The malacologicalsocietymalacological society of Japan

fin VENUS (Jap. Jour. Malac.) ･171 VoL 44, No. 3 (1985):159-

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Report on () Recently Dredged in the South-Western Indian Ocean I.

Description of Eight New Speeies

Roland HouART

(St. Jobsstraat, 8, B-3330 Landen (Ezemaal) Belgium: Scientific Collaborator

at the Institut Royal des Seiences Naturelles de Belgique)

Introduction

A recent paper by Vokes (1978) on the Muricidae from the eastern coast of Africa has demonstrated the imperfect state of our knowledge of Indian Ocean muricids. The present paper reports on Muricidae from a region adjacent to the area treated by Vokes: the islands and island groups of the SW Indian Ocean (Madagascar, Mauritius, Mayotte, Glorieuses Is., Reunion and the Sey- chelles). Although the material studied includes reef material, most of it

originates from dredgings and trawlings in deeper water, down to 1150m.

Undoubtedly this must account for' the high number (more than 25%) of new

taxa among the speeies collected. Despite their moderate to large size, and their

predominantly eontinental shelf habitat, murieids have a confused beeause of extensive convergence in shell charaeters. In this report, I am dealing

with the subfamilies Murieinae, , Typhinae and Trophoninae. There

are apparently no representative of the Ocenebrinae in that part of the world.

In addition to the subfamilies studied, Ergalataxinae were also present but are

being studied separately by Dr. E.H. Vokes. Species ar'e illustrated and com-

mented on when the record represents new information on the variation or

geographical distribution.

Material

The material on which this paper is based originates from various eollections

or expeditions in the south-west Indian Ocean and now housed in Mus･eum Na-

tional d'Histoire Naturelle.

* Kccepted: May 1, 1985

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160 VENUS: VoL 44, No.3(I985)

(1) Material collected by Dr. A. Crosnier off NW Madagascar in 1971-73 aboard "Vauban". R. V.

(2) Material collected by Drs. B. A. Thomassin and M. Peyrot-Clausade during

ecological surveys of Tulear reefs (SW Madagascar) in the 1970's.

(3) Material collected by Dr. P. Bouchet during the BENTHEDI expedition

"SuroiV' carried in 1977 aboard R.V. under the direction of Dr. B.A.

Thomassin in the northern Mozambique Channel.

(4) Material colleeted by R.V. Cleva during the REVES-2 expedition carried

in 1980 in the Seychelles.

(5) Material eollected by P. Bouchet, B. Metivier and A. War6n during the

"Marion-Dufresne" eruise 32 of R.V. (MD 32) carried in 1982 around

R-5union Island under the direction of Dr. A. Guille.

(6) Material dredged at Mauritius Island.

Description of New Taxa

Genus Chieorezes Montfort, 1810 Subgenus Chicoretes Montfort, 1810

Type-species: Murex ramosus Linn6, 1758 (ICZN opinion 911, 1970, Bull. Zool. nomencl. 27:20)

C:hicoreus (e]hicoreus) cloveri n. sp. (Figs. 5 and 5A)

Shell small fer the , fr'om 20 to 29.5mm. Aperture roundly ovate. Columellar lip white, smooth, partially erect on the lower portion of the anterior

part. Anal notch deep and relatively large. Outer lip slightly erect, denticulate,

briefly striate interiorly. Spire moderately high, consisting of two glossy

rounded nuclear whorls and five rounded, squamose postnuclear whorls. Suture

slightly appressed. Body whorl bearing three foliate varices, ornamented with

four short epen frondose spines. Shoulder spine larger and slightly longer,

followed by three short, equally distant spines with one or two intermediate

spinelets. Other axial sculpture consisting of one large and one very weak

intervarical eostae. Spiral sculpture consisting of numerous small squamous

cords and intermediate threads as in most of all the Chicoreus s.s. species.

Siphonal canal moderately long for the genus, finely open, backward reeurved

at the tip, bearing three upward recurved spines at its end. Shell of a uniform

light brown color, aperture white.

Dimerzsions: Holotype 23 × 13mm.

Material: Holotype in the Mus/eum National d'Histoire Naturelle, Paris, type

collection; 2 paratypes in the author's eollection.

Type tocality: Dredged in 180 m, muddy sand, at Tamarin Bay, 20 rniles south of Port Louis, Mauritius, 1978.

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Houart:Muricids in SW Indian Ocean-I. 161

Other localities: Flic en Flac, Mauritius, 50m, under rock (J. Drivas) and

Baie du Tom, Mauritius (P.W. Clover).

Etymotogy: Named for Phillip W. Clover, California, USA, who first gave me

the occasien to examine other specimens of this species.

Remutrlts :

Chicoreu$ ctoveri is related to ChicoTeus nobitis. Shikama, 1977. It presents

the same sort of spines and their arrangement is ver'y near. Only the faet

that C. eloveri is of a uniform light brown color differentiates it en first sight,

although some speeimens of C. nobilis are also uniformly coloured. C. cloveri

is smaller and, compared with a young C. nobitis, it shows a more elongate

shell, The seeond spine after that of the shoulder is mueh shorter than for C. nobilis. The most irnportant point (and what makes it surely a different species) is the protoconch: two globulous and glossy whorls, with a shallow terminal varix for C. ctoveri; conical, slightly carinate with a very different

terminal varix for C. nobilis.

thicoreus (Chicor'eus) crasnieri n. sp. (Figs. 4 and 4A)

Shell small for the genus, from 25 to 37mm. Aperture roundly ovate. Colu- mellar lip white, smooth, ereet on 1/4 of its anterior part. Anal notch deep and small. Outer apertural lip ereet and denticulate, briefiy striate interiorly. Spire high, consisting of two glossy flat and rounded nuclear whorls and five to six slightly angular postnuclear whorls. Suture appressed. Body whorl bearing three more-or-less foliate variees. Ornamentation of the body whorl variees variable. Shoulder spine absent or very shallow for the holotype, which present only the last, anterior spine. Both paratypes with carinal spines followed by three others of which the second is almost invisible on the smallest paratype. "forms". Protoconch and first whor]s are indentical for both Other axial sculp-

ture eonsisting of one large and one weak axial node; on holotype both axial nodes alrnost blended into a large single one. Spiral seulpture consisting of 9 or 10 squamous eords, usually with one intermediate squamous thread; sometimes two. Cords are slightly darker coloured on the largest paratype. Siphonal canal long and straight, narrowly open and slightly backward recurved at the at the tip; ornamented with three spines: two very near each other end of the canal and one situated medially, between the aperture and the first of the two Iast spines. The two last ones are strongly downward bent. Shell of a light brown color, aperture white. Dimensions: Holotype 37 × 15.8mm

Materiat: Holotype and one paratype in the Museum National d'Histoire Naturelle, Paris, type eollection; one paratype in the author's collection. Type tocatity; South of Madagascar, 26005'S, 44050'E, 100 m, 7 Mareh 1969,

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R. Plante coll.

Other locatities: Trawl 72, 25ell'S, 47e15'E, 85-90m (A. Crosnier coll.); Tu16ar, Madagascar, st. D71.3, outer slope, dredge, 58-165m (M. Pichon eoll.).

Etymotogy: This new speeies is named after Dr. Alain Crosnier, who eollected

this and rnany other interesting gastropods during his survey of the shrimp

fishing grounds off Madagascar.

Remarks :

C. erosnieri is near two related speeies: C. artemis Radwin & D'Attilio,

1976 and C. rossiteri (Crosse, 1872). From both speeies it differs by its eolor,

its Iarger aperture, its arrangement of spines on the siphonal canal and the

ver'y different protoconeh: eonical and slightly carinate for C. artemis and C.

rosst'teri, flat, slightly rounded and non-carinate for C. cTosnieri. The operculum

is almost identieal with that of C. rossiteTi. It must be noted that C. crosnieri

sometimes lacks the shoulder spine in the same manner as C. TossiteT'i.

Genus Pterynotus Swainson, 1833

Subgenus PteTynotus Swainson, 1833

Type species by S. D.: Mmex ptnnatus Swainson, 1822

Pterunotus (Pterunotus) guittei n. sp. (Figs. 1 and IA)

Shell medium for the subgenus, fusiform. Aperture ovate. Columellar lip

smooth, adherent posteriorly and slightly erect on 3/4 of its anterior part. Anal

notch shallow and large. Outer apertural lip barely erect and crenulated;

briefly striated interiorly. Spir'e high, consisting of 2 rounded nuelear whorls

and 6 slightly shouldered postnuclear whorls. Sutur'e appressed. The body whorl

bearing 3 large varices of which the last one shows short spines, eorresponding

to the spiral cords and connected with a thin webbing, extending also on the

siphonal eanal. The two preeeding varices are ornamented with shorter spines.

Other axial sculpture consisting of a large intervarical node. Spiral sculpture

consisting of 12 to 14 cords, flanked with 2 minor threads. Siphonal canal long,

narrowly open and slightly recurved dorsal]y at the tip; ornamented with

2 or 3 short open spines, conneeted with a thin webbing. Color light brown with

darker coloured intervarical nodes and medium part of the varices, aperture white.

Dimensions: Holotype 33.2 × 18.5mm; paratype 29.5 × 15mm. Material: Holotype and paratype in the Mus6um National d'Histoire Naturelle,

Paris, type collection.

Type tocality: MD32/R6union, st.DC126, 20052'S, 55e38'E, 110m.

"Marion- Etymology: Named after Dr. A. Guille, Chief scientist of the R.V.

Dufresne" eruise MD32 around Reunion Island in 1982.

Remarks :

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Houart : Muricids in SW IndianOcean-I. 163

1

1

Figs. 1-IA. Rerynotus guiUei n. sp,. R6union, holotype. 33.2 × 18.5mm LIA, × 35..2. AspeUa thomassini n. sp.. Tu16ar. holotype, 7, 9 × 4mm.

This species may be compared with 2 more or Iess closely related species: PteTynotus (Pterynotus) la･quea･tus (Sewerby, 1841) and PteTynotzes (Pterynotits) martinetan- (R6ding, 1798). From the first it differs by its aperture, more

rounded and more denticulate for P. Iaqueatus which shows also fewer spiral

cords, a shorter siphonal eanal and in which the varices are ornamented with

longer webbed spines. From P. martinetana it differs by its number of varices,

the spiral eords, the rnore denticulate aperture for P. ma･rtinetana, which also

has a different columellar lip and protoconch.

Genus Poim'erin Jousseaume, 1880

Subgenus Pazinotus Vokes, 1970

Type-speeies by O.D.: Euptema stimpsonii Dall, 1889

Pbirieria (Rizinofus) brevispIendoris n. sp.

(Figs. 6 to 6B)

lip Shell moderately large for the subgenus. Aperture ovate. Columellar

completely erect, bearing a small single but strong dentiele on its anterior part,

smooth elsewhere. Anal notch shallow and large, delimitated on the right by a

strong denticle, followed by a deep interspace and 4 other strong dentieles on

the inner side of the outer lip; this last weakly undulate. Spire high, consisting

of one and one-half glossy rounded nuelear whorls and five angulate pQstnuclear

whorls. Suture deep and impressed. Body whorl triangular, bearing 5 spinose

varices. No other axial sculpture exeept shallow axial growth lines. Spiral

sculpture of the first whorls consisting of 2 spiral eords. The body whorl shows

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164 VENUS: VoL 44, No.3(1985)

4 spiral cords with 2 or 3 intermediate shallow threads on upper part, 4 spiral

threads on lower part and numerous fine striae on centre surfaee. Where the

cords intersect the varices they give rise to short and acute spines. Carinal

longest and r'ecurved upward on the last varix; almost horizontal on the other

varices. Spines of ]ast varix connected by a fimbriate webbing which does not

appear on other varices. Carinal spine of last varix also conneeted to body

whorl by a horizontal webbed flange. Siphonal eanal long and slender, finely

open, strongly backward recurved and smooth, with the varical flange of last

varix coming to its mid-point. Color reddish-brown, aperture light pink.

Dimensions: Holotype 14 × 8.8mm

Materiat: Holotype and 4 paratypes (juveniles) in the Museum National d'His-

toire Naturelle, Paris, type eolleetion; 1 paratype in the Natal Museum, Pieter-

maritzburg, South Afriea, nr.K311/T3043.

Type tocality: MD32, St.CP129, 2005YS, 55036'E, 290-300m.

Other loeaeities: MD32, St.DC26, 21e22'S, 55"47'E, 310m; St.DC128, 20'51'S,

55036tE, 280-340 m.

Remarics :

Four speeies of Pazinotus are known from the Indo-Pacific: Paz-inotu･s sm･ith･i

(Schepman, 1911), Eastern Sulu Archipelago, Philippines; Pazinotus fatcatiformis

(Thiele, 1925), Indian Oeean; Pazinotus oliveTai (Kosuge, 1984), Philippines;

Pazinotus phitcloveri Houart, 1984, Philippines. However, this new species may

be reasonably only compared with 2 species: P. smithi and P. philctoveri, the

other 2 speeies are very different. P. smithi is smooth, exeept for 6 large and

shallow spiral cot-ds giving rise to 6 large, ]ong spines. On the last varix these

spines are eonnected by a webbing. Ne webbing is present on the other varices.

Carinal spine longest and recurved upward on all the variees. The new species

shows 4 good visible spiral cords on the upper part of the body whorl with

intermediate threads and 4 thr'eads on lower part. The carinal spine is Iongest

but still fairly short and aeute; only the apertural is reeurved upward, on the

other varices it is almost horizontal and straight. The eolumel]ar' lip of the

new species bears a small but strong denticle on its anterior part while that of

P. smithi is srnooth. The apertural denticles of the new speeies are stronger,

showing deeper interspaces between them. The siphonal eanal is slender.

P. philcloveri has shorter spines. The suture is not so deep. The columellar ]ip shows 3 to 4 small denticles anteriorly. The presenee of heavy growth

]amellae gives an imbr'icate appearanee to the shell seulpture. The shoulder is

Iower. There are more numerous and finer spiral threads. Finally, the pro-

Figs. 3"3D. Poirieria (Flifexopteron) Primanova n. sp. Madagascar, holotype. 14.7 × 11.4mm C3B. × 20;3C. × 3500;3D. × 2500). 4-4A. Chicoreus crosnieri n. sp. Mada- gascar. holotype, 37 × 15.3mm. (4A. × 20), 5-5A. Chicoreus cloveri n. sp.. Mauritius, holotype, 23 × 13mm, (5A, × 27).

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Houart : Muricids in SW Indian Ocean-I. 165

3A

3B 3D 3C

4A /,-･t.

L di. /･k.f twfittt

5A 5

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166 VENUS: Vol. 44, No.3(1985)

toeonch is different in being smaller and more acute in P. philcloveTi.

Subgenus Rtexopteron Shuto, 1969 Type-species by O.D.: Ii'lexopteron phitimpinensis Shuto, 1969

.Pbirieria (IiZexoptei'orz) Ill'imanova n. sp. (Figs. 3 to 3D) Shell medium sized for the subgenus, from 14.7 to 17 mm, rounded, rather fragile. Aperture ovate, columella lip eompletely adherent to the shell and

smooth. Anal notch shallow and fairly Iarge. Outer apertural lip weakly undulate,

inner side bearing 5 strong dentieles. Spire moderately low, eonsisting of one and one-half fiat, glossy nuclear whorls and five rounded, large and slightly angulate postnuclear whorls. Suture impressed. The body whorl bears 7 or 8 sharp, low winglike varices; no other axial sculpture. Spiral seulpture eon-

sisting of 6 cords and numerous fine spiral striae covering the whele shell. Wher'e

the cords intersect the varieel flange, short open spines formed. Carinal spine

longest and upward recurved. Siphonal canal short, open, very slightly bent

backward. Color of the shell white to light br'own. Aperture of the same color,

usually Iight brown, the dentieles and space between them and apertural edge

white. The rather small muricine radula eonsists of a raehidian tooth and a

long and slender lateral. The raehidian has strong, prominent endpoints and

bears five cusps: a long central with long laterals, intermediate eusps are short.

Dimensions: Holotype 14.7 × 11.4mm

Mnterinl: Holotype and 2 paratypes in the Museum National d'Histoire

Naturelle, Paris, type eollection ; 1 paratype in the Natal Museum, Pietermaritz-

burg, South Afriea, nr. J9362fT3038; 1 paTatype in the author's collection.

Type locality: Crosnier, Madagascar, trawl 13, 12041'S, 48e16'E, 308-314m.

Other l'oc/alities: Crosnier, Madagasear dredge 2, 12038'S, 48a16'E, 240m;

trawl 10, 12043'S, 48043'E, 300-340 m; trawl 31, 12e34'S, 48015'E, 395 m; trawl

12!, 12040rS, 48014,E, 410m.

Remarks :

Only known from about 10 fossil species: 8 European, 1 Caribbean and 1 from the Philippines (Vokes, 1971), this genus was ereeted by Shuto (1969: 111) in the family Coralliophilidae. Strueture of the shell and presenee of a murieine radula (unknown in the Coralliophilidae) shows its aMnities with the

Murieidae. The discovery of live specimens of a new species in the Indian

Oeean is undoubtedly the most exeiting news of this report. P'lexopteron pri-

manova･ n. sp. may be compared with Flexop･teron pahitippinensis Shuto, 1969.

The new species has 6 large spiral eords on the body whorl, including the

carinal one; Rlexopteron phil'ippinensis has finer spiral cords and threads, 10 in

total, on the body whorl. The new species has a flatter shoulder', finer variees,

a flatter protoeonch with a straighter terminal varix.

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Houart:Muricids in SW Indian Ocean-I. 167

Genus AspeUa M6rch, 1877 Type-speeies by monotypy: RanetLa anceps Lamarck, 1822 Aspella thomassini n. sp.

(Fig. 2)

Shell medium sized for the genus, elongate. Aperture roundly ovate with a

completely adherent, smooth, columellar lip. Inner side of outer lip denticulate,

with 6 short denticles. Very shallow and Iarge anal sulcus. Spire high, con-

sisting of a protoconch of undetermined nature and 6 angulate postnuclear whorls. Suture impressed and obscured by varical buttresses. Body whorl broad, bearing

4 strong variees and two strong costae, whieh represent the remains of the single ventral and dorsal varices of the first whorls. One single shallow cord

representing the only spiral sculpture on penultimate and last whorl ; other whorls

smooth. Siphonal canal moderately long for the genus, open, bent to the right and reeurved dorsally at the tip. Most of the intritacalx removed but traces

of it showing a cancellate seulpture.

Dimensiens: 7.9 × 4mm

Ma-teriat: Holotype in the Mus6um National d'Histoire Naturelle, Par'is, type

colleetion.

Type locality: Tu16ar, St.217, Gran dR/6cif, outer reef slope, 17m, B.A.

Thomassin coll. Etymology: Named after Dr'. B. A. Thomassin, who colleeted the type material and many other valuable specimens during his survey of Tu16ar reefs.

Remarks :

Related only to the American species, sueh as Aspetta senex Dall, 1903; Aspetla castor Radwin & D'Attilio, 1976 and Aspetla potLux Radwin & D'Attilio, 1976, this shell may be eompared with A. ptatylaevis Radwin & D'Attilio, 1976 from which it differs by its more angulate variees, by its Iarger and more angulate form and by its siphonal canal. From AspeLlen pToducta (Pease, 1861) it differs also by its more angulate variees, by its longer and straighter siphonal canal, by its more triangular form, its finer dorsal costa and its smaller size.

Genus Mzarex$ul Iredale, 1915 Type-speeies by O.D.: MuTex octogon2es Quoy & Gaimard, 1832 Murexsul reunionensis n. sp.

(Figs. 7 to 7B)

Shell small for the genus, from 7 to 10mm, angulate. Aperture lenticular. Columellar lip smooth, adherent to the shell. Anal notch large, barely visible. Inner side of outer apertural lip smooth, or generally bearing 3 to 5 very low denticles briefiy extended interiorly; these denticles becoming stronger on the anterior part of the aperture. Spire high and acute, consisting of one and one-

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168 VENUS: Vol. 44, No.3(1985)

half rounded, glossy nuelear whorls and five angulate postnuclear whorls. Suture

covered by the preeeding whorl. The body whorl bearing 5 or 6 large, slightly

rounded varices, ornamented which short, largely open spines. No other axial

sculpture. Spiral sculpture eonsisting of 6 slightly squamous grooved eords and

fine intermediate spiral threads; these cords and threads corresponding to short

spines at the varices. Other fine threads on the shoulder and 3 on the siphonal

canal. Siphonal canal moderately long to long, open and bent backward. Color

ehalky white or very light brewn to light orange;aperture of the same color.

Dimensions: Holotype 10 × 6mm

Ma･teriae: Holotype and 13 paratypes in the Museum National d'Histoire Naturelle, Paris, type colleetion ; 2 paratypes in the Natal Museum, Pietermaritz-

burg, South Africa, nr.J9361/T3037; 2 paratypes in the author's eolleetion.

Type Iocality: MD32, St.DC128, 2005YS, 55036'E, 280-340m.

Other localities: MD32, St.DC41, 21e21'S, 55027'E, 75m; St.DC56, 21e05'S,

55012iE, 170--225 m.

RemaTks :

Two related speeies may be eompared: MurexsuL toicubeii Nakamigawa &

Habe, 1964 from Japan and Taiwan. This shell is different in being rnuch larger

for the same nurnber of postnuclear whorls; 5 or 6 barely visible threads on

the shoulder and one short spine on the siphonal eanal. Another similar species,

Murexsut elatensis (Emerson & D'Attilie, 1979) from the Red Sea is very near,

but the new species has finer and more grooved spiral eords and more marked

cords en the shoulder. Moreover, it has intermediate spiral threads, while M.

etatensis has none. M. elatensis has also mor'e upward recurved and Ionger

earinal spines. Although named as Favartia I here suggest removing F. etatensis

to the more appropriate genus Mrurexsul.

Genus Siphonoehelus Jousseaume, 1880

Subgenus Siphonoehetus Jousseaume, 1880 Siphonochelus (SiPhonochetus) stillacandidus n. sp. (Figs. 8 and 8B)

Shell small for the genus, about 7 to 8.5mm in length, elongate, fusiform,

translucent or milky white. Aperture ovate. Columellar' ]ip smooth, ereet on

213 of anterior part and adherent posteriorly; no traee of anal noteh. Body

whorl fusoid, bearing 4 sharp varices. A short upward reeurved and rounded

anal tube originating near the succeeding varix and conneeted with it. Spire

high and acute, eonsisting of one and one-ha]f rounded, polished nuelear whorls

tt

."'kg,s)-.61fi-,Bi.t'.ree.'.la.,(e?.zi.",.`.".s;.b..r":isp.Le.rd.og･ts.;6.?pff.,R.f.","g?"i,h.ogol/?hp?･,;`.if8k8,fgsl.i･,E,6.B:

Siphonochel"s stillacandidus n. sp.. Madagascar. holotype, 8, 5 × 4 mm. (8 B. × 45).

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Houart :Muricids in SW Indian Ocean-I. 169

6B

7B

7

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170 VENUS: Vol. 44, No.3(1985)

and 4 or 5 slightly angulate postnuclear whorls. Suture deeply impressed. Siphonal canal moderately long and narrow, barely bent to the right or straight.

Dimensions: Holotype 8.5 × 4mm.

Materiat: Holotype and 3 paratypes in the Mus6um National d'Histoir'e

Naturelle, Paris, type eollection ; 1 paratype in the Natal Museum, Pietermaritz- burg, South Africa, nr.J9363fT3039; 1 paratype in the author's collection.

Type locality: Madagascar: Benthedi, st.3, W. Banc du Leven, 12036'S, 47e38'E,

1100-1150 m.

Other tocalit?ies: Mayotte: Benthedi, St.33, E. Passe Longegori, 12054'S,

45016'E, 275-400m; St.38, E. Recif Bandele, 12055'S, 45016'E, 200-500m; St.

DS42, S. R･6cif Sud, 13005'S, 45005'E, 4oo-s2om; St.72, NNE, Reeif Nord,

12e31'S, 4seo2'E, 35e-700 m; St.120, SE GIorieuses, 11030'S, 47025'E, 335-390 m.

Remarks :

Siphonoehetnts nipponensis Keen and Campbell, 1964 differs in having more flattened anal tubes while these of the new species are more rounded, it has

also a flatter' pt'otoconch ; the new species is a more fusiform shell. Sip･honochelus (Laevityphis) tzaelrookae Keen and Campbell, 1964, an Australian fossil, is a much Iarger and more triangular shell with a larger aperture and more perpen- dieular tubes. Moreover in the subgenus Laevityphis the anal tubes are not

incorporated into the varices. SiphonocheLu's generosus (Ir'edale, 1936) is also

a larger shell with more flat and upward eurved anal tubes; it has also a more globular protoconeh. Siphonochelzas sotus Vella, 1961 is larger, for the same number of postnuelear whorls, it is more slender, has more rounded varices and

a flatter protoconch. Acknowtedgements;-First of all, I wish to thank partieularly Dr. Philippe Bouchet (Museum National d'Histoire Naturelle, Paris) who gave me the opportunity to study

this very interesting material and who suggested me some useful corrections on my

manuscript. I am most indebted also to Dr. E.H. Vokes (Tulane UniveTsity) for her

very welcome suggestions and eomments. I am very thankful to Dr. B.A. Thomassin and Dr. P. Arnaud (Centre d'Oe6anologie de Marseille) for their useful remarks

and eorreetions. Thanks also to Dr. J. Van Goethem

naturelles de Belgique) and to the staff of the Department of Invertebrates for their

collaboration, and to Mrs. J. Buyle (BTussels) for processing some of my photographs. I wish also to thank the following people whe loaned me comparative material and

types; without them valuable work could not have been done: Mr. P. H. Cloyer (Calif., Zoologie, USA), Dn H. E. Coomans and R. G. Moolenbeek (Instituut voor Taxonomisehe Amsterdam), J. Drivas (Reunion), B. Foster and C. Glas･s (Calif. USA), Dr. R.N. Kilburn (Natal Museurn, Pietermaritzburg, S. Africa), Mrs. A. Lesage (Brussels), Dr. W. F. Ponder and Mr. I. Loch (Australian Museum, Sydney), Dr. R. Van Syoe (California Academy of Seiences, San Francisco), Ms. K.H. Way (British Museum Natural History, London) and Dr. T. Shuto (Kyushu UniveTsity, Japan).

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− Houart ： Muricids in SW Indian Ocean 1． 171

要 約

ソ ア ガ る に イ ド 洋 南 西 部 の ッ キ イ 科 に 属 す る 8 新 種 を 記 載 す 。 化 石 属 FlexoPteron Shuto ，1969 属

こ Poirieria の て す る と 思 わ れ る 1 種 が 含 ま れ ， れ を 亜 属 と し 扱 っ た 。

1． Chicoreus （Chicoreus） cloveri n ． sp ．（第 5 ， 5A 図 ）。 殻 は 小 型 で ，C ． nobitis Shikama ク レ ナ イ セ ソ ジ ュガ イ に 似 る が ， 殻 が 褐 色 単 色 で あ る こ と， 殻 ー い こ × モ シ ャ ス が や や 細 長 と ，胎 殻 が 丸 く終 層 に 低 い 縦 張 肋 が あ る 点 で 異 な る 。 23 13mm 。 リ ， ポ ー ト ル イ ス 20 哩 南 の タ マ リ ン 湾 ， 180m の 砂 泥 底 。

crosnieri n sp 4A 2． C ．（C ．） ． ．（第 4 ， 図 ）。 ’ コ セ ソ マ コ 殻 は 小 型 で C ．artemis Radwin ＆ d Attilio （ ジ ュ ガ イ ） や C ． rossiteri （Crosse ） ィ

の で い 口 で る セ ン ジ ＝ ガ イ に 似 る が ，淡 褐 色 の 色 彩 や 棘 の 配 列 み な く， 大 き 殻 と 平 滑 な 胎 殻 異 な 。

マ ガ ス カ ル の 37x15 ．8mm 。 ダ 南 方 水 深 100m 。

3． Aerynotus （Rer ンnotus ） guiUei n ． sp ．（第 1 ， 1A 図 ）。

で の は い ミ ズ カ ヨ ラ ク ガ 殻 は 中型 ， 淡 褐 色 結 節 と 縦 張 肋 中 ほ ど 濃 。 P ．laqueatus （Sowerby ） キ ウ イ に 似 る が ，殻 口 が よ り 細 か く刻 ま れ ，前 水 管 溝 が 長 く縦 張 肋 上 の 棘 が 短 い 。 P ．〃martinetana （R6ding ）

コ ヨ レ ユ ニ ン ウ シ ウ ラ ク ガ イ と は 縦 張 肋 の 数 ， 螺 肋 ， 胎 殻 の 特 徴 で 異 る 。 33．2x18 ．5mm 。 オ ， 水 深

110m 。

4． Poirieria （Pazinotus ） brevisPlendoris n ．sp ．（第 6 ， 6B 図 ）。

P smithi こ の 類 と し て は や や 大 き く，全 体 赤 褐 色 。 ． （Schepman ） ｝よ螺 肋 か ら 6 本 の 棘 を 生 じ ， 〜 P ．Philcloveri Houart は 軸 唇 に 3 4 の 歯 が あ り， 殻 は 鱗 状 と な り， ま た 胎 殻 も小 さ く尖 る の で 本

° ’ ° ’ S 36 E 〜 種 と 異 る 。 14x8 ．8mm 。 20 51 ， 55 ． 290 300m 。

〜 5． R （FtexoPteron ） Primanova n ．sp ．（第 3 3D 図 ）。 殻 は 短 く ， カ プ ラ ガ イ 科 の 化 石 属 と し て Shuto （1969 ） が 創 設 し た FlexoPteron に 属 す る と思 わ れ

Shuto で つ る 。EPhilippinensis と は ，螺 肋 が 粗 く， 肩 が 平 ら ， 平 担 な 胎 殻 を 持 点 で 区 別 さ れ よ う 。

マ ス ル 〜 14 ．7 × IL 4mm ， ダ ガ カ 沖 308 314m 。

6． AsPetta thomassini n ． sp ．（第 2 図 ） 本 種 は 磨 減 し た 標 本 に 基 い て い る た め ，縦 張 肋 間 に は 僅 か に 格 子 状 彫 刻 を 認 め る に す ぎ な い 。 A ． ’ っ の ラ ン platylaevisRadwin ＆ D Attilio と は 角 張 た 縦 張 肋 と 水 管 様 子 が 異 な る 。 7．9x4mm 。 グ

レ シ フ の 外 礁 17m 。

7． M 〃 6 κ 祕 reunionensis n ，sp ．（eg7 ， 7B 図 ）。

本 種 は M ，toleubeii Nakamigawa ＆ Habe オ ナ ガ ア ラ レ ツ ブ リ ボ ラ と 極 め て 似 て い る が ，本 種 は

〜 つ こ 一 つ Radwin 大 型 で 体 層 で 5 6 条 の 螺肋 を も と と 水 管 溝 に 小 短 棘 を 持 。 紅 海 産 の M ．elatensis

’ ° ＆ D Attilioは 極 め て 近 い が ，本 種 の 方 が 螺 状 彫 刻 は 細 か く， 間 肋 を 持 つ 点 で 異 る 。10x6mm 。20

’ ° 「 E 〜 340m 51 S ．55 36 ．280 。

8． SiPhonechetus （SiPhonochelus ） stillacandidus n ．sp 。（第 8 ， 8B 図 ）。

エ ソ ツ ヨ ・・ ソ エ ン ッ ヨ 本 種 は 乳 白 色 の ト ウ ラ ク 類 で ， ッ ポ ト ウ ラ ク ガ イ S、niPPonensis Keen ＆

マ ダ ス ル パ ン Campbell に 似 る が ， 本 新 種 は 後 溝 に 丸 み が あ り， 胎 殻 は 尖 る 。 8．5x4mm 。 ガ カ ，

レ パ ン 〜 150m デ ュ 西 沖 1，100 1， 。

References

Bibliographical references wi11 be listed at the end of the second par 七 〇 f this paper ．

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