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UNESCO : Man and the Biosphere National Committee for Sri Lanka k.. Publication No: 13

A CHECK - LIST OF LAND SNAILS OF SRI LANKA by Ranjini Ratnapala & K. D. Arudpragasam

Published by the Natural Resources, Energy &Science Authority of Sri Lanka. 4715, Maitland Place, . , Colombo 7. UNESCO : Man and the Biosphere Natianai Cnmaitloc for Sri lanka Pllblisrtion No: 13

A CHECK - LlST OF LAND SNAILS OF SRI LAWKA by Ranjini Ratnapala & K. D. Arudpragasam

Published by the Natural Resources, Energy 8: Science Authority of Sri Lanka" 47/5, Maitlandl Place, Colombo 7. A Check- list of Land Saaills of Sri Lanka

RAIVJXNI RATNAPALA & K. D. AWUDPRAGASAM

There is no published check list of the land snails of Sri Lanka. They have been described along with the terrestrial snails of India, Burma, Nepal etc. in three volumes on Mollusca which form part of the Fauna of British India Series. These three volmes still remain our standard source of reference on terrestrial gastropods. They record 12 families, 38 genera and 241 species including 23 varieties, from Sri Lanka. Subsequent to the publication of these volumes our land snails have received very little attention and recent publications on them have been extremely scarce.

The present check list puts together, for the first time the families, genera and species of land snails occuring in Sri Lanka. Thc classification of snails followed in Fauna of British India has undergone considerable change with the alteration of many family and generic nzrnes. Therefore they have been re-arranged according to the scheines of classification of Pulmonata and Prosobran- chia by Zilch (1960) and Wenz (1944) respectively. The check list includes 23 families, 54 genera and 266 species and 22 varieties of land snails.

Sri Lznka is the only recorded locality of the species and genera of land snails marked with an asterisk in this check list.

LIST OF GENERA

Streptaxis Oligospira Ptycho.rema Flanispira Cryptozona Beddomea Hemiplecta Braciybaena Ravana Plectotropis Euplecta Clausiliia Ratnadvipia Achatina Mariaella Subulina Satie!la Opeas Sitala Zootecus Macrochlamys Glessula Microcystina Digoniaxis Kaliella Succinea Eurychlmys Vaginulus Trochomorpha Leptopoma Thysmota Japonia Philalanka Leptopomoidcs Ruthvenia Micraulax Corilla Scabrina Pupisoma Theobaldius ?yramniduln Cyclophorus Ena Aulopoma Rachis so. Pterocyclus Pupoides '51. Cyathopoma Pupilla Tortulosa Gastrocopta Nicida Acavus Tuncatella A C.HECB LIST OF THE LAAD SNAILS OF S3WIE[ LANKA

PHYLUM - MOLLUSCA CLASS - GASTROPODA SUBCLASS - PULMONATA ORDER - STYLOMMATOPXORA SERIES - STREPTAXACEA FAMILY - STWTAXIDAE SUBFAMILY - STREPTAXINAE Genus - STREPTAXIS Gray, 1837

*S. layardinalrs Benson *S. yeroteti Petit * S. cingalensis Benson * S.. gracilis Collett +S. ravanae Blaaford SUBFAMILY - ENNEINAB Genus - PTYCHOTREMA (Morch) 1. Pfeiffer, 1853 Subgenus - Ennea H & A Adam, 1855 P. (Ennea) bicolor Hutton P. (Ennea) planguncula Benson SERIES - ARIOPHANTACEA FAMILY - ARIOPHANTIDAE SUBFAMILY - ARIOPHANTINAE Genus - CRYPTOZONA Morch, 1872 Subgenus - Nilgiria Godwin-Austen, 1888 C (Nilgiria) semirugata Beck

*C (Nilgiria) novella Pfeiffer * C (Nilgiria) ceraria Benson Subgenus - Xestinu Pfeiffer, 1878 C (Xestina) bistrialis Beck * C (Xestina) ceylanica Blanford + C (Xestina) taprobanensis Blanford + C (Xestina) cyix Blanford Genus - HEMIPLECTA Albers, 1850 * H. cherzui Pfeiffer +H. juliana Gray *Haganoma Pfeiffer Genus - RAVANA Godwin-Austen, 1901 R. politisha Pfeiffer Genus - EUPLECTA Semper 1870 E. subopaca Pfeiffer *E. layardi Pfeiffer E. semidecussnta Pfeiffer * E. rosamonda Benson E. travencorica Benson E. praeeminens Blanford E. indica PfeiEer E. albizonata Dohrn *E. emiliana PfeiEer * E. laevis Blanford *E. binoyaensis Godwin - Austen *E. verrucufa Pfeiffer *E. prestoni Godwin - Austen SE. scobinoides Sykes % E. gardneri Pfeiffer *E. partita Pfeifier QE. trimeni Jousscaume E. acuducta Benson * * E. isabellina Pfeiffer SE. colletti Sykes *E. hyphasma (Pfeiffer) rE. turritella (H. Adams) *E. phidias (Thorp) *E. concavospira (F'feiffer)

Genus - RATNADVIPIA Godwin - Austen, '1899 R. irridians Pfeiffer *R. edgariana Benson

SUBFAWTLY - GIRASIINAE

Genus - MARIAELLA Gray, 1855 * M. dussr4mieri Gray SUBFAMILY - DURGELLINAE Genus - SATIEELA Godwin Austen, 1908 QS. membranacea Benson Gems - @TALA H. Adams, 1865 *S. phyllophila Benson *S. (?) pyramidalis Sykes *S. (4) operiens Sykes

SUBFAMILY

Genus - MACROCHLAMYS, Benson, 1832 M. indica Godwin - Austen *M. umbrina Pfeiffer *M. nepas Pfeiffer M. woodinna Pfeiffer M. vilipensa Benson *M. perfucata Benson *M. tratanensis Jousseaume * M. lcandiensis Godwin - Austen

Genus - MICROCYSTINA Morch, 1872 .Yc M. bintennensis Godwin - Austen +M. lita Sykes

SUBFAMILY - SESARINAE Genus - KALIELLA Blandford, 1863 K. barrakporensis Pfeiffer + K. salicensis Godwin - Austen *K. colletti Sykes +K. delectabiiis Sykes *K. leithiana Godwin - Austen

FAMILY - EUCONULIDAE SUBFAMILY - EUCOWULINAE Genus - EURY CWLAMYS Godwin - Austm 1899 QE. regrrlata Benson

FAMILY - TROCHOMORPHIDAE Genus - TROCHOMORPHA Albers, 1850 ST. galerus Benson *T. hyptiocyclos Benson

SERIES - ENDODONTACEA FAMILY - ENDODONTIDAE SUBFAMILY - ENDODONTINAE

Genus - THYSANOTA Albers, 1880 IT. eumita Sykes * T. Rispida Sjkes ST. elegans Preston

Genus - PHILALANKA Godwin - Austen, 1898 QP. szcessa Godwin - Austen, *P. (7) liraiula Pfeiffer *P. (9) laincabensis Jousseatune SP. trifilosa Pfeiffer 4(?) morronema Benson *P. thlvaitesi Pfeiffer Jc Var. suavis Joilssmume *P. depvessn Preston *P. circumsccllpta Sykes +P. (?) sin!lila Godwin - Austen Genm - RUTHVENIA Gude, 1911 *R. clathrctll!a Pfeiffer * Vor. compressa Syklies *R.caligiiwsa Sykes *R. biciliafa Heiffer SERIES - POLYGYRACEA FAMILY - CORILLIDAE Genus - CORILLA H & A Adams, 1855 +C. adamsi Gude * Var. hinidrmensis Nevi11 * C. berldomeae (HanIey) Q C, gzldei Sykes * C. humbevri Brot. * C. fryae Gude C. carabinata (Ferussac) *C. colletti Sykes C. ervonea Albers C.enonea, Var. eronclla, Gude *C. Iesleyae Barnacle C. odontophora, Benson SEWTES - PUPILLACEA FAMILY - VALLOWIIDAE SUBFAWLLY - ACANTHINULINAE Genus - PUPISOMA Stolitzka, 1873 JcP. miccyla Benson *P. longstaffoc Godwin Austen FAMILY - PYiZAMIDULDAE Genus - PYRAMIDULA Fitzinger, 1833 +P. (?) halyi Jousseaunle

FAMILY - ENIDAE SUBFAMILY - ENINAE

Genus - ENA (leach) Turton, 1831) * E. proletarin Meiffer +E. stalix Benson +E. panos Benson

SUBFAMILY - CERASTUINAE Genus - RACEXIS Albers, 1850

Genus - RACHIS Albers, 1850 (=Raclzisellus Bourguignat, 1889) *R. adurnbratus (Pfeiffer) R. pulcher (Gray) R. punetatus (Anton) FAMILY - PUPILEIDAE SUBFAMILY - PUPILLINAE Genus - PUPOIDES Pfeiffer, 1854 P. coenopictus Hutton Genus - PUPILLA Fleming 1828 P. cinghalensis Gude Pi mz!scerd'n Benson

FAMILY - CHONDRINWAE SUBFAMILY - GASTROCOPTINAE Genus - GASTROCOPTA Wollaston 1878 (= BIFIDARIA Sterki 1891) G. rninlzila Senson SERIES - ACAVACEA FAMILY - ACAVLDAE SUBFAMILY - ACAVINAE Genus - ACAVUS Montfort, 1810

*A. haemastoma Lime SVar. rnelanotvagus Born War. conus Pilsbry War. concolor Pilsbry *A. fastosus Albers *A. prosperus P,!bers *A. phoenix Pfeiffer *A. superbus PfeiKer * Var. grevillei Pfeifler *A. roseolabiat~~sNevill "Genus - OLIGOSPIRA Ancey, 1887 SO. waltoni (Reeve) *Var. polei (Collett) * 0.skinneri (Iieveeve)

SERIES FAMILY SUBFAMILY - PLANISPIRA Beck 1837 P. fallaciosa (Feri~ssac) P. vittata Muller Var. albina Grateloup

FAMILY - CAMAENIDAE SUBFAMILY - CAMAENIDAE Genus - BEDDOMEA G. Nevill, 1878 QB. ceylanicus (Pfeiffer) *B. trifasciatus (Gmelin) War. rufoptica (Benson) *B. albizonatus (Reeve) *Var. simoni (Jousseanme) *B. intermedius (Pfeiffer)

FAMILY - BRADYBAENIDAE Genus - BRADYBAENA Beck, 1837 (= EULOTA Hartmann, 1843) B. similaris (Ferussac) Genus - AEGISTA Albers 1850 Subgenus - PLECTOTROPIS Von Martens, 1860 A. (F'lectotropis) hutfoni Pfeiffer War. radleyi Jousseaume

SERIES - CLAUSILIACEA FAMILY - CLAUSILIIDAE SUBFAMILY - CLAUSILIINAE Genus - PHAEDUSA H & A ADAMS, 1855 *P. ceylanica Benson SERIES - ACHATINACEA FAMILY - ACHATINIDAE Genus - ACHATINA Lamarck 1799 A fulica Ferussac FAMILY - SUBULINDAE SUBFAMILY - SUBULININAE Genus - SUBULINA Beck, 1837 S. octona Bruguiere Genus - OPEAS Albers, 1850 0.gracile (Hutton) War. panayaensis Pfeiffer SO. prestoni; Sykes * 0. mariae Jousseaume * 0. layardi Benson SO.sykesi Pilsbry * 0. (?) pussilus H. Adams

SUBFAMILY - RUMININAE Genus - ZOOTECUS Westerlund, 1887 2. insularis (Ehrenberg) FAMILY - FERUSSACIIDAE Genus - GLESSULA Von Martens, 1860 + G. ceylanica Pfeiffer G. inornata Pfeiffer War. minor Beddome G. lankana Pilabry * G. reynelli Var. immitis * G. pallens Beddome * G. prestoni *G. parabilis Benson *S. punctogallana Pfeiffer *G. nitens Gray * G. jirlgens Pfefler *G. panaetha Benson *G. serena Benson +G. sinhila Preston *G. layardi Pilsbry G. deshayesi Pfeifier * G. pachycheila Benson War. taprobanica Pilsbry * G. veruina Benson G. sattaraensis Pfeifffer G. capillacea PfeiiTer * G. collettae Sykes pussila Beddome . . G. + G. sirnoni Jousseaume Genus - DIGOMAXIS Jousseaume 1889- *D. cingalensis Benson

SERIES - SUCCINEACEA FAMILY - SUCCINEIDAE SUBFAMLLY - SUCCINEINAE anus - SUCCINEA Drapamaud, 1801 S. ceylanica Pfeiffer SERIES - SOLEOLIFERA FAMILY - VAGINULIDAE Genus - VAGINULUS Ferussac 1821 * V. tenzpletoni Humbert * V. maculautus Templeton * V, reticulatus Westerlund

SUBCLASS - PROSOBRANCHIA ORDER - MEGAGASTOROPODA SERIES - ARCHITAEMOGLOSSA FAMILY - CYCLOPHORIDAE SUBFAMILY - CYCLOPHORMAE Genus - LEPTOPOMA Pfeiffer 1847 Subgenus - Leptopoma H & A Adams, 1855 *L. (Leptopoma) semiclausurn Pfeiffer *L. (Leptopoma) taprobanensis Preston

Subgenus - TROCHOLEPTOPOMA Kobelt & Mollendorff, 1897 *L. (Trocholeptopoma)apicatum Benson .. 5 : 2 +L. (Trocholeptoporna) elaturn Pfeiffer .. 5 : 4 Genus - JAPONA Gould, 1859 * J. binoyae Sykes .. 5 : 6 * J. occulta Sykes .. 5 : 9 * J. vesca Sykes .. 5:13 Genus - LEPTOPOMOIDES Nevill 1878 L. halophilus Benson .. 5:20 *L. conulus Pfeiffer .. 5:19 *L. flammeus Pfeiffer .. 5:19 *L. orophilus Benson .. 5:20 *L. poecilus Pfeiffer .. 5:21 Genus - MICRAULAX Theobald, 1876 M. coeloconus Benson Genus - SCABRINA BlanCord 1863 SS. brounae Sykes

Genus - THEOBALDIUS NeviU, 1873 ST. annulatus PfeifYer ST. bairdi PfeiRer %T. cadiscus Benson *T. cratera Benson ST. cytopoma Benson ST. Iayardi H. Adarns ST. liliputiailcrs Preston ST. Ioxostoma Pfeiier *T. pnrapsis Benson ST. parma Benson ST. subplicatus Beddome ST. thwnitesi Pfeiffer

Gems - CYCLOPIlORUS Montfort, 1810 Subgenus - Litostylus Kobelt & Mollendorff 1897 C. (Litostylus) invo1vul~~sMuller * C. (Litostylus) ceylanicus Pfeiffer a C. (Litostylus) liratula Preston C. (Litostybls) menkeanus Philipps

Subgenus - Annularia Schumacher 1817 *-C. (Salpingophorus) alabastrianus Pfeiffer C. (Salpingophorccs)punctatzrs Grateloup

*Genus - AULOPBhfA Troschel, 1847 PA. hofmeisteri Troschel A. grade Pf ieffer *A. helicinum Chernnitz *A. itieri Guerin *A. sphaeroideum Dohrn

Genus - PTEROCYCLUS Benson 1832 %P. bifrons Pfeiffer P. bilabiatus Sowerby War. conica Nevill *P. ciilgalensis Benson P. czrmingi Pfeiffer +P. froscheli Benson

Genus - CYATHOPOMA W & H Blanford 1861 Subgenus - Cyathopoma W & H Blanford 1861 FFC. (Cynthopoma) artatum Sykes C. (Cyathopoma) album Beddome SC. (Cyathoporia) leptopo,ttita Sykes *C. (Cyathopoma) marine Jousseaurne * C. (Cycrthopomn) prestoni Sykes

Subgenus - Jerdonia W & H BlanFord 1861 1Y- C. (Jerdonia) ceylnnicum Beddome *C. (Jerdonia) collefti Sgkes * C. (Jerdonia) conoideum Sykes QC. (Jerdonia) innocens Sykes *C. (Jerdonia) ogdeniamrrm Preston *C. (Jerdonia) perconoider~nt Preston * C. (Jerdonia) sereitdibense Prestona *C. (Jerdonia) turbinatrrm Sykes *C. (Jerdonia) uvaense Preston

SUBFAMILY - PUPININAE Genus - TORTULOSA Gray, 1847 Subgenus - Eucataulus Kobelt, 1902 ST. aurea Pfeiffer ST. austeniana Benson ST. blanfordi Dohrn ST. colletti Sykes ST. congetzer Sykes ST. connectens Fulton +T. c~drningiPfeiffer ST. decora Benson ST. drqlicata Pfeiffer ST. eurytrema Pfeiffer * T. greeni Sykes *T. robtrsta Fulton *T. recurvata (Pfeiffer) ST. haemastotnu Pfeiffer *T. layardi Pfeiffer *T. nrarginata Pfeiffer War. crenulata Fulton *Var. rtotata Sykes ST. nevilli Sykes +Var. ffaveob Fulton *T. nietneri Ned1 *T. caperata Collett *T. unicolor Collett *T. prestoni Sykes *T. pyramidaia Pfeiffer *T. rugosa Fulton *T. smithi Sykes ST. sykesi Fulton ST. tenple~nailiPfeiffes * T. ti'zrvaitesi Pfeiffer

SUBFAMILY - DJPLOMMATRJMAE Genus - NICIDA Blanford, 1868 +N. catathymia Sykes *1V. cee)~lanicuBeddolne *N. delcctabilis Pesston *N, lankaerrsis Preston *N. pedrorris Beddome *N. prestorli Beddome

SER[ES FAMILY SUBFAMILY Gnus REFERENCE

1. BARVACLE,G: A. S. (1956) - A new spies of Corllla from Ceylon ,J. Conch London. Vol. 24 : 95-96, pl. 4

2. (1962) - Notes on the Genus Acavus. Montfo~t, J. Conch VOI.25 :128-132.

3. BLANFORD,W. T. & GODWLN- Mollusca, Testacellidae & Zonitidae : Fauna of British AUSTBN,R. H. (1905) India.

4. GUDB,G. K. (1914) - Mollusca Vol. 11, Trochomorphid.ae, Jandlidae, Fauna of British India Series.

5. (1921) - Mollusca Vo'o!. 111, Land Operculates, Fauna of British India Series.

6. SATYAMURTI,S. T. (1960) - Land and Fresh Water Mollusca in the collection of the Madras Museum. Bull. Madras Govt. Mus. New. Ser. Nat. Hist. Sec.

7. WENZ,W. (1938 - 1944) - Handbuch der Palaozoologie, Band 6, Gastropoda Teil 1: Allgemeiner Teil und Prosobranchia.

8. ZZ&, A. (1959 - 1960) - Kandbuch der PalaozoologieBand 6, Gastropoda Teil2, Euthynewa.