Taxonomy and Biostratigraphy of Ordovician Brachiopods from Northeastern Ny Friesland, Spitsbergen
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Zootaxa 3076: 1–122 (2011) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Monograph ZOOTAXA Copyright © 2011 · Magnolia Press ISSN 1175-5334 (online edition) ZOOTAXA 3076 Taxonomy and biostratigraphy of Ordovician brachiopods from northeastern Ny Friesland, Spitsbergen JESPER HANSEN1 & LARS E. HOLMER2 1 Department of Natural Sciences, Tromsø University Museum, NO-9037 Tromsø, Norway. Correspondence: Akvaplan-niva AS, Framsenteret, NO-9296 Tromsø, Norway. E-mail: [email protected] 2 Department of Earth Sciences, Palaeobiology, Uppsala University, Villavägen 16, SE-752 36 Uppsala, Sweden. E-mail: [email protected] Magnolia Press Auckland, New Zealand Accepted by D. Lee: 8 Aug. 2011; published: 28 Oct. 2011 JESPER HANSEN & LARS E. HOLMER Taxonomy and biostratigraphy of Ordovician brachiopods from northeastern Ny Friesland, Spitsbergen (Zootaxa 3076) 122 pp.; 30 cm. 28 Oct. 2011 ISBN 978-1-86977-789-0 (paperback) ISBN 978-1-86977-790-6 (Online edition) FIRST PUBLISHED IN 2011 BY Magnolia Press P.O. Box 41-383 Auckland 1346 New Zealand e-mail: [email protected] http://www.mapress.com/zootaxa/ © 2011 Magnolia Press All rights reserved. No part of this publication may be reproduced, stored, transmitted or disseminated, in any form, or by any means, without prior written permission from the publisher, to whom all requests to reproduce copyright material should be directed in writing. This authorization does not extend to any other kind of copying, by any means, in any form, and for any purpose other than private research use. ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition) 2 · Zootaxa 3076 © 2011 Magnolia Press HANSEN & HOLMER Table of contents Introduction . 6 Material and methods . 7 Geological setting and sampling area . 8 Lithology and depositional environment. 8 Chrono- and biostratigraphy . 11 Preservation and taphonomy . 13 Biogeography. 13 Palaeoecology . 13 Comments on the Great Ordovician Biodiversification Event . 15 Systematic palaeontology . 15 Phylum Brachiopoda Duméril, 1806 . 16 Subphylum Linguliformea Williams et al., 1996 . 16 Class Lingulata Gorjansky & Popov, 1985 . 16 Order Lingulida Waagen, 1885 . 16 Superfamily Linguloidea Menke, 1828 . 16 Family Obolidae King, 1846. 16 Subfamily Obolinae King, 1846. 16 Genus Lenticulella gen. nov.. 16 Lenticulella amphora (Krause & Rowell, 1975) . 16 Genus Palaeoglossa Cockerell, 1911 . 19 Palaeoglossa? sp. 19 Genus Ectenoglossa Sinclair, 1945 . 21 Ectenoglossa? oviforma sp. nov.. 21 Subfamily Glossellinae Cooper, 1956. 23 Genus Glossella Cooper, 1956 . 23 Glossella? sp. 23 Genus Pachyglossella Cooper, 1960. 24 Pachyglossella? sp.. 24 Subfamily Elliptoglossinae Popov & Holmer, 1994 . 24 Genus Elliptoglossa Cooper, 1956 . 24 Elliptoglossa vulgaris sp. nov. 24 Obolid sp. 1. 27 Obolid sp. 2. 28 Obolid sp. 3. 28 Obolid sp. 4. 30 Obolid sp. 5. 30 Family Zhanatellidae Koneva, 1986 . 31 Genus Rosobolus Havlíček, 1982 . 31 Rosobolus? elongatus sp. nov. 31 Zhanatellidae indet.. 33 Family Elkaniidae Walcott & Schuchert in Walcott, 1908 . 34 Genus Broeggeria Walcott, 1902 . 34 Broeggeria obscura sp. nov. ..