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Gymnosperm Foliage from the Upper Triassic of Lunz, Lower Austria: an Annotated Check List and Identification Key
Geo.Alp, Vol. 7, S. 19–38, 2010 GYMNOSPERM FOLIAGE FROM THE UPPER TRIASSIC OF LUNZ, LOWER AUSTRIA: AN ANNOTATED CHECK LIST AND IDENTIFICATION KEY Christian Pott1 & Michael Krings2 With 7 figures and 1 table 1 Naturhistoriska riksmuseet, Sektionen för paleobotanik, Box 50007, SE-104 05 Stockholm, Sweden; [email protected] 2 Department für Geo- und Umweltwissenschaften, Paläontologie und Geobiologie, Ludwig-Maximilians-Universität, and Bayerische Staatssammlung für Paläontologie und Geologie, Richard-Wagner-Straße 10, 80333 München, Germany; [email protected] Abstract The famous Lunz flora from Lower Austria is one of the richest and most diverse Late Triassic floras of the Northern He- misphere. The historical outcrops (mainly coal mines) are no longer accessible, but showy fossils can still be collected from natural exposures around the town of Lunz-am-See and from several of the old spoil tips. This paper presents an annotated check list with characterisations of all currently recognised gymnosperm foliage taxa in the Lunz flora. The descriptions are exemplified by illustrations of typical specimens and diagnostic features of the leaf morphology and epidermal anatomy. Moreover, a simple identification key for the taxa based on macromorphological features is provided that facilitates identification of newly collected specimens. 1. Introduction The Carnian (Late Triassic) flora from Lunz in Lo- ments (i.e. reproductive structures) among the fossils wer Austria is one of only a few well-preserved flo- (see e.g., Krasser, 1917, 1919; Kräusel, 1948, 1949, ras from the Alpine Triassic (Cleal, 1993; Dobruskina, 1953; Pott et al., 2010), the most striking feature of 1998). -
Variation in Sex Expression in Canada Yew (Taxus Canadensis) Author(S): Taber D
Variation in Sex Expression in Canada Yew (Taxus canadensis) Author(s): Taber D. Allison Source: American Journal of Botany, Vol. 78, No. 4 (Apr., 1991), pp. 569-578 Published by: Botanical Society of America Stable URL: http://www.jstor.org/stable/2445266 . Accessed: 23/08/2011 15:56 Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at . http://www.jstor.org/page/info/about/policies/terms.jsp JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact [email protected]. Botanical Society of America is collaborating with JSTOR to digitize, preserve and extend access to American Journal of Botany. http://www.jstor.org AmericanJournal of Botany 78(4): 569-578. 1991. VARIATION IN SEX EXPRESSION IN CANADA YEW (TAXUS CANADENSIS)1 TABER D. ALLISON2 JamesFord Bell Museumof Natural History and Departmentof Ecology and BehavioralBiology, Universityof Minnesota, Minneapolis, Minnesota 55455 Sex expressionwas measuredin severalCanada yew (Taxus canadensisMarsh.) populations of theApostle Islands of Wisconsinand southeasternMinnesota to determinethe extent of variationwithin and among populations. Sex expression was recorded qualitatively (monoecious, male,or female) and quantitatively (by male to female strobilus ratios or standardized phenotypic gender).No discernibletrends in differencesin sex expressionamong populations or habitats wererecorded. Trends in sexexpression of individuals within populations were complex. Small yewstended to be maleor, if monoecious, had female-biasedstrobilus ratios. -
Earliest Record of Megaphylls and Leafy Structures, and Their Initial Diversification
Review Geology August 2013 Vol.58 No.23: 27842793 doi: 10.1007/s11434-013-5799-x Earliest record of megaphylls and leafy structures, and their initial diversification HAO ShouGang* & XUE JinZhuang Key Laboratory of Orogenic Belts and Crustal Evolution, School of Earth and Space Sciences, Peking University, Beijing 100871, China Received January 14, 2013; accepted February 26, 2013; published online April 10, 2013 Evolutionary changes in the structure of leaves have had far-reaching effects on the anatomy and physiology of vascular plants, resulting in morphological diversity and species expansion. People have long been interested in the question of the nature of the morphology of early leaves and how they were attained. At least five lineages of euphyllophytes can be recognized among the Early Devonian fossil plants (Pragian age, ca. 410 Ma ago) of South China. Their different leaf precursors or “branch-leaf com- plexes” are believed to foreshadow true megaphylls with different venation patterns and configurations, indicating that multiple origins of megaphylls had occurred by the Early Devonian, much earlier than has previously been recognized. In addition to megaphylls in euphyllophytes, the laminate leaf-like appendages (sporophylls or bracts) occurred independently in several dis- tantly related Early Devonian plant lineages, probably as a response to ecological factors such as high atmospheric CO2 concen- trations. This is a typical example of convergent evolution in early plants. Early Devonian, euphyllophyte, megaphyll, leaf-like appendage, branch-leaf complex Citation: Hao S G, Xue J Z. Earliest record of megaphylls and leafy structures, and their initial diversification. Chin Sci Bull, 2013, 58: 27842793, doi: 10.1007/s11434- 013-5799-x The origin and evolution of leaves in vascular plants was phology and evolutionary diversification of early leaves of one of the most important evolutionary events affecting the basal euphyllophytes remain enigmatic. -
Palaeogeograph Y, Palaeoclimatology, Palaeoecology , 17(1975): 157--172 © Elsevier Scientific Publishing Company, Amsterdam -- Printed in the Netherlands
Palaeogeograph y, Palaeoclimatology, Palaeoecology , 17(1975): 157--172 © Elsevier Scientific Publishing Company, Amsterdam -- Printed in The Netherlands CLIMATIC CHANGES IN EASTERN ASIA AS INDICATED BY FOSSIL FLORAS. II. LATE CRETACEOUS AND DANIAN V. A. KRASSILOV Institute of Biology and Pedology, Far-Eastern Scientific Centre, U.S.S.R. Academy of Sciences, Vladivostok (U.S.S.R.) (Received June 17, 1974; accepted for publication November 11, 1974) ABSTRACT Krassilov, V. A., 1975. Climatic changes in Eastern Asia as indicated by fossil floras. II. Late Cretaceous and Danian. Palaeogeogr., Palaeoclimatol., Palaeoecol., 17:157--172. Four Late Cretaceous phytoclimatic zones -- subtropical, warm--temperate, temperate and boreal -- are recognized in the Northern Hemisphere. Warm--temperate vegetation terminates at North Sakhalin and Vancouver Island. Floras of various phytoclimatic zones display parallel evolution in response to climatic changes, i.e., a temperature rise up to the Campanian interrupted by minor Coniacian cooling, and subsequent deterioration of cli- mate culminating in the Late Danian. Cooling episodes were accompanied by expansions of dicotyledons with platanoid leaves, whereas the entire-margined leaf proportion increased during climatic optima. The floristic succession was also influenced by tectonic events, such as orogenic and volcanic activity which commenced in Late Cenomanian--Turonian times. Major replacements of ecological dominants occurred at the Maastrichtian/Danian and Early/Late Danian boundaries. INTRODUCTION The principal approaches to the climatic interpretation of fossil floras have been outlined in my preceding paper (Krassilov, 1973a). So far as Late Creta- ceous floras are concerned, extrapolation (i.e. inferences from tolerance ranges of allied modern taxa) is gaining in importance and the entire/non-entire leaf type ratio is no less expressive than it is in Tertiary floras. -
Ginkgo Biloba Maidenhair Tree1 Edward F
Fact Sheet ST-273 November 1993 Ginkgo biloba Maidenhair Tree1 Edward F. Gilman and Dennis G. Watson2 INTRODUCTION Ginkgo is practically pest-free, resistant to storm damage, and casts light to moderate shade (Fig. 1). Young trees are often very open but they fill in to form a denser canopy. It makes a durable street tree where there is enough overhead space to accommodate the large size. The shape is often irregular with a large branch or two seemingly forming its own tree on the trunk. But this does not detract from its usefulness as a city tree unless the tree will be growing in a restricted overhead space. If this is the case, select from the narrow upright cultivars such as ‘Princeton Sentry’ and ‘Fairmont’. Ginkgo tolerates most soil, including compacted, and alkaline, and grows slowly to 75 feet or more tall. The tree is easily transplanted and has a vivid yellow fall color which is second to none in brilliance, even in the south. However, leaves fall quickly and the fall color show is short. GENERAL INFORMATION Scientific name: Ginkgo biloba Pronunciation: GINK-go bye-LOE-buh Common name(s): Maidenhair Tree, Ginkgo Family: Ginkgoaceae Figure 1. Middle-aged Maidenhair Tree. USDA hardiness zones: 3 through 8A (Fig. 2) Origin: not native to North America Uses: Bonsai; wide tree lawns (>6 feet wide); drought are common medium-sized tree lawns (4-6 feet wide); Availability: generally available in many areas within recommended for buffer strips around parking lots or its hardiness range for median strip plantings in the highway; specimen; sidewalk cutout (tree pit); residential street tree; tree has been successfully grown in urban areas where air pollution, poor drainage, compacted soil, and/or 1. -
Dr. Sahanaj Jamil Associate Professor of Botany M.L.S.M. College, Darbhanga
Subject BOTANY Paper No V Paper Code BOT521 Topic Taxonomy and Diversity of Seed Plant: Gymnosperms & Angiosperms Dr. Sahanaj Jamil Associate Professor of Botany M.L.S.M. College, Darbhanga BOTANY PG SEMESTER – II, PAPER –V BOT521: Taxonomy and Diversity of seed plants UNIT- I BOTANY PG SEMESTER – II, PAPER –V BOT521: Taxonomy and Diversity of seed plants Classification of Gymnosperms. # Robert Brown (1827) for the first time recognized Gymnosperm as a group distinct from angiosperm due to the presence of naked ovules. BENTHAM and HOOKSER (1862-1883) consider them equivalent to dicotyledons and monocotyledons and placed between these two groups of angiosperm. They recognized three classes of gymnosperm, Cyacadaceae, coniferac and gnetaceae. Later ENGLER (1889) created a group Gnikgoales to accommodate the genus giankgo. Van Tieghem (1898) treated Gymnosperm as one of the two subdivision of spermatophyte. To accommodate the fossil members three more classes- Pteridospermae, Cordaitales, and Bennettitales where created. Coulter and chamberlain (1919), Engler and Prantl (1926), Rendle (1926) and other considered Gymnosperm as a division of spermatophyta, Phanerogamia or Embryoptyta and they further divided them into seven orders: - i) Cycadofilicales ii) Cycadales iii) Bennettitales iv) Ginkgoales v) Coniferales vi) Corditales vii) Gnetales On the basis of wood structure steward (1919) divided Gymnosperm into two classes: - i) Manoxylic ii) Pycnoxylic The various classification of Gymnosperm proposed by various workers are as follows: - i) Sahni (1920): - He recognized two sub-divison in gymnosperm: - a) Phylospermae b) Stachyospermae BOTANY PG SEMESTER – II, PAPER –V BOT521: Taxonomy and Diversity of seed plants ii) Classification proposed by chamber lain (1934): - He divided Gymnosperm into two divisions: - a) Cycadophyta b) Coniterophyta iii) Classification proposed by Tippo (1942):- He considered Gymnosperm as a class of the sub- phylum pteropsida and divided them into two sub classes:- a) Cycadophyta b) Coniferophyta iv) D. -
X. the Conifers and Ginkgo
X. The Conifers and Ginkgo Now we turn our attention to the Coniferales, another great assemblage of seed plants. First let's compare the conifers with the cycads: Cycads Conifers few apical meristems per plant many apical meristems per plant leaves pinnately divided leaves undivided wood manoxylic wood pycnoxylic seeds borne on megaphylls seeds borne on stems We should also remember that these two groups have a lot in common. To begin with, they are both groups of woody seed plants. They are united by a small set of derived features: 1) the basic structure of the stele (a eustele or a sympodium, two words for the same thing) and no leaf gaps 2) the design of the apical meristem (many initials, subtended by a slowly dividing group of cells called the central mother zone) 3) the design of the tracheids (circular-bordered pits with a torus) We have three new seed plant orders to examine this week: A. Cordaitales This is yet another plant group from the coal forest. (Find it on the Peabody mural!) The best-known genus, Cordaites, is a tree with pycnoxylic wood bearing leaves up to about a foot and a half long and four inches wide. In addition, these trees bore sporangia (micro- and mega-) in strobili in the axils of these big leaves. The megasporangia were enclosed in ovules. Look at fossils of leaves and pollen-bearing shoots of Cordaites. The large, many-veined megaphylls are ancestral to modern pine needles; the shoots are ancestral to pollen-bearing strobili of modern conifers. 67 B. -
Gymnosperms on the EDGE Félix Forest1, Justin Moat 1,2, Elisabeth Baloch1, Neil A
www.nature.com/scientificreports OPEN Gymnosperms on the EDGE Félix Forest1, Justin Moat 1,2, Elisabeth Baloch1, Neil A. Brummitt3, Steve P. Bachman 1,2, Stef Ickert-Bond 4, Peter M. Hollingsworth5, Aaron Liston6, Damon P. Little7, Sarah Mathews8,9, Hardeep Rai10, Catarina Rydin11, Dennis W. Stevenson7, Philip Thomas5 & Sven Buerki3,12 Driven by limited resources and a sense of urgency, the prioritization of species for conservation has Received: 12 May 2017 been a persistent concern in conservation science. Gymnosperms (comprising ginkgo, conifers, cycads, and gnetophytes) are one of the most threatened groups of living organisms, with 40% of the species Accepted: 28 March 2018 at high risk of extinction, about twice as many as the most recent estimates for all plants (i.e. 21.4%). Published: xx xx xxxx This high proportion of species facing extinction highlights the urgent action required to secure their future through an objective prioritization approach. The Evolutionary Distinct and Globally Endangered (EDGE) method rapidly ranks species based on their evolutionary distinctiveness and the extinction risks they face. EDGE is applied to gymnosperms using a phylogenetic tree comprising DNA sequence data for 85% of gymnosperm species (923 out of 1090 species), to which the 167 missing species were added, and IUCN Red List assessments available for 92% of species. The efect of diferent extinction probability transformations and the handling of IUCN data defcient species on the resulting rankings is investigated. Although top entries in our ranking comprise species that were expected to score well (e.g. Wollemia nobilis, Ginkgo biloba), many were unexpected (e.g. -
Curriculum Vitae
CURRICULUM VITAE ORCID ID: 0000-0003-0186-6546 Gar W. Rothwell Edwin and Ruth Kennedy Distinguished Professor Emeritus Department of Environmental and Plant Biology Porter Hall 401E T: 740 593 1129 Ohio University F: 740 593 1130 Athens, OH 45701 E: [email protected] also Courtesy Professor Department of Botany and PlantPathology Oregon State University T: 541 737- 5252 Corvallis, OR 97331 E: [email protected] Education Ph.D.,1973 University of Alberta (Botany) M.S., 1969 University of Illinois, Chicago (Biology) B.A., 1966 Central Washington University (Biology) Academic Awards and Honors 2018 International Organisation of Palaeobotany lifetime Honorary Membership 2014 Fellow of the Paleontological Society 2009 Distinguished Fellow of the Botanical Society of America 2004 Ohio University Distinguished Professor 2002 Michael A. Cichan Award, Botanical Society of America 1999-2004 Ohio University Presidential Research Scholar in Biomedical and Life Sciences 1993 Edgar T. Wherry Award, Botanical Society of America 1991-1992 Outstanding Graduate Faculty Award, Ohio University 1982-1983 Chairman, Paleobotanical Section, Botanical Society of America 1972-1973 University of Alberta Dissertation Fellow 1971 Paleobotanical (Isabel Cookson) Award, Botanical Society of America Positions Held 2011-present Courtesy Professor of Botany and Plant Pathology, Oregon State University 2008-2009 Visiting Senior Researcher, University of Alberta 2004-present Edwin and Ruth Kennedy Distinguished Professor of Environmental and Plant Biology, Ohio -
Seasonal Growth of the Female Strobilus in Pinus Radiata
No. 1 15 SEASONAL GROWTH OF THE FEMALE STROBILUS IN PINUS RADIATA G. B. SWEET and M. P. BOLLMANN Forest Research Institute, New Zealand Forest Service, Rotorua (Received for publication 12 November, 1970) ABSTRACT Growth of female strobili of Pinus radiata D. Don from central North Island of New Zealand is described and illustrated with photographs. The two-and-a-half year period from strobilus emergence until cone maturity comprises a seasonal period of growth in which pollination occurs, a second period of seasonal growth in which fertilisation occurs, and finally a period of cone maturation. The periods of rapid growth do not appear to result directly from either pollination or fertilisation, and the seasonal growth periods have some similarity to those of vegetative growth. The time taken to reach cone maturity in P. radiata (a closed-cone pine) is six months longer than that frequently described for other species of Pinus. INTRODUCTION The general pattern of female strobilus development in Pinus is well documented (e.g., Ferguson, 1904; Stanley, 1958; Sarvas, 1962). Broadly, a total period of two-and- a-half years is involved, leading through from strobilus determination one summer, to anthesis the following spring, to fertilisation late in the subsequent spring and finally to maturation the succeeding autumn. Seed production in Pinus radiata D. Don apparently follows within general limits the typical pattern for Pinus, but few details have been published either of its strobilus or its ovule development. As part of a comprehensive study of the processes leading to seed production in this species, material was collected in 1968 and 1969 to enable details of strobilus development to be determined. -
Ginkgoales Dr Moni Kumari Date: 28 and 29Th September B.Sc 2Nd Year
Ginkgoales Dr Moni Kumari Date: 28 and 29th September B.Sc 2nd Year PG Botany Department Gaya College, Gaya Ginkgoales is a gymnosperm order containing only one extant species: Ginkgo biloba, the ginkgo tree It is classified in its Division: Ginkgophyta Class: Ginkgoopsida Order: Ginkgoales Family: Ginkgoaceae Genus: Ginkgo The only extant species within this group Ginkgo biloba is a highly adaptable plant that can grow in almost any temperate or Mediterranean climate. Resistant to pollution and pests. Long lived due to a beneficial combination of disease-resistant characteristics It can grow up to 30 meters tall and can live for a millenium. It is also well-known for its unique seeds, which have long been used as a food source in Asia. Cultivated predominantly in China Parts of the ginkgo tree are commonly added to foods and drinks or taken as a supplement due to its desirable health benefits. Some old ginkgoes produce aerial roots, known as chichi (Japanese; "nipples") or zhong-ru (Mandarin Chinese), which form on the undersides of large branches and grow downwards. Chichi growth is very slow, and may take hundreds of years to occur Ginkgoales was, however, very abundantly represented in the world by several species of about 16 genera during the Triassic period of Mesozoic age, i.e. about 200,000,000 years ago. Today, all the genera, except Ginkgo biloba, are extinct. Due to the presence of a number of primitive characters, as well as because of its long geological records, Ginkgo is called a “living fossil”. Details of the geological history of Ginkgoales indicate that its members started appearing on the earth during Permian Abundance worldwide distribution during Triassic and Jurassic periods of Mesozoic age Started fading out of existence during Cretaceous Now represented only in some parts of Southern and Eastern China by only one living member i.e. -
And Ginkgo Biloba Linn. Leaves
Comparative study between Ginkgoites tigrensis Archangel~ky and Ginkgo biloba Linn. leaves Liliana Villar de Seoane Liliana Villar de Seoane 1997. Comparative study between Ginkgoites tigrensis Archangelsky and GinA!go biIoba Linn. leaves. Palaeobotanist46(3) : 1-12. ' . A comparative study between leaves of Ginkgoites rigrrmsis Archangelsky'1965, which belong to the familf K.ark.e,ni:aceae and those of Ginkgo biloba L. included within the family Ginkgoaceae is realized, using lighI microscopy (LM) and scanning and transmission electron microscopy (SEM and TEM). The fossil cuticles occur in the sediments located in Baquero Formation (Lower Cretaceous), Santa Cruz Province, Argentina. The morphological, anatomical and ultrastructural analyses indicate great similarities between fossil and extant leaves. Key-words----Ginkgoalean leaves, Baquer6 Formation, Lower Cretaceous, Santa Cruz Province, Argentina. Liliana Villar de Seoane, CONICEr, Divisi6n Paleobotanica, Museo Argentino de Ciencias Naturales "B. Rivadavia ". Av. A. Gallardo 470, (1405) Buenos Aires, Argentina. RESUMEN En el presente trabajose realiza el estudio comparado entre hojas de la especie f6sil Ginkgoites tigrrmsis Archangelsky 1965, pel1eneciente a la familia Karkeniaceae, y de la especie actual Ginkgo biloba L. pel1eneciente a la familia Ginkgoaceae, ambas familias incluidas dentro del orden Ginkgoales. Para efectuar el anAlisis cuticularse utiliz6 microscopia 6ptica (MO) y microscopia electronica de barrido y transmision (MEB y MEn. Los reslOS f6siles analizados fueron extraidos de sedimentitas organ6genas hlladas en la Formacion Baquer6 de la Provincia de Santa Cruz (Cret:!icico Inferior). Los resultados obtenidos con los anAlisis realizados, han demostrado la existencia de grandes semejanzas morfologicas, anat6micas y ultraestructurales entre las hojas de las especies Ginkgoites tigrrmsis y Ginkgo biloba.