AFROTROPICAL . MARK C. WILLIAMS. http://www.lepsocafrica.org/?p=publications&s=atb Updated 12 December 2020

Genus Kirby, [1871] Bush Browns

In: Kirby, [1871], [1877]. A synonymic catalogue of diurnal 47 (690 pp. (1871); Suppl. 691-883 (1877). London). [Replacement name for Idiomorphus Doumet.]

= Idiomorphus Doumet, 1861. Revue et Magasin de Zoologie Pure et Appliqué (2) 13: 174 (171- 178). Type-species: Idiomorphus hewitsonii Dourmet, by monotypy. [Invalid; junior homonym of Idiomorphus de Chaudoir, 1846.]

= Monotrichtis Hampson, 1891. Annals and Magazine of Natural History (6) 7: 179 (179-184). Type-species: Mycalesis safitza Westwood, by original designation.

= Dicothyris Karsch, 1893. Berliner Entomologische Zeitschrift 38: 203 (1-266). Type- species: Mycalesis sambulos Hewitson, by subsequent designation (Hemming, 1935. Stylops 4: 1 (1-3).).

The Bicyclus belongs to the Family Rafinesque, 1815; Subfamily Boisduval, 1833; Tribe Satyrini Boisduval, 1833; Subtribe Mycalesina Reuter, 1896. The other genera in the Subtribe Mycalesina in the Afrotropical Region are Hallelesis, Heteropsis and Brakefieldia.

Bicyclus (Bush Browns) is an Afrotropical genus containing 102 species (October, 2016). Generic review by Condamin (1973). An updated phylogenetic hypothesis was published by Aduse-Poku et al. in 2016.

CURRENT PHYLOGENETIC HYPOTHESIS FOR THE GENUS BICYCLUS (after Aduse-Poku et al., 2016)

SPECIES-GROUP INCLUDED TAXA evadne-group evadne, howarthi, xeneas, alboplaga, xeneoides nobilis-group nobilis, larseni trilophus-group trilophus, jacksoni, sealeae ignobilis-group ignobilis, maesseni, brakefieldi, vandeweghei, ottossoni, rileyi (Brattstrom et al., 2015) hewitsoni-group hewitsoni, medontias, wakaensis, iccius, ephorus, sweadneri, sebetus, graueri, choveti, italus, zinebi, bergeri sciathis-group heathi, ivindo, uniformis, procora, subtilisurae, hyperanthus, amieti, feae, analis, sciathis, elishiae, sigiussidorum, makomensis (Brattstrom et al, 2016) taenias-group taenias milyas-group milyas, pavonis saussurei-group uzungwensis, danckelmani, simulacris, pareensis, dentata, neustetteri, albocincta, aurivillii, anisops, saussurei, suffusa safitza-group safitza, cottrelli

1 funebris-group funebris martius-group dekeyseri, dubia, persimilis, buea, martius, sanaos, matuta, golo, smithi, madetes, istaris, lamani, brunnea, sophrosyne, similis ena-group ena, campina, condamini, vansoni dorothea-group sandace, vulgaris, jefferyi, moyses, dorothea rhacotis-group technatis, nachtetis, rhacotis, mesogena, mesogenina, sambulos, sangmelinae angulosa-group anynana, auricruda, mandanes, kenia, collinsi, mollitia, sylvicolus, abnormis, angulosa, campa, tanzanicus, cooksoni, kiellandi

Relevant literature: Duplouy & Brattstrom, 2017 [Wolbachia in Bicyclus]. Aduse-Poku et al., 2016 [Phylogenetic hypothesis]. Brattstrom et al 2016 [Revision of Bicyclus species group] Brattstrom et al 2015 [Revision of Bicyclus species group] Hedenstrom et al., 2015 [Chemical analysis of pheromones in 17 Bicyclus species]. Brakefield & Joran, 2010 [Evolution of flexibility of wing patterns]. Oliver et al., 2009 [Natural and sexual selection in wing pattern evolution]. Arbesman et al., 2003 [Evolution of wing patterns].

evadne-group

*Bicyclus evadne (Cramer, [1779]) Small Stately Bush Brown

Papilio evadne Cramer, [1779] in Cramer, [1779-80]. Die Uitlandsche Kapellen voorkomende in de drie waerrelddeelen Asia, Africa en America 3: 71 (176 pp.). Amsteldam & Utrecht.

Type locality: . Diagnsos: There are two female morphs, one with, and another without a pale apical band (Aduse-Poku et al., 2016). Distributi