GROUND OF MACARONESIA, AN OVERVIEW (COLEOPTERA, CARABIDAE)

By ANTONIO MACHADO '

With I table

Our knowledge afthe Carabid fauna ofilie Macaronesian archipelagoes has increased considerably in the last few years, due to the conlributions ofseveral authors: GILLERFORS (1986), BORGES (1992), ISRAELSON (1990) and BORGES &. OROMI (199 1) for the Azores; SERRANO (1987), ERBER &. WHEATER (1988), ERBER &. H INTERSEHER (1988, 1990) and ERBER (1990) for Madeira; SERRANO (1983, 1987) and ERBER& HINTERSEHER (1987) for the Salvages; MACHADO (1992) with a revisionary work, for the Canaries; and FRANZ (1987), GElSrnARDT(1982, 1988) and MATEU (1990) for the Cape Verde. The present contribution is a compilation of the species rccorded in Ihe works metioned, ofTering a synoplic lisl of all Carabid beetles inhabiting Macaronesia, in its geographicaJ sense. No such global lisl has been published, if we exc\ude Ihe c\assic WOLLASTON'S ( 1865) "Coleoplera Atlanlidurn" where the Cape Verde was rnissing, and the individuallists incorporated in MACHADO (1992). Only the Cananan fauna can be eonsidered as revised; our knowledge ofthe Azorean and Cape Verdean is rather accurate, bUI Madeiran fauna requires a revision, in particular, sorne conflictive genera such as Trechus. The catalogue below has been cornpiled following the faunal lists inc\uded in Machado's rnonograph ( 1992), with sorne additions and correetions.

Additions: Cedrorum azoricus s. l. BORGES & SERRANO (in liU.) fo r Ihe Azores, and /inearis for Madcira, rccorded by ERBER ( 1990).

Corred ions: MACHADO (o.c.) recorded Acupalpus brunneipes for Madeira, bul it was not inc\uded io the faunal list (all Acupalpus species in Madeira should be Ihoroughly revised). He also mentioned Ocydromus obliquum for Azores, bul BORGES (in press) found Ihat Ihe species is in reality o. semipunclalum (Donovan).

Dcletions: Ocydromus siculumwas mentiooed for Madeira by JUNK ( 1938) probably

, IBCER, Urb. Aguen: 2, 38208 La Laguna, Tenerife

Bol. Mus. Mun. Funcbal, Supo nO4 : 395-410. 1995 ISSN 0870-3876 3.6 Boletim do Museu Municipal do Funchal (História Natural) Sup. ne4

based on the assumption ofNETOLlTZKY ( 19 18) that at/anlicwn could be the westem race oC siculum, and other authors have considered atlonJicum as a "Mischart". Based on MOll..ER­ MOTZFELD'S ( 1986) revision oC the siculum-complex, il is bener to remQve tbe species from Ihe Madeiran Iisl. Acupalpus flavico/lis (Sturm) listed by BORGES (1990) with no furth er reference has beco also removed from Ihe present catalogue, as well as severa! Canarian species which have ao uncertain status: Dyschirius uyltenboogaarti KLVNSTIlA J93 7. Bembidion inconspicuum WOLLASTON. 1864, S/enofophus /anzarolensis KL YNSTIlA 1937 and Syn/omus brevipennis (WOLLASTON, 1864).

CONCLUSIONS

The total number of carabids species present in Macaronesia is 380, 19 ofwhich are clearl y introduced ones. Using Ihese numbers one could deduct a native fauna of361 spocies. but the amount of introduced species is probably higher than statOO . A total of 28 species have developed ínter· or íntrainsular subspecies, which lotal 68. Therefore. if one adds species and subspecies as separale faunistic elements, the gross lotal is 420. Table I summarizes ¡he composition oflhe fauna of each archipelago.

T ABLE I . Faunistic elemcnts in the Carabidae ofMacaronesia

CONC EPT Amr ~s r.hd ~ irl SaIYa¡ts Ca .. arln Cape V~rde Mac.aro_la SP~CIESTOTAL 109 • 6/ JI' introdllCed species "11 9 , '"12 S ,. NA TIV~ S PECIES J 7 ,.0 I ]U J61 emic 1 , " , monoly¡m: specles ,. 201 po ytyplC speCles - , • . aeogr. races - - ftlCflronuian 6 moroot)'plc - - - po yt)'plC 5p«1e$ 2 onen emic , , _. European - , - a eartlc 3 , - , Iterrancan - , 16• , 41 - A tlcan , l 2 , '" A tlcan -- - 3 Sahelian - - 22 22 A ro-tropical - - - - 1 199' Proctedings oflhe M 1st Symposium Fauno &: Flora oft~ AllQ1Itic Islands- 397

The degree ofendemicity orlhe fauna is highesl in the eananes (70%), decreasing in Madei ra (63%), Sa lvagcs (50%), Azores (46%) and Cape Verde (32%). The Hule archipelago ofSelvagens has a poor carabid faunula, and could be considered as an appendix of either the Canarian or Madeiran fauna . There is a trend from North lo South in which the percenlage ofPalearctic and European elements decrease, while the importance ofNorth­ African and Ethiopian species ¡ncreases. The Mediterranean clernent is maximun in the intennediate archipelagoes: the Canaries and Madeira. Each archipelago has a rather "closed" fauna; Ihat ¡s, very few species inhabit two (8%) or more Ihan two (5%) archi pelagoes al one time. Most of Ihem are inlr'Oduced or widely spread common species (winged). However, there is also a small group of "Macaronesian" species (usually wilh archipelago-endemic vicariant subspecies): Aepw gracilicornis, Ocydromus schmidti. Paradromius imu/aris, angustw, etc. The carabid rauna of the Azores, Madeira and the Canaries can be regarded as a subset ofthe Wesl-Palearctic fauna. with a nOlorious entity and affinity between the endemic component of Madeira and the Canaries. The proportion ofCape Verdean paleoendemics which show relations 10 post common colonization with Ihe other archipelagoes is today ver)' smatl and the main charactcr oflhe rauna is clearly Ethiopian. Therefore, the geographica1 Macaronesia can be splil in t\Vo zoogeographical realms, the Palea.rctis and Ethiopis. The author expresses his gratitude to Dr. G. ERBER and P.A.V. BORGES for their commenls and collaboralion.

Ll ST OF THE CA RABID BEETLES OF MACARONES1A

Carabinae Caminaro ch/orastica "ugnllfa (CHAUOOIR. 1850) v Cape Verdean geographical mee Cuminara imbricata (KLUG. 1832) V Sahel-Sudanian Campolito olillier; ( D EJEAN. 1831) ACV No nh-Arrican Cam/mlito maderae madera/! (FAB Rl eIUS. 1775) MC West-Medilerranean Ctenosfa senegalensis (D EJEAN. 1831) V Sahel-S udanian Carubus abbrev;afus BR UL LÉ. 1835 C Canarian Carabusfauslusfausfus BR UL LE. 1838 e Canarian subspecies Curabusfauslus cabrera; ENDERLE1N. 1929 C Canarian subspecies Carabus coarctafus BRULLÉ. 1838 e eanarian

Nebriinae Ld.vfll.~· el/i,Jficus WOLLASTON. 1857 M Madciran Leisfus nubillagus WOLLASTON, J 864 C Canarian Nehria dilarara DEJEAN. 1836 e Canarian Nehr;u C:llrrax WOLLASTON. 1864 e Canarian 398 Boletim do Museu Municipal do Funchal (História Natural) Supon O4

NotiophjJinae No/iophi[us geminorus DEJEA/'l. 1831 Me West-Mediterranean NotiophiJus quadripunclatus DEJEAN, 1826 AM West-European

Lorocerinoe Elliptosoma wollastoni JAVET, 1852 M Madeiran

Sioganinae Siagona europaea DEJEAN, 1826 e Indo-Mediterranean

Scarüinae Scarites buparius (FORSTER. 177\) e West-Mediterrane.:ln Scarites abhreviafus abbrevio/lls DEJEAN, 1825 M Madeimn subspecies Scarites abbreviatus humeralis WOLl.. ASTON. 1854 M Madeiran subspecies &arites abbreviatus dmensis COCKERELL, 1923 M Madeiran subspecies Scarites abbreviatus deserlarum COCKERELL, 1923 M Madeiran subspecies Dyschir:ius numidicus PUTZEY S, 1846 e Saharian Dyschirius chalybeus PUTZEYS, 1846 e Mcditerrancan Dyschirius pouxillus WOLLASTON, 1864 e Nonhwest-African Dyschirius zanzibaricus C HAUDOIR, 1878 V Sahel-Sudanian Dyschirius ouriculatus WOLLASTON. 1867 ev Mediterranean Dyschirius samharicus PUrLEYS, 1873 ev Saharian-lndian

Apotominae Apofomus anguslicollis MOLLER, 1946 e North-African Apotomus chaudairii WOLlASTON, 1860 Me African Apotamus velox CHAUDOIR. 1876 V Sahclian

Broscinae Brascus crassimargo WOLlASTON. 1865 e Canarian Broscus rutilans WOLLASTON. 1862 e Canarian Brascus g/aber (BRULLÉ. 1838) e Canarian

Trechinae Canarobius chusyae M ACHADO. 1987 e Canari an Canarobius aramii MACHAOO, 1987 e Canarian Thalassaphilus Q20ricus OROM í & BORGES, 199 1 A Azorean Thalassaphilus caecus J EANNE l , 1938 M Madciran Thalassaphilus pieperi ERBER, 1990 M Madciran Tha/assaphilus whilei WOLLASTON, 1854 Me Macaronesian Thalassaphilus subterrallells MACHAOO, 1990 e Canarian Spelaeavlllcallia COllarien.\'i.\' MACIMOO. 1987 e Canarian I99S Procecdings oflhc -/st Symposium Fauna &, Flora O/Ihe Allan/ie /Slands-

Aepus graci!icornis grocificornis WOLLASTON, 1860 AM Macaronesian geographical race Aepus graci/icornis deserlarum MA TEU &. COLAS. 1959 M Madeiran geographical race Aepus gracilicornis meridionalis MACHADO. 1992 e Canarian geographical race Perileptus areola/Uf (CREUTZER. 1799 e Wesl·Palearctic Perileptus wollastoni JEANNEL. 1925 V Cape Verdean Perileptus hesperidum JEANNEL, 1925 V Sahelian? Trechus lorrelassol JEANNEL. 1937 A Azorean Trechus montanheirorum OROMi &. BORGES. 1991 A Azorean Truhus jorgensi$ OROMI &. BORGES. 1991 A Azorean Trechus terceiranus MACHADO, 1988 A Azorean Trechus picoensis MACHADO. 1988 A Azorean Trechus maderae FRANZ, 1981 M Madeiran Trechus alecrinensis FRANZ, 1981 M Madeiran Trechus lundbladi JEANNEl.. 1938 M Madeiran Trechusjlavomarginofus WOLLA5TON, 1854 M Madeiran Trechus debilis WOLLASTON. 1871 M Madeiran Trechus laevis WOLLA$TON, 1857 M Madeiran Trechus signo/us WOLLASTON, 1857 M Madeiran Trechus dilulus WOLLASTON, 1854 M Madeiran Trechus custos custos WOLl.ASTON, 1854 M Madciran subspecies Trecnus custos quadricollis WOLLASTON, 1854 M Madeiran subspecies Trechus alficola WOLLASTON, 1854 M Madeiran Trechus caufUS WOLLASTON, 1854 M Madeiran Trechus nigrocruciatus WOLLASTON, 1854 M Madeiran Trechus umbrico/a WOLLASTON, 1854 M Madeiran Trechus wollasfoni JEANNEL, 1922 M Madeiran Trechus minyops WOLLASTON, 1862 M Madciran Trechus lindbergi COLAS, 1963 M Madciran Trechus /aurisilvae FRANZ, 1981 M Madciran (bon.sp.?) Trechus ribeiranus FRANZ, 1981 M Madeiran (bon.sp.?) Trechus rabar;aleruis FRANZ. 1981 M Madeiran (bon.sp.?) Trechus ObfUSUS obtusus ERrCHSON, 1837 M Wcsl· Palcarctic pecolldiaml~' JEANNEL, 1952 M Madeiran gcographieal mce (?) Trechus gomeren.ris FRANZ, 1986 C Canarian Trechus laureticola JEANNEL, 1936 C Canarian Trechus cabrerai (JEANNEL, 1936) C Canarian Trechus machadoensis FRANZ, 1985 C Canarian Trechus uytlellOOogaar/i JEANNEL. 1936 C Canarian Trechllsfortuna/UJ' JEANNEL. 1927 C Canarian Trechus fenoeruis ISRLAELSON &; PALM, 1979 C Canarian Trechus fe/ix WOLLASTON, 1864 C Canarian Trechus antonU JEANNEL, 1936 C Canarian 400 BoJetim do Museu Municipal do Funchal (História Natural) SuponO 4

Trechus flavocircumdalus JEANNEL, 1922 e Canarian Trechus minioculalus MACHAOO, \987 e Canarian Trechus f1avocin clus jIavocinclus JEANNEL. 1922 e Canarian subspecics Trechus f1avocinclus gomerae JEANNEL, 1936 e Canarian subspecies Trechus delersus WOLLA STON, 1864 e Cananan Trechus jlavofimbatus WOLLA STON, 1863 e Canarian Trechus benahoar ilus M AC HADO, \ 990 e Cananan AnchOfrechus pune/llJennis JEANNEl. 1927 e Cananan Lymnaslis goudini goudini JEANNEL, 1929 e Canarian subspecics Lymntulis gaudini gomerae FRANZ, 1965 e Canarian subspecics Lymntutis subovolus MACHADO, 1992 e Canarian Lymnastis Ihoracicus MAC HADO. 1992 e Canarian Lymnasfis ni/orieus MOTSCHQUl.SKV, 1862 v North-Af'rican Tachys centromaculatus WOLLASTON, 1864 CV West-Mediterranean Tachys dimidiolUs MOTSCHOULS KY, 1849 e West-Mediterranean Tachysforlunatus MACHADO. 1989 e ¿Canarian? Tachys po/lidulus ANTOINE, 1943 e Northwesl-African Tachy.f micros F1SCHER. 1828 A European Tachys bisu!calus NICOLAJ. 1822 M Euro-Mediterranean Tachys bistriafus DUFTSCHMID. 181 2 M Mediterranean Tachys algiricus LUCAS. 1848 C West-Mediterranean Tachys impressipennis atomarills (WOLLASTON. 1867) V Cape Verdean geographica.l race Tachylophafeai (ALLUAUD. 1925) V Sahelo-Sudanian Tachyura II/casi DUVAL. 1852 MV Mediterranean Tachyura ~'ariabilis (CHAUOO IR. 1876) V Sahel-Sudanian Tachyura inaeqllalis ( KOLENATI. 1845) A Medilerranean Tachyura cllrvimana ( WOLLASTON. 1854) AMCV North-Af'rican-Sahelian TachyuTO haemorrhoida/is (PONZA. 1805) e West-Mediterranean Tachyla nana s. l. (GYLLENSVARD. 1810) e Palearctic (i) Asaphidion delatorrei UVlTENBOOGAART. 1928 e Canarian Orzolina thalassophila MA CHAOO. 1987 e Canarian Metallina ambigua (DEJEAN. 183 1) A West-Medilerranean (i) Bembidion varium (OUVIER. 1792) e Palearclic Bembidion mixtum (SCHAUM. 1863) v African Bembidion vicinllm (LUCAS. 1846) e Medilerranean Bembidion iricolor B EDEL. 1879 e At1antic-Mediterranean BembidiOIl bigllUatum FABRICIUS. 1779 M Euro-siberian Bembidion semipunclatllm (DoNOVA 'l. 1806) A West-Palearctic SynechOslictlls don¡!lI/lIm .d. DEJ EAN. 183 I M West-Euro-Medilerranean Phyla lethys (NETOUTZK y _ 1(26) M Mediterranean Ocydromlls la e lu.~ ( BRULLÉ. 1838) C Medilerranean Ucydromus crOlchii (WOLLASTON. 1864) C Canarian 199' Proceedings ofthe w 1st Symposium Fauno el Flora oflhe Allantie /s/ands- 401

Ocydromus fortunatus (WOLLASTON, 1871 ) e eanarian Ocydromus derelic!us (ALLUAUO. 1926) A Azorean (paIeoendemic) Ocydromus labellatus (WOLLASTON. 1854) M Madeiran Ocydromus hesperidum (WOLLASTON, 1867) V Cape Verdean Ocydromus schmidti azoricu$ LJNDRQTH, 1960 A Azorean geographlcal race Ocydromus schmidti schmidli (WOLLASTON. 1854) M Madeiran geographical cace Ocydromus schmidli subcallosus (WOLLASTON. 1864) e eananan geographical race Ocydromus schmidti ssp. V Cape Verdean geographical racc? Ocydromus a/lamiclIs (Jllan/iCIIS (WOLLA STON. 1854) Me Macaronesian geographical race Ocydromus andreae (FABR I C I U~, 1781) e Palearctic Ckys harpa/oides (SERVILLE. 182 1) AM West·Palearctic Pogonus chafceu$ MARSHAM. 1802 e South west·Palcarctic Pogonus gilllipes DEJEAN. 1828 ev Atlanto-Mediterranean Syrdenus grayi (WOL.LASTON. 1862) ev West-Mediterranean

Puigoninae nigriceps DEJ EAN. 183 1 AMCV OrientaUcosmopolitan (i)

Pleroslicllínae Olis/hopus ericae W.OllASTON. 1854 M Madeiran Olis/hopus madeirensis WOLlASTON. 1854 M Madeiran Olis/hopus humerosus WOLLASTON. 1858 M Madeiran Olis/hopus glabra/lis glabra/lis BRULL~. 1838 e Canarian subspecies Olis/hoplls glabra/lIS /amaranus MACHAOO. 1992 e Canarian subspecies Olislhopus palmensis WOLLASTON. 1864 e Canarian Olislhopus elongallls WOl.LASTON. 1854 Me WeSI Medilerranean Olislhopus inclavatus ISR l AE l SON. 1983 A Azorean Agonum marginatum (LlNN AF.US. 1758) AMe West- Palearclic (i) Agonum muelleri HERBST. 1784 A Palearctic (i) Pseudanchomenus aptinoides TARNIER. 1860 A Azorean (paleoendemic) Paranchus nichollsii (WOLLA STON. 1864) e Canarian Paranchus debtlis (WOlLASTON. 1864) e -<::anarian Paranchus ruficornis GoETZE. 1777 AMe Palearctic (i) Abuce/us nalalensis CHAUDOIR, 1869 V Afro-tropical Pediusfiguralus (WOLLASTON. 1864) e Canari ... Angoleus crenalus freyanus (UYTIENOOOGAART. 1940) C Northwcst-Afiican Angoleus wollastoni (WOLLASTON. 1854) MeV North-African Lagarus vernalis s.l. (PAN ZER. 1796) A Palearctic (1) Oma~·ell.f aterrimus nigerrimll.~ DEJEAN, 1828 AM SOUlwcst-Palearctic Or/hamus berylensü' (REICHE &. SOULCY. 1854) MSe Mediterranean Or/homus discors (WOLLASTON. 1864) e Canarian Cedrorum azoricus azoricus BORGES &. SERRANO i.p A Azorean subspecies 402 Boletim do Museu Municipal do Funchal (Hist6ria NalU",l) Supo ne 4

Cedrorum azor. cl1l'eirensis BORGES &. SERRANO i.p A Azorean subspecies Nesorthomw berraj BATTONI, 1987 M Madeiran Nesorthom us robuslus (WOl.LASTON, 1854) M Madeiran Nesorlhomus dilaticollis (WOlLASTON, 1854) M Madeiran Nesorlhomus curlUS (WOLLASTON . 1854) M Madeiran Nesorthomus gracilipes (WOLLASTON. 1854) M Madeiran Nesorlhomus pecoudi JEANNEL, 1942 M Madeiran Nesorthomus lundbladi JEANNEL, 1938 M Madeiran EUlrichopus canariensis (BRULL~. 1838) e Canarian Eulrichopus gonzalezi MI. TEU. 1954 e Canarian Walltinerfia Qnllgae MEDINA &. OROMI, 1991 e eanarian Wolltinerfio tenerifae (MACHA OO. 1984) e Canarian Wolllinerjia marti,,; (MACHADO. 1984) e Canarian Pseudomyas doramasensis UVTfENBOOGAART, 1929 e Canarian Gomerina calathiformis (WOLLASTON. 1865) e Canarian Gomerina nitidicollis (LlNDBERG. 1953) e Canarian Pseudop/atyderus amb/YOP!i BOLIVAR. 1940 e Canarian Paraeutrichopu:; harpa/oides (WOLLASTON. 1864) e Canarian Paraeu/riehopu:; peeoudi MA TEU. 1954 e Canarian Platyderus altieo/a alrieola (WOLLASTON. 1864) e Canarian subspedes Platyderus al/ieola hierroensis MACHADO. 1992 e Canarian subspecies Platyderus alricola gamerensi.r MACHADO. 1992 e Canarian subspecies Platyderw ailicola deseendens BEDEL. 1902 e Canarian subspecies Plotyderus lancerollensis ISRAELSON. 1990 e Canarian Platyderw gregarius REICHE. 1861 e Nonhwest-A rrican Amaroschema gaudini JEANNEL. 1943 e Canarian Ca/o/hu:; complana/w complana/u.~ DEJEAN. 1828 M Madeiran subspccics Calarhu:; comp/anaws vandeli COLAS &. MA TEU. 1959 M Madeiran subspccies Cala/hu:; fimbria/u:; WOLLASTON. 1858 M Madeiran Cala/hu:; eo/asianus MATEU. 1970 M Madeiran Cala/hu:; vividus F ABRICIUS. 180 I M Madeiran Cala/hus subfoscus WOLLA$TON. 1865 M Madeiran Cala/hu:; refluimargo MACHADO, 1992 e Canarian Colathw obli/era/us WOLLASTON. 1865 e Canarian Cala/hus pilosipennis MACHADO. 1992 e Canarian Calathus spretw WOLLASTON, 1862 e Canarion Cala/hus gomerensis COLAS. 1943 e Canarian Calo/hus eogna/u:; WOLLASTON. 1864 e Canarian Cala/hu:; marcellae COLAS. 1943 e Canarian Calathus /aure/ieo/a WOLLASTON , 1865 e Canarian Calathus appendieula/us WOLLASTON. 1862 e Canarian Calathus angularis BR ULLE. 1838 e Canarian 199' Proceedings Of lhe M 1st Symposium Fauna & Flora ofthe Allantic Island$" 403

Calathus canariensis HAROl.DS. 1868 e eanarian Ca/ath us rufocastaneus WOLLASTON, 1862 e Canarian Calalhus ciliotus WOl.. l.ASTON, 1862 e Canarian Calathus Que/us WOLl.ASTON, 1862 e eananan Colathus cariTUltus BRULLÉ, 1838 e Canarian Calalhus angustulw WOlLASTON. 1862 e eananan Calathus amplius EscALERA, 1921 e Canarian Calathus depressus BRULLÉ, 1838 e Canarian Cala/hus ¡rey; COLAS. 194 1 e Canarian Cala/hus asceruJens WOLLASTON. 1862 e Canarían Calathus rectus WOLLA STON. 1862 e eanarian Calathus abaxoides BRULLÉ, 1838 e Canarian Calarhus goltUllezi MA TEU, 1956 e Canarian eofalhus simplicicollis WQU.ASTON, 1862 e Canarian Colathus extensicollis PUTZEYS, 1863 A Azorean Colo/hus /undbladi COLA S. 1938 A Azorean Calathus vicenteorum SCHATZMAYR, 1937 A Azorean Calalhus corva/hai SERRANO &: BORGES. 1986 A Arorean Calathidius brevithorax MA CHADO. 1992 e Canarian Calathidius acuminalus (WOU.ASTON. 1862) e Canarian Calathidius sphadroides (WOLLASTON, 1862) e Canarian Licinopsis allernans (OEJEAN. 1828) e Canarian Licinopsis obliterara obliterara (WOLLASTON, 1865) e Canarian subspecies Licinopsis obliterotafranzi MA CHADO, 1987 e Canarian subspecies Licinopsis gaudini JEANNEL. 1937 e Canarian Licinopsis anguslula MACHADO. 1987 e Canarian Licinopsis picescens picescens (WOLLASTON. 1864) e Canarian subspecies Licinopsis picescens gomeri/a MACHADO. 1987 e Canarian subpsecies Licinopsis schurmanni MA CliADO. 1987 e Canarian Sphadrus leucophlhalmu$ (LI NNAEUS. 1758) MSe Indo-European (i) Laemoslenus complanalus (OEJEAN. 1828) AMe NOr1h-A frican (i) Amara aenea (OEGEER. 1774) AMCV Palearctic (i) Amara superans WOLLASTON. 1854 M Madeiran Celia cOltyi (COQUEREL. 1859) MSe No rthwesl-African Amalhilis rufescens (OEJEAN. 1829) e WeSI -Medilerranean Zabrus crassus OEJ EAN. 1828 e Canarian Zabrus laevigatus ZIMMERMANN, 183 1 e Canarian Zabrus canaricus MACHADO. 1992 e Canarian

Hllrpa/inae Crasadactylus punClalus GUé RI N. 1847 e African Anisodaclylus binO/aIUS (F AI1RI CIUS. 1787) AM Palearcti c (i) '04 Boletim do Museu Municipal do Funehal (Hi stóri. NlI.lUraI) Supo ne 4

Notiobia cupripennis (GERMAR. 1824) e Neotropical (i) Odonlocarus cordatus (DEJEAN. 1825) e West-Meditenanean Ditomus opacus (ERICHSON. 1841) e North-Afiican Daptus viua/us labialus MOTSCHOULSKY. 1849 e West-Meditenanean Groniger femoralis (COQUEREL. 1858) e Mediterranean Nesarpalus gregarius gregarius F AUVEL, 1897 M Madeiran subspccies Nesarpalus gregarius al/anlieus SCHAUBERBERG, 1932 M Madeiran subspccies Nesarpalus vividus (DEJEAN. 1829) M Madeiran Nesarpalus pelagicus (WOLLASTON, 1860) S Selvagens Nesarpalus solitarius (WOLLASTON. 1863) e Canarian Nesarpalus !ortunatus (WOLLASTON, 1863) e Canarian Nesarpalus uyltenboogaarti (VANEMOEN. 1929) e Canarian Nesarpa/us micans (WOLLASTON. 1864) e Canarian Nesarpalus empiricus (WOLLASTON, 1985) e Canarian Nesarpalus sanctaecrucis (WOLLASTON. 1864) e eanarian Platymetopus tesse/atus Dejean, 1829 V Sahe1ian Nesiopelus serienotatus (WOl.L.ASTON, 1867) V Cape Verdean Aulacoryssus acicu/atus (DEJEAN. 1829) V Sahe1-Sudanian Cryptophonusjaninae (JEANNE. 1984) e Canarian Cryptophonus schaumii (WOLLASTON, 1864) e Canarian Cryptophonus paivanus paivanus (WOLLASTON. 1867) V Cape Verdean subspecies Cryptophonus paivallus meirai (MATEU. 1958) V Cape Verdean subspecies Cryptophonus tenebrosus (DEJEAN. 1829) Me EIII1)-Med.iterranean Cryptophonusfolvus (DEJEAN. 1829) e Meditemutean Ophonus ardosiacus (lUTSCHNIK. 1922) AMS West·Palearetic (7) Ophonus stictus (STEPHENS. 1828) AM West-Palearctic Pseudophonus rufipes (DEGEER. 1774) AM West·Palearctic (i) Pseudophonus griseus (PANZER. 1797) A Palaeartic Harpa/us attenuatus STEPHENS. 1828 M Atlanlo-Med.ilemutean (7) Harpa/us distinguendus (DUFTSCHMID. 1812) AMe Palaeartic (i) Nesarpax labiatus (ERICHSON. 1843) V Cape Verdean Bradybaenus scalaris (OLlVIER. 1808) V Sahel·Sudanian Dicheirotrichus obso/etus (DEJEAN. 1829) e Atlanlo·Medilerranean Dicheirotrichus punicus BEDEL. 1899 e Mediterranean Bradycellus ventrlcosus WOL LASTON. 1864 e eanarian Bradycelllls excultus WOLLASTON. 1854 M Madeiran Bradycellus mudeirensi.\" MATEU. 1958 M Madeiran Bradycelllls fe/ol MA CIIAOO. 1992 e Canarian 8radycellus harpalinu.f SERVILLE. 1821 M European Bradyt:ellus chave.fi ALLUAUD. 19 19 A Aro""" Brudycellw' distinctus (DEJ EAN. 1867) A West·Palearetic (7) Acupalpus hrunneipe.f (STURM, 1825) AM Wesl-Palearctic 199' Proceedings ofthe ~ Isl Symposium Founo & Floro of/he AlIOlI/k /slands- ..,

Acupalpu.~ dubius SCHILSKY. 1888 AM European Acupalpus notatus MULSANT &. REY, 1861 MC Mediterranean Acupalpus maculatus SCHAUM, 1860 e Euro-Meditemmean Egoploa crenuloto (DEJEAN. 1829) V Sahel·Sudanian Egadroma marginolo (DEJEAN. 1829) Me Mediterranean Egodromo reluscens (ERICHSON. 1845) V Sahel-Sudanian S/enolophus ¡eutontls (SCHRANK. 1781 ) AMe WeSI-Palaeartic Anomoslomus torridus LAFERrE. 1853 V Sahelian Amblystomus orpheus (LAFERTE. 1853) V Sahel-Sudanian Amblyslomus viridulu$ (ERICHSON. 1843) V Sahelian

Udninae Zilrgus schoumii WOLLASTON, 1854 M Madeiran ZarKus deSf!rtae WOLLASTON, 1854 M Madeiran Z(lrgus pellucidus WOI..I.. ASTON, 1854 M Madeiran Zargus monizii WOLLASTON. J 860 M Madeiran Zorgus cralchianus WOLLASTON. 1865 e eananan Eurygnolhus la/reille; la/reille; (LAPORTE. 1834) M Madeiran subspecies Eurygno/hu$ lalreille; 1I'0IlU$IOn; COCKERELL. 1923 M Madeiran subspecies Licim/$ munr;qu;on"s WOLLASTON, 1862 C Canarian Licim/s pune/alulus pllneflt/ullls (F ABRIC IUS, 1792) A West-Palearetic (i) Licinll$ pune/olulus gronulufIIs DEJEAN, 1926 e North-African (i)

Pentngonicin oe Penll1goniea hexogona (WOLLASTON, 1867) V Cape Verdean

Cnllistinoe Pachydinodes conformis UncosiglllllUS WOLLASTON,I867 V Cape Verdean geographicaJ race Slenodinodes baisdllvoli (DEJEAN. 1831) V Sahelo-Sudanian Slenodinades as.~ec/a (LAFERTE. 1851) V Sahelian Epumi.\· drcumscripllls (DUFTSCHM ID. 1812) C Afro-Palearctic Chluenius ~poliatus (ROSSI. 1790) C Mediterranean Chlaenius cunarierlS;s eanar;erlS;s DEJEAN. 183 1 C North-African e hlaenius laelicollis CHAUOOIR. 1876 V Sahel-Saharian

/lfosoreinoe Mal"arel/S ullicola WOLLASTON. 1864 C Canarian Masureul' a.Dinis urenieolu WOL LASTON. 1863 CS Macaronesian geographicaJ mee orienluU.f Sl·p. M Madeiran geograpbicaJ mee? Masoreus orienloli.f nohili.~ WOLLASTON. 1864 C Canarian geograpbicaJ mee Mal'urew' ar;enlali.\' aseemJen.\' WOLLASTON. 1867 V Cape Verdean geograpbicaJ race Aephinidius madagllScar;ensi,\' (CHAUOOIR. 1850) V African 406 Boletim do Museu Municipal do Funchal (Hisl6ria Natural) Supon O4

übünae elevatus (FABRICIUS, 1792) v Cosmopolitan (i) Plarytarus lamin; (DEJEAN, 1826) ev Mediterranean Plarytarus tesselatus (DEJEAN, 1826) V Ethiopian Platytarus gracilis occidenlalis ANTOINE, 1933 e Nonhwest-African Deme/rías o/ricapillus (lINNAEUS. 1758) e West-palearclic (i) maculata KLUGER. 1832 V Medilerranean Plocionus pallens (FABRI CIUS. 1775) V Cosmopolitan (i) zargoides (WOLLASTON. 1863) e Canarian Cymindis veloto (WOLl.ASTON, 1865) e Canarian Cymindis simillima (WOLLASTON, 1865) e Canarian Cymindis amicla (WOLLASTON, 1864) e Canarian Cymindis cinclo BRULLÉ, 1838 e Canarian Cymindis anchomenoides (WOLLASTON, 1867) V Cape Verdean Cymindis alutaceus (WOLLASTON. 1867) V Cape Verdean Cymindis dohrnii (WOLLASTON. 1867) V Cape Verdean Cymindis lindbergi MA TEU, 1964 V Cape Verdean Cymimiis marginella BRULLE, 1838 e Canarian Cymindis maderae (WOLLASTON, 1857) M Madeiran Cymindis paivana (WOLl.. ASTON, 1860) s Salvages Cymindis moralesi MATEU, 1979 e Canarian Cymindis suturalis pseudosu/uralis BEDEL, 1906 Mes Nonhwest-African Cymindis discophora CHAUDOIR, 187) e Nonhwest-African Paradromius iucundus iucundus MACHADO, 1992 e Canarian subspecies Paradromius iucundus gillerforsi MACHADO, 1992 e Canarian subspecies Paradromius amoenus (WOLLASTON, 1863) e Canarian Paradromius tamaranu,\' MACHADO, 1992 e Canarian Paradromius amplius MA CHADO , 1992 e Canarian Paradromius exornatu.~ exornatus MA CHADO, 1992 e Canarian subspecies Paradromius exorna/us furvu~' MACHADO, 1992 C Canarian subspecies Paradromius purpurariu~' MA CHADO. 1992 C Canarian Paradromius slrigifrons s/rigifrons (WOLLASTON, 1865)C Canarian subspecies Paradromius strigifrons linerfen.\ü (MA TEU, 1961) C Canarian subspecies Paradromius strig. grancanariem'Ú' MACI-lA DO, 1992 C Canarian subspecies Paradromius linearis (OLIV IER. 1795) MC Euro-Mcditcrrancan Paradromius hariensis MACHA DO, 1992 C Canarian Paradromiw¡ insularis in sulari,~ (WOLL ASTON. 1854) M Madeiran subspecies Paradromius insularis oceanicu.\' (WOLLASTON. 1865) MS Macaronesian subspecies Paradromius insulari:. il/au/us MACIIADO. 1992 C Canarian geogra;>hical race Paraaromiusscholzi MACHADO, 1992 C Canarian Dromius meridiona/is DEJEAN. 1825 A West-Palcarctic (?) Dromius angw/us alutaceu~' WOLLASTON. 1857 M Madciran geographi cal race 1995 Proeeedings oflhe ~ ¡st Symposillm Forma el Flora O/lIJe Allanfic Islonds- 407

Dromius ongustus plagipennis WOLLASTON, 1865 e Canarian geographical race Dromius angustus dissimilis MACHADO. 1992 e Canarian geographicaJ mee Dromius brittoni brittoni MATEU. 1957 e Canarian geographical race Dromius geuthardti MATEU. 1990 V Cape Verdean Philorhizusferranius MATEU, 1956 e eanarian Phi/orhirus bravoorum MATEU, 1957 e eanarian Phi/ornizus Iindbergi MATEU. 1956 e Canarian Phi/orllirus elliptipennis (WOLLASTON. 1864) e Canarian Phi/orh/zus longicollis (WOLLASTON, 1865) e eanarian Phi/orhizus moteu; MACHADO. 1992 e Canarian Phi/orllizus umb,atus (WOLLASTON, 1865) M Madeiran Phi/Drhjzus wollastoni wollQJlon; (FAUVEL , 1905) M Madeiran subspecies Phi/orhizus wollastoni nilidus MATEU, 1957 M Madeimn subspecies Phi/orhizus conicipennis (FAUVE L, 1905) M Madeiran Phi/orhi:us vieira; MATEU. 1957 M Madeiran allanlicus OIlamicus MA TEU. 1957 e Cananan subspecies Philorhizus allanlicus forlunolUS MA TEU. 1957 e Canarian subspecies Philorhizus ailamicus palmensis MACHADO, 1992 e Cananan subspecies Philorhizus fumalus MA TEU. 1961 e Cananan Philorhizus parvicollis (WOLLASTON, 1865) e Canarian Philorhizus franzi MACHADO. 1992 e Cananan Philorhizus incerlus incerlus (WOl.l.ASTON , 1864) e Canarian subspecies Philorhizus incerlus jandiensis MACHADO, 1992 e Cananan subspecies Philorhizus insignis LUCAS. 1846 e Nonh-Afncan Philorhizus beidensis'beidensis ANT01NE. 1940 e Nonhwest-Afncan Philorhizus melanouphalus (OEJEAN. 1825) A Wesl-Palearclic Philorhizus notatus (STEPHENS. 1827) A Nonh-Pa1earctic Philorhizus attenuatus (WOLLASTON. 1867) V Cape Verdean Philorhlzus sigma (ROSSI. 1790) V European (i?) Metadromius pervenustus (WOLLASTON. 1864) e Cananan Metodromius palmi MACHADO. 1992 e Canarian S)lmomus lundbladi JEANNEL. 1938 M Madeiran S)lntomus inaequalis (WOLLASTON, 1863) e Cananan lancerolensis (WOLLASTON. 1864) e Cananan Syntomus foscomaculatus (MOTSCHOULSKY, 1845) Me South-Palearclic Syntomus laleralis schrammi ANTOINE, 1940 e Nonh-African Syntomus grayi (WOLLASTON. 1867) V Cape Verdean Syntomus submaculatus (WOLLASTON. 1867) V Sahe1ian Microlesles /UClUOSUS chobauli JEANNEL. 1942 MC West-Meditcrranean Micro/esles negrila WOLLASTON. 1854 AM Medilenanean MicroüsltS ongusteforcipatus ANTOINE. 1940 C Wesl-Mediterranean Micro/esles corücalis DUFOUR. 1820 Me Mediterranean 408 Boletim do Museu Municipal do Funchal (Hist6ria Natural) Supo nO"

Micro/estes abeillei brisau/; HOLDHAUS, 1912 e North·African Micro/esles gomerensis LlNOBERG, 1953 e Saharian Micro/esles discoidalis (F A1RMAlRE, 1892) V Sahclian Aprislus holo/euros LINOBERG. 1950 e eanan", Zolotarewskiella slrigicollis (WOLLASTON, 1867) v Sahcl-Sudanian

Zuplri;nae Dicrodontu$ brunneu$ brunneus (DESEAN. 1831) e Canarian subspc:cies Dicrodontus brunneus exilis MACHADO. 1992 e CANA RlAN SUBS PECIES Dicrodontus aplinoides (WOLLASTON. 1865) e CANARlAN Dicrodontus olluoudi MATEU. 1952 e Canarian Parazuphium baeticum maure/anioe HÚRKA, 1982 M North-African Parazuphium blandum MA TEU. 1990 v Ethiopian Parazuphium debi/e harten; MATEU, 1990 V Ehtiopian Porazuphium damascenum canariense MACHADO, 1992 e eanarian geographical race

Braclrininae Pheropsophus hispanicus (DEJEAN. 1823) e Northwest-Amcan (i) 1995 Proceedings ofthe ~ /11 Symposi ~ m Fauna el Flora O/1M A/lantic Jsland..r ~ 409

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WO LLASTON, T. V.: 1865. Coleoplera A/lanlidum. being an enumeralion oflhe Coleoplerous insecls ofthe Madeiras. Sa/vages and Canaries. London: Taylor& Fraocis. 47 + 526 pp. Appcndix 140 pp.