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Selachians from the Late Cretaceous (Turonian) Atarque Sandstone

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Selachians from the Late Cretaceous (Turonian) Atarque Sandstone Selachiansfrom the Late Gretaceous (Turonian)Atarque Sandstone Member, TresHermanos Formation, Sevilleta Grant, SocorroGourty, New Mexico byDonald L. Wolberg,Paleontologist, New Mexico Bureau ol Minesand Mineral Resources, Socorro, NM 87801 Introduction found at the locality. On the Sevilleta Grant, 3.4m (11ft) to 13m (42ft) and canbe divided The Cretaceousfish faunasof NorthAmer- more than 396 m (1,300ft) of Upper Creta- into lower, middle, and upper parts. The basal ica are still poorly known (Applegate,1970). ceous rocks overlie shales of the Dockum Atarque consists of light-gray, fine-grained, Selachians,although not uncommon in Cre- Formation (Upper Triassic).The Upper Cre- calcareoussandstones; the middle Atarque taceousrocks, are especiallypoorly under- taceoussequence consists largely of shales consistsof yellow-orange,fine-grained, thinly stood and in need of extensive study and sandstonesand includes units from the bedded, calcareoussandstone with shale (Cappetta,1973). Although Marcou de- Dakota Sandstoneto the CrevasseCanyon partings;the upper Atarque consistsof very scribed Ptychoduswhipplei from New Mexico Formation (Baker and Wolberg, 1981).The fine grained, gray-orangesandstones (Baker, as early as 1858, very little work on New Tres Hermanos Formation overlies the Rio 1981). Mexico Cretaceousselachians has been pub- SaladoTongue of the Mancos Shale and un- Selachiansare very abundant in lenses lished. derlies the D-Cross Tongue of the Mancos within the middle part of the Atarque. In While conducting a geologic study of Up- Shale.The fossil-richlenses that yielded the addition to selachianfossils, turtle shell ma- per Cretaceousrocks exposedon the Sevil- selachiansdescribed below are found in the terial, crocodilianarmor and teeth, and two leta Grant near La foya, SocorroCounty, New middle part of the Atarque SandstoneMem- plesiosaurteeth have been recovered.Car- Mexico(Fig. 1), BruceBaker discovered a se- ber, the basalunit of the TresHermanos (Fig. bonate-cementedmollusk shell material oc- ries of fossil-richlenses in sandstonesof the 2). The stratigraphicnomenclature for the curs abundantly as well. The vertebrate lower part of the Tres Hermanos Formation region has been revisedby Hook and others material was recoveredby breaking down (Baker,1981). Baker and Wolberg(1981) pro- (1983).In the SevilletaGrant, the Atarque the rock matrix with dilute aceticand formic vided an interim report on the vertebrates SandstoneMember varies in thickness from acids followed by washing, screening, and drying the resultantconcentrate. Specimens then were picked with and without the use of a binocular microscope. rF Lodron /Bernord\--.' Pinos-i -- Alsoin this issue .t\/-Mis.;/,.\s .{' ConchasLake State Park o. 8 l1l.^..Chupoderoj -: Enhancedoil recoverywith sMY veso--j- a. AKLA '.-_ CO2flooding p. 10 Magdoleno Z /"i?N llruy I ru = Gastropodsfrom the Solse lnitts . '1- MeteMember p. 11 Socorro Jornod o = Service/News p. 16 ^ dal--' 50nn Staffnotes p.20 tonio Muerto Z.S Gomingsoon Baritein north-centralNew Mexico Sedimentsorting in gravelly megaripplesfrom the RioGrande EspanolaSubsidence Project -u, ltl Abstractsfrom the 1984Mineral I 13 km Symposium FIGURE l-Location map of the study area, Socorro County, New Mexico (after Baker, 1981). This paper is an abbreviated version of a The genus Hybodusis a common Mes- nus in north-central Texas. Leriche (1939) larger work to be published as part of a vol- ozoic form that is known from the Triassic- reported S, kaupifrom the Coniacian,San- ume on Late Cretaceouspaleontology Cretaceousin North America (Romer,1966). tonian, and early Campanian of Africa. S. (NMBMMR, Circular195), and it attemptsto Hybodusistypical of hybodont sharks,a group falcatusis known from the Turonian of Texas acquaint the readerwith the diversity of New that had a Paleozoicorigin and were cos- (Bilelo, 1969)and South Dakota (Cappetta, Mexico's Cretaceousselachians; economies mopolitan in distribution. Hybodonts 1973).S. falcatus also is known from the Cen- have been made in taxonomic descriptions underwent their own adaptiveradiation and omanian,Santonian, and Campanianof Eu- and historic data for various taxa. The clas- were not at the origin of modern selachians. rope (Priem, 1.972). sification schemeused generally follows S. pristodontusranges from the late San- Romer (L966),Cappetta (1973),Cappetta and Family PrvcHoooNrrDAEWoodward, L9L2 tonian to the end of the Maestrichtian, al- Case(1975), and Cappetta(pers. comm. 1984). Genus PtychodusAgassiz, 1839 though a pre-Campanianage may be doubted Ptychoduszohipplei Marcou, 1858 on geologic grounds (Bilelo, 1969). S. pris- iTAGI ROCXUNITS ITHOLOGI Fig. 3-A todontushas a cosmopolitandistribution. De- spite a stratigraphic overlap, the S. kaupi-9. .9 P. whippleiis a very distinctive taxon, and pristodontussequence forms a suit- =t ;E exceptfor P. anonymus,not easily confused falcatus-9. U :1 with other species of Ptychodus.P. whrpplei able evolutionary series. The broad geo- H:-5 was first named for specimensfound in the graphic distribution of these taxa enhances Cretaceousof New Mexico. Marcou (1858) their utility for stratigraphic application. z J. F S. is not abundantly represented Sondstone reported P. whippleifrom "the gray sandy falcatus tr marls, three miles north of Galisteo,on the in the Joyita Hills fauna. No completespec- Sond ston€ imens have been recoveredvet. S. falcatus z f--t Srllslone road from Galisteoto Pecos,New Mexico." Distinguishedby their raisedcrown, teeth of was an active predator as evidencedby its e trenchant,serrated teeth. T -l Sho Ptychoduswere adaptedto crushing mollusk I shells.Although the genus has a cosmopol- Order GalEronvns Colcorenile 9c E itan distribution, P. zohipplei,which occurs SuborderIsunorplI 3F commonly when it is found, seemsto be re- Family Isunroer Garman, 1913 t4l Con g omerote t'.l stricted to the Late Cretaceousof North Genus CretolamnaGlikman, 1958 America. Cretolamnaappendiculata Agassiz, 1835 tr I olurboied Ptychodusanonylrtus Williston, 1900 Fig. 3-R-T Eurrowed )t) Fig. 3-C, D .9 tr C. appendiculatahas cosmopolitan distri- o bution and occurs in rocks of Cenomanian E Concreiions P. anonymusis similar to P. uthipplei;both l e to late Paleoceneage. The family Isuridae td taxa display strongly raised apical crowns. z t-;l Fossi lized (Lamnidaein older literature) includes the g t__J However, the crown of P. anonymusis com- E mackerel sharks and consists of large, vo- U z Cross- parativelymore elongatedthin that of P o J> t I -tE strotificolion racious fish with lunate caudal The d. L,,I zohippleiand is less acutely conicai. Like P fins. l - F E whipplei,P. anonymusis restrictedto the Late E Coo L! E o_ Ss Cretaceousof North America and was orig- l E Unconforhrly inally describedby Williston (1900)from the t__l "Benton'"and "Niobrara" of Kansas.It seems New AAexnc@ 9 Foss L ocoiily likely that these two speciesare closely re- F lated,although the natureof the relationship is as yet unclear. GEOLOGY - I ::--51 Bardack(1968) figured a ptychodont from r Scionceand Service tt. ;T;. .9 lTi c to1.o the Boyne member of the Vermillion River Volume 7, No. 1, February 1985 Formationin Manitoba, Canada.I concurwith EdifolrDeborah A. ShaF Published quarterly by .9 I :1" Evetts (1979)that this specimencan be re- New Mexico Bureau of Mines and Mineral Resources P' ferred to P. anonymus. a division o(New MexicoInstitute ofMining & Technology z ;e BOARD OF REGENTS 3 Ptychoduspolygyrus Agassiz, 1839 z Ex Officio Fig. 3-B Toney Anaya, Goaernorof Nru Mexico zz Leonard Delayo, Superintend.entof Public lnstruction z P.polygyrusis a cosmopolitanspecies known ADDointed LtJ t ! from the Turonian through the Santonianof Donald W Monis, Pres.,1983-1989, Los Alqmos Robert Lee Sanchez,Sec.lTreas., 1983-1989, Albuquqque Dokolo the (Woodward, 1911), English Chalk the William G. Abbott, Hobbs Sondslone 1951 7985, "Niobrara" of Kansas(Williston, 1900),and Judy Floyd, Dn-1987, Ins Cruces E6 0ockum the Selma Group of Alabama (Applegate, Steve Torres, 1967-7985, SNorrc tg 1970). P. polygyrusalso is known from Bel- New Mexico lnstitute of Mining & Technology gium and the U.S.S.R.The abundanceand Pt$idilt.. .. LaurenceH.Lattman FIGURE2-Stratigraphy the New Mexico Bureau of Mines & Mineral Resources of SevilletaGrant, diversityof the ptychodontfauna in theAtar- D;reclor.. ....... FrankE.Kottlowski SocorroCounty, New Mexico(after Baker and que Member clearly reflects the abundance DeputyDirector ...... Georges.Austin Wolberg,1981). of the mollusks that formed the food source Subwiptions:Issued quarterly, February, May, August, for theseselachians. November; subsaiption price 96.00/calendaryear. Ed.itorialruttq: Conhibutions of material for consideration Systematicpaleontology in future issuesof NMG arewelcome. Articles submitted Class CHONDRICHTHYES Order LarnrNrFoRMES for publietion should be in the editor's hands a minimm Suborder LeuNoroEr of five (5) months before date of publication (February SubclassElasN,rosRAl'rcHr May, August, or November). Address inquiries to Order Strecnu Family Al.qconactpen Casier, 1947 Deborah A. Shaw, editor of Nru Mexico Geology,New MexicoBureau of Mines & Mineral Resources,Socono, Suborder Hynooorurorora Genus SqualicoraxWhitley, 1939 Squalicorax (Agassiz),1843 NM 87801. Family HysoooNnoer Owen, 1846 falutus Publishedas public domain, fhereforereproducible without Genus HybodusAgassiz, 1,837 Fig. 3-P permission.Source credit requested. 1,4N Hybodussp. The frenchant rec- Circulation: teeth of S. t'alcatusare
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