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An annotated checklist of Wisconsin Mordellidae (Coleoptera)

Anneke E. Lisberg University of Wisconsin-Madison

Daniel K. Young University of Wisconsin-Madison

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Lisberg, Anneke E. and Young, Daniel K., "An annotated checklist of Wisconsin Mordellidae (Coleoptera)" (2003). Insecta Mundi. 39. https://digitalcommons.unl.edu/insectamundi/39

This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 195

An annotated checklist of Wisconsin Mordellidae (Coleoptera)

Anneke E. Lisberg and Daniel K. Young Department of Entomology University of Wisconsin-Madison 445 Russell Labs 1630 Linden Dr. Madison, WI 53706, U.S.A.

Abstract: A three-year survey of Wisconsin Mordellidae (Coleoptera) encompassing a compilation of data from literature records and local collections as well as field work including trapping, hand-collecting, and rearing yielded 68 species comprising 14 genera in three tribes. Sixty-three species (92% of Wisconsin fauna) represent new state species records, not previously recorded from the state in the literature. Plant-associations and state- specific temporal and spatial distribution data for larvae and adults are noted as available. Distributional records suggest 16 additional species and one additional genus are likely to occur in Wisconsin.

Introduction Methods

The family Mordellidae is a widely distributed, A general, statewide survey of Wisconsin Mordel- speciose, and commonly encountered beetle taxon. lidae was conducted from literature searches, exam- There are approximately 1500 species worldwide com- ination of previously collected material [e.g. private prising 113 genera. In North America north of Mexico collections, Wisconsin Department of Natural Re- there are 17 genera comprising 201 species (Jackman sources (WI-DNR) specimens and the University of and Lu 2002, Lisberg 2004). Despite the relative Wisconsin-Madison Insect Research collection (WIRC)] abundance of Mordellidae, natural history and distri- as well as field sampling (particularly in historically bution data for North American species are scant. undersampled regions of the state.) Sampling meth- Adult mordellids are phytophagous and feed from the ods included the use of passive traps, particularly flowers of many taxonomically diverse plants (Robert- flight-intercept traps, hand collecting from flowers son 1928, Lu 1997, Jackman and Lu 2002, present and foliage, and a small-scale rearing project from study), however, species-specific records of adult prairie floral stem cuttings. Passive traps were used mordellid food plants are rare. Larvae are stem and to maximize the overall number as well as geograph- wood-borers whose food consists largely of pithy plant ical and taxonomic breadth of species sampled, where- stems and rotten wood (Lawrence 1991, Ford and as hand collecting and rearing provided more in-depth Jackman 1996). Larvae of Mordellini are usually natural history data. Field work from other ongoing collected from woody plants whereas larvae of Mordel- studies [e.g. from the WI-DNR and other insect listenini are more common in stems of pithy herba- inventory projects] also contributed significant data ceous plants (Jackman and Lu 2002), but species- to this study. specific plant-larval associations are also rare. Distri- butional data for North American Mordellidae point Results out many gaps. Published Wisconsin records of Mordel- lidae are particularly sparse, historically accounting This survey yielded 68 Wisconsin species com- for merely five species in three genera: Tomoxia prising 24 genera. A permanent voucher collection of lineella LeConte (Lisberg and Young 2003), Mordella Wisconsin Mordellidae was developed and is housed in atrata Melsheimer, Mordella melaena Germar, the WIRC. Data for these specimens (species, distri- Mordella quadripunctata (Say), and Hoshihanano- bution, and natural history data) were compiled and mia octopunctata (Fabricius) (Liljeblad 1945). By recorded in an established WIRC database which comparison, neighboring Illinois boasts 75 species in utilizes BIOTA software (Colwell 1996). 13 genera (Bright 1986, Jackman 1991, Jackman and The following checklist of Wisconsin Mordellidae Lu 2001, 2002). No statewide surveys for Mordellidae includes 63 new state species records, indicated have previously been conducted in the United States. in boldface. To aid interpretation of county records for each species listed, Wisconsin has been divided into nine regions (Map 1, after Kriska and Young 2002 196 Volume 17, No. 3-4, September-December, 2003, INSECTA MUNDI

and Hilsenhoff 1995). Plant associations taken from Mordellaria undulata (Melsheimer) – Recov- Wisconsin specimen labels in pre-existing collections ered from flight intercept and malaise traps in oak are listed using the plant names in the exact form, barrens. Late June to late September. NC: Oneida; common name or scientific name, given in the origi- NE: Shawano; WC: Jackson; EC: Fond du Lac; SW: nal source. Although we concur that it is preferable to Grant; SE: Waukesha. consistently provide scientific rather than common names, the variability and occasional duplicity of Mordella albosuturalis Liljeblad – Collected from common names makes their arbitrary replacement a flowers of Daucus carota Linnaeus. Late June to mid- potential source of introducing inaccurate data. Many July. SC: Dane, Green. Wisconsin species still lack detailed natural history information. To reduce redundance of reporting local- Mordella atrata Melsheimer – Hand collected from ity and temporal distributions, all records reported flowers of Amorpha canescens Pursh, Anemone ca- below should be assumed to pertain specifically to nadensis Linnaeus, Coreopsis lanceolata Linnaeus, adults unless otherwise noted. Cornus racemosa Lamarck, Erigeron anuus Linnae- us, Eupatorium perfoliatum Linnaeus, Opuntia com- Tribe Conaliini pressa Salisbury, Rosa carolina Linnaeus, Spiraea alba DuRoi, Trifolium repens Linnaeus, and white Conalia helva (LeConte) – Late July. C: Wood. clover. Also recorded from barrier pitfall, flight inter- cept, malaise, and yellow pan traps and sweep nets in Tribe Mordellini prairies, prairie remnants, and occasionally oak sa- vannas, sand barrens and oak barrens. Early June to Tomoxia inclusa LeConte – Collected at flight late September. NW: Bayfield, Burnett; NC: Oneida; intercept traps, malaise traps, and sweep nets in oak NE: Oconto, Shawano; WC: Jackson, Pierce; C: Ju- savannas and sandy oak barrens. Mid-June to mid- neau; EC: Sheboygan; SW: Grant, Richland; SC: August. NW: Bayfield; WC: Jackson, Monroe; SW: Columbia, Dane, Green, Iowa, Sauk; SE: Walworth. Grant; SC: Sauk; SE: Waukesha. Mordella lunulata Helmuth – Hand collected Tomoxia lineella LeConte – Larvae collected from a from rose and recovered from a yellow pan trap in big standing, dead, large-toothed aspen (Populus grandi- bluestem (prairie). Early June to late July. SC: Dane; dentata Michaux) (Lisberg and Young 2003). Adults SE: Jefferson. collected using flight intercept, malaise, and Lindgren funnel traps in southern mesic hardwood, mesic oak, Mordella marginata Melsheimer – Collected from and mixed forests and prairie-forest ecotones. Late flowers of Amorpha canescens Pursh, Anemone ca- June to early August. NE: Oconto; WC: Monroe; EC: nadensis Linnaeus, Berteroa incana Linnaeus, Cor- Fond du Lac; SC: Green, Lafayette, Sauk. nus racemosa Lamarck, Daucus carota Linnaeus, Eupatorium maculatum Linnaeus, Euphorbia corro- Mordellaria borealis (LeConte) – Early July. lata Linnaeus, Parthenium integrifolium Linnaeus, NW: Bayfield. Physocarpus opulifolius Linnaeus, Rhus glabra Lin- naeus, Sambucus canadensis Linnaeus, Spiraea alba Mordellaria serval (Say) – Hand collected from Du Roi, Trifolium pratense Linnaeus, Viburnum daisy fleabane and elderberry flowers, and collected opulus Linnaeus, orange milkweed, daisy, daisy flea- using flight intercept, malaise, and Lindgren funnel bane, and wild rose, and one specimen was found on traps in northern and southern mesic hardwood a Cirsium altissimum Linnaeus stem. Also sampled forests, northern dry-mesic forests, oak savannas, by using sweep nets and flight intercept, malaise, and oak barrens, an aspen grove, a cut Pinus banksiana yellow pan traps. Late May through early September. Lambert, and oak, oak-basswood, oak-pine, and pine- NW: Bayfield; NC: Oneida, Vilas; NE: Oconto, Sha- poplar forests. Mid-June to mid-September. NC: Onei- wano; WC: Jackson; C: Marquette, Waushara, Wood; da; NE: Oconto; WC: Jackson, Monroe; C: Portage, EC: Fond du Lac, Sheboygan; SW: Crawford, Grant, Waupaca, Wood; EC: Fond du Lac; SW: Grant; SC: LaCrosse, Richland; SC: Columbia, Dane, Green, Dane, Green, Iowa, Lafayette; SE: Ozaukee, Racine, Iowa, Lafayette, Rock, Sauk; SE: Ozaukee, Wal- Waukesha. worth, Waukesha. INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 197

Map 1. Regions of Wisconsin (after Kriska and Young 2002, and Hilsenhoff 1995). Mordella melaena Germar – Recorded from flowers of Mordella oculata Say – Collected from forests, mid- Erigeron anuus Linnaeus, Eupatorium perfoliatum July to early August. SC: Dane, Green; SE: Washing- Linnaeus, rose, and Queen Anne’s lace. Also recov- ton. ered from barrier pitfall, flight intercept, and malaise traps from oak savannas, sandy oak barrens, and Mordella quadripunctata (Say) –From flowers of prairies. Early June through late August. NW: Bay- Cicuta maculata Linnaeus and Monarda sp. One field; NC: Iron; NE: Shawano; WC: Monroe, Clark; specimen was collected in a malaise trap. Late June SW: Grant; SC: Columbia, Dane, Lafayette, Sauk; to late July. NC: Oneida; WC: Clark; SW: Grant; SC: SE: Jefferson, Washington, Waukesha. Dane.

Hoshihananomia octopunctata (Fabricius) – At flow- ers of Daucus carota Linnaeus, Monarda sp., and 198 Volume 17, No. 3-4, September-December, 2003, INSECTA MUNDI

Ceanothus sp. Also collected by hand, using sweep Erigeron sp. Adults collected from flowers of Achillea nets, and in flight intercept, malaise, and yellow pan millefolium Linnaeus, Cornus racemosa Lamarck, traps in oak savannas, oak barrens, and prairie-forest Erigeron sp., black-eyed Susan, daisy fleabane, and ecotones. Late May to late August. NW: Bayfield; WC: goldenrod, as well as from box-elder and daisy. Also Monroe; C: Marquette; EC: Fond du Lac; SW: Grant, collected using sweep nets, flight intercept traps, and Richland, Trempealeau; SC: Columbia, Dane, Green, malaise traps from prairies, oak savannas, and prai- Iowa, Lafayette, Rock, Sauk; SE: Washington, Wauke- rie-forest ecotones. NW: Bayfield; NC: Oneida, Vilas; sha. NE: Oconto; WC: Monroe; C: Marquette; SW: Craw- ford, Grant, Richland; SC: Dane, Green, Iowa, Rock, Tribe Mordellistenini Sauk; SE: Jefferson, Waukesha.

Mordellochroa scapularis (Say) – At flowers of Mordellistena comata (LeConte) – A single spec- Heracleum maximum Bartram, Prunus sp., and imen from late June. SC: Green. Queen Anne’s Lace, as well as leaves of birch and cranberry. From sweep nets and flight intercept and Mordellistena convicta LeConte – Larvae reared malaise traps in southern-mesic forests, northern from stems of Helenium autumnale Linnaeus, Silph- dry-mesic forests, and sandy oak barrens. Mid-May ium perfoliatum Linnaeus, and Solidago gigantea through mid-July. NC: Oneida; NE: Oconto; WC: Aiton (Andrew Williams, personal communication). Eau Claire, Jackson; EC: Fond du Lac; SW: Grant, Adults collected from prairies and mesic oak forests Richland; SC: Dane, Iowa, Sauk; SE: Milwaukee, using sweep nets, flight intercept traps, and malaise Racine, Waukesha. traps. Late May through early August. NC: Oneida; WC: Jackson, Monroe; EC: Fond du Lac; SW: Grant, Mordellistena aemula LeConte – Collected using SC: Dane, Iowa, Rock; SE: Walworth. malaise traps from mid-June to early August. WC: Jackson; EC: Fond du Lac; SE: Grant. Mordellistena dimidiata Helmuth – Collected by hand and from malaise traps in southern mixed Mordellistena aspersa (Melsheimer) – Collected deciduous hardwood forests and sandy oak barrens. primarily from sweep samples; one specimen recov- Mid-June to mid-August. SC: Dane, Columbia, Lafay- ered from a flight intercept trap. Most commonly ette; SE: Waukesha. collected in dry, dry mesic, and sand prairies; single specimens found in mesophytic and wet prairies. Mordellistena divisa LeConte – Commonly found Early June through late August. NE: Door, Oconto; feeding on flowers of Helianthus (Helianthus gigan- WC: Dunn, Pierce; C: Green Lake, SW: Buffalo, teus Linnaeus, Helianthus grosseserratus Martens, Grant, La Crosse, Pepin; SC: Columbia, Dane, Green, Helianthus occidentalis Riddell, and Helianthus tu- Iowa, Lafayette, Rock, Sauk; SE: Jefferson, Kenosha, berosis Linnaeus). Commonly collected by hand, Walworth, Waukesha. sweeping and from a yellow pan trap. Early August through late September. NW: Barron; WC: Chippewa; Mordellistena badia Liljeblad – One specimen SW: Grant, La Crosse; SC: Dane, Green, Lafayette, collected from an inflorescence of Queen Anne’s Lace. Suak; SE: Milwaukee, Racine. Additional specimens from flight intercept and mal- aise traps from sandy oak barrens and southern mesic Mordellistena fulvicollis (Melsheimer) – Collect- and northern dry mesic hardwood forests. Late June ed using malaise traps in oak savanna and sand to early August. C: Wood; SC: Dane, Green, Lafay- prairie ecotones. Early July to late August. WC: ette; SE: Waukesha. Monroe; EC: Fond du Lac; SC: Iowa.

Mordellistena bicinctella LeConte – Recovered Mordellistena fuscipennis (Melsheimer) – At from flight intercept and malaise traps in sandy oak blacklight; also from flight intercept and malaise barrens. Mid-June to mid-August. NW: Bayfield; EC: traps in sandy oak barrens and northern mesic and Fond du Lac; SW: Grant; SC: Iowa, Sauk; SE: dry mesic hardwood forests. Early July to mid-Au- Waukesha. gust. NW: Bayfield; NE: Oconto; C: Waupaca, Wood; SC: Dane; SE: Waukesha. Mordellistena cervicalis LeConte – Larvae reared from stems of Achillea millefolium Linnaeus and INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 199

Mordellistena gigas Liljeblad – From malaise oak and sand barrens. Early June to mid July. NW: trap samples. Mid-July through early August. NW: Bayfield; WC: Eau Claire, Monroe; SW: Grant, La- Bayfield; EC: Fond du Lac. Crosse; SC: Iowa, Rock; SE: Jefferson.

Mordellistena incommunis Liljeblad – From flow- Mordellistena rufilabris Liljeblad – Hand-col- ers of Cornus racemosa Lamarck and Rosa sp. Also lected in late-June while feeding on flowers of Eri- collected by hand from a prairie-forest ecotone, using geron sp. C: Marquette; SC: Rock. sweep nets, and in malaise traps. Late May through early July. NE: Oconto; WC: Jackson; SW: Grant; Mordellistena sexnotata Dury – Collected using SC: Columbia, Green, Iowa. malaise traps from mid-June to early August. NW: Bayfield; WC: Jackson. Mordellistena limbalis (Melsheimer) – At flow- ers of Ceanothus sp. and from malaise traps on the Mordellistena smithi (Dury) – Sampled at flight edge of a southern mixed deciduous hardwood forest intercept traps in a mesic hardwood forest from mid- and in an oak savanna. Late June through early to late July. SC: Sauk. September. NC: Oneida; WC: Jackson, Monroe; SW: Grant; SC: Lafayette. Mordellistena suturella Helmuth – Collected from Coreopsis sp., harebell, black eyed Susan, and flowers Mordellistena liturata (Melsheimer) – A single of Aster sp., and Rosa sp. Also taken using sweep nets specimen collected from herbaceous vegetation in a and flight intercept traps from dry and mesophytic mesic-hardwood forest in late June. SC: Sauk. prairies. Mid-June to late July. SC: Dane, Green, Iowa; SE: Jefferson, Walworth, Waukesha. Mordellistena nigella Liljeblad – Recovered from flight intercept and malaise traps. One specimen Mordellistena syntaenia Liljeblad – Taken with collected from a sandy prairie. Early July to late flight intercept and malaise traps primarily from oak August. SW: Grant; SE: Ozaukee. savannas, but also from mesic oak and southern deciduous hardwood forests. Mid-June to mid-August. Mordellistena ornata (Melsheimer) – Collected NW: Bayfield; WC: Monroe; SW: Grant; SC: Lafay- from sumac, poplar, Ceanothus sp., and flowers of ette. Cornus racemosa Lamarck and “Cornus femina” (likely Cornus foemina Miller.) Also taken at flight Mordellistena tosta LeConte –From flowers of intercept and malaise traps in oak savannas, sandy sumac. Also collected using sweep nets, blacklights, oak barrens, and northern and southern hardwood and malaise traps. One specimen trapped in a malaise forests. Early June to late July. NC: Oneida; NE: trap along edge of southern mixed deciduous hard- Oconto; WC: Eau Claire, Jackson, Monroe; C: Wau- wood forest. Late June to mid-August. NE: Langlade, paca, Wood; SW: Grant; SC: Dane, Green, Iowa, Oconto; EC: Fond du Lac; SW: Richland; SC: Dane, Lafayette, Sauk; SE: Milwaukee, Racine. Iowa, Lafayette; SE: Milwaukee.

Mordellistena pauxilla Liljeblad – From sweep Mordellistena trifasciata Say – Primarily collect- samples, flight intercept traps, and yellow pan traps ed using malaise traps; a single specimen collected in in dry and dry mesic prairies and dry prairie/ savanna a barrier pitfall trap; from sandy oak barrens, sand ecotones. Early June to late July. SC: Dane, Green; prairie/oak savanna ecotones, and mixed deciduous SE: Ozaukee. hardwood forests. Late June to early August. NW: Bayfield; NC: Oneida; WC: Jackson, Monroe; C: Mordellistena pulchra Liljeblad – Reared from Wood; SW: Grant; SC: Dane, Iowa, Lafayette; SE: stems of Achillea millefolium Linnaeus and Erigeron Waukesha. sp. Adults swept from prairies. Mid-July to early August. SC: Dane, Green. Mordellistena vera Liljeblad – A single specimen hand collected in late July from Ceanothus sp. in an Mordellistena rubrilabris Helmuth – Taken from oak savannah. WC: Monroe. flowers of Achillea millefolium Linnaeus, Coreopsis lanceolata Linnaeus, Erigeron sp., and Rudbeckia hirta Linnaeus. Also recovered from malaise traps in 200 Volume 17, No. 3-4, September-December, 2003, INSECTA MUNDI

Mordellistena vilis (LeConte) – Swept from a dry Iowa, Lafayette, Sauk; SE: Kenosha, Ozaukee, Wauke- mesic hill prairie. Early June- early July. SC: Colum- sha. bia, Iowa. Mordellina minutalis (Liljeblad) – Found in late Mordellina ancilla (LeConte) – Collected using July using flight intercept and yellow pan traps. SC: malaise traps from the edge of a southern mix decid- Dane; SE: Jefferson. uous hardwood forest. Early June through late Au- gust. NW: Bayfield; NC: Oneida; NE: Oconto; WC: Mordellina nigricans (Melsheimer) – Associated Jackson; EC: Fond du Lac; SW: Grant; SC: Iowa, with sand prairies, dry mesic prairies, prairie-hard- Lafayette; SE: Waukesha. wood forest ecotones, and sandy oak barrens, prima- rily using sweep nets, but occasionally in flight Mordellina blatchleyi (Liljeblad) – Swept from intercept or malaise traps. Early June through early sand, dry, dry mesic, and mesic prairies and prairie September. WC: Dunn; C: Marquette; SW: Grant; remnants. Late June to late August. SW: Buffalo, SC: Dane, Green, Iowa, Lafayette, Sauk; SE: Wauke- Grant; SC: Dane, Rock, Sauk. sha.

Mordellina floridensis (Ray) – Recovered from Mordellina parva (Liljeblad) – From dry/sand sandy oak barrens using a flight intercept trap. Early prairies, and sandy oak barrens using sweep nets, and to mid-July. SC: Sauk. barrier pitfall, flight intercept, and malaise traps. Late May to late August. NC: Oneida; WC: Jackson; Mordellina guttulata (Helmuth) – Collected from C: Marquette; EC: Sheboygan; SW: Grant; SC: Dane, a restored sandy prairie and an oak forest/marsh Rock, Sauk; SE: Waukesha. ecotone using a flight intercept trap. Also collected using malaise traps and sweep nets. Late May through Mordellina pustulata (Melsheimer) – Larvae late August. NC: Oneida; WC: Jackson; SW: Grant; reared from stems of Eupatorium maculatum Lin- SC: Iowa, Lafayette; SE: Ozaukee. naeus, Eupatorium perfoliatum Linnaeus, Gentiana andrewsii Grisebach, Helenium autumnale Linnae- Mordellina impatiens (LeConte) – Primarily as- us, Helianthus tuberosus Linnaeus, Heliopsis he- sociated with dry, dry mesic, sand prairies, and lianthoides Linnaeus, Rudbeckia laciniata Linnae- prairie remnants, but also collected from sandy oak us, Solidago canadensis Linnaeus, Solidago gigantea barrens. Usually collected in sweep nets, but also Aiton, and Vernonia fasciculata Michaux (Andrew using flight intercept and malaise traps. Early June Williams, personal communication), and Achillea through mid-September. NW: Bayfield; WC: Dunn, millefolium Linnaeus. Adults collected from prairies Eau Claire, Jackson; C: Marquette; SW: Grant; SC: using barrier pitfall, flight intercept, light, and mal- Dane, Green, Iowa, Lafayette, Sauk; SE: Walworth. aise traps and by sweeping. Late May through early August. NW: Bayfield; NC: Oneida; NE: Oconto; WC: Mordellina infima (LeConte) – Found in sweep Jackson; EC: Fond du Lac, Sheboygan; SW: Grant; samples and barrier pitfall, flight intercept, and SC: Dane, Green, Iowa, Lafayette, Rock; SE: Jeffer- malaise traps from sand prairies, sand and oak son, Kenosha, Ozaukee, Racine, Walworth. barrens, and oak savannas. Early June through late August. NW: Bayfield; NC: Oneida; WC: Eau Claire, Mordellina semiusta (LeConte) – Collected from Jackson, Monroe; C: Green Lake; SW: Grant; SC: sandy oak barrens using malaise traps. Mid-June to Dane, Lafayette, Sauk; SE: Jefferson. early September. WC: Jackson; SW: Grant; SC: Iowa; SE: Waukesha. Mordellina lecontei (Ermisch) – Collected prima- rily from prairies (sandy, dry, dry mesic, and wet Mordellina testacea (Blatchley) – Recovered from mesic) but also from sandy oak barrens, an oak/ malaise traps and by sweeping dry mesic and sand maple/pine forest, a dry mesic northern hardwood prairies. Late May through late August. NE: Oconto; forest, and a prairie-forest ecotone. Taken using WC: Jackson, Pierce; SW: Grant; SC: Dane, Iowa, sweep nets, flight intercept traps, and malaise traps. Sauk. Early June to mid-August. WC: Eau Claire, Jackson; C: Marquette, Wood; SW: Grant; SC: Dane, Green, Mordellina wickhami (Liljeblad) – Taken in dry prairies and sandy oak barrens using sweep nets and INSECTA MUNDI, Vol. 17, No. 3-4, September-December, 2003 201 flight intercept and malaise traps. Late June through serval, M. scapularis and M. ornata were collected mid-August. NW: Bayfield; WC: Jackson; SW: Grant; primarily from hardwood and mixed conifer-hard- SC: Iowa, Lafayette, Sauk. wood forests. Other species, including T. inclusa, M. undulata, T. discoidea discoidea, H. octopunctata, Pseudotolida arida (LeConte) – Collected from M. syntaenia, P. arida, and P. lutea were collected sandy oak barrens, oak savannas, and southern primarily from sand barrens and oak savannas. mesic hardwood forests using flight intercept, light, Several species, including M. melaena, M. margina- and malaise traps. Late June through late August. ta, M. divisa, M. aspersa, M. pustulata, and M. NW: Bayfield; NC: Oneida; WC: Jackson, Monroe; impatiens, were collected almost exclusively from EC: Fond du Lac; SW: Grant; SC: Dane, Green, Iowa; Wisconsin prairie remnants, which are particularly SE: Waukesha. prevalent in the southwest region of the state. Collecting methods appeared to be correlated with Pseudotolida lutea (Melsheimer) – Taken from certain species of Mordellidae, as well. Three species, flight intercept and Lindgren funnel traps and sweep- Mordella atrata, M. marginata, and M. cervicalis, ing in sandy oak barrens, oak savannas, and oak were readily hand-collected from a diverse array of forests. WC: Monroe; SW: Richland; SC: Lafayette, flowers while feeding, but were less frequently repre- Sauk; SE: Waukesha. sented in passive traps. Hoshihananomia octopunc- tata was hand-collected from flowers with relative Glipostenoda ambusta (LeConte) – Collected by frequency, but this is likely due more to collecting bias hand and using flight intercept and malaise traps. relating to the large size and conspicuous color of the One specimen collected from Queen Anne’s Lace, species than to actual species abundance. Mordellis- another from oak barrens between sand barrens and tena divisa was also hand-collected in relatively large a field. Early July through early August. NE: Sha- numbers during the course of this study as it can wano; SC: Dane, Iowa. reliably be found feeding on Helianthus sp. flowers. Some species of smaller mordellids common to prai- Falsomordellistena bihamata (Melsheimer) – A ries and savannas, particularly M. aspersa and spe- single specimen collected in August from the edge of cies of Mordellina, were most frequently collected a southern mixed deciduous hardwood forest using a using sweep nets. While the majority of mordellid malaise trap. SC: Lafayette. species were infrequently to rarely encountered by collectors in the field, many were obtained, sometimes Falsomordellistena discolor (Melsheimer) – in large numbers, using passive traps, especially Taken from sandy oak barrens and a mixed southern flight intercept and Townes’ malaise traps. forest with sandy soil using flight intercept and The limited larval rearing portion of this study malaise traps. Late June through early August. WC: yielded three species of Mordellidae from three plant Jackson; SW: Grant; SC: Iowa, Lafayette; SE: Wauke- species: M. cervicalis and M. pulchra were reared sha. from stems of A. millefolium and Erigeron sp., and M. pustulata was reared from A. millefolium, Erigeron Falsomordellistena hebraica (LeConte) – From sp., and H. grosseserratus. The mordellids reared by the edges of southern mixed deciduous hardwood Williams (1999) and reported as “unidentified” were forests using malaise traps. Late June through mid- determined: M. convicta was reared from S. perfolia- July. NC: Oneida; SC: Iowa, Lafayette. tum, S. gigantea, and H. autumnale, and M. pustu- lata was reared from E. maculatum, V. fasciculata, S. Falsomordellistena pubescens (Fabricius) – canadensis, R. laciniata, S. gigantean, H. helian- Collected from malaise traps along the edge of a thoides, H. tuberosus, G. andrewsii, H. autumnale, southern mixed deciduous hardwood forest. Late June and E. perfoliatum. to late July. WC: Jackson; SC: Dane, Lafayette. Acknowledgments Discussion We thank our labmates (Craig Brabant, Jeff Species of Mordellidae are common in woodlands Gruber, Kerry Katovich, Nadine Kriska, Michele like Wisconsin’s hardwood forests and grasslands like Price, Alistair Ramsdale, and Andrew Williams) for Wisconsin’s prairie and savanna remnants. During generous support and contribution of material to this this study, mordellid species such as T. lineella, M. study. We would also like to acknowledge the assis- 202 Volume 17, No. 3-4, September-December, 2003, INSECTA MUNDI tance of Steve Krauth (WIRC), Rich Henderson (WI- Jackman, J. A. and W. Lu 2001. Nomenclatural DNR), Ed Riley (Texas A&M University), Gary Par- changes for Mordellidae (Coleoptera) in North sons (Michigan State University) and the private America. Insecta Mundi 15: 31-34. collection of Bill Smith. We also thank John Jackman Jackman, J. A. and W. Lu. 2002. 101. Mordellidae (Texas A&M University) for graciously allowing ac- Latreille 1802, p.423-30. In Arnett Jr., R. H., M. cess to his library, notes, and private synoptic mordel- C. Thomas, P. E. Skelley and J. H. Frank (eds.) lid collection. Field work for this study was supported American Beetles. Volume 2. CRC Press LLC, in part by grants from the Zoological Society of Boca Raton, FL. Milwaukee and Prairie Biotic Research. The Wiscon- Kriska, N. L. and D. K. Young. 2002. An annotated sin chapter of The Nature Conservancy, the WI-DNR, checklist of Wisconsin Scarabaeoidea (Coleoptera). and several generous private landowners (Kriska, Insecta Mundi 16: 31-48. Norman, Uttech, and Wedel families) permitted ac- Lawrence, J. F. 1991. Mordellidae (Tenebrionoidea), cess to valuable collecting sites across the state. p. 508-509. In Stehr, F. W. (ed.) Immature In- sects. Volume 2. Kendall-Hunt Publishing Co., Literature Cited Dubuque, IA. Liljeblad, E. 1945. Monograph of the family Mordel- Bright, D. E. 1986. A catalog of the Coleoptera of lidae (Coleoptera) of North America, north of America North of Mexico, Family: Mordellidae. Mexico. Miscellaneous Publications, Museum of USDA, ARS, Agriculture Handbook 529-125, 22 Zoology, University of Michigan, No. 62, 229 p. p. Lisberg, A. and D. K. Young. 2003. Descriptions of Colwell, R. K. 1996. Biota: The Biodiversity Data- larva and pupa of Tomoxia lineella LeConte with base Manager. Sinauer Associates, Sunderland, notes on larval habitat (Coleoptera: Mordellidae). MA, 574 p. Coleopterists Bulletin 57: 17-22. Ford, E. J. and J. A. Jackman. 1996. New larval Lisberg, A. E. 2004. Taxonomic changes for fifteen host plant associations of tumbling flower beetles species of North American Mordellidae (Co- (Coleoptera: Mordellidae) in North America. Co- leoptera). Insecta Mundi 17: 191-194. leopterists Bulletin 50: 361-368. Lu, W. 1997. Notes on identification and ecology of Hilsenhoff, W. L. 1995. Aquatic Hydrophilidae and tumbling flower beetles (Mordellidae) from Ossa- Hydraenidae of Wisconsin (Coleoptera). I. Intro- baw Island, Georgia. Florida Entomologist 80: 95- duction, key to genera of adults, and distribution, 98. habitat, life cycle, and identification of species of Robertson, C. 1928. Flowers and Insects: Lists of Helophorus Fabricius, Hydrochus Leach, and visitors of four hundred and fifty three flowers. Berosus Leach (Hydrophilidae), and Hydraenidae. Published by the author, Carlinville, IL, 222 p. The Great Lakes Entomologist 28: 25-53. Williams, A. H. 1999. Fauna overwintering in or on Jackman, J. A. 1991. Notes on the nomenclature of stems of Wisconsin prairie forbs. Proceedings of Mordellidae of America north of Mexico. Coleop- the North American prairie conference 16: 156- terists Bulletin 45: 323: 330. 161.