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1.5.3 Environmental legislation
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1.6 Research overview
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Figure 1.2 Location of study sites within Thames Gateway area of South-east England. Basemap source: Esri, DigitalGlobe, GeoEye, i-cubed, USDA FSA, USGS, AEX, Getmapping, Aerogrid, IGN, IGP, swisstopo, and the GIS user community.
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Figure 1.3 Location map for Canvey Wick Site of Special Scientific Interest, Essex. Basemap source: Esri, DigitalGlobe, GeoEye, i-cubed, USDA FSA, USGS, AEX, Getmapping, Aerogrid, IGN, IGP, swisstopo, and the GIS user community. Contains public sector information licensed under the Open Government Licence v3.0. HFRORJLFDOVXUYH\VUHYHDOHGQDWLRQDOO\RXWVWDQGLQJLQYHUWHEUDWHDVVHPEODJHV SDUWLFXODUO\DVVRFLDWHGZLWKGLVWXUEHGEDUHJURXQGKHUEULFKJUDVVODQGVFUXE HGJHDQGEUDFNLVKKDELWDWV &DWKHULQH%LFNPRUH$VVRFLDWHV $VDUHVXOW KDDW&DQYH\:LFNZHUHGHVLJQDWHGDVD6LWHRI6SHFLDO6FLHQWLILF,QWHUHVW (QJOLVK1DWXUHD 7KHUHFUHDWHGDOHJDOREOLJDWLRQWRPDLQWDLQWKH FRPSOH[PRVDLFRIHDUO\VXFFHVVLRQDOKDELWDWV (QJOLVK1DWXUHE
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Figure 2.8 Biplot scores for staphylinid species (Ɣ), environmental variables (ĺ), and treatment plots, calculated with Canonical Correspondence Analysis of the staphylinid assemblage at Canvey Wick in 2015, as determined from captures in pitfall traps. Management treatments are coded ( ) unmanaged control; ( ) phase one restoration cleared to mineral substrate in May 2014; ( ) phase two cleared in May 2015; ( ) open mosaic habitat
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Phytophagous
Omnivorous Body size Autumn Macropterous
Threat
Brachypterous
Dimorphic PCoA axis 2 (30.3%) Spring
Predatory -1.0 -1.0 1.0 PCoA axis 1 (30.3%)
Figure 3.1 Principal Coordinates Analysis (PCoA) of the similarity between 80 species of Carabidae, based on 10 traits. Wing morphology is given as macropterous, dimorphic, or brachypterous. Trophic group is given as predatory, omnivorous, or phytophagous. Breeding period is given as spring or autumn. Body size is the median from the range given in Luff (2007). Conservation status is based on the most recent published review (Telfer, 2016). Vectors represent the Spearman correlation of traits, with the length and direction of each arrow indicating the relationship with PCoA axes. Samples were collected from Canvey Wick, Essex, in 2015 from areas of closed canopy scrub (n=6), scrub cleared and substrate scraped to mineral soil in May 2014 (phase one, n=3), scrub cleared and substrate scraped to mineral soil in May 2015 (n=6), and open mosaic habitat reference plots (n=5). Samples were pooled in each year from six pitfall traps per monitoring plot, open for nine days in June, July, and August. Species within the six most numerically abundant carabid tribes are displayed with symbols: Bembidiini ( ), Harpalini ( ), Lebiini ( ), Pterostichini ( ), Scaritini ( ), and Zabrini ( ). The remaining species are pooled into one symbol ( ).
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1.0 Phytophagous PCoA axis 2 (15.6%) Litter
Myrmecophilous Body size Mycetophilous
Threat Dung/carrion -0.4 -1.0 0.6 PCoA axis 1 (58.1%)
Figure 3.2 Principal Coordinates Analysis (PCoA) of the similarity between 99 species of Staphylinidae, based on three traits. Body size is the median from the range given in Lott (2009), Lott & Anderson (2011), and Freude et al. (1974). Adult resource preference is given as litter, myrmecophilous, phytophagous, mycetophilous, or dung and carrion. Conservation status is based on the most recent published review (Hyman, 1994). Vectors represent the Spearman correlation of traits, with the length and direction of each arrow indicating the relationship with PCoA axes. Samples were collected from Canvey Wick, Essex, in 2015 from areas of closed canopy scrub (n=6), scrub cleared and substrate scraped to mineral soil in May 2014 (phase one, n=3), scrub cleared and substrate scraped to mineral soil in May 2015 (n=6), and open mosaic habitat reference plots (n=5). Samples were pooled in each year from six pitfall traps per monitoring plot, open for nine days in June, July, and August. Species within the six most numerically abundant staphylinid tribes are displayed with unique symbols: Athetini ( ), Oxypodini ( ), Oxytelini ( ), Paederini ( ), Staphylinini ( ) and Thinobiini ( ). The remaining species are pooled into one symbol ( ).
1.0 Ballooning
Threat Running
Size
Web Ambush Moisture
PCoA axis 2 (22.4%) Shade -1.0 -1.0 1.0 PCoA axis 1 (43.3%)
Figure 3.3 Principal Coordinates Analysis (PCoA) of the similarity between 67 species of spider, based on six traits. Hunting strategy is given as running, ambush, or web, based on the guild structure in Uetz (1999). Dispersal is based on species-specific records of ballooning in Bell (2005) and Blandenier (2009). Moisture and shade perference is taken from Entling (1997). Body size is the median from the range given for females in Roberts (1985, 1987). Conservation status is based on the most recent available review (Merrett, 1990). Vectors represent the Spearman correlation of traits, with the length and direction of each arrow indicating the relationship with PCoA axes. Samples were collected from Canvey Wick, Essex, in 2015 from areas of closed canopy scrub (n=6), scrub cleared and substrate scraped to mineral soil in May 2014 (phase one, n=3), scrub cleared and substrate scraped to mineral soil in May 2015 (n=6), and open mosaic habitat reference plots (n=5). Samples were pooled in each year from six pitfall traps per monitoring plot, open for nine days in June, July, and August. Species within the six most numerically abundant spider families are displayed with unique symbols: Clubionidae ( ), Gnaphosidae ( ), Liocranidae ( ), Linyphiidae ( ), Lycosidae ( ), and Thomisidae ( ). The remaining species are pooled into one symbol ( ).
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Scraping Excavator disturbance Legend Undisturbed grassland Pitfall trap layout (3 per plot)
Figure 4.1 Experimental design for ground disturbance plots at Canvey Wick, Essex. Scraping removed all vegetation and organic soil horizons, which were taken away from the experimental area. Vegetation and humus layers were shallowly buried during excavator disturbance. Treatments were applied in November 2015. Basemap source: Esri, DigitalGlobe, GeoEye, i-cubed, USDA FSA, USGS, AEX, Getmapping, Aerogrid, IGN, IGP, swisstopo, and the GIS user community.
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Figure 4.5 Sample-based rarefaction curves of species richness for beetles and spiders, based on species likely to be captured by pitfall traps in excavator disturbance, scraping, and unmanaged grassland plots at Canvey Wick in 2016 (n=27). Shaded areas represent 95% confidence intervals. Curves become more shallow approaching the asymptotes of the estimated faunas. 7KHQXPEHURILQGLYLGXDOVGHFUHDVHGDWWKHFRQWUROSORWVWKDWZHUHVXEVHTXHQWO\ VFUDSHGIURPLQWRLQ)URPVSHFLHVWUDSSHGLQ WKHPRVWDEXQGDQWZHUHAmara lunicollis Drusilla canaliculata DQGAmara communis 7KHUHZHUHVSHFLHVUHSUHVHQWHGE\WKUHH RUIHZHULQGLYLGXDOVIRUPLQJRIWKHFDWFK)URPVSHFLHVWUDSSHGLQ WKHPRVWDEXQGDQWZHUHBledius gallicus Calodera protensa DQGXantholinus longiventris 7KHUHZHUHVSHFLHVUHSUHVHQWHGE\ WKUHHRUIHZHULQGLYLGXDOVIRUPLQJ RIWKHFDWFK
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Figure 4.6 Biplot scores for stenotopic early successional beetle species (Ɣ) and treatment plots, calculated from Non-Metric Multidimensional Scaling of abundance data at Canvey Wick ground disturbance plots, in 2015 and 2016 (stress value = 0.1448, 3 dimensions). Ordination was based on a matrix of Jaccard dissimilarities, with a maximum of 100 random starts to search for the lowest stress solution. Samples were pooled in each year from three pitfall traps per monitoring plot (n=27), open for sixteen days in May, July, and September. All plots were monitored before management in 2015. Excavator disturbance and scraping treatments were applied in November 2015 (both n=9), and monitoring was repeated in 2016. Management treatments are coded ( ) unmanaged grassland in 2015, ( ) unamanged grassland in 2015, ( ) unmanaged grassland in 2015, ( ) unmanaged grassland in 2016, ( ) excavator disturbance plots in 2016, ( ) scraped plots in 2016. Numerical references for species names: 1 = Agonum marginatum, 2 = Amara aenea, 3 = Amara convexior, 4 = Amara eurynota, 5 = Amara tibialis, 6 = Anthicus antherinus, 7 = Anthracus consputus, 8 = Bembidion lunulatum, 9 = Bembidion varium, 10 = Bledius gallicus, 11 = Bledius opacus, 12 = Brachinus crepitans, 13 = Calathus ambiguus, 14 = Carpelimus corticinus, 15 = Carpelimus erichsoni, 16 = Carpelimus gracilis, 17 = Carpelimus incongruus, 18 = Carpelimus rivularis, 19 = Curimopsis maritima, 20 = Curtonotus convexiusculus, 21 = Dyschirius aeneus, 22 = Dyschirius luedersi, 23 = Dyschirius politus, 24 = Elaphrus riparius, 25 = Harpalus rubripes, 26 = Harpalus tardus, 27 = Hippodamia variegata, 28 = Leiodes rufipennis, 29 = Melanophthalma suturalis, 30 = Neobisnius procerulus, 31 = Notiophilus substriatus, 32 = Notoxus monoceros, 33 = Otiorhynchus ovatus, 34 = Paederus littoralis, 35 = Panagaeus bipustulatus, 36 = Paradromius linearis, 37 = Philonthus quisquiliarius, 38 = Platystethus alutaceus, 39 = Platystethus nitens, 40 = Poecilus cupreus, 41 = Sitona cylindricollis, 42 = Sunius propinquus, 43 = Syntomus foveatus, 44 = Tasgius globulifer, 45 = Tasgius winkleri, 46 = Thecturota marchii, 47 = Thinonoma atra, 48 = Xantholinus elegans.
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Figure 4.7 Biplot scores for stenotopic early successional beetle species (Ɣ) and treatment plots, calculated from Non-Metric Multidimensional Scaling of abundance data at Canvey Wick ground disturbance plots, in 2015 and 2016 (stress value = 0.1295, 3 dimensions). Ordination was based on a matrix of Bray-Curtis dissimilarities, with a maximum of 100 random starts to search for the lowest stress solution. Samples were pooled in each year from three pitfall traps per monitoring plot (n=27), open for 16 days in May, July, and September. All plots were monitored before management in 2015. Excavator disturbance and scraping treatments were applied in November 2015 (both n=9), and monitoring was repeated in 2016. Management treatments are coded ( ) unmanaged grassland in 2015, ( ) unamanged grassland in 2015, ( ) unmanaged grassland in 2015, ( ) unmanaged grassland in 2016, ( ) excavator disturbance plots in 2016, ( ) scraped plots in 2016. Numerical references for species names: 1 = Agonum marginatum, 2 = Amara aenea, 3 = Amara convexior, 4 = Amara eurynota, 5 = Amara tibialis, 6 = Anthicus antherinus, 7 = Anthracus consputus, 8 = Bembidion lunulatum, 9 = Bembidion varium, 10 = Bledius gallicus, 11 = Bledius opacus, 12 = Brachinus crepitans, 13 = Calathus ambiguus, 14 = Carpelimus corticinus, 15 = Carpelimus erichsoni, 16 = Carpelimus gracilis, 17 = Carpelimus incongruus, 18 = Carpelimus rivularis, 19 = Curimopsis maritima, 20 = Curtonotus convexiusculus, 21 = Dyschirius aeneus, 22 = Dyschirius luedersi, 23 = Dyschirius politus, 24 = Elaphrus riparius, 25 = Harpalus rubripes, 26 = Harpalus tardus, 27 = Hippodamia variegata, 28 = Leiodes rufipennis, 29 = Melanophthalma suturalis, 30 = Neobisnius procerulus, 31 = Notiophilus substriatus, 32 = Notoxus monoceros, 33 = Otiorhynchus ovatus, 34 = Paederus littoralis, 35 = Panagaeus bipustulatus, 36 = Paradromius linearis, 37 = Philonthus quisquiliarius, 38 = Platystethus alutaceus, 39 = Platystethus nitens, 40 = Poecilus cupreus, 41 = Sitona cylindricollis, 42 = Sunius propinquus, 43 = Syntomus foveatus, 44 = Tasgius globulifer, 45 = Tasgius winkleri, 46 = Thecturota marchii, 47 = Thinonoma atra, 48 = Xantholinus elegans.
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Figure 4.8 Biplot scores for stenotopic early successional spider species (Ɣ) and treatment plots, calculated from Non-metric Multidimensional Scaling of presence/absence data at Canvey Wick ground disturbance plots, in 2015 and 2016 (stress value = 0.1184, 3 dimensions). Ordination was based on a matrix of Jaccard dissimilarities, with a maximum of 100 random starts to search for the lowest stress solution. Samples were pooled in each year from three pitfall traps per monitoring plot (n=27), open for 16 days in May, July,and September. All plots were monitored before management in 2015. Excavator disturbance and scraping treatments were applied in November 2015 (both n=9), and monitoring was repeated in 2016. Management treatments are coded ( ) unmanaged grassland in 2015, ( ) unamanged grassland in 2015, ( ) unmanaged grassland in 2015, ( ) unmanaged grassland in 2016, ( ) excavator disturbance plots in 2016, ( ) scraped plots in 2016. Numerical references for species names: 1 = Arctosa leopardus, 2 = Argenna subnigra, 3 = Clubiona neglecta, 4 = Drassodes cupreus, 5 = Drassyllus pusillus, 6 = Dysdera crocata, 7 = Erigone arctica, 8 = Haplodrassus signifer, 9 = Mermessus trilobatus, 10 = Ozyptila simplex, 11 = Pardosa agrestis, 12 = Sibianor aurocinctus, 13 = Trachyzelotes pedestris, 14 = Xysticus kochi, 15 = Zelotes apricorum, 16 = Zelotes latreillei, 17 = Zodarion italicum.
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Figure 5.6 Experimental design for aggregate mound creation at Rainham Marshes, Essex. Basemap source: Esri, DigitalGlobe, GeoEye, i-cubed, USDA FSA, USGS, AEX, Getmapping, Aerogrid, IGN, IGP, swisstopo, and the GIS user community.
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Legend Grassland control Thanet sand mound Recycled bulk fill mound
Figure 5.7 Experimental design for aggregate mound creation at Hadleigh Park, Essex. Basemap source: Esri, DigitalGlobe, GeoEye, i-cubed, USDA FSA, USGS, AEX, Getmapping, Aerogrid, IGN, IGP, swisstopo, and the GIS user community.
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Legend Grassland control Thanet sand mound Recycled bulk fill mound
Figure 5.8 Experimental design for aggregate mound creation at Pitsea Landfill, Essex. Basemap source: Esri, DigitalGlobe, GeoEye, i-cubed, USDA FSA, USGS, AEX, Getmapping, Aerogrid, IGN, IGP, swisstopo, and the GIS user community.
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43 11 29 52 18 58 17 71 35 20 16 34 45 51 23 63 69 7 62 66 60 46 13 10 6 4 1 14 2 37 53 24 26 64 72 56 NMDS axis 2 3 59 28 61 32 9 12 47 73 44 55 8 49 54 33 31 38 22 21 27 30 70 25 5 48 19 68 15 50 41 65 67 40 39
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Figure 5.11 Biplot scores for stenotopic early successional beetle and spider species (Ɣ) and treatment plots, calculated from Non-metric Multidimensional Scaling of presence-absence data at south Essex aggregate mounds, 2015-2016 (stress value = 0.13994, 3 dimensions). Ordination was based on a matrix of Jaccard dissimilarities, with a maximum of 100 random starts to search for the lowest stress solution. Samples were pooled in both years from two pitfall traps per monitoring plot, open for 14 days in May, July, and September. Treatments were 5-tonne mounds of recycled bulk fill (n=9) or Thanet sand (n=9), and unmanaged grassland controls (n=9). Treatments are coded recycled bulk fill ( ), Thanet sand ( ), unmanaged grassland ( ). Sites are coded Hadleigh Park ( ), Pitsea Landfill ( ), Rainham Marshes ( ). Numerical references for species names: 1 = Amara aenea, 2 = Amara convexior, 3 = Amara eurynota, 4 = Amara familiaris, 5 = Amara tibialis, 6 = Anotylus inustus, 7 = Anthicus antherinus, 8 = Argenna subnigra, 9 = Astenus lyonessius, 10 = Bembidion femoratum, 11 = Bembidion lunulatum, 12 = Brachinus crepitans, 13 = Calathus ambiguus, 14 = Carpelimus corticinus, 15 = Carpelimus incongruus, 16 = Carpelimus rivularis, 17 = Centromerus capucinus, 18 = Crustulina guttata, 19 = Curtonotus convexiusculus, 20 = Drassodes cupreus, 21 = Drassyllus pusillus, 22 = Dysdera crocata, 23 = Elaphrus riparius, 24 = Haplodrassus signifer, 25 = Harpalus anxius, 26 = Harpalus attenuatus, 27 = Harpalus rubripes, 28 = Harpalus tardus, 29 = Helophorus rufipes, 30 = Hypera fuscocinerea, 31 = Lobrathium multipunctum, 32 = Melanophthalma suturalis, 33 = Microlestes minutulus, 34 = Neobisnius villosulus, 35 = Notiophilus substriatus, 36 = Omphalapion hookerorum, 37 = Ophonus ardosiacus, 38 = Ophonus azureus, 39 = Otiorhynchus ligneus, 40 = Otiorhynchus ovatus, 41 = Otiorhynchus raucus, 42 = Oxypoda brachyptera, 43 = Oxypoda procerula, 44 = Ozyptila simplex, 45 = Paederus littoralis, 46 = Panagaeus bipustulatus, 47 = Paradromius linearis, 48 = Pardosa agrestis, 49 = Philhygra debilis, 50 = Philhygra palustris, 51 = Philorhizus melanocephalus, 52 = Phrurolithus minimus, 53 = Platyderus depressus, 54 = Platystethus alutaceus, 55 = Platystethus nitens, 56 = Poecilus cupreus, 57 = Romualdius angustisetulus, 58 = Rugilus orbiculatus, 59 = Scybalicus oblongiusculus, 60 = Sibianor aurocinctus, 61 = Sunius propinquus, 62 = Tasgius ater, 63 = Tasgius globulifer, 64 = Tasgius winkleri, 65 = Tegenaria agrestis, 66 = Trachyzelotes pedestris, 67 = Troxochrus scabriculus, 68 = Walckenaeria capito, 69 = Xantholinus elegans, 70 = Xysticus kochi, 71 = Zelotes apricorum, 72 = Zelotes latreillei, 73 = Zodarion italicum.
29
17 16
3 52 45
71 42 66 63 7 35 62 10 18 49 47 64 11 31 51 2 6 46 69 56 20 9 32
NMDS axis 2 43 54 28 22 73 61 59 53 67 19 24 4 60 13 12 37 1 23 33 55 30 50 40 57 58 44 34 65 38 72 28 70 68 15 41 39 36 26 14 21 8 48 25
5 -2 2.8 -2 3 NMDS axis 1
Figure 5.12 Biplot scores for stenotopic early successional beetle and spider species (Ɣ) and treatment plots, calculated from Non-metric Multidimensional Scaling of abundance data at south Essex aggregate mounds, in 2015- 2016 (stress value = 0.13228, 3 dimensions). Ordination was based on a matrix of Bray-Curtis dissimilarities, with a maximum of 100 random starts to search for the lowest stress solution. Samples were pooled in each year from two pitfall traps per monitoring plot, open for 14 days in May, July, and September. Treatments were 5-tonne mounds of recycled bulk fill (n=9) or Thanet sand (n=9), and unmanaged grassland controls (n=9). Treatments are coded recycled bulk fill ( ), Thanet sand ( ), unmanaged grassland ( ). Sites are coded Hadleigh Park ( ), Pitsea Landfill ( ), Rainham Marshes ( ). Numerical references for species names: 1 = Amara aenea, 2 = Amara convexior, 3 = Amara eurynota, 4 = Amara familiaris, 5 = Amara tibialis, 6 = Anotylus inustus, 7 = Anthicus antherinus, 8 = Argenna subnigra, 9 = Astenus lyonessius, 10 = Bembidion femoratum, 11 = Bembidion lunulatum, 12 = Brachinus crepitans, 13 = Calathus ambiguus, 14 = Carpelimus corticinus, 15 = Carpelimus incongruus, 16 = Carpelimus rivularis, 17 = Centromerus capucinus, 18 = Crustulina guttata, 19 = Curtonotus convexiusculus, 20 = Drassodes cupreus, 21 = Drassyllus pusillus, 22 = Dysdera crocata, 23 = Elaphrus riparius, 24 = Haplodrassus signifer, 25 = Harpalus anxius, 26 = Harpalus attenuatus, 27 = Harpalus rubripes, 28 = Harpalus tardus, 29 = Helophorus rufipes, 30 = Hypera fuscocinerea, 31 = Lobrathium multipunctum, 32 = Melanophthalma suturalis, 33 = Microlestes minutulus, 34 = Neobisnius villosulus, 35 = Notiophilus substriatus, 36 = Omphalapion hookerorum, 37 = Ophonus ardosiacus, 38 = Ophonus azureus, 39 = Otiorhynchus ligneus, 40 = Otiorhynchus ovatus, 41 = Otiorhynchus raucus, 42 = Oxypoda brachyptera, 43 = Oxypoda procerula, 44 = Ozyptila simplex, 45 = Paederus littoralis, 46 = Panagaeus bipustulatus, 47 = Paradromius linearis, 48 = Pardosa agrestis, 49 = Philhygra debilis, 50 = Philhygra palustris, 51 = Philorhizus melanocephalus, 52 = Phrurolithus minimus, 53 = Platyderus depressus, 54 = Platystethus alutaceus, 55 = Platystethus nitens, 56 = Poecilus cupreus, 57 = Romualdius angustisetulus, 58 = Rugilus orbiculatus, 59 = Scybalicus oblongiusculus, 60 = Sibianor aurocinctus, 61 = Sunius propinquus, 62 = Tasgius ater, 63 = Tasgius globulifer, 64 = Tasgius winkleri, 65 = Tegenaria agrestis, 66 = Trachyzelotes pedestris, 67 = Troxochrus scabriculus, 68 = Walckenaeria capito, 69 = Xantholinus elegans, 70 = Xysticus kochi, 71 = Zelotes apricorum, 72 = Zelotes latreillei, 73 = Zodarion italicum.
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3
13 3 15 38 37 49 50 8 5 39 45 6 43 14 9 11 22 23
20 52 41 44 10 18 53 27 NMDS axis 2 26 28 51 7 30 17 1 32 4 42 21 47 2 31 46 36 24 33 48 25 34 16 19 29 40 12 35 -2 -1.5 2.5 NMDS axis 1
Figure 5.13 Biplot scores for aculeate Hymenoptera species (Ɣ) and treatment plots, calculated from Non-metric Multidimensional Scaling of presence-absence data at aggregate mounds in south Essex, 2015-2016 (stress value = 0.1417, 3 dimensions). Ordination was based on a matrix of Jaccard dissimilarities, with a maximum of 100 random starts to search for the lowest stress solution. Samples were pooled from both years, based on 30 minutes hand netting per plot (n=27), during three visits in May, June, and August. Treatments were 5-tonne mounds of recycled bulk fill or Thanet sand, and unmanaged grassland. There was zero data from unmanaged grassland plots, and these were not included in the ordation. Treatments are coded recycled bulk fill ( ), Thanet sand ( ). Sites are coded Hadleigh Park ( ), Pitsea Landfill ( ), Rainham Marshes ( ). Numerical references for species names: 1 = Ammophila sabulosa, 2 = Andrena chrysosceles, 3 = Andrena dorsata, 4 = Andrena flavipes, 5 = Andrena haemorrhoa, 6 = Andrena labialis, 7 = Andrena minutula, 8 = Andrena nigroaenea, 9 = Andrena nitida, 10 = Andrena praecox, 11 = Andrena scotica, 12 = Anoplius concinnus, 13 = Anthophora bimaculata, 14 = Arachnospila anceps, 15 = Auplopus carbonarius, 16 = Cerceris arenaria, 17 = Cerceris quinquefasciata, 18 = Cerceris rybyensis, 19 = Crossocerus wesmaeli, 20 = Entomognathus brevis, 21 = Episyron rufipes, 22 = Gorytes quadrifasciatus, 23 = Halictus rubicundus, 24 = Hylaeus communis, 25 = Hylaeus hyalinatus, 26 = Lasioglossum albipes, 27 = Lasioglossum calceatum, 28 = Lasioglossum leucozonium, 29 = Lasioglossum malachurum, 30 = Lasioglossum minutissimum, 31 = Lasioglossum morio, 32 = Lasioglossum parvulum, 33 = Lasioglossum pauperatum, 34 = Lasioglossum pauxillum, 35 = Lasioglossum puncticolle, 36 = Lasioglossum villosulum, 37 = Melitta leporina, 38 = Mimumesa unicolor, 39 = Nomada flava, 40 = Nomada flavoguttata, 41 = Nomada flavopicta, 42 = Nomada goodeniana, 43 = Nomada marshamella, 44 = Oxybelus uniglumis, 45 = Philanthus triangulum, 46 = Priocnemis pusilla, 47 = Sphecodes ephippius, 48 = Sphecodes geoffrellus, 49 = Sphecodes gibbus, 50 = Sphecodes longulus, 51 = Sphecodes monilicornis, 52 = Sphecodes puncticeps, 53 = Trypoxylon medium.
3
12 35 34 29 25 40 31 32 36 19 21 26 42 24 2 18 46 33 16
NMDS axis 2 17 48 7 1 47 41 30 44 23 4 20 28 53 52 39 10 5 27 22 6 14 8 50 13 351 374549 15 43 11 9
38 -2 -2 3 NMDS axis 1
Figure 5.14 Biplot scores for aculeate Hymenoptera species (Ɣ) and treatment plots, calculated from Non-metric Multidimensional Scaling of abundance data at aggregate mounds in south Essex, 2015-2016 (stress value = 0.1261, 3 dimensions). Ordination was based on a matrix of Bray-Curtis dissimilarities, with a maximum of 100 random starts to search for the lowest stress solution. Samples were pooled from both years, based on 30 minutes hand netting per plot, during three visits in May, June, and August. Treatments were 5-tonne mounds of recycled bulk fill (n=9) or Thanet sand (n=9), and unmanaged grassland (n=9). There was zero data from unmanaged grassland plots, and these were not included in the ordation. Treatments are coded recycled bulk fill ( ), Thanet sand ( ). Sites are coded Hadleigh Park ( ), Pitsea Landfill ( ), Rainham Marshes ( ). Numerical references for species names: 1 = Ammophila sabulosa, 2 = Andrena chrysosceles, 3 = Andrena dorsata, 4 = Andrena flavipes, 5 = Andrena haemorrhoa, 6 = Andrena labialis, 7 = Andrena minutula, 8 = Andrena nigroaenea, 9 = Andrena nitida, 10 = Andrena praecox, 11 = Andrena scotica, 12 = Anoplius concinnus, 13 = Anthophora bimaculata, 14 = Arachnospila anceps, 15 = Auplopus carbonarius, 16 = Cerceris arenaria, 17 = Cerceris quinquefasciata, 18 = Cerceris rybyensis, 19 = Crossocerus wesmaeli, 20 = Entomognathus brevis, 21 = Episyron rufipes, 22 = Gorytes quadrifasciatus, 23 = Halictus rubicundus, 24 = Hylaeus communis, 25 = Hylaeus hyalinatus, 26 = Lasioglossum albipes, 27 = Lasioglossum calceatum, 28 = Lasioglossum leucozonium, 29 = Lasioglossum malachurum, 30 = Lasioglossum minutissimum, 31 = Lasioglossum morio, 32 = Lasioglossum parvulum, 33 = Lasioglossum pauperatum, 34 = Lasioglossum pauxillum, 35 = Lasioglossum puncticolle, 36 = Lasioglossum villosulum, 37 = Melitta leporina, 38 = Mimumesa unicolor, 39 = Nomada flava, 40 = Nomada flavoguttata, 41 = Nomada flavopicta, 42 = Nomada goodeniana, 43 = Nomada marshamella, 44 = Oxybelus uniglumis, 45 = Philanthus triangulum, 46 = Priocnemis pusilla, 47 = Sphecodes ephippius, 48 = Sphecodes geoffrellus, 49 = Sphecodes gibbus, 50 = Sphecodes longulus, 51 = Sphecodes monilicornis, 52 = Sphecodes puncticeps, 53 = Trypoxylon medium.
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detritivorous
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phytophagous -0.6 0.8 -1.5 1.5 Axis 1
Figure 5.15 Biplot scores for invertebrate functional groups (ĺ), environmental variables (ĺ), and treatment plots, calculated with Redundancy Analysis (RDA) of beetle and spider assemblages from the aggregate mound creation experiment in 2015. Samples were pooled from two pitfall traps per monitoring plot (n=27), open for 14 days in May, July, and September. Treatments were 5-tonne mounds of recycled bulk fill or Thanet sand, and unmanaged grassland controls. Treatments are coded recycled bulk fill ( ), Thanet sand ( ), unmanaged grassland ( ). Sites are coded Hadleigh Park ( ), Pitsea Landfill ( ), Rainham Marshes ( ). Environmental variables are coded B, bare ground; H, herbaceous plant species richness; A, annual plant species richness; P, perennial plant species richess; G, graminoid species richess; L, litter depth. RDA scaling was focused on inter-species correlations and centred by species. Treatments and sites were included in the final triplot but were not used to influence the analysis. Relationships between environmental variables and distribution of functional groups were tested by forward selection (Monte Carlo test, 500 permutations). Permutations were made within but not between whole plots that were
defined by sites. Significant variables were percentage of bare ground (Padj = 0.014), graminoid
species richness (Padj = 0.024) and herbaceous plant species richness (Padj = 0.014). Other environmental variables either correlated with these three, or did not describe a significant proportion of the variation in the ordination.
phytophagous
P zoophagous predatory spider HG A Axis 2
detritivorous B
predatory beetle L -0.8 0.8 -1.0 1.5 Axis 1
Figure 5.16 Biplot scores for invertebrate functional groups (ĺ), environmental variables (ĺ), and treatment plots, calculated with Redundancy Analysis (RDA) of beetle and spider assemblages from the aggregate mounds experiment in 2016. Samples were pooled from two pitfall traps per monitoring plot (n=27), open for 14 days in May, July, and September. Treatments were 5-tonne mounds of recycled bulk fill or Thanet sand, and unmanaged grassland controls. Treatments are coded recycled bulk fill ( ), Thanet sand ( ), unmanaged grassland ( ). Sites are coded Hadleigh Park ( ), Pitsea Landfill ( ), Rainham Marshes ( ). Environmental variables are coded B, bare ground; H, herbaceous plant species richness; A, annual plant species richness; P, perennial plant species richess; G, graminoid species richess; L, litter depth. RDA scaling was focused on inter-species correlations and centred by species. Treatments and sites were included in the final triplot but were not used to influence the analysis. Relationships between environmental variables and distribution of functional groups were tested by forward selection (Monte Carlo test, 500 permutations). Permutations were made within but not between whole plots that were defined by sites.
The significant variable was litter depth (Padj = 0.014). Other environmental variables either correlated with this, or did not describe a significant proportion of the variation in the ordination.
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6WHSKHQV 6WHQRWRSLF 3K\WRSKDJRXV &DUDELGDH Amara eurynota 3DQ]HU 6WHQRWRSLF 3K\WRSKDJRXV &DUDELGDH Amara lunicollis 6FKL|GWH *HQHUDOLVW 3K\WRSKDJRXV &DUDELGDH Amara ovata )DEULFLXV *HQHUDOLVW 3K\WRSKDJRXV &DUDELGDH Amara tibialis 3D\NXOO 6WHQRWRSLF 3K\WRSKDJRXV &DUDELGDH Anisodactylus binotatus )DEULFLXV *HQHUDOLVW 3K\WRSKDJRXV &DUDELGDH Anthracus consputus 'XIWVFKPLG 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Badister bullatus 6FKUDQN *HQHUDOLVW =RRSKDJRXV &DUDELGDH Bembidion assimile *\OOHQKDO *HQHUDOLVW =RRSKDJRXV &DUDELGDH Bembidion biguttatum )DEULFLXV *HQHUDOLVW =RRSKDJRXV &DUDELGDH Bembidion lunulatum *HRIIUR\LQ)RXUFUR\ 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Bembidion minimum )DEULFLXV *HQHUDOLVW =RRSKDJRXV &DUDELGDH Bembidion properans 6WHSKHQV *HQHUDOLVW =RRSKDJRXV &DUDELGDH Bembidion quadrimaculatum /LQQDHXV *HQHUDOLVW =RRSKDJRXV &DUDELGDH Bembidion varium 2OLYLHU 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Brachinus crepitans /LQQDHXV 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Bradycellus harpalinus $XGLQHW6HUYLOOH (XU\WRSLF 3K\WRSKDJRXV &DUDELGDH Bradycellus verbasci 'XIWVFKPLG *HQHUDOLVW 3K\WRSKDJRXV &DUDELGDH Calathus ambiguus 3D\NXOO 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Calathus fuscipes *RH]H *HQHUDOLVW =RRSKDJRXV &DUDELGDH Calathus melanocephalus /LQQDHXV *HQHUDOLVW =RRSKDJRXV &DUDELGDH Carabus nemoralis 0OOHU2) *HQHUDOLVW =RRSKDJRXV )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO &DUDELGDH Carabus violaceus /LQQDHXV *HQHUDOLVW =RRSKDJRXV &DUDELGDH Clivina fossor /LQQDHXV *HQHUDOLVW =RRSKDJRXV &DUDELGDH Curtonotus aulicus 3DQ]HU *HQHUDOLVW 3K\WRSKDJRXV &DUDELGDH Curtonotus convexiusculus 0DUVKDP 6WHQRWRSLF 3K\WRSKDJRXV &DUDELGDH Demetrias atricapillus /LQQDHXV *HQHUDOLVW =RRSKDJRXV &DUDELGDH Dyschirius aeneus 'HMHDQ 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Dyschirius luedersi :DJQHU 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Dyschirius politus 'HMHDQ 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Elaphrus riparius /LQQDHXV 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Harpalus affinis 6FKUDQN *HQHUDOLVW 3K\WRSKDJRXV &DUDELGDH Harpalus rubripes 'XIWVFKPLG 6WHQRWRSLF 3K\WRSKDJRXV &DUDELGDH Harpalus rufipes 'H*HHU (XU\WRSLF 3K\WRSKDJRXV &DUDELGDH Harpalus tardus 3DQ]HU 6WHQRWRSLF 3K\WRSKDJRXV &DUDELGDH Leistus ferrugineus /LQQDHXV *HQHUDOLVW =RRSKDJRXV &DUDELGDH Leistus fulvibarbis 'HMHDQ *HQHUDOLVW =RRSKDJRXV &DUDELGDH Loricera pilicornis )DEULFLXV :RRGODQG =RRSKDJRXV &DUDELGDH Microlestes maurus 6WXUP *HQHUDOLVW =RRSKDJRXV &DUDELGDH Nebria brevicollis )DEULFLXV *HQHUDOLVW =RRSKDJRXV &DUDELGDH Notiophilus biguttatus )DEULFLXV (XU\WRSLF =RRSKDJRXV &DUDELGDH Notiophilus palustris 'XIWVFKPLG *HQHUDOLVW =RRSKDJRXV &DUDELGDH Notiophilus substriatus :DWHUKRXVH*5 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Ophonus puncticeps 6WHSKHQV *HQHUDOLVW 3K\WRSKDJRXV &DUDELGDH Ophonus rufibarbis )DEULFLXV *HQHUDOLVW 3K\WRSKDJRXV &DUDELGDH Panagaeus bipustulatus )DEULFLXV 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Paradromius linearis 2OLYLHU 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Poecilus cupreus /LQQDHXV 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Pterostichus gracilis 'HMHDQ *HQHUDOLVW =RRSKDJRXV )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO &DUDELGDH Pterostichus melanarius ,OOLJHU *HQHUDOLVW =RRSKDJRXV &DUDELGDH Pterostichus niger 6FKDOOHU *HQHUDOLVW =RRSKDJRXV &DUDELGDH Pterostichus nigrita 3D\NXOO *HQHUDOLVW =RRSKDJRXV &DUDELGDH Pterostichus strenuus 3DQ]HU *HQHUDOLVW =RRSKDJRXV &DUDELGDH Pterostichus vernalis 3DQ]HU *HQHUDOLVW =RRSKDJRXV &DUDELGDH Stenolophus mixtus +HUEVW *HQHUDOLVW 3K\WRSKDJRXV &DUDELGDH Stenolophus teutonus 6FKUDQN *HQHUDOLVW 3K\WRSKDJRXV &DUDELGDH Stomis pumicatus 3DQ]HU *HQHUDOLVW =RRSKDJRXV &DUDELGDH Syntomus foveatus *HRIIUR\LQ)RXUFUR\ 6WHQRWRSLF =RRSKDJRXV &DUDELGDH Trechus obtusus (ULFKVRQ (XU\WRSLF =RRSKDJRXV &DUDELGDH Trechus quadristriatus 6FKUDQN (XU\WRSLF =RRSKDJRXV &HUDPE\FLGDH Pseudovadonia livida )DEULFLXV *HQHUDOLVW 3K\WRSKDJRXV &KU\VRPHOLGDH Altica palustris :HLVH *HQHUDOLVW 3K\WRSKDJRXV &KU\VRPHOLGDH Bruchus atomarius /LQQDHXV *HQHUDOLVW 3K\WRSKDJRXV &KU\VRPHOLGDH Bruchus loti 3D\NXOO *HQHUDOLVW 3K\WRSKDJRXV &KU\VRPHOLGDH Bruchus rufipes +HUEVW *HQHUDOLVW 3K\WRSKDJRXV &KU\VRPHOLGDH Chaetocnema concinna 0DUVKDP (XU\WRSLF 3K\WRSKDJRXV &KU\VRPHOLGDH Chaetocnema hortensis )RXUFUR\ (XU\WRSLF 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus dorsalis )DEULFLXV *HQHUDOLVW 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus parvulus 3D\NXOO (XU\WRSLF 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus suturellus 'XIWVFKPLG (XU\WRSLF 3K\WRSKDJRXV &KU\VRPHOLGDH Neocrepidodera ferruginea 6FRSROL (XU\WRSLF 3K\WRSKDJRXV &KU\VRPHOLGDH Podagrica fuscipes )DEULFLXV (XU\WRSLF 3K\WRSKDJRXV &ODPELGDH Clambus pubescens 5HGWHQEDFKHU *HQHUDOLVW 8QNQRZQ &RFFLQHOOLGDH Coccinella septempunctata /LQQDHXV (XU\WRSLF 3UHGDWRU\ &RFFLQHOOLGDH Harmonia axyridis 3DOODV (XU\WRSLF 3UHGDWRU\ &RFFLQHOOLGDH Hippodamia variegata *RH]H 6WHQRWRSLF 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO &RFFLQHOOLGDH Nephus redtenbacheri 0XOVDQW *HQHUDOLVW 3UHGDWRU\ &RFFLQHOOLGDH Psyllobora vigintiduopunctata /LQQDHXV (XU\WRSLF 3UHGDWRU\ &RFFLQHOOLGDH Rhyzobius litura )DEULFLXV *HQHUDOLVW 3UHGDWRU\ &RFFLQHOOLGDH Subcoccinella vigintiquattuorpunctata /LQQDHXV *HQHUDOLVW 3UHGDWRU\ &RFFLQHOOLGDH Tytthaspis sedecimpunctata /LQQDHXV (XU\WRSLF 3UHGDWRU\ &RFFLQHOOLGDH Tytthaspis sedecimpunctata /LQQDHXV *HQHUDOLVW 3UHGDWRU\ &RU\ORSKLGDH Corylophus sublaevipennis -DFTXHOLQGX9DO *HQHUDOLVW 'HWULWLYRURXV &U\SWRSKDJLGDH Atomaria testacea 6WHSKHQV (XU\WRSLF 'HWULWLYRURXV &U\SWRSKDJLGDH Cryptophagus distinguendus 6WXUP (XU\WRSLF 'HWULWLYRURXV &U\SWRSKDJLGDH Ootypus globosus :DOWO 8QNQRZQ 'HWULWLYRURXV &XUFXOLRQLGDH Barypeithes pellucidus %RKHPDQ (XU\WRSLF 3K\WRSKDJRXV &XUFXOLRQLGDH Ceutorhynchus contractus 0DUVKDP (XU\WRSLF 3K\WRSKDJRXV &XUFXOLRQLGDH Euophryum confine %URXQ (XU\WRSLF 3K\WRSKDJRXV &XUFXOLRQLGDH Glocianus punctiger *\OOHQKDO *HQHUDOLVW 3K\WRSKDJRXV &XUFXOLRQLGDH Hypera meles )DEULFLXV *HQHUDOLVW 3K\WRSKDJRXV &XUFXOLRQLGDH Hypera plantaginis 'H*HHU *HQHUDOLVW 3K\WRSKDJRXV &XUFXOLRQLGDH Otiorhynchus ovatus /LQQDHXV 6WHQRWRSLF 3K\WRSKDJRXV &XUFXOLRQLGDH Otiorhynchus rugosostriatus *RH]H (XU\WRSLF 3K\WRSKDJRXV &XUFXOLRQLGDH Rhinoncus pericarpius /LQQDHXV (XU\WRSLF 3K\WRSKDJRXV &XUFXOLRQLGDH Rhinoncus perpendicularis 5HLFK (XU\WRSLF 3K\WRSKDJRXV &XUFXOLRQLGDH Sitona cylindricollis )nKUDHXV 6WHQRWRSLF 3K\WRSKDJRXV &XUFXOLRQLGDH Sitona humeralis 6WHSKHQV *HQHUDOLVW 3K\WRSKDJRXV &XUFXOLRQLGDH Sitona lepidus *\OOHQKDO (XU\WRSLF 3K\WRSKDJRXV &XUFXOLRQLGDH Sitona lineatus /LQQDHXV (XU\WRSLF 3K\WRSKDJRXV &XUFXOLRQLGDH Sitona sulcifrons 7KXQEHUJ (XU\WRSLF 3K\WRSKDJRXV &XUFXOLRQLGDH Tanymecus palliatus )DEULFLXV *HQHUDOLVW 3K\WRSKDJRXV &XUFXOLRQLGDH Tychius junceus 5HLFK *HQHUDOLVW 3K\WRSKDJRXV )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO &XUFXOLRQLGDH Tychius picirostris )DEULFLXV *HQHUDOLVW 3K\WRSKDJRXV '\WLVFLGDH Agabus bipustulatus /LQQDHXV (XU\WRSLF 3UHGDWRU\ '\WLVFLGDH Agabus nebulosus )RUVWHU *HQHUDOLVW 3UHGDWRU\ '\WLVFLGDH Hydroporus planus )DEULFLXV *HQHUDOLVW 3UHGDWRU\ (ODWHULGDH Agriotes lineatus /LQQDHXV (XU\WRSLF 3K\WRSKDJRXV (ODWHULGDH Agriotes sputator /LQQDHXV (XU\WRSLF 3K\WRSKDJRXV (ODWHULGDH Agrypnus murinus /LQQDHXV *HQHUDOLVW 3K\WRSKDJRXV (ULUKLQLGDH Grypus equiseti )DEULFLXV *HQHUDOLVW 3K\WRSKDJRXV (ULUKLQLGDH Notaris acridulus /LQQDHXV *HQHUDOLVW 3K\WRSKDJRXV (ULUKLQLGDH Notaris scirpi )DEULFLXV *HQHUDOLVW 3K\WRSKDJRXV (URW\OLGDH Cryptophilus integer +HHU (XU\WRSLF 8QNQRZQ +HORSKRULGDH Helophorus aequalis 7KRPVRQ&* *HQHUDOLVW 'HWULWLYRURXV +HORSKRULGDH Helophorus alternans *HQp *HQHUDOLVW 'HWULWLYRURXV +HORSKRULGDH Helophorus brevipalpis %HGHO *HQHUDOLVW 'HWULWLYRURXV +HORSKRULGDH Helophorus grandis ,OOLJHU *HQHUDOLVW 'HWULWLYRURXV +HORSKRULGDH Helophorus minutus )DEULFLXV *HQHUDOLVW 'HWULWLYRURXV +HORSKRULGDH Helophorus obscurus 0XOVDQW *HQHUDOLVW 'HWULWLYRURXV +LVWHULGDH Hister unicolor /LQQDHXV *HQHUDOLVW 3UHGDWRU\ +LVWHULGDH Kissister minimus $XEp *HQHUDOLVW 3UHGDWRU\ +LVWHULGDH Margarinotus purpurascens +HUEVW *HQHUDOLVW 3UHGDWRU\ +LVWHULGDH Saprinus aeneus )DEULFLXV *HQHUDOLVW 3UHGDWRU\ +\GUDHQLGDH Ochthebius minimus )DEULFLXV *HQHUDOLVW 'HWULWLYRURXV +\GURSKLOLGDH Anacaena limbata )DEULFLXV *HQHUDOLVW 'HWULWLYRURXV +\GURSKLOLGDH Cercyon lateralis 0DUVKDP (XU\WRSLF 'HWULWLYRURXV +\GURSKLOLGDH Cercyon sternalis 6KDUS *HQHUDOLVW 'HWULWLYRURXV +\GURSKLOLGDH Cercyon tristis ,OOLJHU *HQHUDOLVW 'HWULWLYRURXV +\GURSKLOLGDH Hydrobius fuscipes /LQQDHXV (XU\WRSLF 'HWULWLYRURXV )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO +\GURSKLOLGDH Megasternum concinnum 0DUVKDP (XU\WRSLF 'HWULWLYRURXV +\GURSKLOLGDH Sphaeridium bipustulatum )DEULFLXV (XU\WRSLF 'HWULWLYRURXV .DWHUHWLGDH Brachypterus glaber 1HZPDQ (XU\WRSLF 3K\WRSKDJRXV /DWULGLLGDH Corticarina minuta )DEULFLXV (XU\WRSLF 'HWULWLYRURXV /DWULGLLGDH Enicmus transversus 2OLYLHU (XU\WRSLF 'HWULWLYRURXV /DWULGLLGDH Melanophthalma suturalis 0DQQHUKHLP 6WHQRWRSLF 'HWULWLYRURXV /HLRGLGDH Agathidium laevigatum (ULFKVRQ *HQHUDOLVW 'HWULWLYRURXV /HLRGLGDH Catops fuscus 3DQ]HU *HQHUDOLVW 'HWULWLYRURXV /HLRGLGDH Catops grandicollis (ULFKVRQ *HQHUDOLVW 'HWULWLYRURXV /HLRGLGDH Catops morio )DEULFLXV *HQHUDOLVW 'HWULWLYRURXV /HLRGLGDH Catops nigricans 6SHQFH *HQHUDOLVW 'HWULWLYRURXV /HLRGLGDH Choleva agilis ,OOLJHU *HQHUDOLVW 'HWULWLYRURXV /HLRGLGDH Choleva oblonga /DWUHLOOH *HQHUDOLVW 'HWULWLYRURXV /HLRGLGDH Leiodes rufipennis 3D\NXOO 6WHQRWRSLF 'HWULWLYRURXV /HLRGLGDH Ptomaphagus subvillosus *RH]H *HQHUDOLVW 'HWULWLYRURXV /HLRGLGDH Sciodrepoides watsoni 6SHQFH *HQHUDOLVW 'HWULWLYRURXV 0DODFKLLGDH Cordylepherus viridis )DEULFLXV *HQHUDOLVW 3UHGDWRU\ 1LWLGXOLGDH Meligethes aeneus )DEULFLXV (XU\WRSLF 3K\WRSKDJRXV 1LWLGXOLGDH Meligethes carinulatus )|UVWHU *HQHUDOLVW 3K\WRSKDJRXV 1LWLGXOLGDH Meligethes nigrescens 6WHSKHQV (XU\WRSLF 3K\WRSKDJRXV 2HGHPHULGDH Oedemera lurida 0DUVKDP (XU\WRSLF 3K\WRSKDJRXV 3WLOLLGDH Acrotrichis fascicularis +HUEVW *HQHUDOLVW 'HWULWLYRURXV 6FDUDEDHLGDH Aphodius plagiatus /LQQDHXV (XU\WRSLF 'HWULWLYRURXV 6FDUDEDHLGDH Aphodius sphacelatus 3DQ]HU (XU\WRSLF 'HWULWLYRURXV 6FDUDEDHLGDH Hoplia philanthus )HVVO\ *HQHUDOLVW 'HWULWLYRURXV 6LOSKLGDH Nicrophorus vespillo /LQQDHXV (XU\WRSLF 'HWULWLYRURXV 6LOSKLGDH Silpha laevigata )DEULFLXV *HQHUDOLVW 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO 6LOSKLGDH Silpha tristis ,OOLJHU *HQHUDOLVW 3UHGDWRU\ 6LOSKLGDH Thanatophilus sinuatus )DEULFLXV *HQHUDOLVW 'HWULWLYRURXV 6WDSK\OLQLGDH Aleochara bipustulata /LQQDHXV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Aleochara brevipennis *UDYHQKRUVW (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Aleochara curtula *RH]H (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Aloconota gregaria (ULFKVRQ (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Amischa analis *UDYHQKRUVW (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Amischa decipiens 6KDUS (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Anotylus insecatus *UDYHQKRUVW (XU\WRSLF 'HWULWLYRURXV 6WDSK\OLQLGDH Anotylus nitidulus *UDYHQKRUVW (XU\WRSLF 'HWULWLYRURXV 6WDSK\OLQLGDH Anotylus rugosus )DEULFLXV (XU\WRSLF 'HWULWLYRURXV 6WDSK\OLQLGDH Anotylus sculpturatus *UDYHQKRUVW (XU\WRSLF 'HWULWLYRURXV 6WDSK\OLQLGDH Atheta aquatica 7KRPVRQ (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Bledius gallicus *UDYHQKRUVW 6WHQRWRSLF 3K\WRSKDJRXV 6WDSK\OLQLGDH Bledius opacus %ORFN 6WHQRWRSLF 3K\WRSKDJRXV 6WDSK\OLQLGDH Brachygluta fossulata 5HLFKHQEDFK *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Callicerus obscurus *UDYHQKRUVW *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Callicerus rigidicornis (ULFKVRQ *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Calodera protensa 0DQQHUKHLP *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Carpelimus corticinus *UDYHQKRUVW 6WHQRWRSLF 'HWULWLYRURXV 6WDSK\OLQLGDH Carpelimus erichsoni 6KDUS 6WHQRWRSLF 'HWULWLYRURXV 6WDSK\OLQLGDH Carpelimus gracilis 0DQQHUKHLP 6WHQRWRSLF 'HWULWLYRURXV 6WDSK\OLQLGDH Carpelimus incongruus 6WHHO 6WHQRWRSLF 'HWULWLYRURXV 6WDSK\OLQLGDH Carpelimus rivularis 0RWVFKXOVN\ 6WHQRWRSLF 'HWULWLYRURXV 6WDSK\OLQLGDH Cypha longicornis 3D\NXOO (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Dimetrota atramentaria *\OOHQKDO (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Dinaraea angustula *\OOHQKDO (XU\WRSLF 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO 6WDSK\OLQLGDH Drusilla canaliculata )DEULFLXV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Gabrius appendiculatus 6KDUS *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Gabrius breviventer 6SHUN *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Gyrohypnus wagneri 6FKHHUSHOW] 8QNQRZQ 3UHGDWRU\ 6WDSK\OLQLGDH Ilyobates bennetti 'RQLVWKRUSH *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Ischnosoma splendidum *UDYHQKRUVW *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Lamprinodes saginatus *UDYHQKRUVW *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Lathrobium brunnipes )DEULFLXV *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Lathrobium fulvipenne *UDYHQKRUVW *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Lathrobium geminum .UDDW] *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Liogluta longiuscula *UDYHQKRUVW (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Liogluta microptera 7KRPVRQ&* (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Liogluta pagana (ULFKVRQ *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Lobrathium multipunctum *UDYHQKRUVW *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Meotica exilis .QRFK *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Microdota aegra +HHU 8QNQRZQ 3UHGDWRU\ 6WDSK\OLQLGDH Mycetoporus /XQGVWU|P 8QNQRZQ 3UHGDWRU\ 6WDSK\OLQLGDH Neobisnius procerulus *UDYHQKRUVW 6WHQRWRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Ochthephilum collare 5HLWWHU *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Ocypus aeneocephalus 'H*HHU *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Ocypus fuscatus *UDYHQKRUVW *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Ocypus olens 0OOHU2) (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Oligota pumilio .LHVHQZHWWHU (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Ontholestes murinus /LQQDHXV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Othius laeviusculus 6WHSKHQV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Othius punctulatus *RH]H *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Oxypoda brevicornis 6WHSKHQV (XU\WRSLF 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO 6WDSK\OLQLGDH Oxypoda elongatula $XEp *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Oxypoda procerula 0DQQHUKHLP *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Paederus littoralis *UDYHQKRUVW 6WHQRWRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Pella limbata 3D\NXOO *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Philhygra elongatula *UDYHQKRUVW *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Philhygra volans 6FULED *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus carbonarius *UDYHQKRUVW *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus cognatus 6WHSKHQV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus cruentatus *PHOLQLQ/LQQDHXV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus debilis *UDYHQKRUVW (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus marginatus 0OOHU2) *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus micans *UDYHQKRUVW *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus quisquiliarius *\OOHQKDO 6WHQRWRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus succicola 7KRPVRQ&* *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus varians 3D\NXOO (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Platydracus stercorarius 2OLYLHU *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Platystethus alutaceus 7KRPVRQ&* 6WHQRWRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Platystethus nitens 6DKOEHUJ 6WHQRWRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Quedius curtipennis %HUQKDXHU *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Quedius levicollis %UXOOp *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Quedius maurorufus *UDYHQKRUVW *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Quedius molochinus *UDYHQKRUVW (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Quedius nemoralis %DXGL (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Quedius nigriceps .UDDW] *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Quedius persimilis 0XOVDQW 5H\ (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Quedius picipes 0DQQHUKHLP (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Quedius schatzmayri *ULGHOOL (XU\WRSLF 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO 6WDSK\OLQLGDH Quedius semiaeneus 6WHSKHQV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Quedius semiobscurus 0DUVKDP (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Rugilus erichsonii )DXYHO *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Rugilus rufipes *HUPDU *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Sepedophilus immaculatus 6WHSKHQV *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Sepedophilus marshami 6WHSKHQV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Sepedophilus nigripennis 6WHSKHQV *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Staphylinus dimidiaticornis *HPPLQJHU (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Stenichnus collaris 0OOHU3:- .XQ]H 8QNQRZQ 3UHGDWRU\ 6WDSK\OLQLGDH Stenichnus scutellaris 0OOHU3:- .XQ]H 8QNQRZQ 3UHGDWRU\ 6WDSK\OLQLGDH Stenus bimaculatus *\OOHQKDO *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Stenus brunnipes 6WHSKHQV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Stenus clavicornis 6FRSROL (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Stenus fulvicornis 6WHSKHQV *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Stenus juno 3D\NXOO *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Stenus nanus 6WHSKHQV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Stenus ossium 6WHSKHQV *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Stenus pusillus 6WHSKHQV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Sunius melanocephalus )DEULFLXV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Sunius propinquus %ULVRXWGH%DUQHYLOOH 6WHQRWRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Tachinus flavolimbatus 3DQGHOOp *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Tachinus laticollis *UDYHQKRUVW *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Tachinus rufipes /LQQDHXV *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Tachyporus atriceps 6WHSKHQV *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Tachyporus chrysomelinus /LQQDHXV *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Tachyporus dispar 3D\NXOO (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Tachyporus hypnorum )DEULFLXV (XU\WRSLF 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO 6WDSK\OLQLGDH Tachyporus nitidulus )DEULFLXV (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Tasgius globulifer *HRIIUR\ 6WHQRWRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Tasgius melanarius +HHU *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Tasgius morsitans 5RVVL *HQHUDOLVW 3UHGDWRU\ 6WDSK\OLQLGDH Tasgius winkleri %HUQKDXHU 6WHQRWRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Thecturota marchii 'RGHUR 6WHQRWRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Thinonoma atra *UDYHQKRUVW 6WHQRWRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Tinotus morion *UDYHQKRUVW (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Xantholinus elegans 2OLYLHU 6WHQRWRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Xantholinus linearis 2OLYLHU (XU\WRSLF 3UHGDWRU\ 6WDSK\OLQLGDH Xantholinus longiventris +HHU (XU\WRSLF 3UHGDWRU\ 7HQHEULRQLGDH Helops caeruleus /LQQDHXV 8QNQRZQ 'HWULWLYRURXV 7KURVFLGDH Trixagus obtusus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gelena labyrinthica &OHUFN *HQHUDOLVW 3UHGDWRU\ $UDQHLGDH Hypsosinga pygmaea 6XQGHYDOO *HQHUDOLVW 3UHGDWRU\ &OXELRQLGDH Clubiona diversa 23&DPEULGJH *HQHUDOLVW 3UHGDWRU\ &OXELRQLGDH Clubiona neglecta 23&DPEULGJH 6WHQRWRSLF 3UHGDWRU\ &OXELRQLGDH Clubiona reclusa 23&DPEULGJH (XU\WRSLF 3UHGDWRU\ 'LFW\QLGDH Argenna subnigra 23&DPEULGJH 6WHQRWRSLF 3UHGDWRU\ 'LFW\QLGDH Dictyna latens )DEULFLXV *HQHUDOLVW 3UHGDWRU\ '\VGHULGDH Dysdera crocata &/.RFK 6WHQRWRSLF 3UHGDWRU\ *QDSKRVLGDH Drassodes cupreus %ODFNZDOO 6WHQRWRSLF 3UHGDWRU\ *QDSKRVLGDH Drassodes pubescens 7KRUHOO *HQHUDOLVW 3UHGDWRU\ *QDSKRVLGDH Drassyllus pusillus &/.RFK 6WHQRWRSLF 3UHGDWRU\ *QDSKRVLGDH Haplodrassus signifer &/.RFK 6WHQRWRSLF 3UHGDWRU\ *QDSKRVLGDH Micaria pulicaria 6XQGHYDOO (XU\WRSLF 3UHGDWRU\ *QDSKRVLGDH Trachyzelotes pedestris &/.RFK 6WHQRWRSLF 3UHGDWRU\ *QDSKRVLGDH Zelotes apricorum /.RFK 6WHQRWRSLF 3UHGDWRU\ *QDSKRVLGDH Zelotes latreillei 6LPRQ 6WHQRWRSLF 3UHGDWRU\ +DKQLLGDH Antistea elegans %ODFNZDOO *HQHUDOLVW 3UHGDWRU\ +DKQLLGDH Hahnia nava %ODFNZDOO (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Bathyphantes gracilis %ODFNZDOO (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Bathyphantes parvulus :HVWULQJ (XU\WRSLF 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO /LQ\SKLLGDH Centromerita bicolor %ODFNZDOO (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Centromerita concinna 7KRUHOO *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Centromerus sylvaticus %ODFNZDOO *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Ceratinella brevipes :HVWULQJ *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Ceratinopsis stativa 6LPRQ *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Dicymbium /LQQDHXV (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Dicymbium nigrum %ODFNZDOO (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Erigone arctica :KLWH 6WHQRWRSLF 3UHGDWRU\ /LQ\SKLLGDH Erigone atra %ODFNZDOO (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Erigone dentipalpis :LGHU (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Gongylidiellum vivum 23&DPEULGJH *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Meioneta beata 23&DPEULGJH *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Meioneta rurestris &/.RFK (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Mermessus trilobatus (PHUWRQ 6WHQRWRSLF 3UHGDWRU\ /LQ\SKLLGDH Micrargus subaequalis :HVWULQJ *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Microlinyphia pusilla 6XQGHYDOO *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Microneta viaria %ODFNZDOO *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Oedothorax apicatus %ODFNZDOO (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Oedothorax fuscus %ODFNZDOO (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Oedothorax gibbosus %ODFNZDOO *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Oedothorax retusus :HVWULQJ (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Palliduphantes ericaeus %ODFNZDOO *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Panamomops sulcifrons :LGHU *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Pelecopsis parallela :LGHU *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Pocadicnemis juncea /RFNHW 0LOOLGJH (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Porrhomma pygmaeum %ODFNZDOO *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Prinerigone vagans $XGRXLQ (XU\WRSLF 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO /LQ\SKLLGDH Tapinocyba praecox 23&DPEULGJH (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Tenuiphantes tenuis %ODFNZDOO (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Tiso vagans %ODFNZDOO (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Walckenaeria acuminata %ODFNZDOO (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Walckenaeria antica :LGHU (XU\WRSLF 3UHGDWRU\ /LQ\SKLLGDH Walckenaeria atrotibialis 23&DPEULGJH *HQHUDOLVW 3UHGDWRU\ /LQ\SKLLGDH Walckenaeria nudipalpis :HVWULQJ (XU\WRSLF 3UHGDWRU\ /LRFUDQLGDH Agraecina striata .XOF]\QVNL *HQHUDOLVW 3UHGDWRU\ /LRFUDQLGDH Agroeca proxima 23&DPEULGJH *HQHUDOLVW 3UHGDWRU\ /LRFUDQLGDH Phrurolithus festivus &/.RFK (XU\WRSLF 3UHGDWRU\ /\FRVLGDH Alopecosa pulverulenta &OHUFN (XU\WRSLF 3UHGDWRU\ /\FRVLGDH Arctosa leopardus 6XQGHYDOO 6WHQRWRSLF 3UHGDWRU\ /\FRVLGDH Pardosa agrestis :HVWULQJ 6WHQRWRSLF 3UHGDWRU\ /\FRVLGDH Pardosa nigriceps 7KRUHOO (XU\WRSLF 3UHGDWRU\ /\FRVLGDH Pardosa palustris /LQQDHXV *HQHUDOLVW 3UHGDWRU\ /\FRVLGDH Pardosa prativaga /.RFK (XU\WRSLF 3UHGDWRU\ /\FRVLGDH Pardosa pullata &OHUFN *HQHUDOLVW 3UHGDWRU\ /\FRVLGDH Pirata piraticus &OHUFN *HQHUDOLVW 3UHGDWRU\ /\FRVLGDH Pirata uliginosus 7KRUHOO *HQHUDOLVW 3UHGDWRU\ /\FRVLGDH Trochosa ruricola 'H*HHU (XU\WRSLF 3UHGDWRU\ /\FRVLGDH Trochosa terricola 7KRUHOO (XU\WRSLF 3UHGDWRU\ 0LPHWLGDH Ero cambridgei .XOF]\QVNL :RRGODQG 3UHGDWRU\ 3KLORGURPLGDH Thanatus striatus &/.RFK (XU\WRSLF 3UHGDWRU\ 3KLORGURPLGDH Tibellus oblongus :DOFNHQDHU *HQHUDOLVW 3UHGDWRU\ 3LVDXULGDH Pisaura mirabilis &OHUFN (XU\WRSLF 3UHGDWRU\ 6DOWLFLGDH Euophrys frontalis :DOFNHQDHU (XU\WRSLF 3UHGDWRU\ 6DOWLFLGDH Heliophanus cupreus :DOFNHQDHU (XU\WRSLF 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 0DQDJHPHQWJXLOG 7URSKLFJURXS & &>'@ &>6@ & ' 6 7RWDO 6DOWLFLGDH Heliophanus flavipes +DKQ (XU\WRSLF 3UHGDWRU\ 6DOWLFLGDH Sibianor aurocinctus 2KOHUW 6WHQRWRSLF 3UHGDWRU\ 7HWUDJQDWKLGDH Pachygnatha degeeri 6XQGHYDOO (XU\WRSLF 3UHGDWRU\ 7KHULGLLGDH Enoplognatha thoracica +DKQ (XU\WRSLF 3UHGDWRU\ 7KHULGLLGDH Episinus angulatus %ODFNZDOO *HQHUDOLVW 3UHGDWRU\ 7KHULGLLGDH Robertus arundineti 23&DPEULGJH *HQHUDOLVW 3UHGDWRU\ 7KHULGLLGDH Robertus lividus %ODFNZDOO *HQHUDOLVW 3UHGDWRU\ 7KRPLVLGDH Ozyptila brevipes +DKQ *HQHUDOLVW 3UHGDWRU\ 7KRPLVLGDH Ozyptila sanctuaria 23&DPEULGJH *HQHUDOLVW 3UHGDWRU\ 7KRPLVLGDH Ozyptila simplex 23&DPEULGJH 6WHQRWRSLF 3UHGDWRU\ 7KRPLVLGDH Pachygnatha degeeri 7XUQHU (XU\WRSLF 3UHGDWRU\ 7KRPLVLGDH Xysticus cristatus &OHUFN (XU\WRSLF 3UHGDWRU\ 7KRPLVLGDH Xysticus kochi 7KRUHOO 6WHQRWRSLF 3UHGDWRU\ =RGDULLGDH Zodarion italicum &DQHVWULQL 6WHQRWRSLF 3UHGDWRU\ =RULGDH Zora spinimana 6XQGHYDOO *HQHUDOLVW 3UHGDWRU\ $SSHQGL[ 7DEOH$ &DSWXUHGDWDIRUEHHWOHVIURPDJJUHJDWHPRXQGVH[SHULPHQWDW+DGOHLJK3DUN3LWVHD/DQGILOODQG5DLQKDP0DUVKHVLQDQG 7UHDWPHQWVZHUHWRQQHPRXQGVRIUHF\FOHGEXONILOO Q RU7KDQHWVDQG Q DQGXQPDQDJHGJUDVVODQG Q 7KHUHZHUHWZRSLWIDOOWUDSV SHUPRQLWRULQJSORWLQHDFK\HDURSHQIRUGD\VLQ0D\-XO\DQG6HSWHPEHU3OXVV\PERO Ά E\VSHFLHVQDPHGHQRWHVVWDWXVDVDVWHQRWRSLF LQGLFDWRURIHDUO\VXFFHVVLRQDOKDELWDWLQWKLVVWXG\6SHFLHVZHUHJURXSHGE\WURSKLFJXLOGIURPDGXOWIHHGLQJKDELWVDVGHWULWLYRURXVSK\WRSKDJRXV SUHGDWRU\]RRSKDJRXVRUXQNQRZQ )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO $OH[LLGDH Sphaerosoma pilosum 3DQ]HU 'HWULWLYRURXV $QWKLFLGDH Anthicus antherinus /LQQDHXV 'HWULWLYRURXV $SLRQLGDH Aspidapion radiolus 0DUVKDP 3K\WRSKDJRXV $SLRQLGDH Ceratapion carduorum .LUE\ 3K\WRSKDJRXV $SLRQLGDH Ceratapion gibbirostre *\OOHQKDO 3K\WRSKDJRXV $SLRQLGDH Ceratapion onopordi .LUE\ 3K\WRSKDJRXV $SLRQLGDH Malvapion malvae )DEULFLXV 3K\WRSKDJRXV $SLRQLGDH Omphalapion hookerorum .LUE\ 3K\WRSKDJRXV $SLRQLGDH Protapion assimile .LUE\ 3K\WRSKDJRXV $SLRQLGDH Protapion trifolii /LQQDHXV 3K\WRSKDJRXV $SLRQLGDH Stenopterapion tenue .LUE\ 3K\WRSKDJRXV $SLRQLGDH Taeniapion urticarium +HUEVW 3K\WRSKDJRXV %\UUKLGDH Byrrhus pilula /LQQDHXV 3K\WRSKDJRXV %\UUKLGDH Simplocaria semistriata )DEULFLXV 3K\WRSKDJRXV &DQWKDULGDH Cantharis decipiens %DXGL 3UHGDWRU\ &DQWKDULGDH Cantharis lateralis /LQQDHXV 3UHGDWRU\ &DQWKDULGDH Cantharis pallida *RH]H 3UHGDWRU\ &DQWKDULGDH Cantharis rufa /LQQDHXV 3UHGDWRU\ &DQWKDULGDH Cantharis rustica )DOOpQ 3UHGDWRU\ &DQWKDULGDH Rhagonycha fulva 6FRSROL 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO &DQWKDULGDH Rhagonycha limbata 7KRPVRQ 3UHGDWRU\ &DUDELGDH Acupalpus dubius 6FKLOVN\ 3UHGDWRU\ &DUDELGDH Acupalpus meridianus /LQQDHXV 3UHGDWRU\ &DUDELGDH Agonum fuliginosum 3DQ]HU 3UHGDWRU\ &DUDELGDH Amara aenea 'H*HHU 3K\WRSKDJRXV &DUDELGDH Amara communis 3DQ]HU 3K\WRSKDJRXV &DUDELGDH Amara convexior 6WHSKHQV 3K\WRSKDJRXV &DUDELGDH Amara eurynota 3DQ]HU 3K\WRSKDJRXV &DUDELGDH Amara familiaris 'XIWVFKPLG 3K\WRSKDJRXV &DUDELGDH Amara lunicollis 6FKL|GWH 3K\WRSKDJRXV &DUDELGDH Amara ovata )DEULFLXV 3K\WRSKDJRXV &DUDELGDH Amara plebeja *\OOHQKDO 3K\WRSKDJRXV &DUDELGDH Amara similata *\OOHQKDO 3K\WRSKDJRXV &DUDELGDH Amara tibialis 3D\NXOO 3K\WRSKDJRXV &DUDELGDH Anchomenus dorsalis 3RQWRSSLGDQ =RRSKDJRXV &DUDELGDH Anisodactylus binotatus )DEULFLXV =RRSKDJRXV &DUDELGDH Badister bullatus 6FKUDQN =RRSKDJRXV &DUDELGDH Bembidion biguttatum )DEULFLXV =RRSKDJRXV &DUDELGDH Bembidion femoratum 6WXUP =RRSKDJRXV &DUDELGDH Bembidion lampros +HUEVW =RRSKDJRXV &DUDELGDH Bembidion lunulatum *HRIIUR\LQ)RXUFUR\ =RRSKDJRXV &DUDELGDH Bembidion minimum )DEULFLXV =RRSKDJRXV &DUDELGDH Bembidion obtusum $XGLQHW6HUYLOOH =RRSKDJRXV &DUDELGDH Bembidion properans 6WHSKHQV =RRSKDJRXV &DUDELGDH Bembidion quadrimaculatum /LQQDHXV =RRSKDJRXV &DUDELGDH Brachinus crepitans /LQQDHXV =RRSKDJRXV &DUDELGDH Bradycellus verbasci 'XIWVFKPLG 3K\WRSKDJRXV )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO &DUDELGDH Calathus ambiguus 3D\NXOO =RRSKDJRXV &DUDELGDH Calathus fuscipes *RH]H =RRSKDJRXV &DUDELGDH Calathus melanocephalus /LQQDHXV =RRSKDJRXV &DUDELGDH Carabus nemoralis 0OOHU2) =RRSKDJRXV &DUDELGDH Carabus violaceus /LQQDHXV =RRSKDJRXV &DUDELGDH Clivina fossor /LQQDHXV =RRSKDJRXV &DUDELGDH Curtonotus aulicus 3DQ]HU 3K\WRSKDJRXV &DUDELGDH Curtonotus convexiusculus 0DUVKDP 3K\WRSKDJRXV &DUDELGDH Demetrias atricapillus /LQQDHXV =RRSKDJRXV &DUDELGDH Elaphrus riparius /LQQDHXV =RRSKDJRXV &DUDELGDH Harpalus affinis 6FKUDQN 3K\WRSKDJRXV &DUDELGDH Harpalus anxius 'XIWVFKPLG 3K\WRSKDJRXV &DUDELGDH Harpalus attenuatus 6WHSKHQV 3K\WRSKDJRXV &DUDELGDH Harpalus rubripes 'XIWVFKPLG 3K\WRSKDJRXV &DUDELGDH Harpalus rufipes 'H*HHU 3K\WRSKDJRXV &DUDELGDH Harpalus tardus 3DQ]HU 3K\WRSKDJRXV &DUDELGDH Leistus ferrugineus /LQQDHXV =RRSKDJRXV &DUDELGDH Loricera pilicornis )DEULFLXV =RRSKDJRXV &DUDELGDH Microlestes maurus 6WXUP =RRSKDJRXV &DUDELGDH Microlestes minutulus *RH]H =RRSKDJRXV &DUDELGDH Nebria brevicollis )DEULFLXV =RRSKDJRXV &DUDELGDH Notiophilus biguttatus )DEULFLXV =RRSKDJRXV &DUDELGDH Notiophilus palustris 'XIWVFKPLG =RRSKDJRXV &DUDELGDH Notiophilus substriatus :DWHUKRXVH*5 =RRSKDJRXV &DUDELGDH Ophonus ardosiacus /XWVFKQLN 3K\WRSKDJRXV &DUDELGDH Ophonus azureus )DEULFLXV 3K\WRSKDJRXV &DUDELGDH Ophonus puncticeps 6WHSKHQV 3K\WRSKDJRXV )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO &DUDELGDH Ophonus rufibarbis )DEULFLXV 3K\WRSKDJRXV &DUDELGDH Oxypselaphus obscurus +HUEVW =RRSKDJRXV &DUDELGDH Panagaeus bipustulatus )DEULFLXV =RRSKDJRXV &DUDELGDH Paradromius linearis 2OLYLHU =RRSKDJRXV &DUDELGDH Philorhizus melanocephalus 'HMHDQ =RRSKDJRXV &DUDELGDH Platyderus depressus $XGLQHW6HUYLOOH =RRSKDJRXV &DUDELGDH Poecilus cupreus /LQQDHXV =RRSKDJRXV &DUDELGDH Pterostichus macer 0DUVKDP =RRSKDJRXV &DUDELGDH Pterostichus madidus )DEULFLXV =RRSKDJRXV &DUDELGDH Pterostichus nigrita 3D\NXOO =RRSKDJRXV &DUDELGDH Pterostichus strenuus 3DQ]HU =RRSKDJRXV &DUDELGDH Scybalicus oblongiusculus 'HMHDQ 3K\WRSKDJRXV &DUDELGDH Stenolophus mixtus +HUEVW 3K\WRSKDJRXV &DUDELGDH Stomis pumicatus 3DQ]HU =RRSKDJRXV &DUDELGDH Syntomus obscuroguttatus 'XIWVFKPLG =RRSKDJRXV &DUDELGDH Trechus quadristriatus 6FKUDQN =RRSKDJRXV &HUDPE\FLGDH Gracilia minuta )DEULFLXV 3K\WRSKDJRXV &HUDPE\FLGDH Phytoecia cylindrica /LQQDHXV 3K\WRSKDJRXV &HUDPE\FLGDH Pseudovadonia livida )DEULFLXV 3K\WRSKDJRXV &KU\VRPHOLGDH Altica carinthiaca :HLVH 3K\WRSKDJRXV &KU\VRPHOLGDH Altica palustris :HLVH 3K\WRSKDJRXV &KU\VRPHOLGDH Aphthona euphorbiae 6FKUDQN 3K\WRSKDJRXV &KU\VRPHOLGDH Aphthona nigriceps 5HGWHQEDFKHU 3K\WRSKDJRXV &KU\VRPHOLGDH Batophila aerata 0DUVKDP 3K\WRSKDJRXV &KU\VRPHOLGDH Batophila rubi 3D\NXOO 3K\WRSKDJRXV &KU\VRPHOLGDH Bruchidius imbricornis 3DQ]HU 3K\WRSKDJRXV &KU\VRPHOLGDH Bruchus loti 3D\NXOO 3K\WRSKDJRXV )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO &KU\VRPHOLGDH Cassida rubiginosa 0OOHU2) 3K\WRSKDJRXV &KU\VRPHOLGDH Cassida vibex /LQQDHXV 3K\WRSKDJRXV &KU\VRPHOLGDH Cassida viridis /LQQDHXV 3K\WRSKDJRXV &KU\VRPHOLGDH Chaetocnema concinna 0DUVKDP 3K\WRSKDJRXV &KU\VRPHOLGDH Chaetocnema hortensis )RXUFUR\ 3K\WRSKDJRXV &KU\VRPHOLGDH Chaetocnema picipes 6WHSKHQV 3K\WRSKDJRXV &KU\VRPHOLGDH Chrysolina banksii )DEULFLXV 3K\WRSKDJRXV &KU\VRPHOLGDH Chrysolina oricalcia 0OOHU 3K\WRSKDJRXV &KU\VRPHOLGDH Lema cyanella /LQQDHXV 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus dorsalis )DEULFLXV 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus gracilis .XWVFKHUD 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus jacobaeae :DWHUKRXVH*5 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus luridus 6FRSROL 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus melanocephalus 'H*HHU 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus parvulus 3D\NXOO 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus pellucidus )RXGUDV 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus pratensis 3DQ]HU 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus rubiginosus )RXGUDV 3K\WRSKDJRXV &KU\VRPHOLGDH Longitarsus suturellus 'XIWVFKPLG 3K\WRSKDJRXV &KU\VRPHOLGDH Neocrepidodera ferruginea 6FRSROL 3K\WRSKDJRXV &KU\VRPHOLGDH Neocrepidodera transversa 0DUVKDP 3K\WRSKDJRXV &KU\VRPHOLGDH Phaedon tumidulus *HUPDU 3K\WRSKDJRXV &KU\VRPHOLGDH Phyllotreta atra )DEULFLXV 3K\WRSKDJRXV &KU\VRPHOLGDH Phyllotreta consobrina &XUWLV 3K\WRSKDJRXV &KU\VRPHOLGDH Phyllotreta cruciferae *RH]H 3K\WRSKDJRXV &KU\VRPHOLGDH Phyllotreta diademata )RXGUDV 3K\WRSKDJRXV &KU\VRPHOLGDH Phyllotreta nemorum /LQQDHXV 3K\WRSKDJRXV )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO &KU\VRPHOLGDH Phyllotreta nigripes )DEULFLXV 3K\WRSKDJRXV &KU\VRPHOLGDH Phyllotreta undulata .XWVFKHUD 3K\WRSKDJRXV &KU\VRPHOLGDH Phyllotreta vittula 5HGWHQEDFKHU 3K\WRSKDJRXV &KU\VRPHOLGDH Podagrica fuscipes )DEULFLXV 3K\WRSKDJRXV &KU\VRPHOLGDH Psylliodes affinis 3D\NXOO 3K\WRSKDJRXV &KU\VRPHOLGDH Psylliodes chrysocephala /LQQDHXV 3K\WRSKDJRXV &KU\VRPHOLGDH Sphaeroderma testaceum )DEULFLXV 3K\WRSKDJRXV &RFFLQHOOLGDH Coccinella septempunctata /LQQDHXV 3UHGDWRU\ &RFFLQHOOLGDH Hippodamia variegata *RH]H 3UHGDWRU\ &RFFLQHOOLGDH Nephus redtenbacheri 0XOVDQW 3UHGDWRU\ &RFFLQHOOLGDH Psyllobora vigintiduopunctata /LQQDHXV 3UHGDWRU\ &RFFLQHOOLGDH Rhyzobius litura )DEULFLXV 3UHGDWRU\ &RFFLQHOOLGDH Scymnus haemorrhoidalis +HUEVW 3UHGDWRU\ &RFFLQHOOLGDH Subcoccinella vigintiquattuorpunctata /LQQDHXV 3UHGDWRU\ &RFFLQHOOLGDH Tytthaspis sedecimpunctata /LQQDHXV 3UHGDWRU\ &RU\ORSKLGDH Corylophus sublaevipennis -DFTXHOLQGX9DO 'HWULWLYRURXV &RU\ORSKLGDH Orthoperus brunnipes *\OOHQKDO 'HWULWLYRURXV &RU\ORSKLGDH Sericoderus brevicornis 0DWWKHZV$ 'HWULWLYRURXV &RU\ORSKLGDH Sericoderus lateralis *\OOHQKDO 'HWULWLYRURXV &U\SWRSKDJLGDH Atomaria rubella +HHU 'HWULWLYRURXV &U\SWRSKDJLGDH Atomaria testacea 6WHSKHQV 'HWULWLYRURXV &U\SWRSKDJLGDH Cryptophagus distinguendus 6WXUP 'HWULWLYRURXV &U\SWRSKDJLGDH Cryptophagus puncticollis /XFDV 'HWULWLYRURXV &U\SWRSKDJLGDH Ootypus globosus :DOWO 'HWULWLYRURXV &XUFXOLRQLGDH Anthonomus rubi +HUEVW 3K\WRSKDJRXV &XUFXOLRQLGDH Barypeithes pellucidus %RKHPDQ 3K\WRSKDJRXV &XUFXOLRQLGDH Ceutorhynchus contractus 0DUVKDP 3K\WRSKDJRXV )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO &XUFXOLRQLGDH Ceutorhynchus erysimi )DEULFLXV 3K\WRSKDJRXV &XUFXOLRQLGDH Ceutorhynchus obstrictus 0DUVKDP 3K\WRSKDJRXV &XUFXOLRQLGDH Ceutorhynchus pallidactylus 0DUVKDP 3K\WRSKDJRXV &XUFXOLRQLGDH Ceutorhynchus typhae +HUEVW 3K\WRSKDJRXV &XUFXOLRQLGDH Euophryum confine %URXQ 3K\WRSKDJRXV &XUFXOLRQLGDH Gymnetron pascuorum *HUPDU 3K\WRSKDJRXV &XUFXOLRQLGDH Hadroplontus litura )DEULFLXV 3K\WRSKDJRXV &XUFXOLRQLGDH Hypera fuscocinerea 0DUVKDP 3K\WRSKDJRXV &XUFXOLRQLGDH Hypera meles )DEULFLXV 3K\WRSKDJRXV &XUFXOLRQLGDH Hypera nigrirostris )DEULFLXV 3K\WRSKDJRXV &XUFXOLRQLGDH Hypera plantaginis 'H*HHU 3K\WRSKDJRXV &XUFXOLRQLGDH Hypera postica *\OOHQKDO 3K\WRSKDJRXV &XUFXOLRQLGDH Larinus planus )DEULFLXV 3K\WRSKDJRXV &XUFXOLRQLGDH Leiosoma deflexum 3DQ]HU 3K\WRSKDJRXV &XUFXOLRQLGDH Mecinus pyraster +HUEVW 3K\WRSKDJRXV &XUFXOLRQLGDH Nedyus quadrimaculatus /LQQDHXV 3K\WRSKDJRXV &XUFXOLRQLGDH Neliocarus faber +HUEVW 3K\WRSKDJRXV &XUFXOLRQLGDH Neophytobius quadrinodosus *\OOHQKDO 3K\WRSKDJRXV &XUFXOLRQLGDH Orthochaetes setiger %HFN 3K\WRSKDJRXV &XUFXOLRQLGDH Otiorhynchus ligneus 2OLYLHU 3K\WRSKDJRXV &XUFXOLRQLGDH Otiorhynchus ovatus /LQQDHXV 3K\WRSKDJRXV &XUFXOLRQLGDH Otiorhynchus raucus )DEULFLXV 3K\WRSKDJRXV &XUFXOLRQLGDH Otiorhynchus rugosostriatus *RH]H 3K\WRSKDJRXV &XUFXOLRQLGDH Otiorhynchus sulcatus )DEULFLXV 3K\WRSKDJRXV &XUFXOLRQLGDH Parethelcus pollinarius )RUVWHU 3K\WRSKDJRXV &XUFXOLRQLGDH Pelenomus quadrituberculatus )DEULFLXV 3K\WRSKDJRXV &XUFXOLRQLGDH Phyllobius pyri /LQQDHXV 3K\WRSKDJRXV )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO &XUFXOLRQLGDH Phyllobius viridicollis )DEULFLXV 3K\WRSKDJRXV &XUFXOLRQLGDH Rhinocyllus conicus )U|OLFK 3K\WRSKDJRXV &XUFXOLRQLGDH Rhinoncus pericarpius /LQQDHXV 3K\WRSKDJRXV &XUFXOLRQLGDH Romualdius angustisetulus +DQVHQ 3K\WRSKDJRXV &XUFXOLRQLGDH Sitona cylindricollis )nKUDHXV 3K\WRSKDJRXV &XUFXOLRQLGDH Sitona hispidulus )DEULFLXV 3K\WRSKDJRXV &XUFXOLRQLGDH Sitona humeralis 6WHSKHQV 3K\WRSKDJRXV &XUFXOLRQLGDH Sitona lepidus *\OOHQKDO 3K\WRSKDJRXV &XUFXOLRQLGDH Sitona lineatus /LQQDHXV 3K\WRSKDJRXV &XUFXOLRQLGDH Trichosirocalus troglodytes )DEULFLXV 3K\WRSKDJRXV &XUFXOLRQLGDH Tychius picirostris )DEULFLXV 3K\WRSKDJRXV &XUFXOLRQLGDH Tychius tibialis %RKHPDQ 3K\WRSKDJRXV &XUFXOLRQLGDH Zacladus exiguus 2OLYLHU 3K\WRSKDJRXV 'HUPHVWLGDH Megatoma undata /LQQDHXV 'HWULWLYRURXV '\WLVFLGDH Colymbetes fuscus /LQQDHXV 3UHGDWRU\ '\WLVFLGDH Hydaticus seminiger 'H*HHU 3UHGDWRU\ '\WLVFLGDH Rhantus frontalis 0DUVKDP 3UHGDWRU\ (ODWHULGDH Agriotes lineatus /LQQDHXV 3K\WRSKDJRXV (ODWHULGDH Agriotes sputator /LQQDHXV 3K\WRSKDJRXV (ODWHULGDH Agrypnus murinus /LQQDHXV 3K\WRSKDJRXV (ODWHULGDH Athous campyloides 1HZPDQ 3K\WRSKDJRXV +HORSKRULGDH Helophorus brevipalpis %HGHO 'HWULWLYRURXV +HORSKRULGDH Helophorus rufipes %RVFG $QWLF 'HWULWLYRURXV +HWHURFHULGDH Heterocerus fenestratus 7KXQEHUJ 3K\WRSKDJRXV +LVWHULGDH Hister unicolor /LQQDHXV 3UHGDWRU\ +LVWHULGDH Kissister minimus $XEp 3UHGDWRU\ +LVWHULGDH Margarinotus brunneus )DEULFLXV 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO +LVWHULGDH Margarinotus purpurascens +HUEVW 3UHGDWRU\ +\GURSKLOLGDH Cercyon melanocephalus /LQQDHXV 'HWULWLYRURXV +\GURSKLOLGDH Cercyon pygmaeus ,OOLJHU 'HWULWLYRURXV +\GURSKLOLGDH Cercyon tristis ,OOLJHU 'HWULWLYRURXV +\GURSKLOLGDH Megasternum concinnum 0DUVKDP 'HWULWLYRURXV +\GURSKLOLGDH Sphaeridium lunatum )DEULFLXV 'HWULWLYRURXV +\GURSKLOLGDH Sphaeridium marginatum )DEULFLXV 'HWULWLYRURXV .DWHUHWLGDH Brachypterus glaber 1HZPDQ 3K\WRSKDJRXV .DWHUHWLGDH Brachypterus urticae )DEULFLXV 3K\WRSKDJRXV /DWULGLLGDH Cartodere bifasciata 5HLWWHU 'HWULWLYRURXV /DWULGLLGDH Corticaria impressa 2OLYLHU 'HWULWLYRURXV /DWULGLLGDH Corticarina minuta )DEULFLXV 'HWULWLYRURXV /DWULGLLGDH Cortinicara gibbosa +HUEVW 'HWULWLYRURXV /DWULGLLGDH Enicmus transversus 2OLYLHU 'HWULWLYRURXV /DWULGLLGDH Melanophthalma suturalis 0DQQHUKHLP 'HWULWLYRURXV /HLRGLGDH Agathidium laevigatum (ULFKVRQ 'HWULWLYRURXV /HLRGLGDH Catops fuscus 3DQ]HU 'HWULWLYRURXV /HLRGLGDH Catops grandicollis (ULFKVRQ 'HWULWLYRURXV /HLRGLGDH Catops morio )DEULFLXV 'HWULWLYRURXV /HLRGLGDH Choleva agilis ,OOLJHU 'HWULWLYRURXV /HLRGLGDH Choleva oblonga /DWUHLOOH 'HWULWLYRURXV /HLRGLGDH Nargus anisotomoides 6SHQFH 'HWULWLYRURXV /HLRGLGDH Nargus velox 6SHQFH 'HWULWLYRURXV /HLRGLGDH Ptomaphagus medius 5H\ 'HWULWLYRURXV /HLRGLGDH Ptomaphagus subvillosus *RH]H 'HWULWLYRURXV /HLRGLGDH Ptomaphagus varicornis 5RVHQKDXHU 'HWULWLYRURXV /HLRGLGDH Sciodrepoides watsoni 6SHQFH 'HWULWLYRURXV )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO 0DODFKLLGDH Cordylepherus viridis )DEULFLXV 3UHGDWRU\ 0RUGHOOLGDH Mordellistena parvula *\OOHQKDO 3K\WRSKDJRXV 1LWLGXOLGDH Glischrochilus hortensis *HRIIUR\LQ)RXUFUR\ 3K\WRSKDJRXV 1LWLGXOLGDH Meligethes aeneus )DEULFLXV 3K\WRSKDJRXV 1LWLGXOLGDH Meligethes fulvipes %ULVRXWGH%DUQHYLOOH 3K\WRSKDJRXV 1LWLGXOLGDH Meligethes rotundicollis %ULVRXWGH%DUQHYLOOH 3K\WRSKDJRXV 1LWLGXOLGDH Soronia grisea /LQQDHXV 3K\WRSKDJRXV 1LWLGXOLGDH Thalycra fervida 2OLYLHU 3K\WRSKDJRXV 2HGHPHULGDH Oedemera nobilis 6FRSROL 3K\WRSKDJRXV 3KDODFULGDH Olibrus aeneus )DEULFLXV 3K\WRSKDJRXV 3KDODFULGDH Phalacrus championi *XLOOHEHDX 3K\WRSKDJRXV 3KDODFULGDH Phalacrus fimetarius )DEULFLXV 3K\WRSKDJRXV 3KDODFULGDH Stilbus oblongus (ULFKVRQ 3K\WRSKDJRXV 3WLOLLGDH Acrotrichis fascicularis +HUEVW 'HWULWLYRURXV 3WLOLLGDH Acrotrichis sitkaensis 0RWVFKXOVN\ 'HWULWLYRURXV 3WLOLLGDH Ptenidium fuscicorne (ULFKVRQ 'HWULWLYRURXV 3WLOLLGDH Acrotrichis fascicularis +HUEVW 'HWULWLYRURXV 3WLOLLGDH Acrotrichis sitkaensis 0RWVFKXOVN\ 'HWULWLYRURXV 3WLOLLGDH Ptenidium intermedium :DQNRZLF] 'HWULWLYRURXV 3WLOLLGDH Ptenidium pusillum *\OOHQKDO 'HWULWLYRURXV 6FDUDEDHLGDH Aphodius erraticus /LQQDHXV 'HWULWLYRURXV 6FDUDEDHLGDH Aphodius foetidus +HUEVW 'HWULWLYRURXV 6FDUDEDHLGDH Aphodius plagiatus /LQQDHXV 'HWULWLYRURXV 6FDUDEDHLGDH Aphodius prodromus %UDKP 'HWULWLYRURXV 6FLUWLGDH Cyphon coarctatus 3D\NXOO 3K\WRSKDJRXV 6FLUWLGDH Cyphon laevipennis 7RXUQLHU 3K\WRSKDJRXV 6LOSKLGDH Nicrophorus interruptus 6WHSKHQV 'HWULWLYRURXV )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO 6LOSKLGDH Nicrophorus vespillo /LQQDHXV 'HWULWLYRURXV 6LOSKLGDH Silpha atrata /LQQDHXV 3UHGDWRU\ 6LOSKLGDH Silpha laevigata )DEULFLXV 3UHGDWRU\ 6LOSKLGDH Silpha tristis ,OOLJHU 3UHGDWRU\ 6LOSKLGDH Thanatophilus sinuatus )DEULFLXV 'HWULWLYRURXV 6LOYDQLGDH Psammoecus bipunctatus )DEULFLXV 'HWULWLYRURXV 6WDSK\OLQLGDH Acrolocha sulcula 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Acrotona muscorum %ULVRXWGH%DUQHYLOOH 3UHGDWRU\ 6WDSK\OLQLGDH Acrotona parens 0XOVDQW 5H\ 3UHGDWRU\ 6WDSK\OLQLGDH Aleochara bipustulata /LQQDHXV 3UHGDWRU\ 6WDSK\OLQLGDH Aleochara brevipennis *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Aleochara curtula *RH]H 3UHGDWRU\ 6WDSK\OLQLGDH Aleochara lanuginosa *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Aleochara sparsa +HHU 3UHGDWRU\ 6WDSK\OLQLGDH Aloconota gregaria (ULFKVRQ 3UHGDWRU\ 6WDSK\OLQLGDH Amarochara forticornis /DFRUGDLUH 3UHGDWRU\ 6WDSK\OLQLGDH Amischa analis *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Amischa decipiens 6KDUS 3UHGDWRU\ 6WDSK\OLQLGDH Anotylus insecatus *UDYHQKRUVW 'HWULWLYRURXV 6WDSK\OLQLGDH Anotylus inustus *UDYHQKRUVW 'HWULWLYRURXV 6WDSK\OLQLGDH Anotylus rugosus )DEULFLXV 'HWULWLYRURXV 6WDSK\OLQLGDH Anotylus sculpturatus *UDYHQKRUVW 'HWULWLYRURXV 6WDSK\OLQLGDH Anotylus tetracarinatus %ORFN 'HWULWLYRURXV 6WDSK\OLQLGDH Astenus immaculatus 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Astenus lyonessius -R\ 3UHGDWRU\ 6WDSK\OLQLGDH Atheta aquatica 7KRPVRQ 3UHGDWRU\ 6WDSK\OLQLGDH Atheta crassicornis )DEULFLXV 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO 6WDSK\OLQLGDH Atheta triangulum .UDDW] 3UHGDWRU\ 6WDSK\OLQLGDH Atheta xanthopus 7KRPVRQ 3UHGDWRU\ 6WDSK\OLQLGDH Bolitobius castaneus 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Brachida exigua +HHU 3UHGDWRU\ 6WDSK\OLQLGDH Brachygluta fossulata 5HLFKHQEDFK 3UHGDWRU\ 6WDSK\OLQLGDH Brachygluta haematica 5HLFKHQEDFK 3UHGDWRU\ 6WDSK\OLQLGDH Brachygluta waterhousei 5\H 3UHGDWRU\ 6WDSK\OLQLGDH Callicerus obscurus *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Callicerus rigidicornis (ULFKVRQ 3UHGDWRU\ 6WDSK\OLQLGDH Calodera protensa 0DQQHUKHLP 3UHGDWRU\ 6WDSK\OLQLGDH Carpelimus corticinus *UDYHQKRUVW 'HWULWLYRURXV 6WDSK\OLQLGDH Carpelimus incongruus 6WHHO 'HWULWLYRURXV 6WDSK\OLQLGDH Carpelimus rivularis 0RWVFKXOVN\ 'HWULWLYRURXV 6WDSK\OLQLGDH Chaetida longicornis *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Cypha longicornis 3D\NXOO 3UHGDWRU\ 6WDSK\OLQLGDH Dinaraea angustula *\OOHQKDO 3UHGDWRU\ 6WDSK\OLQLGDH Drusilla canaliculata )DEULFLXV 3UHGDWRU\ 6WDSK\OLQLGDH Falagrioma thoracica 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Gabrius breviventer 6SHUN 3UHGDWRU\ 6WDSK\OLQLGDH Gyrohypnus fracticornis 0OOHU2) 3UHGDWRU\ 6WDSK\OLQLGDH Ilyobates propinquus $XEp 3UHGDWRU\ 6WDSK\OLQLGDH Ischnosoma splendidum *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Lathrobium fulvipenne *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Lathrobium geminum .UDDW] 3UHGDWRU\ 6WDSK\OLQLGDH Lesteva longoelytrata *RH]H 'HWULWLYRURXV 6WDSK\OLQLGDH Lesteva sicula (ULFKVRQ 'HWULWLYRURXV 6WDSK\OLQLGDH Liogluta microptera 7KRPVRQ&* 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO 6WDSK\OLQLGDH Liogluta pagana (ULFKVRQ 3UHGDWRU\ 6WDSK\OLQLGDH Lobrathium multipunctum *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Megalinus glabratus *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Metopsia clypeata 0OOHU3:- 3UHGDWRU\ 6WDSK\OLQLGDH Micropeplus staphylinoides 0DUVKDP 3UHGDWRU\ 6WDSK\OLQLGDH Mocyta fungi *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Mycetoporus nigricollis 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Neobisnius villosulus 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Ocypus brunnipes )DEULFLXV 3UHGDWRU\ 6WDSK\OLQLGDH Ocypus fuscatus *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Ocypus olens 0OOHU2) 3UHGDWRU\ 6WDSK\OLQLGDH Ocyusa picina $XEp 3UHGDWRU\ 6WDSK\OLQLGDH Oligota pumilio .LHVHQZHWWHU 3UHGDWRU\ 6WDSK\OLQLGDH Ontholestes murinus /LQQDHXV 3UHGDWRU\ 6WDSK\OLQLGDH Othius laeviusculus 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Oxypoda brachyptera 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Oxypoda procerula 0DQQHUKHLP 3UHGDWRU\ 6WDSK\OLQLGDH Paederus littoralis *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Pella limbata 3D\NXOO 3UHGDWRU\ 6WDSK\OLQLGDH Philhygra debilis (ULFKVRQ 3UHGDWRU\ 6WDSK\OLQLGDH Philhygra elongatula *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Philhygra palustris .LHVHQZHWWHU 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus carbonarius *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus cognatus 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus cruentatus *PHOLQLQ/LQQDHXV 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus laminatus &UHXW]HU 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus marginatus 0OOHU2) 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO 6WDSK\OLQLGDH Philonthus sanguinolentus *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus splendens )DEULFLXV 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus succicola 7KRPVRQ&* 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus tenuicornis 0XOVDQW 5H\ 3UHGDWRU\ 6WDSK\OLQLGDH Philonthus varians 3D\NXOO 3UHGDWRU\ 6WDSK\OLQLGDH Platydracus stercorarius 2OLYLHU 3UHGDWRU\ 6WDSK\OLQLGDH Platystethus alutaceus 7KRPVRQ&* 3UHGDWRU\ 6WDSK\OLQLGDH Platystethus nitens 6DKOEHUJ&5 3UHGDWRU\ 6WDSK\OLQLGDH Quedius curtipennis %HUQKDXHU 3UHGDWRU\ 6WDSK\OLQLGDH Quedius fuliginosus *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Quedius levicollis %UXOOp 3UHGDWRU\ 6WDSK\OLQLGDH Quedius molochinus *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Quedius nemoralis %DXGL 3UHGDWRU\ 6WDSK\OLQLGDH Quedius nigriceps .UDDW] 3UHGDWRU\ 6WDSK\OLQLGDH Quedius nitipennis 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Quedius picipes 0DQQHUKHLP 3UHGDWRU\ 6WDSK\OLQLGDH Quedius semiaeneus 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Quedius semiobscurus 0DUVKDP 3UHGDWRU\ 6WDSK\OLQLGDH Rugilus erichsonii )DXYHO 3UHGDWRU\ 6WDSK\OLQLGDH Rugilus orbiculatus 3D\NXOO 3UHGDWRU\ 6WDSK\OLQLGDH Rugilus rufipes *HUPDU 3UHGDWRU\ 6WDSK\OLQLGDH Sepedophilus immaculatus 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Sepedophilus marshami 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Sepedophilus nigripennis 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Staphylinus dimidiaticornis *HPPLQJHU 3UHGDWRU\ 6WDSK\OLQLGDH Stenichnus scutellaris 0OOHU3:- .XQ]H 3UHGDWRU\ 6WDSK\OLQLGDH Stenus brunnipes 6WHSKHQV 3UHGDWRU\ )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO 6WDSK\OLQLGDH Stenus clavicornis 6FRSROL 3UHGDWRU\ 6WDSK\OLQLGDH Stenus juno 3D\NXOO 3UHGDWRU\ 6WDSK\OLQLGDH Stenus nanus 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Stenus nitens 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Stenus ossium 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Stenus providus (ULFKVRQ 3UHGDWRU\ 6WDSK\OLQLGDH Sunius propinquus %ULVRXWGH%DUQHYLOOH 3UHGDWRU\ 6WDSK\OLQLGDH Tachinus flavolimbatus 3DQGHOOp 3UHGDWRU\ 6WDSK\OLQLGDH Tachinus marginellus )DEULFLXV 3UHGDWRU\ 6WDSK\OLQLGDH Tachinus rufipes /LQQDHXV 3UHGDWRU\ 6WDSK\OLQLGDH Tachyporus atriceps 6WHSKHQV 3UHGDWRU\ 6WDSK\OLQLGDH Tachyporus chrysomelinus /LQQDHXV 3UHGDWRU\ 6WDSK\OLQLGDH Tachyporus dispar 3D\NXOO 3UHGDWRU\ 6WDSK\OLQLGDH Tachyporus hypnorum )DEULFLXV 3UHGDWRU\ 6WDSK\OLQLGDH Tachyporus nitidulus )DEULFLXV 3UHGDWRU\ 6WDSK\OLQLGDH Tasgius ater *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Tasgius globulifer *HRIIUR\ 3UHGDWRU\ 6WDSK\OLQLGDH Tasgius melanarius +HHU 3UHGDWRU\ 6WDSK\OLQLGDH Tasgius morsitans 5RVVL 3UHGDWRU\ 6WDSK\OLQLGDH Tasgius winkleri %HUQKDXHU 3UHGDWRU\ 6WDSK\OLQLGDH Thinonoma atra *UDYHQKRUVW 3UHGDWRU\ 6WDSK\OLQLGDH Xantholinus elegans 2OLYLHU 3UHGDWRU\ 6WDSK\OLQLGDH Xantholinus linearis 2OLYLHU 3UHGDWRU\ 6WDSK\OLQLGDH Xantholinus longiventris +HHU 3UHGDWRU\ 7HQHEULRQLGDH Helops caeruleus /LQQDHXV 'HWULWLYRURXV 7HQHEULRQLGDH Scaphidema metallicum )DEULFLXV 'HWULWLYRURXV 7KURVFLGDH Trixagus carinifrons GH%RQYRXORLU XQNQRZQ )DPLO\ 7D[RQ $XWKRULW\ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO 7KURVFLGDH Trixagus obtusus &XUWLV XQNQRZQ 7DEOH$ &DSWXUHGDWDIRUVSLGHUVIURPDJJUHJDWHPRXQGVH[SHULPHQWDW+DGOHLJK3DUN3LWVHD/DQGILOODQG5DLQKDP0DUVKHVLQDQG 7UHDWPHQWVZHUHWRQQHPRXQGVRIUHF\FOHGEXONILOO Q RU7KDQHWVDQG Q DQGXQPDQDJHGJUDVVODQG Q 7KHUHZHUHWZRSLWIDOOWUDSV SHUPRQLWRULQJSORWLQHDFK\HDURSHQIRUGD\VLQ0D\-XO\DQG6HSWHPEHU3OXVV\PERO Ά E\VSHFLHVQDPHGHQRWHVVWDWXVDVDQVWHQRWRSLF LQGLFDWRURIHDUO\VXFFHVVLRQDOKDELWDWLQWKLVVWXG\6SHFLHVZHUHJURXSHGE\WURSKLFJXLOG DOOVSLGHUVSUHGDWRU\ )DPLO\ 7D[RQ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO $JHOHQLGDH Tegenaria agrestis :DOFNHQDHU 3UHGDWRU\ $JHOHQLGDH Tegenaria silvestris /.RFK 3UHGDWRU\ &OXELRQLGDH Cheiracanthium erraticum :DOFNHQDHU 3UHGDWRU\ &OXELRQLGDH Clubiona phragmitis &/.RFK 3UHGDWRU\ &OXELRQLGDH Clubiona reclusa 23&DPEULGJH 3UHGDWRU\ &OXELRQLGDH Clubiona subtilis /.RFK 3UHGDWRU\ 'LFW\QLGDH Argenna subnigra 23&DPEULGJH 3UHGDWRU\ '\VGHULGDH Dysdera crocata &/.RFK 3UHGDWRU\ *QDSKRVLGDH Drassodes cupreus %ODFNZDOO 3UHGDWRU\ *QDSKRVLGDH Drassyllus pusillus &/.RFK 3UHGDWRU\ *QDSKRVLGDH Haplodrassus signifer &/.RFK 3UHGDWRU\ *QDSKRVLGDH Micaria pulicaria 6XQGHYDOO 3UHGDWRU\ *QDSKRVLGDH Trachyzelotes pedestris &/.RFK 3UHGDWRU\ *QDSKRVLGDH Zelotes apricorum /.RFK 3UHGDWRU\ *QDSKRVLGDH Zelotes latreillei 6LPRQ 3UHGDWRU\ +DKQLLGDH Hahnia nava %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Bathyphantes gracilis %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Bathyphantes parvulus :HVWULQJ 3UHGDWRU\ /LQ\SKLLGDH Centromerus capucinus 6LPRQ 3UHGDWRU\ /LQ\SKLLGDH Ceratinella brevipes :HVWULQJ 3UHGDWRU\ /LQ\SKLLGDH Ceratinopsis stativa 6LPRQ 3UHGDWRU\ /LQ\SKLLGDH Cnephalocotes obscurus %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Dicymbium nigrum %ODFNZDOO 3UHGDWRU\ )DPLO\ 7D[RQ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO /LQ\SKLLGDH Diplocephalus graecus 23&DPEULGJH 3UHGDWRU\ /LQ\SKLLGDH Diplostyla concolor :LGHU 3UHGDWRU\ /LQ\SKLLGDH Erigone atra %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Erigone dentipalpis :LGHU 3UHGDWRU\ /LQ\SKLLGDH Gnathonarium dentatum :LGHU 3UHGDWRU\ /LQ\SKLLGDH Gongylidiellum vivum 23&DPEULGJH 3UHGDWRU\ /LQ\SKLLGDH Lophomma punctatum %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Maso sundevalli :HVWULQJ 3UHGDWRU\ /LQ\SKLLGDH Meioneta beata 23&DPEULGJH 3UHGDWRU\ /LQ\SKLLGDH Meioneta rurestris &/.RFK 3UHGDWRU\ /LQ\SKLLGDH Meioneta saxatilis %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Meioneta simplicitarsis 6LPRQ 3UHGDWRU\ /LQ\SKLLGDH Micrargus herbigradus %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Micrargus subaequalis :HVWULQJ 3UHGDWRU\ /LQ\SKLLGDH Microlinyphia pusilla 6XQGHYDOO 3UHGDWRU\ /LQ\SKLLGDH Monocephalus fuscipes %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Neriene clathrata 6XQGHYDOO 3UHGDWRU\ /LQ\SKLLGDH Oedothorax apicatus %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Oedothorax fuscus %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Oedothorax retusus :HVWULQJ 3UHGDWRU\ /LQ\SKLLGDH Ostearius melanopygius 23&DPEULGJH 3UHGDWRU\ /LQ\SKLLGDH Palliduphantes ericaeus %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Palliduphantes pallidus 23&DPEULGJH 3UHGDWRU\ /LQ\SKLLGDH Panamomops sulcifrons :LGHU 3UHGDWRU\ /LQ\SKLLGDH Pelecopsis parallela :LGHU 3UHGDWRU\ /LQ\SKLLGDH Pocadicnemis juncea /RFNHW 0LOOLGJH 3UHGDWRU\ /LQ\SKLLGDH Porrhomma pygmaeum %ODFNZDOO 3UHGDWRU\ )DPLO\ 7D[RQ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO /LQ\SKLLGDH Stemonyphantes lineatus /LQQDHXV 3UHGDWRU\ /LQ\SKLLGDH Syedra gracilis 0HQJH 3UHGDWRU\ /LQ\SKLLGDH Tenuiphantes tenebricola :LGHU 3UHGDWRU\ /LQ\SKLLGDH Tenuiphantes tenuis %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Tenuiphantes zimmermanni %HUWNDX 3UHGDWRU\ /LQ\SKLLGDH Tiso vagans %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Troxochrus scabriculus :HVWULQJ 3UHGDWRU\ /LQ\SKLLGDH Walckenaeria acuminata %ODFNZDOO 3UHGDWRU\ /LQ\SKLLGDH Walckenaeria antica :LGHU 3UHGDWRU\ /LQ\SKLLGDH Walckenaeria atrotibialis 23&DPEULGJH 3UHGDWRU\ /LQ\SKLLGDH Walckenaeria capito :HVWULQJ 3UHGDWRU\ /LQ\SKLLGDH Walckenaeria dysderoides :LGHU 3UHGDWRU\ /LQ\SKLLGDH Walckenaeria nudipalpis :HVWULQJ 3UHGDWRU\ /LQ\SKLLGDH Walckenaeria unicornis 23&DPEULGJH 3UHGDWRU\ /LRFUDQLGDH Agraecina striata .XOF]\QVNL 3UHGDWRU\ /LRFUDQLGDH Agroeca inopina 23&DPEULGJH 3UHGDWRU\ /LRFUDQLGDH Agroeca proxima 23&DPEULGJH 3UHGDWRU\ /LRFUDQLGDH Phrurolithus festivus &/.RFK 3UHGDWRU\ /LRFUDQLGDH Phrurolithus minimus &/.RFK 3UHGDWRU\ /\FRVLGDH Alopecosa pulverulenta &OHUFN 3UHGDWRU\ /\FRVLGDH Pardosa agrestis :HVWULQJ 3UHGDWRU\ /\FRVLGDH Pardosa nigriceps 7KRUHOO 3UHGDWRU\ /\FRVLGDH Pardosa palustris /LQQDHXV 3UHGDWRU\ /\FRVLGDH Pardosa prativaga /.RFK 3UHGDWRU\ /\FRVLGDH Pardosa pullata &OHUFN 3UHGDWRU\ /\FRVLGDH Pirata uliginosus 7KRUHOO 3UHGDWRU\ /\FRVLGDH Trochosa ruricola 'H*HHU 3UHGDWRU\ )DPLO\ 7D[RQ 7URSKLFJURXS &RQWURO 5%) 7KDQHW &RQWURO 5%) 7KDQHW 7RWDO /\FRVLGDH Trochosa terricola 7KRUHOO 3UHGDWRU\ 0LPHWLGDH Ero furcata 9LOOHUV 3UHGDWRU\ 3KLORGURPLGDH Philodromus cespitum :DOFNHQDHU 3UHGDWRU\ 3KLORGURPLGDH Thanatus striatus &/.RFK 3UHGDWRU\ 3KLORGURPLGDH Tibellus oblongus :DOFNHQDHU 3UHGDWRU\ 3LVDXULGDH Pisaura mirabilis &OHUFN 3UHGDWRU\ 6DOWLFLGDH Euophrys frontalis :DOFNHQDHU 3UHGDWRU\ 6DOWLFLGDH Sibianor aurocinctus 2KOHUW 3UHGDWRU\ 7HWUDJQDWKLGDH Pachygnatha clercki 6XQGHYDOO 3UHGDWRU\ 7HWUDJQDWKLGDH Pachygnatha degeeri 6XQGHYDOO 3UHGDWRU\ 7KHULGLLGDH Crustulina guttata :LGHU 3UHGDWRU\ 7KHULGLLGDH Enoplognatha latimana +LSSD 2NVDOD 3UHGDWRU\ 7KHULGLLGDH Enoplognatha thoracica +DKQ 3UHGDWRU\ 7KHULGLLGDH Robertus arundineti 23&DPEULGJH 3UHGDWRU\ 7KHULGLLGDH Robertus lividus %ODFNZDOO 3UHGDWRU\ 7KHULGLLGDH Theridion varians +DKQ 3UHGDWRU\ 7KRPLVLGDH Ozyptila brevipes +DKQ 3UHGDWRU\ 7KRPLVLGDH Ozyptila praticola &/.RFK 3UHGDWRU\ 7KRPLVLGDH Ozyptila sanctuaria 23&DPEULGJH 3UHGDWRU\ 7KRPLVLGDH Ozyptila simplex 23&DPEULGJH 3UHGDWRU\ 7KRPLVLGDH Ozyptila trux %ODFNZDOO 3UHGDWRU\ 7KRPLVLGDH Xysticus cristatus &OHUFN 3UHGDWRU\ 7KRPLVLGDH Xysticus kochi 7KRUHOO 3UHGDWRU\ =RGDULLGDH Zodarion italicum &DQHVWULQL 3UHGDWRU\ =RULGDH Zora spinimana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ndrena chrysosceles .LUE\ $QGUHQLGDH Andrena dorsata .LUE\ $QGUHQLGDH Andrena flavipes 3DQ]HU $QGUHQLGDH Andrena haemorrhoa )DEULFLXV $QGUHQLGDH Andrena labialis .LUE\ $QGUHQLGDH Andrena minutula .LUE\ $QGUHQLGDH Andrena nigroaenea .LUE\ $QGUHQLGDH Andrena nitida 0XOOHU $QGUHQLGDH Andrena praecox 6FRSROL $QGUHQLGDH Andrena scotica 3HUNLQV5&/ $SLGDH Anthophora bimaculata 3DQ]HU $SLGDH Nomada flava 3DQ]HU $SLGDH Nomada flavoguttata .LUE\ $SLGDH Nomada flavopicta .LUE\ $SLGDH Nomada goodeniana .LUE\ $SLGDH Nomada marshamella .LUE\ &ROOHWLGDH Hylaeus communis 1\ODQGHU &ROOHWLGDH Hylaeus hyalinatus 6PLWK &UDEURQLGDH Cerceris arenaria /LQQDHXV &UDEURQLGDH Cerceris quinquefasciata 5RVVLXV &UDEURQLGDH Cerceris rybyensis /LQQDHXV &UDEURQLGDH Crossocerus wesmaeli 9DQGHU/LQGHQ )DPLO\ 7D[RQ $XWKRULW\ 5%) 7KDQHW 5%) 7KDQHW 7RWDO &UDEURQLGDH Entomognathus brevis 9DQGHU/LQGHQ &UDEURQLGDH Gorytes quadrifasciatus )DEULFLXV &UDEURQLGDH Mimumesa unicolor 9DQGHU/LQGHQ &UDEURQLGDH Oxybelus uniglumis /LQQDHXV &UDEURQLGDH Philanthus triangulum )DEULFLXV &UDEURQLGDH Trypoxylon medium GH%HDXPRQW +DOLFWLGDH Halictus rubicundus &KULVW +DOLFWLGDH Lasioglossum albipes )DEULFLXV +DOLFWLGDH Lasioglossum calceatum 6FRSROL +DOLFWLGDH Lasioglossum leucozonium 6FKUDQN +DOLFWLGDH Lasioglossum malachurum .LUE\ +DOLFWLGDH Lasioglossum minutissimum .LUE\ +DOLFWLGDH Lasioglossum morio )DEULFLXV +DOLFWLGDH Lasioglossum parvulum 6FKHQFN +DOLFWLGDH Lasioglossum pauperatum %UXOOH +DOLFWLGDH Lasioglossum pauxillum 6FKHQFN +DOLFWLGDH Lasioglossum puncticolle 0RUDZLW] +DOLFWLGDH Lasioglossum villosulum .LUE\ +DOLFWLGDH Sphecodes ephippius /LQQDHXV +DOLFWLGDH Sphecodes geoffrellus .LUE\ +DOLFWLGDH Sphecodes gibbus /LQQDHXV +DOLFWLGDH Sphecodes longulus YRQ+DJHQV +DOLFWLGDH Sphecodes monilicornis .LUE\ +DOLFWLGDH Sphecodes puncticeps 7KRPVRQ 0HOLWWLGDH Melitta leporina 3DQ]HU 3RPSLOLGDH Anoplius concinnus 'DKOERP 3RPSLOLGDH Arachnospila anceps :HVPDHO )DPLO\ 7D[RQ $XWKRULW\ 5%) 7KDQHW 5%) 7KDQHW 7RWDO 3RPSLOLGDH Auplopus carbonarius 6FRSROL 3RPSLOLGDH Episyron rufipes /LQQDHXV 3RPSLOLGDH Priocnemis pusilla 6FKLRHGWH 6SKHFLGDH Ammophila sabulosa /LQQDHXV