INSECTA MUNDI, Vol. 12, Nos. 1 & 2, March-June, 1998 139 Redescriptions, new combinations, synonymies, and new records of South American Lepturini (Coleoptera: Cerambycidae: Leptur'inae)

Osvaldo R. Di Iorio

Entomologia, Facultad de Ciencias Exactas y Naturales, 1428 Ciudad Universitaria, Buenos Aires, Argentina. Internet: diiorio®biolo.bg.fcen.uba.ar

Abstract: Some Neotropical Lepturini are studied: Euryptera latipennis Audinet-Serville, 1828 (type species) redescription, Strangalia dimidiata (Redtenbacher, 1868) new combination (= Strangalia melanophthisis (Berg, 1889) new combination, new synonymy, and lectotype designation), Strangalia fulvicornis (Bates, 1872) new record from Argentina; Strangalia melaniira (Redtenbacher, 1868) new combination from Brazil; Strangalia rubricollis (Bates, 1870) new record from Peru. "Leptura' bonariensis Burmeister, 1865 is not a true Leptura, being exotic for Argentina and for the Neotropical fauna; the type-specimen was mislabelled as coming from Buenos Aires. Elytral patterns, posterior tarsi, antennae, 5th apparent urosternite, and elytral apices of E. latipennis, and S. dimidiata are illustrated. A key to Argentinian genera and species is provided. Key words: Coleoptera, Cerambycidae, Lepturini, systematics.

Introduction Facultad de Humanidades y Ciencias, Montevideo, Uruguay; GP: Glorialdo Pellerano (MACN); MACN: Linsley & Chemsak (1971) characterized the Museo Argentino de Ciencias Naturales "Bernar­ genera Euryptera Audinet-Serville, 1828 (type dino Rivadavia," Buenos Aires; MLP: Museo de La species E. latipennis Audinet-Serville, 1828) and Plata, Buenos Aires; ODI: Osvaldo Di Iorio, Buenos Strangalia Audinet-Serville, 1828 (type-species Aires; MV: Manuel Viana, Rosario de Lerma, Salta; Leptura luteicornis F.: Thomson, 1860 designation), MZ: Mateo Zelich, Liebig, Entre Rios. and transferred many Central American species from Euryptera to Strangalia. The revision of the Euryptera Audinet-Serville, 1828 North American fauna of , started by Linsley and Chemsak (1972), was completed in 1976 Type-species: E. latipennis Audinet-Serville, by the same authors, who included Strangalia and 1828 (by monotypy). Leptura among the genera with the sides of the Diagnosis (from Linsley and Chemsak, 1971): prothorax inerm. elytra more or less flattened, costate, expanded Within the South American fauna, not all the posteriorly; apices transversely sinuate; sutural and species originally described in Euryptera and still external angles dentiform (Fig. 6); presternal placed in this (Menne and Giesbert, 1995) process narrow, more or less laminiform, arcuately belong to it. Of the 3 species recorded for Argentina, declivous behind; coxae exerted, extending well Euryptera latipennis, Euryptera dimidiata Red­ above the intercoxal process; mesosternal process tenbacher (1868) and Euryptera melanophthisis prominent, elevated above the coxae, arcuately but Berg (1889), the last 2 belong in the genus subvertically declivous in front. Strangalia, the last being a synonym of the second. One new record of species of Strangalia for Euryptera latipennis Audinet-Serville, 1828 Argentina is added here, the first species of this (Figs. 1, 5-6, 8, 11-13,27) genus is recorded for Peru, and the diagnosis of the genus Euryptera is amplified. Euryptera latipennis: Bruch, 1912; Buck, 1959. Leptura bonariensis Burmeister (1865), de­ Euryptera latipennis val'. virgata Gounelle, 1911; scribed from Buenos Aires, is an exotic species for Bruch, 1915; Buck, 1959. the Argentinian fauna: it was mentioned with an Redescription: Body depressed, wider than its interrogation mark (Morine and Giesbert, 1995), height. Antennae 11-segmented (Fig. 1): scape and besides it is no true Leptura (Linsley and gradually broadening toward apex, somewhat Chemsak, 1976; Villiers, 1978). arched; antennomere II very short, wider than long; Collections examined: CB: Carlos Bruch antennomere III 3/4 length of scape; antennomere (MACN); HB: Hermann Burmeister (MACN); FHC: IV shorter than III; antennomere V as long as III, 140 Vol. 12, Nos. 1 & 2, March-June, 1998, INSECTA MUNDI with narrow base and broadening gradually toward Two lengthwise spots on pronotum and those on apex, with outer apical angle acute, but not distal 1/3 of elytra, black in colour, are found in all produced; antennomeres VI, VII, and VIII, specimens. Black spot on the apical 1/3 of elytra subequal, becoming steadily broader at base and may form band, broader toward suture and less angular at outer apex; antennomeres IX and X following along it up to scutellum (Fig. 11); it also somewhat shorter than the VIII, with inner and may be reduced (Fig. 12) or absent (Fig. 13). In this outer margins arcuate, as wide as VIII in middle of last case, apical black band on elytra is reduced to their length; antennomere XI longer than X, with half its size. distal 1/3 acuminate, acute at apex. Scape and Known geographical distribution: Brazil: antennomeres II, III, and IV coarsely, more or less Espiritu Santo: Fazenda Jerusalem (Zajciw, 1973); densely punctured; antennomeres V to XI with very Rio de Janeiro: Corcovado (Zajciw, 1958), Parque fine punctures and microsculpture. Nacional do Itatiaia (Zajciw, 1972); Rio Grande do Fore tibiae as long as femora; first tarsomere Sul: Porto Alegre, Sao Leopoldo, Cerro Largo, San little shorter than second and third taken together. Francisco da Paula (Buck, 1959); Paraguay: Middle tibiae slightly shorter than femora; first Villarica, Estancia Aguerito (Rio Aquidaban) tarsomere as long as second and third taken (Viana, 1972); Argentina: 'I'ucuman (Bruch, 1912; together. Hind tibiae longer than femora, narrow, Viana, 1972). slender, with apex not exceeding the posterior Material examined: Brazil: 16-X-1901, 1 ex. margin of elytra, bearing 2 apical spurs, inner one (CB); Bolivia: "Sur-Yungas", Suiqui, 1-1951, Suilar longer than outer; tarsi shorter than tibiae, first leg., 1 ex. (ODI); Argentina: Jujuy: 1 ex. (ODI); tarsomere half as long as second and third taken Salta: Tabacal 29-XII-1944, Bridarolli leg., 1 ex. together (Fig. 5). (ODI); 1-1930, Witte leg., 1 ex. (CB); Departamento Pronotum trapezoidal, gradually broadening La Villa, Talapampa, XII-1992, Martinez A. leg., 1 backwards; lateral margins slightly convex in ex. (ODI); Rosario de Lerma, Martinez A leg., 1­ anterior half; anterior margin somewhat concave; 1988, 1 ex. (ODI), II-1992, 1 ex. (ODI); Sumalao, posterior margin strongly convex on median line Martinez A. leg., XII-1982, 1 ex. (ODI), 1-1993, 1 ex. (not covering scutellum); posterior angles acute, (ODI); Tucuman: IV-1897, 1 ex. (CB), 26-1-1908, 2 projecting backward and going around humeri, exs. (CB). which are rounded. Elytra slightly narrowed behind Taxonomic discussion: Linsley and Chemsak humeri, behind this, lateral margins diverge (1971) give more characters for Strangalia than for gradually backward, broadening on apical 1/4 (Figs. Euryptera, probably because they did not have 11-13); outer apical angle produced into more or less specimens of E. latipennis. To the genus diagnosis triangular spine; posterior margin truncate, first (Linsley and Chemsak, 1971), the following outwardly concave and then inwardly convex; characters are added, different from Strangalia: III sutural angle straight, slightly projecting (Fig. 6). antennomere slightly shorter than scapus (Fig. 1); Elytra with 5 costas, not reaching anterior and hind tibiae not carinate, not exceeding the posterior posterior margins of each elytron; 3 very weak margin of elytra; first segment of hind tarsus twice veins, which coincide with first, second, and fourth as long as the second and third taken together (Fig. costae, counting from suture. 5); posterior angles of the prothorax going round the Prosternal process little less strongly raised humeri; posterior margin of the pronotum strongly than fore coxae (globose, prominent), strongly convex on the median line; elytra with 5 costae; narrowed between these and backward, not ex­ posterior angles .of the 5th apparent urosternite ceeding posterior margin of coxae; mesosternal strongly projecting (Fig. 8). process slightly raised above middle coxae, with Biogeography: distribution of E. latipennis anterior margin subvertical. covers from the north-east and central-east of Brazil Urosternites subequal in length, gradually (Zajciw, 1958, 1972, 1973) to Paraguay (Viana, narrowing backward, with setigerous punctures 1972), and from there along the Yungas in Bolivia, evenly spaced; posterior margin straight in first 3 and the northwest of Argentina (material ex­ urosternites and slightly concave on median line of amined), outside the Chaco sensu stricto (prado, fourth; posterior margins with microscopically small 1993). teeth; posterior angles of fifth projecting into acute, triangular spines, posterior margin between them "Leptura" bonariensis Burmeister, 1865 concave (Fig. 8), exceeding posterior margin of elytra in 1/3 of its length. Ophioetomis bonariensis (Burm.): Bruch, 1912. INSECTA MUNDI, Vol. 12, Nos. 1 & 2, March-June, 1998 141

Leptura bonaeriensis: Blackwelder, 1946. Euryptera dimidiata: Redtenbacher, 1868; Gou­ ? Leptura bonaerensis: Menne & Giesbert, 1995. nelle, 1911; Bruch, 1912; Blackwelder, 1946; Known geographical distribution: Argen­ Buck, 1959; Zajciw and Ruffinelli, 1962; Morine tina: Buenos Aires (Menne & Giesbert, 1995). and Zajciw, 1970; Morine & Giesbert, 1995. Material examined: "Buenos/Ayres" (printed, Euryptera dimidiata var. atra Gounelle, 1911; black on green paper), 1 ex. (HB), labelled "Leptura Buck,1959. bonariensis Burm., 1865 / Typus" (hand-written by Diagnosis: Form long and slender; males: Axel Bachmann, reddish colour). totally black, with golden hairlike setae at 2 Taxonomic discussion: The type specimen longitudinal bands in lateral margins of prothorax; shows that this is not a true Leptura (Linsley and females: prothorax always with 2 lateral yellow Chemsak, 1976; Villiers, 1978) or a species of Stran­ bands, humeri of same color and black scutellum; galia (= Ophistomis) (Bruch, 1912). This species is dark and light forms, varying in elytral pattern of exotic for the Argentinian fauna, and it is not black and yellow (Figs. 15-24); elytra finely and known whether the specimen was wrongly labelled densely punctate, with 3 veins, not elevated; lateral by Burmeister as coming from Buenos Aires or margins of elytra straight, not narrowed at middle; whether it is an introduced individual (although the both elytra narrowing progressively posteriorly. species was never found again in Argentina). The Redescription: Body not depressed, as high as generic and nomenclatorial status of this species wide at middle of its length, narrowing gradually remains uncertain, as it should be determined towards both ends. whether the specimen belongs to some Palearctic Female: Antennae 11-segmented (Fig. 2): and/or Nearctic genus and species (undescribed or antennomere III little longer than scape; anten­ not), which is outside the scope of the present paper. nom ere IV shorter than V; antennomere VI sub­ equal to IV; following antennomeres decrease Strang alia Audinet-Serville, 1828 gradually in length, except antennomere XI longer Ophistomis Thomson, 1857 (type-species O. flaoo­ than X, with apex acuminate. cinctus Thomson, 1864 by monotypy). Pronotum trapezoidal; anterior margin straight; Diagnosis (modified from Linsley and Chem­ lateral margins diverging backwards, convex at sak, 1971): Form slender, tapering posteriorly. Head middle of their length; posterior margin slightly oblique, abruptly and deeply constricted behind the convex on middle line; posterior angles acute, eyes; front rather short; palpi subequal, apical slightly projecting backward, shorter than elytral segment slender, cylindrical; eyes large, finely fac­ width between humeri. eted, notched. Antennae inserted on front along an­ Humeri rounded; lateral margins of elytra terior margin of eyes; scape slender, distal segments straight, narrowing gradually backwards, slightly usually with distinct poriferous areas, (females); III curved towards median line at apical 1/4; outer antennomere longer than scape; IV shorter than III; apical angle projecting into short broad spine; V longer than IV (Figs. 2-3). Pronotum trapezoidal, posterior margin concave; sutural angle projecting about as long as basal width; hind angles acute. into very short spine; suture dehiscent on apical 1/4 Prosternal process narrow, expanded at apex; front (Fig. 7). Elytra not costate, with 3 veins in dorsal coxal cavities open behind; mesosternal process view and 1, parallel to margin, in lateral view. broad, gradually arcuate anteriorly, narrowing pos­ Middle tibiae slightly shorter than femora; 1th teriorly. Elytra usually strongly cuneiform (Figs. 15­ tarsomere little longer than 2nd and 3rd taken 26), sometimes sides narrowing at middle (Fig. 14); together. Hind tibiae little longer than femora; 1st obliquely or transversely truncate at apex (Figs. 7, tarsomere twice as long as 2nd and 3rd taken 10, 14-26). Legs slender; posterior tibia often car­ together; 5th tarsomere little shorter than 2nd and inate along inside (males); posterior tarsi slender, 3rd taken together (Fig. 4). elongate (Fig. 4); apical segment narrow, cleft at First apparent urosternite longer than each of apex. Abdomen usually with last sternite deeply following ones; 2nd, 3rd, and 4th decreasing excavated and margins expanded (males) or elon­ gradually in length and width; posterior margin of gate, with posterior margin convex and posterior each sternite with small teeth; 5th urosternite angles not projected (Fig. 9). nearly twice as long as the 4th, trapezoidal, with posterior angles projecting into very small, but Strangalia dimidiata (Redtenbacher, 1868) evident spine, with slightly convex margin (Fig. 10). new combination Prosternal process very narrow and placed at (Figs. 2-4, 7, 9-10, 15-24, 27) base of fore coxae; mesosternal process broad, with 142 Vol. 12, Nos. 1 & 2, March-June, 1998, INSECTA MUNDI

lateral margins somewhat converging backward, at Grande do Sul: Porto Alegre, Pared Novo, Cerro anterior end very little raised above level of coxae, Largo, San Francisco da Paula (Buck, 1959); Uru­ with anterior margin oblique. guay: Maldonado: Cerro Lemos (Zajciw and Ruf­ Entirely black, except for lateral margins of finelli, 1962); Artigas: Rio Cuareim, Sepulturas, prothorax, which are yellow. According to color of Picada del Negro Muerto; Treinta y Tres: Arroyo elytra, light or dark forms may be distinguished, Avestruz Grande; Tacuaremb6: Puntas Arroyo yellow dominating in former, and black in latter. Laureles; (Zajciw and Morine, 1970); Argentina: Dark forms (Figs. 15-18): elytra black on most of Corrientes (Bruch, 1912). their surface, except for following parts: 1) narrowly Material examined: Argentina: Misiones: no yellow humeri (Fig. 15); 2) narrowly yellow humeri; more data, 1 a (CB); Bonpland, Jorgensen leg., 1 'i1 lengthwise spot behind scutellum, common to both (CB), "Euryptera dimidiata Rdtb." (hand-written by sutural edges (Fig. 16); 3) narrowly yellow humeri, Bruch), "Ophioetomis Gounelle det." (hand-written spot against lateral margin of each elytron behind by?); Parque Nacional Iguazu, 22-XI-1989, Cueto humerus; 4) narrowly yellow humeri; both spots leg., 1 a (ODI); Corrientes: Santo Tome, Pellerano present, sutural and posthumeral marginal; 5) leg., X-1925, 1 'i1a (GP), X-1928, 1 a (GP), XI-1928, posthumeral marginal spot merged with humeral 1 'i1 (GP); Entre Rios: Pronunciamiento, XI-1969, spot into band which does not reach middle of Zelich M. leg., 1 a (ODI); Liebig, without date, 1 'i1 elytral length; sutural spot extending backward up (ODI), "U. Martins det. 1991 Strangalia succincta to middle of elytral length. Light forms (Figs. 19­ Redt.," XII-1988, Zelich M. leg., 6 a a, 15 'i1'i1 (ODI); 24): scutellum always black; humeri broadly yellow; Buenos Aires: without locality, Bruch C. leg., 16­ apical half of elytra always black; basal half of XII-1922, 2 aa, 2 'i1'i1 (CB), without date, 1 a (CB); elytra yellow, spotted with black with following Otamendi, Estaci6n LN.T.A., 7-1-1979, Di Iorio leg., pattern variations: oval spot, common to both elytra 3 aa, 6 'i1'i1 (ODI); San Isidro, XI-1973, Martinez A. and disposed transversely surrounding scutellum, leg., 1 a (ODI); Hudson (F.C.8.), 6-XII- (1941 ?), 1 'i1 extending up to basal 1/3 of the elytra, not reaching (ODI); Uruguay: Artigas: Sepulturas, Cuareim lateral margins (Fig. 19); basal black spot reduced river, Picada del Negro Muerto, 10-XII-1966, gradually from behind forward (Figs. 20 to 23); Carbonell, Morine and San Martin leg., 1 female basal 1/3 of elytra entirely yellow (Fig. 24). (FHC). Male: narrower than female; legs little longer Taxonomic discussion: The original descri­ in proportion. Shape of prothorax, elytra and legs ption (Redtenbacher, 1868) is based, although this similar to those of females. Antennae II-segmented is not stated, on a female, as can be deduced by the (Fig. 3). Antennomere III little longer than scape; yellow and black color of the elytra. As char­ antennomeres IV and V little shorter than III; acterized by Linsley and Chemsak (1971), E. antennomere VI shorter than IV; antennomeres VII dimidiata belongs to Strangalia. to X subequal in length; antennomere XI longer Strangalia succincta has been found only at Rio than X, with apex elongate, acuminate. Antenno­ de Janeiro (Morine and Giesbert, 1995), and can be meres IV to VII broadening towards apex; distinguished from S. dimidiata by the reddish antennomeres III and VIII to XI elongate, more or prothorax, and the elytra entirely black except for a less linear (Fig. 3). transverse yellow band about middle of their length Last apparent sternite not modified, slightly (Redtenbacher, 1868). Some specimens of Stran­ exceeding the posterior margin of elytra, 1.5 times galia dimidiata (Fig. 17) ressemble S. succincta, but width of preceding segments, narrowing gradually in the first species the humeri are always yellow backwards; posterior margin somewhat convex on and the pronotum is black and yellow. median line, with posterior angles slightly pro­ jecting. Biogeography: E. latipennis and S. dimidiata Cuticle entirely black except sometimes small are sympatric in Brazil (Zajciw, 1972; Buck, 1959), yellow spot against each lateral margin at base of but the distribution of S. dimidiata extends pronotum; lateral margins of prothorax with golden southward, as far as Uruguay (Zajciw & Ruffinelli, hairlike setae, from base to anterior margin. 1962) and Argentina, only east of the Chaco sensu stricto, in the so-called mesopothamic region, from Known geographical distribution: Brazil: Misiones to Entre Rios (along the Gallery Forest of Rio de Janeiro (Type-locality: Redtenbacher, 1868): the Uruguay river), and Buenos Aires (coast of the Parque Nacional do Itatiaia (Zajciw, 1972); Rio de Rio de la Plata) (Fig. 27). Janeiro: Serra do Caraca (Gounelle, 1911); Rio INSECTA MUNDI, Vol. 12, Nos. 1 & 2, March-June, 1998 143

Strang alia melanophthisis (Berg, 1889), melanophthisis, "the sides of the prothorax alone new combination, and new synonymy bear golden yellow hairs". One of Bosq's classic nomenclatorial comments (in Bosq and Ruffinelli, Known geographical distribution: Brasil: 1951) says that "Melzer, when describing his E. Rio Grande do Sul: Porto Alegre (Buck, 1959); melanura Rdtb., 1868, abo nigripennis Melz., 1930, Argentina: Corrientes (cerca ciudad de Corrientes) expressed doubts about specimens from Rio Grande (Berg, 1889; Bruch, 1912); Uruguay: Tacuaremb6: do Sul [Brazil], seen by him, which also correspond Puntas Arroyo Laureles (Zajciw & Ruffinelli, 1962); to the description of E. melanophthisis Berg, from Rocha: Parque San Miguel (Zajciw and Menne, Argentina" [translated from Spanish]. By the 1968); Florida: Paso de Pache (Bosq and Ruffinelli, characters discussed further on, S. melanura is a 1951; Zajciw & Ruffinelli, 1962). different species from S. melanophthisis and, Material examined: Argentina: Misiones: therefore, from S. dimidiata. without locality, 20-1-1910, Bruch C. leg., 1

Known geographical distribution: Brazil Biology of Lepturini (Morine and Giesbert, 1995): Rio de Janeiro, Minas Gerais, Sao Paulo, Santa Catharina (Melzer, 1930); Larval host plants are unknown for any species Rio Grande do Sul: Porto Alegre (Melzer, 1930; of Lepturini from Argentina and nearby countries. Buck, 1959), Pared Novo, Cerro Largo (Buck, 1959). Adults are often captured by daylight on Material examined: Brazil: 1 0' (CB), "Ophi­ flowers, feeding on nectar or pollen or copulating: E. stotnis melanura Rdtb." (hand-written by Bruch). latipennis from Rosario de Lerma on Baccharis Taxonomic discussion: The combination set medullosa DC. (Asteraceae); S. dimidiata from by Bruch on the determination label, in spite of Liebig, LN.T.A Station, and Uruguay (Zajciw and being correct (the genus was put into synonymy of Menne, 1970) on Eryngium sp. (Apiaceae); S. Strangalia), never was recorded in the literature. fulvicornis on Eryngium sp. (Zajciw and Morine, The species remained on record under Euryptera 1970). Deliosoma xerophila in Salta (Di Iorio, 1996), (Menne & Giesbert, 1995), although the characters with the same elytral pattern in black and yellow, are unmistakeably those of Strangalia, down to the was also captured together with E. latipennis on the elytra narrowed about the middle (Fig. 14), a same flowers of Baccharis (Viana, pers. com.). character which is not shown by all the species of Variations in elytral patterns of E. latipennis Strangalia, and that distinguished this one from S. and S. dimidiata are concordant with those dimidiata. observed in Lycid and other mimic , Some females of S. dimidiata are similar in visiting the same flowers. Dark and light forms may coloring. In S. melanura, the basal black spot on the correspond to color pattern of different genera and elytra is placed lengthwise and the humeri are species of Lycid models or to a single species of extensively yellow (Fig. 14), while in S. dimidiata Lycid with the same variations. the same spot is placed transversely, while the humeri are narrowly yellow (Fig. 19). Entirely black Key to the Argentine species of Lepturini males of S. melanura (described as var. nigripennis Melzer, 1930) appear to differ from males of S. 1. Elytra broadened backwards, flattened, with 5 dimidiata by their greater size and the shape of ribs; posterior margin of elytra wide, and their elytra. transversely truncate (Fig. 6); body more or less depressed; posterior tibiae not carinate along Strang alia rubricollis (Bates, 1870) inside, and not exceeding posterior margins of elytra; antennomere III shorter than scape (Fig. Ophistomis rubricollis Bates, 1870. 1); first segment of posterior tarsi 2 times as Strangalia rubricollis: Menne and Giesbert, 1995. long as following 2 taken together (Fig. 5): Diagnosis: Entirely black, except for the base Euryptera (Bolivia, northwestern Argentina: of the head behind the eyes, the prothorax, Jujuy, Salta, Tucuman E.latipennis) scutellum and anterior margin of elytra, all of which I'. Elytra narrowed backward, wedge-shaped are reddish. (Figs. 14-26), sometimes narrowed at middle Known geographical distribution: Brazil (Fig. 14), convex, without ribs; posterior margin (Menne and Giesbert, 1995): Amazonas: Ega, now of elytra short and obliquely truncate (Fig. 7); Tefe (Bates, 1870: 312). body not depressed; posterior tibiae longer than Material examined: Peru: Rio Marafion, elytra, carinate along their inner face, and Montenegro, 12-XI-1961, Willinerleg., 10' (ODI). exceeding largely posterior margin of elytra; Remarks: First record of a Lepturini for Peru. antennomere III longer than scape (Figs. 2-3); Similar to Strangalia succincta Redtb. 1868 [var. 1st segment of posterior tarsi twice as long as nigra Redtb.: Brazil: 9-IX-1901, 1 0' (CB)], but in the following 2 taken together (Fig. 4) Strangalia ... latter the base of the head, behind the eyes, is black; 2 the light red pronotum bears about the center an U­ shaped spot; the general shape of the body is more 2. Ventral side of the body entirely black 3 elongate and slender. Strangalia tristis Melzer, 2'. Ventral side of the body reddish brown (except 1922 differs in being entirely black, still more metacoxae, and posterior margins of uro­ elongate and slender than Strangalia succincta sternites 1st to 4th, (5th black); elytral pattern (var. nigra), with the abdomen longer than the as in Fig. 26. Northeastern Argentina: Misiones, elytra, and the last apparent sternite modified Corrientes (Fig. 27) . (males). . Strangalia fulvicornis (females) INSECTA MUNDI, Vol. 12, Nos. 1 & 2, March-June, 1998 145

3. Elytra totally black, including humeri, with small Burmeister, H. 1865. Longicornia argentina. Stet­ basal yellow spot (Fig. 25); all femora and tibiae tiner entomologische Zeitung 26: 162. reddish. Northeastern Argentina: Corrientes Di Iorio, O. R. 1996. The genus Deliosoma (Fig. 27) Strangalia fulvicornis (male) Thomson,1864 (Coleoptera: Cerambycidae) in 3'. Elytra totally black, excluding humeri (females), Argentina: D. lacordairei Thomson 1864, D. without small basal yellow spot (Figs. 15-16), or :x:erophila n. sp., with a revision of larval yellow area increasing progressively (Figs. 17 to galleries and pupal chambers in Neotropical 24); if elytra entirely black, including humeri . Giornale Italiano di Ento­ (males), all femora and tibiae also black. mologia (in press). Northeastern Argentina: Misiones, Corrientes Gounelle, E. 1911. Liste des cerambycides de la to Entre Rios, and Buenos Aires (Fig. 27) . region de Jatahy, Etat de Goyaz, Bresil. ... Strangalia dimidiata Annales de la Societe Entomologique de France 80:103-252. References Lewis, J. P., and M. B. Collantes. 1973. El Espinal periestepico, Ciencia e Investigaci6n 29: Audinet-Serville, Jean Guillaume 1828. Encyl. 360-377. Meth.l0(2):345-832. Linsley, E. G., and J. A Chemsak. 1971. An Bates, H. W. 1870. Contributions to an fauna attempt to clarify the generic status of some of the Amazon Valley (Coleoptera, Ceram­ Neotropical species currently assigned to bycidae). Trans. Entomol. Soc. London 1870: Euryptera, Chontalia and Ophistomis (Coleo­ 243-335, 391-444. ptera, Cerambycidae). Arquivos de Zoologia, Sao Bates, H. W. 1872. On the longicorn Coleoptera of Paulo 21(1):1-40. Chontales, Nicaragua. Trans. Entomol. Soc. Linsley, E. G., and J. A Chemsak. 1972. London 1872: 163-238. Cerambycidae of North America. Part VI, N° 1. Belon, R. P. 1897. Remarques sur le genre Eury­ Taxonomy and classification of the subfamily ptera Serv., du group des lepturides et Lepturinae. University of California, Publi­ description d'une especie nouvelle de Bolivie. cations in Entomology 69:1-138. Annales Societe entomologique de Belgique 41: Linsley, E. G., and J. A. Chemsak. 1976. 339-342. Cerambycidae of North America. Part VI, N° 2. Berg, C. 1889. Quadraginta Coleoptera nova Taxonomy and classification of the subfamily Argentina. Anales de la Universidad de Buenos Lepturinae. University of California, Publi­ Aires 6:105-157. cations in Entomology 80:1-186. Blackwelder, R. E. 1946. Checklist of the Coleo­ Melzer, J. 1922. Longicornios (Coleoptera) do pterous insects of Mexico, Central America, the Brasil novos ou pouco conhecidos. Notas West Indies, and South America. Part 4. Bull. Preliminares do Museu Paulista 2(2):1-12. U. S. Nat. Mus., Smithsonian Institution 185: Melzer, J. 1927. Longicornios do Brasil, novos ou 551-763. pouco conhecidos. Revista do Museu Paulista, Bosq, J. M., and A Ruffinelli. 1951. Notas para el Sao Paulo 15:137-202. catalogo de Cerambicidos del Uruguay. Comun­ Melzer, J. 1930. Longicornios do Brasil, novos ou icaciones en Zoologia del Museo de Historia pouco conhecidos. II. Arquivos do Instituto Natural de Montevideo 3(62):1-32. Biologico, Sao Paulo 3:189-208. Bruch, C. 1912. Catalogo sistematico de los cole6­ Morine, M. A, and E. F. Giesbert. 1995. Checklist pteros de la Republica Argentina. Pars VIII. of the Cerambycidae and Disteniidae (Coleo­ Familia Cerambycidae (prionini, Cerambycini, ptera) of the Western Hemisphere. Wolfsgarden Lamiini). Revista del Museo de La Plata 18: 19­ Books, Burbank, California, xiv + 420 p. 226. Monne, M. A, and D. Zajciw. 1970. Cerambicidos Bruch, C. 1915. Suplemento al catalogo sistematico del Uruguay, nuevos 0 poco conocidos. II. Atas de los cole6pteros de la Republica Argentina. I da Sociedade Bio16gica do Rio de Janeiro 13:29­ (addenda, corrigenda y resumen). Revista del 32. Museo de La Plata 19:538-573. Morello, J. 1958. La provincia fitogeografica del Buck, P. von. 1959. Cerambycidae in del' Monte. Opera Lilloana 2:5-155. Sammlung des Instituto Anchietano de Prado, D. E. 1993. Contribution to the study of the Pesquisas, Porto Alegre, Brasilien. Pesquisas 3: flora and vegetation of the Chaco. VII. What is 577-609. the Gran Chaco vegetation in South America? 146 Vol. 12, Nos. 1 & 2, March-June, 1998, INSECTA MUNDI

II. A redefinition. Candollea 48:615-629. Zajciw, D. 1972. Contribuicao para 0 estudo da Redtenbacher, L. 1868. Reise des osterreichischen fauna dos longicornios do Parque Nacional do Fregatte Novara urn die Erde in den Jahren Itatiaia (Coleoptera, Cerambycidae). Brasil 1857, 1858, 1859 unter den befehelen des FlorestalI2:40-72. Commodore B. von Wullerstorf-Urbair. Wien, Zajciw, D. 1973. Contribuicao para 0 estudo da Zoologischer Theil, Zweiter Band, Coleopteren, fauna dos longicornios (Coleoptera, Ceram­ 249p. bycidae) das florestas do estado do Espiritu Thomson, J. 1857. Description de cerambycides Santo e principalmente da Reserva Biologica nouveaux ou peu connus de rna collection. Arch. "Soore-tama". Ministerio da Agricultura, Ins­ Ent. Paris 1: 291-320. tituto Brasileiro de Desenvolvimento Florestal, Thomson, J. 1860. Essai d'une classification de la Boletim Tecnico 4:37-91. famille des cerambycides et materiaux pour Zajciw, D. and A. Ruffinelli. 1962. Fauna de los servir a une monographie de cette famille. Paris cerambicidos del Uruguay. Boletin de la 404p. Universidad de la Republica, Facultad de Viana, M. J. 1972. Aporte al catalogo de Agronomia de Montevideo 60:1-89. Cerambycidae del Paraguay (Insecta, Coleo­ Zajciw, D. and M.A. Menne. 1968. Cerambicidos ptera). Revista del Museo de Ciencias Naturales del Uruguay, nuevos 0 poco conocidos. Revista "Bernardino Rivadavia," Entomologia 3(4): de la Sociedad Uruguaya de Entomologia, Mon­ 207-405. tevideo 7:51-61. Villiers, A. 1978. Faune des Coleopteres de France. I. Cerambycidae. Encyclopedia Entomologique. XLII. Lechevalier. Paris 611 pp. Zajciw, D. 1958. Fauna do Distrito Federal. XLVIII. Contribuicao para 0 estudo dos longicornios do Rio de Janeiro (Coleoptera, Cerambycidae). Boletim do Museu Nacional, Nova Serie, Zoologia 189:1-26. INSECTA MUNDI, Vol. 12, Nos. 1 & 2, March-June, 1998 147

5

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v- __ 6

6 o10

Figs. 1, 5-6, 8 Euryptera latipennis: 1, right Figs. 11-26, Elytral patterns: 11-13, Euryptera antenna; 5, hind tarsi; 6, apical portion of left latipennis; 14, Strangalia melanura (male); 15-24, elytron; 8, 5th visible sternite; Figs. 2, 4, 7, 10, Strangalia dimidiata: 15-18, dark forms; 19-24 light Strangalia dimidiata (female): 2, right antenna; 4, forms; 25-26 Strangalia fulvicornis: 25, males; 26, posterior tarsi; 7, apical portion of left elytron; 10, females. 5th apparent urosternite; Figs. 3, 9, Strangalia dimidiata (male): 3, right antenna; 9, 5th visible sternite. 148 Vol. 12, Nos. 1 & 2, March-June, 1998, INSECTA MUNDI

72'

) )

rt 28:'- I

32° 32

f\ 27

Fig. 27, Geographical distributions of Lepturini: stars, Euryptera latipennis; triangles, Strangalia dimidiata; arrow, Strangalia fulvicornis, wide horizontal stripes: Province of Chaco (according to Prado, 1993); diagonal stripes: Province of Monte (according to Morello, 1958); vertical stripes: Province of the Espinal (according to Lewis and Collantes, 1973); narrow horizontal stripes: Paranaense Province (a: Transition Austro-brazilian Forest; b: Paranaense Forest; c: Gallery Forest).