dmitry Logunov, Oleg Lyakhov, Ilia cations. The valid names are on the right: Lyubechansky, Sergei Ovtchinnikov, Alexander Ryvkin, Synonyms: Veryaskina and Vadim Zinchenko. All available mate- Agyneta parasaxatilis Marusik, Hippa et Koponen, rial, upon which the papers Marusik et al. [1996] and 1996 = Agyneta pseudosaxatilis Tanasevitch, 1984 [Ta- Levina, Mikhailov [2004] were based, kept at the Zoo- nasevitch, 2011b] logical Museum, University of Turku, Finland (ZMT) Lepthyphantes tes Marusik, Hippa et Koponen, 1996 and the Zoological Museum, Moscow State University = Oryphantes geminus (Tanasevitch, 1982) [Ta- (ZMMU), have been re-examined. nasevitch, 2011b] Type and non-type material is shared between the Misidentifications, re-examined: collections of the ZMMU and the Institute of System- Agyneta rurestris (C.L. Koch, 1836) [Savelyeva, atics and Ecology of Animals, Novosibirsk (ISEA); 1979] = Agyneta saaristoi Tanasevitch, 2000 some more samples used here are also housed in the Agyneta similis (Kulczyñski, 1926) [Marusik et al., Muséum dhistoire naturelle, Geneva, Switzerland 1996; Levina, Mikhailov, 2004] = Agyneta saaristoi (MHNG) and in the private collection of Andrei Ta- Tanasevitch, 2000 nasevitch (CAT). Agyneta subtilis (O. Pickard-Cambridge, 1863) Distribution patterns of linyphiids are mainly giv- [Savelyeva 1970 1979] = Agyneta allosubtilis Loksa, en, based on the latest publications of Tanasevitch 1965 [2005b, 2006, 2008, etc]. Bolyphantes luteolus (Blackwall, 1833) [Levina, In the descriptions, the chaetotaxy of Erigoninae Mikhailov, 2004] = Bolyphantes alticeps (Sundevall, is given in a formula (e.g., 2.2.1.1) which refers to the 1832) number of dorsal spines on tibiae IIV. For Micronet- Hilaira tatrica Kulczyñski, 1915 [Levina, inae, the chaetotaxy is presented in a different formu- Mikhailov, 2004] = Oreoneta eskovi Saaristo et la, e.g., Ti I: 2-1-1-2(1), which means that tibia I has Marusik, 2004 two dorsal spines, one pro-, one retrolateral spine, Lepthyphantes bipilis (Kulczyñski, 1885) [Levina, and two or one ventral spine (the apical spines are Mikhailov, 2004] = Oryphantes geminus (Tanasevitch, disregarded). The sequence of leg segment measure- 1982) ments is as follows: femur + patella + tibia + metatar- Minicia uralensis Tanasevitch, 1983 [Levina, sus + tarsus. All measurements are in mm. All scale Mikhailov, 2004] = Silometopus uralensis Tanasevitch, lines in the figures correspond to 0.1 mm, unless 1985 indicated otherwise. The linyphiid spiders of the Altais 269 The terminology of genitalic structures in Micro- RECORDS FROM THE ALTAIS. RUSSIA: Arty- netinae follows that

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