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and D~uue~h~inn,J. B.: Forest vegetation of eastern Specialization and flexibility in genetic systems of forest . and northern . Wash. Agric. Exp. Sta. Tech. Bull. No. 60 Silvae Genet. 18 (4): 118-123 (1969). - ROWE, K. E. and CHING, K. K.: (1968). - KIIIBBII.,H. B.: Parental and provenance effects on Provenance study of Douglas- in the Region. growth of red oak seedlings. Proc. 4th Cent. States For. Imp. 11. Field performance at age nine. Silvae Genet. 22 (4): 115-119 Conf. p. 19-25 (19664). - JI~NKINSON,J. L.: Ponderosa pine progenies: (1973). - SII.EN,R. R.: Personal communication (1976). - SNEDECO~, differential response to ultramafic and granitic soils. USDA Forest W. and COCHIIAN,W. G.: Statistical Methods. The Iowa State Uni- Service Res. Pap. PSW-101 (1974). - HERMANN,R. K. and LAVENDER, versity Press, Ames, Iowa (1968). - SQIJILLACL:,A. E.: Geographic D. P.: Early growth of Douglas-fir from various altitudes and variation in slash pine. For. Sci. Mono. 10 (1966). - S~iiii.r.~ce,A. E. aspects in southern . Silvae Genet. 17: 143-151 (1968). - LANG- and SII.IIN,R. R.: Racial variation in ponderosa pine. For. Sci. I.ET,0.: TWO hundred years of genecology. Taxon 20 (56): 653-722 Monog. 2 (1962). - STEEL,R. G. D. and Toani~,J. H.: Principles and (1971). - LEVINS,R.: Theory of fitness in a heterogeneous environ- Procedures of Statistics. McGraw Hill, N.Y. (1960). - STERN, K.: ment. I. The Fitness set and adaptive function. Amer. Natur. 96: 361 Herkunftsversuche für die Forstpflanzenzüchtung - erläutert am -373 (1962). - LEVINS, R.: Theory of fitness in a heterogeneous envi- Beispiel zweier Herkunftsversuche mit Birken. Der Züchter 34: ronment. 11. Developmental flexibility and niche selection. Amer. 181-219 (1964). - STERN,K. and ROCIIK,L.: Genetics of Forest Eco- Natur. 97: 75-90 (1963). - LI~VINS,R.: Theory of fitness in a hetero- systems. Spring-Verlag, N.Y. (1974). - TOWNSINI>,J. 1.: Genetics geneous environment. 111. The response to selection. J. of Theoret. of marginal populations of Drosophila willistoni. Evol. 6: 428442 Biol. 7: 224-240 (1964). - LI, C. C.: Population Genetics. Univ. of (1952). - VAN VAI.EN,L.: Morphological variation and width of eco- Chigao Press, Chicago (1955). - LIBRY,W. J. and JLIND,E.: Variance logical niche. Amer. Natur. 99: 377-390 (1965). - WANT., CH. W. and associated with Cloning. Heredity 17: 533-540. - LITTLB,E. L., JR.: PATTEE,R. K.: Variation in characteristics and seedling growth Atlas of Trees. USDA Forest Service Misc. Pub. No. of open-pollinated ponderosa pine progenies. FWR Exp. Sta., Univ. 1146 (1971). - MAINAIII>SMITII, J.: Disruptive selection, polymor- of Idaho Sta. Pap. No. 15 (1974). - VAN DI:I~SEN,J. L.: Five-year phism, and sympatric speciation. Nature. 195: 60-2 (1962). - MAYR, results of a ponderosa pine provenance study in the Black Hills. E.: Anima1 Species and Evolution. Harvard Univ. Press, Cambridge, USDA Forest Service Res. Note RM-27Q (1974). - WELLS,0. 0.: Mass. (1963). - NAMK~ON<:,G., USANIS, R. A. and SILEN, R. R.: Age- Geographie variation in ponderosa pine. I. The ecotypes and their related variation in genetic control of height growth in Douglas- distribution. Silvae Genet. 13 (4): 89-103 (1964 a). - WELI.S,0. 0.: fir. Theoret. and Appl. Gen. 42: 151-9 (1972). - PFISTER, R., KOVAL- Geographic variation in ponderosa pine. 11. Correlations between <.IIII<,P. D. L., AIINO,A. F. and PIIISBY,R. C.: Forest habitat types progeny performance and characteristics of the native habitat. oP . Intermt. For. and Con. Exp. Sta. and Northern Region Silvae Genet. 13 (5): 125-133 (1964 b). - WRIGHT,J. W.: Genetic USFS. Mise. Pub. (1974). - R~:HI:I:I.DT,D. G.: Genetic variation of variation among 140 half-sib Scotch pine families derived from Douglas-fir in the northern , USDA Forest Service 9 stands. Silvae Genet. 12: 83-9 (1963). - WIIIGI~T,J. W.: Introduc- Res. Note INT-184 (1974). - REI~FELDT,G. D. and LISTER,D. T.: tim to Forest Genetics. Academic Press, N.Y. (1976).

An Intergrated Population of Abies grandis - in Central Idaho and its Relation to Decay')

Cooperative Tree Improvement and Hardwood Research Programs North Carolina State University Raleigh, North Carolina 27667 USA

(Received July 1976 1 February 1977)

Introduction Several authors recognize A. concolor as occurring in these two regions (DAVIS,1952 and PECK, 1961). Many dis- Grand fir (Abies grandis [DOUGL.ex. D. DON] LINDL.) and claim its occurrence (MCMINNand MAINO, 1947; FOWELLS, white or concolor fir (Abies concolor [GORDON and GLEND.] 1965; and E. L. LI~TLE,personal communication, 1975). LINDL. ex. HILDEBR.) are two important and widely distribut- ed true in the Western United States. Grand fir occurs These two species hybridize freely in areas of overlap in primarily in the Pacific Northwest and in the Rocky Moun- California and western Oregon (HAMRICICand LIBBY, 1972). tains from central Idaho to southern . There are no reports, however, of verified hybrids occur- White fir grows in the southern Cascade, Sierra Nevada, and ring in the northern Rocky Mountains. Rocky Mountains south of the range of grand fir (LITTLE, In central Idaho* south of the Salmon River (approxi- 1971). The southern Rocky Mountains white fir is often mately 46O 30' N. Lat.) an intergraded population apparent- referred to as A. concolor var. concolor and the Sierran to ly exists between concolor fir and grand fir. JOHNSON(1966) A. concolor var. lowiana. Presently grand fir and concolor recognized this morphological complex in west-central fir are regarded as morphologically distinct (Table 1). Be- Idaho and tentatively interpreted it as an intergraded pop- tween the undisputed ranges of grand fir and white fir lies ulation of A. concoLor, var. lowiana, A. grandis and trees an area in northeastern Oregon and west central Idaho sup- which exhibited characteristics of both species. DANIELS porting fir populations that are morphologically inter- (1969) reported typical grand fir types, intermediate forms, mediate and not easily assigned to either spccies. and concolor-like individuals growing together near McCall, Idaho. He studied the phenotypic variability of the

I) Supported by McIntire-Stennis, U.S. Department of Agricul- population and generally outlined its composition and ex- ture. tent using a hybrid index technique. Individuals in his

Silvae Genetica 26, 1 (1977)