Scientific Papers

Home , Ammobatini, Ammobatoidini, Ancylandrena, Andrena banksi, Andrena bradleyi, Andrena caerulea

QL 566 .M43 1997 Scientific Papers

Natural History Museum The University of Kansas

25 July 1997 Number 1:1-81

Genus-Group Names of Bees and Supplemental Family-Group Names^

By Charles D. Michener MCZ LIBRARY Division of Entomologi/, Natural History Museum The Kansas University of QCOLU 1997 Lawrence, Kansas 66045, USA CONTENTS HARVARD UNIVERSITY ABSTRACT 1 INTRODUCTION 2 Acknowledgments 2 GENUS-GROUP NAMES 2 SUPPLEMENT TO THE LIST OF FAMILY-GROUP NAMES 52 LITERATURE CITED 63 ADDENDUM 81

ABSTRACT This paper includes an alphabetical list of generic and subgeneric names that have been proposed for bees, with bibliographic references and indications of type species. Synonymies are not indicated, but the major group (family subfamily tribe or sometimes merely "fossil") is indicated for each. Synonymies will be indicated in a future account of world bees. New nomenclatural details include the following: Apis tlioracica Fabricius designated as type species o( Melanchxnia Perez; Paracella new subgenus of Megachile, with M. seniiivjiusta Cockerell as type species, for Paracella Pasteels, a name proposed without designation of a type species; Callistochlora new name for Callochlora Moure, 1964, not Packard, 1864; Gyrodroniella new name for Gyrodronia Thomson, 1872, not Klug, 1807; and Lithurgus Berthold, 1827, given preference over the French vernacular Lithurge Latreille, 1825. The list of family-group names in bees is corrected and up-dated.

Key words: Apoidea; Generic and Family names; Type species.

' Contribution Number 3167 from the Snow Entomological Division Natural History Museum, and Department of Entomology, The University of Kansas

QL 566 .M43 1997 Scientific Papers

Natural History Museum The University of Kansas

25 July 1997 Number 1:1-81

Genus-Group Names of Bees and Supplemental Family-Group Names^

By Charles D. Michener MCZ LIBRARY Division of Enkvnologi/, Natural History Miisfuin

The University of Kansas QcnLO 1 7 1997 Lawrence, Kansas 66045, USA CONTENTS HARVARD UNIVERSITY ABSTRACT 1 INTRODUCTION 2 Acknowledgments 2 GENUS-GROUP NAMES 2 SUPPLEMENT TO THE LIST OF FAMILY-GROUP NAMES 52 LITERATURE CITED 63 ADDENDUM 81

ABSTRACT This paper includes an alphabetical list of generic and subgeneric names that have been proposed for bees, with bibliographic references and indications of type species. Synonymies are not indicated, but the major group (family, subfamily, tribe or sometimes merely "fossil") is indicated for each. Synonymies will be indicated in a future account of world bees. New nomenclatural details include the following: Apis thoracica Fabricius designated as type species oi Melandrena Perez; Paracella new subgenus of Megachile, with M. seuiivenusta Cockerell as type species, for Paracella Pasteels, a name proposed without designation of a type species; Callistochlora new name for Callochlora Moure, 1964, not Packard, 1864; Gyrodromella new name for Gyrodroma Thomson, 1872, not Klug, 1807; and Litlntrgiis Berthold, 1827, given preference over the French vernacular Lithiirge Latreille, 1825. The list of family-group names in bees is corrected and up-dated.

Key zvords: Apoidea; Generic and Family names; Type species.

' Contribution Number 3167 from the Snow Entomological Division Natural History Museum, and Department of Entomology, The University of Kansas

Table 1. The last list of genus-group names of bees was that of Sandhouse (1943). Since that time many new taxa have been described; there were about 977 entries in her list, compared with about 2057 herein. It is of some interest to examine the entire list from the viewpoint of production of genus-group names or actual entries in the list, including nomina nuda and the like. From the initiation of Lin- naean nomenclature until the end of 18th Century, five generic names were proposed for bees, Andrena, Apis, Eucera, Hylaeus, and Nomada. But for the first decade of the next century, 1800 to 1809, there are 66 entries. The result of a tabulation by decade, from 1800 to the present, is shown in Table 1. In a reevaluation in preparation, I expect to recognize about 867 genera and subgenera. the exten- Melittologists are sometimes criticized for sive use of subgenera, because species-groups, named in- formally with species names, could be used instead of subgenera. Such a procedure would avoid burdening the literature with numerous subgeneric names with their associated formalities such as type species, problems of homonymy, etc. Such a criticism is justified for certain groups of bees, especially in certain regions of the world.

I believe strongly, however, that large genera are commonly practical and desirable. Nearly all biologists have some idea of what is meant by generic names like Droso- philn, Cidex, and Anopheles, and many would recognize major bee genera like Aiidreun or MegachUe. If the subgenera of each of these were elevated to generic rank, the av- erage biologist would often lose track of the sort of organ- ism involved. For many large genera with hundreds or thousands of species, the choice is not between recognizing the genus with subgenera vs. the genus with species groups. It is between recognizing the genus with subgenera vs. several or many genera, because many of the current subgenera are quite different from one another, sometimes recognizable in flight, and many specialists prefer to recognize them as genera. I prefer to retain inclusive genera, for the reason indicated above. These decisions, however, are subjective regardless of one's views on systematic methodology. Some recent bee specialists have used generic names in much the same way that I would use tribal names. More, however, split "excessively," Robertson (1918b) being one of the first to be overtly and purposely a "splitter." One of the advantages of subgenera is that they need not be cited, i.e., unlike genera, which are required, they Genus-Group Names of Bees

The first 1807) shows no author either in the table rent tribe, subfamily, or faniily name is provided (in bold- (Illiger, of contents of the volume or in the text an indication face type) for each entry, following the classification put itself, that it was written The second 1807b) is forward by Roig-Alsina and Michener (1993) and by Illiger. (Klug, Alexander and Michener (1995). attributed to Klug in both places. The two articles were published simultaneously; page 199 of the first article is I have cited the names of type species in their original on the same leaf as page 200 of the second. Some of the combinations. This means that I have ignored the same generic names were proposed in both articles. All of complexities that could have been cited when an author, in them have been attributed in the past to Klug, and Illiger's designating a type species, placed it in a genus different from article in most cases credits Klug with such names. Given the original one or cited an erroneous author. This policy is that publication was simultaneous, I have adjusted the generally in accord with the Code, ed. 3, Art. 67d, f. references to retain Klug's authorship, in different ways Of course there have been errors, especially probable in different cases because of differing treatments in the two printers' errors, from time to time. For example, Westwood articles concerned. (1840a) instead of Coelioxi/s. published Caelioxys Except Westwood (1840a) listed a typical British species for when such errors have been or have nomencla- repeated each genus. These have been taken as type designations, tural I have not listed them. Also, nomina significance, in accordance with Opinion 71 of the Commission (1922). nuda that have not been as to a recognized relating par- Westwood listed synonymous generic names in his ac- ticular taxon are omitted. counts of many genera, often with the notation "p." for I follow Sandhouse (1943) in using the term monoba- "part." It seems clear that he was merely listing synonyms sic to indicate that there was only one originally included and that his type designations did not apply to these syn- species, and autobasic to mean that a replacement name onymous names. has the same as its antecedent, automatically type species Mehely's monumental work (1935) on palearctic name. When a is monobasic usually preoccupied, genus Hylaeus included descriptions of various subgenera. When and in the the was also original publication single species monobasic, such subgenera are valid, but those including as the I use the designated type species, only expression more than one species are invalid because of his failure to since the action the "by original designation," positive by designate a type species, a requirement after 1930 (Code, author of the name seems more than the mere significant 3rd ed.. Art. 13b). Popov (1939) rectified the sih.iation by fact that there was but one included originally species. designating type species as needed, except for certain of which he (and authors) Sandhouse (1943) used the term isogenotypic for taxa Mehely's subgenera subsequent as format included a having the same type species. Since genotype is no longer regarded junior synonyms. Popov's for a followed a list of used to mean the type species of a genus, I have changed heading subgenus, by synonyms, material, and then a statement of the the term to isotypic. When two generic names are isotypic, descriptive type species. It is clear that his intention was to a they are objectively synonynious. Isotypic by synonymy, designate type for the indicated in the however, means that the taxa are the same only by syn- species subgenus heading. Sandhouse (1943), however, considered that same onymy of their type species, which is usually subjective. species to be the of each of the names, result- The synonymy of such generic names therefore may de- type synonymized in the same for more than one of pend on the accuracy of the synonymy of the type species. ing type species Mehely's subgenera. Popov's paper cannot be read to justify this; Sandhouse cited "Blanchard, 1849" as a repeatedly he did not designate type species for Mehely's names that source of Thanks to Cowan, 1976 (see type designations. he considered synonyms. Sandhouse herself could be in- Blanchard, 1845-1849), we now know that the of this parts terpreted to have designated type species for the junior work (Cuvier's edition) on bees were Disciples' published synonyms, although such types would not be species in- in 1845, 1846, and 1847. When 1 first examined it, I con- cluded under those names by Mehely. Thus, for example. cluded that this work did not contain type designations Imperfecta Sandhouse, 1943, would have as type species a was listed for each in the although single species genus Hylaeus brevicornis Nylander (a species of Dentigera); it of the However, as out to me explanations plates. pointed would not be the same as Imperfecta Mehely, and there- Dr. D. B. Baker, the title contains in small the by page type, fore, not being accompanied by a description or relevant notation de "accompagne planches gravees, representant bibliographic reference, would be a nomen nudum. More- les de tous les Thus, were types genres,...." types desig- over, Sandhouse's type designations do not validate these nated, but as of the date of of the title publication pages, subgeneric names, since she considered these names to be 1849. 1 indicate the of of the in year publication plates junior synonyms and they were not adopted as the names The authors' names attached to in parentheses. species of taxa before 1961 (Code, 3rd ed.. Art. lie). They thus Blanchard's of are explanations plates commonly wrong; remain invalid. The names involved are Aiiricularia, as elsewhere, I have such errors and cited the cor- ignored Imperfecta, Pectinata, Pseudobranchiata, and Trichota. All this rect authors. has no effect whatever on the current nomenclature and I Another source of confusion has been two articles have included these names below only as proposed by to or published in 1807 in Illiger's Magazin fiir Insekteiikunde. Mehely, not as attributed Popov Sandhouse. Scientific Papers, Natural History Museum, The University of Kansas

Abda Sandhouse, 1943:521, replacement for Anthedon Acentron Mitchell, 1 934:307. Type species: Megachile albitarsis Cresson, 1872, Robertson, 1900. Type species: Melissodes compta Cresson, by original designation. [Megachile]. 1878, autobasic. See Anthedonia. Eucerini. Megacliilini. Aceratosmia Schmiedeknecht, 1885:19. Type species: Osmia Abromelissa Snelling and Brooks, 1985:23. Type species: Mel- emarginata 1841, by designation of Sandhouse, issa lendliana Friese, 1910, by original designation. Lepeletier. 1939:9. [Osmia]. Osmiini. Ericrocidini. Aciandrena Warncke. 1968:62. Type species: Andremi aciculata Abrupta Mehely, 1935:32, 137. Type species: Hylaeus coniufus Morawitz. 1886, by original designation. [Andrena]. Curtis. 1831. monobasic. Sandhouse ( 1943) attributed Abnipta Andreninae. to Popov (1939). perhaps because Mehely. while including 1988:508. as a Ackmonopsis Brooks, Type species: Anihophora only one species, listed another specific name synonym. mimadvena Cockerell, 1916, by original designation. She may thus have considered that Mehely included two spe- [Amegilla). Antliopliorini. cies without designating a type. [Prosopis]. Hylaeinae. Acritocentris Snelling, 1974:36. Type species: Centris ruthannae Acalcaripes Pesenko, 1984:347. Type species: Halictus patellatiis Snelling, 1966, by original designation. [Centris]. Centridini. Morawitz. 1873, by original designation. [HalictusJ. Halictini. Acrocoelioxys Mitchell, 1973: 71. Type species: Coelioxys Acamptopoeum Cockerell, 1905a:320. Type species: otomita Cresson, 1 878, by original designation. [Coelioxys]. 1851, sensu Cockerell, Camptopoeum trifasciatum Spinola, Megachilini. 1905a = Camptopoeum submetallicum Spinola, 1851, Acrosmia Michener, 1947:298. Type species: Hoplitis monobasic. Because ofCockerelFs misidentification, the type plagiostoma Michener, 1947, by original designation. species was designated as Camptopoeum submetallicum [Hoplitis]. Osmiini. Spinola. 1851. by Commission Opinion 1759 (1994). Hurd and Moure, 1963:236. Type species: Panurginae. Acro.xylocopa Xyiocopa capitata Smith. 1854, by original designation. Acanthalictus Cockerell, 1924a: 184. Type species: Halictus [Xylocopa]. Xylocopini. dyboMskii Radoszkowski, 1877, by original designation. Acunomia Cockerell, 1930, in Cockerell and Blair, 1930: 1 1 . Type Halictini. species: Nomia nortoni Cresson, 1868. by original designa- Acanthidiiim Michener and Griswold, 1994a:305. Type species: tion. [Nomia]. Nomiinae. Acanthidium batrae Michener and Griswold, 1994 = Adanthidium Moure, 1965:29. Type species: Anthidium texanum Dianthidium flavoclypeatum Gupta, 1993, by original desig- Cresson, 1878, by original designation. Antliidiini. nation. Anthidiini. Addendella Mitchell, 1 980:24. Type species: Megachile addenda Acatitbomelecta Lieftinck, 1972:316. Type species: Crocisa Cresson, 1878, by original designation. [Megachile]. bicuspis Stadelmann, 1898, by original designation. Megachilini. Meiectini. Adventoribombus Skorikov, 1922a:25. Type species: Apis Acanthonomada Schwarz, 1966:383. Type species: Nomada 1971:28. sylvarum Linnaeus, 1 76 1 , by designation of Yarrow, Kohl, 1905, designation. Nomadini. = odontophora by original Agrabombus [sic J adventor Skorikov, 1914 B. filclmerae Sandhouse, was not Acanthopus Klug, 1807, in Illiger, 1807:199; Klug, 1807b:226. Vogt, 1908, designated by 1943:522, included. with Type species: Apis splendida Fabricius, 1793 -Apispalmata originally [Agrobombus]. Isotypic Olivier, 1789, monobasic. Acanthopus is consistently attrib- Thoracobombus. Bombini. literature. It was in two con- uted to Klug in the published Aeganopria Moure, 1949a:442. Type species: Lonchopria nivosa thus secutive articles in the same journal and simultaneously; Vachal, 1909, by original designation. Colletinae. the first was who, however, attributed the name to by Illiger, Aenandrena Warncke, 1968:64. Type species: Andrena the second was Dalla Torre ( 1 896) listed both Klug; by Klug. aeneiventris Morawitz, 1872, by original designation. references under the but attributed the name species generic [Andrena]. Andreninae. to Klug, and may therefore be regarded as the first reviser, Aethammobates Baker, 1 994: 1 55. Type species: Aethammobates and to have decided the question as to authorship of prionogaster Baker, 1994, by original designation. Acanthopus. Ericrocidini. Ammobatoidini. Acanthosmia Thomson, 1872:233; no first in- species. Species Aethechlora Moure and Hurd, 1987:275. Type species: cluded Schmiedeknecht, 1885:21. Osmia by Type species: Augochloru matucanensis Cockerell, 1914, by original des- Morawitz, 1872, of Sandhouse, montivaga by designation ignation. [Augochlora]. Augochlorini. 1943:522. [Osmia]. Osmiini. Aframegilla Popov, 1 950b:260. Type species: Anthophora nubica Acanthosmiades Titus. 1904b: 101, incorrect subsequent spell- Lepeletier. 1841, by original designation. [Amegilla]. ing of Acanthosmioides Ashmead, 1899. Osmiini. Anthophorini. Acanthosmioides Ashmead, 1899a:76. Type species: Osmia Afranihidium Michener. 1948a:24. Type species: Hypanthidium = odonlogaster Cockerell, 1897, by original designation. ludophilum Cockerell. 1936 Anthidium reicherti Brauns, Osmiini. 1929, by original designation. [Anthidium]. Anthidiini. Genus-Group Names of Bees

Afrodialictiis Pauly, 1984b: 142. Type species: Halictus heUulus Agrobombus Vogt, 1911 :52. Type species: Apis agrorum Fabri- = Vachal. 1909. by original designation. [Lasioglossum]. cius, 1787 (not Schrank, 1781 ) Apis pascuorwn (Scopoli), Haiictini. 1763, by designation of Sandhouse, 1943:523. [Bombus]. Bombini. Afrodufourea Ebmer, 1984:362. Type species: Dufoitrea pimica Ebmer, 1976, by original designation. [Dufourea]. Alayoapis Michener, 1966a:728. Type species: Megacilissa Rophitinae. nigrescens Cresson, 1869, by original designation. Afroheriades Peters, 1970:157. Type species: Pseudoheriades [Caupolicana]. Caupolicanini. primus Peters, 1970, by original designation. Albocolletes Warncke. 1978:353. Type species: Halictus Osmiini. [Pseudoheriades]. albomaculatus Lucas, 1849, by original designation. Afroinelecla Lieftinck, 1972:309. Type species: Crocisa [Colletes]. CoIIetinae. Cameron, 1905, Melectinl. fulvoliirta by original designation. Alcidamea Cresson. 1 864:385. Type species: Alcidamea producta Afronomia Pauly, 1990:126. Type species: Nomia picardi Cresson. 1864. by designation of Michener. 1941:158. Gribodo, 1894, by original designation. Nomiinae. Isotypic with Aiitoclielostoma by synonymy. Osmiini.

Afrostelis Cockerell, 1931b:340. Type species: Afrostelis Alepidosceles Moure. 1947b:244. Type species: Ancyloscelis tegularis Cockerell, by original designation. Anthidiini. imitatrix Schrottky. 1909. by original designation. Afwxylocopa Hurd and Moure. 1963:264. Type species: Apis Emphorini. nigrita Fabricius, 1775. by original designation. [Xylocopa]. Alfkenella Bomer. 1919: 168. Type species: Apis quadrifasciala Xylocopini. Villers. 1789. by original designation. Isotypic W\Xh Amegilla. Agaesischia Moure and Michener, 1955b:273. Type species: Anthophorini. Eucera Ducke, patellicomis 1910, by original designation. Alfkenomia Hirashima. 1956:33. replacement for Epinomia [Gaesischia]. Eucerini. Alfken. 1 939. Type species: Nomia andrenoides Vachal. 1 903 = Agandrena Wamcke. 1968:56. Type species: Apis agilissiina Nomia a it dre i Vachal, 1897, autobasic. Nomiinae. 1770. [Andrena]. Scopoli, by original designation. Alfkenylaeus Snelling, 1985:13. Type species: Hylaeus Andreninae. namaquensis Cockerell, 1942, by original designation. Agapanthiints LaBerge, 1957:35. Type species: Melissodes [Hylaeus]. Hylaeinae. callophila Cockerell, 1923. by original designation. Eucerini. AUanihidium Moure, 1 947a:2 1 . Type species: Anthidium rodolfi 1844:448. Agapostemon Guerin-Meneville. Type species: Apis Ruiz, 1938, by original designation. Anthidiini. femoralis Guerin-Meneville. 1844 = Apis viridula Fabricius. AllochalicodomaTka\cu, 1969:359. Type species: Chalicodoma 1793, monobasic. [Andrena]. Haiictini. lefebvrei Lepeletier. 1841. by original designation. Roberts and Brooks, 1987:364. Agapostemoiwides Type species: [Chalicodoma]. Isotypic with Heteromegachile. Megachilini. Agapostemonoides liiirdi Roberts and Brooks, 1987. by origi- Allocoelioxys Tkalcu. 1974a:340. Type species: Coelioxys afra nal designation. Haiictini. Lepeletier. 1841. by original designation. Isotypic with Sitdikov and Pesenko. 1988:87. Agatheitcera Type species: Coelioxita. Megachilini. Eucera bidentata Perez, 1887, by original designation. AUodapa Schulz. 1906:244. unjustified emendation of A//o<^ape [Eucera]. Eucerini. Lepeletier and Serville, 1825. Allodapini. Agemmonia Michener, 1 98 1 :26. Type species: Nomia tsavoensis Allodape Lepeletier and Serville, 1825:18. Type species: Strand, 1920, by original designation. Meganomiinae. Allodape rufogastra Lepeletier and Serville, 1825, monoba- Schulz, 1906:258. emendation of A Aglaa Unjustified glae sic. Allodapini. Lepeletier and Serville, 1825. Euglossini. AUodapula Cockerell, 1934c:220. Type species: Allodape and Serville. 1825:105. Aglae Lepeletier Type species: Aglae variegata Smith, 1854, by original designation. [Allodape]. cacriilca and monobasic. Lepeletier Serville, 1825, Allodapini. Euglossini. Allodapulodes Michener. 1969:291. Type species: Allodape 1901b:262. Aglaoapis Cameron, Type species: Aglaoapis palliceps Friese. 1924. by original designation. [AUodapula]. monobasic. brevipennis Cameron, 1901, Dioxyini. Allodapini. and Brooks. 1985:25. Me- Aglaomelissa Snelling Type species: Allodioxys Popov. 1947:87. Type species: Dioxys schulthessi lissa diickei Friese. 1906. by original designation. Popov. 1936. by original designation. Dioxyini. Ericrocidini. Allomacrotera Timberlake. 1960:131. Type species: Perdita Agogenoliylaeus Michener, 1965a: 148. Type species: Prosopis stephanomeriae Timberlake. 1954. by original designation. nul?ili>sellus Cockerell. 1910. by original designation. [Perdita]. Panurginae.

[Amphylaeus]. Hylaeinae. Allomegachile Rebmann. 1970:42. Type species: Megachile Agrihombus Skorikov, 1938a: 145. unjustified emendation of asiatica Morawitz, 1875. by original designation. Agrobombus Vogt, 1911. Bombini. [Megachile]. Isotypic with Euchalicodoma. Megachilini. Scientific Papers, Natural History Museum, The University of Kansas

AUoperdita Viereck, 1917:241. Type species: Perdita Amegachile Friese, 1909a:326. Type species: Megachile novaeangliae Viereck, 1907, monobasic. [Perdita]. sjoestedti Friese, 1901 = Megachile bituberculata Ritsema, Panurginae. 1880, by designation of Cockerell, 1931c;167. Mitchell's (1934:298) of nasicomis Friese, 1903, Allopsitliynis Popov. 193 la: 136. Type species: Apis harhuteUa designation Megachile as the is and therefore invalid. Kirby, 1802. by original designation. [Psithyrus]. Bombini. type species subsequent [Megachile]. Megachilini. Alloscirtetica Holmberg, 1909a:77, replacement for Sciretetica 1 897a: 1 24. Holmberg. 1903. Type species: Scirtetica antarctico Amegilla Friese, 8, Type species: Apis quadrifasciata 1789, of 1931e:277. Holmberg. 1903, autobasic. Isotypic with Holinbergiapis and Villers, by designation Cockerell, Neoscirtetica. Eucerini. [Podalirius]. Isotypic W\Xh Alfkenella. Anthophorini.

Allosmia Tkalcu, I974a:331. Type species: Osmia rufohirta Ammobates Latreille, 1809:169. Type species: Ammobates Latreille, 181 1, by original designation. [Osmia]. Osmiini. rufiventris Latreille, 1809, by designation of Latreille, 1 8 1 0:439. Ammobates bicolor Lepeletier and Serville, 1825, Alloxylocopa Maa. 1939:155. Type species: Xylocopa designated by Blanchard, 1849:pl. 127 (1846), was not appendiculata Smith, 1852, by original designation. Nomen origi- included. Ammobatini. nudum because no characters given; see the Code, 3rd ed.. nally Art. 13(a)(i). [Xylocopa]. Xylocopini. Awi/^oiiflro/V/c^ Radoszkowski, 1868:82, not Schenck, 1869. Type species: Phileremus abdominalis Eversmann, 1852, des- Alloxylocopa Hurd and Moure, 1963:239. Type species: Xylocopa by of Sandhouse, 1943:525. with Phiarus and appendiculata Smith, 1852, by original designation. ignation Isotypic Paidia. Ammobatoidini. [Xylocopa]. Xylocopini. Ammobatoides Schenck, 1869:349, not Radoszkowski, 1868. Alocandreua Michener, 1986c:68. Type species: Alocandrena Ammobates bicolor and porteri Michener, 1986, by original designation. Andreninae. Type species: Lepeletier Serville, 1825 = Epeolus punctatus Fabricius, 1804, monobasic. Not Alphaneiira Gray, 1832:pl. 1 1 1, alternative for Aphaneura Gray. replaced because synonym of Ammobates. Isotypic with Type species: Aphaneura rufescens Gray, autobasic. Phileremus by synonymy. Ammobatini. Meliponini. Amphipedia Michener and Moure, 1957:413. Type species: Alpigenihomhus Skorikov. 1938a: 145, unjustified emendation haeckeli Friese. 1910, of Alpigenobombits Skorikov, 1914. Bombini. Tetrapedia by original designation. [Paratetrapedia]. Tapinotaspidini. Alpigenobombus Skorikov, 1914a: 128. Type species: Micfxcncr. 1965a: 147, 149. Alpigenobombus pulcherhmus Skorikov, 1914 = Bombus Amphylaeus Type species: Prosopis morosa Smith, 1879, kashmirensis Friese, 1909, by designation of Williams, by original designation. Hylaeinae. 1991:65. Bombus alpigenus Morawitz, designated as the type AnacoUetes Michener. 1965a:59. Type species; Lamprocolletes species by Prison, 1927:54, was not an originally included bimaculatus Smith, 1879, by original designation. species; see Williams, 1991. [Bombus]. Bombini. [Leioproctus]. Colletinae.

Alpinibombus Skorikov, 1937:53, unjustified emendation of Analastoroides Rayment, 1 950:20. Type species: Analasteroides Alpinohombus Skox'\ko\, 1914. Bombini. foveata Rayment, 1950, by original designation. Hylaeinae.

Alpinobombus Skorikov, 1914a: 122. Type species: Apis alpimis Ananthidiellum Pasteels, 1969a:49. Type species: Anthidium Linnaeus, 1758, by designation of Prison, 1927:66. [Bombus]. anale Friese, 1914, by original designation. [Anthidiellum]. Bombini. Anthidiini. Ebmer, 1984:360. Alpinodufourea Type species: Dufourea alpina Ananthidium Urban, 1991:73. Type species: Anthidium inerme Morawitz, 1865, by original designation. [Dufourea]. Friese, 1908, by original designation. Anthidiini. Rophitinae. Anchandrena LaBerge, 1986:496. Type species: Andrena Amalthea 1815:123. for Rafinesque, unjustified replacement angustella Cockerell, 1936, by original designation. Jurine. amalthea Olivier, 1789, Trigona Type species; Apis [Andrena]. Andreninae. autobasic and by absolute tautonomy. Meliponini. Auchirhiza Michener. 1965a: 147. Type species: Palaeorhiza Amblyapis Cockerell, 1910b:362. Type species: Halictoides mandibularis Michener, 1965, by original designation. ilicifolioe Cockerell, 1910, by original designation. [Palaeorhiza]. Hylaeinae. [Halictoides]. Dasypodinae. Ancyla Lepeletier, 1841:294. Type species: Ancyla oraniensis Amblys Klug, 1807, in Illiger, 1807:198; Klug, 1807b:226. Type Lepeletier, 1841, monobasic. Isotypic with Plistotrichia by species: Apis bicomis Linnaeus, 1758 = Apis rufa Linnaeus, synonymy. Ancylini. 1758, by designation of Latreille, 1811:577. Monobasic in Cockerell, 1930d:5. Andrena Illiger, 1807; Klug, 1807b was published simultaneously and Ancylandrena Type species: listed two species. Isotypic with Osmia and Ceratosmia; heterodoxa Cockerell, 1930, not Perez, 1903 =A. atoposoma isotypic with Pachyosmia by synonymy. Osmiini. Cockerell, 1934, monobasic. [Andrena]. Andreninae.

Amblyspatulariella Pittioni. 1950:80. Type species: Prosopis Ancylocopa Maa, 1954: 198. Type species: Xylocopa nitidiventris sulphuripes Gribodo, 1894, by original designation. Smith, 1878, by original designation. [Proxylocopa]. [Prosopis]. Hylaeinae. Xylocopini. Genus-Group Names of Bees

Ancylosceles Haliday, 1836:320, unjustified emendation of Anhylaeiis Heider, 1 926: 1 84. unjustified emendation of Anylaeus Ancyloscelis Latreille, 1829. Emphorini. Bridwell, 1919. Hylaeinae.

Ancyloscelis Berthold, 1827:466, nomen nudum. Latinization Annosmia Wamcke. 1991b:307. Type species: Osmia annulata of Anc\7o5ce/f Latreille. 1825:463. Emphorini. Latreille, 181 1. by original designation. [Osmia]. Osmiini.

Ancyloscelis Latreille, 1829:355, no included species: Haliday, Anodonteutricharaea Tkalcu, 1993b:807. Type species: = 1836:320, one included species. Type species: Ancylosceles Megachile larochei Tkalcu, 1993 Megachile lanigera (sic) iirsinus Haliday, 1836. first included species. Blanchard, Alfken, 1933. by original designation. [Megachile]. 1849:pl. 128 (1847), designated Ancyloscelis brasiliensis Megachilini.

Blanchard, 1847, as but it was not the first in- type species Anodontobomhiis Kriiger, 1917:61. Type species: Apis hypnorum cluded species. Emphorini. Linnaeus, 1758, by designation of Williams. 1991:69. A sec- Ancyloscelus Berthold, 1827:565, nomen nudum. Emphorini. tional name treated as a genus-group name in view of the Code, 3rd ed.. Art. 10(e). [Bombus]. Isotypic with Ancyiosoma Dalla Torre, 1896:206. This name appeared in the Pyrobombus, Uncobombus. and Hypnorobombus. Bombini. synonymy of Xylocopa hrasilianonim (Linnaeus), as ''Apis (Ancyiosoma) Brasilianoriim Blanchard. ...1849...." The Anoediscelis Toro and Moldenke. 1979:131. Type species: is Oediscelis herhsti Friese, 1906, Blanchard reference to pi. 128 ( 1847), where Ancyiosoma by original designation. Xeromelissinae. does not appear. Blanchard. however, did propose Ancyloscelis brasiliensis for a bee on the and probable tapinotaspine plate Anomalohesma Exley, 1977:39. Type species: Brachyhesma numbers listed Dalla Torre. Dalla Torre figure by Evidently scapata Exley, 1977. by original designation. [Brachyhesma]. erred in genus and species names, and in placing the refer- Euryglossinae. ence among genuine Xylocopa references. The species name Anthedon Robertson, 1900:53, not Agassiz, 1846. Type species: Apis (Ancyiosoma) brasilianorum Blanchard refers to no spe- Melissodes compta Cresson, 1 878, monobasic. SttAbda and Dalla Torre in cies; Ancyiosoma appeared only synonymy Anthedonia. Eucerini. under Xylocopa. associated with a nonexistent species, and Anthedonia Michener, 1942:282, replacement for Anthedon therefore also is not available for use. [Apis]. Not available. Robertson, 1900. Type species: Melissodes compta Cresson, Eickwort. 1969:407. Andinaugochlora Type species: 1878, autobasic. Eucerini. Andinaugochlora micheneri Eickwort, 1 969, by original des- AnthemoessaKobtvl&on, 1905:372, not Agassiz, 1 846. Type spe- ignation. Augochlorini. cies: Anthophora abrupta Say. 1837, by original designation. Anrf/neta Ashmead, 1899a:71, incorrect for subsequent spelling See Melea. Anthophorini. Audinetia Lepeletier, 1841. Xylocopini. Anthemois Roh&nson, 1903a: 168, 172. Type species: Megac/ij/e Andrena Fabricius, 1775:376. species: Apis helvola Type infragilis Cresson, 1 878 =Apis centuncularis Linnaeus, 1758, Linnaeus, 1758, of Viereck, 1912:613. by designation Apis by original designation. Isotypic with Megachile by syn- c(;ieran'fl Linnaeus, 1758, Latreille, 1810:439; designated by onymy. Megachilini. Apis niiida Fourcroy, 1785, designated by Curtis, 1826:pl. 1 902b:32 1 . = Anthemitrgus Robertson, Type species: Anthemurgus 129; and Andrena fiinebris Panzer. 1 798 Apis albopunctata passiflorae Robertson. 1902. monobasic. Panurginae. Rossi, 1792, designated by Blanchard, 1849:pl. 125 (1847) Anthidianum Michener. 1948a: 13, for were not originally included species. Andreninae. replacement Trichanthidinm Moure. 1947. Type species: Anthidiiim Andrenella Hedicke, 1933:210. Type species: Melitta minutula subpetiolatum Schrottky, 1910, autobasic. [Allanthidium]. Andreninae. Kirby, 1802, by original designation. [Andrena]. Anthidiini. Andrenopsis Cockerell, 1905a:363. Type species: Amirenopsis Anthidiellum Cockerell, 1904c:3. Type species: Trachusa strigata flavorufus Cockerell, 1905, monobasic. Colletinae. Panzer, 1805, by original designation. [Anthidium]. Isotypic Androgynella Cockerell, 1911c:313. Type species: Megachile with Cerianthidium. Anthidiini. detersa Cockerell, 1910, by original designation. Megachilini. Anthidioma Pasteels, 1984:34. Type species: Anthidioma Andronicus Cresson, 1864:384. Type species: Andronicus chalicodomoides Pasteels, 1984, by original designation. cylindricus Cresson. 1864 (not Osmia cylindrica Giraud. Anthidiini. 1857) = Osmia spoliata Provancher, 1888, monobasic. Anthidium Fabricius, 1804:364. Type species: Apis manicata Osmiini. Linnaeus. 1758, by designation of Latreille. 1810:439. Anepicharis Moure, 1 945b:302. Type species: Epicharis dejeanii Anthidiini. Lepeletier, 1841, by original Centridini. designation. Anthidulum Michener. 1948a: 19. Type species: Dianthidium Angochlora Schrottky, 1901:213, incorrect subsequent spelling currant Schwarz, 1933, by original designation. (repeated) ior Angochlora Smith, 1853. Augochlorini. [Dianthidium]. Anthidiini.

Angochloropsis Schrottky. 1901:213, incorrect subsequent spell- Anthocharessa Gistel, 1850:82, unjustified replacement for ing for Augochloropsis Cockerell, 1897. [Angochlora]. Andrena Fabricius, 1775. Type species: Apis helvola Linnaeus, Augochlorini. 1758, autobasic. Andreninae. 8 Scientific Papers, Natural History Museum, The University of Kansas

Anthocopa Lepeletier and Serville, 1 825:3 14. Type species: Apis Aplumeiira Gray. 1832:575. Type species: Aphaneiira nifescens papaveris Latreille, 1799, monobasic. Isotypic with Gray. 1 832. monotypic. SeeAlphaneiira Gray, and Michener, Phylloloma and Furcosmia. Osmiini. 1995. Meliponini. Anthodioctes Holmberg, 1887b:36. Nomen nudum. Anthidiini. Aphelonomada Snelling. 1986:9. Type species: Noinada cruruiis Moure. 1960. by original designation. [Hypochrotaenia]. Anthodioctes Holmberg, 1903:435. Type species: Anthodioctes Nomadini. megachiloides Holmberg, 1903, by designation of Cockerell, 1969a:41. Anthidium 1927b:2. A. dasygasthmis Holmberg, 1903, designated as type Apiauthidiiim Pasteels. Type species: 1914. species by Isensee, 1927:375, was not an originally included apifonne Meade-Waldo, by original designation. Anthidiini. species name and is not a published name so far as known. Anthidiini. Apiaria Germar. 1 839:2 10. Type species: Apiaria dubia Germar, 1839. monobasic. Fossil. Anthoglossa Smith. 1 853: 16. Type species: Anthoglossa phimata

Smith, 1 853, monobasic. Colletinae. Apiants Rafinesque. 1815: 123, unjustified replacement for Apis

Linnaeus. 1 758. Linnaeus, 1758. Anthomegilla Marikovskaya, 1976:688. Type species: Type species: /4/j/.?/»e//(/era autobasic. Apini. Anthophoni arctica Morawitz, 1 883. by original designation. Anthophorini. Apicula Rafinesque. 1814:27. unjustified replacement for Apis Linnaeus. 1 758. Type species: Apis mellifera Linnaeus. 1 758, Anthophora Latreille, 1803:167, replacement for Podaliriiis = autobasic. Apini. Latreille, 1802. Type species: Apis pilipes Fabricius, 1775 Linnaeus, 1758:343, 574. Apis pliimipes Pallas, 1 772, designated by Commission Opin- Apis Type species: Apis mellifica Linnaeus, 1761 = A. Linnaeus, 1758, ion 151 (1944). Apis retusa Linnaeus, 1758, designated as mellifera by designation of Latreille, 1810:439. type species by Curtis. 1831:pl. 357, is not an originally in- Apini. cluded species. Anthophorini. ApistaSmxXh, 1861:148, not Hiibner, 1816. Type species: y4pwM

opalina Smith. 1 86 1 . monobasic. See Egapista. Dissoglottini. Anthophorisca Michener and Moure, 1957:433, Type species: Melissodes pygmaea Cresson, 1872, by original designation. Apomelissodes LaBerge. 1956:1 175. Type species: Melissodes [Exomalopsis]. Exomalopsini. fimbriata Cresson. 1878. by original designation. [Melissodes]. Eucerini. Anthophorites Heer, 1850:97. Type species: Anthophorites inellona Heer. 1850, designated by Cockerell. 1909d:315. Aporandrena Lanham. 1949:201. Type species: Andrena Fossil. coactipostica Viereck. 1917, by original designation. [Andrena], Andreninae. Anthophoroides Cockerell and Cockerell, 1901:48. Type spe- Moure, 1961:233. Trigona nebulata cies: Podalirius valloriim Cockerell, 1896, by original desig- Apotrigona Type species: Smith, 1854. nation. Anthophorini. by original designation. Meliponini. Apoxylocopa Hurd and Moure. 1963:226. Type species: Xylocopa Anthophorula Cockerell. 1 897d:44. Type species: Anthophorula litgubris Gerstaecker. 1857. by original designation. compactida Cockerell, 1897, monobasic. Exomalopsini. [Xylocopa]. Xylocopini. Anthrena Illiger. 1801:127, unjustified emendation oi' Aiuhrna Archeriades Peters. 1978a:337. species: Eriades laiTcitiis Fabricius, 1775. Andreninae. Type Friese, 1909, by original designation. Osmiini. Anthrenoides Ducke, 1907:368. Type species: Anthrenoides Archiandrena LaBerge. 1986:482. Type species: Andrena banksi alfkeni Ducke, 1907 = Protandrena meridionalis Schrottky. Malloch. 1917. by original designation. [Andrena]. 1906. monobasic. Panurginae. Andreninae. Anylaeiis Bridwell, 1919: 129. Type species: Nothylaeus aberrans Archianthidium Mavromoustakis, 1939:91. Type species: Bridwell. 1919. by original designation. [Nothylaeus]. Anthidium laticeps Morawitz, 1873. by original designation. Hylaeinae. Anthidiini. Aparatrigona Moure. 195Ia:60. species: Melipona Type Archihalictiis Pauly. 1984b: 132. Type species: Halictits jojfrei Ducke. 1916, impunctata by original designation. Benoist, 1962, by original designation. Halictini. [Paratrigona]. Meliponini. Archimegachile Alfken, 1933:56. Type species: Megachile Newman, 1834:404 footnote, Apathus unjustified replacement flavipes Spinola, 1838, by original designation. [Megachile j. for Psithynis Lepeletier. 1832. Type species: Apis nipestris Megachilini. Fabricius. 1793. autobasic. Bombini. Arctosmia Schmiedeknecht. 1885:21. Type species: Megachile Apeulaema Moure, 1950b: 184. Type species; Eulaema fasciata villosa Schenck, 1853 (not Apis villosa Fabricius. 1775) = = Lepeletier. 1841 Centris cingidata¥abr\c'ms. 1 804. by origi- Osmia plar\cera Gerstaecker, 1869, monobasic. [Osmia]. nal designation. [Eulaema]. Euglossini. Isotypic with Lithosmia. Osmiini.

Aphalictus Cockerell, 1930e:40. Type species: Parasphecodes Ardenthidium Pasteels, 1969a: 103. Type species: Anthidium brihiensis Cockerell. 1916. by original designation. ardens Smith, 1879, by original designation. [Anthidium]. [Parasphecodes]. Halictini. Anthidiini. Genus-Group Names of Bees

1984:132. Argalictiis Pesenko. 1984:348. Type species: Hylaeus senilis Atropium Pasteels. Type species: Megachile airopos Anthidiini. Eversmann. 1852. by original designation. [Halictus]. Smith. 1853. by original designation.

Halictini. Atrosamba Michener. 198 1 :65. Type species: Haplomelitta atra Michener. 1981. Argohesma Exley, 1969:528. Type species: Argohesma eremica by original designation. [Haplomelitta]. Exiey, 1969, by original designation. Euryglossinae. Dasypodinae. Audineta Ashmead. 1 899a:97, incorrect subsequent spelling for Argyropile Mitchell. 1934:308. Type species: Megachile Audinetia Lepeletier. 1 84 1 . Xylocopini. parallcia Smith. 1853. by original designation. [Megachile]. Megachilini. Audinetia Lepeletier. 1 84 1 :203. Type species: Apis latipes Drury, 1773, by designation of Sandhouse, 1943:529. [Xylocopa], Argyroselenis Robertson, 1903c:284. Type species: Triepeolus Isotypic with Platynopoda. Xylocopini. minimus Robertson. 1902. by original designation. Epeolini. Augandrena LaBerge, 1986:557. Type species: Andrena Arhysocehle Moure, 1948:335. Type species: Arhysuceble plumiscopa Timberlake, 1951, by original designation. xanthopoda Moure, 1948, by original designation. [Andrena]. Andreninae. Tapinotaspidini. Augochlora Smith. 1853:73. Type species: Halictus punts Say, Arhysosage Brethes, 1922:121. Type species: Arhysosage 1837, designated by Cockerell, 1923b:448. Augochlorini. johnsoni Brethes, 1922 - Camptopoeitm ochraceum Friese, Augochlorella Sandhouse, 1937:66. Type species: Augochlora 1908, monobasic. Isotypic with Riiiziella and Ruziapis by gratiosa Smith, 1853, by original designation. Augochlorini. synonymy. Panurginae. Augochlorodes Moure, 1958:53. Type species: Augocblorodes Ariphanarthm Moure, 195 lb: 137. Type species: Ariphanarthra turrifaciens Moure. 1958, by original designation. palpalis Moure, 195 1 . by original designation. Augochlorini. Augochlorini. Arogochila Michener. 1939:58. Type species: Ashmeadiella Augochloropsis Cockerell. 1 897b:4. Type species: Augochlora tiniberlakei Michener. 1936. by original designation. subignita Cockerell. 1897 = Augochlora ignila Smith. 1861, [Ashmeadiella]. Osniiini. by original designation. Augochlora spinolae Cockerell. 1900, Asaropoda Cockerell. 1926b:216. Type species: Saropoda designated by Schrottky. 1909b:481. was not originally in- bomhiformis Smith, 1854, by original designation. cluded. [Augochlora]. Augochlorini. Anthophorini. Auricularia Mehely, 1935:41, 147. Invalid because no type spe- Ascirteticu Moure and Michener, 1955b:260. Type species: cies was designated. Sandhouse"s (1943:529) view that Popov a and thus validated the name Eiicera herbsti Friese, 1906, by original designation. ( 1939) designated type species [Alloscirtetica]. Eucerini. is incorrect. [Prosopis]. Hylaeinae. Anthidiellum Ashmeadiella Cockerell, 1897c: 197. Type species: Heriades Ausanthidium Pasteels. 1969a:60. Type species: ausense 1934, opiintiae Cockerell, 1897, by original designation. Osmiini. Mavromoustakis. by original designation. Anthidiini. Ashtonipsithyrus Frison, 1927:69. Type species: Apathus ashtoni Australictus Michener. 1965a: 165. Halictus Cresson. 1 864, by original designation. [Psithyrus]. Bombini. Type species: peraustralis Cockerell. 1904. by original designation. Asianthidium Popov, 1950a:315. Type species: Anthidium [Lasioglossum]. Halictini. glasunovi Morawitz. 1894. by original designation. Austrandrena Cockerell. 1906c:37. Andrena [Meganthidium]. Anthidiini. Type species: modesta Smith. 1 879. by original designation. [Protandrena]. Aspidosnua Brauns. 1926:208. Type species: Osmia arnoldi Panurginae. Brauns, 1926. monobasic. [Osmia^j. Osmiini. Austrevylaeus Michener, 1965a: 170. Type species: Halictus Astewnomada Broemeling. 1988:336. Type species: Nomada sordidus Smith, 1853, by original designation. additcta Cresson, 1878, by original designation. [Nomada]. [Lasioglossum]. Halictini. Nomadini. Austrochile Michener, 1965a:202. Type species: Megachile Atopeucera Tkalcu, 1984b:71. Type species: Eiicera seminuda resinifera Meade-Waldo, 1915, by original designation. Brulle. 1832. [Eucera]. Eucerini. by original designation. [Chalicodoma]. Megachilini. Cockerell, 1935a:50. Type species: Osmia thodonta Atoposmia Aiistrodioxys Cockerell. 1916a:432. Type species: Austrodioxys monobasic. Osniiini. Cockerell. 1935. [Osmia]. thomasi Cockerell, 1916 = Caenoprosopis crabronina Atrocinctohfambus] Skorikov. 1933b:244. nomen nudum. Holmberg, 1887, monobasic. Isotypic with Caenoprosopis Bombini. by synonymy. Caenoprosopidini.

Atmdufourea Ebmer. 1984:360. Type species: Rophites atrata Austromegachile Mitchell, 1943:666. Type species: Megachile Warncke. 1979. by original designation. [Dufourea]. moittezuma Cresson, 1878, by original designation. Rophitinae. [Megachile]. Megachilini. Atronomioides Pesenko. 1983:186. Type species: Ceylalictus Austronomia Michener, 1965a: 156. Type species; Nomia warnckei Pesenko. 1983. by original designation. australica Smith, 1875. by original designation. [Nomia]. [Ceylalictus]. Nomioidinae. Nomiinae. 10 Scientific Papers, Natural History Museum, The University of Kansas

Moure, Austropanurgus Tom, 1980:209. Type species: Austropanurgus Belopria Moure, 1956:305. Type species: Belopria zonata Colletinae. punctatus ToTO. 1980, monobasic. Panurginae. 1956, by original designation. Austropeponapis Hurd and Linsley, 1970:21. Type species: Benanthis Pasteels, 1969a:61. Type species: Anthidium Tetralonia melonis Friese, 1925, by original designation. madagascariensis Benoist. 1963, by original designation. [Peponapis]. Eucerini. Antiiidiini. 1961:195. cassiae Austroplebeia Moure, Type species: Trigona Berna Friese, 1911c:668. Type species: Bema africana Friese, = Cockerell, 1910, by original designation. Meliponini. 1911 (combination as though Berna were a genus) Austrosphecodes Michener, 1978b:327. Type species: Sphecodes Megachile africanibia Strand. 1912. monobasic. [Megachile]. chiknsis Spinola, 1 85 1 . by original designation. [Sphecodes], Megachilini. Halictini. Betheliella Cockerell, 1924b: 169. Type species: Betheliella Austrostelis Michener and Griswold, 1994b:676. Type species: calocharti CockereW, 1924, monobasic. Rophitinae. Stelis aliena Cockerell, 1919, by original designation. Biareolina Dours, 1873:288. Type species: Andrena neglecta [Hoplostelis]. Anthidiini. Dours, 1873 = Andrena lagopus Latreille, 1809, designated Autochelostoma Sladen, 1916a:270. Type species: by Sandhouse, 1943:530. Andreninae. Autochelostoma canadensis Sladen, 1916 = Alcidamea brevicornis vj'nh Alcidamea Biastes Panzer, 1806:239. Type species: Tiphia producta Cresson, 1 864, monobasic. Isotypic Panzer, 1798, monobasic. with Rhineta by synonymy. by synonymy. Osmiini. Isotypic Biastini. Avandrena Wamcke, 1968:31. Type species: Andrena avara Pasites Warncke, 1967, by original designation. [Andrena]. Biastoides Schenck, 1873:252. Type species: punctatus Andreninae. Schenck, 1870 = Phileremus emarginatus Schenck, 1853, monobasic. Biastini. Avpanurgus Warncke, 1972:70. Type species: Panurgus flavofasciatus Warncke, 1972, by original designation. Bicolletes Friese. 1908b:341 (p. 11 in reprint). Type species: [Panurgus]. Panurginae. Bicolletes neotropica Friese, 1908, by designation of Axestotrigona Moure, 1961:237. Type species: Melipona Cockerell, 1915:342. Colletinae. Lepeletier, 1836, by original designation. ferniginea Bicornelia Friese, 1899a:239. Type species: Bicornelia serrata Meliponini. Friese, 1899, monobasic. Dissoglottini. Axillanthidium Pasteels, 1969a:39. Type species: Axillanthidium Biglossa Friese, 1906c:374. Type species: Biglossa thoracica axiUare Pasteels, 1969 = Anthidium diicale Morawitz, 1876, Friese, 1906, by designation of Cockerell, 1914:328. by original designation. Antiiidiini. Colletinae. Aztecanthidium Michener and Ordway, 1964:70. Type species: Moure, 1948:313. Biglossa chalybaea Aztecanthidium xochipillium Michener and Ordway, 1964, Biglossidia Type species: Friese, 1906, designation. Colletinae. by original designation. Anthidiini. by original nasalis Westwood. Baana Sandhouse, 1943:530, replacement for Ctenopoda Ma, Biluna Ma, 1938:276. Type species: Xylocopa 1938. Type ?,^tc\e:^:Apisfenestmta Fabricius, 1798, autobasic 1842, by original designation. [Xylocopa]. Xylocopini. and by original designation. Xylocopini. Binghamiella Cockerell, 1907c:235. Type species: Sphecodes Baeocolletes Michener, 1965a:70. Type species: Leioproctiis antipodes Smith, 1853. monobasic. Colletinae. calcaratus Michener. 1965, by original designation. Birkmania Viereck, 1909:50. Type species: Birkmania Colletinae. [Leioproctus], andrenoides Viereck, 1909 = Panurginus polytrichus Barbata 1935:32, 138, not 1797. Mehely, Humphrey, Type spe- Cockerell, 1909, by original designation. Panurginae. cies: Prosopis friesei M^tn, 1904, monobasic. [Prosopis]. Bluethgenia Pesenko, 1986a: 136. Type species: Halictus dynastes See Mehelya and Mehelyana. Hylaeinae. Bingham, 1898, by original designation. [Lasioglossum]. Bathanthidium Mavromoustakis, 1953:837. Type species: Halictini. Dianthidium bifoveolatum Alfken, 1937, by original desig- Bombias Robertson, 1903a: 176. Bombias nation. [Dianthidium]. Antiiidiini. Type species: auricomus Robertson, 1903 = B. nevadensis auricomus Bekilia Benoist, 1962:220. Type species: Bekilia mimetica (Robertson), by original designation. Isotypic with Benoist, 1962, by original designation. Osmiini? Boopobombus. Bombini. Belandrena Ribble, 1968b:221. Type species: Andrena Bomboixylocopa Maa, 1939:155. Type species: Xylocopa nemophilae Ribble, 1968, by original designation. [Andrena]. Andreninae. bomboides Smith, 1879, by original designation. [Xylocopa]. Xylocopini. Bellanthidiiini Pasteels, 1969a:38. Type species: Anthidium 1879b:370. Melecta infuscatum Erichson, 1835, by original designation. Bombomelecta Patton, Type species: Anthidiini. thoracica Cresson, 1876, monobasic. Melectini Genus-Group Names of Bees 11

Bomhus Latreille. 1802a:437. Type species: Apis terreslris BrasUagapostemon Moure and Sakagami, 1984:4. Type species: 191 1, monobasic. Linnaeus, 1 758, monobasic. A subsequent designation is Apis Pseudagapostemon fluminensis SchroUky, Halictini. inuscorum Linnaeus, 1758, by Westwood. 1840a:86; it was [Pseudagapostemon|. included with Bremus not an originally species. Isotypic Braunsapis Michener, 1969:290. Type species: Allodapefacialis Jurine, Bremus Panzer, Leucobombus. and Terrestribomhiis. Gerstaecher. 1857, by original designation. [Allodape]. Bombini. Allodapini. 1856:28. Bowbiisoides Bombusoides Motschulsky. Type species: Bremus Jurine, 1 80 1 : 1 64. Type species: Apis terrestris Linnaeus, inengei Motschulsky, 1856. monobasic. Fossil. 1758, by designation of Morice and Durrant, 1915:428. In- validated Commission 135, 1939. with Boopobombtis Prison, 1927:62. Type species: Bombias by Opinion Isotypic and Terrestribombus. aiiricomits Robertson, 1903 = B. nevadensis auricomus Bremus Panzer, Bombus, Leucobombus, Bombini. (Robertson), by designation of Williams, 1994:339. A sec-

tional name that, according to the Code, 3rd ed.. Art. 10(e), Bremus Panzer, 1 804a: 1 9. Type species: Apis terrestris Linnaeus, must be treated as a subgenus. Isotypic with Bombias. 1758. by designation of Benson, Ferriere and Richards, [Bombus], Bombini. 1937:93. Sandhouse's ( 1943) subsequent designation oiApis agrorum Fabricius, 1787, as the type species is invalid. Panzer Boreocoelioxys Mitchell, 1973:37. Type species: Coelioxys (1804a, dated 1801 by Benson, Ferriere and Richards) did rufitarsus Smith, 1854, by original designation. [Coelioxys]. not include terrestris. but he did include Bremus faciatus Megachilini. A/?/'.v Panzer. Because Benson, Ferriere and Richards list that name Ikudome, 1991 :790. Type species: Hylaeus macilentus Boieopsis as a synonym of Apis terrestris. the latter was available for Ikudome, 1989, original designation. [HylaeusJ. by designation as type species of Bremus Panzer [see the Code, Hylaeinae. in 3rd ed.. Art. 69(a)(v)l. I follow Sandhouse ( 1943) the dat- Bothraiiiliiditim Moure, I947a:23. Type species: Bothranthidiitm ing of Bremus Panzer; the real date of publication could have Iciiiroi Moure, 1947. by original designation. Anthidiini. been from 1 801 to 1 804. Isotypic with Bombus. Bremus Jurine, Leucobombus, and Terrestribombus. Bombini. Brachyandrena Pittioni, 1948:54. Type species: Andrena colletifoniiis Morawitz, 1874, by original designation. Bremus Kirby, 1837:272 footnote, not Panzer, 1804. Type spe- of [Andrena.] Andreninae. cies: Apis rupestris Fabricius, 1793, by designation Milliron, 1961:59. Not replaced because a junior synonym of Brachycephalapis Viereck, 1909:47. Type species: Melitta Psith\rus. Isotypic with Psithyrus. Bombini. califomica Viereck, 1909, by original designation. Melittinae. Brevineura Michener, 1965a:224. Type species: Exoneura Brachycephalibombits Williams, 1985:247. Type species: concinnula Cockerell, 1913, by original designation. Bombus brachycepludus Handlirsch. 1888, by original des- [Exoneura]. Allodapini. ignation. [Bombus]. Bombini. Bythandretia Lanham, 1 950: 1 40, replacement for Cryptandrena Biachyglossa Friese. 1922a:577, not Boisduval, 1829. Type spe- Lanham, 1949. Type species: Amlremi carlini Cockerell, 1901 , cies: Brachyglossa rufocaerulea Friese, 1922, monobasic. See autobasic. [Andrena]. Andreninae. Brachyglossula. Colletinae. Bytinskia Mavromoustakis, 1954:269. Type species: Bytinskia Brachyglossula Hedicke, 1922:427, replacement for erythrogastra Mavromoustakis, 1954, by original designa- Friese, 1922. Brachyglossa Type species: Brachyglossa tion. Osmiini. riifocaeridea Friese, 1922, autobasic. Colletinae. Cacosoma Smith, 1879:39, not Felder, 1874. Type species: Brachyhesma Michener, 1965a: 1 12. Type species: Euryglossina Cacosoma (//.sYr;/(*r Smith, 1 879, by original designation. See Cockerell, 1913, siiiphurella by original designation. Neocorymira. Augochlorini. Euryglossinae. Cadeguala Reed, 1892:234. Type species: Colletes chilensis 1939a:458. Melecta = Brachymelecta Linsley, Type species: Spinola, 1851 Colletes occidentalis Ha\ida.y. 1836, by des- miicida Cresson, 1879, Meiectini. by original designation. ignation of Sandhouse, 1943:532. Diphaglossinae. 1956:926. Eucera Brachymelissodes LaBerge, Type species: Cadegualina Michener, 1986b: 187. Type species: Bicornelia cressonii Dalla Torre, 1896, by original designation. andina Friese, 1925, by original designation. Diphaglossinae. [Melissodes]. Eucerini. Caenaugochlora Michener, 1954:76. Type species: 272. Brachymmiada Holmberg, I886b:233. 239. Type species: Caencmgochlora macswaini Michener, 1 954. by original des- Brachynomada urgeiitina Holmberg, 1886, designated by ignation. Augochlorini. Sandhouse, 1943:531. Brachynomadini. Caenohalictus Cameron, I903a:23l. Type species: Brachyspatiilarietta Pittioni, 1950:79. Type species: Spatulariella Caenohalictus trichiothalmus Cameron, 1903, monobasic. helemie Pittioni, 1950, by original designation. [Prosopis]. Halictini.

Hylaeinae. Caenonomada k?hmtad, 1 899a:68. Type species: Caemmomada with Branthidiiim Pasteels, I969a:88. Type species: Antliidiwu hraitnsi fen(H<'r; Ashmead, 1899, by original designation. Isotypic Friese, 1904, by original designation. Anthidiini. Chacoana by synonymy. Tapinotaspidini. 12 Scientific Papers, Natural History Museum, The University of Kansas

Caenoprosopina Roig-Alsina, 1987:312. Type species: Callochlora Moure, 1964b:269. not Packard. 1864. Type spe-

cies: chloris 1 1 Caenoprosopina holmbergi Roig-Alsina. 1987, by original Halictus Spinola. 85 , by original designation. designation. Caenoprosopidini. See Callistochlora. Augochlorini.

Caenoprosopis Holmberg, 1886b:235, no included species; Callocolletes Michener, 1965a:80. Type species: Trichocolletes 1887a:22, one included species. Type species: Caenoprosopis pulcherrimus Michener. 1965, by original designation. crabroniim Holmberg. 1887. first included species. Isotypic [Trichocolletes]. Colletinae. with Aiistrodioxys by synonymy. Caenoprosopidini. Ccdlohesma Michener, 1965a:95. Type species: Euryglossa Caerulosmia Zanden, 1988:123, nomen nudum. calliopsifonnis Cockerell, 1905, by original designation.

Caerulosmia Zanden 1989:83. Type species: Osmia gallanim [Euryglossa]. Euryglossinae. Spinola, 1808, by original designation. [Osmia]. Osmiini. Callomacrotera Timberlake, 1954:413. Type species: Perdita ;««nY//)M Timberlake. 1954, Caesarea Friese, 191 Id: 142. Type species: Ammobates depressa by original designation. [Perdita]. Friese, 1911, by designation of Sandhouse, 1943:532. Panurginae.

[Ammobates]. Ammobatini. Callomegachile Michener, 1 962:2 1 . Type species: Chalicodoma Michener. 1962. Calchalictits Wamcke. 1975b:99. Type species: Apis caheata mystaceana by original designation. Scopoli, 1763. by original designation. [Halictus]. Halictini. [Chalicodoma]. Megachilini.

Callomelecta Cockerell. 1926c:621 . Callomelecta Caliendra Gistel 1848:viii, unjustified replacement for Type species: Cockerell, 1926, Chrysantheda Perty. 1 833. Type species: Chnsantheda niiida pendlchuryi by original designation. Melectini. Perty, 1^33= Apis dentaia Linnaeus. 1758, autobasic. Isotypic with Exaerete by synonymy. Euglossini. Callomelitta Smith, 1853:85. Type species: Callomelitta picta Smith, 1853. monobasic. Colletinae. Callalictiis Michener, 1965a: 170. Type species: Parasphecodes tooloomensis Cockerell. 1929, by original designation. Callonychium Brethes, 1922:120. Type species: Callonychium [Lasioglossum]. Halictini. argentinum Brethes, 1922, monobasic. Panurginae. Callandreiui Cockerell, 1898b: 186. Type species: Pamirgus Calloprosopis Snelling. 1985:27. Type species: Hylaeus manifestus Fox, 1894, monobasic. Andreninae. magnificus Cockerell, 1942, by original designation.

Callanthidium Cockerell, 1925a:365. Type species: Antliidiiini Hylaeinae. illustre Cresson. 1879, by original designation. Anthidiini. Callorhiza Hirashima, 1989:2. Type species: Prosopis apicatus Smith. 1863, [PalaeorhizaJ. Calleupetersia Cockerell, 1938:329. Type species: Halictus by original designation. lasureus Friese, 1910, by original designation. Halictini. Hylaeinae. Friese. 1909b: 182. CaUimelissodes LaBerge, 1961:294. Type species: Melissodes Callosphecodes Type species: Sphecodes ralunensis Friese. 1909, monobasic. Halictini. lupina Cresson, 1878, by original designation. [Melissodes]. [Sphecodes]. Eucerini. Calloxylocopa Hurd and Moure, 1963:142. Type species: tenuata Smith. 1874. Callinomada Rodeck, 1945: 181. Type species: Nomada antonita Xylocopa by original designation. Cockerell. 1909, by original designation. [Nomada]. [Xylocopa]. Xylocopinl. Nomadini. Calohoplitis TkaXcu, 1995:129. Type species: Osmia laboriosa Smith, 1878, Osmiini. Calliopsima Shinn. 1967:834. Type species: Calliopsis rozeni by original designation. [Hoplitis], Shinn. 1967, by original designation. [Calliopsis]. Calomelissa Hirashima and LaBerge, 1963, in Hirashima,

Panurginae. 1963:241 . Type species: Andrena prostomias Perez, 1 805, by Andreninae. Calliopsis Smith, 1853:128. Type species: Calliopsis original designation. [Andrena]. andreniformis Smith, 1853, by designation of Calospiloma Brethes, 1909b:68 footnote. Type species: Epeolus Ashmead, 1899a: 85. Panurginae. viperinus Holmberg. 1 886. by original designation. Isotypic with with Callistochlora Michener. new name, replacement for Palinepeolus: isotypic Isepeolus by synonymy. Callochlora Moure. 1964b:269. not Packard, 1864. Type spe- Isepeolini cies: Halictus chloris Spinola. 1851, autobasic. When in- Calyptapis Cockerell. 1906a:41. Type species: Calyptapis formed of the homonymy. Padre J. S. Moure suggested the florissantensis Cockerell, 1906, monobasic. Fossil.

name here but asked that I it. proposed publish Augochlorini. Camargoia Moure. 1989a:72. Type species: Camargoia Callobombus Dalla Torre, 1896:503, unjustified emendation of camargoi Moure, 1989, by original designation. Meliponini. Kallobombus Dalla Torre. 1880. Bombini. [Bombus], Campanularia Mehely, 1935:157. not Lamarck, 1816. Invalid because no also Calloceratina Cockerell, 1924d:77. Type species: Ceraiina type species was designated; homonym. = See amabilis Cockerell, 1897 C. exima Smith. 1862, by origi- [Prosopis]. Paraprosopis. Hylaeinae. nal designation. [Ceratina]. Ceratinini. Camptopaeum Spinola, 1851:192. unjustified emendation of 1843. Callochile Michener, 1962:27. Type species: Megachile Camptopoeum Spinola. Panurginae. = ustulatiformis Cockerell, 1910 Apis mystacea Fabricius, Camptopoeum Spinola, 1843:139. Type species: Prosopis fron- 1775, by original designation. [Megachile]. Megachilini. talis Fabricius, 1804, by original designation. Panurginae. Genus-Group Names of Bees 13

Campyloi^aster DouTS. 1873:286. Type species: Campylogaster Caupolicanoides Michener, 1966a:725. Type species: = fulvo-cntstatus Dours, 1873 Audrena erheri Morawitz, Caupolicana pubescens Smith. 1879. by original designation. 1871. designated by Sandhouse, 1943:534. Andreninae. Caupoiicanini.

Caiiephoni Fnese. 1908b:94.not Hiibner, 1 822, replacement for Ceblurgus Urban and Moure. 1993: 102. Type species: Ceblurgus Corhiciila Friese, 1908. Type species: Corhicula apifonyns longipalpis Urban and Moure, 1993, by original designation. Friese, 1908. autobasic. See Caiiephonilci. Eucerini. Rophitinae.

Canephontki Jorgensen. 1909:212. replacement for Canephora Celetandrena LaBerge and Hurd, 1965:188. Type species: Friese. 1908. Type species: Corbicula apiformis Friese, 1908, Andrena vinnula LaBerge and Hurd, 1965, by original desig- autobasic. Eucerini. nation. [Andrena]. Andreninae. 1969a:85. Anthidium Capanthidium Pasteels, Type species: Celetrigona Moure, 1950a:246. Type species: Trigona "capicole Friese," lapsus for capicola Brauns, 1 905, by origi- longicornis Friese. 1903, by original designation. nal Antliidiini. designation. [Hypotrigona]. Meliponini. 191 la:672. braimsiana Capicola Friese. Type species: Capicola Cellaria Friese, 1913a:575, not Ellis and Solander, 1786. Type Friese, 1911, monobasic. Dasypodinae. species: Nomioides arnoldi Friese, 1913 = Nomioides CapicoloidesMidncntv. 1981:83. Type species: Capicola aliciae somalica Magretti, 1899, monobasic. [Nomioides]. See Cockerel], 1932, by original designation. [Capicola]. Cellariella. Nomioidinae. Dasypodinae. Cellariella Strand. 1926:53. replacement for Cellaria Friese, = Caposmia Peters, 1984:378. Type species: Osiuia brauiisi Pe- 1913. Type species: Nomioides arnoldi Friese. 1913 ters, 1984, by original designation. [Osmia]. Osmiini. Nomioides somalica Magretti. 1899, autobasic. [Nomioides]. Nomioidinae. Carandrena Warncke, 1968:94. Type species: Audrena aeriuifrons Dours, 1873, by original designation. [Andrena]. Cemolobus Robertson, 1902b:324. Type species: Xenoglossa Andreninae. ipomoeae Robertson, 1 89 1 , by original designation. Eucerini.

Caiwithophora Brooks, 1988:470. Type species: Autlwphora Centrias Robertson, 1903d: 174, 176. Type species: Nomada diifoiiiii Lepeletier, 1841, by original designation. erigeronis Robertson, 1897, by original designation. [Anthophora]. Anthophorini. Nomadini. Carimmtludium Pasteels. 1969a:42. Type species: Megachile Centris Fabricius, 1 804:354. Type species: Apis haemorrhoidalis cariniveiitris Friese, 1904, Anthidiini. by original designation. Fabricius, 1775, designation by Commission Opinion 567 Carinapis Stage. 1981. in Michener, 1981:98. Type species: (1959). The designation o^ Apis dimidiata Fabricius, 1793, a Hesperapis carinata Stevens, 1919, by original designation. euglossine bee, by Blanchard, 1 840:405, and the designation [Hesperapis], Dasypodinae. of Apis hiriipes Fabricius, 1793 =Apis rustica Olivier, 1789, Blanchard, 1849:pl. 128 bis (1846), are invalidated. Carinella Pasteels. 1965:447. not Johnston, 1833. Type species: by with Hemisia. Centridini. Megachile tonida Smith. 1853, by original designation. Isotypic [Chalicodoma]. See Cannula. Megachilini. Centrosmia Robertson. 1903a: 165. 166. Type species: Osmia Cresson. 1864, Osmiini. Carinellitm Pasteels, 1969a:80. Type species: Anthidium bucepliala by original designation. ochrognathum Alfken. 1932, by original designation. Cephalapis Cockerel], 1910e:23. Type species: Chelostoma [Pseudoanthidium]. Anthidiini. jacintanum Cockerel], 1910, monobasic. [Chelostoma]. Osmiini. Carinowphites Warncke. 1979:136. Type species: Dufourea

rufiventris Friese. 1898. by original designation. [Rophites]. Cephalictoides Cockerel], 1 924c:244. Type species: Halictoides Rophitinae. paradoxus Morawitz. 1867. by original designation. [Halictoides]. Cannula Michener, McGinley, and Danforth, 1 994: 1 74, replace- Rophitinae. ment for Carinella Pasteels. 1965. Type species: Megachile Cephalocolletes Michener. 1989:656. Type species: Biglossa torrida Smith, 1853, autobasic. [Megachile]. Megachilini. laticeps Friese. 1906. by original designation. [Leioproctus]. Carloticola Moure and Urban. 1990:90. Type species: Colletinae. Dianthidium paraguayense Schrottky. 1908, by original des- Cephalosmia Sladen, 1916a:270. Type species: Osmia

Anthidiini. = 1 ignation. armaticeps Cresson, 1 878 Osmia montana Cresson, 864, monobasic. Osmiini. Catoceratina Vecht. 1952:30. Type species: Ceratina perforatrix Ceratinini. Smith, 1879, by original designation. [Ceratina]. Cephalotrigona Schwarz, 1940:10. Type species: Trigona Smith. 1854. Caupolicana Spinola, 1 85 1 :2 1 2. Type species: Caupolicana gayi capitata by original designation. [Trigona]. Spinola, 1851, designated by Sandhouse, 1943:534. IVleliponini. Caupoiicanini. Cephalurgus Moure and Lucas de Oliveira, 1962:2. Type spe- Caupolicania Schulz. 1906:238, unjustified emendation of cies: Cephalurgus anomalus Moure and Lucas de Oliveira, Caupolicana Spinola, 1851. Caupoiicanini. 1962, by original designation. Panurginae. 14 SciENTinc Papers, Natural History Museum, The University of Kansas

Cephalylaeus Michener, 1942:273. Type species: Prosopis basa- Ceroliopoeum Ruz, 1991:237. Type species: Camptopoeum lts Smith. 1853. by original designation. [Hylaeus]. laetum Vachal, 1909, by original designation. [Calliopsis]. Hylaeinae. Panurginae. Cephen Robertson, 1903d: 174, 176. Type species: Nomada Cestella Pasteels, 1965:547. Type species: Megachile cestifera texaiui Cresson, 1872, by original designation. Nomadini. Benoist, 1954, by original designation. [Chalicodoma]. Megachilini. Cephylaeus Moure, 1972:280. Type species: Hylaeus larocai Moure, 1972, by original designation. [Hylaeus]. Hylaeinae. Ceylalictus Strand, 1913a:137. Type species: Halictus horni Strand, 1913, monobasic. [Halictus]. Nomioidinae. Cemheriades Tkalcu.l984a:5. Type species: Archeriades petersi Tkalcu, 1984, by original designation. [Archeriades]. Osmiini. Ceylonicola Friese. 1918:501. Type species: Ceylonicola atra Friese, 1918 = Siuiila bidentata Cameron, 1898, by designa- Ceraplastes Gistel, 1848:x, unjustified replacement for Stelis tion of Sandhouse, 1943:536. Isotypic with Sudila by syn- Panzer, 1806. Type species: Apis aterrima Panzer, 1798, not Halictini. Christ. 1791 = Apis pimctidatissima Kirby, 1802, autobasic. onymy. Isotypic with Gyiuniis. Stelis. Trachusa iurine. and Gyrodroma Chacoana Holmberg, 1887b: 225, nomen nudum. Klug. Anthidiini. Tapinotaspidini.

Ceratalictiis Moure, 1943b:463. Type species: Oxystoglossa tlieia Chacoana Holmberg, 1903:432. Type species: Chacoana = Schrottky, 191 1, by original designation. Augochlorini. melanoxantha Holmberg, 1903 Caenonomada bruneri Ashmead, 1899, monobasic. Isotypic with Caenonomada by Ceraiias Robertson, 1 903a: 1 72, not Kroyer, 1 845. Type species: Tapinotaspidini. Megachile pugnatiis Say, 1837, by original designation. See synonymy. Sayapis. Isotypic with Gnatliocera by synonymy. Chaetalictus Michener, 1978a:509. Type species: Halictus Megachilini. pearstonensis Cameron, 1905, by original designation. [Patellapis]. Halictini. Ceratina Latreille, 1 802b:380. Type species: Hylaeus albilabris Fabricius, 1793 = Apis cucurbitina Rossi, 1792, monobasic. Chaetochile Mitchell, 1980:63. Type species: Cressoniella Westwood, 1840a:85, designated /4/j;'.yrofn

and Blanchard, 1840:409, designated A/?/.s callosa Fabricius, [Cressoniella]. Megachilini. 1794, as the Placed on Official List of Generic type species. Chaetohesma Exley, 1978:373. Type species: Chaetohesma Names in 1011 of the Commission 1 Zoology, Opinion ( 973). tuberculata Exley. 1978, by original designation. with Claviceni. Ceratinini. Isotypic Euryglossinae. Ceratinidia Cockerell and Porter, 1899:406. Type species: Chaetostetha Michener, 1942:281. Type species: Exomalopsis Ceratina hieroglypluca Smith, 1854, by original designation. pyropyga Friese, 1925, by original designation. Ceratinini. [Ceratina]. Tapinotaspidini. Ceratinula 1941a:78. Ceratina lucidula Moure, Type species: Chaeturginus Lucas de Oliveira and Moure, 1963:575. Type Smith. 1854, Ceratinini. by original designation. species: Rhophitulus testaceus Ducke, 1907, by original des- Ceratocolletes Michener, 1965a:63. Type species: ignation. Panurginae. antennatus Smith, 1879, Laniprocolletes by original desig- Chalcobombus Cockerell. 1908d:326. Type species: nation. Colletinae. [Leioproctus]. Chalcobombus hiiinilis Cockerell, 1908, designated by Ceratoinonia Michener, 1981:20. Type species: Ceratonwnia Cockerell, 1909e:l 1. Fossil. rozenoruin Michener, 1981, by original designation. Chalcosmia Schmiedeknecht. 1885:20. Type species: Apis = Meganomiinae. fulviventris Panzer, 1798 (not Scopoli, 1763) Apis niveata Ceratopsithyrus Pittioni, 1949:270. Type species: Psithyrus Fabricius, 1804, by designation of Sandhouse, 1939:13. klapperichi Pittioni, 1949 - Psithyrus cornutus Prison, 1933. [Osmia]. Osmiini. monobasic. Bombini. [Psithyrus]. Chalepogenoides Michener, 1942:279. Type species: Ceratorhiza Hirashima, 1978:81. Type species: Palaeorhiza Chalepogenus leucostoma Cockerell, 1923, by original des- conica Michener, 1965, by original designation. [Palaeorhiza]. ignation. Tapinotaspidini. Hylaeinae. Chalepogenus Holmberg, 1903:416. Type species: Chalepogenus Ceratosmia Thomson, 1872:232. Type species: Apis bicornis incertus Holmberg, 1903 = Tetrapedia muelleri Friese, 1899, = Linnaeus, 1758 Apis rufa Linnaeus, 1758, designated by by designation of Sandhouse, 1943:537. Sandhouse consid- Sandhouse,1939: 9. [Osmia]. Isotypic with Osniia and ered Chalepogenus to be monobasic, but Holmberg named Arnblys; isotypic with Pachyosmia by synonymy. Osmiini. two "varieties'" of C. incertus, and thus included three taxa. with Cercorhiza Hirashima, 1982b:88. Type species: Palaeorhiza Isotypic Desmotetrapedia by synonymy. gressittorum Hirashima, 1975, by original designation. Tapinotaspidini.

[Palaeorhiza]. Hylaeinae. Chalicodoma Lepeletier, 1 84 1 :309. Type species: Apis muraria muraria Cerianthidium Friese, 1923:304. Type species: Trachusa strigata Olivier, 1789 (as Xylocopa Fabricius, 1804) =Apis Panzer, 1805, by designation of Cockerell, 1925a:361. parietina Fourcroy, 1785, by designation of Girard, 1879:778. [Anthidium]. Isotypic with Anthidielluni. Anthidiini. Megachilini. Genus-Group Names of Bees 15

Chalicodomoides Michener, 1962:24. Type species: Megachile Chlerogella Michener, 1954:75. Type species: Chlerogella aethiops Smith, 1853. by original designation. [Chalicodoma]. elongaticeps Michener, 1954, by original designation. Megachilini. Augochlorini. Chariiandrena Hedicke, 1933:210. Nomada Type species: Chloralictus Robertson, 1902c:245, 248. Type species: Halictus Fabricius, 1775, by original luittorfiana designation. cressoni Robertson, 1890, by original designation. Halictini. [Andrena]. Andreninae. Chlorandrena Perez, 1890:172. Type species: Andrena humilis Chuuhindrena LaBerge, 1964:314. Type species: Andrena Imhoff, 1832, by designation of Hedicke, 1933:211. Cockerell, 1900, portenie by original designation. [Andrena]. Andreninae. Andreninae. [Andrena]. Chloranthidiellum Mavromoustakis, 1963b:491. Type species: Cheesmania Hirashima, 1981b:27. Type species: Palaeorhiza Anthidium flavescens Friese, 1925, by original designation. amahilis Hirashima, 1981, by original designation. [Anthidiellum]. Anthidiini. [Palaeorhiza]. Hylaeinae. Chlorunthidium Pasteels, 1969a:48, for Chloranthidiellum Chelostoma Latreille, 1809:161. Type species: Apis maxUlosa lapsus Linnaeus, 1767 =Apw/?on50/?i/7w Linnaeus, 1758, monoba- Mavromoustakis, 1963. [Anthidiellum]. Anthidiini. sic. Osmiini. Chloroceratina Cockerell, 191 8a: 143. Type species: Ceratina Chelostomoda Michener, 1962:24. Type species: Megachile cyanura Cockerell, 1918, by original designation. [CeratinaJ. spissula parvula Strand, 1913 = M. spissula Cockerell, 1911, Ceratinini. by original designation. [Chalicodoma]. Megachilini. Chlorosmia Sladen, 1916a:270. Type species: Osmia fulgida Chelostonwidella Snelling, 1990:36. Type species: Megachile Cresson, 1864, monobasic. Osmiini. spinotulata Mitchell, 1934, by original designation. Chromobombus Dalla Torre, 1880:40. Type species: Apis [Chalicodoma]. Megachilini. muscorum Linnaeus, 1758, by designation of Sandhouse, Chelostomoides Robertson, 1901:231. Type species: Megachile 1943:538. [Bombus]. Bombini. rufimanns Robertson, 1891 = Chelostoma rugifrons Smith, Hedicke, 1933:211. 1854, monobasic. Megachilini. Chrysandrena Type species: Apis fidvago 1 1 Christ, 79 , by original designation. [Andrena]. Andreninae. Chelosiomopsis Cockerell, 1925b:205. Type species: Chelynia 1833:147. ruhifloris Cockerell, 1898, by original designation. Isotypic Chrysantheda Perty, Type species: Chrysantheda - with Raphidostoma by synonymy. Osmiini. nitida Perty, 1833 Apis dentata Linnaeus, 1758, monobasic. Isotypic with Caliendra; isotypic with Exaerete by syn- Chelynia Provancher, 1888:322. Type species: Chelynia labiata Provancher, 1888, monobasic. Anthidiini. onymy. Euryglossini. Michener, 1965a:71. Paracolletes Chenosmia Sinha, 1958:233. Type species: Osmia penstemanis Chrysocolletes Type species: moretonianus Cockerell, 1906, by original designation. Osmiini. Cockerell, 1905, by original designation. [Leioproctus]. Colletinae. Chilalictiis Michener, 1965a: 174. Type species: Halictus subinclinans Cockerell, 1915, by original designation. Chrysopheon Titus, 1901:256. Type species: Chrysopheon [Lasioglossum]. Halictini. aurifuscusTiXu^, 1901, monobasic. Dioxyini. Chilicola 1851:210. Chilicola rubriventris Spinola, Type species: Chrysosarus Mitchell, 1943:664. Type species: Megachile 1851, of Sandhouse, 1943:537. Spinola, by designation guaranitica Schrottky, 1908, by original designation. Xeromelissinae. [Megachile]. Megachilini. Chilicolletes Michener, 1989:640. Type species: Leioproctus Cilissa Leach, 1815: 155. Type species: Andrena haemorrhoidalis delahozHTom, 1973, by original designation. [Leioproctus]. Fabricius, 1775, by designation of Westwood, 1840a:84. Colletinae. Melitta tricincta Kirby, 1802, was designated subsequently Chilinialopsis Toro, 1976:73. Chilimatopsis Type species: by Taschenberg, 1883:52. Melittinae. parvula Toro, 1976, by original designation. Exomalopsini. Cingulata Mt\\e\y, 1935:43, 149. Type species: Vespa pratensis Chilimelissa Toro and Moldenke, 1979:149. Type species: Geoffroy, 1785 (not Miller, 1759) = Sphex signata Panzer, Chilimelissa iiiisa Toro and Moldenke, 1 979, by original des- 1798, monobasic. [Prosopis]. Isotypic with Prosopis by syn- ignation. Xeromelissinae. onymy. Hylaeinae. Chilioediscelis Toro and Moldenke, 1979:104. Type species: Cirroxylocopa Hurd and Moure, 1963:102. Type species: Chilicola andina Toro and Moldenke, 1979, by original des- vestita Hurd and Moure, 1963, ignation. [Chilicola]. Xeromelissinae. Xylocopa by original designation. [Xylocopa]. Xylocopini. Chilosima Michener, 1939:78. Type species: Ashmeadiella rhodognatha Cockerell, 1924, by original designation. Citrinopsithyrus Thorp. 1983, in Thorp, Homing and Dunning, [Ashmeadiella]. Osmiini. 1983:50. Type species: Apathus citrinus Smith, 1854, by original Substitute for Chlerogas Vachal, 1904:127. Type species: Halictus chlerogas designation. LxiboriopsithyrusYnson, 1927, Vachal, 1904, monobasic and absolute tautonomy. the type species of which turns out to be an anthophorine bee Augochlorini. of the genus Habropoda. Bombini. 16 Scientific Papers, Natural History Museum, The University of Kansas

Cladocerapis Cockerell, 1904b:292. Type species: Coelioxula Pasteels, 1982:110. Type species: Coelioxys = LampmcoUetes cladocenis Smith. 1862 = LcimprocoUetes ntfocaiidata Smith, 1854 Coelioxys ruficaiida Lepeletier, 1841, bipectimitiis Smith. 1 836, by original designation. Colletinae. by original designation. Megachilini. Claremoutiella Cockerell 1933c:26. Coelioxys Latreille, 1809:166. Type species: Apis conica Type species: Spinoliella = = Linnaeus, 1758 Apis quadridentata Linnaeus, 1758, by eiixcmtha Cockerell. 1916 Calliopsls zonalis Cresson, 1 879, designation of Curtis, 1831, pi. 349 and Westwood, I840a:85. by original designation. [Spinoliella]. Isotypic with Latreille's designation. 1810:439. is not acceptable because Nomadopsis by synonymy. Panurginae. he listed two species names, not indicating that they were Clavicera Latreille, 1 802a:432. Type species: Hylaeus albilabris synonyms. He treated them as he treated dissimilar species in = Fabricius, 1793 Apis cuciirhitinci Rossi, 1792, monobasic. certain other cases, e.g., Centris. Megachilini. Commission 1011 Suppressed by Opinion (1973). Isotypic Colax Lepeletier and Serville, 1825:4, 213, nomen nudum. with Cenitina. Ceratinini. Colax Lepeletier and Serville, 1 828:448, not Hubner, 1819. Type Clavinomia Wamcke, 1980:372. Type species: Nomia clavicomis species: Rhathynuis hicolor Lepeletier and Serville, 1828, monobasic. Colax was as a of It Warncke, 1980, by original designation. [Nomia]. Nomiinae. shown synonym Rhathymns. is not only a junior homonym of Colax Hubner but is not Clavipanurgus Warncke 1972:95. Type species: Pamirgus available according to the Code, ed. 3, Art. 1 1(e) because it c/avaf«5 Wamcke, 1972, by original designation. [Panurgus]. was proposed in synonymy. Isotypic with Rhathymiis; isotypic Panurginae. with Liogastra by synonymy. Rhathymini. Moure, 1961:219. Lesthmelitta Cleptotrigona Type species: Colletellus Michener, 1965a:70. Type species: Andreiiopsis cubiceps Friese, 1912, by original designation. [Lestrimelitta]. velittinus Cockerell, 1929 (not Paracolletes velutinus Meliponini. Cockerell, 1929, homonym in Leioprottits) = Leioproctiis veliitinellus Michener, 1965, by original designation. Clisodon Patton, 1879a:479. Type species: Authophora tenninalis [Leioproctus]. Colletinae. Cresson, 1869, by original designation. Anthophorini. Colletes Latreille, I802a:423. Type species: Apis sitccincta Clisranthidium Michener and Griswold, 1994a:315. Type spe- Linnaeus, 1758, monobasic. Colletes hirta Lepeletier and cies: Dianthidium tiirnehcitm Mavromoiistakis, 1934, by Serville, 1825 =A/7WCM/H'cM/ana Linnaeus, 1758, designated original designation. [EoanthidiumJ. Anthidiini. as type by Blanchard, 1849, pi. 125 (1847), was not origi- Clypanthidium Pasteels, 1968a: 1060. Type species: Anthidium nally included. Isotypic with Evodia by synonymy. nificeps Friese, 1914, by original designation. Also described Colletinae. as new Pasteels, 1969a:53. Anthidiini. by [Bathanthidium]. Colletopsis Michener. 1965a:58. Type species: Leioproctus contrarius 1965, Cnemidandrena Hedicke, 1933:213. Type species: Melitta Michener, by original designation. Colletinae. nigriceps Kirby, 1802, by original designation. [Andrena]. [Leioproctus]. Andreninae. Colocyuthophila Moure, 1948:342. Type species: Tetralonia fen-ens Smith, 1879, by original Cneiiiidiiiiu Perty, 1833:148, not Goldfuss, 1826. Type species: designation. [Peponapis]. Eucerini. Cnemidium viride Perty, 1 833, monobasic. See Euglossella. Euglossini. Comeptila Moure and Michener, 1955b:304. Type species: Thyreothremma paraguayensis Brethes, 1909, by original Cnemidorliizci Hirashima, 1981a:l. Type species: Prosopis designation. [Melissoptila]. Eucerini. elegans Smith, 1 864 = Prosopis elegantissiina Dalla Torre, Moure, 1995:143. 1896, by original designation. [Palaeorhiza]. Hylaeinae. Compsoclepta Type species: Compsoclepta fasciatu Moure. 1995 = Osiriniis lemniscatiis Roig-Alsina. Coccineoboinbiis Skorikov, 1922b: 157. Type species: Boinbus 1989, by original designation. This synonymy was indicated coccineus Friese, 1903, by designation of Sandhouse, by D. Urban in litt. (1996). Isotypic with Osiriniis by syn- 1943:539. Bombini. [Alpigenobombus]. onymy. Osirini. Cockerellia Ashmead, 1898:284. Type species: Perdita hyalina Compsomelissa Alfken, 1 924:25 1 . Type species: Compsomelissa - Cresson, 1878 Perdita albipeimis Cresson, 1865, by origi- borneri Alkfen, 1924, monobasic. Isotypic with Exoneunila nal designation. Panurginae. by synonymy. Allodapini. CockereUiila Strand, 1932:196, for Littziella replacement Conandrena Viereck, 1924:20. Type species: Andrena bradleyi 1922. Perdita Cockerell, Type species: opuntiae Cockerell, Viereck, 1907, by original designation. [Andrena]. 1922, autobasic. Panurginae. Andreninae. Coelioxita 1977:180. Pasteels, Type species: Coelioxys afra Conanthalictus Cockerell, 1901b:209. Type species: Halictiis with Lepeletier, 1841, by original designation. Isotypic conanthi CockertW, 1901, monobasic. [Halictus|. Rophitinae. Allocoelioxys. Megachilini. Confiisibombiis Ball, 1914:78. Type species: Bombiis confiisiis Coelioxoides Cresson, 1878:94. Type species: Coelioxoides Schenck, 1859, monobasic. [Bombus|. Isotypic with pimctipennis Cresson, 1878, monobasic. Tetrapediini. Sulcobombiis and Confiisohombiis. Bombini. Genus-Group Names of Bees 17

Confusobomhiis Skorikov, 1922b: 156. Type species: Bomhiis Smith, 1879, by original designation. [Corynura]. See confiisiis Schenck, 1859, by designation of Richards, Corynuroides. Augochlorini. 1968:214. with Sidcobomhus and Confusibombiis. Isotypic Corythochlla Michener, 1939:74. Type species: Ashmeadiella Bombini. inyoensis Michener. 1939, by original designation. Congotrachusa Pasteels, 1969a:24. Type species: Anthidium [Ashmeadiella]. Osniiini. sclwiitedeni Wddvdl, 1910, by original designation. [TrachusaJ. Crawfordapis Moure, 1964a:448. Type species; Megacilissa Anthidiini. luctuosa Smith. 1861, by original designation. [Zikanapis]. Conohalictoides Viereck, 1904a:245; also described as new in Caupolicanini. 1904b:26]. Type species: Conohalictoides lovelll Viereck, Creightonella Cockerell, 1908b: 146. Type species: Megachile 1904 = Pciiuiigus uovaeangllae Robertson, 1897, by original mitlmia Cockerell, 1908 = Megachile cognata Smith, 1853, designation. Rophitinae. by original designation. [Megachile]. Megachilini. Copoxyla Maa, 1 954:2 1 1 . Type species: "Apis iris Christ," 1 79 1 Creightonlella Pasteels, 1965:10, error for Creightonella = Xylocopa cyanescens Brulle, 1 832. by original designation. Cockerell, 1908. Megachilini. Christ's work is largely binominal but a section on bees is Cremnandrena 1986:554. Andrena trinominal and the names published there are not valid. None- LaBerge, Type species: anisochlora Cockerell, 1936, theless the identity of the type species was clearly indicated by original designation. Andreninae. by Maa's designation using a common appellation, iris Christ, [Andrena]. for the species concerned. [XylocopaJ. Xylocopini. Cressoniella Mitchell, 1934:307. Type species: Megachile Cresson" = M. Cresson, 1878, Coptepeolus Moure, 1955:120. Type species: Coptepeolus "zapoteka zapoteca by original emarginatus Moure, 1955, by original designation. Epeolini. designation. [Megachile]. Megachilini. Coptorthosomci Perez, 1901:3, unjustified emendation of Crewella Cockerell, 1903b:202. Type species: Ceratiiui titusi Koptortosomu Gribodo, 1894. Xylocopini. Cockerell, 1903, by original designation. [Ceratina]. Ceratinini. Coqidllettapis Viereck, 1909:47. Type species: Coqiiillettapis welittoldes Viereck. 1909 = Melissodes nigrifrons Cresson, Crinoglossa Friese, 1925b:502. Type species: Crinoglossa 1878, by original designation. Emphorini. natalensis Friese, 1925, monobasic. Nomiinae. Corbicula Friese, 1908b:59, also 1908c: 170, not Megerle, I81I. Crocisa Jurine, 1801:164. Type species: Nomada scutellata - Type species: Codiicida aplformis Friese, 1908, monobasic. Jurine, 1801 Melecta histrlonica Illiger, 1806, by designa- See Canephonila. Eucerini. tion of Morice and Durrant, 1915:423. Invalidated by Com- mission 135 (1939). Melectini. Cordandreiia Wanicke, 1968:63. Type species: Andrena cordialis Opinion Morawitz, 1878, by original designation. [Andrena]. Crocisa Jurine, 1807:239. Type species: Nomada hisirio Fabri- Andreninae. cius, 1775, by designation of Latreille, 1810:439. Nomada scutellarls Fabricius. 1781, was designated by Michener, Comylaeiis Snelling, 1985:8. Type species: Prosopis aterrima 1 1 1 965a:2 1 8. Crocisa ramosa Lepeletier, 84 , was designated Friese, 191 1, by original designation. [Hylaeus]. Hylaeinae. as type species by Blanchard, 1849:pl. 128 (1847) but was Coiynogaster Sichel, 1867:146. Type species: Coryiuira gayi not originally included. Melectini. Spinola, 1851 = Hcdictiis ntbellusHdV\&dy. 1836, designated Crac/.«i.s7;/W(c; Ashmead, 1899a:68. Crocisaspidia by Daly, Michener, Moure, and Sakagami, 1987:104; these Type species; clumdleri Ashmead, 1899, by original designation. Nomiinae. authors explained the invalidity of Sandhouse's (1943:540) designation due to her confusion of Halictus gayi Spinola Crocissa Panzer, 1806:263. Type species: Nomada scutellarls and Corynura gayi Spinola. Isotypic with Corynura. Fabricius, 178l,by designation of Sandhouse, 1943:541. This Augochlorini. name was published as a synonym of Thyreus and was credited to Jurine; presumably it was an error for Crocisa. Isotypic Coiyiiiira Spinola. 1851:296. Type species: Corynura gayi with Thyreus. Melectini. Spinola, 185! - Halictus rubellits Hdhday, 1836, designated by Alfken, 1926a; 146. Sandhouse, 1943:540, subsequently Crotchiihombus Franklin, 1954:51. Type species: Bombus designated the same species. Isotypic with Coiynogaster crotchii Cresson, 1878, by original designation. [Bombus], Augochlorini. Bombini.

Corynurella Eickwort, 1969:398. Type species: CorynureUa Cryptandrena Pittioni, 1948:49, not Lanham, 1949. Type spe- mourei Eickwort, 1969, by original designation. cies: Andrena ventricosa Dours, 1 873, by original designa- Augochlorini. tion. [Andrena]. Andreninae. Corynuroides Sandhouse, 1943:540, replacement for Cryptandrena Lanham, 1949:222, not Pittioni, 1948. Type spe- Corymiropsis Cockerell, 1901. Type species: Corynura cies: Andrena carlini Cockerell, 1901, by original designa- (/flnv//;/ Cockerell, 1901 = Augochlora briseis Sim\\h.. 1879, tion. [Andrena]. See Bythandrena. Andreninae. autobasic. Augochlorini. Cryptohalictoides Viereck, 1 904b: 261. Type species; Corynuropsis Cockerell, 1901a:220, not Scott, 1894. Type spe- Cryptohalictoides spiniferiis Viereck, 1904, by original des- cies; Coo7!Mrac/a/-n7>(; Cockerell, 1901 = Augochlora brisels ignation. Rophitinae. 18 SciENTipyc Papers, Natural History Museum, The University of Kansas

1950:14. Cubiandrena Wamcke. 1 968:76. Andrena ciibiceps Crv7"ti-$'"'a Yasumatsu and Hirashima. Type species; Type species: Osmia satoi Yasumatsu and Hirashima, 1950, by original Friese, 1914, by onginal designation. [Andrena]. Andreninae. Osmiini. designation. [Osmia]. Cubitalia Friese, 191 Id: 136. Type species: Eiicera breviceps Ctenanthidium Urban, 1993:85. Type species: Ctenaiuhidium Friese, 1911. monobasic. [Eucera]. Eucerini. Urban, 1991, Anthidiini. gracile by original designation. Cubitognatha Michener. 1939:81. Type species: Ashmeadiella Ctenaugochlora Eickwort. 1969:435. Type species: Neocorynura xenomastax Michener. 1939, by original designation. perpectinata Michener. 1954. by original designation. [Ashmeadiella]. Osmiini. [Caenaugochlora]. Augochlorini. Cidlumanihombus Skorikov. 1938a: 145. unjustified emendation Ctenioschelus Romand, 1840:336. Type species: Acanthopus of Cullumanobombits Vogt. 1911. Bombini. Romand. 1840, monobasic. Taschenberg, 1883:78, des- gorv'/ Ciilliimanohombiis 1911 :57. cullwmina 1841 Wogl. Type species: Apis ignated as type species Melissoda hitreillei Lepeletier. Kirby. 1802. by designation of Frison, 1927:66. [Bombus]. = Acanthopus Romand. 1 840. Isotypic with Ischnocera goni Bombini. and Melissoda by synonymy. Ericrocidini. 1918a:91. Halictiis lutea Curtisapis Robertson, Type species: Ctenoapis Cameron, 1901a: 1 16. Type species: Ctenoapis coriaceus Smith, 1853, by original designation. Halictini. Cameron, 1901, by designation of Sandhouse, 1943:542. Pararhophitini. Curvinomia Michener, 1944:251. Type species: Nomia Michener. 1937 = Nomia tetrazonata Cockerell. Ctenoceratina Daly and Moure, 1988, //; Daly, 1988:12. Type californiensis 1910. original designation. [Nomia]. Nomiinae. species: Ceratina annata Smith. 1854, by original designa- by tion. Ceratinini. Ciispidella Pasteels. 1965:544. Type species: Chalicodoma Pasteels. 1965. Ctenocolletes Cockerell, 1929c: 35 8. Type species: Stenothtus qiiadraticauda by original designation. nicholsoni Cockerell. 1929. monobasic. [Stenotritus]. [Chalicodoma]. Megachilini. Stenotritidae. Cyaneoderes k%\\mt^.A, 1899a:70. Type species: Cyaneoderes = Bombus coeruleus Fabricius, Ctenocorynura Schrottky. 1914:628. Type species: fairchildi Ashmead. 1899 = Ctenocorynura vernoniae Schrottky. 1914 Halictiis 1804. by original designation. Xylocopini. with inflaticeps Ducke, 1906, by original designation. Isotypic Cyanocentris Friese, 1900:244, 251. Type species: Apis versi- Rhinocorymira by synonymy. Augochlorini. color Fahncius. 1775. by designation of Sandhouse. 1943:543. Ctenonomia Cameron, 1903b: 178. Type species: Ctenonomia [Centris]. Centridini. monobasic. Halictini. carinata Cameron, 1903, Cyathocera Smith. 1875:47. Type species: Cyathocera no Ctenoplectra Kirby. 1826. in Kirby and Spence. 1826:681. nodicornis Smith. 1875. monobasic. Nomiinae. chalyhea Smith. 1858. species. Type species: Ctenoplectra Cyphanthidium Pasteels. 1969a:57.Type species: Cyphanthidiiun and of Sandhouse, 1943:542. by inclusion designation Isotypic intermedium Pasteels. 1969, by original designation. with Ctenoplectra Smith. Ctenoplectrini. Anthidiini. Smith. 1858:44. Ctenoplectra Ctenoplectra Type species: Cyphepicharis Moure. 1945b:306. Type species: Cyphepicharis Smith. 1858. monobasic. and chalybea Homonym synonym borgmeieri Moure. 1945. by original designation. Centridini. of Ctenoplectra Kirby. 1826. Isotypic with Ctenoplectra Cyphomelissa Schrottky. 1902a:493. Type species: Cyphomelissa Kirby. Ctenoplectrini. = pemigra Schrottky. 1902 Melissa diabolica Friese, 1900, Cockerell. 1909d:314; also described as new by Ctenoplectrella by original designation. Ericrocidini. Cockerell. 1909e: 19. Type species: Ctenoplectrella viridiceps Robertson. 1 903a: 1 69. 1 72. Type species: Megachile Cockerell, 1909. monobasic. Fossil. Cyphopyga montivaga Cresson, 1878. by original designation. Cockerell. 1930a:360. Ctenoplectra Ctenoplectrina Type species: Megachilini. politula Cockerell, 1930. monobasic. [Ctenoplectra]. Cxphoxylocopa Hurd and Moure. 1963:283. Type species: Ctenoplectrini. Xylocopa ocularis Perez, 1901, by original designation. Ma. 1938:285, not McAtee and Malloch. 1933, Type Ctenopoda [Xylocopa]. Xylocopini. species: Apisfenestrata Fabricius, 1798. by original designa- Cvprirophites Warncke. 1979:135. Type species: Dufourea tion. [Xylocopa]. See Ctenoxylocopa Michener and Baana caeruleocephalu cypria Mavromoustakis, 1952, by original Sandhouse. Xylocopini. designation. [Rophites]. Rophitinae. Ctenosibyne Moure. 1956:311. Type species: Lonchopria Cvr/fl/JM Cockerell. 1908c:339. Type species: Cyrtapis anomalus cingulata Moure, 1956, by original designation. [Lonchopria]. monobasic. Fossil. Colletinae. Cockerell, 1908, Mitchell. 1973:106. Type species: Coelioxys Ctenosmia Thomson. 1872:233. Type species: Apis adunca Cyrtocoelioxys costaricensis Cockerell. 1914. by original designation. Panzer, by designation of Michener. 1941:158. Isotypic with Megachilini. Hoplitis. Osmiini. [Coelioxys]. Mavromoustakis. 1963d:753. Ctenoxylocopa Michener. 1942:282, replacement for Ctenopoda Cxrtopasites Type species: Schmiedeknechtia Mavromoustakis. 1959. by origi- Ma, 1938. Type species: Apis fenestrata Fabricius, 1798. verhoeffi Ammobatoidini. autobasic. [Xylocopa]. Xylocopini. nal designation. [Schmiedeknechtia]. Genus-Group Names of Bees 19

and 1955:218. Cyrtosmia Michener, 1947:292. Type species: Osmia hypocrita Dasyscirtetica Michener. LaBerge, Moure, Type Cockerell, 1906. by original designation. [Hoplitis]. Osmiini. species: Tetralonia gilva Hoimberg, 1884, by original designation. [Alloscirtetica]. Eucerini. Dactylandrena Viereck, 1924:20. Type species: Andrena maiira

= 1 1 villosa Viereck. 1 924 Andrena caliginosa Viereck. 9 6, by origi- Dasystilbe Dressier, 1978:193. Type species: Euglossa nal designation. [Andrena]. Andreninae. Moure, 1968, by original designation. [Euglossa]. Euglossini. Dactylomegachile Mitchell, 1943:670. Type species: Megachile Dasyxylocopa Hurd and Moure, 1963:113. Type species: parsonsiae Schrottky. 1914. by original designation. Xylocopa bimaculata Friese, 1903, by original designation. [Megachile]. Megachilini. [Xylocopa]. Xylocopini. Daclylurina Cockerell, 1934b:47. Type species: Trigona Delomegachile Viereck, 1916:745. Type species: MegachUe stainlingeri Gribodo, 1893, by original designation. vidua Smith, 1853 = M. kitimaims Say, 1823, monobasic. Meliponini. [Megachile]. Isotypic with Xanihosarus by synonymy. Dalloapida Michener, 1975:246. Type species: Allodape dichroa Megachilini. Strand. 1915, by original designation. [Allodapula]. Deitoptila LaBerge and Michener. 1963:211. Type species: Allodapini. Habropoda moittezumia Smith, 1879, by original designa- Dasiapis Cockerell, 1903c:450. Type species: Dasiapis ochracea tion. Anthophorini. 1903. monobasic. Cockerell, Emphorini. DenticoUetes Noskiewicz, 1936:25, 486. Type species: CoUetes Dasyandrena LaBerge, 1977:71. Type species: Andrena graejfei Miken. 1900, monobasic. [Colletes]. Colletinae. Viereck, 1916, obscuripostica by original designation. Dentigera Mehely. 1935:45, 151. Invalid because no type spe- Andreninae. [Andrena]. cies was designated. [Prosopis]. Hylaeinae. and 1985:49. species: Dasybombus Labougle Ayala, Type Dentigera Popov, 1939:168. Type species: Hylaeus brevicornis Bombus and 1985, by original macgregori Labougle Ayala, Nylander, 1852, by original designation. [Prosopis]. [Bombus]. Bombini. designation. Hylaeinae. Mitchell, 1973:99. Coelioxys Dasycoelioxys Type species: Dentirophites Warncke, 1979:130. Type species: Ditfourea pergandei Schletterer, 1890, by original designation. gaullei Vachal, 1897, by original designation. [Rophites]. Megachilini. [Coelioxys]. Rophitinae. Smith, 1853:14. Type species: DasycoUetes DasycoUetes Deranchanthidiiim Griswold and Michener, 1988:34. Type spe- metallicus Smith, 1853, by designation of Cockerell, cies: Dianthidium chaniela Griswold and Michener, 1988, by 1905a:347. Colletinae. original designation. [Dianthidium]. Anthidiini. Dasyglossa lUiger, 1807, in Klug, 1807b:217 footnote, no valid Deranchylaeus Bridwell. 1919:136. Type species: Prosopis species names; Klug, 1810:44, included valid species. Type ciirvicarinata Cameron, 1905, by original designation. species: Oxaeaflavescens Klug, by designation of Sandhouse, [Hylaeus]. Hylaeinae. 1943:544. Isotypic with Oxaea. Oxaeinae. Derandrena Ribble, 1968a:333. Type species: Andrena vandykei Dasyhalonia Michener, LaBerge, and Moure, 1955:226. Type Cockerell, 1936, by original designation. [Andrena]. Tetralonia Friese, 1910 = Tetralonia species: spiniventris Andreninae. mimeiica Brethes, 1910, by original designation. Eucerini. Dermatohesina Michener. 1965a:91. Type species: Dasyhesma Michener. 1965a: 102. Type species: Dasyhesma Euryglossimorpha ubnormis Rayment, 1935, by original des- robiista Michener, 1965, by original designation. ignation. [Euryglossa]. Euryglossinae. Euryglossinae. Derotropis Mitchell, 1936:156. Type species: Megachile Dasymegachile Mitchell, 1943:669. Type species: MegachUe pascoensis Mitchell, 1934, by original designation. saulcyi Guerin, 1845, by original designation. [Megachile]. JMegachile]. Megachilini. Megacliilini. Desmotetrapedia Schrottky, 1909d:223. Type species: Tetrapedia Dasyniegilla Brooks, 1988:486. Type species: Apis niuelleri Friese, 1899, by original designation. Isotypic with quadrimacuhaa Panzer. 1798. by original designation. Chalepogenus by synonymy. Tapinotaspidini. [Anthophora]. Isotypic with Lasius Jurine. Antiiopliorini. Diadasia Patton, 1879a:475. Type species: Melissodes enavata Dasyosmia Michener. 1947:294. Type species: Alcidamea Cresson, 1872, by original designation. Emphorini. biscHtellae Cockerell. 1897, by original designation. Diadasiana Michener, 1954:130, incorrect subsequent spelling [Hoplitis]. Osmiini. of Diadasina Moure, 1 950. Emphorini. Dasypoda Latreille, 1802a:424. Type species: Andrena hirtipes Diadasiella Ashmead, 1899a:64. Diadasiella Fabricius, 1793, Blanchard, 1840:414. Me/Zm; Type species: designated by = coqidlletti Ashmead, 1 899 Synhalonia albicans Provancher, swammerdamiella Kirby, 1802, designated as the type spe- 1896, by original designation. Exomalopsini. cies by Curtis, 1831:367, was not originally included in Melitoma Dasypoda but is a synonym of Andrena hirtipes Fabricius. Diadasina Moure, 1950c:392. Type species: paraensis Dasypodinae. Ducke, 1912, by original designation. Emphorini. 20 SciENxmc Papers, Natural History Museum, The University of Kansas

Diagonozus Endeilein, 1903:35. Type species: Diagonozus Dilobopeltis Fairmaire, 1858:266. Type species: Dilobopeltis bicometes Enderlein, 1903, monobasic. Halictini. fiiscipennis Fairmaire, 1858 = Thynniis abdomiiwlis Fabri- cius, 1793, monobasic. with Dialictiis Robertson. 1 902d:48. Type species: Halictus anomalus Isotypic Euaspis by synonymy. Anthidiini. Robertson, 1892, by original designation. Halictinae. Moure and Hurd, 1982:46. Dialonia Robertson, 1903b: 104. Type species: Sphecodes Dinagapostemou Type species: Halictus sicheli Vachal, 1901. antenuariae Robertson, 1891, by original designation. by original designation. Halictini. Halictini. Dinavis Pasteels, 1965:549. miiansae Diaiulrena Cockerell, 1903a:75. Type species: Paniirgus Type species: Megachile Friese, 1911. chalybaeits Cresson, 1878, by original designation. by original designation. [Chalicodoma]. Andreninae. Megachilini. Dinomia Hirashima. 1956:29. Nomia taiwana Dianthidiiim Cockerell. 1900b:412. Type species: Anthidiiim Type species: Hirashima, 1956, by curvatuiu Smith, 1 834, by original designation. [Anthidium]. original designation. [Nomia]. Anthidiini. Nomiinae. Diiioxylocopa Hurd and Moure. 1963:230. Type species: Diaxylocopa Hurd and Moure, 1963: 129. Type .species: Xylocopa Xylocopa absiirdipes Enderlein. 1903 = Xylocopa io Vachal. tnixali Hurd and Moure, 1963. by original designation. 1898, by original designation. [Xylocopa], [Xylocopa]. Xylocopini. Xylocopini. Diomulopsis Michener and Moure, 1957:431. Type species: D/cera/05/»/(7 Robertson, 1903a:166. 171. Type species: Osmia Exomalopsis bicellularis Michener and Moure, 1 957, 4-deutata Cresson, 1878 = Osmia conjimcta Cresson, 1864, by original designation. [Exomalopsis]. Exomalopsini. by original designation. Osmiini. Dioxoides Popov, 1947:89. Type species: Coelioxys trideutata Dichanthidiiiiu Moure, 1947a:30 (January): also described as Nylander. 1848, by original designation. Dioxyini. new by Moure. 1947b:235 (June). Type species:

and 1 825: 1 09. Dicluiiuhidium exile Moure, 1947, by original designation. Dioxys Lepeletier Serville, Type species: Trachusa Anthidiini. cincta Jurine, 1807, monobasic. Dioxyini. Friese. 1906d:92. Ancvloscelis Dichroa Illiger, 1806:46. Type species: Sphex gihba Linnaeus. Dipedia Type species: armatus Smith. 1854. ofLutz and 1920:592. 1758. by designation of Sandhouse. 1943:545. Isotypic with by designation Cockerell, Sphecodes. Halictini. Emphorini.

185 1 : 168. Dicranthidiiim Moure and Urban. 1975:837. Type species: Diphaglossa Spinola, Type species: Diphaglossa gayi 1851. monobasic. Anthidium aremihum Ducke, 1907, by original designation. Spinola. Diphaglossinae. Anthidiini. Diphysis Lepeletier, 1 841 :307. Type species: Diphysis pyreuaica 1841 Panzer, 1798. monobasic. Dicnmiouia Michener and Brooks. 1987:100. Type species: Lepeletier, -Apis byssina with Trachusa Panzer and Agemmonici wenzeli Michener and Brooks, 1987, by original Isotypic Megachileoides by syn- Anthidiini. designation. [Agemmonia]. Meganomiinae. onymy. 1981, in Michener. 1981:96. Dictyohalictiis Michener, 1978a:518. Type species: Halictus Disparapis Stage. Type species: arenicola Crawford, 1917, retigeriis Cockerell, 1940, by original designation. Hesperapis by original designation. [Pachyhalictus]. Isotypic with Rugalictiis by synonymy. [Hesperapis]. Dasypodinae. Halictini. Dissoglotta Moure, 1945a: 144. Type species: Dissoglotta stenoceratina Moure. 1945, Didoiiia Gribodo, 1894:106. Type species: Didonia piinica by original designation. Gribodo. 1894 = Andreiui miicida Kriechbaumer, 1873, Dis.soglottini. monobasic. Andreninae. Distandrcna Warncke, 1968:60. Type species: Andrena hmgibarbis Perez, 1895, Diepeohis Gribodo, 1894:79. Type species: Epeolus giaiinellii by original designation. [Andrena]. Andreninae. Gribodo, 1894, monobasic. [Epeolus]. Epeolini. Moure and 1955b:309. Dietinomia Cockerell, 1899a: 14. replacement for Eunotnia Dithygater Michener, Type species: seabiai Moure and Michener, 1955, Cresson, 1875. Type species: Eiinomia margitiipennis Dithygater by original Eucerini. Cresson, 1875 - Nomia kirbii Smith. 1865, autobasic and by designation. designation of Cockerell. 1910f:290. Nomiinae. Diversibonihus Skorikov, 1938b: I, unjustified emendation of Diversobombus 1914. Bombini. Digitella Pasteels, 1965:191. Type species: Megachile Skorikov, digiticauda Cockerell, 1937. by original designation. Diversobombus Skorikov, 1914b:406. Type species: Bombus [Megachile]. Megachilini. diversusSmnh. 1869. by designation of Sandhouse, 1943:546. Bombini. Digressobombiis Laverty. Plowright, and Williams. 1984:1051. [Bombus]. Type species: Megabambiis digressiis Milliron, 1 962, by origi- Dizonamegilla Brooks, 1988:505. Type species: Megilla nal designation. [Bombusj. Bombini. sesquicincta Erichson and Klug, 1842, by original designation. [Amegilla]. Dignmoceras Cockerell. 1931c: 134. Type species: Megachile Anthophorini. combusta 1853 Smith, =Apiscincta Fabricius. 1771, by origi- Doeringiella Holmberg. 1886a: 151: 1886b:233. Type species: nal designation. [Gronoceras]. Megachilini. Doeringiella bizonata Holmberg, 1 886, monobasic. Epeolini. Genus-Group Names of Bees 21

Dolichochile Viereck. 1909:49. Type species: Doliclioclille Effactapis Michener. 1977:6. Type species: Ejfactapis furax melitioulesViereck. 1909, by original designation. Melittinae. Michener, 1977, by original designation. Allodapini.

Dolichosteiis Parker and Bohart, 1979:138. Type species: Stelis Egapista Cockerell, 1904a:357. replacement for Apista Smith, laticiiicta Cresson, 1878. by original designation. Anthidiini. 1861. Type species: Apista opalina Smith. 1861, autobasic. Dolichotrigona Moure, 1950a:248. Type species: Triguna Dissoglottini. longitarsis Ducke, 1916, by original designation. Eickwortapis Michener and Poinar, 1997:354. Type species: [Hypotrigona]. Meliponini. Eickwortapis dominicana Michener and Poinar. 1997, by Fossil. Halictinae. Domautludiiim Pasteels, 1969a:95. Type species: Anthidium original designation. abdominale Friese, 1904, Anthidiini. by original designation. Elandrena Lanham, 1 949:203. Type species: Andrena ampUficata Doxauthidiiim Pasteels, 1969a:28. Type species: Anthidium Cockerell, 1910, by original designation. [Andrena|. parcidoxum Mocsary, 1884, by original designation. Andreninae. Anthidiini. Elaphropoda Lieftinck, 1966:148. Type species: Habropoda Drepauium Robertson. 1903b: 103. Type species: Sphecodes impatiens Lieftinck. 1944. by original designation. = fcilcifer Patton, 1880 Sphecodes conferius Say, 1837, by Anthophorini. original designation. Halictini. Elecolletes Warncke, 1978:330. Type species: CoUetes elegans 1960:2. Duckeanthidiiim Moure and Hurd. Type species: Noskiewicz. 1936, by original designation. [Colletes]. Anthidium megachiliforme Ducke, 1907, by original desig- Colletinae. nation. Anthidiini. Electrapis Cockerell, 1908d:326; also described as new by

Duckeola Moure, 1 944a:7 1 . huberi Friese, Type species: Trigona Cockerell, 1909e:7. Type species: Apis melipouoides Buttel- 1901 = 1853, Trigona ghilianii Spinola, by original designa- Reepen, 1906, designated by Cockerell, 1909e:8. Fossil. tion. Meliponini. Emphor Patton, 1879a:476. Type species: Melissodes 1841:227. minuta Diifourea Lepeletier, Type species: Dufourea bombiformis Cresson, 1878, by original designation. 1841. of Richards, 1935:172 and Lepeletier, by designation Emphorini. Timberlake, 1939:396. D. vulgaris Schenck, 1859, was not Emplioropsis Ks\\me:id. 1 899a:60, no included species; Cockerell designated by Cockerell, 1924c:244, as stated by Sandhouse and Cockerell, 1901:48, included species. Type species: ( 1943:547), and was not included. Rophitinae. originally = Anthophorafloridana Smith, 1 854 Bombus laboriosus Fab- ricius, 1804, by designation of Cockerell and Cockerell, Ebmeria Pesenko, 1 986a: 1 36. Type species: Halictus costulatus 1901:48. Anthophorini. Kriechbaumer. 1873. by original designation. [Lasioglossum]. Halictini. Energoponus Holmberg, 1903:406. Type species: Energoponus strenuus Holmberg, 1903 - Ptilothrix plumatus Smith. 1853, Ebmeriana Pagliano and Scaramozzino, 1 990: 1 27, replacement by designation of Sandhouse, 1943:547. Isotypic with for Micropasites Warncke, 1983. Type species: Ammohates Ptilothrix by synonymy. Emphorini. minutissimus Mavromoustakis, autobasic. [Pasites].

Ammobatini. Ensliniana Alfken, 1 938:43 1 . Type species: Ensliniana cuspidata = Alfken, 1 938 Stelis bidentata Friese, 1 899, by original des- Ecclitodes Roig-Alsina, 1989:9. Type species: Epeolus stuardi ignation. Dioxyini. Ruiz, 1935, by original designation. Osirini. Enteclmia Patton. 1879a:476. Type species: Anthophora taurea Echiiuinthidium Pasteels, 1969a: 101. Type species: Anthidium Say, 1837, by original designation. Emphorini. ec/iiiuitum Kiug, 1832, by original designation. [Anthidium]. Anthidiini. Eoanthidiellum Pasteels, I969a:51. Type species: Anthidium = elongatum Friese, 1 897 Anthidium clypeare Morawitz, 1 873, Echthralictus Perkins and Cheesman, 1928:14. Type species: by original designation. [Eoanthidium]. Anthidiini. Halictus extraordinarius Kohl, 1 908, by original designation. Halictini. Eoanthidium Popov, 1950a:316. Type species: Anthidium insulare Morawitz, 1873, designation. Ecplectica Holmberg, 1884:123. Type species: Ecplectica by original Anthidiini. tintinnans Holmberg, 1884, monobasic. Eucerini. [Dianthidium]. Moure, 1992:35. Edriohylaeus Michener, 1965a: 124. Type species: Hylaeus Eomelipona Type species: Melipona marginata ofarrelli Michener, 1965, by original designation. [Hylaeus]. Lepeletier, 1836, by original designation. [Melipona]. Hylaeinae. Meliponini. and 1970:20. Edwyiiia Moure, 1951c:195, not Aldrich, 1930. Type species: Eopeponapis Hurd Linsley, Type species: utahensis Cockerell. 1905, Pasiphae flavicornis Spinola, 1851, by original designation. Xenoglossa by original designa- See Edwyniana. Colletinae. tion. [Peponapis]. Eucerini.

1 1 a: 1 34. tibetanus Edwyniatui Moure. 1 954a: 1 65. replacement for Edwynia Moure, Eopsithyrus Popov, 93 Type species: Apathus 1951. Type species: Pasiphae flavicornis Spinola. 1851, Morawitz, 1886, by original designation. fPsithyrus]. autobasic. Colletinae. Bombini. 22 SciENTinc Papers, Natural History Museum, The University of Kansas

Eothrincostoma Bliithgen, 1930:496, 501. Type species: Halictus Epimethea Morawitz. 1876:61. Type species: Epimethea lorriihis Smith. 1 879. by designation of Sandhouse. 1943:548. variegata Morawitz, 1876, by designation of Cockerell, Halictini. 1922c:l. Panurginae. Eoxenoglossa Hurd and Linsley. 1970:34. Type species: Epimonispractor Holmberg, 1887b:225, nomen nudum. Melissodes strenua Cresson, 1 878, by original designation. Exomalopsini. [Xenoglossa]. Eucerini. Epimonispractor Holmberg, 1903:426. Type species: Eoxylocopa Sakagami and Yoshikawa, 1 96 1 :4 1 3, nomen nudum. Epimonispractor gratiosus Holmberg, 1903 = Exomalopsis [Xylocopa]. Xylocopini. «;;«fa Spinola, 1853,by designation of Sandhouse, 1943:548. Sandhouse Epanthidium Moure, 1947a:33. Type species: Hypauthidium erroneously reported original designation. tigrinum Schrottky, 1905, by original designation. Anthidiini. Exomalopsini. Ashmead. 1899a:88. Nomia Epeicharis Radoszkowski, 1884:18. Type species: Epeichahs Epinomia Type species: persimilis = mexiccmiis Radoszkowski, 1884, monobasic. See Fiorentinia. Cockerell, 1898 A'o«!/fl triangulifera Vachal, 1897, by origi- Tapinotaspidini. nal designation. Nomiinae.

Alfken, 1 939: 1 1 3, not 1 899. Epeoloides Giraud, 1 863:44. Type species: Epeoloides ainhigmis Epinomia Ashmead, Type species: = Giraud, 1863 = Apis coecutiens Fabricius, 1775, monobasic. Nomia andrenoides Vachal, 1903 Nomia andrei Vachal, Osirini. 1897, by original designation. [Nomia]. See Alfkenomia. Nomiinae. Epeoliis Latreille, 1802a:427. Type species: Apis variegata Linnaeus, 1758, monobasic. Isotypic with Monoepeolus. Epixylocopa Hurd and Moure, 1963:223. Type species: Xylocopa Epeolini. rufitarsis Lepeletier, 1841. by original designation. [Xylocopa]. Xylocopini. Epichanma Michener, 1954:144. Type species: Apis rusticu Olivier. 1789. by original designation. [Epicharis]. Centridini. Erandrena LaBerge, 1986:505. Type species: Andrena principalis 1986. [Andrena]. Epicharis Klug, 1807, in llliger, 1807:197, nomen nudum. LaBerge. by original designation. Centridini. Andreninae. Eremandrena 1964:295. Pterandrena Epicharis Klug, 1807b:226. Type species: Centris unibraculata LaBerge, Type species: Viereck. 1904, Fabricius, 1804, by designation of Lutz and Cockerell, pallidiscopa by original designation. [Andrena]. Andreninae. 1 920:562. Latreille (1810) listed Apis hirtipes Fabricius, 1 793 as the type species. This was not an originally included spe- Eremaphanta Popov, 1940:53. Type species: Rhophites vitellinus cies but is a subjective synonym o^Apis nistica Olivier, 1789, Morawitz, 1876, by original designation. Dasypodinae. as is Centris dasypoda Klug. 1807, which was originally in- Eremapis Ogloblin, 1956:149. Type species: Eremapis panula cluded. Moure (1945b, 1960) considered the type to be Apis Ogloblin, 1956, by original designation. Exomalopsini. rustica Olivier, the type species of Epicharana, because Eremopasites Linsley, 1942:131. Type species: Townsendiella Latreille listed a synonym of rustica as the type species of californica Michener, 1936, by original Epicharis, but the Code, 3rd ed.. Art. 69(a)(v), makes clear designation. [Townsendiella]. Townsendiellini. that this reasoning applies only if the author concerned (Latreille) indicated the synonymy, which he did not. Snelling Eremoplosmia Zanden, 1991:164. Type species: Osmia (1984:59) correctly explained the situation. Centridini. eremoplana Mavromoustakis, 1949, by original designation. [Anthocopa]. Osmiini. Epicharitides Moure, 1945b:311. Type species: Epicharis cockerelli Friese, 1900, by original designation. Centridini. Eremosmiu Michener, 1943:66. Type species: Osmia robustida Cockerell, 1935, by Epicharoides Radoszkowski, 1884:20. Type species: original designation. [Anthocopa]. Osmiini. Epicharoides hipiinctatus Radoszkowski, 1884 = Epicharis macidatus Smith, 1874, monobasic. Centridini. Eriades Dalla Torre and Friese. 1895:69, unjustified emendation of Heriades 1808. Osmiini. Epiclopus Spinola, 1851:183. Type species: Epiclopus gayi Spinola, Spinola, 1851. monobasic. Ericrocidini. Enc/or/i- Cresson, 1887:131, 134. Type species: Crocisa '.'lata Cresson, 1878, monobasic. Ericrocidini. Epicolpiis Ashmead, 1899a:61, lapsus for Epiclopus Spinola, 1851. Ericrocidini. Eriops Klug, 1807, in llliger, 1807:197; Klug. 1807b:207, 227.

Andrena lohata Panzer, 1 799 = calcarata Epihalictoides Cockerell and Porter, 1899:420. Type species: Type species: Apis Panurgus marginatus Cresson. 1878. monobasic. Scopoli, 1763, monobasic. Isotypic with Panurgus by syn- [Halictoides]. Rophitinae. onymy. Panurginae.

Epimacrotera Timberlake, 1954:377. Type species: Perdita Eryops Latreille. 1811:716. unjustified emendation of Eriops ainsliei Crawford, 1932, by original designation. [Perdita]. Klug, 1807. Panurginae. Panurginae. Enthrandrena Zavortink, 1972:61. Type species: Megandrena Epimelissodes Ashmead, 1899a:63. Type species: Melissodes mentzeliae Zavortink, 1972, by original designation. atripes Cresson, 1872. by original designation. Eucerini. [Megandrena]. Andreninae. Genus-Group Names of Bees 23

Erythronomioides Pesenko. 1 983: 1 76. Type species: Nomioides Euglossella Moure 1 967a:401 , replacement for Cnemidium Perty, socotraniis Bliithgen. 1925, by original designation. 1833. Type species: Cnemidium viride Perty, 1833, autobasic. [Nomioides]. Nomioidinae. [Euglossa]. Euglossini.

Enthrosmia Schmiedeknecht. 1885:20. Type species: Osmia Euherbstia Frie.se, 1925a:8. Type species: Euherbstia excellens andrenoides Spinola. 1808. by designation of Cockerell, Friese, 1925, monobasic. Andreninae. 1922a:6. Osmiini. [Osmia]. Euhesma Michener, 1965a:88. Type species: Euryglossa Ethalouchopha Michener, 1989:670. Type species: Apisia gaidlei wahlenbergiae Michener, 1965, by original designation. Vachal, 1909, by original designation. [Eulonchopria]. [Euryglossa]. Euryglossinae. Colletinae. Eulaema Lepeletier, 1 84 1 : 1 1 . Type species: Apis dimidiata Fab- 1793 = meriana Olivier, 1789, of Euandrena Hedicke, 1933:213. Type species: Andreim hicolor ricius, Apis by designation 1883:85. Fabricius, 1775, by original designation. [AndrenaJ. Taschenberg, Euglossini. Andreninae. Eulaenia Spinola, 1851:167, incorrect subsequent spelling of

Eulaema 1 84 1 . Euaspis Gerstaecker, 1857:460. Type species: Thynnus Lepeletier, Euglossini. abdomiualis Fabricius, 1793, by original designation. Lsotypic Eulema Smith, 1854:380, unjustified emendation of Eulaema with Dilobopeltis by synonymy. Anthidiini. Lepeletier, 1841. Euglossini.

Eiicam Friese, 1905b:241. Type species: Anthophom laticeps Eulonchopria Brethes, 1909a:247. Type species: Eulonchopria Friese, 1905, by designation of Cockerell, 1933a:465. psaenythioides Brethes, 1909, monobasic. Colletinae. Eucerini. [Anthophora]. Eumegachilana Michener, 1965a: 191. Type species: Megachile Eiicera Scopoli. 1770:8. Type species: Apis longiconus Linnaeus. clotho Smith, 1861, by original designation. [Chalicodoma]. 1758, by designation of Latreille, 1810:439. Eucerini. Megachilini. Euceratina Hirashima. Moure, and Daly. 1971. in Hirashima, Eumegachile Friese, 1898b: 198, no included species; Friese, included 1971a:369. Type species: Apis callosa Fabricius. 1794, by 1899c:36, species. Type species: Megachile

1 of original designation. [Ceratina]. Ceratinini. bomb\cina Radoszkowski, 874, by designation Cockerell, 1930c:209. Megachilini. Eucerinoda Michener and Moure, 1957:445. Type species: Eumelissa and Brooks, 1985:21. Melissa Anthophora gayi Spinola. 1851, by original designation. Snelling Type species: Eucerini. decorata Smith, 1854, by original designation. [Mesoplia]. Ericrocidini. Euchalicodoma Tkalcu. 1969:358. Type species: Megachile Eumelissodes 1956:1177. Melissodes asiatica Morawitz, 1875, by original designation. LaBerge, Type species: agilis Cresson, 1878, original designation. [Melissodes]. [Chalicodoma]. Isotypic wkh Allowegachile. Megachilini. by Eucerini. Eucharis Dalla Torre, 1896:300. Listed as a junior synonym of Friese 1899b: 126, not Hiibner, 1807. in Eumorpha Type species: Epichahs and Eucharis hiriipes listed synonymy on p. 301 , Euglossa pulchra Smith, 1854, by designation of Cockerell, and both attributed to Lepeletier [and Serville], 1825 (cor- 1908a: 41 footnote. See Eufriesea. Euglossini. rectly 1828): 794 and 795. Search of these and other pages of Eunomia not 1818. that work reveals no use of Eucharis. which therefore ap- Cresson, 1875:723, Hiibner, Type species: Eunomia 1 875 = Nomia kirbii pears to be an error by Dalla Torre, and not available accord- marginipennis Cresson, Smith, 1 865, of Cockerell, 1910f:290. See Dieunomia. ing to the Code, 3rd ed.. Art. 1 1(c). Centridini. by designation Nomiinae. Eucondylops Brauns, 1902:377. Type species: Eucondylops Eunomioides 1937:3, nomen nudum, = konowi Brauns, 1902, monobasic. Allodapini. Bliithgen, Ceylalictus. Type species: Andrena variegata Olivier, 1789, by original Eudioxys Mavromoustakis, 1963c:443. Type species: Dioxys designation. [Nomioides]. Nomioidinae. quadrispinosa Friese, 1 899, by original designation. Dioxyini. Eupalaeorhiza Meade-Waldo, 1914b:403. Type species: Euflorilegus Ogloblin, 1955:231. Type species: Florilegus Eupalaeorhiza papuana Meade-Waldo, 1914, by original riparius Ogloblin, 1955, by original designation. [Florilegus]. designation. Hylaeinae. Eucerini. Eupavknskia Popov, 1955c:330. Type species: Melectafuneraria

Cockerell 1 908a:4 1 , for Friese, Eufriesea replacement Eumorpha Smith, 1854, by original designation. Melectini. 1 899. Type species: Euglossa pulchra Smith, 1854, autobasic. Eupetersia Bliithgen, 1928a:49. Type species: Eupetersia neavei Euglossini. Bluthgen, 1928. by original designation. Also published as Lutz and emendation Eufriesia Cockerell, 1920:544, unjustified new by Bluthgen, 1928b: 165. Halictini. of Eufriesea Cockerell. 1908. Euglossini. Euphileremus Popov, 1951:906. Type species: Phileremus Euglages Gerstaecker, 1 869: 1 49. Type species: Eiiglages scripta oraniensis Lepeletier, 1841, by original designation. Gerstaecker, 1 869, monobasic. Ammobatoidini. [Ammobates]. Ammobatini. Euglossa Latreille, 1802a:436. Type species: Apis cordata Euplusia Moure 1943a: 189, replacement for Plusia Linnaeus, 1758, by designation of Blanchard, 1849:pl. 129 Hoffmannsegg, 1817. Type species: Plusia superba (1847). Euglossini. Hoffmannsegg, 1817, autobasic. Euglossini. 24 SciENTinc Papers, Natural History Museum, The University of Kansas

Euprosopellus Michener, 1965a: 132. Type species; Prosopis Euxylocopa Hurd and Moure, 1963:234. Type species: Xylocopa = dromedaria Cockerel], 1910. by original designation. fraudulenta Gribodo. 1 894 Xylocopa eiythrina Gribodo, [Hylaeus]. Hylaeinae. 1894. by original designation. [Xylocopa]. Isotypic with

Euprosopis Perkins, 1912:106. Type species: Prosopis liusela Perixylocopa by synonymy. Xylocopini. Cockerell. 1910, by original designation. Hylaeinae. Eversmannihoinhus Skorikov. 1938a: 145. Type species: Mucidohomhus eversmanniellus Skorikov, 1922 = Bomhus Eiiprosopoides Michener, 1965a: 131. Type species: Prosopis eversmanni Friese, 1911 (not B. modestus eversmanni fulvicornis Smith, 1853 = Prosopis ruficeps Smith, 1853, by Skorikov, \9 = Bomhus Rado&zkowfki, 1881, original designation. [Hylaeus]. Hylaeinae. 10} persicus by designation of Richards, 1968:214. [Agribombus]. Bombini Eiiiyapis Sandhouse, 1943:551, replacement for Orbitella Ma, Evodia Panzer. 1 806:207. calendarum Panzer, 1938. Type species: Xylocopa confusa Perez, 1901 . autobasic Type species: Apis 1802 succincta Linnaeus, monobasic. and original designation. Xylocopini. -Apis 1758, Isotypic with Colletes by synonymy. CoIIetinae. Euryglossa Smith, 1853:17. Type species: Euryglossa

Robertson. 1 902c:247. Halictus arcuatus cupreochalyhea Smith, 1 853, by designation of Meade-Waldo, Evylaeus Type species: 1923:6. Euryglossinae. Robertson, 1893, by original designation. Halictini. Exaerete 1817:53. dentata Euryglosseila Cockerell, 1910c:263. Type species: Euryglossella Hoffmannsegg, Type species: Apis minima Cockerell, 1910, monobasic. Euryglo.ssinae. Linnaeus, 1758, monobasic. Isotypic with Caliendra and Chiysantheda by synonymy. Euglossini. Euiyglossidia Cockerell, 1 9 1 Ob:358. Type species: Eiiiyglossidia rectangiilaia Cockerell, 1910, by original designation. Exallocentris Snelling, 1974:35. Type species: Centris anomala CoIIetinae. Snelling, 1966, by original designation. [Centris]. Centridini.

Eiiiyglossiinorplia Strand, 1910:40. Type species: Eiiiyglossa Exanlhidium Pasteels, 1969a:82. Type species: Antliidiuni nigra Smith, 1879, monobasic. [Euryglossa]. Euryglossinae. eximium Giraud, 1863, by original designation. Anthidiini.

Euryglossina Cockerell, 1910a:211. Type species: Euryglossa Exanthocopa Tkalcu, 1993a:55. Type species: Osmia oxypyga semipurpiirea Cockerell, 1910, monobasic. [Euryglossa]. Benoist, 1927, by original designation. [Anthocopa]. Osmiini. Euryglossinae. Excolletes Michener. 1965a:56. Type species: Leiopructus Eitiyglossula Michener, 1965a: 111. Type species: Euryglossina impatellatus Michener, 1965, by original designation. ciialcosonia Cockerell, 1913, by original designation. [Leioproctus[. CoIIetinae. Euryglossinae. Exilobombus Skorikov, 1 922b: 1 50. Type species: Mucidohomhus Eurymella Pasteels, 1965:64. Type species: Megachile eurymera exit [misprinted exiln] Skorikov, 1922, monobasic. Smith, 1854, by original designation. [Megachile]. [Mucidobombus]. Bombini. Megachilini. Exomalopsis Spinola, 1853:89. Type species: Exomalopsisfidvo- = Euiypariella Tkalcu, 1995:1 1 1. Type species: Osmia denudata pilosa Spinola, 1 853 Exomalopsis auropilosa Spinola. 1 853, Morawitz. 1880, by original designation. Osmiini. by designation of Taschenberg, 1883:82. Exomalopsini.

Euiytis Smith, 1854:279. Type species: Eutytisfimereiis Smith, Exoneuru Smith, 1854:232. Type species: Exoneura bicolor 1854, monobasic. Ericrocidini. Smith, 1 854. monobasic. Allodapini.

Eusphecogastra Hirashima, 1992:395. replacement for Exoneurellu Michener, 1963:257. Type species: Exoneura Sphecogaster Hirashima, 1978. Type species: Palaeorhiza lawsoni Payment, 1946, by original designation. Allodapini. paradisea Hirashima, 1978. autobasic and by original desig- Exoneuridia Cockerell. 1911d:232. Type species: Exoneura nation. Hylaeinae. libanensis Friese, 1 899, by original designation. [Exoneura]. Eusynhalonia Ashmead, 1899a:63. Type species: Melissodes Allodapini. edwardsii Cresson, 1878, Eucerini. by original designation. Exoneurula Michener, 1966b:573. Type species: Exoneurula Eiitechnia Holmberg, 1903:400. lapsus for Entechnia Patton. zavattarii Michener, 1966 - Compsomelissa fconicn Alfken, 1879. Emphorini. 1924, by original designation. Isotypic with Compsomelissa by synonymy. Allodapini. Euthosniia Sinha, 1958:235. Type species: Heriades glaucum Fowler, 1899. by original designation. [Osmia]. Osmiini. Exosmia Tkalcu, 1979:321. Type species: Osmia dijficilis Morawitz, 1875, [Osmiuj. Osmiini. Euthyglossa Radoszkowski, 1884:21. Type species: Euthyglossa by original designation. = fasciata Radoszkowski, 1884 Eucera euthyglossa Dalla Falirhalictus Warncke, 1975b:95. Type species: Halictus Torre, 1896, monobasic. Osirini. fahringeri Friese, 1921, by original designation. [Halictus]. Halictini. Eutricharaea Thomson, 1 872:228. Type species: Apis argentata Fabricius, 1793, monobasic. [Megachile]. Isotypic with Fasciata Mehely, 1935:44, 150. Type species: Prosopis facialis Paramegachile. Megachilini. Perez, 1895 = Prosopis irinotata Perez, 1895. monobasic. Eutrypetes Popov, 1955a:280. Type species: Heriades [Prosopis]. Hylaeinae. turcomanica Popov, 1955, by original designation. [Heriades]. Fascista Popov, 1939:168, incorrect subsequent spelling for

Osmiini. Fasciata Mehely , 1935. Genus-Group Names of Bees 25

Fermildaepsithynts Prison. 1927:70. Type species: Psitbyriis prinii Holmberg, 1884, monobasic. See Parafriesea. fenialdae Franklin, 1911, monobasic. [Psithyrus]. Bombini. Panurginae.

1. Fertonella Cockerell. 1920a:257, replacement for Perezio Ferton, Friesella Moure. 1946a:441 : 1946b:61 Type species: Melipona = 1914. Type species: Perezia mama Ferton, 1914 Megachile schrottkyi Friese, 1900, by original designation. Meliponini. leachella autobasic and Curtis, 1828, original designation. Frieseomelitta Ihering, 1912:5. Type species: Trigona silvestrii Megachilini. Friese, 1902, monobasic. Meliponini. Fervidohomhus Skorikov, 1922b: 123, 153. Type species: Apis Fiimandrena Warncke. 1974:98. nomen nudum. [Andrena]. fenida Fabricius, 1798, by designation of Frison, 1927:69. Andreninae. Bombini. Fumandrena Warncke, 1975a:57. Type species: Andrena fumida 1972:27. Bomhits Festivohombus Tkalcu, Type species: festivus Perez, 1895. by original designation. [Andrena]. Andreninae. Smith, 1861, by original designation. [Pyrobombus]. Funebribombus Skorikov, 1922b: 157. Type species: Bombus Bombini. funebris Smith, 1854, monobasic. [Alpigenobombus]. Fidelia Friese, 1 899a:244. Type species: Fidelia parado.xa Friese, Bombini. 1899, monobasic. Fideiiini. Furcosmia Schmiedeknecht, 1885:22. Type species: Apis Filiglossa Rayment, 1959a:324. Type species: Filiglossa papaveris Latreille, 1799, by designation of Cockerell, filamentosa Rayment. 1959, by original designation. 1922a:6. [Osmia]. Isotypic with Anthocopa and Phyllotoma. Colletinae. Osmiini.

Fiorentinia Dalla Torre. 1896:334, replacement for Epeicharis Gaesischia Michener, LaBerge, and Moure, 1955:220. Type spe- Radoszkowski, 1884, not Epicharis Klug. Type species: cies: Svastra fiilgiirans Holmberg, 1903, by original desig- Epeicharis mexicanus Radoszkowski, 1884. autobasic. nation. Eucerini. Tapinotaspidini. Gaesischiana Michener, LaBerge, and Moure, 1955:224. Type FlavipaiwrgusV^amcke:, 1972:69. Type species: Panurgiisflavus species: Gaesischia e.xul Michener, LaBerge, and Moure, Friese, 1897, by original designation. [Panurgus]. 1955, by original designation. [Gaesischia]. Eucerini.

Panurginae. Gaesischiopsis Michener, LaBerge, and Moure, 1955:22 1 . Type Flavodiifourea Ebmer, 1984:373. Type species: Diifourea species: Gaesischia flavoclypeata Michener, LaBerge, and flavicornis Fnese, 1913, by original designation. [Dufourea]. Moure, 1955, by original designation. [Gaesischia]. Eucerini. Rophitinae. Gaesochira Moure and Michener. I955b:283. Type species: = Flavomeliturgula Warncke, 1985:229. This name is not valid Gaesochira complanata Moure and Michener, 1955 since no type species or even included species was indicated. Melissodes obscurior Dalla Torre, 1 896, by original designa- From descriptions one can guess that Warncke intended this tion. Eucerini. to be a name for deserta Warncke. subgeneric Meliturgula Gastrohalictus Ducke, 1902:102. Type species: Halictits [Meliturgula]. Panurginae. osmioides Ducke, 1902, monobasic. [Halictus]. Halictinae. for Fiorentinia Dalla Florentiiui Ashmead, 1899a:67. lapsus Gastropsis Smith. 1868b:xxxix, replacement for Oestropsis 1896. Torre, Tapinotaspidini. Smith, 1868. Type species: Oestropsis pubescens SmiX\\, 1868, Florilegus Robertson, 1900:53. Type species: Melissodes autobasic. Stenotritidae. Cresson. 1879. monobasic. Eucerini. condigna Geandrena LaBerge, 1 964:3 13. Type species: Andrena caerulea Andreninae. Floriraptor Moure and Michener, 1955b:268. Type species: Smith, 1879, by original designation. [Andrena]. = Melissodes atropos Smith, 1879 Tetralonia melectoides Geissandrena LaBerge and Ribble, 1972:302. Type species: Smith, 1879, by original designation. [Florilegus]. Eucerini. Andrena trevoris Cockerell, 1897. by original designation. Andreninae. Foersterapis Moure, 1964a:441. Type species: Zikanapis [Andrena].

foersteh Moure and Seabra, 1962, by original designation. Geniotrigona Moure, 1 96 1 : 2 1 2. Type species: Trigona thoracica [Zikanapis]. Caupolicanini. Smith, 1857, by original designation. Meliponini. Formicapis Sladen, 1916a:271. Type species: Formicapis Genyandrena LaBerge, 1986:544. Type species: Andrena = clypeata Sladen, 1916 Osmia robusta Nylander, 1848, mackieae CockereU, 1937, by original designation. [Andrena], monobasic. Osmiini. Andreninae.

Foveosmia Warncke, 1991c:267. Type species: Heriades Geoperdita Cockerell and Porter, 1899:415. Type species: Perdita foveolatus Morawitz, 1868, by original designation. [Osmia]. chamaesarachae Cockerell, 1896, monobasic. [Perdita]. Osmiini. Panurginae. Mexalictiis Fraternubombus Skorikov, 1922b: 156. Type species: Apathiis Georgealictiis Packer. 1993:1656. Type species: fratermis Smith, 1854, by designation of Frison, 1927:63. polybioides Packer, 1993, by original designation. [Alpigenobombus], Bombini. [Mexalictus], Halictini.

1 943a: 146. nwmbiica Friesea Schrottky, 1902a:418, not Dalla Torre, 1895. Type spe- Geotrigona Moure, Type species: Trigona cies: Friesea brasiliensis Schrottky, 1902 = Camptopoeum Smith, 1863, by original designation. [Trigona]. Meliponini. 26 Scientific Papers, Natural History Museum, The University of Kansas

Gephyrohylaeus Michener, 1965a: 138. Type species: Heterapis Gmithanthidium Pasteels. 1969a:92. Type species: Anthidium sandiiccmensis Cockerell, 1919, by original designation. prionognathuni Mavromoustakis, 1935, by original designa- Hylaeinae. tion. Anthidiini. Glossadufourea Ebmer, 1993:32. Type species: Dufourea Gnatlumthidium Urban, 1994b:337, not Pasteels, 1969. Type Gnathanthidiwn longiglossa Ebmer, 1993, by original designation. species: sakagamii Urban, 1992, by original See Michanthidiuin. Anthidiini. Rophitinae. designation. G«fl//;/rt.s Robertson, 1903d: 173. 174. 175. Mi/hwJh Glossalictus Michener, 1965a: 173. Type species: Halictiis Type species: bella Nomadini. etheridgei Cockerell, 1916, by original designation. Cresson, 1863, by original designation. [Lasioglossum]. Halictini. Gnathocera Provancher, 1882:232. not Kirby, 1825. Type species: Guathocera cephalica Provancher. 1882 = Megachile GlossamegiUa Brooks, 1988:512. Type species: Anthophoni 1837, monobasic. See with mesopynhu Cockerell, 1930, by original designation. pugnatus Say, Sayapis. Isotypic Ceratias by synonymy. Megachilini. [AmegillaJ. Anthophorini. Gnathodon Robertson, 1903a: 168, not Oken, 1816. etc. Type Glossodialictiis Pauly, 1984a:704.Type species: Glossodialictits species: Megachile georgica Cresson, 1878, monobasic. See wittei Pauly, 1984. by original designation. Halictini. Sarogaster. Megachilini. Glossopasipliae Michener, 1989:643. Type species: Leioproctus Gnatliopasites Linsley and Michener. 1939:272. Type species: plaumanni Michener, 1989, by original designation. Philereiinis fulviventris Cresson, 1878, by original designa- [Leioproctus]. Colletinae. tion. Isotypic with Neopasites. Biastini. Glossoperdita Cockerell, 1916c:43. Type species: Glossoperdita Gnathoprosopis Perkins. 1912:104. Type species: Prosopis pelargoides Cockerell. 1916, monobasic. Panurginae. .xanthopoda Cockerell, 1910 = P. euxantha Cockerell, 1910, Glossosmia Michener, 1943:84. Osmia Type species: singularis by original designation. Hylaeinae. Morawitz, 1875, by original designation. [OsmiaJ. Osmiini. Gnathoprosopoides Michener, 1 965a: 1 27. Type species: Prosopis Glossura Cockerell, 1917b:144. Type species: Euglossa eburniella Cockerell, 1912 = Prosopis philoleucus Cockerell, piliventris Guerin-Meneville, 1845, by original designation. 1910, by original designation. [Hylaeus]. Hylaeinae. Euglossini. [Euglossa]. Gnathosmia Robertson, 1903a: 165. Type species: Osmia Glossurella Dressier. 1982:131. Type species: Euglossa georgica Cresson, by original designation. Osmiini. Moure, 1970. bursigera by original designation. [Euglossa]. Gnathoxylocopa Hurd and Moure, 1963:182. Type species: Euglossini. Xylocopa sicheli Vachal. 1898. by original designation. GlossiimcoUetes Michener, 1965a:60. Type species: Leioproctus [Xylocopa]. Xylocopini. bilobatus Michener, 1965. by original designation. Giiathylaeus Bndwell, 1919:126, 133. Type species: Gnaf/2y/aeM.y Colletinae. [Leioproctus]. willianisi Bridwell, 1919, by original designation. Hylaeinae. Glossuropoda Moure, 1989c:387. Type species: Euglossa Gonandrena Viereck. 1917:390. Type species: Andrena intersecta Latreille, 1837, by original designation. [Euglossa]. persiiuulata Viereck, 1917, monobasic. [Andrena]. Euglossini. Andreninae.

Glyphaiulrena Hedicke, 1933:2 1 2. Type species: Apis carbonaha Gongyloprosopis Snelling, 1982:16. Type species: Prosopis Linnaeus, 1767, by original designation. [Andrena]. cruenta Vachal, 1910, by original designation. [Hylaeus]. Andreninae. Hylaeinae.

Glyptapis Cockerell, 1909d:314; also described as new by Goniocolletes Cockerell, 1 907c:23 1 . Type species: Goniocolletes Cockerell, 1909e:13. Type species: Glyptapis mirabilis morsus Cockerell, 1907, by original designation. Colletinae. monobasic and Cockerell, Cockerell, 1909, designated by Graecandrena Warncke, 1968:61. Type species; Andrena 1909e:14. FossiL gracella Warncke, 1965, by original designation. [Andrena]. Glyptobasia Moure, 1941a:98, replacement for Glyptobasis Andreninae.

1940. chloera 1 940, Moure, Type species: Augochlora Moure, Grafanthidiuni Urban, 1995b:435. Type species: Grafanthidium autobasic. [Augochlora]. Augochlorini. ainazonense Urban, 1995, by original designation. Anthidiini.

Glyptobasis Moure, 1940:48, not M'Lachlan. 1871. Type spe- Grafella Mitchell, 1980:56. Type species: Pseudocentron cies: Augochlora chloera Moure, 1940, by original designa- crotalariae Schwimmer, 1980, by original designation. tion. See Glyptobasia. [Augochlora]. Augochlorini. [Pseudocentron]. Megachilini.

Glyptochlora Moure, 1959: 188. Type species: Megalopta onuitu Greeleyella Cockerell, 1904d:235. Type species: Greeleyella Smith, 1879, by original designation. [Augochloropsis]. heardsleyi Cockerell, 1904, by original designation. Augochlorini. Panurginae.

Glyptocoelioxys Mitchell, 1973: 92. Type species: Coelioxys Gressittapis Hirashima, 1978:65. Type species: Palaeorhiza vidua Smith 1854, by original designation. [Coelioxys]. mirauda Hirashima, 1978, by original designation. Megachilini. [Palaeorhiza]. Hylaeinae. Genus-Group Names of Bees 27

is an error for Ginnoceras Cockerell, 1907b:65. Type species: Gronoceras zonata" by Taschenberg, 1883:79, probably wellmani Cockerell. 1907 - Megachile honihlfonnis Habrophora ezonata Smith, 1854. Gerstaecker, 1857, by original designation. Megachilini. Hackeriapis Cockerell. 1922b:267. Type species: Megachile rhodura Cockerell. 1906. Grosapis Mitchell. 1980:46. Type species: MegacliiU' cockerelli by original designation. Rohwer. 1923. by original designation. |Eumegachile]. [Megachile]. Megachilini. Megachilini. Hadrorhiza Hirashima. 1980:108. Type species: Prosopis

Smith. 1 863. ]Palaeorhiza]. Gi(l(i)itliiiliiii)i Pasteels, 1969a: 101. Type species: Anthidiiim imperialis by original designation.

ciii,iiiilivc'nlre Morawitz, 1888. by original designation. Hylaeinae. [Anthidium]. Anthidiini Haetosmia Popov, 1952:104. Type species: Osmia latipes 1875 = Osmia 1875, Gundlachia Cresson, 1865:195. not Pfeiffer. 1850. Type spe- Morawitz. brachyiira Morawitz, Osmiini. cies: Centris ? conuita Cresson, 1865. monobasic. See monobasic. ]Anthocopa]. Heterocentris. Centridini. Halictanthrena Ducke. 1907:364. Type species: Halictanthrena Ducke, 1907, monobasic. Colletinae. Gymmmdrena Hedicke. 1933:212. Type species: Apis ihoracica nuilpighiacearuni Fabricius. 1775, by original designation. [Andrena]. Halictilliis Moure, 1947a:7. Type species: Chloralictus loureiroi Andreninae. Moure, 1941, by original designation. Augochlorini.

Gynuuis Spinola, 1808:9. Type species: Apis aterrima Panzer. Halictoides Nylander. 1848:195. Type species: Halicioides 1798, not Christ. 1791 = Apis pitnctidutissima Kirby, 1802. dentiventris Nylander. 1848. by designation of Cockerell and monobasic. Isotypic with Ceniplastes. Stelis. Tmclnisa Jurine, Porter, 1899:420. Rophitinae. and Anthidiini. Gywdroma Klug. Halictomorpha Schrottky, 191 1:81. Type .species: Halictomorpha 1807b:225. 191 Halictini. Gymdroma Klug, 1807. in Illiger, 1807:198: Klug. phaedra Schrottky. 1, by original designation. aterrima Panzer, 1798. not Christ. 1791 = Type species: Apis Halictonomia Pauly, 1980:123. Type species: Halictus 1802, Sandhouse, Apis punctidatissima Kirby, designated by decemmaculatus Friese, 1900. by original designation. who considered as monoba- 1943:555, incorrectly Gyrodroma [Nomia]. Nomiinae. sic: two species are listed by Klug in Illiger. 1807, which has Halictus Latreille, 1804:182. Type species: Apis quadricincta 1 Traclnisa page priority over Klug, 807b. Isotypic with Stelis, Fabricius, 1776, by designation of Richards, 1935:170. Apis Jurine, and Gymnus. Anthidiini. rubicunda Chhst. 1791, designated by Curtis, 1833:pl. 448a, Thomson, 1872:259. not 1807. Gyrodroma Klug. Type species: was not an originally included species. A discus.sion of type = Hcriades 1848 Heriades rapuncitli is in Omitted from that dis- ni}>ricornis Nylander, designations Michener ( 1978a). Lepeletier. 1841. by designation of Cockerell, 1925b:205. See cussion was Westwood's (1840a:84) and Cockerell's Osmiini. Gyrodromella. (1931f:529) designation of Apis se.xcincta Fabricius. 1775, noted in that discussion, Gyrodromella Michener. new name, replacement for Gyrodroma as the type species of Halictus. As A. se.\ci?icta was not an included Blanchard, Thomson. 1 872. Type species: Heriades nigricornis Nylander, originally species. Panz." as = 1 1 and 125 (1847), "Halictus 1 848 Heriades rapunciili Lepeletier, 84 . autobasic 1849:pl. designated flavipes the of the He could have meant present designation. Gyrodromella is considered a subgenus type species genus. Apis a dubium to Ebiner of Chelostoma. [Chelostoma]. Osmiini. flavipes Panzer, 1798, nomen according (1988). Blanchard. however, was careless about the authors" HabraUctelhis Moure and Hurd, 1982:46. Type species: Halictus names placed after species names in the explanations of plates, aMrafMs Ashmead, 1900, by original designation. Halictini. and probably he actually meant to cite the well-known Apis Hahralictus Moure, 1941a:59. Hahralictiis Type species: flavipes Fabricius, 1787 {=Apis tumulorum Linnaeus, 1758), Moure, 1941, designation. Halictini. of flavopictus by original a species of the subgenus Seladonia. as the type species Habromelissa Hirashima and LaBerge, 1964, //; Hirashima, Halictus. Fortunately, the name Halictus, with Apis

1 1 as the is on the 1 964:7 1 . Type species: Andrena omogensis Hirashima. 953, quadricincta Fabricius, 776, type species, in by original designation. [Andrena]. Andreninae. Official List of Generic Names Zoology (Commission 1713, 1993). Halictini. Habrophora Smith, 1854:318, not Erich.son, 1846. Type spe- Opinion = cies: Habriiphora ezonata Smith. 1854 Tctralonia tarsata Halterapis Michener. 1969: 289. Type species: Allodape Spinola. 1838, inversed autobasic, because of type designa- nigrinervis Cameron. 1905, by original designation. tion for replacement name, Habropoda. by Patton, 1 879a:477. Allodapini. silvai Anthophorini. Haniatothri.x Urban. 1 989: 121. Type species: Hamatothrix Urban, 1989, Eucerini. Habrophorula Lieftinck, 1974:217. Type species: Habropoda by original designation. nubilipennis Cockerell, 1930. by original designation. Haplocoelioxys Mitchell, 1973:85. Type species: Coelioxys Anthophorini. me.xicana Cresson. 1 878. by original designation. [Coelioxys]. Habropoda Smith. 1854:320, replacement for Habrophora Megachilini. 1934a:446. Rliinochaetula Smith, 1854. Type species: Habrophora ezonata Smith, 1854 Haplomelitta Cockerell. Type species: = Cockerell, 1932, Tetralonia tarsata Spinola, 1 838, by designation of Patton. ogilviei by original designation. of "H. 1 879a:477. Anthophorini. The subsequent designation Dasypodinac. 28 SciENTinc Papers, Natural History Museum, The University of Kansas

227. Haplosamba Michener, 1981:66. Type species: Haplomelitta Hemisia Klug, 1807b:213, Type species: Apis tridentata Michener, 1981, by original designation. haemorrhoidalis Fabricius, 1 775, by designation of Cockerell, [Haplomelitta]. Dasypodinae. 1906d:105. Isotypic with Centris. Centridini. Haujfapis Armbruster. 1938:37. Type species: Hauffapis Hemisiella Moure. 1945c:407. Type species: Apis lanipes Fab-

= 1 1 93 , scheuthlei Armbruster, 1 938 Apis armbmsteri Zeuner, ricius, 1775, by original designation. Centridini. of Zeuner and Manning, 1976:243. Haujfapis by designation Henicohesma Exley, 1968b: 199. Type species: Brachyhesma is not a valid name because no type species was designated macdonaldensis Exley, 1968, by original designation. by Armbruster; original type designation is required for ge- [Brachyhesma]. Euryglossinae. nus-group names proposed after 1930 [Code. 3rd ed.. Art. Herbstiella Friese. 1 9 1 6: 1 68. not Stimpson. 1871. Type species: 13(c)]. The subsequent type designation by Zeuner and Man- Herhstiella chilensis Friese, 1916, monobasic. See Kelita. ning was in the synonymy oiApis and thus does not validate the name as Hauffapis Zeuner and Manning [Code, 3rd ed.. Brachynomadini. Art. 1 1(c)]. Fossil. Apini. Heriades Spinola, 1808:7. Type species: Apis truncorum 1810:439. A subse- Helicosmia Thomson, 1872:233, no included species; Linnaeus, 1758. designated by Latreille, was Heriades 1841. Schmiedeknecht, 1885:22. included species. Type species: quent designation rapunculi Lepeletier, 126 Apis aurulenta Panzer. 1799, by designation of Michener. by Blanchard, 1849:pl. (1846). Taschenberg, 1883:62, 1941:163. [Osmia]. Osmiini. listed two species names without indicating that they are synonyms and therefore failed to designate a type species. Isotypic Heliomelissodes LaBerge, 1956: 1 172. Type species: Melissodes with Trypetes. Osmiini. desponsa Smith, 1854, by original designation. [Melissodes]. Eucerini. Heriadopsis Cockerell, 1931b:338. Type species: Heriadopsis striarulus Cockerell, 1931. by original designation. Heliophila Klug, 1807, in Illiger, 1807:197; Klug. 1807b:227. Type species: Apis bimaculata Panzer, 1798. monobasic. Megachilini. Heliophila is attributed in the literature to Klug. It was pub- Hesperandrena Timberiake, 1949. in Lanham, 1949:208. Type in consecutive articles in the same and thus lished two journal species: Andrena escondida Cockerell, 1938, by original des- The first article was who attrib- simultaneously. by Illiger. ignation. [Andrena]. Andreninae. uted the name to Klug; the second was by Klug. Dalla Torre Hesperapis Cockerell, 1898a: 147. Type species: Hesperapis (1896) listed both references under the species bimaculata elegantula Cockerell, 1898. monobasic. Dasypodinae. but attributed the genus-group name to Klug, and may there- Michener, 1965a:75. fore be regarded as the first reviser, who decided the author- Hesperocolletes Type species: Michener. 1965, ship of Heliophila. Anthophorini. HespemcoUetes douglasi by original designation. CoUetinae. Hemicoelioxys Pasteels. 1968b: 133. Type species: Coelioxys = 1939b:5. grac/7/5 Pasteels. 1968 (not Schenck. \%6%) Hemicoelioxys Hesperonomada Linsley, Type species: Hesperonomada gracillima Pasteels. 1977, by original designation. melanantha Linsley, 1939. by original designation. [Coelioxys]. Megachilini. Brachynomadini. Hemicotelles Toro and Cabezas, 1977:46. Type species: Hesperoperdita Timberiake. 1954:374. Type species: Perdita Lonchopria ruizii Herbst, 1923. by original designation. ntficaiida Cockerell. 1916. by original designation. [Perdita]. CoUetinae. Panurginae.

Hemidiellum Pasteels, 1972:112. Type species: Eoanthidium Heteranthidium Cockerell, 1904b:292. Type species: Anthidium semicahnatum Pasteels, 1972, monobasic. [Eoanthidium]. dorsale Lepeletier. 1841, by original designation. Anthidiini. [Anthidium]. Anttiidiini.

Hemieucera Sitdikov and Pesenko, 1988:88. Type species: Heterapis Cockerell. 191 la: 140. not Linston. 1889. Type spe- Eucera paraclypeata Sitdikov. 1988, by original designation. cies: Heterapis delicata Cockerell. 191 1. by original desig- [Eucera]. Eucerini. nation. See Heterapoides. Hylaeinae. Hemihalictus Cockerell, 1897a:288. Type species: Pamirgus Heterapoides Sandhouse, 1943:557, replacement for Heterapis lustrans 1897. Halictini. Cockerell, by original designation. Cockerell. 1911. Type species: Heterapis delicata Cockerell, Heminomada Cockerell, 1902, in Cockerell and Atkins, 1902:42. 1911. autobasic. Hylaeinae. footnote. Nomada obliterata Cresson, 1863, Type species: Hetereucera Tkalcu, 1978:167. Type species: Eucera hispana Nomadini. by original designation. [Nomada]. Lepeletier, 1841. by original designation. [Eucera]. Eucerini. Hemiosmia Tkalcu, 1975a:34. Osmia Type species: argyropyga Heterocentris Cockerell. 1 899a; 14, replacement for Gundlachia 1879, Osmiini. Perez, by original designation. [Osmia]. Cresson, 1865. Type species: Centris ? comuta Cresson, 1865 = with Hemirhiza Michener. 1965a: 147. Type species: Palaeorhiza Centris difformis Smith. 1854. autobasic. Isotypic melliceps Cockerell, 1918, by original designation. Rhodocentris by synonymy. Centridini. Hylaeinae. Heterocolletes Rayment, 1935:184. Type species: Paracolletes Hemisia Klug, 1807. in Illiger, 1807:198, nomen nudum. capillatus Rayment. 1935, by original designation. Centridini. [Paracolletes] CoUetinae. Genus-Group Names of Bees 29

Heteroediscelis Toro and Moldenke. 1979:112. Type species; Holmbergeria Jorgensen, 1 9 1 2: 1 00. Type species: Holmbergeria monobasic. Colletinae. Chilicola numtagiia Toro and Moldenke. 1979, by original cristariae Jorgensen, 1912, designation (misspelled Heteroesdiscelis in heading but cor- Holmbergiapis Cockerell, 1918b:36, replacement for Scirtetica rect elsewhere). Xeromelissinae. Holmberg, 1903. Type species: Scirtetica antarctica 1903, autobasic. See Alloscinetica. Eucerini. Heterohesma Michener. 1965a:97. Type species: Stilpnosoma Holmberg, clypeata Rayment. 1954. by original designation. Holohesma Michener, 1965a: 102. Type species: Stilpnosoma Euryglossinae. semisericea Cockerell. 1905 = Sericogaster fasciatus Westwood, 1835, Isotypic with 1 . designation. Heteroinegachile Rebmann. 1 970:4 Type species: Chalicodoma by original Euryglossinae. lefehvrei Lepeletier. 1841, by original designation. Sericogaster by synonymy. [Megachile]. Isotypic wilh Allochalicodowa. Megachilini. Holonomada Robertson, 1903d: 174. 175, 176. Type species: Nomada Cresson, 1863, original designation. Heteropeniita Timberlake, 1954:365. Type species: Perdita superba by Nomadini. rhodogastni Timberlake, 1954. by original designation. emendation of [Perdita]. Panurginae. Homachthes Dalla Torre, 1896:499, unjustified Omachthes Gerstaecker, 1 869. Ammobatini. Hetewrhiza Cockerell, 1929b;316. Type species: Palaeorhiza melaniira Cockerell. 1910. by original designation. Homalictus Cockerell. 1919b: 13. Type species: Halictus [Palaeorhiza]. Hylaeinae. taclobanensis Cockerell, 1915, by original designation. [Halictus]. Halictini. Heterosarus Robertson, 1918a:91. Type species: Calliopsis Moure. 1961 :200. parvus Robertson. 1892, by original designation. Panurginae. Homotrigomi Type species: Trigona fimbiiata Smith. 1857. by original designation. Meliponini. HetemsniiaTkaku. 1993a:56. nomen nudum. [Osmia]. Osmiini. Honanthidium Pasteels, 1969a:88. Type species: Anthidium Hetewstelis Timberlake. 1941:125. Type species: Stelis honestum Cockerell. 1936. by original designation. anthidioides Timberlake. 1941. by original designation. Anthidiini. [Stelis]. Anthidiini. Hoplandrena Perez, 1 890: 1 70. Type species: Melitta trimmerana Heterothgona Schwarz. 1939a;96. Type species: Trigona itama Kirby, 1802,by designation of Hedicke, 1933:214. [Andrena]. Cockerell. 1918. by original designation. [Trigona]. Andreninae. Meliponini. Hoplepicharis Moure, 1945b;300. Type species; Epicharis Hexcmthedu Ogloblin. 1948:172. Type .species: Hexantheda fasciata Lepeletier and Serville, 1825. by original designa- missionlca 1948. Ogloblin. by original designation. tion. Centridini. Colletinae. Hopliphora Lepeletier. 1841:458. Type species; Mesocheira Timberlake. 1954:416. Type species: Perdita ignota Hexaperdita velutina Lepeletier and Serville. 1825. monobasic. Isotypic Cockerell. 1896. [Perdita]. by original designation. with Oxynedys by synonymy. Ericrocidini. Panurginae. Hoplitella Cockerell. 1910d; 169. not Levinsen. 1909. Type spe- Hexataenhes Pesenko, 1984:348. sexcincta = Type species: Apis cies; Hoplitella pentamera Cockerell, 1910 Ashmeadiella Halictini. Fabricius, 1775. by original designation. [Halictus]. //ovvar^/ Cockerell, 1910, monobasic. See Hoplitina. Osmiini. and Michener. 1937:77. 81. Hexepeolus Linsley Type species: Hoplitina Cockerell. 1913:34, replacement for Hoplitella 1937 = Hexepeohis mojavensis Linsley and Michener, Cockerell, 1910. Type species; Hoplitella pentamera Hexepeolus rhodogyne Linsley and Michener. 1937, by origi- Cockerell, 1910 = Ashmeadiella howardi Cockerell. 1910. nal designation. Hexepeolini. autobasic and by original designation. Osmiini.

Hexosmia Michener. 1943:74. Type species: Osmia copelandica Hoplitis Klug, 1807b:225. Type species; Apis adunca Panzer, Cockerell, 1908, by original designation. [Anthocopa]. 1798, monobasic. Isotypic with Ctenosmia. Osmiini. Osmiini. assimilis Hoplitocopa Lieftinck, 1 955:27. Type species; Xylocopa Hofferia Tkalcu, 1984a: 10. Type species: Chelostoma Ritsema. 1880. by original designation. [Xylocopa[. schmiedeknechti Schletterer, 1889, by original designation. Xylocopini. Osmiini. Hoplocolletes Michener. 1965a:42. Type species: Dasycolletes Holandrena Perez, 1890:176. Type species: Melitta labialis ventralis Friese, 1924. by original designation. [Leioproctus]. Colletinae. Kirby, 1 802, by designation of Hedicke. 1933:214. [Andrena]. Andreninae. Hoplonomia Ashmead, 1904b;4. Type species; Hoplonomia Ashmead, 1904, of Cockerell, Holcomegachile Moure. 1953c: 119. Type species: Megachile quadrifasciata by designation 1910f;289. Nomiinae. (;/>(#( Schrottky. 1913, by original designation. [Megachile]. Megachilini. Hoplopasites Ashmead, 1898:284. Type species: Phileremus Cresson, 1879, by original designation. Dioxyini. Holcopasites Ashmead. 1899a:82. no species. Crawford (1915) productus included species. Type species: Phileremus illinoiensis Hoploprosopis Hedicke, 1926:415. Type species; Prosopis Robertson. 1891. by designation of Crawford, 1915:123. quadricornis Hedicke, 1926, by original designation. Ammobatoidini. [Prosopis]. Hylaeinae. 30 Scientific Papers, Natural History Museum, The University of Kansas

1904b:292. Anthidium Hoplosmia Thomson, 1872:233. no included species; Hypanlhidium Cockerell. Type species: Smith, 1879, Schmiedeknecht. 1884:23. included species. Type species: flavomarginatum by original designation. of Michener, Anthidiini. Apis spiimlosa Kirby. 1802, by designation 1941:161. lOsmia]. Osmiini. Hyperandrena Pittioni, 1948:58. Type species: Apis hicolorata included Rossi. 1790, [Andrena]. Andreninae. Hoplostelis Dominique, 1898:60. no valid species; by original designation. Griswold and Michener, 1988:36, included species. Type spe- Hyphesma Michener. 1965a: 103. Type species: Pachyprosopis = hivittatum cies: Stelis abnonnis Friese, 1925 Anthidium atromicans Cockerell. 1913. by original designation. of Griswold and Michener. Cresson, 1878, by designation Euryglossinae. 1988:36. Isotypic with Odontostelis. Anthidiini. Hypnorobombus Quilis, 1927:97. Type species: Apis hypnorum Hoploxylocopa Hurd and Moure, 1963:260. Type species: Linnaeus, 1758, monobasic. [Bombus]. Isotypic with Xylocopa acutipennis Smith. 1854. by original designation. Pxrobombus. Anodontobombus. and Uncobombus. Bonibini. [Xylocopal. Xylocopini. Hypochrotaenia Holmberg. 1886b:234. 273. Type species: emendation of Hortibombus Skorikov, 1938a: 146, unjustified Hypochrotaenia parvula Holmberg, 1886, monobasic. Hortoboinbiis Vogl. 1911, Bonibini. Nomadini. 1911:56. hortonim Hortobombiis Vogt, Type species: Apis Hypomacrotera Cockerell and Porter, 1899:418. Type species: of Sandhouse. 1943:559. Linnaeus, 1761, by designation Hypomacrotera callops Cockerell and Porter, 1899, by origi- Bombini. [Bombus]. nal designation. Panurginae. Hylaeana Michener, 1954:28. Type species: Hylaeus panamensis Hypotrigona Cockerell. 1934b:47. Type species: Trigona 1954. designation. [Hylaeus]. Michener, by original gribodoi Magretti, 1884. by original designation. [Trigona]. Hylaeinae. Meliponini. 1896:51. emendation of Hylaeoides Dalla Torre, unjustified Icteranthidium Michener, 1948a;25. Type species: Anthidium 1853. Hyleoides Smith, Hylaeinae. limbiferum Morawitz. 1875, by original designation. mexiccma Hvlaeopsis Michener. 1954:27. Type species: Prosopis Anthidiini. 1869, Hylaeinae. Cresson, by original designation. [Hylaeus]. Idiomelissodes LaBerge. 1956:1027. Type species: Melissodes H\laeorhiza Michener, 1965a:141. Type species: Prosopis duplocincta Cockerell, 1905, by original designation. mtbilosa Smith. 1 853, by original designation. Hylaeinae. [Melissodes]. Eucerini. Hylaeosoma Ashmead. 1898:284. Type species: Hylaeosoma Idioprosopis Meade-Waldo. 19l4a:451. Type species: longiceps Ashmead, 1898, by original designation. Idioprosopis chakidiformis Meade-Waldo. 1914, by original Xeromelissinae. designation. Xeromelissinae. Hxlaeleroii Michener. 1965a: 126. Type species: Prosopis Immanthidlum Pasteels. 1969a:89. Type species: Anthidiimi Cockerell, pidchricnis Cockerell. 1915= Eiiiyglossa semirufa immaculutum Smith. 1854. by original designation. 1914, by original designation. [Hylaeus]. Hylaeinae. Anthidiini. anmdata because no Hylaeus Fabricius, 1793:302. Type species: Prosopis Impeifecta Mehely. 1935:48. 154. Invalid type spe- = Linnaeus, 1758, view Fabricius. 1 804 /l/?/.sflWH//a?a by designa- cies was designated. [Prosopis]. Sandhouse's (1943:560) 1798 = tion of Latreille, 1810:438. Sphex sigmita Panzer. that Popov (1939) designated a type species and thus vali- Melliims bipiinclatits Fabricius, 1798. designated by dated the name is incoiTect. Hylaeinae. Blanchard. 125 (1847) was not originally included. 1849:pl. //«/«/(//»Halictidae. Isotypic with Halictus. Halictinae. Halictini. Fabri- Hyleoides Smith, 1853:32. Type species: Vespa concinna liihalictus Warncke, 1975b:96. Type species: Hylaeus interrup- cius, 1775. by designation of Taschenberg, 1883:45. tus Panzer, 1798, by original designation. [Halictus]. Hylaeinae. Halictini. Ashmead, 1899b:376. error for Hylaeosoma Hyloeosoma excavata 1 96 1 : 1 79. Exoneura Ashmead. 1898. Xeromelissinae. Inquilma Michener, Type species: Cockerell. 1922, Allodapini. Anthidium by original designation. Hypaiuhidiodes Moure, 1947a:35. Type species: lonu'lissa Robertson. 1900:50. Type species: Andrena violae flavofasciatum Schrottky, 1902. by original designation. monobasic. Andreninae. Anthidiini. Robertson. 1891.

Hurd and Moure, 1 963: 1 1 6. Type species: Xylocopa Hypanihidioides Michener. 1948a:22; Moure and Urban, loxylocopa 1902. designation. 1975:837; Urban, 1993:28. error for //.vpc»?r/;iVf/oc?« Moure, chrysopoda Schrottky. by original 1947. Anthidiini. [Xylocopa]. Xylocopini. Genus-Group Names of Bees 31

Isclmoceni Shuckard. 1840:166, no included species; Smith, 1943:561. Vitzthum (1930) implied that Apis aestuans should be the of 1 854:284 included Ischnocera as a synonym of Cu'iilosclwliis, Linnaeus, 1758, type species Koptorthosoma, with the single species latreillii Lepeletier. Type species: but his wording does not unambiguously make a type desig- Melissoda latreillii Lepeletier. 1841 = Acantliopiis goiyi nation. Both gabonica and aestuans were originally included Remand, 1840, first included species. Ctenioscheliis Romand species and both fall in the subgenus Koptortosoma as now was published in November. 1840. [schnocerii Shuckard prob- understood. Xylocopini. in December. 1840. for in the of the ably appeared preface Kozlovibombus Skorikov. 1922b; 152. Type species: Bombus volume in which this name was is a note dated "Nov. proposed kozlovi Skorikov, 1909 = Bombus keriensis Morawitz, 1886, 1 840." and a of the volume in the Natural History Mu- copy by designation of Sandhouse, 1943:561. Bombini. seum. London, bears the handwritten notation "Dec. 1840" Kozlowibombus Bischoff, 1936:10, unjustified emendation of (information thanks to Julie Harvey. Librarian). Clcnioschehis Kozlovibombus Skorikov, 1922. [Bombus]. Bombini. therefore appears to have priority, as well as universal usage 1962:300. Osmia tenuicornis since 1840. Isotypic with Ctenioscehis by synonymy. Kumobia Popov. Type species: Ericrocidini. Morawitz. 1875, by original designation. Osmiini. hepeolus Cockerell. 1907b:64. Type species: Isepeolus Kylopasiphae Michener. 1989:641. Type species: Leioproctus alhopictiis Cockerell, 1907 = Epeolus viperimis Holmberg. pruinosus Michener. 1989, by original designation. Colletinae. 1 886. monobasic. Isotypic with Palinepeolus and Calospiloma [Leioproctus]. by synonymy. Isepeolini. Laboriopsithyrus Frison. 1927:69. Type species: Bombus Isomalopsis Michener and Moure, 1957:434. Type species: laboriosus Fabricius. 1804. by original designation. The spe- Tetralonia niveata Friese, 1908, by original designation. cific name was misapplied by Frison and others, who consid- Exomalopsini. ered it to be a species of Psithyrus. See Citrinopsithyrus and

Williams ( 1991 ). [Psithyrus]. Anthophorini. Isosmia Michener and Sokal. 1957: 159. Type species: Anthocopa rubrella Michener. 1949. by original designation. Labrohalictus Pauly. 1981:719. Type species: Lasioglossum [Anthocopa]. Osmiini. saegeri Pauly, 1981, monobasic. [Lasioglossum]. Halictini. Jaxartiniila Popov. 1963:865. Type species: Jaxaniniila Laccohylaeus Houston. 1981:88. Type species: Prosopis malyshevi Popov, 1963. by original designation. Osmiini. cyanophila Cockerell. 1910, by original designation. [Hylaeus]. Hylaeinae. Kallobombiis Dalla Torre, 1 880:40. Type species: Apis someensis Fabricius, 1776, by designation of Sandhouse, 1943:561. Laesibombus Skorikov, 1938a: 145, unjustified emendation of [Bombus]. Isotypic with Soroeensibombiis. Bombini. Laesobomhus Kriiger, 1920 or Skorikov, 1922. Bombini.

Katamegachile Rebmann. 1970:43. Type species: Megachile Laesobombus Kriiger, 1920:350. Type species: Bombus laesus

niaiiiciita G'naud. 1 86 1 , by original designation. [Megachile]. Morawitz, 1875, monobasic. [Agrobombus]. Isotypic with Megachilini. Laesobombus Skorikov. Bombini.

Kelitci Sandhouse. 1943:561. replacement for Herbstiella Friese, Laesobombus Skorikov. 1 922b: 1 50. Type species: Bombus laesus 1916. Type species: Herbstiella chllensis Friese, 1916, Morawitz, 1875. monobasic. [Agrobombus]. Junior synonym autobasic. Brachynomadini. and homonym of Laesobomhus Kriiger. Isotypic with Laesobombus Bombini. Kelneriapis Sakagami, 1978(June):232. Type species: Trigona Kriiger. with eocenica Kelner-Pillault, 1970, monobasic. Isotypic Lagobata Smith. 1861:151. Type species: Lagobata diligens Kelnermelia. Fossil. Meliponini. Smith. 1861 = Ancyloscelis omata Sp'moXa, 1853. monoba- Kebiennelia Moure and Camargo, l978(Nov. 17): 565. Type sic. Tetrapediini. species: Trigona eocenica Kelner-Pillault. 1970. by original Lagripocla Vachal. 1909b:7. Latinization of Lagripode Lepeletier designation. Isotypic with Kelneriapis. Fossil. Meliponini. and Serville. 1828; these authors gave a few characters but included no 1 Unrecognizable. Kirbya Lepeletier, 1 84 1 : 145, not Robineau-Desvoidy, 830. Type species. species: Melitia tricincta Kirby. 1802. by designation of Lagripode Lepeletier and Serville. 1828:795. French vernacu- Sandhouse, 1943:561. Not replaced because synonym of lar; see Lagripoda Vachal. 1909. Unrecognizable. Melitta. with Melitta. Melittinae. Isotypic LambdopsisM€\\t\y. 1935:65, 171. Invalid because no type spe- Koptobaster Popov. 1939:168. error for Koptogaster Alfken. cies was designated. [Prosopis]. Hylaeinae. 1912. [Prosopis] Hylaeinae. Lcmibdopsis Popov. 1939:169. Type species: Melitta annularis Koptogaster M^en. 1912:23. Type species: Prosopis bifasciata Kirby. 1802, by original designation. [Prosopis[. Hylaeinae. Jurine. 1807. of Meade- Waldo. 1923:16. by designation Laminomada Rodeck, 1947:266. Type species: Nomada hesperia [Prosopis]. Hylaeinae. Cockerell. 1903. by original designation. [Nomada]. Koptorthosoma Dalla Torre. 1896:202. unjustified emendation Nomadini.

of 1 894. Koptortosoma Gribodo, Xylocopini. Lampralictus Pesenko, 1984:348. Type species: Halictus Koptortosoma Gribodo. 1894:271. Type species: Koptortosoma modermis Morawitz. 1876, by original designation. [Halictus]. gabonica Gribodo, 1894, by designation of Sandhouse, Halictini. 32 SciENTinc Papers, Natural History Museum, The University of Kansas

Lamproapis Cameron, 1902a:419. Type species: Lamproapis Leioproctus Smith. 1853:8. Type species: Leioproctus imitatiis maculipennis Cameron. 1902. monobasic. Nomadini. Smith, 1853, by designation of Cockerell, 1905a:348. Colletinae. Lamprocolletes Smith, 1853:10. Type species: Andrena Hedicke, 1933:215. Andrena chalybeata Erichson, 1851, by designation of Cockerell. Lepidandrena Type species: Thomson, 1870. 1905a:345. Isotypic with Nodocolletes by synonymy. ciirriingiila by original designation. Colletinae. [Andrena]. Andreninae. Lepidorhopalomelissa Wu. 1985:58. Nomia Lanthanella Michener and Moure, 1957:417. Type species: Type species: burmica Cockerell, 1920, by original designation. Lanthanomelissa complete! Michener and Moure, 1957, by [Rhopalomelissa]. Nomiinae. original designation. [Lanthanomelissa]. Tapinotaspidini. Lepidotrigona Schwarz, 1939a: 132. Type species: Trigona Lanthanomelissa Holmberg, 1903:418. Type species: nitidiventris Smith. 1857. by original designation. [Trigona]. Lanthanomelissa discrepans Holmberg, 1903, monobasic. Meliponini. Tapinotaspidini. Leptergates HolmhcTg, 1887b:224. nomen nudum. Eniphorini. Lapidaiiibombiis Skonkov, 1938a: 145. unjustified emendation Leptergatis Holmberg, 1903:422. Type species: Leptergatis of Lapidariobombus Vogt, 1911. Bombini. halictoides Holmberg, 1903. by designation of Lutz and 191 1:58. Lapidariobombus Vogt, Type species: Apis lapidaria Cockerell, 1920:592. Emphorini. Linnaeus, 1758, by designation of Sandhouse, 1943:562. Leptoglossa Friese, 1925a:9, not Klug, 1839. Type species: [Bombus]. Isotypic with Mekmobombus. Bombini. Leptoglossa paradoxa Friese, 1925 = Orphana inqidrenda 63. Lapponicobombus Quilis, 1927:19, 22, Type species: Apis Vachal, 1909, monobasic. See Orphana. Isotypic with of lapponica Fabricius, 1793. by designation Milliron. Ptoleglossa: isotypic with Orphana by synonymy. 1961:58. IBombus]. Bombini. Andreninae.

Larandrena LaBerge, 1964:304. Type species: Andrena Leptonietria Holmberg, 1903:409. Type species: Leptometria miserabilis Cresson, 1872, by original designation. [Andrena]. perexrae Holmberg, 1903, by designation of Brethes, Andreninae. 1910:213. Emphorini.

Largella Pasteels, 1965:534. Type species: Megachile semivestita Leptophanthiis Ruz and Toro, 1983:277. Type species: = Smith. 1853. by original designation. [Chalicodoma]. Psaenythia nigra Friese, 1916 (not Frie.se, 1908) Liphanthus nitidus and Megachilini. Ruz Toro, 1983, by original designation. [Liphanthus]. Panurginae. Larinostelis Michener and Griswold, 1994a:317. Type species: Mitchell, 1980:56. species: Megachile laeta Larinostelis scapiilata Michener and Griswold, 1994, by origi- Leptorachina Type Smith, 1853, original designation. [Pseudocentron]. nal designation. Anthidiini. by Megachilini. Lasanthidium Romankova, 1 988:26. Type species: Stelis malaisei Leptorachis Mitchell, 1934:301, 308. Type species: Megachile Popov, 1941, by original designation. Anthidiini. petulans Cresson. 1878, by original designation. [Megachile]. Curtis, 1833:448. Lasioglossum Type species: Lasioglossum Megachilini. tricingulum Curtis, 1833 = Melitta xanthopus Kirby, 1802. Lestis Lepeletier and Serville, 1828:799. Type species: Apis by original designation. Halictini. bomhylans Fabricius. 1775 (misidentified as Apis nmscaria Lasiiis Jurine, 1801:164. Type species: Apis qiiadrimaculata Fabricius, 1775; see Hurd and Michener, 1961 ). monobasic, Panzer, 1798, monobasic. Suppressed by Commission Opin- fixed by Commission Opinion 657 (1963). Xylocopini. ion 135 (1939), Direction 4. Isotypic with Dasymegilla. Lestrimelitta Friese, 1903a:361. Type species: Trigona liniao Anthophorini. Smith, 1863, monobasic. [Trigona]. Meliponini. Lasius Panzer, 1804b:tab. 16. Lasius salviae Type species: Leucandrena Hedicke, 1933:215. Type species: Apis sericea Panzer, 1804 = Smith, 1854, monoba- Anthophora crinipes Christ, 1791 (notForster, 111 \) = Melitta barbilabris Kirby,

sic. Commission 151 ( Direc- Suppressed by Opinion 1944). 1802, by original designation. [Andrena]. Andreninae.

tion 4. to 1 5 1 the correct for According Opinion , date Lasius Leuchalictits'Wamcke. 1975b:98. Type species: Apis leucozonia Panzer is [1801-1802]; see Literature Cited. Anthophorini. Schrank. 1781. by original designation. [Halictus]. Halictini. Griswold and Michener, 1988:27. Legnanthidiiim Type species: Leiicobombus Dalla Torre, 1 880:40. Type species: Apis terrestris Anthidiiini ridingsii Cresson, 1878, by original designation. Linnaeus, 1758, by designation of Sandhouse, 1943:564. [Trachusa]. Anthidiini. [Bombus]. Isotypic with Brennis Jurine. Bremiis Panzer, Bombus. and Terrestribombus. Bombini. Leimelissa Osychnyuk, 1984b:20. Type species: Andrena bairucumensis Morawitz. 1876, by original designation. Leuconomia Pauly, 1980:124. Type species: Nomia Candida [Andrena]. Andreninae. Smith, 1875, by original designation. [Nomia]. Nomiinae.

Leiopodus Smith, 1 854:252. Type species: Leiopodus lacertinus Leucosmia Robertson, 1903a: 166, 171. Type species: Osmia Smith, 1854, monobasic. Protepeolini. albiventris Cresson, 1864, by original designation. Osmiini. Genus-Group Names of Bees 33

Leiicostelis Noskiewicz. 1961:126, 132. Type species: Lithanthidiiini Cockerell. 1911d:225. Type species: Gymdroma omatiila Klug, 1807. by original designation (p. Litlumthidium pertriste Cockerell, 1911, monobasic. Fossil. Anthidiini. 132). [Stelis]. Lithosmia Alfken, 1935:188. Type species: Megachile villosa = Leurotrigona Moure. 1950a:244. Type species: Trigona miielleri Schenck. 1853, not Apis villosa Fabricius, 1775 Osmia Friese. 1900, by original designation. [Hypotrigona]. platycera Gerstaecker, 1869. [Osmia]. Isotypic with Meliponini. Arctosmia. Osmiini.

Lihelhilapis Cockerell. 1906a:42. Type species: Libellulapis Lithurge Latreille, 1825:463. Type species: Andrena conmta antiquorum Cockerell, 1906, monobasic. Fossil. Fabricius, 1787, monobasic. Latreille's original spelling (1825) has been regarded by many authors as French ver- Lieftinckella Hurd and Moure, 1963:286. Type species: Xylocopa nacular unavailable as a scientific name Fox, 1902). sinirhii Ritsema, 1876, by original designation. [Xylocopa]. (e.g.. Such authors use the emended Latinized form Xylocopini. Lithurgus Berthold (1827), published in a translation of Latreille's work Lioceratina Vecht, 1952:32. Type species: Ceratina flavopicta into German. The spelling Litharge, as noted by Fox ( 1902), Smith, 1858, by original designation. [Ceratina]. Ceratinini. was used by Lepeletier and Serville ( 1 828:795 ) but they were 1 833: 146. Liogastni Perty, Type species: Liogastra bicolorPerty, writing in French and used other French vernacular names. 1833 = bicolor and Serville, 1828, Rhathymus Lepeletier Lithurge was shown as the correct name by Sandhouse ( 1 943 ) monobasic. Isotypic with Colax and Rhathymus by synonymy. and has since been used by Michener (1944, 1951), Hurd Rhathymini. (1979), and others in major studies and catalogues. I now that the authors are cor- Liopodus Schulz, 1906:258, unjustified emendation of Leiopodus believe, however, using Lithurgus Smith. 1854. Protepeolini. rect. Lithurgini.

Liopoeodes Ruz. 1991:236. Type species: Calliopsis xenopous Lithurgomma Moure, 1949b:277. Type species: Lithurgomma Ruz, 1991. by original designation. [Calliopsis]. Panurginae. wagenknechti Moure, 1949. by original designation. Trichothurgus Moure and Lithurgomma were published si- Liopoeiim Friese, 1906b: 176. Type species: Camptopaeiim multaneously. Michener (1983) synonymized these names as hirsutulum Spinola, 1851, by designation of Sandhouse, first reviser and selected Trichothurgus as the generic name. 1943:564. [Camptopoeum]. Panurginae. Lithurgini. Lioproctiis Smith, 1 879:6. unjustified emendation of Leiopwctiis Fox, 1902:138. Smith, 1853. Colletinae. Lithurgopsis Type species: Lithurgus apicalis Cresson, by original designation. Lithurgini. Liosmia Thomson, 1872:233, no included species; Friese, Lithurgus Berthold. 1 827:467. Latinization of Lithurge Latreille. 191 lb:438, included species. Type species: Osmia claviventris 1825. Type species: Andrena cornuta Fabricius, 1787. Thomson, 1872. by designation of Michener, 1941:159. monobasic. Blanchard, 127 (1846). designated as [Osmia]. Osmiini. 1849:pl. 1 the type species Lithurgus fuscipennis Lepeletier, 184 , which Liothgmphis Sandhouse, 1943:564. incorrect subsequent spell- was not originally included. Lithurgini. ing of Liothgrapis Cockerell. 1932. Megacliilini. Litomegachile Mitchell. 1934:301 . 308. Type species: Megachile Cockerell, 1932:26, incorrect Liothgrapis subsequent spelling brevis Say, 1837, by original designation. [Megachile]. of Cockerell, 191 1. Liothyrapis [Coelioxys]. Megacliilini. Megachilini. Liothyrapis Cockerell, 1911b:246. species: Coelioxys Type L;fH;;?/(.v Ashmead, 1899a:77, error for L;'r/!iLipotriches Gerstaecker, 1857:460. Type species: Lipotriches annectens Michener. 1989. by original designation. = abdominalis Gerstaecker. 1 857 Sphecodes cribrosa Spinola, [Lonchopria]. Colletinae. 1843, monobasic. Nomiinae. LonchopriaWachal. 1905a:204. Type species: Lonchopria herbsti Lisotrigona Moure, 1961:194. Type species: Melipona cacciae Vachal, 1905 = Colletes zonalis Reed, 1892, monobasic. Nurse, 1907, by original designation. Meliponini. Colletinae.

Lissopedia Moure, 1994b:306. Type species: Tetrapedia Lonchorhyncha Michener, 1989:667. Type species: Diphaglossa globulosa Friese, 1899. by original designation. ecuadoria Friese, 1925, by original designation. Colletinae.

Tapinotaspidini Lophalictus Pesenko, 1986a: 125. Type species: Lasioglossum Lithandrena Cockerell. 1906a:44. Type species: Lithandrena acuticrista Pesenko, 1986, by original designation. saxorum Cockerell, 1906. monobasic. Fossil. [Lasioglossum]. Halictini. 34 SciENTinc Papers, Natural History Museum, The University of Kansas

Lophanthophora Brooks, 1988:464. Type species: Anthophora Macronomadopsis Rozen, 1958:93. Type species: Nomadopsis porterae Cockerell. 1900. by original designation. micheneri Rozen. 1958, by original designation. [Anthophora]. Anthophorini. [Nomadopsis]. Panurginae.

Lophopedia Michener and Moure, 1957:413. Type species: Macronomia Cockerell, 1917c:468. Type species: Nomia Tetrapedia pygmaea Schrottky. 1902. by original designation. platycephala Cockerell, 1917, by original designation. [Paratetrapedia]. Tapinotaspidini. [Nomia]. Nomiinae. 1809:no. 16. labiata Lophothygater Moure and Michener, 1955b:313. Type species: Macropis Panzer, Type species: Megilla Fabricius, 1805, monobasic. has been attributed to Tetralonia decorata Smith. 1879. by original designation. Macropis Eucerini. Klug (/;; Panzer, 1809) but as pointed out in Commission Opinion 1383 (1986), Panzer provided the description. Lophotrigona Moure, 1961 :205. Type species: Trigona canifrons Melittinae. Smith, 1857, by original designation. Meliponini. Macrotera Smith, 1853:130. Type species: Macrotera bicolor Loxoptiliis LaBerge, 1957:28. Type species: Loxoptiliis Smith, 1853, monobasic. Panurginae. longifellator LaBerge, 1957, by original designation. Eucerini. Macroterella Timberlake. 1954:360. Type species: Perdita mortuaria Timberlake. 1954, by original designation. Liicaselhis Schulz, 191 1:202, replacement for Lucasiits Dours, [Perdita]. Panurginae. 1 872. Type species: Halictiis clavipes Dours, 1 872, autobasic. Mwcrofera/w/j- Ashmead, 1899a:85. species: Perdita latior [Halictus]. Halictini. Type Cockerell, 1896, by original designation. Panurginae. Liicasielliis Cockerell, 1905d:272, replacement for Lucasius Maculonomia Wu, 1982:275. Type species: Nomia terminata Dours, 1872. Type species: Halictus clavipes Dours, 1872, Smith, by original designation. [Nomia]. Nomiinae. autobasic. [Halictus]. Halictini. Madagalictus Pauly, 1984b: 125. Type species: Halictus Lucasius Dours, 1872:350, not Kinahan, 1859. Type species: suarezensis Benoist, 1962, by original designation. Halictini. Halictus clavipes Dours, 1 872, by designation of Sandhouse, Madrosonm A^\\mtad. 1899a:94. for Smith. 1943:566. [Halictus]. See Lucasellus and Lucasiellus. lapsus Mydrosonia 1 879. Halictini. Dissoglottini. Maiella Michener, 1942:282, replacement for Orbitella Ma, L«rc;>//fl Cockerell. 1922d:l,notEnderlein. 1922. Type species: 1938. Type species: Xylocopa confusa Perez. 1901 . autobasic. Perdita opuntiae Cockerell. 1922. by original designation. [Xylocopa]. Xylocopini. [Perdita|. See Cockerellula. Panurginae. Malanthidium Pasteels, 1969a:26. Type species: Anthidium Lysicolletes Rayment, 1935:208. Type species: Paracolletes malaccense Friese. 1914, by original designation. Anthidiini. singularis Rayment, 1935, by original designation. [Paracolletes]. Colletinae. Malay apis Baker, 1995a:67. Type species: Andrcna chrysochersonesus Baker, 1995, by original designation. Machaeris Robertson, 1903b: 104. Type species: Sphecodes [Andrena]. Andreninae. stygius Robertson, 1893, by original designation. Halictini. Manthidiuni Pasteels, 1969a:43. Type species: Anthidium Macrocera Latreille. 1810:339. 439. not Meigen, 1803. Type binghanii Friese. 1901 (first described under the homonymous species: Euceru untennaia Fabricius, 1793 = Apis inalvae name Anthidium fraternum Bingham. 1897), by original des- Rossi, 1 790, by original designation. See Tetralonia. Eucerini. ignation. Anthidiini.

Macmgalea Cockerell. 1930f:291 . Type species: Allodape Can- ManueliaVachal, 1905b:25. Type species: Halictus gayi Spinola. dida Smith, 1879, by original designation. Allodapini. 1851, by designation of Daly, Michener, Moure, and

1 884: 1 7, not 1816. 1 987: 1 04. 1 Macroglossa Radoszkowski, Ochsenheimer, Sakagami, Sandhou,se ( 943 ) designated "Halictus = Type species: Macroglossa oribazi Radoszkowski. 1884 (Coiynura) gayi Spinola" as the type species, but this is mean- Macrocera analis Lepeletier, 1841, monobasic. See ingless since both Halictus gayi and Coiynura gayi were de- Macroglossapis. Eucerini. scribed by Spinola in the same work. Manueliini. Macroglossapis Cockerell. 1899c: 14, replacement for Margandrena Warncke, 1968:91. Type species: Andrena Macroglossa Radoszkowski, 1884. Type species: niarginata Fabricius. 1776, by original designation. Macroglossa oribazi Radoszkowski, 1 884 = Macrocera analis [Andrena]. Andreninae. 1841, autobasic. with Lepeletier, Isotypic Thygater by syn- Marghalictus Warncke, 1975b:95. Type species: Hylaeus Eucerini. onymy. marginellus Schenck. 1853, by original designation. Halictini. Macrohylaeus Michener, 1965a: 133. Type species: Prosopis [Halictus]. vidua 1853 = Smith, Prosopis alcyonea Erichson, 1842, by Martinapis Cockerell, 1929f:19, replacement for Martinella original designation. [Hylaeus]. Hylaeinae. Cockerell, 1903. Type species: Melissodes luteicornis Cockerell, 1896, autobasic. Eucerini. Macromegachile Noskiewicz, 1948:48. Type species: Apis lagopoda Linnaeus, 1761, by original designation. Mitchell. Martinella Cockerell, 1903c:450, not Jousseaume, 1887. Type 1980:23, incorrectly \hitd Apis maritinui Kirby, 1802, as the species: Melissodes luteicornis Cockerell, 1896, monobasic. type species. [Megachile], Megachilini. [Melissodes]. See Martinapis. Eucerini. Genus-Group Names of Bees 35

of Massanthidiiim Pasteels. 1969a:24. Type species: Trachiisa Megalochila Schulz, 1906:263, unjustified emendation fhivorufiila Pasteels. 1969, by original designation. Megachile Latreille. 1802. Megachilini. [Trachusa]. Anthidiini. Megalocilissa Schulz, 1906:243. unjustified emendation of 1853. Mastnicatobonihus Kriiger. 1917:66. Type species: Boinbus Megacilissa Smith, Caupolicanini. 1869 = Boinhus mustriicatus Gerstaecker, uiirflenii Megaloheriades Peters, 1984:366. Type species: Osmia sclmltzei Bombini. Radoszkowski, 1859, monobasic. [Bombus|. Friese, 1909, by original designation. Osmiini. Maximegachile Guiglia and Pasteels, 1961:27. Type species: Megalopta Smith. 1853:83. Type species: Megulopta idaliu Megachile maxillosa Guerin-Meneville, 1845. by original Smith, 1853, designated by Commission Opinion 788(1 966 ). designation. [Megachile]. Megachilini. The earlier type designation of Megalopta hituherculaia Smith, 1853, Cockerell, 1900a:374, is invalidated. Ma.xschwarzici Pagliano and Scaramozzino, 1990:5, unnecessary by replacement tor Megaiuviiia Cockerell, 1909: .see Michener, Augochlorini. 1992. Type species: Noiuia hinghaini Cockerell, 1909, Megaloprella Schrottky, I906a:3l2. Type species: Halictus autobasic. Meganomiinae. ochrias Vachal. 1904, by original designation. [Megalopta] . Augochlorini. Maynenomia Pauly, 1984d:698. Type species: Noinia inaynei Cockerell, 19.^7 - Nomia testacea Friese, 1914, by original Megaloptidia Cockerell, I900a:373. 374. Type species: designation. Nomiinae. Megalopta contradicta Cockerell, 1900, by original designation. [Megalopta 1. Augochlorini. Mecanthidhim Michener, 1942:278. Type species: Panintliidiiiii, sononim Michener. 1942. by original designation. Megaloptilla Moure and Hurd, 1 987:24 1 . Type species: Halictus [Paranthidium]. Anthidiini. callopis Vachal, 1911, by original designation. [Megommation]. Augochlorini. Mediocrolictiis Pauly, 1984b: 143. Type species: Halictus 1969:441. mediocris Benoist, 1962, by original designation. Megaloptilla Eickwort, Type species: Augochlora Cockerell, 1930, [Lasioglossum]. Halictini. ogilviei by original designation.

. [Megommation I Augochlorini. Megahombiis Dalla Torre, 1880:40. Type species: Bombiis Moure, 1959:183. Ugusticus Spinola. 1805 = Apis argillacca Scopoli, 1763, Megaloptodes Type species: Megalopta bitubercutata Smith, 1 853, Nomiinae. monobasic. [Bombus]. Isotypic with Odontohombus by syn- by original designation. onymy. Bombini. Megalosmia Schmiedeknecht, 1885:23. Type species: Osmia Morawitz, 1873, designation of Sandhouse, Megacemtina Hirashima, I971b:251. Type species: Ceratma grandis by = 1943:568. Osmiini. buuyssoiii \Acha\, 1903 Ceratiuu sciilpliirata Smith, 1854, acraensis by oiiginal designation. Ceratinini. Megamegilla Brooks, 1988:505. Type species: Apis Fabricius, 1793, by original designation. [Amegilla]. Megachile Latreille, 1 802a:434. Type species: Apis centiinciilaris Anthophorini. Linnaeus, 1758, by designation of Curtis, 1828:pl. 218. A I927a:42. Andrena subsequent designation, Xylocopa muraria Fabricius, 1804 Megandrena Cockerell, Type species: = Apis parietinaFomcroy. 1785, wasby Blanchard, 1840:408. euceliac Cockerell. 1927, by original designation. [Andrena|. Andreninae. Isotypic with Antliemois by synonymy. Megachilini. I909a:402. Noinia Megachileoides Radoszkowski, 1874b: 132. Type species: Meganomia Cockerell, Type species: = Cockerell, 1909, monobasic. Trachusa serratidae Panzer, 1 804 Apis hyssina Panzer. biiighami [Nomia|. 1798, by designation of Michener, 1995:375. Isotypic with Meganomiinae. Trachusa Panzer: isotypic with Diphysis by synonymy. Megauomioides Pesenko. 1983:183. Type species: Nomioides Anthidiini. karachensis Cockerell, 1911, by original designation. Nomioidinae. MegachiloidesMiichcW, 1924: 154. Type species: Meguchiloides [Ceylalictus]. oenotherae Mitchell, 1924, by original designation. Meganthidium Popov. I950a:3l5. Type species: Anthidium Meguchiloides Saussure (1890) is an incorrect sub.sequent christophi Morawitz, 1884 = Anthidium supcrbum spelling of Megachileoides Radoszkowski and does not pre- Radoszkowski, 1 876, by original designation. Anthidiini. Mitchell. occupy Megachiloides Megachilini. Megapis Ashmead, 1 904a: 1 20. Type species: Apis dorsaia Fab- Megachiloides Saussure, 1890:35, incorrect subsequent spell- ricius, 1793, by original designation. Apini.

of Radoszkowski, 1 874, see Michener, ing Megachileoides Megascirtetica Moure and Michener, I955b:264. Type species: 1995. Anthidiini. Macrocera mephistophelica Schrottky, 1902, by original des-

Megacilissa Smith. 1 853: 1 23. Type species: Megacilissa superba ignation. Eucerini.

Smith, 1 853 = 1 85 1 , monoba- Ccmpolicaiiafulvicollis Spinola, Megaxylocopa Hurd and Moure. 1963:151. Type species: Apis sic. Caupolicanini. frontalis Olivier. 1789, by original designation. [Xylocopa]. Megahoplitis Tkalcii, I993a:55. Type species: Osiiiia rigrina Xylocopini. Morawitz, 1872, Osniiini. by original designation. [Hoplitisj. Megella Pasteels, 1 965: 167. Type species: Megachile malimbaiui Megalobombus Schulz. 1906:267. unjustified emendation of Strand. 1911, by original designation. [Megachile]. Megubombus DAUlont, 1880. [Bombus]. Bombini. Megachilini. 36 Scientific Papers, Natural History Museum, The University of Kansas

Cockerell, 1929b:314. Meghylaeiis Type species: Palaeorhiza Melanomada Cockerell. 1903d:587. Type species: Nomada 1926 = gigantea Cockerell, Prosopisfijieusis Cockerell, 1909, grindeliae Cockerell, 1903, by original designation. by original designation. [Hylaeus]. Hylaeinae. [Nomada|. Brachynomadini. Fabricius. 1804:328. Fabri- Megilla Type species: Apis pilipes Melanomia Pauly. 1990:149; also described as new by Pauly, cius, ni5=Apisphimipes'Pa\Vd^. 1772, designated by Com- 1991:310. Type species: Nomia melanosoma Benoist, 1963, mission Opinion 1383 (1986). Megilla labiata Fabricius, by original designation. Nomiinae. 1804, was designated by Westwood, 1840a: 158, and Apis Melanosarus Mitchell, 1934:303. 307. Type species: Megachile acerx'orum Linnaeus, 1758, was designated by Richards, xylocopoides Smith, 1853, by original designation. 1935:172, as the type species of Megilla. These designations [Megachile]. Megachilini. would have made Megilla the correct name for Macropis or a Melanosmia Schmiedeknecht. 1885:19[885]. Type species: junior synonym of Bombus. respectively, although Apis OsmiafiiciformisLiAxQxWe. 1811 - Apis xanrhomelana acen'orum was interpreted as an Anthophora at the time of Kirby, 1802, by designation of Sandhouse, 1939:33. [Osmia]. Richard's work. Isotypic with Podaliriiis and Anthophora. Osmiini. Anthophorini. Melanostelis Ashmead, 1898:283. Melanosrelis Megomalopsis Michener and Moure, 1957:430. Type species: Type species; = berheli Aahmead. 1 898 5'?f//.yn/W Cockerell, 1898, E.xomalopsis diversipes Cockerell, 1949 = E.xomalopsis by original designation. Anthidiini. mellipes Cresson, 1878. by original designation. Exomalopsini. Melanrhidium Cockerell, 1 947: 106. Type species: Melanrhidium carri Cockerell, 1947, by original designation. Anthidiini. Megommation Moure, 1943b:479. Type species: Halictus insignis Smith, 1853, by original designation. Augochlorini. Melanrhidium Pasteels, 1969a:90. not Cockerell. 1947. Type species: Anrhidium concolor Friese. 1913, Mehelya Popov, 1939:167, not Csiki, 1903, replacement for by original designation. See Nigranthidium and Wamckeia. Anthidiini. Barbara Mehely, 1935. Type species: Prosopisfriesei Mfken, Melea 1904, by original designation. [Prosopis]. See Mehelyana. Sandhouse, 1943:569, replacement for Anrhemoessa Hylaeinae. Robertson. 1905. Type species; Anrhophora abrupra Say, 1 837, autobasic. Anthophorini. Mehelyana Sandhouse, 1943:569, replacement for Barbara Melecta Latreille. Mehely, 1935, and Mehelya Popov, 1939. Type species: 1802a:427. Type species: Apis punctata Fab- Prosopis friesei A\fker\. 1904. autobasic. Hylaeinae. ricius, 1775= Apis albifrons Forster, 1 77 1 , by designation of Latreille, 1810:439. Isotypic w'Wh Symmorpha. Melectini. Melaliphanthits Ruz and Toro, 1983:271. Type species: Melectoides Liphanrhus arratiis Ruz and Toro. 1983, by original designa- Taschenberg, 1883:75. Type species: Melectoides tion. [Liphanthus]. Panurginae. ieHf.vTaschenberg, 1883, by original designation. Isepeolini.

Melanapis Cameron, 1902a:420. Type species: Melanapis Melectomimus Linsley, 1939a:448. Type species: Melecta violaceipennis Cameron. 1902 = Andrena fuscosa Erichson. edwardsii Cresson. 1878. by original designation. [Melecta]. 1835, monobasic. Andreninae. Melectini.

Melandrena Perez, 1890: 170. Type species: Apis rhoracica Fab- Melectomorpha Linsley, 1939a:451. Type species: Melecta ricius, 1775. by present designation. Andrena morio Brulle. califomica Cresson. 1878. by original designation. [Melecta]. 1832, designated by Hedicke, 1933:216. was not among the Melectini. species included Perez. Andreninae. originally by [Andrena]. Melikerria Moure, 1992:34. Type species: Apis compressipes Melanempis Saussure, 1890:84. Type species: Phileremus ater Fabricius, 1804, by original designation. [Melipona]. Saussure, 1890, monobasic. [Phileremus]. Ammobatini. Meliponini.

Melaneurricharaea Tkalcu, 1993b:803. Type species: Megachile Meliphila Schrottky, 1902b:310. Type species: Meliphila = hohmanni Tkaku. 1983. by original designation. [Megachile]. ipomoeae Schrottky, 1902 Meliroma euglossoides Lepeletier Megachilini. and Serville. 1828, by original designation. Isotypic with Melitoma by synonymy. Emphorini. Melanoanrhidiitm Tkalcii. 1967:91. Type species: Anrhidium monraniim Morawitz, 1864, by original designation. Meliphilopsis Roig-Alsina, 1994:183. Type species: [Anthidium]. Anthidiini. Meliphilopsis melanandra Roig-Alsina, 1994. by original designation. Emphorini. Melanobombus Dalla Torre, 1880:40. Type species: Apis lapidaria Linnaeus, 1758, by designation of Sandhouse, Meliplebeia Moure, 1961:229. Type species: Trigona beccarii 1943:569. [Bombus]. Isotypic with Lapidariobombits. Gribodo, 1879, by original designation. Meliponini. Bombini. Melipona Illiger, 1806: 157. Type species; Apisfavosa Fabricius, Melanocenrris Friese, 1900:244, 246. Type species: Cenrris atra 1798. by designation of Latreille. 1810:439. Melipona fulva Friese, 1899, by designation of Sandhouse, 1943:569. Lepeletier, 1841, designated by Blanchard, 1849:pl. 129 [Centris]. Centridini. (1845), was not originally included. Meliponini.

Melanocoelioxys Mitchell, 1973:78. Type species: Coelioxxs Meliponorytes Tosi, 1896:352. Type species: Meliponorytes rolreca Cresson, 1878, by original designation. [Coelioxys]. succini Tosi, 1896, by designation of Sandhouse, 1943:570. Megachilini. Fossil. Meliponini. Genus-Group Names of Bees 37

Melipomila Cockerell. 1934b:47. Type species: Melipona Melitiirga Lepeletier and Serville. 1828:799, unjustified emen- bocaiulei Spinola, 1851. by original designation. [Trigona]. dation of Melitturga Latreille, 1809. Panurginae. Meliponini. Meliturgopsis Ashmead, 1899a:62, no included species; Melissa Smith. 1854:279. Type species: Mesacheira aziirea Cockerell, 1909c(May 13):414, includedaspecies. Type spe- Lepeletier and Serville. 1825. by designation of Sandhouse. cies: Einphoropsis murihirta imirina Cockerell. 1909, first 1943:570. Isotypic with Mesoplia. Ericrocidini. included species, designated by Sandhouse, 1943:572. The inclusion and of Anlliopliora aiirulento-caiidata Melissina Cockerel]. 191 le:670. Type species: Melissina viator designation was in and Cockerell. 1911. monobasic. Eucerini. Dours, 1869, by Vachal, 1909b:9, published July is junior to Cockerell ( 1909c). Anthopliorini. Melissoda Lepeletier. 1 84 1 :508. Type species: Melissoda latreillii = Friese, 1 903b:33. species: Meliturgida hruunsi Lepeletier. 1841 Acanthopiis goni Romand, 1840, monoba- Meliturgiila Type Friese, 1903, monobasic. sic. Isotypic with Ischnocera; isotypic with Ctenioschelus by Panurginae. synonymy. Ericrocidini. Meliwillea Roubik. Segura. and Camargo, 1997:102. Type spe- and Melissode Latreille, 1825:464, nomen nudum. French vernacu- cies: Meliwillea hivea Roubik, Segura, Camargo, 1997,

lar. Eucerini? by original designafion. Meliponini.

Melissodes Berthold. 1827:468, nomen nudum, Latinization of Mellitidia Guerin-Meneville. 1830:270. Type species: Andrena aiistralis Guerin-Meneville, 1830, monobasic. Nomiinae. Melissode Latreille. There is no certainty that these names are listed here to refer to Melissodes Latreille. 1829. They Melliturga Latreille. 1809:177. lapsus for Melitturga Latreille, avoid possible confusion. Eucerini? 1809. Panurginae.

Melissodes Latreille, 1829:354, no included species. Romand, Mendaciboinlnts Skorikov, 1914a: 125. Type species: Boinbits 1841:6, included Melissodes fonscolombei Romand, 1841; mendax Gerstaecker, 1869, by designation of Sandhouse, this specific name was suppressed by Commission Opinion 1943:572. [Bombus]. Bombini. 750 (1965). Type species: Melissodes lepheiiri Blanchard. Mermiglossa Friese, 1912: 188. Type species: Meriniglossa ritfa 1846. first valid included species and designated as type spe- Friese, 1912, monobasic. Panurginae. cies by Opinion 750. Taschenberg, 1883:78, subsequently Smith, 1853:33. species: Meroglossa designated Macrocera rustica Say, 1837, as the type species. Meroglossa Type monobasic. Eucerini. canaliculata Smith, 1853, Hylaeinae. Perkins, 1912:99. Type species: Meroglossa eu- Melissoptila Holmberg, 1884:119. Type species: Melissoptila Meroglossula Cockerell, 1910, monobasic. Hylaeinae. tandilensis Holmberg, 1884, monobasic. Isotypic with calypti [Meroglossa]. Thyreothremma and Thyreotremata by synonymy. Eucerini. Merrophites Wamcke. 1 979: 1 33. Type species: Dufomea inerceti Melitoina Berthold, 1827:468. nomen nudum, Latinization of Vachal, 1907, by original designation. [Rophites]. Rophitinae. Melitome Latreille. Emphorini? Mesanthidiellum Pasteels. 1969a:83. Type species: Mesanthidiellum amoemim Pasteels. 1969. des- Melitoina Lepeletier and Serville. 1828:529. Type species: by original Anthidiini. Melitoina eiiglossoides Lepeletier and Serville. 1 828. monoba- ignation. sic. Isotypic with Meliphila by synonymy. Emphorini. Mesanthidium Popov, 1950a:316. Type species: Anthidium 1930. Melitome Latreille, 1825:464, nomen nudum. French vernacu- pentagonum Gussakovskij, by original designation. Anthidiini. lar. Empliorini? Mesocheira and Serville, 1825:106. Melitribus Rayment, 1930a:217. Type species: Melitribus Lepeletier Type species: Melecta bicolor Fabricius. 1804. of greavesi Rayment. 1930. by original designation. Rayment. by designation 1883:72. Ericrocidini. 1930b:61. subsequently designated Gastropsis victoriae Taschenberg. Cockerell, 1906, as the type species. Stenotritidae. Mesoc/z/ra Agassiz, 1846:20; Schulz, 1906:257, unjustified emendation of Mesocheira and Serville, 1825. Melitta Kirby. 1802:117. Type species: Melitta tricincta Kirby. Lepeletier Ericrocidini. 1802, by designation of Richards, 1935:172. Isotypic with Melittinae. Kirbya. Mesonychium Lepeletier and Serville, 1825:107. Type species: coerulescens and Serville. 1825, Melittidia Dalla Torre. 1896:99. lapsus for Mellitidia Guerin- Mesonychiitin Lepeletier Meneville. 1830. Nomiinae. monobasic. Ericrocidini.

1 84 1 :457. Mesocheira azurea Melittoides Friese, 1921a: 177. Type species: Andrena melittoides Mesoplia Lepeletier, Type species: monobasic. with Me- Friese. 1899. by absolute tautonymy. Andreninae. Lepeletier and Serville. 1825. Isotypic lissa. Ericrocidini. Melittosmitliia Schulz, 1906:244, replacement for Sinithia 1838:112. Mesotrichia Vachal, 1897. Type species: Scrapter carinala Smith, 1862, Mesotrichia Westwood, Type species: monobasic. by designation of Cockerell, 1910b:358. Euryglossinae. torrida Westwood. 1838. Xylocopini. 1976:41. Oxaea Melittoxena Morawitz, 1873b: 154. Type species: Noinada Mesoxaea Hurd and Linsley, Type species: Oxaeinae. truncata Nylander. 1848, monobasic. Biastini. nigerrima Friese, 1912, by original designation.

1 Metadioxxs 1947:88. Melitturga Latreille. 1 809: 76. Type species: Eucera clavicomis Popov. Type species: Dioxys forinosa Latreille, 1806, monobasic. Panurginae. Morawitz, 1875, by original designation. Dioxyini. 38 Scientific Papers, Natural History Museum, The University of Kansas

Metallinella Tkalcu, 1966:200. Type species: Osmia Micranthophora Cockerell. 1 906d:66. Type species: Anthophora = atwcaenilea Schilling. 1 849 Eiicera hrevicornis Fabricius. curta Provancher. 1895. by original designation. 1798, by original designation. Osmiini. Anthophorini.

Metainegachile Tkalcu. 1967:102. Type species: Antliopliora Micrapis Ashmead, 1904a: 122. Type species: Apisflorea Fabri- albisecta Klug, 1817. by original designation. [Megachile]. cius, 1787, by original designation. Apini. Megachilini. Micraugochlora Schrottky. 1909b:483, no included species; 1969:89. MetapsaenythiaT\mbtr\ake. Type species: Calliopsis Schrottky, 1909a: 138, included one species. Type species: abdomiinilis Cresson, 1878, by original designation. Micraugochlora sphaerocephala Schrottky, 1909, first in- Panurginae. cluded species. Isotypic with Temnosomula by synonymy.

Metapsithynis Popov. 1 93 1 a: 1 35 . Type species: Apis campestris Augochlorini. Bombini. Panzer, 1801, by original designation. [Psithyrus]. Micreriades Mavromoustakis, 1958:444. Type species: Heriades Metasamba Michener, 1981:69. Type species: Haplomelitta parnesica Mavromoustakis, 1958, by original designation. fasciata Michener. 1981. by original designation. [Heriades]. Osmiini. [Haplomelitta]. Dasypodinae. Microcolletes Michener, 1965a:55. Type species: ParacoUetes Metatrachiisa Pasteels, 1969a:22. Type species: Anthidium halictifonnis Cockerell, 1916 (not Euryglossa halictiformis pendlebiiryi Cockerell, 1927, by original designation. Smith. 1879, homonym in Leioproctus) = Leioproctus [Trachusa]. Anthidiini. halictomimus Michener, 1965 = ParacoUetes minutus Cockerell, 1916, Metylaeus Bridwell, 1919:126, 131. Type species: Met\'laeus by original designation. [Leioproctus]. Colletinae. cribratus Bridwell, 1919, by original designation. Hylaeinae. Microdontura Cockerell, 1929b: 322. Microdontura Metziella Michener, 1942:273. Type species: Prosupis patens Type species: mellea Cockerell, 1929, monobasic. Metz, 1911 = Prosopis sparsa Cresson, 1869, by original Euryglossinae. designation. [Hylaeus]. Hylaeinae. Microhalictus Warncke. 1975b:85. Type species; Melitta miniitissinia Mexalictits Eickwort, 1978:567. Type species: Mexalictiis Kirby, 1802, by original designation. [Halictus]. Halictini. micheneri Eickwort, 1978, by original designation. Halictini.

Michanthidium Urban. 1994a:28l. replacement for Microhesnia Michener. 1 965a: 1 13. Type species: Brachyhesma Gnatluintliidiuiu Urban. Type species: Gnathantliidium incompleta Michener. 1965. by original designation. sakagamii Urban, autobasic. Anthidiini. [Brachyhesma]. Euryglossinae.

Michenerapis Hirashima and Lieftinck. 1982:5. Type species: MicrohoplitisTkalcu. 1993b:81 1 . Type species: Osmia hohmanni Osmiini. Pcdaeorhiza fe(co/<;/- Hirashima and Lieftinck, 1982, by origi- Tkalcu. 1993, by original designation. [Osmia]. nal designation. [Palaeorhiza]. Hylaeinae. Micromelecta Baker, 1906: 143. Type species: Melecta maculata Michenerella Krombein, 1950:122. Type species: Heriades Cresson, 1879. by original designation. Isotypic with paganensis Yasumatsu, 1942. by original designation. Zacosmia. Melectini. Osmiini. [Heriades]. Micrommation Moure, 1969:247. Type species: Micrommation

Micheneria Kerr. Pisani, and Ally. 1 967: 1 39. not Orfila and Rossi, larocai Moure, 1969. by original designation. Augochlorini. 1956. scutellaris Latreille, 181 1, Type species: Melipona by Micronomada Cockerell and Atkins, 1902:44. Type species: See Michmelia. original designation. [Melipona]. Meliponini. Nomada modesta Cresson, 1863, by original designation. Micheneriila Bohart. 1965:320. Type species: Michencrula [Nomada]. Nomadini. beameri Bohart, 1965, by original designation. Rophitinae. Micronomadopsis Rozen, 1958:107. Type species: Nomadopsis

1 for Michmelia Moure, 1975:62 , replacement Micheneria Kerr, fracta Rozen, 1958, by original designation. [Nomadopsis]. Pisani. and Aily. Type species: Melipona scutellaris Latreille. Panurginae. 1811, autobasic. [Melipona]. Meliponini. Micronychapis Moure and Michener, 1955b:286. Type species: Mcra//cto(^es Timberlake, 1939:397. Type species: Halictoides Tetralonia duckei Friese, 1908, by original designation. ruficaiidiis Michener, 1937, by original designation. Eucerini. [Dufourea]. Rophitinae. Micropasites Linsley, 1942:130. Type species: Neopasites Micramegilla Brooks, 1988:508. Type species: Anthophora cressoni Crawford, 1916. by original designation. niveata Friese, 1905. by original designation. [Amegilla]. [Gnathopasites|. Biastini. Anthophorini. Micropasites Warncke, 1983:283, not Linsley, 1942. Type .spe- Micrandrena Ashmead, 1899a: 89. Type species: Micrandrena cies: Ammobates minutissimus Mavromoustakis, 1959, re- pacifica Ashmead, 1899 = Andrena melanochroa Cockerell. named tunensis Warncke, 1983, because of homonymy in 1898, by original designation. Andreninae. Pasites. [Pasites]. See Ebweriana. Ammobatini.

Micranthidium Cockerell, 1930b:45. Type species: Anthidium Microrophites Warncke, 1979:133. Type species: Rophites truncatum Smith. 1854. by original designation. Cjuadridentatus Warncke, 1979, by original designation. [Pachyanthidmm]. Anthidiini. [Rophites]. Rophitinae. Genus-Group Names of Bees 39

Microspbecodes Eickwort and Stage, 1972:501. Type species: Monosiniu Tkalc, 1974a:337. Type species: Osmia apicata Smith, Sphecoiles kutlileenae Eickwort, 1972, by original designa- by original designation. [Osmia]. Osmiini. tion. Halictini. ISphecodes], Monoxylocopa Hurd and Moure, 1963:127. Type species: Micwstelis Robertson, 1903a:l70, 175. Type species: Stelis Xylocopa abhreviata Hurd and Moure, 1963, by original des- lateralis Cresson. 1864, by original designation. Anthidiini. ignation. [Xylocopa]. Xylocopini.

Micwihiirge Michener, 1983:181. Type species: Lithurgus Monumetha Cresson, 1864:387. Type species: Chelostoma phanidontiis Moure, 1948, by original designation. albifrons Kirby, 1 837, by designation of Titus, 1 904a:26. Lithurgini. Osmiini.

Mielkcaniludimn Urban, 1996:121. Type species: Morawitzella Popov, 1957:916. Type species: Epimethea nana Mielkeanthidiiim nigripes Urban, 1996, by original designa- Morawitz, 1 880, by original designation. Rophitinae. tion. Anthidiini. Morawitzia Friese, 1902:185. Type species: Morawitzia Miimmdrena Lanham, 1 949:2 1 7. Type species: Andrena iinitatrix panurgoides Friese, 1902, by designation of Sandhouse, Cresson, 1872, by original designation. [Andrena]. 1943:574. Sandhouse (1943) considered this genus monoba- Andreninae. sic, but two names (one called a variety) were included by Friese Mimoxylocopa Hurd and Moure, 1963:203. Type species: (1902). Rophitinae.

Xylocopa rufipes Smith, 1852, by original designation. Morgania Smith, 1 854:253. Type species: Pasites dichroa Smith, [Xylocopa]. Xylocopini. 1854, monobasic. Ammobatini.

Mimulapis Bridwell, 1919:162. Type species: Mimiilapis Morphanthidiiim Pasteels, 1969b:423. Invalid because no type versatilis Bridwell, 1919, by original designation. Rophitinae. species designated: three included species. Synonym of to two of the included and Mimitodufourea Ebmer, 1984:361. Type species: Diifourea Ardenthidiiim. judge by species, was intended Pasteels to be the same as niiiuirissiimi Ebmer, 1976, by original designation. [Dufourea]. presumably by Anthidiini. Rophitinae. Ardenthidiiim. [Anthidium[. Pasteels, 1965:537. hiseta Milchellapls Michener, 1 965a:2 1 1 . Type species: Megachilefab- Morpbella Type species: Megachile ricator Smith, 1868. by original designation. [Megachile]. Vachal, 1903, by original designation. [Chalicodoma]. Megachilini. Megachilini.

Mitsiikuriapis Hirashima, LaBerge, and Ikudome, //( Ikudome, Moureana Mitchell, 1980:56. Type species: Megachile 1994:6, replacement for Mitsiikuriella Hirashima and anthidioides Radoszkowski, 1874, by original designation. LaBerge. Type species: Ntmiia japonica Smith, 1873, [Pseudocentron]. Megachilini. autobasic. Andreninae. [Andrena]. Moiireanthidiuni Urban, 1995a:37. Type species: Dianthidium Mitsiikuriella Hirashima and LaBerge, 1965, in Hirashima, siibarenarium Schwarz, 1933, by original designation. 1965a:472, not Heding and Panning, 1954. Type species: Anthidiini. Nomia Smith, 1873, japonica by original designation. Mourecotelles Toro and Cabezas, 1977:50. Type species: See Andreninae. [Andrena]. Mitsiikuriapis. Moiirecotelles mixta Toro and Cabezas, 1977, by original Monia Westwood, 1875:221, not Gray, 1850. Type species: designation. Colletinae. Monia Westwood, 1875, monobasic. See Monidia. grisea Mourella Schwarz, 1946. in Moure, 1946a:442. Type species: Colletinae. Melipona caeriilea Friese, 1900, by original designation. Monidia Cockerell, 1905c:9, replacenjent for Monia Westwood, Meliponini.

1 875. Monia Westwood, 1 875, autobasic. Type species: grisea Mucidobombus Kriiger, 1920:350. Type species: Bonibus Colletinae. mucidus Gerstaecher, 1869, monobasic. Kriiger gave no Monilapis Cockerell, 1931f:529. Type species: Hylaeus subgeneric characters, but according to the Code, 3rd ed.. Art. tomentosiis Eversmann, 1852, by original designation. 12(b)(5), the name is nonetheless valid. Isotypic with [Halictus]. Halictini. Mucidobombus Skorikov. Bombini.

Monilosmia Robertson, 1903a:166, 171. Type species: Osmia Mucidobombus Skorikov, 1922b: 149. Type species: Bombiis = canadensis Cresson, 1864 Osmia simillima Smith, 1853, mucidus Gerstaecker, 1 869, by designation of Sandhouse, by original designation. Osmiini. 1943:574. Homonym and junior synonym of and i.sotypic with Mucidobombus Kriiger, 1 920. Bombini. Monoeca Lepeletier and Serville, 1828:528. Type species: Halictus Monoeca hrasiliensis Lepeletier and Serville, 1 828, monoba- Mycterochlora Eickwort, 1969:423. Type species: sic. Tapinotaspidini. repandirostris Vachal, 1911, by original designation. [Augochlora]. Augochlorini. Monoepeolus Bischff, 1930:3. Type species: Apis variegata

1 879:5. metalliciim Linnaeus, by original designation. Named in synonyiny of Mxdrosoma Smith, Type species: Mydrosoma = monobasic. Epeolus s. str. as contrast to Diepeolus. [Epeolus]. Isotypic Smith, 1879 Apista opalina Smith, 1861, with Epeolus. Epeolini. Dissoglottini. 40 ScffiNTiFic Papers, Natural History Museum, The University of Kansas

Mydrosomella Michener, 1986b: 194. Type species: Diphaglossa Neochalicodoma Paslesls, 1970:231. Type species: Chalicodoma (?) gaiillei Vachal, 1904. by original designation. pseudohiminata Pasteels. 1965. by original designation. Dissoglottini. [Chalicodoma]. Megachilini.

Mystacanthophora Brooks, 1988:466. Type species: Anthophora Neochelynia Schrottky, 1920:187. Type species: Neochelynia montana Cresson, 1869. by original designation. paidista Schrottky. 1920. monobasic. Megachilini. [Anthophora], Anthophorini. Neocoelioxys Mitchell. 1973:64. Type species: Coelioxys 1967: 104. Osmia nemoris Mystacosmia Snelling. Type species: assumptionis Schrottky, 1911. by original designation. Osmiini. Sandhouse. 1924. by original designation. [Osmia]. [Coelioxys]. Megachilini. Nananthidium Moure. 1947a:26. Nammthidium Type species: Neocoryniira Schrottky. 1910:540. replacement for Cacosoma Moure. 1947. by original designation. Anthidiini. hettyae Smith, 1879. Type species: Cacosoma discolor Smith, Nannothgona Cockerell. 1922e:9. Type species: Melipona autobasic. Augochlorini. testaceiconiis Lepeletier, 1836, by original designation. Neoconniiroides Eickwort. 1969:404. Type species: Halictus Meliponini. rhytis Vachal. 1904. by original designation. [Neocorynura]. nanus Nanocolletes Warncke, 1978:341. Type species: Colletes Augochlorini. Friese, 1898. by original designation. [Collates]. Colletinae. Neocressoniella Gupta. 1993:172. Type species: Megachile Hurd and Moure. 1963:99. Nanoxylocopa Type species: Xylocopa carbonaria Smith. 1853, by original designation. ciliata Burmeister, 1876. by original designation. [Xylocopa]. [Cressoniella]. Megachilini. Xylocopini. Neoeutricharaea Rebmann. 1967:36. Type species: Apis Michener. 1970:208. Nasutapis Type species: Nasutapis rotundata Fabricius, 1787. by original designation. straussonim Michener. 1970. original designation. by [Megachile]. Megachilini. Allodapini. Neofidelia Moure and Michener. 1955a:202. Type species: Navicularia Mehely, 1935:34. 140. Invalid because no type spe- Neofidelia profiiga Moure and Michener, 1955, by original cies was designated. [Prosopis]. Hylaeinae. designation. Fideliini. Navicularia Popov, 1939: 168. Type species: Mellinus variegatus Neohalidoides Viereck. 1904b:261. Type species: Pcmiirgus Fabricius, 1798, by original designation. [Prosopis]. maiinis Cresson. 1878. by original designation. Rophitinae Hylaeinae. Neolarra Ashmead. 1890:8. Type species: Neolarra pruinosa Neagapostemon Cure, 1989:295. Type species: Pseudagapo- Ashmead. 1890, monobasic. Neolarrini. stemon cyanomelas Cure, 1989, by original designation. Cure Ruz andToro, 1983:274. (1989) attributes the specific name cyanomelas to Moure, but Neoliphanthus Type species: Liphanthus hicelhdaris Ruz and Toro. 1983. earlier usages were nomina nuda; Cure was the first to de- by original designation. scribe the species. [Pseudagapostemon]. Halictini. [Liphanthus]. Panurginae. Mitchell. 1934:302, 306. Nealictus Pesenko, 1984:346. Type species: Halictus parallelus Neomegachile Type species: Megachile chichimeca 1878. Say, 1837. by original designation. [Halictus]. Halictini. Cresson, by original designation. [Megachile]. Megachilini. Neanthidium Pasteels, 1969a:93. Type species: Anthidium 1971:2. octodentatum Perez. 1895, by original designation. Neopanurgus Schwammberger, Type species: 1 1 des- Anthidiini. Neopaniirgus richteri Schwammberger, 97 , by original Panurginae. Nectarodiaeta Holmberg, 1887b:225, nomen nudum. Eucerini. ignation. Neopasiphae Perkins. 1912:114. Type species: Neopasiphae Nectarodiaeta Holmberg. 1903:420. Type species: Nectarodiaeta mirabilis Perkins. 1912. monobasic. Colletinae. oliveirae Holmberg. 1903. monobasic. Eucerini. Neopasites Ashmead. 1898:284. Type species: Phileremus Neffapis Ruz, 1995, in Rozen and Ruz, 1995:3. Type species: fulviventris Cresson. 1878. by original designation. Isotypic Nejfapis longilingua Ruz. 1995. by original designation. with Biastini. Panurginae. Gnathopasiies. Ashmead, 1899a:85. Perdita zebrata Neglectella Pasteels, 1965:431. Type species: Megachile Neoperdita Type species: Cresson. 1878. armatipes Friese. 1909, by original designation. by original designation. Panurginae. [Chalicodoma]. Megachilini. Neoscirtetica Schrottky, 1913:256, replacement for Scirteiica 1903. Scirtetica antarctica Nemandrena LaBerge, 197 la:48. Type species: Andrena torulosa Holmberg. Type species: autobasic. W\\.\\ AUoscirtetica and LaBerge. 1971. by original designation. [Andrena]. Holmberg. 1903, Isotypic Andreninae. Holmhergiapis. Eucerini. Osmia Neoashmeadiella Gupta, 1990:56. Type species: Ashmeadiella Neosmia'YkaXcu. 1974a:333.Type species: gracUicornis Osmiini. indica Gupta. 1990. by original designation. [Ashmeadiella]. Perez, by original designation. [Osmia]. IVIegachilini. Neotrypetes Robertson. 1918a:91. Type species: Trypetes = Neoceratina Perkins, 1912:117. Type species: Neoceratina productus Robertson, 1905 Megachile variolosa Cresson, australensis Perkins. 1912. monobasic. Ceratinini. 1872. by original designation. Osmiini. Genus-Group Names of Bees 41

Neoxylocopa Michener. 1954:157. Type species: Apis 1936:12, and Milliron. 1961:54, designated type species but brasilianorwn Linnaeus. 1767. by original designation. placed Nobilihomhus as a junior synonym of other names,

[Xylocopa]. Xylocopini. and thus did not validate it (see Williams, 1991 ). Bombini.

Neshylaeus Heider, 1935:2245. unjustified emendation of Nohilihombus Richards, 1968:216, 222. Type species: Bomhus Nesylaeus Bridwell, 1919. Hylaeinae. nohilis Friese, 1904, by original designation. [Bombus]. Bombini. NesocoUetesMichtTicr, 1965a:52. Type species: Lampmcolletes

Smith. 1 876. fiilvesceiis by original designation. [Leioproctus]. Nodocolletes Rayment, 1931:164. Type species: Nodocolletes Colletinae. dentatus Rayment, 1931 = Andrena chalybeata Erichson, Nesoeiipetersia Bliithgen, 1936:182. Type species: Sphecodes 1851. by original designation. Isotypic with Lamprocolletes

scotti Cockerell. 1912. by original designation. [Eupetersia] . by synonymy. Colletinae. Halictini. Nodula Ma, 1938:290. Type species: Apis amethystina Fabri- Nesohalictus Crawford. 1 9 10: 1 20. Type species: Halictus robhii cius, 1793, by original designation. [Xylocopa]. Xylocopini. = Crawford. 1910 Nomia halictoides Smith. 1858. by origi- Nogueirapis Moure. 1953b:247. Type species: Trigona butteli nal designation. [Halictus]. Halictini. Friese, 1900, by original designation. [Partamona]. Nesohylaeus Ikudome, 1989:125. Type species: Hylaeus niger Meliponini. Bridwell, 1919, by original designation. [Hylaeus]. Nomada Scopoli. 1770:44. Type species: Apis ruficornis Hylaeinae. Linnaeus, 1758, by designation of Curtis, 1832:pl. 419. Apis Nesomelecta Michener, 1948b: 15. Melecta Type species: /afenr/fl/!fl Linnaeus, 1767, designated by Latreille, 1810:439, haitensis Michener. 1948, [Melecta]. by original designation. was not originally included. Nomada succincta Panzer, 1 798, Melectini. was designated as the type species by Blanchard. 1849:pl. Nesomonia Michener, Brooks, and Pauly, 1990:135. Type spe- 128 (1847) but was not originally included. Nomadini. cies: Nomia flaviveiUris Benoist. 1963. by original designa- Nomadita Mocsary, 1894:37. Type species: Nomadita montana tion. [Uromonia]. Meganomiinae. Mocsary, 1894, monobasic. Nomadini. Nesoprosopis Perkins, 1899:75. Type species: Prosopis facilis Nomadopsis Ashmead, 1898:285. Type species: Perdita zonalis Smith, 1879, by designation of Popov, 1939:168. Hylaeinae. Cresson, 1879, lapsus for Calliopsis zonalis Cresson, 1879, Nesothrincostoma 1933:364. Bliithgen, Type species: by original designation. Isotypic with Claremontiella by syn- Thrincostoma serricorne Bliithgen, 1933, monobasic. onymy. Panurginae. Halictini. Nomadosoma Rohwer, 1911:24. Type species: Pasites pilipes Bridwell, 1919:147. Nesylaeus Type species: Hylaeus Cresson, 1865. by original designation. Nomadini. nesoprosopoides Bridwell, 1919, by original designation. Nomadula Cockerell, 1903d:611. Type species: Nomada [Hylaeus], Hylaeinae. americana Kirby of Robertson, 1903d = Nomada articulata Nevadensibombus Skorikov, 1922b: 149. Bombiis Type species: Smith, 1854, by original designation. See Code, ed. 3, Art. nevadensis of Prison, 1927:64. Ciesson, 1874, by designation 70(c). [Nomada]. Nomadini. Bombini. Nomia Latreille, 1 804: 1 82. Type species: Andrena cun'ipes Fab- Pasteels, 1984:57, for Melanthidium Nigrantliidium replacement ricius, 1781, monobasic. Nomia diversipes "Fabr.," actually Pasteels. 1969. Anthidium concolor Friese, Type species: Latreille, 1806, designated by Blanchard, 1849:pl. 125(1847), 1913, and autobasic. Anthidiini. by original designation was not originally included. Nomiinae. 1993:235. Nigrocoelioxys Gupta, Type species: Coeiioxys Nomiapis Cockerell, 1919a:208. Type species: Nomia diversipes Smith, 1854, fuscipennis by original designation. [Coeiioxys]. Latreille, 1806, by original designation. [Nomia]. Nomiinae. Megachilini. Nomiocolletes Brethes, 1909c:455. Type species: Nomia Niltonia Moure, 1964c:52. Niltonia Moure, Type species: virgilii joergenseni Friese, 1908, by original designation. Colletinae. 1964, by original designation. Colletinae. Nomioides Schenck, 1866:333. Type species: Apis minutissima Nitocris 1815:123. for Rafinesque. unjustified replacement Rossi, 1790, designated by the Commission, Opinion 1319 Nomia Latreille, 1804. Andremi Fab- Type species: cun'ipes (1985). Misidentification of the type species as Andrena ricius, 1781, autobasic. Nomiinae. pulchella Jurine, 1807, by Schenck and associated nomen- Nivanthidium Pasteels, 1969a: 106. Type species: Anthidium clatural problems are explained by Pesenko and Kerzhner niveocinctum Gerstaecker, 1857, by original designation. (1981). Nomioidinae. [Anthidium]. Anthidiini. Noonadania Hirashima, 1978:68. Type species: Palaeorhiza Nobandrena Wamcke, 1968:45. Type species: Andrena mibilis sculpturalis Hirashima. 1978, by original designation. Morawitz, 1874, by original designation. [Andrena]. [Palaeorhiza]. Hylaeinae. Andreninae. Notandrena Perez, 1 890: 1 73. Type species: Andrena nitidiuscula Nobilibombus Skorikov, 1933a:62. Invalid because no type spe- Schenck, 1 853, designated by Hedicke, 1933:216. [Andrena]. cies was designated [Code, 3rd ed.. Art. 13(b)]. Bischoff, Andreninae. 42 SciENTinc Papers, Natural History Museum, The University of Kansas

Notanthidiiim Isensee, 1927:373. Type species: Anthidium Odantopasites Linsley. 1942:128. Type species: Neopasites sieloides Spinola. 1851. by original designation. arizonicus Linsley. 1942. by original designation. [Dianthidium|. Anthidiini. [Neopasites]. Animobatoidini.

Noteopsis Ikudome, 1991 : 791 . Type species: Hylaeus nanseiensis Odontostelis Cockerell, 1 93 ld:542. Type species: Stelis abnormis Ikudome, 1989, by original designation. [Hylaeus]. Friese, 1925= Anthidium bivittatum Cresson, 1878. by origi- Hylaeinae. nal designation. Isotypic with Hoplostelis. Anthidiini.

Nateriades Cockerell, 1931b:332. Type species: Megachile Odantotrigona Moure, 1961:213. Type species: Trigona triciirinatci Bingham. 1903. by original designation. liaematoptera Cockerell. 1919. by original designation. [Heriades]. Osmiini. Meliponini.

Nothosmia Ashmead, 1899a:75. Type species: Osmia disdncta Odyneropsis Schrottky, 1902a:432. Type species: Odyneropsis Cresson. 1864. by original designation. Osmiini. holosericea Schrottky. 1902. by original designation. Epeolini. Nuthykieus Bridwell. 1919:125, 126. Type species: Prosopis hemldica Smith, 1853, by original designation. Hylaeinae. Oediscelis Philippi, 1866: 109. Type species: Oediscelis vernalis Philippi. 1866. by designation of Cockerell, 1919a:185. Toro NatocoUeles Cockerell, 1916b:44. Type species: NolocoUetes and Moldenke ( 1979) list the as O. hetewdoxus Cockerell, 1916, monobasic. Colletinae. enoneously type species plebeia Philippi. Xeromelissinae. Notolonia Popov, 1962:294. Type species: Notolonia astragali Oediscelisca Moure, 1946c:243. Type species: Oediscelis friesei Popov. 1962. by original designation. Eucerini. Ducke. 1907. by original designation. Xeromelissinae. Notomegilla Brooks. 1988:511. Type species: Aiithophora Ocstropsis Smith. 1868a:253, not Brauer, 1868. Type species: aeruginosa Smith, 1 854. by original designation. [Amegilla], Oestropsis pubescens Smith. 1868, monobasic. See Gastropis. Anthophorini. Stenotritidae. Notoiuclirta Cockerell, 1933b: 128. Type species: Rediviva Oligandrena Lanham, 1949:207. Type species: Andrena neliami Cockerell. 1 93 1 . by original designation. Melittinae. macrocepluila Cockerell, 1916. by original designation. Notoxaea Hurd and 1976:21. Oxaea Linsley. Type species: [Andrenaj. Andreninae. ferruginea Friese, 1 898, by original designation. Oxaeinae. Oligeucera Sitdikov and Pesenko. 1988:83. Type species: Eucera Hurd. 1956:2. Notoxylocopa Type species: Xylocopa popovi Sitdikov. 1988. by original designation. [Eucera]. tabaniformis Smith. 1854. by original designation. Eucerini. [Xylocopa]. Xylocopini. Oligochlora Engel. 1997:336. Type species: Oligochlora Nitbenomiu 1980:122. Noiuia nubecula Pauly, Type species: eickworii Engel. 1997, by original designation. Fossil. Nomiinae. Smith. 1875. by original designation. [Nomia]. Augochlorini. Cockerell. 1929e:303. Bonibus Nycinmelitta Type species: Oligotropus Robertson, 1903a: 168. Type species: Oligotropus tranquebaricus Fabricius. 1804, by original designation. campanulae Robertson, 1903. monobasic. Megachilini. [Xylocopa]. Xylocopini. Olmecanthidium Peters, 1972:377. Type species: Ukmthidium Obertobombus 1930:107. Bomlnis oberti Reinig. Type species: interdisciplinaris Peters. 1972. by original designation. monobasic. Bombini. Morawitz. 1883. [Bombus]. [UlanthidiuiTi|. Anthidiini. Ochreriades Mavromoustakis, 1956:226. Type species: Eriades Omachtes Friese. 1909a:436, unjustified emendation of fasciatits Friese, 1899, by original designation. Osmiini. Omachthes Gcvsinscktr, 1869. Ammobatini.

Odontalictus 1 9 1 8a:9 1 . Halictus Robertson. Type species: ligatus Omachthes Gerstaecker. 1869:154. Type species: Omachthes Say, 1837. by original designation. Halictini. carnifex Gerstaecker, 1869, designated by Sandhouse, 1943:580. Ammobatini. Odontanthocopa Tkalcu, 1974b: 125. Type species: Osmia bidentata Morawitz, 1876. by original designation. Onagrandrena Linsley and MacSwain. 1 956: 111. Type species: [Anthocopaj. Osmiini. Andrena oenotherae Timberlake. 1937. by original designation. [Andrena]. Andreninae. Odonterythrosmia Tkalcu, 1974b: 131. Type species: Osmia fallax Perez. 1895. by original designation. [Anthocopa]. Opacula Pesenko and Sitdikov, 1988:849. Type species: Opacula Osmiini. donatica Sitdikov, 1988. by original designation. Eucerini. 1902a: 193. Odontobombus Kriiger. 1917:61. Type species: Apis argiilacea Opandrena Robertson, 187, Type species: Andrena Scopoli. 1763, by designation of Williams, 1994:339. A sec- cressonii Robertson, 1871, by original designation. tional name, to be treated as a subgenus according to the Code. Andreninae. 3rd ed.. Art. 10(e). [Bombus]. Isotypic with Megabombus by Oragapostemon Cure, 1989:312. Type species: Halictus synonymy. Bombini. divaricatus Vachal, 1903, by original designation. Halictini.

Odontochlora Schrottky. 1909a: 141. Type species: Augoehlora Orandrena Warncke. 1968:36. Type species: Andrena oralis nuilierl Cockerell, 1900, by original designation. Morawitz. 1876. by original designation. [Andrena]. Augochlorini. Andreninae. Genus-Group Names of Bees 43

Onuuhidiitm Pasteels. 1969a:95. Type species: Anthidium Oxaea Klug. 1807a:261. Type species: Oxaea flavescens Klug, oraiiieiise Brauns. 1905 = Anthicliuni folliculosum Buysson. monobasic. The names Oxaea and Dasyglossa appear to have 1897, by original designation. Anthidiini. been published at appro.ximately the same time in late 1807. has not been used since, and in 1810 was dis- Orhitella Ma. 1938:305. not Douville. 1915. Type species: Dasyglossa cussed under the of Oxaea 1 8 10). Even if Xxlocopa confusa Perez. 1901. by original designation. See heading (Klug, has it should not be resurrected. Maiella and Eiiiyapis. [Xylocopa]. Xylocopini. Dasxgiossa priority, Isotypic with Dasyglossa. Oxaeinae. Oreomelissa Hirashima and Tadauchi. 1975: 1 76. Type species: 1977:135. Andrena Andremi mitcikensis Hirashima. 1963. by original designa- Oxyandrena LaBerge. Type species: tion. [Andrena]. Andreninae. longifovea LaBerge. 1 977, by original designation. [Andrena]. Andreninae. Oieopasites Cockerell. 1906e:442. Type species: Oreopasites scituli Cockerell. 1906, monobasic. Ammobatini. Oxyanthidium Mavromoustakis, 1963a:653. Type species: Anthidium aculeatum 1832. Orfilana Moure, 1954b:266. Type species: Doeringiella Klug. by original designation. Anthidiini. variegciia Holmberg. 1886. by original designation. [Meganthidium]. [Doeringiella]. Epeolini. O.ni'/fl.srei Mavromoustakis. 1954:260. Type species: Oxybiastes Mavromoustakis, 1954, OrieiUalibombus Richards, 1929:378. Type species: Bombus bischoffi by original designation. orientalis Smith. 1854. by original designation. [Bombus]. Epeolini.

Bombini. Oxyhalictus Cockerell and Ireland, 1 935, in Cockerell, 1 935b:9 1 . Cockerell and Orientalobombiis Kriiseman, 1952:102, unjustified emendation Type species: Halictus acuiferus Ireland, 1935, of 0/7>«?rt///wwZ)(/5 Richards. 1929. [Bombus]. Bombini. monobasic. [Halictus]. Halictini. for Orientocoelio.xys Gupta. 1992:73. Type species: Coelioxys Oxynedis Moure, 1946d:18. 27. 31, pi. 3, error Oxynedys quadrifasciatus Gupta. 1992. by original designation. Schrottky. 1902. Ericrocidini.

[Coelioxys]. Megachilini. Oxynedys Schrottky, 1902a:491 . Type species: Oxynedys heroni = OrientocressonieUa Gupta. 1993:165. Type species: Megachile Schrottky. 1 902 Mesocheira vehitina Lepeletier and Serville. relata Smith, 1879. by original designation. [Cressoniella]. 1825, by original designation. Isotypic with Hopliphora by Megachilini. synonymy. Ericrocidini. Oricnioheriades Gupta, 1987:68. Type species: Heriades Oxystoglossa Smith, 1853:83. Type species: Oxystoglossa orientalis Gupta, 1987, by original designation. [Heriades]. decorata Smith, 1853, monobasic. Augochlorini. Osmiini. Oxystoglossella Eickwort. 1969:422. Type species: Augochlora 1978b:332. Osmia nuixillaris Onentosmia Peters. Type species: cordiaefloris Cockerell. 1907, by original designation. Morawitz, 1875, Osmiini. by original designation. [Osmia]. [Augochlora]. Augochlorini. 1993:50. Anthidiuiu Orientotracluisa Gupta. Type species: Oxystoglossidia Moure, I943b:473. Type species: orientale 1897, [Trachusa]. Bingham, by original designation. Oxystoghissidia uranieUa Moure. 1943, by original designa- Antliidiini. tion. Augochlorini. Vachal. 1909a:35. 38. Type species: Orphana Orpluina Oxytrigona Cockerell, 1 9 1 7a: 1 24. Type species: Trigonaflaveola inquirenda Vachal. 1909, monobasic. Isotypic with medioriifa Cockerell. 1 9 1 3, by original designation. [Trigona]. and Ptoleglossa by synonymy. Andreninae. Leptoglossa Meliponini. Orthunthidium Mavromoustakis. 1953:837. Type species: Oxyxylocopa Hurd and Moure, 1963:275. Type species: Xylocopa Anthidiumfonnosanwu Friese, 191 7, by original designation. varipes Smith. 1854. by original designation. [Xylocopa]. [Paraanthidium]. Anthidiini. Xylocopini. Osirimis Roig-Alsina. 1989:17. Type species: Osirinus Ozbekosmia Zanden, 1994:168. Type species: Osmia avosetta lemniscatiis Roig-Alsina. 1989, by original designation. Wamcke. 1988. by original designation. [Osmia]. Osmiini. Isotypic with Compsoclepta by synonymy. Osirini. Pacbyanthidium Friese. I905a:66. Type species: Anthidium hi- Osiris Smith. 1854:288. Type species: Osiris pallidiis Smith. color Lepeletier. 1841, by designation of Cockerell, 1854. by designation of Sandhouse. 1943:580. Osirini. 1920b:298. Also described as new^by Friese, 1910b: 158. Osmia 1806:230. bicornis Linnaeus, Panzer, Type species: Apis [Anthidium]. Anthidiini. I15'&-Apis riifa Linnaeus, 1758, by designation of Latreille, Pachyceble Moure, 1940:54. Type species: Pachyceble lanei 1810:439. Osmia tricornis Latreille. 1811. designated by Moure. 1940, Halictini. not an included by original designation. Blanchard. 1849:pl. 127 ( 1846), was originally Friese, 1902:186. species. Isotypic with Amblys and Ceratosmia; isotypic with Pachycentris Type species: Pachycentris monobasic. Pachyosmia by synonymy. Osmiini. schrottkyi Friese. 1902, Tapinotaspidini. 1922d:4. Olhinosmia Michener. 1943:86. Type species: Thaiimatosoma Pachxcerapis Cockerell, Type species: Exomalopsis motuliferiim Cockerell. 1932. by original designation. cornigera Cockerell, 1922, by original designation. [Anthocopa]. Osmiini. [Exomalopsis]. Exomalopsini. 44 ScENTinc Papers, Natural History Museum, The University of Kansas

Pachycolletes Bischoff, 1954, w Stoeckhert, 1954:73. Type spe- Palinepeolus Holmberg, 1909b:77. Type species: Epeolus cies: Apis ciinicitlaria Linnaeus. 1761. by original designa- viperinus Holmberg. 1 886, by original designation. Isotypic tion. [Colletes]. Colletinae. with Calospikmui: isotypic with Isepeolus by synonymy.

Pachyhulictus Cockerell, 1929d:589. Type species: Halictus Isepeolini. merescens Cockerell. 1919. by original designation. Pallandrenu Warncke, 1968:39. Type species: Andrena [Halictus]. Halictini. pallidicincta Brulle. 1832. by original designation. [Andrena]. Andreninae. Pachyhalonia Moure and Michener, 1955b:281. Type species: = Dasyhalonia jiisti Moure and Michener. 1955 Svastra Pallhatictus Warncke, 1975b:92. Type species: Halictus pollens sapucacensis Cockerell. 1918. by original designation. Brulle. 1832. by original designation. [Halictus]. Halictini. Eucerini. [Dasyhalonia]. Panomalops is Timberlake. 1980:82. Type species: Exomalopsis

Pachymelopsis Cockerell, 1 905a:33 1 . Type species: Pachymelus linsleyi Timberlake. 1980. by original designation. conspicuus Smith. 1879, by original designation. [Exomalopsis]. Exomalopsini.

[Pachymelus]. Anthophorini. Panurgimis Nylander, 1 848:223. Type species: Pamirgimis niger Pachymelus Smith. 1879:116. Type species: Pachymelus Nylander. 1848, monobasic. Panurginae. Smith. 1879. of Sandhouse, micrelephas by designation Pamvgomia Viereck, 1909:48. Type species: Panurgomia fuchsi 1943:581. Sandhouse considered this to be the species type Viereck. 1909. monobasic. Dasypodinae. because the other species originally included had been re- Panurgus Panzer, 1806:209. Type species: Andrena lobata moved to Pachymelopsis. but see Code, 3rd ed.. Art. 69(b). Panzer. 1799 =Apis calcarata Scopoli, 1763, by designation Her conclusion is nonetheless accepted. Anthophorini. of Latreille, 1810:439. Later and therefore invalid type des- Rodeck. 1945:180. Nomada vincta Pachynomada Type species: ignations were Apis ursina Gmelin, 1790, by Westwood, 1837. original [Nomada]. Nomadini. = Say. by designation. 1840a: 85, and/4/ji/,sflrra Panzer, \?>05 Apis banksiana Kirby, Pachynomia Pauly, 1980:124. Type species: Nomia amoenula 1802,byBlanchard, 1849:pl. 126 (1846). Isotypic with fnopi Gerstaecker. 1870. by original designation. [Nomia], by synonymy. Panurginae. Nomiinae. Papualictus Michener, 1980:8. Type species: Homalictus = Pachyodonta Rayment, 1954:48. Type species: Binghamiella megalochilus Michener, 1980 Halictus lorentzi Friese, 1911, fulvicornis Rayment, 1954, monobasic. [Binghamiella]. by original designation. [Homalictus]. Halictini. Colletinae. Papuanorhiza Hirashima, 1996:80. Type species: Xenorhiza Pachyosmia Ducke, 1900: 18. Type species: Osmia rufa Linnaeus, krombeini Hirashima. 1996, by original designation. 1758, designated by Sandhouse, 1939:9. [Osmia]. Isotypic [Xenorhiza]. Hylaeinae. with Osmia, and Ceratosmia Osmiini. Amblys. by synonymy. Papuatrigona Michener and Sakagami, 1990, in Michener, Pachyprosopina Michener, 1965a: 108. Type species: Euryglossa 1990:153. Type species: Trigona genalis Friese, 1908, by paupercula Cockerell. 1915, by original designation. original designation. [Trigona]. Meliponini. [Pachyprosopis]. Euryglossinae. Paraanthidiellum Pasteels, 1969a:79, unnecessary emendation Pachyprosopis Perkins, 1908:29. Type species: Pachyprosopis of Paranlhidiellum Michener. [Pseudoanthidium]. Anthidiini. mirabilis Perkins, 1908, monobasic. Euryglossinae. Paraanthidium Friese, 1898a: 101. Type species: Apis interrupta Pachyprosopula Michener, 1965a: 106. Type species: Fabricius, 1781. by designation of Cockerell, 1909b: 269. Pachyprosopis kellyi Cockerell, 1916, by original designa- [Anthidium]. Anthidiini. tion. [Pachyprosopis]. Euryglossinae. Paracella Pasteels. 1965:277, invalid because no type species Pachysvastra Moure and Michener, 1955b;284. Type species: was designated. [Megachile]. Megachilini. Tetralonia Bertoni and leucocephala Schrottky, 1910, by origi- Paracella Michener, new subgenus of Megachile for Paracella nal Eucerini. designation. Pasteels, 1965:277. Type species: Megachile semivenusta

Paedia Dalla Torre. 1 89 1 : 1 47. unjustified emendation of Paidia Cockerell, 1931. This taxon was characterized by Pasteels, Radoszkowski, 1872. Ammobatoidini. 1965:227. Megachilini.

Paidia Radoszkowski, 1872b: 10, not Herrich-Schaffer, 1847, Paracentris Cameron, 1903a:235. Type species: Paracentris unnecessary replacement for Ammobatoides Radoszkowski, fulvohirta Cameron, 1903. monobasic. Centridini. 1868. Phileremus abdominalis Eversmann. Type species: Parachalicodoma Pasteels, 1 966: 1 3, not Tkalcu, 1 969. Type spe- 1 852. autobasic. with Ammobatoides Radoszkowski Isotypic cies: Megachile incana Friese. by original designation. and Phiarus. Ammobatoidini. [Chalicodoma]. Megachilini. 1984:37. Palaeapis Hong, Type species: Palaeapis beiboziensis Parachalicodoma Tkalcu, 1969b:363, not Pasteels, 1966. Type monobasic. Fossil described in Hong, 1984, wasp, Apidae. 1 1 species: Chalicodoma rufitarsis Lepeletier, 84 , by original Palaeorhiza Perkins, 1908:29. Type species: Prosopis peniridis designation. See Xenochalicodoma. [Chalicodoma]. Cockerell, 1905, by original designation. Hylaeinae. Megachilini. Genus-Group Names of Bees 45

Paracoflio.xys Gribodo, 1884:274. Type species: Coelioxys Parammohatodes Popov, 1931b:462. Type species: Phiarus montaiuloui Gribodo, 1884 - Coelioxys cilata Forster, 1853, ininutus Mocsary. 1878, by original designation. monobasic. [Coelioxys]. Megachilini. Ammobatini.

1 Paraiidrena 1897:337. Paracoelioxys Radoszkowski. 1 893:33, not Gribodo. 884. Type Robertson, Type species: Panurgus andrenoides Cresson, 1878, of Cockerell, species; Paracoelioxys barrel Radoszkowski, 1 893, by des- by designation 1897a:288. Andreninae. ignation of Sandhouse, 1943:583. Coelioxys decipiens Spinola. 1838. designated by Alfken, 1938:433, was not an Parandrenella Popov, 1958:112. Type species: Andrena originally included species. See Radoszkowskiana. dentiventris Morawitz. 1874, by original designation. Megachilini. [Andrena]. Andreninae.

Paracolleies Smith. 1 853:6. Type species: Paracolletes crassipes Paranomada Linsley and Michener, 1937:82. Type species: Smith, 1853, monobasic. Coiletinae. Paranoinada nitida Linsley and Michener, 1937, by original

Paracroclsa Alfken. 1937:173. Type species: Paracrocisa designation. Brachynomadini.

siiuiilica Alfken, 1937. monobasic. Melectini. Paranomia Friese, 1 897b:48, not Conrad, 1 860. Type species: Nomia chalybeata Smith, 1875, by designation of Cockerell, Paradialictus Pauly, 1984c:691. Type species: Paradialictus 1910f:290. [Nomia]. See Paranomina. Nomiinae. synavei Pauly. 1984, by original designation. Halictini. Paraiwmina Michener. 1944:251, not Hendel, 1907, replace- Paradioxys Mocsary, 1894:35. Type species: Dioxys pannonica ment for Paranomia Friese, 1897. Type species: Nomia Mocsary, 1877, monobasic. Dioxyini. chalybeata Smith, 1875, by original designation and Brauns, 1926:202. Type species: Parafidelia friesei Parafidelia autobasic. Not replaced because subjective synonym of Brauns, 1926, monobasic. Fideliini. Acunomia. [Nomia]. Nomiinae. 1906b: 118, replacement for Friesea Parafriesea Schrottky. Parcwomioides Pesenko, 1983:175. Type species: Nomioides 1902. Friesea hrasiliensis Schrottky, Type species: Schrottky, steinbergi Pesenko. 1983, by original designation. 1902 = 1884, autobasic. Camptopoeiim prinii Holmberg, [Nomioides]. Nomioidinae. Panurginae. Paranthidiellum Michener. 1948a:25. Type species: Aniliidium Vachal, 1903:89. Halictus Paragapostemon Type species: cribratum Morawitz, 1875, by original designation. podagerV&chiiX. 190?> = Nomia coelestina 'Westwood. 1875, Anthidiini. by designation of Cockerell, 1905a:354 footnote. Halictini. Paranthidium Cockerell and Cockerell. 1901:50. Type species: Parahalictoides Cockerell and 1899:420. Porter, Type species: Anthidium perpicium Cockerell, 1898, by original designa- Halictoides campanulae Cockerell. 1897. by original desig- tion. Anthidiini. nation. [Halictoides]. Rophitinae. Paranthocopa Tkalcu, 1 974b: 1 32. Type species: Osmia pinquis 1965a:92. Parahesma Michener. Type species: Euryglossa Perez, 1895, by original designation. [Anthocopa]. Osmiini. tuberculipes Michener. 1965, by original designation. Paranychium Toro and Herrera, 1 980:2 1 3. Not valid because no [Euryglossa]. Euryglossinae. type species was designated. [Callonychiumj. See Paraheterorhiza Hirashima. 1980: 104. Type species: Palaeorhiza Paranychium Toro. Panurginae. hilara Cheesman, 1948, by original designation. Paranychium Toro, 1989:231. Type species: Camptopoeum [Palaeorhiza]. Hylaeinae. chilense Friese, 1906, by original designation. Paralictus Robertson. 1901:229. Type species: Halictus [Callonychium]. Panurginae. Robertson, 1892 (not-Morawitz, 1813) = Halictus cephalicus Parapachyprosopis Exley, 1972: 17. Type species: Pachyprosopis Dalla Torre, 1896, cephalotes by original designation. angophorae Cockerell, 1912, by original designation. Halictini. [Pachyprosopis]. Euryglossinae. and 1936:287. Paramacropis Popov Guiglia, Type species: Parapartamona Schwarz, 1948:428. Type species: Trigona ussuriana 1936, monobasic. Ctenopleclra Popov, [Macropis]. zonata Smith, 1854, by original designation. [Trigona]. Melittinae. Meliponini. 1898b: 198. Paramegachile Friese, Type species: Apis argentata Parapolyglossa Brauns, 1929:134. Type species: Polyglossa Fabricius, 1793. of Mitchell. 1934:298. by designation heterodoxa Cockerell, 1921, by designation of Sandhouse, with Eutricharaea. [Megachile]. Isotypic Megachilini. 1943:584. Sandhouse recorded the type as by "original des- Paramegalochila Schulz, 1906:71, unjustified emendation of ignation,"" but this was an error. [Polyglossa]. Coiletinae. Friese. 1898. Paramegachile [Megachile]. Megachilini. Paraprosopis Popov, 1939:169. replacement for Campanularia 1852. Paramegilla Friese, 1 897a: 1 8, 24. Type species: Apis ireos Pallas, Mehely. Type species: Hylaeus pictipes Nylander. by 1773, designated by Sandhouse, 1943:584. [Podalirius]. original designation. [Prosopis]. Hylaeinae. Anthophorini. Parapsaenythia Friese, 1908b:42. Type species: Psaenythia 1908 = Caenohalictus Parammohates Friese, 1906e: 1 18. Type species: Parammobates argeutina Friese, serripes Ducke, 1908, hrasiliensis Friese, 1906, monobasic. Epeolini. monobasic. [Psaenythia]. Panurginae. 46 Scientific Papers, Natural History Museum, The University of Kansas

Pararhophites Friese, 1898c:305. Type species: Pamrhophites Pasites Jurine, 1807:224. Type species: Pasites maculata Jurine, qiiadratiis Friese. 1898. monobasic. Although he described 1807, by original designation. Subsequent designation: as a = Pararhophites subgenus of Rhophiles. Friese described Nomada schuttii Fabricius. 1 804 Tiphia hrevicornis Panzer, the species using Pararhophites as a genus. [Rophites]. 1798, by Latreille, 1810:439. Ammobatini. Pararhophitini. Pasitonuichles Sandhouse. 1943:586. unjustified emendation of Parasanis Ruz, 1993, in Ruz and Rozen, 1993:2. Type species: Pasitomachthes Bischoff, 1923. Ammobatini. Parasariis atacamensis Ruz, 1993, by original designation. Pasitoinachthes Bischoff, 1923:596. Type species: Panurginae. Pasitomachthes nigerrimus Bischoff. 1923. by original des-

Parasphecodes Smith. 1853:39. Type species: Parasphecodes ignation. Bischoff ( 1923) confusingly gave the type species hilactiis S,m\\\\. 1 853. by designation of Sandhouse, 1943:585. as nigerrimus (without genus or author) after mentioning Halictini. Pasites nigerrimus Friese, 1922. He included the latter among species of Morgania. however, and clearly intended Paratetrapedia Moure. 1941b:517. Type species: Ancyloscelis Pasitomachthes nigerrimus Bischoff. Ammobatini. lineata Spinola, 1851, by original designation. Tapinotaspidini. Patagiata Bliithgen, 1949:77. Type species: Prosopis dijformis Eversmann. 1852. by original designation. Parathrincostoma Bliithgen, 1933:389. Type species: [Prosopis]. Hylaeinae. Parathrincostoma seyrigi Bliithgen. 1933. by original designation. Halictini. Patellapis Friese, 1909a: 148. Type species: Halictus schultzei Friese, 1909, by designation of Cockerell. 1920b:311. Paratrigoiui Schwarz, 1938:487. Type species: Melipona

] Halictus]. Halictini. prosopiformis Gribodo, 1893. by original designation. Patera [Trigona]. Meliponini. Schwarz. 1938:475, not Lesson. 1837. Type species: Melipona testacea Klug, 1807. by original designation. Paraiigochlora Schrottky, 1910:540. Type species: Aiigoddora [Trigona]. See Partamona. Meliponini. spinolae Cockerell, 1900, by original designation. Augochlorini. Pauphalictus Warncke. 1981:87. Type species: Halictus pcniperatus BruUe. 1832. by original designation. [Halictus]. Paraugochloropsis Schrottky, 1906a:312. Type species: Halictini. Aiigochloropsis lycorias Schrottky, 1906 = Augochlora Pavostelis Sladen. 1916b:313. Stelis montana epipyrgitis Holmberg, 1903, monobasic. [Augochloropsis]. Type species: monobasic. Augochlorini. Cresson. 1864. [Stelis]. Anthidiini. 1970:322. Paremisia Moure. 1945c:406. Type species: Centris lineolala Pectinapis LaBerge, Type species: Pectinapisfasciata = 1970. Eucerini. Lepeletier. 1841 Bojiihiis siniilisFabncms. 1804. by origi- LaBerge. by original designation. nal designation. Centridini. Pectinata Mehely, 1935:54. 161. Invalid because no type species was Sandhouse's (1943:586) view Parepeolus Ducke, 1912:71. 102. Type species: Leiopodiis designated. [Prosopis]. that (1939:169) a and thus lecointei Ducke. 1907 - Epeolus aterrima Friese. 1906, by Popov designated type species validated the name is inconect. designation of Sandhouse, 1943:585. Osirini. Hylaeinae. Pelandrena Cockerell. 1909f:159. Pelandrena Parepicharis Moure, 1945b:307. Type species: Epicharis zonata Type species: reducta Cockerell, 1909, monobasic. Fossil. Smith. 1854, by original designation. Centridini. Pelicandrena and Ribble. 1972:346. Pareucera Tkalcu, 1978:164. Type species: Eiicera caspica LaBerge Type species: Parandrena atypica Cockerell. 1 94 1 . Morawitz, 1873, by original designation. [Eucera]. Eucerini. by original designation. [Andrena]. Andreninae. Parevaspis Ritsema, 1874:lxxi. Type species: Parevaspis basa- Michener. 1965b:324. lis Ritsema. 1874. by designation of Sandhouse. 1943:585. Penapis Type species: Penapis penai Anthidiini. Michener. 1965. by original designation. Rophitinae. Peniella Toro and Ruz, 1972:146. Pariotrigona Moure, 1961:192. Type species: Trigona Type species: Camptopoeum maculatum 1851. pendlebiiryi Schwarz. 1939. by original designation. Spinola. by original designation. Meliponini. ]Spmoliella]. Panurginae. Pentadentosmia 1991a: 14. Osmia Paroxystoglossa Moure, 1940:59. Type species: Oxystoglossa Warncke, Type species: jocasta Schrottky. 1910. by original designation. quinquespinosa Friese, 1899. by original designation. Augochlorini. [Osmia]. Osmiini.

Partamona Schwarz. 1939b:23, replacement for Patera Schwarz, Pentaperdita Cockerell and Porter, 1899:414. Type species: Perdita 1938. Type species: Melipona testacea Klug. 1 807. autobasic albovittata Cockerell. 1895. monobasic. [Perdita]. and original designation. [Trigona]. Meliponini. Panurginae.

Pasiphae Spinola, 1851:226, not Latreille, 1819. Type species: Penteriades Michener and Sokal, 1957: 158. Type species: Osmia Pasiphae caerulescens Spinola, 1851. designated by remotula Cockerell. by original designation. [Proteriades]. Sandhouse, 1943:585. See Spinolapis. Colletinae. Osmiini. Genus-Group Names of Bees 47

Penthemisia Moure, 1950c:390. Type species: Hemisia chilensis Phileremidiis Cockerell, 1895:9. Type species: Philerenndiis Neolarrini. Spinola. 1851, by original designation. (Centris). Centridini. i7i,'(7rt/!.s Cockerell, 1895, by original designation. Peponapis Robertson. 1902b:324. Type species: Macrocera Phileremus Latreille, 1809: 169. Type species: Epeoliis piinctatiis pniinosa Say, 1837. by original designation. Eucerini. Fabricius, 1804. by designation of Latreille. 1810:439. Pluleroniis oraniensis 1841. designated as type Perdita Smith, 1853:128. Type species: Pcnllta lialictoides Lepeletier. Blanchard. 128 (1847) was not Smith, 1853. monobasic. Isotypic with Zaperdita by syn- species by 1849:pl. originally included. v^ith Aninwbaloides Schenk onymy. Panurginae. Isotypic by synonymy. " Ammobatini. PerditelUi Cockerell. 1 899b:3 1 2. Type species: Perdita "laiieae Philotracluisa Pasteels. 1969a:22. species: Aiii/iidiiun Cockerell, 1 899, error for /^ /«/•/«;(' Cockerell, 1896, by origi- Type Strand, 1912, [TrachusaJ. nal designation. [Perdita). Panurginae. ac/iiiflliini by original designation. Anthidiini. Perditomorpha Ashmead, 1899a:86. Type species: Pliiloxanthiis Ashmead, 1898:285. Perdita heatu Perditomorpha hnmerii Ashmead. 1899, monobasic. Type species: Colletinae. Cockerell, 1895, by original designation. Panurginae. P/io/- Robertson. 1903d: 174. 175. 176. Nonuida Penlitopasites Rozen, 1992:49. Type species; Oreopasites linsleyi Type species: Robertson. 1893 (not Brulle. 1832) = Noinuda Rozen, 1992, by original designation. [Oreopasites], ituegra Ammobatini. integerrima Dalla Torre, 1896, by original designation. Nomadini. Pereirapis Moure, 1943b:461. Type species: Pereirapis not 1819. rhizophiUi Moure, 1943, by original designation. Phyllotoma Dumeril. 1860:842. Leach. Type species: 1799. of Michener. Augochlorini. Apis papaveris Latreille. by designation 1941: 160. Not replaced because isotypic whh Anthocopa and Perezia Ferton. 1914:233. not Leger and Dubosc, 1909. Type = with Fiircosmia. Osmiini. species: Perezici iinnira Ferton. 1914 Megachile leucliella Curtis, 1828, monobasic. See Fertonella. Megachilini. Pliysostetha Michener. 1938a:523. Type species: Heriades carinatum Cresson. 1 864. by original designation. [Heriades]. Perissander Michener, 1942:275. Type species: Calliopsis Osmiini. anomoptera Michener, 1942, by original designation. Pileteiicera Sitdikov and 1988:87. Eiicera [Calliopsis]. Panurginae. Pesenko, Type species: cineraria Eversmann, 1852 = ?E. cinerea Lepeletier, 1841, Perixylocopa Hurd and Moure, 1963:232. Type species: by original designation. [Eucera]. Eucerini. Xylocopa eiythrina Gribodo, 1894, by original designation. [Xylocopaj. Isotypic with Euxylocopa by synonymy. Pithitis Klug, 1807, in lUiger, 1807:198: Klug. 1807b:225.Type Xylocopini. species: Apis smaragdida Fabricius. 1787. monobasic. The by Illiger and Klug were published simultaneously: Petalosteruon Brooks. 1988:484. Type species: Aiuhophora papers Illiger credited the name Piiliitis to Klug and it is customary rivolleri Perez. 1895, by original designation. [Anthophora]. and to do so. Ceratinini. Anthophorini. appropriate Phmiandrena 1983:794. Type species: Andrena Pluiceliapis Michener, 1942:277. Type species: Conanthalictiis Osychnyuk Morawitz. 1876, designation. hakeri Crawford, 1907, by original designation. planirostris by original [Andrena|. Andreninae. [ConanthalictusJ. Rophitinae. Houston. 1 98 1 ;96. Prosopis trilobata Phaenosanis Mitchell, 1934:303, 309. Type species: MegachUe Planihylaeiis Type species: Cockerell, 1910, original designation. [Hylaeusj. fortis Cresson. 1872. by original designation. [Megachile]. by Megachilini. Hylaeinae. Plastandrena Hedicke, 1933:217. Melitta tibialis Phaeosmia Michener. 1943:77. Type species: Osinia enceliae Type species: 1802, [Andrena]. Andreninae. Cockerell. 1935, by original designation. [Anthocopa]. Kirby, by original designation. Osmiini. Platandrena Viereck. 1924:21. Type species: Andrena nasonii Robertson, 1895. [Andrena|. Phauomalopsis Michener and Moure. 1957:430. Type species: by original designation. Andreninae. Exomcdopsis jenseni Friese. 1908, by original designation. [Exomalopsis]. Exomalopsini. Platinopoda Dalla Torre. 1896:202, lapsus for Platynopoda

Westwood, 1 840. Phcimhylaeiis Michener. 1965a: 141. Type species: Meroglossa Xylocopini. lactifera Cockerell. 1910. by original designation. Hylaeinae. Platosmia Warncke. 1990:482. Type species: Osniia platalea Wamcke, 1990, [Osmia]. Osmiini. Phelonoimida Snelling. 1986:24. Type species: Nomada helfmgei by original designation. Cresson, 1878, by original designation. [Nomada|. Noniadini. Platychile Michener, 1965a:205, nomen nudum. Type species: Smith, 1861. monobasic. Error for Phenacolletes Cockerell, 1 905b:30 1 . Type species: Plwnacolletes Megachile foliata OTiOTM.? Cockerell, 1905. monobasic. Colletinae. Callochile. [Megachile]. Megachilini. 1973:120. Phianis Gerstaecker. 1869:147. Type species: Phileremus Platycoelioxys Mitchell. Type species: Coeliaxys ahdominalis Eversmann. 1852. monobasic. Isotypic with spatiiliventer Cockerell. 1927, by original designation. Aiuniohatoides Radoszkowski and Pciidlu. Ammobatoidini. [Coelioxysj. Megachilini. of Kansas 48 Scientific Papers, Natural History Museum, The University

Warncke, 1968:71. Andrena polita Pesenko, 1984:347. Type species; Halictits minor Poliandremi Type species: Platyhalictiis Andreninae. Halictini. 1 847, [Andrena]. Morawitz. 1876, by original designation. [Halictus]. Smith, by original designation. herbsti 1 9 1 Oa:65 1 . Colletes Friese, Platynopoda Westwood. I840b;271. Type species: Apis latipes Policana Friese. Type species: 1899a:71. = 1851, of brury. 1773, by designation of Ashmead. 1910 Colletes albopilosiis Spinola, by designation 1943:589. [Xylocopa]. Isotypic with Audinetia. Xylocopini. Sandhouse, Diphaglossinae. 1916d:208. Platvspatiilariella Pittioni, 1950:80. Type species: Prosopis Polybiapis Cockerell, Type species: Polybiapis Nomadini. punctata Brulle, 1832, by original designation. [Prosopis]. mimus Cockerell, 1916, by original designation. Hylaeinae. Polyglossa Friese, 1909a: 123. Type species: Polyglossa capensis 1921a:203. Plat}sta Pasteels, 1965: 171. Type species: Megachile platystoma Friese, 1909. by designation of Cockerell. Pasteels, 1965. by original designation. [Megachile]. Sandhouse's (1943:589) subsequent designation of P. Megachilini. albitarsis Friese, 1909, is invalid. Colletinae. 148. Platysvastm Moure, 1967b: Type species: Platysvastra Pomibombus Skorikov, 1938a: 145. unjustified emendation of 1967, Eucerinl. macraspis Moure, by original designation. Pomobombus Kriiger, 1917. Bombini. Moure, 1961 :203. species: Trigonapkmifwns Platytrigona Type Pomobombus Kriiger, 1917:65. Type species: Bremus pomorum Smith, 1864, by original designation. Meliponini. Panzer. 1804. by designation of Sandhouse, 1943:589. Bonibini. Plebeia Schwarz, 1938:480. Type species: Trigona mosquito [Bombus|. Isotypic with Rhodobombus. Smith, 1863, [Trigona]. Meliponini. by original designation. Pontcmthidium Pasteels. 1969a: 105. Type species: Anthidium 1961:226. lendliana Pleheiella Moure, Type species: Trigona /7win.v Cockerell, 1933. by original designation. [Anthidium|. Friese, 1900, by original designation. Meliponini. Anthidiini. Moure. 1961:228. Meiipona denoiii Plebeina Type species: Popovapis Michener. 1981:80. Type species: Rhophites dispar Vachal, 1903, by original designation. Meliponini. Morawitz. 1892. by original designation. [Eremaphanta]. Plesianthidium Cameron, 1905:256. Type species: Dasypodinae. Cameron, 1905, by original des- Plesianthidiumfulvopilosum Porterapis Michener, 1989:678. Type species: Lonchopria Anthidiini. ignation. porteri Ruiz, 1936, by original designation. [Lonchopria|. Plesiopanurgus Cameron. 1907:130. Type species: Colletinae. cinerarius Cameron, 1907, monobasic. Plesiopanurgus Pratibombus Skorikov. 1937:59, unjustified emendation of Panurginae. Pratobombus Vogt, 1911. Bombini. Plistotriihia Morawitz, 1873a: 134. Type species: Nomia 191 1:49. pratorum 1841, Pratobombus Vogt. Type species: Apis Lucas, 1 846 = oraniensis Lepeletier, flavilabris Ancyla of 1927:67. Linnaeus, 1 761. by designation Frison, [Bombus]. monobasic. with Ancyla by synonymy. Ancylini. Isotypic Bombini. Plusia Hoffmannsegg 1817:52. not Hlibner. 1 806. Type species: Presbia Sandhouse. 1943:590, nomen nudum. See Daly et al., Plusia superba Hoffmannsegg, 1817, monobasic. See 1987. Manueliini. Euplusia. Euglossini. Pressibombus Frison, 1935:342. Type species: Bremus pressus Podalirius Latreille. 1802a:430. Type species: Apis pilipes Fab- Frison. 1935, [Bremus|. Bonibini. ricius. 1775 = Apis plumipes Pallas, 1772. Suppressed by by original designation. Direction 4. See Commission Opinion 151 (1944). Prionohoplitis Tkalcu. 1993a:55. Type species: Osmia curvipes and Osmiini. Anthophora. Isotypic with Megilla Anthophora. Morawitz, 1871, by original designation. [Hoplitis]. Anthopliorini. Proantludium Friese, 1898a: 101. Type species: Anthidium for = Podasxs Rafinesque. 1815:123. unjustified replacement oblongatum Latreille, \?,09 Anthophora oblongata llliger, 1802. Andrena Pasteels Dasypoda Latreille, Type species: lurtipes 1806, by designation of Cockerell, 1909b:269. Fabricius. 1793, autobasic. Dasypodinae. hsted Anthidimn undulatum Dours as the ( 1969a) incoiTectly

1 Andrena labiata Anthidiini. Poecilandrena Hedicke, 1 933:2 8. Type species: type species of Proanthidium. [Anthidium]. Fabricius, 1781. designation. [Andrena]. hirsutus by original Probombus Piton, 1940:218. Type species: Probombus Andreninae. Piton, 1940, monobasic. Fossil. Poecilobombus Dalla Torre, 1882:23. species: Bombus Type Prochelostoma Robertson, 1903a: 167. Type species: Heriades sitkensis Nylander, 1848, by designation of Sandhouse, philadelphi Robertson, 1891, by original designation. 1943:589. Bonibini. [Bombus]. Osmiini. Poecilocentris Friese. 1900:244. 252. Type species: Centris Timberlake. 1954:402. Perdita of Sandhouse. Procockerellia Type species: fasciatella Friese, 1900, by designation albonotata Timberlake. 1954, by original designation. 1943:589. [Centris). Centridini. [Perdita). Panurginae. Poecilomelitta Friese, 1913a:574 and 1913b:585. Type species: Gaesischia of LaBerge, 1958:199. Type species: PoecUomelitta flavida Friese, 1913, by designation Prodasyhalonia 1958, designation. Sandhouse, 1943:589. Poecilomelitta was described as new mexicana LaBerge, by original Eucerini. twice. Panurginae. [Gaesischia]. Genus-Group Names of Bees 49

= Prodiaxys Friese, 1914a:221. Type species: Prodio.xys Anthidium: see Michener and Griswold, 1994a) Ainhidium = ciniuiharina Friese, 1914 Dioxys longiventris Perez, 1 895, loiigiconie Friese, 1902, by original designation. Antliidiini. monobasic. Dioxyini. Protepeolus Linsley and Michener. 1937:75. Type species: and Pwhalictiis Pesenko, 1984:346, not Armbmster, 1938. Type spe- Protepeolus singularis Linsley Michener, 1937, by original cies: Apis riibiciinda Christ. 1791, by original designation. designation. Protepeolini. Halictini. [Halictus]. See PwtohaliclHS. Proteraner Robertson, 1903b: 103. Type species: Sphecodes ra- nunculi Robertson. 1897. monobasic. Halictini. Proniflittci Warncke, 1977:59. Type species: Dufoiirea ulhoclypeata Frie.se, 1900, by original designation. [Melitta]. ProteriadesTxXus, 1904a:25. Type species: Proteriades semirubra Dasypodinae. Cockerell, 1898, monobasic. Osmiini. Praplebeia Michener, 1982:44. Type species: Trigona Protoanthidium Cameron, 1902b: 125. Type species: Joininicana Wille and Chandler, 1964, by original designa- Protoanthidium rufohalteatum Cameron, 1902, by designation. [Trigonal. Fossil. Meliponini. tion of Sandhouse, 1943:591. Anthidiini.

Prosamha Michener, 1981:63. Type species: Haplomelitta Protobombus Cockerell, 1908d:326; also described as new by griseonigra Michener, 1981, by original designation. Cockerell, 1909e:9. Type species: Protobombus indecisus also [Haplomelitta]. Dasypodinae. Cockerell. 1908. monobasic; designated by Cockerell. 1909e: 10. Fossil. Pmsapis Ashmead, 1 894:43, unjustified emendation of Prosopis Protodiscelis 1909a:245. Protodiscelis Fabricius, 1804. Hylaeinae. Brethes, Type species: fiebrigi Brethes. 1909. monobasic. Colletinae. Prosopalictiis Strand, 1913b:26. Type species: Prosopalictus 1955:105. micans Strand. 1913 (not Halictus micans Strand, 1909, se- Protodufourea Timberlake, Type species: wasbaueriTimbevXake.. 1955, des- nior homonym in Lasioglossum) = Lcisioglossum micante Protodufourea by original ignation. Rophitinae. Michener. 1993, by original designation. Halictini.

1 1 for ictus Protohulictus Pesenko. 986b:63 . replacement Prohal Prosopella Snelling. 1966:139. Type species: Hyhwus hurdi Pesenko, 1984. Type species: Apis rubicunda Christ, 1791, Snelling. 1966, by original designation. [Hylaeus]. Hylaeinae. autobasic. [Halictus]. Halictini. Prosopis Jurine, 1801:164. Suppressed by Commission Opin- Protomelecta Cockerell, 1908c:341 . Type species: Protomelecta ion 135(1939). Hylaeinae. brevipennis CockersW, 1908, monobasic. Fossil. Prosopis Fabricius, 1 804:293. Type species: Melliniis hipimctatus Protomelissa Friese, 1914c(June):322. Type species: Fabricius, 1798 = Sphex signata Panzer, 1798, designated by Protomelissa iridescens Friese, 1914(June) - Tetralonia (?) Morice and Durrant, 1915:416. Isotypic with Cingulata by hoozana Strand. 19 i4( Apr.-May ). by designation of synonymy. Hylaeinae. Sandhouse. 1943:592. Lieftinck's (1983) contention that the Hirashima, 1967:134. Prosopisteroides Type species: Hylaeus type species is Anthophora sauteri Friese, 1911(May) = heteroclitis Hirashima, 1967, by original designation. Melecta formosana Cockerell, 1911(Mar.), is not justified. [Hylaeus]. Hylaeinae. Isotypic with Tetralonioidella by synonymy. Melectini.

Prosopisteron Cockerell, 1 906b: 17. Type species: Prosopisteron Protomeliturga Ducke, 1912:63, 90. Type species: Calliopsis serotinellum Cockerell. 1906, monobasic. Hylaeinae. turnerae Ducke, 1907, monobasic. Panurginae.

Prosopoides Friese, 1908b:338 (reprint p. 10). Type species: Protomorpha Rayment. 1959b:334. Type species: Protomorpha Oediscelis paradoxus "Ducke" Friese, 1908 = Oediscelis tarsalis Rayment. 1959, by original designation. Colletinae. Ducke, 1907, monobasic. The name prosopoides Prosopoides Protopithitis Hirashima, 1969:651. Type species: Ceratina was rather Friese and attributed to published casually by aereola Vachal, 1903, by original designation. [Pithitis]. Ducke. However, there was no description by Ducke and I Ceratinini.

1 in the to Friese. agree with Sandhouse ( 943 ) attributing name Protosiris Roig-Alsina, 1989:20. Type species: Osiris obtusus Xeromelissinae. Michener, 1954, by original designation. Osirini. Prosopoxylocopa Hurd and Moure, 1963:215. Type species: Protosmia Ducke. 1900:12. Type species: Heriades glutinosus mirabilis Hurd and Moure, 1963. des- Xylocopa by original Giraud, 1871, designation by Michener. 1943:86 and ignation. [Xylocopa]. Xylocopini. Sandhouse, 1943:592. [Osmia]. Osmiini.

Protandrena Cockerell. 1 896:91 . Andrena maurula Type species: Protostelis Friese. 1895:25. Type species: Stelis freygessneri Cockerell. 1896. of Sandhouse, 1943:591. by designation Friese, 1885 = Anthidium annulatum Lepeletier, 1841, by Panurginae. designation of Popov. 1938:41. [Stelis]. Anthidiini. 1903:337. Protaitdreuopsis Crawford, Type species: Protoxaea Cockerell and Porter. 1899:410. Type species: 1903 = Protandrenopsis fuscipennis Crawford, Pamtrgus Megacilissa gloriosa Fox, 1893. by original designation. aethiops Cres.son, 1872, by original designation. Isotypic with Oxaeinae. Pseudopanurgus by synonymy. Panurginae. Proxylocopa Hedicke. 1 938: 1 92. Type species: Xylocopa olivieri Protanthidium Cockerell and Cockerell, 1901 :49. Type species: Lepeletier, 1841, by original designation. [Xylocopa]. Megachile steloides Bingham, 1896 (preoccupied in Xylocopini. 50 Scientific Papers. Natural History Museum, The University of Kansas

Psaenythia Gerstaecker, 1868:111. Type species: Psaenylhia Pseiidoheriades Peters. 1970: 153. Type species: Eriades moricei philanthoides Gerstaecker. 1868, by designation of Friese. 1897, by original designation. Osmiini. Sandhouse, 1943:592. Panurginae. Pseiidolipliantliiis Ruz and Toro. 1983:253. Type species: Psammandrena LaBerge. 1977:83. Type species: Andrena Liphanthus rozeni Ruz and Toro. 1983, by original designa- cercocarpi Cockerell, 1936, by original designation. tion. [Liphanthus]. Panurginae. [Andrena]. Andreninae. Pseiidoiuacrotera Timberlake, 1954:349. Type species: Perdita 1909a: 145. Pseudagapostemon Schrottky, Type species: tiirgicepsT\mbex\2ke. 1954, by original designation. [Perdita]. Agapostemou aremihus Schrottky. 1902, by original desig- Panurginae. nation. Halictini. Pseiidomegacliile Friese, 1 898b: 1 98, no included species; Friese, W. F. 1900:15. Pseudapis Kirby, Type species: Pseiidapis 1899c:36, included species. Type species: Megachile anomala W. F. 1900. monobasic. Nomiinae. Kirby. ericetonim Lepeletier. 1841. designated by Alfken, 1933:56. Pseiidaiigochlora Michener. 1954:77. Type species: Halictiis [Megachile]. Megachilini. 1851 = Fabri- nigromarginatus Spinola, Megilla graminea Pseudomegalochila Schulz, 1906:71, unjustified emendation of cius. 1804, by original designation. [Caenaugochlora]. Pseiidomegacliile Friese, 1898. [Megachile]. Megachilini. Augochlorini. Pseiidomelecta Radoszkowski, 1865:55. Type species: Melecta 1906a:313. Type species: Pseiidaiigochloropsis Schrottky, diacantha Eversmann, 1852. by designation of Sandhouse. sthena of Augochloropsis Schrottky, 1906, by designation 1943:594. Melectlni. Sandhouse, 1943:593. Halictiis nigromarginatus Spinola. Pseiido-osmia Radoszkowski. 1874a: 137. unjustified emenda- designated as the type species by Moure, 1940:51. was not tion of Pseiidosmia Radoszkowski. 1 872. Osmiini. an originally included species; see Michener. 1995. [Augochloropsis]. Augochlorini. Pseiidoosmia Radoszkowski, 1 874b: 1 52, unjustified emendation oi' Pseiidosmia Radoszkowski. 1872. Osmiini. Pseitdepeohis Holmberg. 1886b:234, 284. Type species: Pseudepeolus fasciatiis Holmberg, 1886, monobasic. Pseudopaniirgus Cockerell. 1897a:290. Type species: Panurgiis Epeolini. aethiops Cresson. 1 872. by original designation. Isotypic with

Pseiideiicera Tkalcu, 1978:157, 158. Type species: Eucera Protandrenopsis by synonymy. Panurginae. pan'iconiis Mtxsai^, 1878, by original designation. Eucerini. Pseiidopasites Bischoff, 1923:593. Type species: Pasites Friese. 1922. of Sandhouse, Pseudhylaeus Cockerell, 1929b:299. Type species: Einyglossa pygmaeus by designation 1943:594. Ammobatini. alhociineata Cockerell, 1913, by original designation. [Sphecodopsis], Hylaeinae. Pseiidopeohis Ashmead, I899a:80. unjustified emendation of 1886. Pseudiscelis Friese, I906a:228. Type species: Pseiidiscelis Pseudepeolus Holmberg. Epeolini. rostratu monobasic. The was de- Friese, 1906, type species Pseiidophilaiuluis Alfken, 1939:121. Type species: scribed in this combination the name although genus-group PseudophiUuitluts taeniatus Alfken, 1939, monobasic. was as a of Oediscelis. proposed subgenus [Oediscelis], Meganomiinae. Xeromelissinae. Pseitdosarus Ruz, 1980:25. Type species: Pseiidosarus virescens Pseudoanthidimu Frie.se, 1898a:101. Anthidium Type species: Ruz, 1980, monobasic. Panurginae. alpiniiniMoTdw'nz. 1 873, designated by Sandhouse. 1943:593. Pseudosmia Radoszkowski, 1872a:xviii, no included [Anthidium]. Anthidiini. species; Radoszkowski, 1874b: 152, included species. Type species: Pseudobranchiata Mehdy. 1935:33. 139. Invalid because no type = Megachile cristatu Fonscolombe. 1 846. not Dufour, 1 84 1 species was designated. Sandhouse's (1943:593) view that Anthocopa cristatula Zanden, 1990, by designation of Popov (1939) designated a type species and thus validated Cockerell. 1922a:6. Osmiini. this name is incorrect. [Prosopis]. Hylaeinae. Pseudostelis Popov, 1 956: 1 67. Type species: Stelis stniiuli Popov, Pseitdocentrou Mitchell, 1934:307. Type species: Megachile 1935, by original designation. Anthidiini. pniina Smith, 1853, by original designation. Megachilini. Psilomelissodes LaBerge. 1 956: 1173. Type species: Melissodes Psciidochilalictiis Michener, 1965a: 170. Type species: intorta Cresson, 1 872. by original designation. [Melissodes]. Lasioglossiim imitator Michener, 1965, by original designa- Eucerini. tion. [Lasioglossum]. Halictini. Psilylaeus Snelling. 1985:28. Type species: Psilylaeus sugiops Pseudocilissa Radoszkowski, 1891:241. Type species: Cilissa 1985 = Cockerell, 1914, robiista Radoszkowski, 1 876 = Melitta dimidiata Morawitz, Snelling. Prosopis pciimmilis by 1876, monobasic. Melittinae. original designation. Hylaeinae. 1832:373. Pseudocosmia Radoszkowski, 1886:14, unjustified emendation Psithyrus Lepeletier, Type species: Apis rupestris of Pseudosmia Radoszkowski, 1872. Osmiini. Fabricius. 1793, by designation of Curtis. 1833:pl. 468. Isotypic with Biennis Kirby. Bombini. Pseudodichroa Bischoff, 1923:595. Type species: Omaclites capensis Friese, 1915. by designation of Sandhouse, 1943:593. Pteneucera Tkalcu, 1984c:72. Type species: Eucera eiicnemidea Ammobatini. Dours, 1873, by original designation. Eucerini. Genus-Group Names of Bees 51

a of RhinocoUetes. Plemndrena Robertson. 1 902a: 1 87. 1 93. Type species: Audreua Pimcticolletes synonym [Colletes]. = piilcheUa Robertson, 1891 (not Jurine, 1807) Andrena Colletinae. Andreninae. accepta Vicveck. 1916, by original designation. Pycnanthidium Krombein, 1951:292. Type species: solonionis des- Pteiosariis Timberlake. 1967:10. Type species: Calliopsis Pycnanthidiiiin Krombein. 1951. by original ntdheckiae Robertson. 1895, by original designation. ignation. Anthidiini. [Pseudopanurgus]. Panurginae. Pyganthophora Brooks, 1988:460. Type species: Apis retusa 1758, Ptikindrena Robertson. 1902a: 1 87. 192. Type species: Andrena Linnaeus, by original designation. [Anthophora]. erigeniae Robertson, 1891, by original designation. Anthophorini. Andreninae. Pyghalictus Warncke, 1975b: 103. Type species: Andrena Fabricius, 1804, Ptiloccntris Snelling. 1984:22. Type species: Centris festiva pygmaea by original designation. Pyghalictus was intended for the of what is called Smith, 1854, by original designation. [Centris]. Centridini. group usually Lasioglossiim politum (Schenck). The identity of Andrena Ptiloclepris Michener. 1978b:315. Type species: Ptilocleptis pygmaea Fabricius is in doubt (see Ebmer, 1988). [Halictus]. ronientosa Michener, 1978, by original designation. Halietini. Halietini. P!il(>i>lossa Smith, 1853:7. Type species: Ptiloglossa diicalis Pygnanthidiellwn Mavromoustakis, 1963b:492. Type species: Smith, 1853, monobasic. Caupolicanini. Anthidiiim zebra Friese. 1904. by original designation. Moure. 1953a:73. Piiloglossidia Type species: Ptiloglossidia [Pygnanthidiumj. Anthidiini. fcdkix Moure, 1953, by original designation. Dissoglottini. Pygnanthidium Mavromoustakis. 1963b:491. unjustified emen- 1945a:153. Ptiloglossodes Moure, Type species: MegacUissa dation oi' Pycnanthidium Krombein. 1951. Anthidiini. tarsata Friese. 1900. by original designation. [Ptiloglossa]. Pygopasiphae Michener, 1989:647. Type species: Leioproctus Caupolicanini. mourellus Michener. 1989, by original designation. Ptilomelissa Moure, 1943b:482. Type species: Melissoptila [Leioproctus]. Colletinae. bonaerensis Holmberg, 1903, by original designation. Pygaperdita Timberlake, 1956:275. Type species: Perdita Eucerini. interrupta Cresson. 1878, by original designation. JPerditaj. 1 Noniia PtUonomia Michener, 1965a: 60. Type species: plumosa Panurginae. Michener, 1965, by original designation. [Nomia]. Nomiinae. Pyrobombus Dalla Torre, 1 880:40. Type species: Apis hypnorum Ptilopoda Friese. 1 92 1 b:83. Type species: CoUetes niaciilipennis Linnaeus. 1758, monobasic. [Bombus]. Isotypic with = Friese. 1921 CoUetes spiloptera CockereW. 1917, monoba- Anodontobombus. Uncobombus, and Hypnorobombus. sic. [Colletes]. Colletinae. Bonibini. 1980:63. Ptilosawides Mitchell. Type species: Megachile Pyrosmia Tkalcu, 1975b: 182. Type species: Osmia ferruginea neo.xcinthoptera Cockerell, 1933, by original designation. Latreille. 181 1, by original designation. [Osmia]. Osmiini. [Cressoniella]. Megachilini. Pyrrhobombus Dalla Torre, 1882:28, unjustified emendation of Ptilosanis Mitchell, 1943:667. Type species: Megachile hertonii Pyrobombus Dalla Torre, 1880. [Bombus]. Bombini. Schrottky. 1908. by original designation. [Megachile|. Pyrrhomelecta Ashmead, 1899a:66. Type species: Epeolus Megachilini. glabratus Cresson, 1878, by original designation. Epeolini. Ptilothrix Smith. 1853:131. Type species: Ptiloihrix pliinmtus Quasihesma Exley. 1968a:228. Type species: Quasihesma Smith. 1 853. monobasic. with Energoponus by syn- Isotypic moonhiensis Exley, 1968, by original designation. onymy. Emphorini. Euryglossinae. Marschall, 1873:269, emendation of Ptilothryx unjustified Quasilictus Walker. 1986:166. Type species: Homalictus Ptilothrix Smith, 1853. Emphorini. breviconiutus Walker. 1986, by original designation. Ptilotopus Klug, 1810:32. Type species: Ptilotopiis uniericamts [Homalictus]. Halietini. monobasic. Centridini. Klug. 1810. Radoszkowskiana Popov. 1955b:547. replacement for Ptilotrigona Moure. 1951a:47. Type species: Trigono heideri Paracoelioxys Radoszkowski. 1893. Type species: Friese, 1900 = Trigona liirida Smith. 1854. by original des- Paracoelioxys barrel Radoszkowski. 1893. autobasic and by ignation. [Trigonaj. Meliponini. original designation. Megachilini. Ptoleglossa Friese, 1930:127, replacement for Leptoglossa Ramalictus Pesenko, 1984:347. Type species: Halictus Friese, 1925. Type species: Leptoglossa paradoxa Friese, 1925 latisignatus Cameron, 1908, by original designation, = Orphana inquirenda Vachal. 1909, autobasic. Isotypic with [Halictus). Halietini. and with Andreninae. Leptoglossa by synonymy Orphana. Ramphorhiiui Michener. 1939:8, lapsus for Arogochila Michener, Piincthalictus Warncke, 1975b:87. Type species: Hylaeus 1939. [Ashmeadiella]. Osmiini. Schenck, 1853. punctatissimiis by original designation. Ranthidiellum Pasteels. 1969a:48. Type species: Protoanthidium Halietini. [Halictus]. rufoniaculatuni Cameron. 1902. by original designation. For Pimcticolletes Noskiewicz. 1936:26, 490. Invalid because no type aspects of this type designation, see Michener and Griswold, species was designated. Warncke (1978) considered 1994a. [Anthidiellum]. Anthidiini. 52 Scientific Papers. Natural History Museum, The University of Kansas

Rapanthidium Michener, 1948a:ll. Type species: Anthiclium Rhinochaetida Friese, 1912:185. Type species: Capicola vespoides Friese. 1925. by original designation. cinctiventris Friese. 1912. by designation of Cockerell, [Parantiiidium]. Anthidiini. 1915:343. Dasypodinae.

Raphidostoma Coctcerell. 1936a: 133. Type species: Rhinocoelioxys Mitchell, 1973:113. Type species: Coelioxys = Raphidostoma ceanothi Cockerell, 1936 Chelynia mhifloris zapoteca Cresson. 1 878, by original designation. [Coelioxys]. Cockerell. 1898, by original designation. Isotypic with Megachiiini. Chelostomopsis by synonymy. Osmiini. Rhinocolletes Cockerell, 1910a:242. Type species: Colietes

Rathymiis Smith. 1 854:278, not Dejean, 1 83 1 . unjustified emen- nasutus Smith, 1853, monobasic. [Colletes|. Colletinae. dation of Rhuthymus Lepeletier and Serville, 1828. Rhinocorynura Schrottky, 1909a; 147. Type species: Halictus Rhathymini. inflaticeps Ducke, 1906, by original designation. Isotypic with Reanthidiuiu Pasteels, 1969a:87. Type species: Anthidium Ctenocoiynuru by synonymy. Augochlorini. reticiilatitm Mocsary, 1884, original designation. by Rhinomegilla Brooks, 1988:482. Type species: Anthophora Anthidiini. megarrhina Cockerell, 1910, by original designation. Redivha Friese. 191 la:671. Type species: Andrena peringueyi [Anthophora]. Anthophorini. Friese. 1911, designation of Cockerell, 1931a:402. by Rhodaniltidiitm Isensee. 1927:374. Type species: Anthidium [Andrena]. Melittinae. sicuhim Spinola, 1838, by original designation. [Anthidium]. Redivivoides Michener. 1981:42. Type species: Redivivoides Anthidiini.

simukms Michener, 198 1 . by original designation. Melittinae. Rhodobombus Dalla Torre, 1880:40. Type species: Bremus Reedapis Michener, 1989:656. Type species: Leioproctus pomorum Panzer, 1804, by designation of Sandhouse, bathycyaneus Toro, 1973, by original designation. 1943:596. [Bombus]. Isotypic with Pomohombus. Bombini. [Leioproctus]. Colletinae. Rhodocentris Friese, 1900:244, 250. Type species: Centris Reepenia Friese, 1909b:205. Type species: Nomia variabilis difformis Smith, 1854, by designation of Sandhouse, Friese, 1909, by designation of Sandhouse, 1943:595, 1943:596. [Centris]. Isotypic with Heterocentris and Michener, 1965a: 159, and Commission Opinion 788 (1966). Gundlachia by synonymy. Centridini.

is not as indicated ( 1 The name monobasic by Sandhouse 943 ) Rhodohylaeus Michener, 1 965a: 1 24. Type species: Prosopis and by the Offcial List of Generic Names of the International cenibera Cockerell. 1910, by original designation. [Hylaeus]. Commission on Nomenclature, since Friese cited Zoological Hylaeinae. additional species-group names as "varieties" of variabilis. Rhodomegachile Michener, 1965a:20I . Type species: Megachile [Nomia]. Nomiinae. abdominalis Smith. 1853, by original designation. Rhacandrena 1977:90. Andrena LaBerge, Type species: [Chalicodoma]. Megachiiini. brevipalpis Cockerell, 1930, by original designation. Rhodosmia Michener, 1943:85. Type species: Osmia paradoxa [Andrena]. Andreninae. Friese. 1899, by original designation. [Anthocopa]. Osmiini. Rhamphorhina Michener and Sokal. 1957: 159, unjustified emen- Rhogepeolus Moure, 1955:117. Type species: Rhogepeohis dation of Ramphorliina Michener, 1939, lapsus for Arogochila bigibbosus Moure. 1955, by original designation. Epeolini. Michener, 1939. [Ashmeadiella]. Osmiini. Rhopalictus Sichel. 1867:146. Type species: Corynura Rhanthidielhtm Pasteels, 1972:102, unjustified emendation of flavofasciata Spinola, 1851 = Halictus chilensis Ranthidiellum Pasteels, 1969. [Anthidiellum]. Anthidiini. Spinola, 1851, designated by Sandhouse, 1943:596. Augochlorini. Rluiphaudrena LaBerge, 1971b:507. Type species: Andrena Rhopalolemma Roig-Alsina, 1991:33. species: prima Casad and Cockerell, 1896, by original designation. Type robertsi des- [Andrena]. Andreninae. Rhopalolemma Roig-Alsina, 1991, by original ignation. Biastini. Rhuthymus Lepeletier and Serville, 1828:448. Type species: Alfken. 1926b:267. Rhathymus bicolor Lepeletier and Serville, 1828, monoba- Rhopalomelissa Type species: Alfken, 1 926, sic. Isotypic with Colax; isotypic with Liogastra by synonymy. Rhopalomelissa xanthogaster by designation Rhathymini. of Sandhouse, 1943:596. Nomiinae.

Rhectomia Moure. 1947a:9. Type species: Rhectomia pumilla Rhophites Agassiz. 1846:29 [Hymenoptera section], unjustified Moure, 1947, by original designation. Augochlorini. emendation of Rophites Spinola, 1808. Rophitinae.

Rhinepeohis Moure, 1955: 1 15. Type species: Epeohis rufiventris Rhophitoides Schenck, 1861:69 (for the date see Michener, Friese, 1908, by original designation. Epeolini. 1986a). Type species: Rhophitoides distinguendus Schenck, 1861 = cana Eversmann, 1852, monobasic. Rhineta Illiger, 1807: 198. Type species: Nomada schottii Fabri- Rhophites cius, 1804 = Tiphiu brcvicornis Panzer, 1798, monobasic. Rophitinae. with Biastes Biastini. Isotypic by synonymy. Rhophitulus Ducke, 1907:366. Type species: Rhophitulus frisei Rhinetula Friese, I922b:581. Type species: Rhinetida denticriis Ducke. 1907, by designation of Sandhouse, 1943:597. Friese, 1922, by designation of Sandhouse, 1943:596. Sandhouse corrected the spelling of the species name lofriesei Halictini. but since Ducke used frisei on pages 366 and 368, he must Genus-Group Names of Bees 53

1980: 121 . Halictus weenenicus have intended that speUing although naming the species for Rugalictus Pauly. Type species: = CockcxtW. 1940, H. Friese. Panurginae. Cockerell, 1941 Halictus retigems by origi- with Dictx'ohalictus nal designation. [Pachyhalicms]. Isotypic Rhynchalictus Moure. 1947a:5. Type species: Rhynchalktus by synonymy. Halictini. rostratus Moure. 1947, by original designation. Halictini. 1990:103. Type species: Nouiia nigiventris Rhynchocolletes Moure. I943b:447. Type species: Ruginomia Pauly, Friese. 1930, Nomiinae. RhxnchocoUetes cilhiciiuliis Moure. 1943. by original desig- by original designation. nation. Colletinae. Ruizantheda Moure. 1964b:265.Type species: Halictus proximus 1851, Halictini. Rhynostelis Moure and Urban. 1994:297. Type species: Spinola. by original designation. Halictus Authidiiim multlplicatiiiii Smith. 1879. by original designa- Ruizanthedella Moure, 1964b:267. Type species: tion. Anthidiini. mutabilis Spinola. 1851, by original designation. Halictini. Rhyso.xylocopa Hurd and Moure. 1963:178. Type species: [Ruizantheda]. Xxlocopa caniabriia Lepeletier. 1841, by original designa- Ruizapis Timberiake, 1953:598, emendation of Ruziapis tion. [Xylocopa]. Xylocopini. Timberiake, 1952. Panurginae. Rhyssomegachile Mitchell. 1980:63. Type species: Megachile Ruiziella Timberiake, 1952a: 105 (April), not Cortes, 1952 Friese. similliimi Smith. 1 853. by original designation. [Cressoniella]. (March). Type species: Camptopoeum ochraceum and Megachilini. 1908, by original designation. See Ruziapis Ruizapis. with Rhxffiiceni Sitdikov and Pesenko. 1988:87. Type species: Eucera Isotypic Arhysosage by synonymy. Panurginae. and parvula Friese, 1895, by original designation. [Eucera]. Ruziapis Timberiake, 1 952b:528, error for Ruizapis replace- Eucerini. ment for Ruiziella Timberiake, 1952. Panurginae.

cirsii Rivali.sia Strand, 1921:270. Type species: Rivalisia metalUca Sfl/j(///c'o/fl Verhoeff. 1890:328. Type species: Sabulicola

= monoba- = 1 793, monoba- Strand. 1921 ? Augochlom argenma¥nt

Robustobombus Skorikov, 1922b: 157. Type species: Bombm Salemanthidium Pasteels, 1969a:5 1 . Type species: Hypanthidium rofo«m.v Smith. 1854. by designation of Sandhouse, 1943:597. salemense Cockerell. 1919. by original designation. [Alpigenobombus]. Bombini. [Eoanthidium]. Anthidiini. Rophites Spinola. 1808:8, 72. Type species: Rophites Samba Friese, 1908a:568. Type species: Samba calcarata Friese. quinquesphwsus Spinola, 1808, monobasic. Rophitinae. 1908, monobasic. Dasypodinae. Hcillctiis mourei Ur- RostratUapis Friese. 1914b:26. Type species: Santiago Urban, 1989:1 17. Type species: Scmtiago macrognathus Friese, 1914, by designation of Sandhouse, ban, 1989, by original designation. Eucerini. 1943:597. Halictini. [Halictus]. Saranthidium Moure and Hurd. 1960:6. Type species: Anthidium Halictus Rostrolmlictus Warncke. 1975b:88. Type species: flavoputum Smith, 1854, by original designation; incon-ectly longirostris Morawitz. 1876. by original designation. stated to be Anthidium furcatum Ducke, 1908, by Griswold [Halictus]. Halictini. andMichener. 1988:321. [Hypanthidium]. Anthidiini.

Roussyana Manning, 1961:306. Type species: Apis Sawcolletes Michener, 1989:643. Type species: Lonchopriu 1937. palmnickenensis Roussy. by original designation. rufipennis Cockerell, 1917, by original designation. Fossil. [Electrapis]. [Leioproctus]. Colletinae. Pasteels, 1969a:86. Anthidiuin Royanthidium Type species: Sarogaster Robertson, 1918a:92, replacement for Gnathodon melunurum 1832, Anthidiini. Klug, by original designation. Robertson. 1903. Type species: Megachile georgica Cresson, 154. Bombus Rubicimdobombus Skorikov, 1922b: Type species: 1 878, autobasic. Megachilini. ntbicundus Smith. 1854, of Sandhouse, by designation Saropoda Latreille. 1809:177, unnecessary replacement for 1943:597. [Fervidobombus]. Bombini. bimaculata Panzer. Heliophila Klug, 1 807. Type species: Apis 1920:350. with two included (but Ruderariobombiis Kruger, Type species: Apis 1 798, autobasic. Saropoda was named of Yarrow, 1971:27. to be a nideraria Miiller. 1776. by designation synonymous) species: it was also stated replacement fixation [Bombus]. Bombini. fox Heliophila Klug. Sandhouse (1943) regarded the Andrena as automatic (autobasic) since is monobasic. Apis Riifaudremi Warncke. 1 968:33. Type species: nifiventris Heliophila 1798 = A. bimaculata Panzer, 1798, Lepeletier. 1841. by original designation. [Andrena]. rotundata Panzer. desig- Andreninae. nated by Westwood, 1840a:86, was not originally included Bombus in Heliophila. Anthophorini. Rufipedibombiis Skorikov. 1922b: 156. Type species: Bombini. for Gnathocera rufipes Lepeletier, 1836, monobasic. Sayapis Titus, 1906:154, replacement Provancher, 1882, and Ceratias Robertson. 1903. Type spe- Rufocinctobombus Prison. 1927:78. Type species: Bombus 1 837, autobasic. rufocinctus Cresson, 1863, monobasic. [Bremus]. Bombini. cies: Megachile pugnatus Say, Megachilini. 54 Scientific Papers, Natural History Museum, The University of Kansas

Scaphandrena Lanham, 1949:200. Type species: Andrena Scrapter Lepeletier and Serville, 1828:403, not Scrapter montroseusis Viereck and Cockerel!, 1914. by original des- Lepeletier, 1841. Type species: Scrapter bicolor Lepeletier Andreninae. ignation. I Andrena]. and Serville, 1828, by designation of Vachal, 1897:63. Ashmead's of Scaptotrigoiia Moure. 1942:315. Type species: Trigomi postica (1899a:84) subsequent designation Scrapter hrullei Latreille. 1807, by original designation. [Trigona]. Lepeletier, 1841, as the type species of "Scrapter Meliponini. Lepel." concerns Scrapter Lepeletier, 1841 (= Panurginus). which was proposed as a new genus, with no reference to Scaiira Schwarz. 1938:479. Type species: Trigona Uiiitarsis Scrapter Lepeletier and Serville, 1828. Cockerell's Frie.se, 1900, by original designation. [Trigona]. Meliponini. (1920a:254) subsequent designation of Andrena lagopus Schizandrena Hedicke, 1933:2 1 8. Type species: Andrena aiilica Latreille, 1809, as the type species of "Scrapter Lepeletier, Morawitz, 1876 - Melitta himacidata 1802. Kirby, by origi- 1825" is invalid. Colletinae. nal designation. [Andrena]. Andreninae. Scrapter Lepeletier, 1841:260, not Scrapter Lepeletier and SchizocoeUoxys Mitchell. 1973:50. Type species: Coelioxxs Serville, 1 828. brullei 1 1 Type species: Scrapter Lepeletier, 84 , funeraria Smith, 1854. by original designation. The spelling by designation of Ashmead, 1899a:84. Not replaced because Schizococoelioxys in the heading of Mitcheirs description a synonym of Panurginus. Panurginae. (p. 50) is corrected on the same page and elsewhere in the work. Scrapteroides Gribodo, 1894:1 12. Type species: Scrapteroides [Coelioxys]. Megachilini. = dijfonnis Gribodo, 1 894 Panurgus cmmdatus Sichel, 1 859, Schizomegachile Michener, 1 965a: 1 99. Type species: Megachile monobasic. Panurginae. monstrosa Smith, 1868, by original designation. Viereck, 1922:42. Andrena [Chalicodoma]. Megachilini. Scrapteropsis Type species: fenningeri Viereck, 1922, monobasic. [Andrena]. Schmiedeknechiia 1896:277. Friese, Type species: Andreninae. Schmiedeknechtia oraniensis Friese, 1896, monobasic. Ammobatoidini. Seladonia Robertson, 1918a:91. Type species: Apis seladonia Fabricius, 1794, by original designation. Halictini. Schoenherria Hind and Moure, 1963:1 18. unjustified emenda- Sellalictus 1980:120. Halictus latesellatus tion of Sclionnherria Lepeletier, 1841. [Xylocopa]. Pauly, Type species: Cockerell. 1937, Xylocopini. by original designation. [Lasioglossum]. Halictini. Sclumherria Dalla Torre, 1896, 202, unjustified emendation of Semicarinella Pasteels, 1984:32. Pseudoanthidium Schonnherria Lepeletier, 1841. [Xylocopa]. Xylocopini. Type species: latitarse Pasteels, 1984, by original designation. Sclionnherria Lepeletier, 1841:207. Type species: Xylocopa . [Pseudoanthidium I Anthidiini. micans Lepeletier, 1841. by designation of Sandhouse, Senexihomhus Prison, 1930:3. Bombus senex 1943:598. [Xylocopa]. Xylocopini.^ Type species: Vollenhoven, 1873, original [Bremus]. Friese. 1908c: 170, no included by designation. Schrottkya species. Type spe- Bombini. cies: Tetrapedia goeldiana Friese. 1899, first included spe- Prison. 1927:64. Bombus cies, by Friese, 1908b:58. Isotypic with Lanthunomelissa by Separatohondnis Type species: 1863 = synonymy. Tapinotaspidini. separatus Cresson, Apis griseocollis DeGeer, 1773, by original designation. [Bremus]. Bombini. Schrottkyapis Mitchell. 1980:46. Type species: Megachile Smith. 1854:218. not Link. 1830. assumplionis Schrottky, 1908, by original designation. Serapis Type species: Serapis [Eumegachile]. Megachilini. denticulatus Smith, 1854, monobasic. See Serapista. Anthidiini. Schwarziana Moure. 1943a: 147. Type species: Melipona

1 for qiiadripunctata Lepeletier, 1836, by original designation. Serapista Cockerell, 904a:357. replacement Serapis Smith, [Trigona). Meliponini. 1854. Type species: Serapis denticulatus Smith, 1854, autobasic. Anthidiini. Schwarzula Moure, 1946a:439. Type species: Trigona tiniida Silvestri, 1902, by original designation. Meliponini. Sericogaste r "Westwood. 1835:71. Type species: Sericogaster fasciatus Westwood, 1835, monobasic. Isotypic with Scirtetica Holmberg, 1903:389. not Saussure, 1884. Type spe- Holohesma cies: Scirtetica antarciica Holmberg, 1903, monobasic. See by synonymy. Euryglossinae. Alloscirtetica. Holmbergiapis. and Neoscirtetica. Eucerini. Sericohalictus Pesenko, 1 986a: 1 37. Type species: Halictus subopacus Smith, 1853, by original designation. Scirteticops Moure and Michener, 1955b:261. Type species: [Lasioglossum]. Halictini. Tetralonia gayi Spinola, 1851. by original designation.

[Alloscirtetica]. Eucerini. Severanthidiuni Pasteels, 1 969a: 1 06. Type species: Anthidiuni severini Vachal, 1903, by original [Anthidium]. Scitandrena Warncke, 1968:51. Type species: Andrena scita designation. Anthidiini. Eversmann, 1852, by original designation. [Andrena]. Andreninae. Shornherria Ashmead. 1899a:7l. error for Schonnherria 1841. Scoliandrena Lanham, 1949:223. Type species: Andrena Lepeletier, Xylocopini. osmioides Cockerell, 1916, by original designation. Sibiricibombus Skorikov, 1938a: 145, unjustified emendation of [Andrena]. Andreninae. Sibiricobombus Vogt, 1911. Bombini. Genus-Group Names of Bees 55

Sihiricohoinhus Vogt, 1911 :60. Type species: Apis sibihca Fab- Sphecodiiiin Robertson. 1903b: 104. Type species: Sphecodiiim ricius. 1781. by designation of Sandhouse, 1943:599. cressonii Robertson, 1 903, by original designation. Halictini. Bombini. [Bombus]. Sphecodogastra Ashmead, I899a:92. Type species: Sphecodes Halictini. Signuitapis Maa, 1933:556. Type species: Apis ceninci Fabri- texatui Cresson. 1872, by original designation. cius. 1793. by original designation. [Apis|. Apini. Sphecodopsis Bischoff. 1923:593. Type species: Omacluhes

191 1 , Ammobatini. Siniaiuliviui Perez. 1890: 174. Type species: Andrena propinqiia capicola Strand, by original designation. = Schenck, 1851 Melitia dorsata Kirby. 1802. by designa- Sphecodosoma Crawford. 1907:182. Type species: tion otHedicke, 1933:218. [AndrenaJ. Andreninae. Sphecodosonia pratli Crawford, 1907, by original designation. Siiiuiiitliedon Zavortink, 1975:232. Type species: Simcmthedon Rophitinae. linsleyi Zavortink, 1975. by original designation. Eucerini. Sphecogaster Hirashima, 1978:72, not Lacordaire, 1869. Type Palaeorhiza Hirashima, 1978, Siiucolletes Warncke, 1978:348. Type species: Colletes similis species: paradisea by original See [Palaeorhiza]. Schenck, 1853, by original designation. [Colletes]. Colletinae. designation. Eusphecogastra. Hylaeinae. Ashmead, 1899a:89, for Siiuioceratiiui Daly and Moure. 1988, in Daly. 1988:42. Type Sphecophala unjustified replacement Sphegocepliala Saussure, 1 890. species: Sphegocephala species: Ceratina moereuhomi Vachal, 1903, by original des- Type philanthoides Saussure, 1 890, autobasic. Nomiinae. ignation. [Ctenoceratina]. Ceratinini. Saussure, 1890:74. Sphegocephala Siiupanurgus Warncke, 1972:67. Type species: Paniirgiis Sphegocephala Type species: philanthoides Saussure, 1890, monobasic. Nomiinae. pliyllopodus Warncke. 1972, by original designation. [Panurgus]. Panurglnae. Sphegodes Mavromoustakis, 1949:553, unjustified emendation of Sphecodes Latreille, 1804. Halictini. Sinonuicropis Michener. 1981:51. Type species: Macropis hedini Pasteels, 1969a:59. Anihidinni Alt"ken, 1936, by original designation. [Macropis J. Melittinae. Spinanihidielliini Type species: volkmanni Friese. 1909. by original designation. Smeathhalictus Warncke, 1975b:88. Type species: Melitta [Spinanthidium|. Anthidiini. snicatlinuinelUi Kirby. 1802, by original designation. 1951:977. [Halictus]. Halictini. Spinanthidiiini Mavromoustakis, Type species: Anthidiiim trachnsifonne Friese. 1913, by original designa- Smithia Vachal, 1897:63, not Milne-Edwards, 1851. Type spe- tion. [Dianthidium]. Anthidiini. cies: Scnipter carinata Smith, 1862, by designation of and Sharma. in 1993:104. Cockerell, 1910b:358. See Melittusmithia. Euryglossinae. Spinasternella Gupta 1993, Gupta, Type species: Spinasternella nienatiis Gupta and Sharma, Solamegilla Marikovskaya, 1980:650. Type species: Anthophora 1993, by original designation. Osmiini. prslii'wcdskyi Morawitz, 1880, by original designation. Moure, 1 95 1 c: 1 93, for Anthophorini. Spinolapis replacement Pasiphae Spinola, 1851. Type species: Pasiphae caerulescens Spinola, 1851, Solenopcdpa Perez, 1 897a:lxvii; also described as new by Perez, by original designation and autobasic. Colletinae. 1897b:260. Type species: Solenopalpa fi'itoni Perez, 1903 Spinoliella Ashmead, 1899a:84. Type species: Camptopoeiini (not Andrena fertoni Perez, 1895) - Andreiui solenopalpa noinioides - nomadoides Benoist, 1945, monobasic. Andreninae. [sic| Spinola Camptopoewn Spinola, 1851, by original designation. Panurginae. Sophrohoinlms Cockerell. 1908d:326: also described as new by Spinopasites Warncke, 1 983:28 1 . Type species: Pasites spinotiis Cockerell, 1909g:21. Type species: Sophrobombiis fatalis Warncke. 1983, original designation. [Pasites]. Cockerell, 1908, monobasic. Fossil. by Ammobatini. Soroeensihamlms Vogt, 191 1:63. Type species: Apis soroeensis Steganomus Ritsema, 1873:224. Type species: Steganomits Fabricius, 1776, monobasic. [BombusJ. Isotypic with Ritsema, 1 873. monobasic. Nomiinae. Kallobomhiis. Bombini. javatms Stegocephala Dalla Torre, 1896: 176, unjustified emendation of Spatidarici Mehely. 1935:69, 175, not Van Deventer, 1904. In- Sphegocephala Saussure, 1 890. Nomiinae. valid because no type species was designated. [Prosopis]. See Stelidiella Timberlake. 1941:133. lapsus for Stelidina Spatularicdla. Hylaeinae. Timberlake. 1941. [Stelis|. Anthidiini. SpatiiUirii'lla Popov, 1939:169, replacement for Spatidaria Stelidina Timberlake, 1 94 1 : 1 3 1 . Type species: Stelis hemirhoda Mehely, 1935. Type species: Hylaeus hyalimitus Smith, 1 842, Linsley. 1939, by original designation. [Stelis]. Anthidiini. by original designation. [Prosopis]. Hylaeinae. Stelidium Robertson, 1902b:323. Type species: Stelidiuni Spatiinoinia Pauly, 1980:121. Type species: Nomia filifcra tiypetiniini Robertson, 1902, monobasic. The subsequent des- Cockerell, 1932, by original designation. [Nomia[. Nomiinae. ignation, Robertson, 1903b: 104, of "S. cresio;!// Robertson" Cockerell, 1929a:217. " Sphaerhylaeits Type species: as the type species of "Stelidiuni was a lapsus: the intention des- Gnathoprosopis globulifera Cockerell, 1929, by original was to designate that species as the type of Sphecodium. ignation. [Gnathoprosopis]. Hylaeinae. Anthidiini.

Sphecodes Latreille. 1804:182. Type species: Noiuada gibba Stelidium Robertson, 1903b: 104, lapsus for Sphecodium: this is - Fabricius. 1804 Sphex gibba Linnaeus, 1758, monobasic. not Stelidium Robertson, 1902b. Type species: Sphecodium Isotypic with Dichroa. Halictini. cressonii Robertson, 1903, by original designation. Halictini. 56 SciENTinc Papers, Natural History Museum, The University of Kansas

Stelidomorpha Morawitz, 1875:131. Type species: Anthidiiim Stilpnosoma Smith, 1879:16. Type species: Stilpnosoma nasutiim Latreille, 1809, monobasic. Anthidiini. laevigatiitn Smith, 1879, monobasic. Euryglossinae.

Stelis Panzer, 1806:246. Type species; Apis ateniina Panzer, Strandiella Friese, 1912: 181. Type species: Strandiella longida 1798, not Christ, 1791 = Apis punctulatissima Kirby, 1802, Friese, 1912, by designation of Cockerell, 1916a:430. monobasic. Isotypic with Trachusa Jurine, Gywdroma Klug, Colletinae. and Ceraplastes. Anthidiini. Gyinnus. Siiandrena Wamcke. 1 968:52. Type species: Andrena siierinensis Stellenigris Meunier, 1888:152. Type species: Stellenigris Friese, 1884, by original designation. [Andrena]. Andreninae. vandeveldi Meunier, monobasic. Megachilini. Siihterraneihomhiis Skorikov, 1938a: 145, unjustified emenda- Stelodides Moure, 1953c: 123. Type species: Megachile euzona tion of Siihterraneobombiis Vogt. 1911. Bonibini. Perez, 1899, Megachilini. by original designation. Subterraneobombus Vogt, 1911:62. Type species: Apis Stenandrena Timberlake. 1949. in Lanham, 1949:2 1 3. Type spe- siibterrunea Linnaeus. 1758, by designation of Frison, cies: Pterandremi pallidifovea Viereck, 1904, by original des- 1927:68. [BombusJ. Bonibini. ignation. [Andrena]. Andreninae. Sudila Cameron, 1898:52. Type species: Siidila bidentata 1968a: 1059. Also described as new Stenanthidiellum Pasteels, Cameron, 1898, by designation of Sandhouse, 1943:602. Pasteels, 1969a:53. Anthidiiim sibiriciim by Type species: Isotypic with Ceylonicola by synonymy. Halictini. Eversmann, 1852, by original designation. [Bathanthidium]. Siilcobombiis Krtiger, 1917:65. Type species: Bonibiis confiisus Anthidiini. Schenck. 1859, designated by Sandhouse, 1943:502. This was 1947a: 16. Anthidiiim Stenanthidium Moure, Type species: a sectional name, but is treated as a subgeneric name in view 1938. Anthidiini. espinosai Ruiz, by original designation. of the Code, 3rd ed., art 10(e). [Bombus]. Isotypic with StenocoUetes Schrottky, 1909c:253. Type species: Stenocoiletes Confitsibombiis and Confiisobombiis. Bombini. 1909, monobasic. pictiis Schrottky, Panurginae? Sundatrigona Inoue and Sakagami, 1993:769, replacement for Stenoediscelis Toro and Moldenke, 1979:135. Type species: Trigonella Sakagami and Moure, 1975. Type species; Trigona Oediscelis inermis Friese, 1908, by original designation. moorei Schwarz, 1937, by original designation and autobasic. Xeromelissinae. [Trigona]. Meliponini.

Stenoheriades Tkalcu, 1 984a: 1 . Type species: Pseudoheriades Svastra Holmberg, 1884:127. Type species: Svastra bombilans hofferi Tkalcu, 1984, by original designation. [Pseudo- Holmberg, 1884, by designation of Sandhouse, 1943:602. heriades). Osmiini. Eucerini.

Stenohesma Michener, 1965a:99. Type species: Stenohesma Svastrides Michener, LaBerge, and Moure, 1955:220. Type spe- nomadiformis Michener, 1965, by original designation. cies: Tetralonia melaniira Spinola, 1851, by original desig- Euryglossinae. nation. Eucerini.

Stenomegachile Pasteels, 1965:507. Type species: Megachile Svastrina Moure and Michener, 1955b;276. Type species: chelostomoides Gribodo, 1894. by original designation. Svastrina anaroliata Moure and Michener, 1955 = Svastra

[Chalicodoma]. Megachilini. siifjapicalis Brethes, 1910, by original designation. Eucerini. Stenomelissa Hirashima and 1965, //; Hirashima, LaBerge. Svastropsis Moure and Michener, 1955b:291. Type species: 1965b:500. Andrena halictoides Smith, 1869, Type species: Tetralonia bipiinctuta Friese, 1908, by original designation. [Andrena]. Andreninae. by original designation. [Martinapis]. Eucerini.

Stenosmia Michener, 1 94 1 : 1 65. Osniia fluvicornis Type species: Symmorpha Klug, 1807, in lUiger, 1807:198, nomen nudum. Morawitz, 1878, by original designation. Osmiini. Melectini. Stenothisa Moure, 1954b:277. species: Doeringieila Type Symmorplia Klug, 1807b:227. Type species: Apis punctata Fab- Moure, 1954. by original designation. angiistata ricius, 1775 = Apis albifrons Forster, 1771, monobasic. [Doeringieila]. Epeolini. Isotypic with Melecta. Melectini. Stenotritus Smith. 1853:1 19. Type species: Stenotritus elegans Synalonia Robertson, 1905:365, unjustified emendation of Smith, 1853, monobasic. Stenotritidae. Synhalonia Patton, 1879. Eucerini. Steno.xylocopa Hurd and Moure, 1960:809. Type species: Synapis Cockerell, 1907a:229. Type species: Apis henshawi Xylocopa artifex Smith, 1874, by original designation. Cockerell, 1907, monobasic. [Apis]. Fossil. Apini. [Xylocopa]. Xylocopini. Synepeolus Cockerell. 1921b:6. Type species: Triepeolus Stictonomia Cameron, 1905:192. Type species: Stictonomia insolitus Cockerdl, 1921, monobasic. [Triepeolus]. Epeolini. punctata Cameron, 1905, monobasic. Nomiinae. Synhalonia Patton, 1879a:473. Type species: Melissodes Stilbeiicera Tkalcu. 1978:162. Type species: Eiicera clypeata tars is Cresson, 1878, by original designation. Eucerini. Erichson, 1835, by original designation. [Eucera]. Eucerini. fiilvi Mitchell, 1973:57. Coelioxys texana Stilhomalopsis Silveira, 1995:450. Type species: Exomalopsis Synocoelio.xys Type species: 1872, solani Cockerel!. 1896, by original designation. Cresson, by original designation. [Coelioxys]. [Exomalopsis]. Exomalopsini. Megachilini. Genus-Group Names of Bees 57

Syntrichalonici LaBerge, 1957:10. Type species: Melissodes Isotypic with Bremiis Jurine, Bremiis Panzer, Bombus. and e.xquisita Cresson. 1878, by original designation. Eucerini. Leucobombus. Bombini.

Systropha lUiger, 1806:145. Type species: Andrena spiralis Tetrachlora Schrottky, 1909b:481. Type species: Halictus mul- 1789 = Eitcera curvicornis Olivier. Scopoli. 1770, monoba- tiplex Vnchal. 1903, monobasic. [Augochlora]. Augochlorini. sic. Rophitinae. Tetraglossula Ogloblin, 1948:165. Type species: Tetraglossula Systrophidia Cockerell, 1936b:477. Type species: Systropha deltivaga Ogloblin, 1948, by original designation. CoIIetinae. ogilviei Cockerell, 1936, monobasic. [Systropha]. Tetragona Lepeletier and Serville, 1828:710. Type species: Rophitinae. Trigona elongata Lepeletier and Serville, 1828 = Centris 1956: 1 170. Melissodes Tachymelissodes LaBerge, Type species: clavipes Fabricius, 1804, by original designation. [Trigona], dagosa Cockerell, 1909, by original designation. Meliponini. [Melissodes]. Eucerini. Tetragonilla Moure, 1961:210. Type species: Trigona atripes Taeniandreiui Hedicke, 1933:219. Melitta ovatida Type species: Smith, 1857, by original designation. Meliponini. Kirby, 1802, by original designation. [Andrena] . Andreninae. Tetragonisca Moure, 1946a:438. Type species: Trigona Jaty Taiiguticobombus Pittioni, 1939:201. Bombiis = Type species: Smith, 1863 Trigona angustula Latreille, 181 1, by original Morawitz, 1886, tanguticus by original designation. designation. Meliponini. [Bombus]. Bombini. Tetragomda Moure, 1 96 1 :206. Type species: Trigona iridipennis Tapinorhina Michener and Moure, 1957:421. species: Type Smith, 1854, by original designation. Meliponini. Exomalopsis caendea Friese, 1906, by original designation. Tetralonia Spinola, 1838:538, replacement for Macrocera [Tapinotaspis]. Tapinotaspidini. Latreille, 1810. Type species: Eucera antennata Fabricius, Tapinorrhina Moure, 1994a:274, emendation of unjustified 1793 = Apis malvae Rossi, 1790, autobasic. See Michener, Michener and Moure, 1957. Tapinorhina Tapinotaspidini. 1997. Eucerini. Tapinotapsis [sic| Holmberg, 1887b:225, nomen nudum. Tetraloniella Ashmead, 1 899a:6 1 . Type species: Macrocera graja Tapinotaspidini. Eversmann, 1 852, by original designation. Eucerini. Tapinotaspis Holmberg, 1903:413. Type species: Tapinotaspis Tetralonioidella Strand, 1914 (Apr.-May):140. Type species: chacabiicensis Holmberg, 1903 = Exomalopsis chalybaea Tetralonia { ?) hoozana Strand, 1914, monobasic. Isotypic with Friese, 1899, by designation of Sandhouse, 1943:603. Protomelissa by synonymy. Melectini. Tapinotaspidini. Tetranthidium Moure, 1947a: 15. Type species: Anthidium latum Tapinotaspoides Moure, 1944b: 10. Type species: Tetrapedia Schrottky, 1902, by original designation. Anthidiini. serraticornis Friese, 1899, by original designation. Dalla emendation of Tapinotaspidini. Tetrapaedia Torre, 1896:299, unjustified Tetrapedia Klug, 1810. Tetrapediini. Tarsalia Morawitz, 1895:9. Type species: Tarsalia hirtipes 1810:33. Morawitz, 1895, monobasic. Ancylini. Tetrapedia Klug, Type species: Tetrapedia diversipes Klug, 1810, monobasic. Tetrapediini. Tarsandrena Osychnyuk, 1984a:24. Type species: Andrena emendation of tarsata Nylander, 1848, by original designation. [Andrena]. Tetrapedium Berthold, 1827:468, unjustified Andreninae. Tetrapedia Klug, 1810. Tetrapediini. Teleuteinnesta Holmberg, 1887b:225, nomen nudum. Tetraperdita Cockerell and Porter, 1899:415. Type species: Emphorini. Perdita sexmaculatu Cockerell, 1895, monobasic. [Perditaj. Panurginae. TeleutemnestaHoXmhcrg, 1903:400. Type species: Teleutemnesta fructifera Holmberg, 1903, by designation of Cockerell, Tetrigona Moure, 1961:215. Type species: Trigona apicalis 19 18b:36. Emphorini. Smith, 1857, by original designation. Meliponini. Temnosoma Smith, 1853:38. Type species: Temnosoma Thalestria Smith, 1854:283. Type species: Thalestria smaragdina = metallicum Smith, 1853, monobasic. Augochlorini. Smith, 1854 Euglossa spinosa Fabricius, 1804, monobasic. Epeolini. Temnosomulu Ogloblin, 1953:2. Type species: Temnosoma platensis Ogloblin, 1953 = Micraugochlora sphaerocephala Thaumatosoma Smith, 1865:394. Type species: Thaumatosoma duboulaii Schrottky, 1909, by original designation. [Temnosoma]. Smith. 1865, monobasic. Megachilini. with Isotypic Micraugochlora by synonymy. Augochlorini. Thectochlora Moure, 1940:51. Type species: Halictus alaris Teratognatha Ogloblin, 1956:154. Type species: Teratognatha Vachal, 1904, by original designation. Augochlorini. modesta 1956, Ogloblin, by original designation. Thoracobombus Dalla Torre. 1880:40. Type species: Apis Exomalopsini. sylvarum Linnaeus, 1761, by designation of Sandhouse, Tergosmia Wamcke, 1988a:390. Type species: Osmia tergestensis 1943:604. [Bombus]. Isotypic with Adventoribombus. Ducke, 1897, by original designation. [Osmia]. Osmiini. Bombini.

Terrestribombus Vogt, 1911:55. Type species: Apis terrestris Thrausmus Buysson, 1900:177. Type species: Thrausmus Linnaeus, 1758,by designation of Prison, 1927:67. [Bombus], grandidieri Buysson, 1900, monobasic. Halictini. 58 Scientific Papers, Natural History Museum, The University of Kansas

Thrinchostoma Saussme, 1890:52. Type species: Thrincliosroiiia Trachusaiunne, 1801:164, not Panzer, 1804. Type species: /4/?w renitantely Saussure, 1890, monobasic. Halictini. aterrima Panzer, 1 798, by designation of Morice and Durrant, 1915:426. Commission 135 (1939), Thrincohcilictiis Bliithgen. 1955:20. Type species: Halictii.s Suppressed by Opinion Direction 4. with Stelis. prognathus Perez. 1912, by original designation. [Halictus]. Isotypic Gyrodroma Klug, Gymniis. Halictini. and Ceraplastes. Anthidiini. Tracliusa Panzer. 14-15. Trachiisa Thrincostoimi DalUTorre. 1896:641, also Bluthgen, 1926, 1928b, 1804b:expl. pi. Type species: serratulae Panzer, Panzer, 1798, des- 1930, unjustified emendation of Thrinchostoma Saussure, \%QA=Apis byssimi by 1890. Halictini. ignation of Sandhouse, 1943:605. Isotypic with Megachileoides: with 1884:133. Tetnilonia isotypic Diphysis by synonymy. Thygater Holmberg, Type species: Anthidiini. terminata Smith. 1854 - Macrocera analis Lepeletier, 1841, monobasic. Eucerini. Trachiisoides Michener and Griswold, 1994a:324. Type species: Trachusoides simplex Michener and Griswold, 1 994. by origi- 77;yga?/'!a Cockerell, 191 lf:237. Type species: Thygatinafiimida nal designation. Anthidiini. Cockerell. 1911, monobasic. Eucerini. Trachiisomimus Popov. 1964:406. Type species: Trachiisa perdita Thyreothremma Holmberg, 1903:391. Type species: Cockerell, 1904, by original designation. Anthidiini. Thyreothrenima rlwpalocera Holmberg, 1903 =Melissoptila Hirashima, 1980:100. Palaeorhiza tandilensis Holmberg. 1884. by designation of Sandhouse. Trachyrhiza Type species: Hirashima. 1980, 1943:604. Isotypic with Thyreotremuta: isotypic with rugosa by original designation. Melissoptila by synonymy. Eucerini. [Palaeorhiza]. Hylaeinae. Trianthidiellum I969a:58. Thyreotremata Holmberg, 1887b:225, nomen nudum. Eucerini. Pasteels, Type species: Hypanthidium sheppardi Mavromoustakis. 1937, by original designation. Thyreotremuta Sandhouse, 1943:604. Type species: — Anthidiini. Thyreothremma rhopahx'era Holmberg. 1 903 Melissoptila Trianthidiiim Mavromoustakis, 1958:435. tandilensis Holmberg. 1884, by designation of Moure and Type species: Anthidium Giraud. 1 863. Michener. 1955b:305. Published as a junior synonym and catiirigense by original designation. Anthidiini. therefore not available according to the Code. 3rd ed.. Art. [Rhodanthidium].

1 1(e). See Moure and Michener, 1955b:305 footnote. Isotypic Trichanthidiodes Michener and Griswold. 1 994a:3 1 9. Type spe- with Thyreothremma: isotypic with Melissoptila by syn- cies: Pachyanthidiiim semiliiteiim Pasteels. 1984, by original onymy. Eucerini. designation. [Pachyanthidium]. Anthidiini.

Thy reus Panzer, 1806:263. Type species: Nomada scutellaris Trichanthidiiim Cockerell, 1930b:52. Type species; Fabricius, 1781. monobasic. Isotypic with Crocissa. Pachyanthidium occipitale Cockerell, 1930 = Anthidium Melectini. benguelense Vachal, 1903, monobasic. [Pachyanthidium]. Anthidiini. Thysandrena Lanham, 1949:213. Type species: Andrena Can- dida Smith. 1879. by original designation. [Andrena]. Trichanthidium Moure. I947a:20. not Cockerell, 1930. Type Andreninae. species: Anthidium subpetiolatum Schrottky, 1910, by original See Anthidiamim. Anthidiini. Titusella Cockerell, I906e:445. Type species: Titusellapronitens designation. Cockerell. 1906, monobasic. Osmiini. Trichchostoma Ashmead. I899a:91. eiTor for Thrinchostoma Saussure, 1890. Halictini. Tmetocoelia Moure. 1943b:481. Type species: Megalopta sulciventris Friese, 1926, by original designation. Trichinosmia Sinha, 1958:244. Type species: Osmia latisulcata Augoclilorini. Michener, 1936, by original designation. [Osmia]. Osmiini.

Torocolletes Michener. 1989:65 1 . Type species: Lonchopriafazii Trichoceutris Snelling. 1956:4. Type species: Centris rhodoleuca Herbst, 1923, by original designation. [Leioproctusj. Cockerell. 1923 = Centris caesalpiniae var. rhodopus Colletinae. Cockerell, 1 897, by original designation. Centridini.

Torridapis Pasteels. 1977:195. Type species: Coelioxys torrida Trichocerapis Cockerell, 1904b:292. Type species: Tetralonia Smith. 1854, by original designation. [Liothyrapis], mirabilis Smith, 1865. by original designation. Eucerini. Megachilini. Trichocolletes Cockerell. 1912:176. Type species: Townsendiella Crawford, 1916:136, 138. Type species: Lamprocolletes venustus Smith. 1862, by original designa- Townsendiella pulchra Crawford, 1916, monobasic. tion. Colletinae. Townsendiellini. Tricholiplninthus Ruz and Toro, 1983:267. Type species:

Tractumdrenu Robertson, 1902a: 1 87, 1 89. Type species: Andrena Liphanthus leucostomus Ruz and Toro, 1 983. by original des- riigosa Robertson. 1891. by original designation. Andreninae. ignation. [Liphanthus]. Panurginae.

Trachina Klug, 1807b:226. Type species: Centris longimana Trichonomada Michener, 1996:90. Type species: Trichonomadu Fabricius. 1804, by designation of Cockerell. I906d:105. roigella Michener, 1996. by original designation. Centridini. Brachynomadini. Genus-Group Names of Bees 59

Trichopasites Linsley, 1942:127. Type species; Neopasites Trinomia Pauly, 1980: 122. Type species: Nomia tridenuita Smith, Noniiinae. insoletiis Linsley, 1942, by original designation. [Neopasites]. 1875, by original designation. [Nomia]. Ammobatoidini. Triopasites Linsley, 1939b:8. Type species: Triopasites Trichoihopalomelissa Wu, 1985:58. Type species: timherlakei Linsley, 1939, by original designation. Rhopalomelissa hainanensis Wu, 1985, by original designa- Brachynomadini. Noniiinae. tion. [Rhopalomelissa]. Troandrena Warncke, 1974:108, nomen nudum. [Andrena]. Trichosioiuu DallaToiTe, 1896:381, also Friese, 1909a: 150, un- Andreninae. justified emendation of Thrinchostoma Saussure, 1890. Troandrena Warncke. 1 975a:8 1 . Type species: Andrena troodica Halictini. Warncke. 1975. by original designation. [Andrena]. Thchota Mehely, 1935:63, 169. Invalid because no type species Andreninae. was designated. [Prosopis]. Sandhouse's (1943:606) view that Tropandrena Viereck. 1924:21. Type species: Andrena fragilis Popov. 1939:169, designated a type species and thus vali- Smith, 1853, by original designation. [Andrena]. Andreninae. dated the name is incorrect. Hylaeinae. Trophocleptria Holmberg. 1886b:233. 275. Type species: Trichothurgus Moure, 1949b:270. Type species: Megachile dii- Trophocleptria variolosa Holmberg. 1886. monobasic. hia Sichel. by original designation. Trichothiirgtis and Epeolini. Lithitrgomma Moure were published simultaneously. 1991:425. Michener (1983). as first reviser, selected Thchotliiirgus as Tropicocoelioxys Gupta. Type species: Coelioxys 1991, the generic name when they are considered synonymous. genoconcavitiis Gupta. by original designation. Lithurgini. [Coelioxys]. Megachilini. Michener and Moure, 1957:411. Triclwtosmia Tkalcu, 1980: 16. Type species: Wainia consinillis Tropidopedia Type species: seahrai Michener and Moure, 1957, Tkalcu, 1980, by original designation. (WainiaJ. Osmiini. Paratetrapedia by original designation. [Paratetrapedia]. Tapinotaspidini. Trichotrigona Camargo and Moure, 1983:421. Type species: Wu, 1985:58. Tricliotrigona extranea Camargo and Moure, 1983, by origi- Tropirhopaloiuelissa Type species: Rhopalomelissa nal designation. Meliponini. nigra Wu, 1985, by original designation. [Rhopalomelissa]. Nomiinae. Trichiirochile Mitchell, 1980:63. Type species: Megachile Andrena thygaterella Schrottky, 1913. by original designation. Truncandrena Warncke. 1968:46. Type species: [Cressoniella]. Megachilini. triincatilahris Morawitz, 1878, by original designation. [Andrena]. Andreninae. Tricomibombus Skorikov, 1922b: 151. Type species: Bombiis triconii.s Radoszkowsky, 1888, monobasic. [Agrobombus]. Trypetes Schenck, 1861:32, 89. [For the date, see Michener, Bombini. 1986a.] Type species: Apis tritncorum Linnaeus, 1758, monobasic. with Heriades. Osmiini. Triclentosiuia Schmiedeknecht, 1885:21. Type species: Osinia Isotypic tridentala Dufour and Perris, 1840, by designation of Tuberanthidium Pasteels, 1969a:87. Type species: Anthidium Michener, 1941:159. [Osmia]. Osmiini. tiiherciiliferuni Brauns, 1905, by original designation. Anthidiini. Triepeohis Robertson, 1901 :23 1 . Type species: Epeohis concaviis Cresson, 1878, by original designation. Epeolini. Titmidihesma Exley, 1996:253. Type species: Tumidihesma tridentata 1996, Triepichahs Moure, 1945b:304. Type species: Epicharis analis Exley, by original designation. Lepeletier, 1841, by original designation. Centridini. Euryglossinae. Titrkanthidiwn Pasteels. 1969a: 103. Anthidium Trigona Jurine, 1807:245. Type species: Apis amaltlwu Olivier, Type species: uniciim 1789. by designation of Latreille, 1810:439. Meliponini. Morawitz. 1875. by original designation. [Anthidiumj. Anthidiini. Trigonella Sakagami and Moure, 1975, /;; Sakagami, 1975:57, Titrnerella Cockerell. 1910c:262. Tnrnerella not da Costa, 1778. Type species: Trigona moorei Schwarz, Type species: 1937, monobasic. See Sundatrigona. Meliponini. gilbe rti CocktTsW. 1910. monobasic. Euryglossinae. 1964:312. Cilissa Trigonisca Moure, 1950a:249. Type species: Trigona diickei Tylandrena LaBerge, Type species: Friese, 1900, by original designation. [Hypotrigona]. erythrogaster Ashmead, 1890, by original designation.

. Andreninae. Meliponini. [Andrena I 1994a:277. Trigonopedia Moure, 1941b:518. Type species: Trigonopedia Tylanthidiiim Urban. Type species: Tylanthidium Anthidiini. oligotriclia Moure. 1941. by original designation. titberigasterVixhdn. 1994. by original designation.

Tapinotaspidini. TylomegachUe Moure. 1 953c: 1 20. Type species: Megachile orha Trilia Vachal. 1899:534. Type species: Diifoiirea miioti Vachal, Schrottky. 1913, by original designation. [Megachile]. 1899, monobasic. [Dufourea]. Rophitini. Megachilini. Trinchosioma Sandhouse, 1943:606. inconectly attributed to Tytthaliclus Pesenko, 1984:348. Type species: Halictiis Sladen. 1915:214. unjustified emendation of Thrinchostoma niaciilatiis Smith, 1848, by original designation. [Halictus].

Saussure, 1 890. Halictini. Halictinae. 60 Scientific Papers, Natural History Museum, The University of Kansas

Xanthidium Robertson, 1903d: 174, 175, 177, not Ehrenberg, Ulandrena Wamcke, 1968:43. Type species: Andrena schulzi Nomada luteola Olivier. 181 1, Strand, 1921, by original designation. [Andrena]. 1833. Type species: by origi- but since it is a Andreninae. nal designation. This name is preoccupied Nomadini. synonym of Nomada. it has not been replaced. Ulanthidiiim Michener. 1948a: 13. Type species: Ulanthidium 1952:39. Ceratina cladura mitchelli Michener, 1948, by original designation. Anthidiini. Xanthoceratina Vecht. Type species: Cockerell, 1919, by original designation. [Ceratina]. Uncobombus Kruger, 1917:65. Type species: Apis hypnorum Ceratinini. Linnaeus. 1758, by designation of Williams, 1991:69. A sectional name, treated as a genus-group name in view of the Xantlwcotelles Toro and Cabezas, 1978:131. Type species: Code, 3rd ed.. Art. 10(e). Kruger attributed the name to Vogt Xanthocotelles adesmiae Toro and Cabezas, 1978, by origi- but Kruger evidently wrote the text of the paper. [Bombus]. nal designation. CoUetinae. with Anodontobombus. and Isotypic Pyrohombus, Xanthopedia Michener and Moure, 1957:411. Type species: Bombini. = Hypnorobombus. Paratetrapedia tricolor Michener and Moure, 1957 UrocoUetes Michener, 1965a:58. Type species: Leioproctiis Tetrapedia iheringii Friese, 1899. by original designation. rhodurus Michener, 1965, by original designation. [Paratetrapedia]. Tapinotaspidini. CoUetinae. [Leioproctus]. Xanthosants Robertson. 1903a: 168, 169. 172. Type species: Urohalictus Urohalictus Michener. 1980:16. Type species: Megacliile latimamts Say. 1823, by original designation. 1980. Halictini. Ueftincki Michener, by original designation. Isotypic with DelomegachUe by synonymy. Megachilini. Uromonia Michener. 1981:23. Type species: Uromonia stagei Xanthosmia Robertson, 1903a: 166. 171. Type species: Osmia Michener, 1981, by original designation. Meganomiinae. cordata Robertson, 1902, by original designation. Osmiini. 1902:44. Verbenapis Cockerell and Atkins. Type species: Xenanthidium Pasteels. 1984:33. Type species: Xenanthidiiuu and 1899. monobasic. Calliopsis verbenae Cockerell Porter. biserratum Pasteels (described as biserrata), 1984, by origi- [Calliopsis]. Panurginae. nal designation. Antiiidiini. Halictus Vestitohalictus Bluthgen. 1961:287. Type species: Xenochalicodoma Tkalcu. 1971:34, replacement for vestitus 1841, [Halictus]. Lepeletier, by original designation. Parachalicodoma Tkalcu. 1969. Type species: Chalicodoma Halictini. nifitarsis Lepeletier. autobasic. [Chalicodoma]. Megachilini. 1907:298. Viereckella obscura Viereckella Swenk. Type species: Xenochilicola Toro and Moldenke. 1979:145. Type species: 1907 = Nomada Cresson. 1878. by original Swenk, pilosula Xenochilicola mamigna Toro and Moldenke, 1979. by origi- Osirini. designation. nal designation. Xeromelissinae. Vihdosmia Wamcke, 1988b:5. Type species: Osmia saxicola Xenoglossa Smith, 1854:315. Type species: Xenoglossa fulva [Osmia]. Osmiini. Ducke, 1899, by original designation. Smith, 1854, monobasic. Eucerini. Voliicellobombiis Skorikov. 1922b:123. 149. Type species: Xenoglossodes Ashmead. 1899a:63. Type species: Melissodes Bombus vohtcelloides Gribodo. 1891, monobasic. Bombini. albata Cresson, 1872, by original designation. Eucerini. Moure, 1950c:389. Type species: Centris ciner- Wagenknechtia Xenoliesma Michener, 1965a:96. Type species: Euryglossa aria Smith. 1854. by original designation. [Centris]. flavicauda Michener, 1965, by original designation. Centridini. [Euryglossa). Euryglossinae. Wainia Tkalcu, 1980:1. Type species: Wainia lonavlae Tkalcu, Xenohylaeus Michener, 1965a: 136. Type species: Hylaeus rieki 1980, by original designation. Osmiini. Michener, 1965, by original designation. [Hylaeus]. Warnckeia and Scaramozzino. 1990:6, replacement for Pagliano Hylaeinae. Melanthidium Pasteels. 1969. Type species: Anthidium Ruz and Toro, 1983:260. Type species: concolor Friese. 1913. autobasic. Anthidiini. Xenoliphantlms Psaenythia parvula Friese, 1916, by original designation. Willinkapis Moure. 1953a:66. Type species: Ptiloglossa [Liphanthus], Panurginae. chalybaea Friese. 1906. by original designation. Rebmann, 1970:44. Type species: Megacliile Caupolicanini. Xenomegachile albocincta Radoszkowski, 1874, by original designation. Xantlmndrena Lanham. 1949:218. Type species: Andrena [Megachile]. Megachilini. auricoma Smith. 1879. by original designation. [Andrena]. Andreninae. Xenopanurgus Michener, 1952:24. Type species: Xenopanurgus readioi Michener, 1952. by original designation. Panurginae. Xanthemisia Moure, 1945c:401. Type species: Centris bicolor Hurd and 1970:29. Lepeletier, 1841, by original designation. Centridini. Xenopeponapis Linsley. Type species: Melissodes crassidentata Cockerell, 1949, by original desig- Xanthepicharis Moure, 1945b:297. Type species: Epicharis bi- nation. Eucerini. color Smith. 1854, by original designation. Centridini. [Peponapis]. Michener. 1965a: 146. Palaeorhiza Xanthesma Michener, 1965a:97. Type species: Euryglossa Xenorhiza Type species: hamada Cheesman, 1948. by original designation. fiircifera Cockerell, 1913. by original designation. Euryglossinae. [Palaeorhiza]. Hylaeinae. Genus-Group Names of Bees 61

Hurd and Moure, 1 963 : 1 60. Xenoxylncopa Hurd and Moure, 1963:243. Type species: Xvlocopina Type species: Xylocopa Hurd and Moure, 1963, Mesotrichia chiyakensis Cockerell, 1908, by original desig- mficollis by original designation. nation. [Xylocopa]. Xylocopini. [Xylocopa]. Xylocopini. Hurd and Moure, 1963; 105. Type species: Xylocopa Xeralictoides Stage. 1981, in Michener. 1981:99. Type species: Xylocopoda elegans Hurd and Moure. 1963, by original designation. Hesperapis laticeps Crawford, 1 9 1 7, by original designation. Xylocopini. Dasypodinae. [Xylocopa]. Xylocopoides Michener, 1 954: 1 55. Type species: Apis virginica Xeralictits Cockerell. 1927a:41. Type species: Xeralictus Linnaeus, 1771, by original designation. [Xylocopa], timbciUikei CockereU. 1927. monobasic. Rophitinae. Xylocopini. Xerammohates Popov, 1951:904. Type species: Ammobates Xylocopsis Hurd and Moure. 1963: 124. Type species: Xylocopa oxiainis Popov, 1951. by designation of the Commission, funesta Maidl. 1912, by original designation. [Xylocopa]. Opinion 1853 (1996). Ammobates biastoides Friese. 1895, Xylocopini. designated as the type species by Popov, 1951, w^as Hurd and Moure, 1963: 109. Type species: Xylocopa misidentified (Baker, 1995b). [Ammobates]. Ammobatini. Xylocospila bambusae Schrottky. 1902. by original designation. Xeroceniris 1974:3. Centris californica Snelling. Type species: [Xylocopa]. Xylocopini. Timberlake, 1940, by original designation. [Centris]. Xylomelissa Hurd and Moure. 1 963:2 1 9. Type species: Xylocopa Centridini. carinata Smith, 1874 = Xylocopa hottentotta Smith. 1854, 1973:44. edita Xemcoelioxys Mitchell, Type species: Coelioxys by original designation. [Xylocopa]. Xylocopini. Cresson, 1872. by original designation. [Coelioxys]. Zacesta Ashmead, 1899a:73. Type species: Zacesta rufipes Megachilini. Ashmead. 1899. monobasic. Dasypodinae. Xerohehades Griswold, 1986:165. species: Xeroheriades Type Zacosmia Ashmead, 1 898:282. Type species: Melecta macidata micheneri Osmiini. Griswold, 1986, by original designation. Cresson, 1879, by original designation. Isotypic with Xeromacrotem Timberlake. 1954:412. Type species: Macwtera Micromelecta. Melectini. cephalotes Cresson. 1878, by original designation. [Perdita]. Zadontomerus A?\\mtdd. 1899a:69. Type species: Ceratina Panurginae. tejonensis Cresson, 1864, by original designation. Ceratinini. Xeromegachile Mitchell, 1934:302. 309. Type species: Megachile Zahgus Cockerell. 1929b:321. Type species: Zaiygus comutus integra Cresson, by original designation. [Megachilej. Cockerell, 1929, monobasic. Euryglossinae. Megachilini. Zaodontomerus Cockerell and Porter, 1899:406, unjustified of Zrt(yo;!ro/Her[« Ashmead, 1899. Ceratinini. Xeromelecta Linsley, 1939a:450. Type species: Bomhomelecta emendation laireac Cockerell, 1900, by original designation. [Melecta]. Zaperdita Robertson, 1918a:91. Type species: Perdita maiira Melectini. Cockerell. 1901 = Perdita lialictoides Smith, 1853. by original with Perdita Xeromelissa Cockerell. 1926a:221. Type species: Xerowelissa designation. Isotypic by synonymy. wibnattae Cockerell, 1926, monobasic. Xeromelissinae. Panurginae. Zarhiopalea Hirashima, 1982a:57. Type species: Palaeorhiza Xeropasites Linsley, 1942:130. Type species: Townsendiella paradoxa Hirashima. 1975, by original designation. rufiventris Linsley, 1942, by original designation. [Palaeorhiza]. Hylaeinae. [Townsendiella]. Townsendiellini. Zebramegilla Brooks, 1988:502. Type species: Anthophora Xeropeponapis Hurd and Linsley. 1970:28. Type species: albigena Lepeletier. 1 84 1 , by original designation. [Amegilla]. Peponapis timberlakei Hurd and Linsley, 1964, by original Anthophorini. designation. [Peponapis]. Eucerini. Zikauapis Moure, 1945a: 147. Type species: Ptiloglossa zikani Cockerell, 1923a: 1. Type species: Xewphusma Xemphasmu Friese, 1925, by original designation. Caupolicanini. bequae rti CocktrtW. 1923 = Perdita bequaertiana Cockerell, Zikaniella Moure, 1941a:57. Type species: Zikaniella crassiceps 1951, by original designation. Panurginae. Moure. 1941, by original designation. Halictini. Xewsmia Michener. 1943:81. Type species: Osmiu xerophila Zonalictits Michener. 1978a:513. Type species: Halictits Cockerell, 1935. by original designation. [Anthocopa]. Halictini. albofasciatus Smith. 1 879, by original designation. Osmiini. Zonalonia Moure and Michener. 1955b:250, 282. Type species: 1802a:432. Commission Xilocopa Latreille, Suppressed by Opin- Eiicera cearensis Ducke. 1910. by original designation. ion 743 See (1965). Xylocopa. Xylocopini. [Dasyhalonia]. Eucerini. 1971b:504. Andrena Xiphaiidrena LaBerge, Type species: Zonamegilla Popov, 1950b:260. Type species: Apis zonata meiidica Mitchell, 1960, by original designation. [Andrena]. Linnaeus, 1758, by original designation. [Amegilla]. Andreninae. Anthophorini. Xylocopa Latreille, 1802b:379. Type species: Apis violacea Zonandrena Hedicke. 1933:220. Type species: Andrena flavipes Linnaeus, 1758, by designation of Westwood, 1840a:86. Panzer, 1799, by original designation. [Andrena]. Xylocopini. Andreninae. 62 ScffiNTiFic Papers, Natural History Museum, The University of Kansas

with Zonohalonia Moure and Michener, 1955b;255, lapsus for by Michener and Moure. 1 956:284. and is in accordance Eucerini. Zonalonia Moure and Michener. 1955. [Dasyhalonia]. the the Code, 3rd ed.. Art. 32(c)(ii). [Dasyhalonia]. Eucerini. Zonomegachite Mitchell. 1980:72. Type species: Megachile Dalla Torre. 1896. Zonohirsuta Ma, 1938:300. Type species: Xylocopa collaris markuuuie by original designation. = Lepeletier. 1841 (not A/7/.s(:Y)//fln5 Olivier. \1%9) Xylocopa [ChrysosarusJ. Megacliilini.

dejeanii Lepeletier, 1841 . by original designation. [Xylocopa]. Zosteranthidium Michener and Griswold, 1994a:3 10. Type spe- Xylocopini. cies: Nigninthidium tergofasciatwn Pasteels. 1984. by original Antliidiini. Zonolonia Moure and Michener. 1 955b:280, lapsus for Zonalonia designation. [Afranthidium]. Moure and Michener, 1955; the latter spelling was selected

SUPPLEIVIENT TO THE LIST OF FAMILY-GROUP NAMES

of 81 .2. and Michener, 1993:159. The authorsbiip and dating of family-group names Tapinotaspini Roig-Alsina The correct bees were dealt with by Michener (1986a). These names, Tnpinotnspis: stem: Tapiiiotaspid-. spell- are is This tribal name was inde- being coordinate at all levels (subtribe to superfamily), ing Tapinotaspidini. 1994b:306. attributed to the same author and date regardless of the pendently proposed by Moure, of the word 3rd Art. 36a; rank and termination [Code, ed.. 81 .3. Paratetrapediini Moure, 1994b:306. Paratetmpedia; also see Art. ll(f)(ii)]. Thus the Megachilini, Megachilinae, stem: Pamtetrapedi-. and all have the same author and date. Megachilidae 81.4. Teratognathini Silveira, 1995:449. Teratognntha; errors and omissions have been dis- Since 1986, a few stem: Temtogunth-. covered. Moreover, a few new family-group names have 85.1. Habropodini Marikovskaya, 1976:688. Habwpodn; been Below is a list of the additions and correc- pubUshed. stem: Habropod-. tions, in the of the earlier work, i.e., each entry con- style stem: and stem. 91. Melitomatae Vachal, 1909b:7. Melitomn; sisting of the name, author, reference, type genus, Melitom-. (See below.) The items in the list are numbered in accordance with Item 101.1. Brachynomadini Roig-Alsina and Michener, the system in the earlier list (Michener, 1986a). Thus, 1993:157. Brachynomada; stem: Brachynomad-. 30.1 could reasonably go after Item 30 in the earlier list, 1993:157. and such items are additions to the 1986 list. Items 3, 18, 104.1. Hexepeolini Roig-Alsina and Michener, 79, 91, 112, and 123 are corrections rather than additional Hexepeoliis; stem: Hexepeol-. of older family-group names. They result from discovery 112. Neolarrini Fox, 1894:302, 304. Ncolarra; stem: usages than those cited in 1986. Neolarr-. 119.9. Eucondylopsinae Friese, 1909a:99, 439. 1813:32. stem: 3. Prosopiariae Fallen, Prosopis; Eiicondylops; stem: Eucoudylops-. See 3. Prosopmi-.\8. 120.1. Manueliini Sakagami and Michener, 1987:439. 30.1. Perditinae Robertson, 1922:160. Perdita; stem: Mamidia; stem: Manueli-. Perdit-. 123. Exaeretinae Cockerell, 1908a:41. Exaerete; stem: stem: 30.2. Calliopsinae Robertson, 1922:160. CaUiopsis; Exaeret-. Calliops-. 125.1. Eufrieseini Moure, 1944b:12. Eufriesea; stem: 1922:160. Antlie- 30.3. Anthemurginae Robertson, Eupiese-. murgus; stem: Anthemurg-. 30.4. Ruz, 1991:215. Protoiueliturga; Protomeliturgini The name for the Emphorini has been a sub- stem: proper Pivtoincliturg-. Robertson (1904) ject of discussion. The name proposed by 1909b:482. 45.1 . Oxystoglossini Schrottky Oxystoglossa; based on Emphor is senior to the name based on Enteclmia stem: As out to me based on Oxystogloss-. pointed by proposed by Cockerell (1906d) and the name Michael S. this name has over Engel, priority Melitomn proposed by Vachal (1909b) (91 above). The name It not been used since it was Augochlorini. has Entechniini has not been used in recent decades, presum- in 1909. We wiU that it of casually published request ably because Eniechnia is a junior subjective synonym be the Commission in the interest of rejected by MeUtoma, and before 1961 the family-group name based of nomenclature. stability on EiitLxiuiin would have been rejected for that reason (Ar- stem: 79. Exomalopses Vachal, 1909b:7. Exomalopsis; ticle 5, International Rules of Zoological Nomenclature, 39:75- Exomalops-. Proceedings of the Biological Society of Washington The current confusion concerns the names 81.1. Ancyloscelina Roig-Alsina and Michener, 104, 1926). and MeUtoma and resulted from the 1993:159. Aiici/loscelis; stem: Aiici/loscelid-. based on Emphor Genus-Group Names of Bees 63 fact that Emplior is a junior subjective synonym of frequency; an incomplete list prepared by A. Roig-Alsina Ptilothrix. Moure (1944c) proposed Melitomini without in 1996 shows 13 usages of Emphorini and 9 usages of 1944. most in the explanation, but Moure (1947b:240) stated that the re- Melitomini since However, usages present placement of Emphorini by Melitomini was necessary decade (1990's) employed the name Emphorini. Thus, cur- because Emplwr had become a synonym of Ptilothrix. rent usage seems to favor Emphorini. The Rules at that time (reference above) re- required It is preferable to have a family-group name based on of a name based on a placement family-group genus an accepted genus rather than on a junior synonym, an name that was a as shown junior synonym. However, argument that favors Melitomini. For this reason and be- the name Melitomini dates from Vachal (1909b), above, cause a replacement for the name Emphorini was in fact who listed as an included Emphor genus. appropriate in 1944 when Melitomini was proposed by The Code, 3rd ed., Art. 40(b), states that if a family- Moure, the matter remains contentious. A request for a de- group name was replaced before 1961 because the genus cision submitted to the Commission elicited the response on which it was based is a junior synonym, and if the re- that Emphorini is the appropriate name and that there is placement has won general acceptance, it is to be main- no need to go to the time and expense of publication in the tained. The name based on Emphor was replaced by that Bulletin of Zoological Nomenclature and evaluation and based on Melitoma by Moure in 1944. Since that date, the decision by the Commission. two family-group names have been used with almost equal

LITERATURE CITED

viii + H. 1904a. bees. of the En- Agassiz, L. 1846. Ncmendatoris Zoologki Index Univcrsniis. 393 pp. Ashmead, W. Remarks on honev Proceedings Soloduri: Jent and Gassmann. tomological Society of Washington 6:120-122. list of the Is- Alexander, B. A., and C. D. Michener. 1995. Phylogenetic studies of the Ashmead, W. H. 1904b. A of the Hymenoptera Philippine with of new of the New York families of short-tongued bees. University of Kansas Science Bulle- lands, descriptions species. Journal tin 55:377^24. Entomological Society 12:1-22. Bericht Baker, C. F. 1906. American bees related toMelectn. Invertebrata Pacifica Alfken, ]. D. 1912. Die Bienenfauna von Westpreussen. des 1:142-145. Westpreussischen Botanisch-Zoologischen Vereins [Danzig] 34:1-94, Baker, D. B. 1994. A new of nomadine bees from north Africa. pis. l-II. genus der mit Tijdschrift voor Entomologie 137:155-159. Alfken, J. D. 1924. Wissenschaftliche Ergebnisse unterstiitzung Andrcna. Deutsche der Akademie der Wissenschaften in Wien aus der Erbschaft Treitl Baker, D. B. 1995a. A new Malayan Entomologische Zeitschrift (n.f.)42:67-69. von E Werner unternommenen zoologischen Expedition nach dem Baker, D. B. 1995b. Xertimnidmtcs 1951 (Insecta, Anglo-Agyptischen Sudan (Kordofan) 1914, XVI Hymenoptera F. Popov, Hymenoptera): of Ammobates (Xcrnnunobates) oxintms Popov, Apidae. Denkschriften der Akademie der Wissenschaften in Wien, Proposed designation Nomenclature 52:157- Mathematische-Naturwissenschaftliche Klasse 99:247-257. 1951 as the type species. Bulletin of Zoological 158. Alfken, J. D. 1926a. Die mir bekannten chilenischen Arten der F. la de la Societe Ball, J. 1914. Les bourdons de Belgique. Annales Bienengattung Corynum M. Spinola. Deutsche Entomologische de 58:77-108, 1 Zeitschrift 1926:145-163. Entomologique Belgique pi de la Treubia Benoist, R. 1962. Nouvelles especes d'apides malgaches. Bulletin Alfken, J. D. 1926b. Fauna Buruana, Hvmenoptera, Fam. Apidae. Societe de France 67:214-223. 7:259-275. Entomologique R. and O. W. Richards. 1937. Part 5. The der Benson, B., C. Ferriere, generic Alfken, J. D. 1933. Beitrag zur Kenntnis Untergattung Pseudomegachile names of the British Aculeata, with a check list of Brit- Friese. Konowia 12:55-59. Hvmenoptera ish 81-149 m Tlie Generic Names Insects. London: von Palastina. species. Pp. ofBritisli Alfken, ]. D. 1935. Beitrag zur Kenntnis der Bienenfauna Royal Entomological Society. Veroffentlichungen des Deutschen Kolonial- und Uebersee-Museums, Berthold, A. A. 1827. Lntreitle's naturliclw Famtlien des Thierreicbs mit Bremen 2:169-192. und Zusntzen. Weimar: 8 + 602 der Crocisn Anmerkungen Industr.-Compt. pp. Alfl^en, J. D. 1937. Zur Unterscheidung Bienengattungen jur. Bischoff, H. 1923. Zur Kenntnis afrikanischer Schmarotzerbienen. und Mck'ctn Latr. Konowia 16:172-175. Bienenfauna Deutsche Entomologische Zeitschrift 1923:585-603. Alfken, J. D. 1938. Ein weiterer Beitrag zur Kenntnis der Bischoff, H. 1930. zur Kenntnis Arten der Gattung von Palastina mit Einschluss des Sinai-Gebirges. Deutsche Beitrag palaarktischer Epeclus. Deutsche Entomologische Zeitschrift 1930:1-15. Entomologische Zeitschrift 1938:418^33. H. 1936. Schwedisch-chinesische wissenschaftliche 111-122 in Missione Bischoff, Expedition Alfken, J. D. 1939. Hymenoptera, Apidae. Pp. Bwlogica nach den nordwestlichen Provinzen Chinas, unter Leitung von Dr. nel Paese dei Bcrana. Vol. 3, Raccolte Zoologiche. Part 2. Roma: Reale Hedin und Prof. Su Insekten vom Accademia d'ltalia. Sven Ping-chang. gesammelt schwedischen Arzt der Dr. David Hummel 1927-1930. Armbruster, L. 1938. Versteinerte Honigbienen aus dem obermiocanen Expedition 10. Arkiv 27:1-27. Randecker Maar. Archiv fiir Bienenkunde 19:1-48, 97-133. 56. Hymenoptera. Bombinae. forZoologi E. 1840. 219-415, 1-7, m F L. N. Ashmead, W. H. 1890. On the Hymenoptera of Colorado; Descriptions Blanchard, Hymenopteres. Pp. pis. Laporte Bul- de Castelnau, Histoire Naturelle des Insectes. . . . Vol. 3. Paris: Dumenil. of new species, notes, and a list of the species found in the state. Castelnau is cited as the name of the editor of this work, letin of the Colorado Biological Association No. 1:1-47. commonlv with de seems Ashmead, W. H. 1894. The habits of aculeate Hvmenoptera II. Psyche but for consistency Lepeletier Saint-Fargeau, Laporte 7:40^6. more appropriate, as indicated by D. B. Baker, in htt. (1996). in G. Ashmead, W. H. 1898. Some new genera of bees. Psyche 8:282-285. Blanchard, E. 1845-1849. Hymenopteres. Pp. 113-227, pis. 107-129, Ashmead, W. H. 1899a. Classification of the bees, or the superfamily L. C. F. D. Cuvier, Le Ri'gne Animal .... ed. 3 ("disciples' edition"). Vol. 2. Masson. dated but see F. C. Apoidea. Transactions of the American Entomological Society 26:49- Paris: Fortin, Usually 1849; Cowan, 100. 1976, Journal of the Society for the Bibliography of Natural History 8:32-64. The text on bees (Les 201-227) in Ashmead, W. H. 1899b. A generic table of the family Panurgidae: A reply Melliferes, pp. appeared 1846. Plates and their of are as foUows: 125, 1847; to Mr. Cockerell's critique on the segregation of Pcrdita Cockerell (sic). years publication 128 1845. Psyche 9:372-376. 126, 1846; 127, 1846; 128, 1847; bis, 1846; 129, 64 Scientific Papers, Natural History Museum, The University of Kansas

Bliithgen, P. 1926. Beitrage zur Kenntnis tier indo-malayischen Halictiiii- Cameron, P. 1901h. Description of a new genus of bees from India. Ento- und Tliriiicostonia-Arten. Zoologische Jahrbiicher, Abteilung fur mologist 34:262-263. 4-5. Systematik, Geographie und Biologie der Tiere 51:375-698, pis. Cameron, P. 1902a. Descriptions of new genera and species of Hy- Bliithgen, P. 1928a. 2. Beitrag zur Kenntnis der athiopischen Halictinae. menoptera collected by Major C. G. Nurse at Deesa, Simla and Deutsche Entomologische Zeitschrift 1928:49-72. Ferozepore, Part II. Journal of the Bombay Natural History Society Bliithgen, P. 192Sb. Beitrage zur Kenntnis der afrikanischen Halictinae. 14:419-449. Zoologische Jahrbiicher, Abteilung fiir Systematik, Geographie und Cameron, P. 1902b. On the Hymenoptera collected by Mr. Robert Shelford Biologie der Tiere 55:163-252. at Sarawak and on the Hymenoptera of the Sarawak Museum. Jour- Bliithgen, P. 1930. Beitrag zur Kenntnis der athiopischen Halictinae. nal of the Straits Branch of the Royal Asiatic Society 37:29-140. Mitteilungen aus dem Zoologischen Museum in Berlin 15:495-542. Cameron, P. 1903a. Descriptions of new species of Hymenoptera taken Bliithgen, P. 1931. Beitrage zur Kenntnis der indomalayischen Hnlictuf- by Mr. Edward Whymper on the "Higher Andes of the Ec|uator." und Thrinaistomti- Arten. Zoologische Jahrbiicher, Abteilung fiir Transactions of the American Entomological Society 29:225-238. Systematik, Geographie und Biologie der Tiere 61:285-346. Cameron, P. 1903b. Descriptions of new genera and species of Hy- Bliithgen, P. 1933. Beitrag zur Kenntnis der athiopischen Halictinae. menoptera taken by Mr. Robert Shelford at Sarawak, Borneo. Journal Mitteilungen aus dem Zoologischen Museum in Berlin 18:363-394. of the Straits Branch of the Royal Asiatic Society 39:89-181. Bliithgen, P. 1936. Beitrag zur Kenntnis der athiopischen Halictinae. Cameron, P. 1905. On the Hymenoptera of the Albany Museum, Deutsche Entomologische Zeitschrift 1935:177-190. Grahamstown, South Africa. Records of the Albany Museum 1:185- Bliithgen, P. 1937. Halictinae (Hymenoptera; Apidae) von den 265. Kanarischen Inseln. Commentationes Biologicae (Societas Scientiarum Cameron, P. 1907. On a new genus and some new species of aculeate Fennica)6(ll):l-ll. Hymenoptera collected by Lieut. -Col. C. G. Nurse in Baluchistan. Bluthgen, P. 1949. Neues oder Wissenswertes uber mitteleuropaische Journal of the Bombay Natural History Society 18:130-136. Aculeaten und Goldwespen I. Apidae. Beitrage zur Taxonomischen Cockerell, T D. A. 1895. New species of bees. Psyche 7, Supplement 1:9-11. Zoologie 1:77-100. Cockerell, T. D. A. 1896. The bees of the genus Andiom found in New Bluthgen, P. 1955. The Halictinae (Hymen., Apoidea) of Israel. I. Genus Mexico. Annals and Magazine of Natural History (6)18:78-92. Halictiis (subgenera Halicttm s. str. and Tlirincohnlictus). Bulletin of the Cockerell, T. D. A. 1897a. On the generic position of some bees hitherto Research Council of Israel (B)5:5-23. referretl to Pniiurgus and Calliopfis. Canadian Entomologist 29:287- Bluthgen, P. 1961. Ergebnisse der Deutschen Afghanistan-E.xpedition 1956 290. der Landessammlungen fiir Naturkunde Karlsruhe. Beitrage zur Cockerell, T D. A. 1897b. On the Mexican bees of the genus Augochhru. Naturkundlichen Forschung in Siidwestdeutschland 19:277-287. Canadian Entomologist 29:4-6. Bohart, G. E. 1965. A new genus of dufoureine bee from Texas (Hy- Cockerell, T D. A. 1897c. A new genus of bees. Entomological News 8:197. menoptera: Halictidae). Annals of the Entomological Society of Cockerell, T. D. A. 1897d. Life-zones in New Mexico. Agricultural Ex- America 58:319-321. periment Station, New Mexico College of Agriculture, Bulletin No. Borner, C. 1919. Stammesgeschichte der Hautflugler. Biologisches 24:1^14. Zentralblatt 39:145-185. Cockerell, T D. A. 1898a. New bees from New Mexico. Canadian Ento-

Brauns, H. 1902. Eucoiidylops n. g. apidarum. Zeitschrift fiir systematische mologist 30:146-148. Hymenopterologie und Dipterologie 2:377-380. Cockerell, T. D. A. 1898b. On some panurgine and other bees. Transac- Brauns, H. 1926. V. Nachtrag zu "Friese, Bienen Afrikas." Zoologische tions of the American Entomological Society 25:185-198. Jahrbiicher, Abteilung fiir Systematik, Geographie und Biologie der Cockerell, T. D. A. 1899a. Notes on nomenclature of some Hymenoptera. Tiere 52:187-230, pi. 5. Entomologist 32:14. Brauns, H. 1929. Neue und auffallende Apiden aus Sud-Afrika. Zeitschrift Cockerell, T D. A. 1899b. Some synonymy Psyche 8:311-312. fiir Wissenschaftliche Insektenbiologie 24:130-143. Cockerell, T. D. A. 1899c. Cntrilogo dc Ins Abejns de Mexico. Ciudad de Brethes, J. 1909a. Hymenoptera Paraguayensis. Anales del Museo Mexico: Secretaria de Fomento. 20 pp. Nacional de Buenos Aires 20:225-256. Cockerell, T D. A. 1900a. Descriptions of new bees collected by Mr. H. H. las del Plata del in Brazil I. of the of of Brethes, J. 1909b. Himenopteros nuevos de Repiiblicas y Smith Proceedings Academy Natural Sciences Brasil. Anales del Museo Nacional de Buenos Aires 19:49-69. Philadelphia 52:356-377. Brethes, J. 1909c. Himenopteros de Mendoza y de San Luis. Anales del Cockerell, T. D. A. 1900b. Observations on bees collected in Las Vegas, Museo Nacional de Buenos Aires 17:455-463. New Mexico, and in the adjacent mountains. Annals and Magazine Brethes, J. 1910. Sur les AncyloccUs et genres voisins [Hym.]. Bulletin de of Natural History (7)5:401^16. la Societe Entomologique de France 1910:211-213. Cockerell, T. D. A. 1901a. Descriptions of new bees collected by Mr. H. H. 1922. varias in Brazil II. of the of Sciences of Brethes, J. Himenopteros y Dipteros de procedencias. Anales Smith Proceedings Academy Natural de la Sociedad Cientifica Argentina 93:119-146. Philadelphia 53:216-222. Bridwell, J. C. 1919. Miscellaneous notes on Hymenoptera. Proceedings Cockerell, T. D. A. 1901b. A peculiar new type of halictine bees. Entomo- of the Hawaiian Entomological Society 4:109-165. logical News 12:208-209. Broemeling, D. K. 1988. A revision of the Nomada subgenus Nomaditn of Cockerell, T. D. A. 1903a. A catalogue of the bees of California. Psyche North America. Pan-Pacific Entomologist 64:321-344. 10:74-78. Brooks, R. W. 1988. Systematics and phylogeny of the anthophorine bees. Cockerell, T D. A. 1903b. New American Hymenoptera, mostly of the University of Kansas Science Bulletin 53:436-575. genus Nomada. Annals and Magazine of Natural History (7)12:200- Buysson, R. du. 1900. Sur quelques hymenopteres de Madagascar. 214. Annales de la Societe Entomologiciue de France 69:177-180. Cockerell, T. D. A. 1903c. North American bees, and a new homopteron. of Camargo, J. M. F, and J. S. Moure. 1983. Tricholrigona, um novo genero Annals and Magazine Natural History (7)12:442^55. de Meliponinae (Hymenoptera, Apidae), do Rio Negro, Amazonas, Cockerell, T. D. A. 1903d. Bees of the genus Nomada from California. Pro- Brasil. Acta Amazonica 13:421-429. ceedings of the Academy of Natural Sciences of Philadelphia 55:559- Cameron, P. 1898. Hymenoptera Orientalia, or contributions to a knowl- 614. edge of the Hymenoptera of the Oriental zoological region. Part VII. Cockerell, T D. A. 1904a. The genus Apista, etc. Canadian Entomologist Memoirs, Manchester Literary and Philosophical Society 42(11 ):1- 36:357. 84, pi. 4. Cockerell, T. D. A. 1 904b. New genera of bees. Entomological News 15:292. Cameron, P. 1901a. Descriptions of three new genera and seven new spe- Cockerell, T. D. A. 1904c. The bees of southern California. 1. Bulletin of cies of Hymenoptera from eastern Asia and Australia. Annals and the Southern California Academy of Sciences 3:3-6. Magazine of Natural History (7)8:116-123. Cockerell, T D. A. 1904d. New records of bees. Entomologist 37:231-236. Genus-Group Names of Bees 65

Cockerell, T. D. A. 1905a. Notes on some bees in the British Museum. Cockerell, T. D. A. 191 le. Descriptions and records of bees. XXXIX. An- Transactions of the American Entomological Society 31:309-364. nals and Magazine of Natural History (8)8:660-673. Cockerell, T. D. A. 1905b. Descriptions and records of bees. III. Annals Cockerell, T. D. A. 19111. New and little known bees. Transactions of the and Magazine of Natural History (7)16:301-308. American Entomological Society 37:217-241. Cockerell, T. D. A. 1905c. Some American Hymenoptera. Entomological Cockerell, T. D. A. 1912. New and little-known bees. Entomologist 45: 1 75- News 16:9-11. 178. Cockerell, T. D. A. 1905d. New Austrahan bees in the collection of the Cockerell, T. D. A. 1913. The bee genus HopUtella. Canadian Entomolo- British Museum. Entomologist 38:270-273, 302-304. gist 45:34. Cockerell, T. D. A. 1906a. Fossil Hymenoptera from Florissant, Colorado. Cockerell, T. D. A. 1914. Bees from Ecuador and Peru. Journal of the New Bulletin of the Museum of Comparative Zoology, Harvard Univer- York Entomological Society 22:306-328. sity 50:33-58. Cockerell, T. D. A. 1915. Descriptions and records of bees. LXVl. Annals Cockerell, T. D. A. 1906b. Four interesting Australian bees, in the collec- and Magazine of Natural History (8)15:341-350. tion of the British Museum. Entomologist 39:15-18. Cockerell, T. D. A. 1916a. Descriptions and records of bees. LXXll. An- Cockerell, T. D. A. 1906c. North American bees of the genera Aiidtriin nals and Magazine of Natural History (8)17:428-435. and Mclittii in the British Museum. Psyche 13:33-37. Cockerell, T. D. A. 1916b. Descriptions and records of bees. LXXIII. An- Cockerell, T. D. A. 1906d. The North American bees of the family nals and Magazine of Natural History (8)18:44-53. Anthophoridae. Transactions of the American Entomological Soci- Cockerell, T. D. A. 1916c. New and little known bees from California. ety 32:63-116. Pomona Journal of Entomology and Zoology 8:43-64. T. D. A. 1906e. bees of Colorado. Bulletin of the Cockerell, The Florissant, Cockerell, T. D. A. 1916d. Some neotropical parasitic bees (Hym.). Ento- of Natural 22:419^55. American Museum History mological News 27:208-210. 40:227-229. Cockerell, T. D. A. 1907a. A fossil honey-bee. Entomologist Cockerell, T. D. A. 1917a. New social bees. Psyche 24:119-128. T. D. A. 1907b. and records of bees. XV. Annals Cockerell, Descriptions Cockerell, T. D. A. 1917b. Some euglossine bees. Canadian Entomologist and Magazine of Natural History 7(2n):59-68. 49:144-146. T. D. A. 1907c. a collection of Australian and Asiatic bees. Cockerell, On Cockerell, T. D. A. 1917c. New records of bees from Natal. Annals of the Bulletin of the American Museum of Natural 23: 221-236. History Durban Museum 1 :460^68. Cockerell, T. D. A. 1908a. Notes on the Exacrcte. 15:41- bee-genus Psyche Cockerell, T. D. A. 1918a. The megachilid bees of the Philippine Islands. 42. Philippine Journal of Science (D)13:127-144. Cockerell, T. D. A. 1908b. A new of African bees. subgenus Entomologist Cockerell, T D. A. 1918b. Neotropical bees, principally collected by Pro- 41:146-147. fessor Bruner in Argentina. Transactions of the American Entomo- Cockerell, T. D. A. 1908c. Descriptions and records of bees. XIX. Annals logical Society 44:25-38. and of Natural (8)1:337-344. Magazine History Cockerell, T D. A. 1919a. Bees in the collection of the United States Na- Cockerell, T. D. A. 1908d. and records of bees. XX. Annals Descriptions tional Museum 3. Proceedings of the United States National Museum and of Natural Magazine History (8)2:323-334. 55:167-221. T. D. A. 1909a. and records of bees. XXIll. Annals Cockerell, Descriptions Cockerell, T D. A. 1919b. The metallic-colored halictine bees of the Phil- and Magazine of Natural History (8)4:393^04. ippine Islands. Philippine Journal of Science 15:9-13. Cockerell, T. D. A. 1909b. Some European bees. Entomologist's Record Cockerell, T. D. A. 1920a. On South African bees, chiefly collected in Na- 21:269-270. tal. Annals of the Durban Museum 2:247-262. Cockerell, T D. A. 1909c. Descriptions of some bees iii the U.S. National Mu- Cockerell, T D. A. 1920b. On South African bees, chiefly collected in Natal. seum. of the United States National Museum 36:411^120. Proceedings Annals of the Durban Museum 2:286-318. Cockerell, T D. A. 1909d. Some European fossil bees. Entomologist 42:313- Cockerell, T. D. A. 1921a. Descriptions and records of bees. XC. Annals 317. and Magazine of Natxiral History (9)7:201-212. Cockerell, T D. A. 1909e. Descriptions of Hymenoptera from Baltic am- Cockerell, T. D. A. 1 92 1 b. The epeoline bees of the American Museum Rocky ber. Schriften der Gesellschaft zu Physikalische-okonomischen Mountain American Museum Novitates no. 23:1-16. 50:1-20. expeditions. Kiinigsberg Cockerell, T. D. A. 1922a. Notes on some western bees. American Mu- Cockerell, T. D. A. 1909f. Two fossil bees. News 20:159- Entomological seum Novitates no. 40:1-7. 161. Cockerell, T. D. A. 1922b. Descriptions and records of bees. XCV. Annals Cockerell, T. D. A. Some additional bees from Prussian amber. 1909g. and of Natural (9)10:265-269. Schriften der Physikalische-okonomischen Gesellschaft zu Magazine History Cockerell, T. D. A. 1922c. Bees of the genus Panurpnus obtained by the Konigsberg 50:21-25. American Museum Mountain expeditions. American Museum Cockerell, T. D. A. 1910a. New and little-known bees. Transactions of the Rocky Novitates no. 36:1-10. American Entomological Society 36:199-249. Cockerell, T. D. A. 1922d. Two new subgenera of North American bees. Cockerell, T. D. A. 1910b. Descriptions and records of bees. XXXIll. An- American Museum Novitates no. 47:1-5. nals and Magazine of Natural History (8)6:356-366. Cockerell, T.D.A. 1922e. Bees in the collection of the United States Na- Cockerell, T. D. A. 1910c. Some very small Australian bees. Entomologist 4. National Mu- 43:262-264. tional Museum. Proceedings of the United States seum 60:1-20. Cockerell, T. D. A. 1910d. Some new American bees. Canadian Entomolo- Cockerell, T. D. A. 1923a. Two nocturnal bees and a minute Pcrciitn. Ameri- gist 42:169-171. can Museum Novitates no. 66:1-4. Cockerell, T. D. A. 1910e. Descriptions and records of bees. XXIV. Annals T. D. A. 1923b. Some bees from British Guiana. Annals and and Magazine of Natural History (8)5:22-30. Cockerell, of Natural Cockerell, T. D. A. 1910f. The North American bees of the genus Noniia. Magazine History (9)11:442^59. T. D. A. 1924a. and records of bees. CI. Annals Proceedings of the United States National Museum 38:289-298. Cockerell, Descriptions and of Natural Cockerell, T. D. A. 1911a. A new genus of Australian bees. Entomologist Magazine History (9)14:179-185. 44:140-142. Cockerell, T. D. A. 1924b. A new genus of bees from California. Entomo-

Cockerell, T. D. A. 1911b. Bees in the collection of the United States Na- logical News 35:169-171 . T. D. A. 1924c. A new bee from 31:243-244. tional Museum, 2. Proceedings of the United States National Mu- Cockerell, Oregon. Psyche seum 40:241-264. Cockerell, T. D. A. 1924d. Notes on the structure of bees. Proceedings of Cockerell, T. D. A. 1911c. Descriptions and records of bees. XXXV. Annals the Entomological Society of Washington 26:77-85. T. A. Anthidiine bees in the collection of the Califor- and Magazine of Natural History (8)7:310-319. Cockerell, D. 1925a. Cockerell, T. D. A. 1911d. Descriptions and records of bees. XXXIV. An- nia Academy of Sciences. Proceedings of the California Academy of nals and Magazine of Natural History (8)7:225-237. Sciences (4)14:345-367. 66 Scientific Papers, Natural History Museum, The University of Kansas

Cockerell, T. D. A. 1925b. Bees in the collection of the California Acad- Cockerell, T. D. A. 1935a. Some California bees. Pan-Pacific Entomolo- emy of Sciences. Proceedings of the California Academy of Sciences gist 11:49-54. (4)14:185-215. Cockerell, T. D. A. 1935b. Scientific results of the Vernay-Lang Kalahari Cockerell, T. D. A. 1926a. Descriptions and records of bees. CVIll. Annals expedition, March to September, 1930, Hymenoptera (Apoidea). An- and Magazine of Natural History (9)17:214-226. nals of the Transvaal Museum 17:63-94. Cockerell, T. D. A. 1926b. Descriptions and records of bees. CXll. Annals Cockerell, T D. A. 1936a. Bees from northern California. Pan-Pacific En- and Magazine of Natural History (9)18:216-227. tomologist 12:133-164. Cockerell, T. D. A. 1926c. Descriptions and records of bees. CXIIl. Annals Cockerell, T. D. A. 1936b. Descriptions and records of bees. CLIV. Annals and Magazine of Natural History (9)18:621-627. and Magazine of Natural History (10)17:477^83. Cockerell, T. D. A. 1927a. Two new types of desert bees. Pan-Pacific En- Cockerell, T. D. A. 1938. Bees of the genus Splwcodes from the Belgian tomologist 4:41-44. Congo. Revue de Zoologie et de Botanique Africaines 30:327-329. Cockerell, T. D. A. 1927h. Megachilid bees from Bolivia collected by the Cockerell, T D. A. 1947. A new genus of bees from Honduras. Proceed- Mulford biological expedition, 1921-22. Proceedings of the United ings of the Entomological Society of Washington 49:106. States National Museum 71(12):l-22. Cockerell, T. D. A., and E. Atkins. 1902. Contributions from the New Cockerell, T. D. A. 1929a. Bees in the Australian Museum collection. Mexico Biological Station. XIII. On the bees of the family Nomaciidae Records of the Australian Museum 17:199-243. of Ashmead. Annals and Magazine of Natural History (7)10:40-46. Cockerell, T. D. A. 1929b. Bees in the Queensland Museum. Memoirs of Cockerell, T. D. A., and B. H. Blair. 1930. Rocky Mountain bees, 1. Ameri- the Queensland Museum 9:298-323. can Museum Novitates no. 433:1-19. Cockerell, T. D. A. 1929c. Descriptions and records of bees. CXV. Annals Cockerell, T. D. A., and W. P. Cockerell. 1901. Contributions from the and Magazine of Natural History (10)3:354-360. New Mexico Biological Station. IX. On certain genera of bees. Annals Cockerell, T. D. A. 1929d. Descriptions and records of bees. CXX. Annals and Magazine of Natural History (7)7:46-50. and Magazine of Natural History (10)4:584-594. Cockerell, T. D. A., and W. Porter. 1899. Contributions from the New Cockerell, T. D. A. 1929e. Descriptions and records of bees. CXIX. Annals Mexico Biological Station. VII. Observations on bees, with descrip- and Magazine of Natural History (10)4:296-304. tions of new genera and species. Annals and Magazine of Natural Cockerell, T. D. A. 1929f. New name for a genus of bees. Entomologist History (7)4:403^21. 62:19. C. 1903. of bees. Crawford, J. A new genus Canadian Entomologist 35:337- Cockerell, T. D. A. 1930a. African bees of the family Ctenoplectridae from 338. et the Belgian Congo and Liberia. Revue de Zoologie de Botanique Crawford, J. C. 1907. New North American Hymenoptera. Journal of the Africaines 18:359-363. New York Entomological Society 15:177-192. T. D. A. Cockerell, 1930b. African bees of the family Megachilidae, Crawford, ]. C. 1910. New Hymenoptera from the Philippine Islands. (Anthidiinae, Sempista and Lithurguf) mainly from Liberia and the Proceedings of the United States National Museum 38:119-133. et 1915. Belgian Congo. Revue de Zoologie de Botanique Africaines 19:43- Crawford, J. C. The bee genus Hokopc^iitcs Ashmead. Insecutor 55. Inscitiae Menstruus 3:123-126.

T. . 1 930c. of Australia . 6:205- C. 1916. Cockerell, D. A The bees Australian Zoologist Crawford, J. Some new American Hymenoptera. Insecutor 236. Inscitiae Menstruus 4:135-144. Cockerell, T. D. A. 1930d. A new subgenus of andrenine bees. Pan-Pacific Cresson, E. T. 1864. On the North American species of several genera of Entomologist 7:5-8. Apidae. Proceedings of the Entomological Society of Philadelphia Cockerell, T. D. A. 1930e. New Australian bees. Memoirs of the 2:373-411. Queensland Museum 10:37-50. Cresson, E. T. 1865. On the Hymenoptera of Cuba. Proceedings of the Cockerell, T. D. A. 19301. A new African genus of Ceratinidae (Hy- Entomological Society of Philadelphia 4:1-200. menoptera, Apoidea). Revue de Zoologie et de Botanique Africaines Cresson, E. T. 1875. Report upon the collections of Hymenoptera made 18:291-293. in portions of Nevada, Utah, Colorado, New Mexico, and Arizona. Cockerell, T. D. A. 1931a. Some African bees. Annals and Magazine of Report of the U.S. Geographical Survey west of the 100th Meridian Natural History (10)8:400^05. 5:705-728. Cockerell, T. D. A. 1931b. Heriadine and related bees from Liberia and Cresson, E. T 1878. Descriptions of new North American Hymenoptera the Belgian Congo. Revue de Zoologie et de Botanique Africaines in the collection of the American Entomological Society. Transactions 20:331-341. of the American Entomological Society 7:61-136. Cockerell, T. D. A. 1931c. African bees of the family Megachilidae from Cresson, E. T 1887. Synopsis of the families and genera of the Hy- Liberia and the Belgian Congo. Revue de Zoologie et de Botanique menoptera of America, north of Mexico. Transactions of the Ameri- Africaines 20:132-179. can Entomological Society, Supplementary Volume, 1887. Pp. 1-350 Cockerell, T D. A. 1931d. Descriptions and records of bees. CXXX. An- + errata page. nals R. e and Magazine of Natural History (10)8:537-553. Cure, J. 1989. Revisao de PseiidagapiKtenwn Schrottky descri(;ao de Cockerell, T. D. A. 1931e. Descriptions and records of bees. CXXVl. An- Oragaptostemcti gen. n. Revista Brasileira de Entomologia 33:229-335. nals of 1826. British Vol. 3. PI. and Magazine Natural History (10)7:273-281. Curtis, J. Eiitoniologx). London: privately printed. Cockerell, T. D. A. 19311. Descriptions and recordsof bees. CXXVlll. An- 99-146. nals of 1828. BriHsli Vol. 5. PI. and Magazine Natural History (10)7:529-536. Curtis, J. Entoiiwlc<^y. London: privately printed. Cockerell, T. D. A. 1932. Bees from the Belgian Congo. Revue de Zoologie 195-241. et Britisii Vol. 8. PI. de Botanique Africaines 23:18-27. Curtis, ]. 1831. Entonwtcg]/. London: privately printed. Cockerell, T. D. A. 1933a. Descriptions and records of bees. CXLIII. An- 338-383. nals of 1832. Britisli Vol. 9. PI. and Magazine Natural History (10)11:456-^68. Curtis, J. Entomology. London: privately printed. Cockerell, T. D. A. 1933b. Descriptions and records of bees. CXLV. An- 384-433. nals British Vol. 10. PI. and Magazine of Natural History (10)12:126-136. Curtis, J. 1833. Entomology. London: privately printed. Cockerell, T. D. A. 1933c. Bees collected at Borego, California, by Wilmatte 434-481. Cockerell and Milene Porter. Pan-Pacific Entomologist 9:25-28. Dalla Torre, K. W. von. 1880. Unsere Hummel-(B[)H//'/is) Arten. Die Cockerell, T. D. A. 1934a. Descriptions and records of bees. CXLVllI. Naturhistoriker 2:30, 40-41. Annals and Magazine of Natural History (10)13:444-456. Dalla Torre, K. W. von. 1882. Bemerkungen zur Gattung Bombus Latr., II. Cockerell, T. D. A. 1934b. Some African meliponine bees. Revue de Bericht des Naturwissenschaftlich-Medezinischen Vereins in Zoologie et de Botanique Africaines 26:46-62. Innsbruck 12:14-31. Cockerell, T. D. A. 1934c. Some new or little-known South African bees Dalla Torre, K. W. von. 1891. Die Gattungen und Arten der Phileremiden. of the genus AUodnpc in the British Museum. Annals and Magazine Bericht des Naturwissenschaftlich- Medezinischen Vereins in of NaUiral History (10)14:220-242. Innsbruck 19:137-159. Genus-Group Names of Bees 67

Dalla Torre, C. G. de. 1896. Catalcgus Hymcmiptcrcrum.... Vol. 10. Apnciae Exley, E. M. 1978. Chaetchesma-A new genus of Austrahan bees. Austra- viii of 26:373-397. (Aiitlwfihiln). Leipzig: Engelmann. + 643 pp. lian Journal Zoology Dalla Torre, C. G. de, and H. Friese. 1895. Synonymischer Katalog der Exley, E. M. 1996. Tumidihesma, a new genus of Australian bees. Austra- europaischen Sammelbienen. Entomologische Nachrichten 21 :21-26, lian Journal of Entomology 35:253-255. 37-50, 53-62, 69-80. Fabricius,J.C. 1775. Systema Enlomologiae.... Flensburgi et Lipsiae: Korte. 832 Daly, H. V. 1988. Bees of the new genus Ctenoceratina in Africa south of pp. C. 1793. Emendata et Aiicta. 2. the Sahara. University of California Publications in Entomology 108:i- Fabricius, J. Entotnohgia Systematica Vol. ix + 1-69. Hafniae: Proft. viii -(-519 pp.

H. D. S. S. Fabricius, J. C. 1804. Piezatorum.... xiv -i- [15|-[440] Daly V., C. Michener, J. Moure, and F Sakagami. 1987. The Systema Braunschweig, + The index in 1805 to relictual bee genus Mamielia and its relation to other Xylocopinae. [11-30 pp. (pp. [l]-30) appeared according Hedicke, 1941, der Deutschen Pan-Pacific Entomologist 63:102-124. Mitteilungen Entomologischen 1898. d'oeil sur les melliferes Gesellschaft 10:82-83. Dominique, J. Coup sud-americains du L. 1858. Ordre ;;( au museum de Nantes. Bulletin de la Societe Naturelles de I'Ouest de la Fairmaire, Hymenopteres, J. Thomson, Voyage Gabon, Histoire Naturelle des Insectes et des Arachnides. Archives France [Nantes] 8(l):57-63. 2:263-267, on 10. Dours, L. 1872. Hymenopteres nouveaux du bassin Mediterraneen. Re- Entomologiques figs, pi. Fallen, C. F. 1813. Novam Metliodiim vue et Magasin de Zoologie (2)23:293-311, 349-359, 396-399, 419- Specimen Hymenoptera Disponendi Exhibens. Lund: Berling. 41 pp., 1 pi. 434, pi. 28. Ferton, C. 1914. Perezia maura-noveau genre Dours, L. 1873. Hymenopteres du bassin Mediterraneen. Andrenn (suite). d'apiaires parasites d'Algerie et observations de ce genre. Annales de la Societe Entomologique de Revue et Magasin de Zoologie (3)1:274-325, pi. 14. France 83:233-237. Dressier, R. L. 1978. An classification of En^Iossn, with notes infrageneric 1894. Fox, W. J. A proposed classification of the fossorial Hymenoptera on some features of special taxonomic importance. Revista de Biologia of North America. Proceedings of the Academy of Natural Sciences Tropical 26:187-198. of Philadelphia 1894:292-307. Dressier, R. L. 1982. New species of Euglossn. 111. The bursigem species Fox, W. 1902. a new of bees. News Revista de 30:131-140. J. Lithurgopsis, genus Entomological group. Biologia Tropical 13:137-140. Ducke, A. 1900. Die Bienengattung Osmia Panz. Bericht des Franklin, H. J. 1954. The evolution and distribution of American bumble- Naturwissenschaftlich-Medezinischen Vereins in Innsbruck 25:1-323. bee kinds. Transactions of the American Entomological Society 80:43- A. 1902. Ein neues von Halictus Latr Zeitschrift fiir Ducke, subgenus 51. 2:102-103. systematische Hymenopterologie und Dipterologie H. Die Bieneu L Friese, 1895. Eiirciprn's. Thiel Berlin: Friedlander. 128 pp. Ducke, A. 1907. zur Kenntnis der Solitarbienen Brasiliens. Beitrag Friese, H. 1896. Neue oder wenig bekannte Stideuropaische Apiden. Zeitschrift fijr und systematische Hymenopterologie Dipterologie Termeszetrajzi Fiizetek 19:277-284. 7:361-368. Friese, H. 1897a. Die Bienen Europa's. Theil 111. Berlin: Friedlander vi + Ducke, A. 1912. Die naturlichen Siidamerikas. Bienengenera Zoologische 316 pp. fur und der Jahrbiicher, Abteilung Systematik, Geographic Biologie Friese, H. 1897b. Monographic der Bienengattung Nomia (Latr.) Tiere 34:51-116. (Palaearctische Formen). Pp. 45-84 in Festschrift zur Feier des Dumeril, A. M. C. 1860. anati/tique. Histoire dassifka- Entonwiogie gencrale, fiinfzigjdhrigen Bestehens des Vereins fiir schlesische Insektenkunde in tioti imtiirelle Insectes a I'aide de tableaux inetlioiliqiie des synoptiques. Paris: Breslau. Breslau: Maruschke and Behrendt. (Mem. Acad. Sci. 31:i-xxii + 1-1239). Friese, H. 1898a. Die Bienen Europya's. Theil IV. Innsbruck: C. Lampe. 303 Ebmer, A. W. 1984. Die Arten der Diifourea westpalaarktischen Gattung pp.; 1 pi. 1841 mit illustrierten Lepeletier Bestimmungstabellen. Friese, H. 1898b. Species aliquot novae vel minus cognitae generis 64:313-379. Senckenbergiana Biologica Megachile Latr. (et Chalicodoma Lap.). Termeszetrajzi Fiizetek 21:198- Ebmer, A. W. 1988. Kritische Liste der nicht-parasitischen Halictidae 202. Osterreichs mit aller Arten. Beriicksichtigung mitteleuropaischen Friese, H. 1898e. Beitrage zur Bienenfauna von Aegypten. Termeszetrajzi Linzer Biologische Beitrage 20:527-711. Fuzetek 21:303-313. A. W. 1993. Die Arten der Dufourea Ebmer, westpalaarktischen Gattung Friese, H. 1899a. Monographic der Bienengattungen Megacilissa, Lepeletier 1841 mit illustrierten Bestimmungstabellen, Dritter Caupolicana und Oxaea. Annalen des K. K. Naturhistorischen Nachtrag. Linzer Biologische Beitrage 25:15^2. Hofmuseums (Wien) 14:239-246. G. C. 1969. and Eickwort, A comparative morphological study generic Friese, H. 1899b. Monographic der Bienengattimg Euglossa. Termeszetrajzi revision of the augochlorine bees. University of Kansas Science Bul- Fuzetek 22:117-170. letin 48:325-524. Friese, H. 1899c. Die Bienen Eurcfm's. Theil V. Innsbruck: C. Lampe. 228 Eickwort, G. C. 1978. Mexalictiis, a new genus of sweat bees from North PP- America (Hymenoptera, Halictidae, Halictinae). Journal of the Kan- Friese, H. 1900. Monographic der Bienengattung Centris (s. lat.). Annalen sas Entomological Society 51:567-580. des K. K. Naturhistorischen Hofmuseums (Wien) 15:237-350. Eickwort, G. C, and G. 1. Stage. 1972. A new subgenus of neotropical Friese, H. 1902. Zwei neue Bienengattungen. Zeitschrift fiir systematische Spihecodes cleptoparasitic upon Dialictus (Hymenoptera: Halictidae, Hymenopterologie und Dipterologie 2:186-187. Halictinae). Journal of the Kansas Entomological Society 45:500-515. Friese, H. 1903a. Neue Meliponiden II. Zeitschrift fiir systematische Enderlein, G. 1903. Drei neue Bienen mit rijsselartiger Verlangenrng des Hymenopterologie und Dipterologie 3:359-361. Kopfes. Berliner Entomologische Zeitschrift 48:35^0. Friese, H. 1903b. Melitnrgula, eine neue Bienengattung aus Sud-Afrika. Engel, M. S. 1997. New augochlorine bees (Hymenoptera: Halictidae) in Zeitschrift fiir systematische Hymenopterologie und Dipterologie Dominican amber, with a brief review of fossil Halictidae. Journal of 3:33-34. the Kansas Entomological Society, 69(1996 suppl.):334-345. Friese, H. 1905a. Die Wollbienen Afrikas. Genus Anthidium. Zeitschrift Exley, E. M. 1968a. Qtmsihesnm-a new genus of Australian bees. Austra- fiir systematische Hymenopterologie und Dipterologie 5:65-75. lian Journal of Zoology 16:227-235. Friese, H. 1905b. Neue AntliofJlwra-Avten aus Afrika (Hym.). Zeitschrift Exley, E. M. 1968b. Revision of the genus Brachyhesma Michener. Austra- fiir systematische Hymenopterologie und Dipterologie 5:233-242. han Journal of Zoology 16:167-201. Friese, H. 1906a. Die Bienengattung Oediscelis Philipp. [sic] und Pseiidiscclis Exley, E. M. 1969. Argohesma-\ new genus of Australian bees. Australian Friese. Zeitschrift fiir systematische Hymenopterologie und Journal of Zoology 17:527-534. Dipterologie 6:225-228. Exley, E. M. 1972. Revision of the genus Pachyprosopis Perkins. Austra- Friese, H. 1906b. Neue Bienenarten aus Chile und Argentina. Zeitschrift lian Journal of Zoology, Supplementary Series No. 10:1^3. fiir systematische Hymenopterologie und Dipterologie 6:169-176. Exley, E. M. 1977. The Australian genus Brachyhesma (Apoidea: CoUetidae) Friese, H. 1906c. Eine neue Bienengattung aus Chile und Argentinien. revised and reviewed. Australian Journal of Zoology, Supplemen- Zeitschrift fiir systematische Hymenopterologie und Dipterologie tary Series No. 53:1-54. 6:374-380. 68 Scientific Papers, Natural History Museum, The University of Kansas

Friese, H. 1906d. Resultate einer Reise des Herrn A. C. Jensen-Haarup in Friese, H. 1923. Die europaischen Bienen. Berlin and Leipzig: Gruyter. xvii Flora + die Gegend von Mendoza (Argentina). og Fauna 8:89-102. 456 pp., 33 pi. Friese, H. 1906e. Neue Schmarotzerbienen aus der neotropischen Region. Friese, H. 1925a. Neue neotropische Bienenarten, zugleich 11. Nachtrag Zeitschrift fiir systematisclie Hvmenopterologie und Dipterologie zur Bienenfauna von Costa Rica. Stettiner Entomologische Zeitung 6:118-121. 86:1^1. Friese, H. 1908a. Neue Bienenarten aus Ostafrika. Deutsche Friese, H. 1925b. IV. Nachtrag zu "Bienen Afrikas." Zoologische Entomologische Zeitschrift 1908:567-572. Jahrbiicher, Abteilung fiir Systematik, Geographie und Biologie der Friese, H. 1908b. Die Apidae (Blumenwespen) von Argentina nach den Tiere 49:501-512. Reisenergebnissen der Herren A. C. Jensen-Haarup und P. Jorgensen Friese, H. 1930. Corrigenda zu "Neue neotropische Bienenarten" in Stettin, in den Jahren 1904-1907. Hora og Fauna 10:331^25, reprint 1-94. Entomologische Zeitung vol. 86, 1925. Stettiner Entomologische Friese, H. 1908c. Fine neue Bienengattung Ccrhicuhi aus Argentina. Zeitung 91:127. Zeitschrift fiir systematische Hymenopterologie und Dipterologie Prison, T. H. 1927. A contribution to our knowledge of the relationships 8:170-174. of the Bremidae of America north of Mexico (Hymenoptera). Trans- Friese, H. 1909a. Die Bienen Afrikas nach dem Stande unserer heutigen actions of the American Entomological Society 53:51-78, pis. XVI, XVll. Kenntnisse. Pp. 83-476, pis. ix-x in L. Schultz (ed.), Zoologische und Prison, T. H. 1930. The bumblebees of Java, Sumatra and Borneo. Treubia Antliropclogisdw Ergcbnisse einer Forscltungsrcise im icesHichen und 12:1-22. zentmleu Siidnfrikn nusgrfuhrt in den jahren 1 903-1 905. Band 2. Lieferung Prison, T. H. 1935. Records and descriptions of Bremu!^ from Asia 1. X Insecta (Ser.3) IJenaische Denkschriften Vol. 14J. Jena: Gustav (Bremidae: Hymenoptera). Records of the Indian Museum 37:339- Fischer. 363, 7 fig. Friese, H. 1909b. Die Bienenfauna von Neu-Guinea. Annales Musei Germar, E. P. 1839. Die versteinerten Insecten Solenhofens. Nova Acta K. Nationalis Hungarici 7:179-288. Leopoldinisch-Carolinische Deutsche Akademie der Naturforscher Friese, H. 1910a. Zur Bienenfauna des siidlichen Argentinien (Hym.). 19:187-222, 3 pis. Zoologische Jahrbiicher, Abteilung fiir Systematik, Geographie und Gerstaecker, A. 1857. [Bees and wasps collected in Mozambique]. Biologie der Tiere 29:641-660. Monatsberichte, Akademie der Wissenschaften, Berlin, 29 October Friese, H. 1910b. 5. Apidae. Pp. 119-168 in Y. Sjostedt, Wi^senschaftliche 1857, pp. 460-464. Ergebnifie der selnoedischen zoclogisehen Exptedition nach dem KiUnianjaro, Gerstaecker, A. 1868. Psaenythia, eine neue Bienengattung mit gezahnten dem Meru und den umgehenden Massaisleppen Deutsch-Ostafrihis 1905- Schienensporen. Archiv fiir Naturgeschichte 34:111-137. 1906. Band 2. Abt. 8 (Hymenoptera). Pt. 5 (Apidae). Stockholm: Gerstaecker, A. 1869. Beitrage zur naheren Kenntniss einiger Bienen- Palmquist. Gattungen. Entomologische Zeitung 30:139-184, 315-367. Friese, H. 1911a. Neue Bienen der athiopischen Region. Zoologische Girard, M. J. A. 1879. Les Insecies. Trade Elementaire d' Entomologie. Vol. 2. Jahrbiicher, Abteilung fiir Systematik, Geographie und Biologie der Paris: Bailliere. [i] + 1028 pp. Tiere 30:671-678. 1863. Giraud, J. Hymenopteres recueillis aux environs de Suse, en Friese, H. 1911b. Apidae I, Megacluliuae. Das Tierreieh. Lieferung 28. Ber- Piemont, et dans le departement des Hautes-Alpes, en France; et de- lin: Friedlander xxvi + 440 pp. scription de c^uinze especes nouvelles. Verhandlungen der Friese, H. 1911c. Nachtrag zu "Bienen Afrikas." Zoologische Jahrbiicher, Zoologisch-Botanischen Gesellschaft in Wien 13:11-46. fiir X. Abteilung Systematik, Geographie und Biologie der Tiere 30:651- Gistel [Gistll, J. N. F 1848. Naturgeschiclde des Thierreichs, fiir Hcihere 670. Scln{k'u. Stuttgart: Hoffmann, xvi + 216 pp., 32 pis. X. 1850. Friese, H. 1911d. Neue Bienen-Arten der palaearktischen Region (Hym.). Gistel, J. N. F Isis, Encyclopadische Zeitschrift, no. 6:82. Archiv fur Naturgeschichte, Abt A 77(1 ):135-143. Gray, G. 1832. [New genera and species.] In E. Griffith and E. Pidgeon. Friese, H. 1912. Neue und wenig bekannte Bienen Siid-Afrikas. Archiv C/i7ss Insecta, Arranged by the Baron Cuvier, with Supplementary Addi- fiir Naturgeschichte, Abt. A 78(5):181-189. tions to Each Order, and Notices of Neir Genera and Species by George Friese, H. 1913a. Neue Bienenarten aus Afrika. Deutsche Entomologische Gray, Esq. Vol. 2. [This is volume 15 of the series: Griffith, E., 1832, The Zeitschrift 1913:573-578. Animal Kingdom Arranged in Conformity with its Organization by the Friese, H. 1913b. II. Nachtrag zu "Bienen Afrikas." Zoologische Baron Cuiiier, Member of the Institute of France, etc., with Supplementary Jahrbiicher, Abteilung fiir Systematik, Geographie und Biologie der Additions to Each Order.] London. Tiere 35:581-598. Gribodo, G. 1884. Sopra alcune specie nuove o poco conosciute di Friese, H. 1914a. Neue Apiden der palaearktischen Region. Stettiner Imenotteri Antofili. BuUettino della Societa Entoniologica Itahana Entomologische Zeitung 75:218-233. (Horence) 16:269-274. Friese, H. 1914b. Die Bienenfauna von Java. Tijdschrift voor Entomologie Gribodo, G. 1894. Note Imenotterologiche, Nota II. Nuovi generi e nuove 57:1-61, 2 pis. specie di Imenotteri antofili ed osservazioni sobra alcune specie gia Friese, H. 1914c. Neue Bienenarten der orientalischen Region. Deutsche conosciute. BuUettino della Societa Entomologica Italiana (Florence) Entomologische Zeitschrift 1914:320-324. 26:76-135,262-314. Friese, H. 1916. Neue Bienen-Arten aus Chile und Sudamerika. Stettiner Griswold, T. 1986. A new heriadine bee from the Mojave Desert. South- Entomologische Zeitung 77:163-174. western Entomologist 11:165-69. Friese, H. 1918. Wissenschaftliche Ergebnisse einer Forschungsreise nach Griswold, T. L., and C. D. Michener 1988. Taxonomic observations on Ostindien, VII, Bienen aus Sumatra, Java, Malakka und Ceylon. Anthidiini of the western hemisphere. Journal of the Kansas Ento- Zoologische Jahrbiicher, Abteilung fiir Systematik, Geographie und mological Society 61:22-45. Biologie der Tiere 41:489-510. Guerin-Meneville, F. E. 1830. Crustaces, arachnides et insectes, Zool. 2, Friese, H. 1921a. Eine neue Bienengattung Melittoides n. gen., pp. 177- div. 1. /); M. L. 1. Duperrey, Voyage Autoiir du Monde, Execute par Ordre !« la 22 in 180, J. Fahringerand H. Friese, 1921, Eine Hymenopteren-Ausbeute du Roi, sur Corvette ...La Coquille. Paris: Bertrand. 319 pp., pis. aus dem Amanusgebirge (Kleinasien und Nord-Syrien, siidl. Atlas. [The introduction is dated 1838 but the text appears to have Armenien). Archiv fiir Naturgeschichte, Abt A 87(3):15(i-180. been published in 1830 and 1831.] Friese, H. 1921b. Nachtrag zur Bienenfauna von Costa Rica. Stettiner Guerin-Meneville, F. E. 1844. Iconographie du Regno Animal de C. Cuvier. Entomologische Zeitung 82:74-98. Vol. 7. Insects. Paris: Bailliere. 576 pp. H. Pasteels. 1961. all'elenco Friese, 1922a. Eine neue Gattung der Urbienen: Bracltx/glcssa. Guiglia, D., and J. J. Aggiunte ed osservazioni Zoologische Jahrbiicher, Abteilung fiir Systematik, Geographie und delle specie di imenotteri descritte da Guerin-Meneville che si trovano Biologie der Tiere 45:577-580. nelle coUezioni del museo di Genova. Annali del Museo Civico di Friese, H. 1922b. Eine neue Bienengattung aus Siidamerika: Rinnctula. Storia Naturale di Genova 72:17-30. Zoologische Jahrbiicher, Abteilung fiir Systematik, Geographie und Gupta, R. K. 1987. On a new subgenus Orientoheriades and two new spe- Biologie der Tiere 45:581-586. cies of genus Heriades Spinola from hidia. Reichenbachia 25(1 5):67-71 . Genus-Group Names of Bees 69

Y. 1 981 b. A of the bee Palaeorhiza Perkins Gupta, R. K. 1990. On a new subgenus Necaslunetiiiidk and two new spe- Hirashima, synopsis genus (Hy- of New Part IV. Cheesmania. cies of genus Ashmeadiella Cock, from India. Reichenbachia 29(10)55- menoptera, CoUetidae) Guinea, Subgenus 59. Kontyu 49:27-36. Hirashima, Y. 1982a. A of the bee Palaeorhiza Perkins Gupta, R. K. 1991. A new subgenus of Coelioxys Latreille (Hymenoptera: synopsis genus (Hy- CoUetidae) of New Guinea, Part VI. Apoidea; Megachilidae) from India. Journal of the Bombay Natural menoptera, Subgenus Zarhiopalea. 50:57-66. History Society 88:425-428. Kontyu Hirashima, Y. 1982b. A of the bee genus Palaeorhiza Perkins (Hy- Gupta, R. K. 1992. On a new subgenus Orientocodioxyf and a new species synopsis CoUetidae) of New Guinea, Part VII. Subgenus Cercorhiza. of genus Coelioxys Latr from India. Reichenbachia 29:73-76. menoptera, Journal of the Faculty of Agriculture, Kyushu University 26:87-109. Gupta, R. K. 1993. Taxonomic Studies oii the Megachilidae of North-Western Hirashima, Y. 1989. A synopsis of the bee genus Palaeorhiza Perkins (Hy- India. New Delhi: Indian Council of Agricultural Research. [4] + 288 menoptera, CoUetidae) of New Guinea, Part VIII. Subgenus Callorliiza. pp. Esakiano. 28:1-9, pi. 1. HaHday, A. H. 1836. Descriptions, etc. of the Hymenoptera. Pp. 316-331 Hirashima, Y. 1992. Anew name of Palaeorhiza. Japanese Journal of Ento- F. etc. the in J. Curtis, A. H. Haliday, and Walker (eds.). Descriptions, of mology 60:395. insects collected (n/ Captain P. P. King, R.N., F.R.S., in the survey of the Hirashima, Y. 1996. Subgeneric classification of the genus Xenorhiza Straits of Transactions of the Linnean Society of London 17. Magellan. Michener of Papua New Guinea. Memoirs of the Entomological So- Hedicke, H. 1922. Nomina nova. Deutsche Entomologische Zeitschrift ciety of Washington No. 17:78-81. 1922:427^28. Hirashima, Y, and M. A. Lieftinck. 1982. Systematic studies on the genus Hedicke, H. 1926. zur der Deutsche Beitrage Apidenfauna Philippinen. Palaeorhiza of New Guinea collected by the third Archbold expedi- Zeitschrift 1926:413-423. Entomologische tion (I). Esakia no. 19:1-50. Hedicke, H. 1933. zur der Andrena F. Beitrage Systematik Gattung Hirashima, Y., and O. Tadauchi. 1975. A new subgenus of the genus aus dem Museum in BerUn 19:199-220. Mitteilungen Zoologischen Andrena (Hymenoptera, Andrenidae) from Japan and allied areas. H. 1938. Uber vom Hindu-kusch. Deutsche Hedicke, einige Apiden Journal of the Faculty of Agriculture, Kyushu University 19:175-186. Zeitschrift 1938:186-196. 1817. bei Entomologische Hoffmannsegg, G. J. C. von. Entomologische Bemerkungen der von und Heer, O. 1850. Die Insektenfauna Tertiargebilde Oenigen Gelegenheit der Abhandlungen iiber amereikanische Insekten. von Radoboj in Croatien. Neue Denkschriften der Allgemeinen Magazin 1:8-56. fiir die Schweizerischen Gesellschaft gesammten Holmberg, E. L. 1884, 1886a. Viajes al Tandil y a La Tinta, 2nd Parte, ll:i-v + I-XVII. la Naturwissenschaften [Zurich] 1-264, pis. Zoologia, Insectos. I. Himenopteros- Hymenoptera. Actas de Heider, K. 1926. Pp. 161-320, in Nomenclator Animalium Generum et Academia Nacional de Ciencias de la Republica Argentina en Cordoba 2 Suhgenerum. Vol. 1. Lief. 2. Berlin: Preussischen Akademie der 5:117-136 (1884), 137-184, pis. (1886). Wissenschaften. Holmberg, E. L. 1886b, 1887a. Sobre Apidos Nomadas de la Republica et Anales la Sociedad Cientifica 22(1886):231- Heider, K. 1935. Pp. 2185-2248, in Nomenclator Animalium Generum Argentina. de Argentina 67-82. Suhgenerum. Vol. 4. Lief 16. Berlin: Preussischen Akademie der 240, 272-286; 23(1887):17-33, Wissenschaften. Holmberg, E. L. 1887b. Viaje a Misiones. Boletin de la Academia Nacional de Ciencias en Cordoba 10:5-391. Hirashima, Y. 1 956. Some bees of the genus Nomia Latreille from Formosa. Insecta Matsumurana 20:29-33. Holmberg, E. L. 1903. Delectus Hymenopterologicus Argentinus ... Anales del Museo Nacional de Buenos Aires (3)2:377-512. Hirashima, Y. 1963. Systematic and biological studies of the family E. L. 1909a. AUoscirtetica, Holmb. n. nom. de Historia Andrenidae of Japan (Hymenoptera, Apoidea) Part 2. Systematics, 2. Holmberg, Apuntes Natural [Buenos Aires] 1:77. Journal of the Faculty of Agriculture, Kyushu University 1 2:241-263. E. L. 1909b. Palinepeolus, Holmb. nov. nom. Apuntes de Hirashima, Y. 1964. Systematic and biological studies of the family Holmberg, Historia Natural [Buenos Aires] 1:77-78. Andrenidae of Japan (Hymenoptera, Apoidea) Part 2. Systematics, 4. Hong, Y.-c. 1984. New fossil insects of the Laiyang Group from the Journal of the Faculty of Agriculture, Kyushu University 13:71-97. Laiyang Basin, Shandong Prov. Professional Papers on Stratigraphy Hirashima, Y. 1965a. Systematic and biological studies of the family and Palaeontology [Beijing] 11:31-41. Andrenidae of Japan (Hymenoptera, Apoidea) Part 2. Systematics, 5. Houston, T. F 1981. A revision of the Australian hylaeine bees. II. Austra- Journal of the Faculty of Agriculture, Kyushu University 13:461-491. Uan Journal of Zoology, Supplementary Series No. 80:1-128. Hirashima, Y. 1965b. Systematic and biological studies of the family Hurd, P. D., Jr 1956. Notes on the subgenera of the new world carpenter Andrenidae of Japan (Hymenoptera, Apoidea) Part 2. Systematics, 6. bees of the genus Xylocopa. American Museum Novitates No. 1776:1- of the of 13:493-517. Journal Faculty Agriculture, Kyushu University 7. Y. of from New Hirashima, 1967. A new subgenus and,species Hylaeus in K. V. P. D. Hurd, P D., Jr. 1979. Apoidea. Pp. 1741-2209, Krombein, Guinea. 35:134-138. Kontyu Hurd, Jr, D. R. Smith, and B. D. Burks (eds.). Catalog of Hymenoptera Hirashima, Y. 1969. of the Pithitis Klug of the world (Hy- Synopsis genus in American North of Mexico. Vol. 2. Washington, D. C: Smithsonian Pacific Insects 11:649-669. menoptera: Anthophoridae). Institution Press. Hirashima, Y 1971a. classification of the Ceratina Subgeneric genus Hurd, P. D., Jr, and E. G. Linsley. 1970. A classification of the squash and Latreille of Asia and West Pacific, with comments on the remaining gourd bees Peponapis and Xenoglossa. University of California Publi- of the world of the Fac- subgenera (Hymenoptera, Apoidea). Journal cations in Entomology 62:i-iv + 1-39. of 16:349-375. ulty Agriculture, Kyushu University Hurd, P D., Jr, and E. G.Linsley 1976. The bee family Oxaeidae with a Y 1971b. a new of bees of Africa Hirashima, Megaceratina, genus (Hy- revision of the North American species. Smithsonian Contributions 5:251-256. menoptera, Anthophoridae). Journal of Natural History to Zoology no. 220:1-75. Y. 1978. of the bee Palaeorhiza Perkins Hirashima, A synopsis genus (Hy- Hurd, P. D., Jr, and C. D. Michener. 1961. testis Lepeletier and Serville, menoptera, CoUetidae) of New Guinea. Part 11. Subgenera Gressittapis, 1828 (Insecta: Hymenoptera); Proposed designation of a type species Esakia no. 12:63-87. Noonadnnia, Spliecogaster, Anchirhizn, Ceratorhiza. under the plenary powers. Bulletin of Zoological Nomenclature Hirashima, Y. 1980. A synopsis of the bee genus Palaeorhiza Perkins (Hy- 18:201-202. of Part III. P. 1960. new-world of bam- menoptera, CoUetidae) New Guinea, Subgenera Trachyrhiza, Hurd, D., Jr., and J. S. Moure. A subgenus Paraheterorhiza, Hadrorhiza. Journal of the Faculty of Agriculture, boo-nesting carpenter bees belonging to the genus Xylocopa Latreille. Kyushu University 25:99-117. Annals of the Entomological Society of America 53:809-821. Palaeorhiza Perkins P. S. 1963. classification of the Hirashima, Y. 1981a. A synopsis of the bee genus (Hy- Hurd, D., Jr, and J. Moure. A large carpen- menoptera, CoUetidae) of New Guinea, Part V. Subgenus ter bees (Xylocopini). University of California Publications in Ento- Cnemidorhiza. Esakia no. 17:1-48. mology 29:i-vi + 1-365. 70 Scientific Papers, Natural History Museum, The University of Kansas

Ihering, H. von. 1912. Zur Biologie der brasilianischen Meliponiden. LaBerge, W. E. 1956. A revision of the bees of the genus Melissodes in Zeitschrift fur Wissenschaftliche Insektenbiologie 8{l):l-5, (2):43-46. North and Central America, Part I. University of Kansas Science Bul- Ikudome, S. 1989. A revision ol^ the family CoUetidae of Japan. Bulletin of letin 37:911-1194. the Institute of Minami-kyushu Regional Science, Kagoshima LaBerge, W. E. 1957. The genera of bees of the tribe Eucerini in North and Women's Junior College no. 5:43-314. Central America. American Museum Novitates No. 1837:1^14. Ikudome, S. 1991. Two new subgenera of Hylaeiis. Japanese Journal of LaBerge, W. E. 1958. Notes on the genus Caesischia Michener, LaBerge Entomology 59:789-792. and Moure, with descriptions of a new species and subgenus from Ikudome, S. 1994. A list of the bee taxa of Japan and their Japanese names. Mexico. Pan-Pacific Entomologist 34:195-201. Bulletin of Kagoshima Women's Junior College no. 29:1-23. LaBerge, W. E. 1961. A revision of the bees of the genus Melissodes in Illiger, K. 1801. Namen der Insekten-Gattungen, ihr Genitiv, ihr North and Central America, Part III. University of Kansas Science grammatisches Geschlecht, ihr Silbenmass, ihre Herleifung; zugleich Bulletin 42:283-663. mit den Deutschen Benennungen. Magazin fur Insektenkunde 1:125- LaBerge, W. E. 1964. Prodromus of American bees of the genus Andrena. 155. Bulletin of the University of Nebraska State Museum 4:279-316. K. 1806. Illiger, William Kirby's Familien der Bienenartigen Insekten mit LaBerge, W. E. 1970. A new genus with three new species of eucerine Zusatzen, Nachweisungen und Bemerkungen. Magazin fiir bees from Mexico. Journal of the Kansas Entomological Society 43:321- Insektenkunde 5:28-175. 328.

K. 1 807. Fabr. Illiger, Vergleichung der Gattungen der Hautfliigler, Piezata LaBerge, W. E. 1971a. A new subgenus of Andrena found in California Hymenoptera Linn. Jur. Magazin fiir Insektenkunde 6:189-199. and Oregon. Pan-Pacific Entomologist 47:47-57. Inoue, T, and S. F. Sakagami. 1 993. A new name of Tngona. Japanese Jour- LaBerge, W. E. 1971b. A revision of the bees of the genus Andrena of the nal of Entomology 61:769. western hemisphere. Part IV. Scrapteropsis, Xiphandrena and Isensee, R. 1927. A study of the male genitalia of certain anthidiine bees. Rhaphandrena. Transactions of the American Entomological Society Annals of the Carnegie Museum 17:371-382, pi. xxxii. 97:441-520. Jorgensen, P. 1909. Beobachtungen iiber blumenbesuch, biologie, LaBerge, W. E. 1977. A revision of the bees of the genus Andrena of the Teil Teil II. VIII. verbreitung usw. der Bienen von Mendoza (Hym.), I, western hemisphere. Part Subgenus Thysandrena, Dasyandrena, Deutsche Zeitschrift 211-228. Rhacandrena , Entomologische 1909:53-65, Psammandrena , Euandrena, and Oxyandrena. Transactions Jorgensen, P. 1912. Revision der Apiden der Provinz Mendoza, Repub- of the American Entomological Society 103:1-143. lica Argentina. Zoologische Jahrbucher, Abteilung fiir Systematik, LaBerge, W. E. 1986. A revision of the bees of the genus Andrena of the Geographie und Biologie der Tiere 32:89-162. western hemisphere. Part XI. Minor subgenera and subgeneric key. Jurine, L. 1801. In [G. W. F. Panzer], Nachricht von einem neuen Transactions of the American Entomological Society 111:441-567. Prof. in P. D. entomologischen Werke, des Hrn. Jurine Geneve. LaBerge, W. E., and Hurd, Jr. 1965. A new subgenus and species of der 1:160-165. This matinal Andrena from the flowers of in Mexico. Intelligenzblatt Litteratur-Zeitung [Erlangen] Stcyos" (Cucurbitaceae) work was suppressed for nomenclatural purposes by Opinion 135 Pan-Pacific Entomologist 41:186-193. (Direction 4) of the Commission (1939). LaBerge, W. E., and C. D. Michener. 1963. Deltoptila, a Middle American Jurine, L. 1807. Nouvclle methodc de Classer les Hymenopieres et les Dipteres. genus of anthophorine bees. Bulletin of the University of Nebraska + 4 14 Geneva: Paschoud. 320 pp, pi. State Museum 4:211-226. Kerr, W. E., J. F. Pisani, and D. Ally. 1967. Aplicagao de principios LaBerge, W. E., and D. W. Ribble. 1972. A revision of the bees of the ge- a a modernos sistematica do genero Melipmim Illiger, com divisao em nus Andrena of the western hemisphere. Part V. Gonandrena, dois subgeneros. Papeis Avulsos de Zoologia, Sao Paulo 20:135-145. Ceissandrena, Parandrena, Pelicandreua. Transactions of the American Kirby, W. F. 1900. Xll. Descriptions of the new species of Hymenoptera, Entomological Society 98:271-358. in The to Sokotra. Bulletin of the 3:13- R. 1985. Expedition Liverpool Museums Labougle, J. M., and Ayala. A new subgenus and species of Bombus 24. (Hymenoptera: Apidae) from Guerrero, Mexico. Folia Entomologica Kirby, W. 1802. Monograpliia Apuni Aiigliac. Ipswich: White. Vol. 1. xxii + Mexicana no. 66:47-55. 2. + 258 pp. Vol. 388 pp 18 pis. Lanham, U. N. 1949. A subgeneric classification of the new world bees of W. 1837. in Kirby, Part IV, Insects, J. Richardson, Fauna Borcali-Amcricana; the genus Andrena. University of California Publications in Entomol- or the Zoology of the Northern Parts of British America. London: Longman. ogy 8: 1 83-238. + I-VIII. xxxix 325 "pp, pi. Lanham, U. N. 1950. A preoccupied subgeneric name in Andrena. Ento- Kirby, W., and W. Spence. 1826. An Introduction to Entomologx/. Vol. 3. Lon- mological News 61:140. cion: Longman. 732 pp. Latreille, P. A. 1802a. Histoire Naturelle des Fourmis.... Paris: Crapelet. xvi F. 1807a. + Klug, Oxaea, eine neue Gattung aus der Ordnung der Piezaten. 445 pp., 12 pis. Gesellschaft Naturforschender Freunde zu Berlin, Magazin fiir Latreille, P. A. 1 802b. Histoire Naturelle Generale et Particuliere des Crustaces neuesten Entdeckungen in der gesammten Naturkunde 1:261-263. et des Insectes. Vol. 3. Paris: Dufart. 467 pp. Klug, F. 1 807b. Kritische Revision der Bienengattimgen in Fabricius neuem Latreille, P. A. 1803. [Several articles in] Noveau Dictioiinaire d'Histoire Piezatensysteme. Magazin fiir Insektenkunde 6:200-228. Naturelle.... Vol. 18. Paris: Deterville. Klug, F. 1810. Einige neue Piezatengattungen. Gesellschaft Latreille, P. A. 1804. Tableau methodique des insectes. Pp. 129-200 in Naturforschender Freunde zu Berlin, Magazin fiir neuesten Noveau Dictionnaire d'Histoire Naturelle. Vol. 24. Paris: Deterville.

in 1. . . . . Entdeckungen der gesammten Naturkunde 4:31—15, pi. Latreille, P. A. 1809. Genera Crustaceorum et Insectorum Vol. 4. Paris: Krombein, K. V. 1950. The aculeate Hymenoptera of Micronesia II. Koenig. 399 pp.

CoUetidae, Halictidae, Megachilidae, and Apidae. Proceedings of the Latreille, P. A. 1810. Considerations Generates . . . des Insectes. Paris: Schoell. Hawaiian Entomological Society 14:101-142. 444 pp. Krombein, K. V. 1951. Additional notes on the bees of the Solomon Is- Latreille, P. A. 1811. [Articles in] M. Olivier (ed.), Encyclopedie Methodique. lands. Proceedings of the Hawaiian Entomological Society 14:277- Histoire Naturelle. Insectes. Vol. 8. Paris: Agasse.

295. Latreille, P. A. 1825. Families Naturelles du Regne Animal ... . Paris: Bailliere. Kriiger, E. 1917. Zur Systematik der mitteleuropaischen Hummeln 570 pp. (Hym.). Entomologische Mitteilungen 6:55-66. Latreille, P. A. 1829. Les Insectes, in G. C. L. D. Cuvier, Regne Animal. 2nd Kriiger, E. 1920. Beitrage zur Systematik und Morphologie der mittel- ed. Vol. 5. Paris: Deterville. xxiv + 556 pp. europaischen Hummeln. Zoologische Jahrbucher, Abteilung fiir Laverty, T. M., R. C. Plowright, and P. H. Williams. 1984. Digressobombus, Systematik, Geographie und Biologie der Tiere 42:289—464. a new subgenus with description of the male of Bombus digressus Kriiseman, G. 1952. Subgeneric division of the genus Bombus Latr. Trans- (Milliron). Canadian Entomologist 116:1051-1056. actions of the Ninth International Congress of Entomology, Leach, W. E. 1815. Entomology, pp. 57-172. In Brewster's Edinburgh Amsterdam 1:101-103. Encyclopaedia. Vol. 9. Edinburgh. Genus-Group Names of Bees 71

sur Vindobonae. 1-482. Lepeletier de Saint-Fargeau, A. L. M. 1832. Observations I'ouvrage ordinem alphabeticuni disposita. [i]-[vi], [Hy- intitule: Bombi Scandinaviae monographice tractato, et., a Gustav menoptera, pp. 254-273.) Dahlbom. Annales de la Societe Entomologique de France 1:366-382. Mavromoustakis, G. A. 1939. New and little-known bees of the subfam- lies Anthidiinae Part I. Annals and of Natural Lepeletier de Saint-Fargeau, A. L. M. 1836, 1841. Histoirc Naturelle ily (Apoidea). Magazine Imedes-Hymmoptires. Paris: Roret. 1836, vol. 1, 1-547; 1841, vol. 2, 1- History (11)3:88-97. 680. Mavromoustakis, G. A. 1949. On the bees (Hymenoptera, Apoidea) of Part I. and of Natural Lepeletier de Saint-Fargeau, A. L. M., and A. Serville. 1825, 1828. [Ar- Cyprus, Annals Magazine History ticles in] M. Diderot et al., Ena/clopedie Mctlwdiqtw, Histoire Naturelle. (12)l(1948):541-587. Inscctes. Vol. 10, P. A. Latreille, ed. Paris. Pp. 1-344, 1825; pp. 345-832, Mavromoustakis, G. A. 1951 . Further contributions to our knowledge of and their classi- 1828, according to C. D. Sherborn and B. B. Woodward, 1906, Annals the Ethiopian Anthidiinae (Hymenoptera: Apoidea) and Magazine of Natural History (7)17:578. fication. Annals and Magazine of Natural History (12)4:962-981. little-known bees of the Lieftinck, M. A. 1955. The carpenter-bees of the Lesser Sunda Islands Mavromoustakis, G. A. 1953. New and subfam- Anthidiinae Part VI. Annals and of Natural and Tanimbar (Hymenoptera, Apoidea). Verhandlungen der ily (Apoidea). Magazine Naturforschenden Gesellschaft in Basel 66:5-32. History (12)6:834-840. Lieftinck, M. A. 1966. Notes on some anthophorine bees, mainly from Mavromoustakis, G. A. 1954. New and interesting bees from Israel. Bul- of Israel 4:256-275. the old world. Tijdschrift voor Entomologie 109:125-161. letin of the Research Council of Lieftinck, M. A. 1968. A review of old world species of Thyreus Panzer (= Mavromoustakis, G. A. 1956. On the bees (Hymenoptera, Apoidea) Cwcisii Jurine) (Hym., Apoidea, Anthophoridae) Part 4, palearctic Siria. Eos 32:215-230. G. A. 1958. Bees of Attica species. Zoologische Verhandelingen no. 98:1-139, pis. 1^. Mavromoustakis, (Hymenoptera, Apoidea) Lieftinck, M. A. 1972. Further studies on old world melectine bees, with (Greece), Part I. Annals and Magazine of Natural History (13)1:433- 447. stray notes on their distribution and host relationships. Tijdschrift G. A. 1963a. the bees of voor Entomologie 115:253-324, 2 pis. Mavromoustakis, On (Hymenoptera, Apoidea) Lieftinck, M. A. 1974. A review of the central and east Asiatic Habropoda Lebanon, Part III. Annals and Magazine of Natural History (13)5:647- F. Smith, with Habwplwrula, a new genus from China. Tijdschrift voor 655. Entomologie 117:157-224. Mavromoustakis, G. A. 1963b. Further contributions to our knowledge Lieftinck, M. A. 1983. Notes on the nomenclature and synonymy of old of the Ethiopian Anthidiinae (Hymenoptera: Apoidea) and their 2. Natural world melectine and anthophorine bees. Tijdschrift voor Entomologie classification. Part Annals and Magazine of History 126:269-284. (13)6:481-499. bees. and Linnaeus, C. 1758. Si/stema Nahirae. Vol. 1, ed. 10. Stockholm: Salvii. 824 pp. Mavromoustakis, G. A. 1963c. A new genus of parasitic Annals of Natural Linsley, E. G. 1939a. A revision of the nearctic Melectinae. Annals of the Magazine History (13)5[1962]:443^145. bees. Annals and Entomological Society of America 32:429-468. Mavromoustakis, G. A. 1963d. On some parasitic Maga- zine of Linsley, E. G. 1939b. Some new genera and species of epeoline and Natural History (13)5[1962]:751-754. nomadine bees. Pan-Pacific Entomologist 15:1-11. Meade-Waldo, G. 1914a. Notes on the Hymenoptera in the collection of the British with of new Annals and Linsley, E. G. 1942. Notes and descriptions of some North American para- Museum, descriptions species. sitic bees. Pan-Pacific Entomologist 18:127-132. Magazine of Natural History (8)14:450-464. 1956. notes on the G. 1914b. Notes on the in the col- Linsley, E. G., and |. W. MacSwain. Further taxonomy Meade-Waldo, Apidae (Hymenoptera) of An- and biology of the andrenine bees associated with Ocnctliem. Pan- lection of the British Museum, with descriptions new species. Pacific Entomologist 32:111-121. nals and Magazine of Natural History (8)13:399^05. G. 1923. fam. subfam. Linsley, E. G., and C. D. Michener 1937. Some new genera and species of Meade-Waldo, Hymenoptera, Apidae, 1-45 P. Genera Insectorum. Ease. North American parasitic bees. Pan-Pacific Entomologist 13:75-84. Prosopidinae. Pp. m Wytsman, (ed.). 181. Brussels. Linsley, E. G., and C. D. Michener 1939. A generic revision of the North American Nomadidae. Transactions of the American Entomological Mehely, L. 1935. Naturgeschlchtc der Urbienen. Budapest: Stephaneum es 214 60 Society 65:265-305, pis. xv-xviii. Nyomda Konyvkiedo. pp., pis. sistematica F. 1888. [sic]. Naturalista Sicihano 7:152. Lucas de Oliveira, B., and J. S. Moure. 1963. Nova posi(;ao Meunier, Megachillidae D. 1938. American bees of the Heriades. Annals of the para Rlwphitulus tesfnceus Ducke, 1907. Anais da Academia Brasileira Michener, C. genus de Ciencias 35:575-584. Entomological Society of America 31:514-531. Lutz, E E., and T. D. A. Cockerell. 1920. Notes on the distribution and Michener, C. D. 1939. A revision of the genus Ashmeadiella. American bibliography of North American bees of the families Apidae, Midland Naturalist 22:1-84.

. of the American Osmiinae with Meliponidae, Bombidae, Euglossidae, and Anthophoridae. Bulletin Michener, C. D. 1941 A generic revision Natural- of the American Museum of Natural History 42:491-641. descriptive notes on old world genera. American Midland Ma, T.-c. 1= Maa]. 1938. The Indian species of the genus Xylocopm Latr. ist 26(1 ):147-166. bees with (Hymenoptera). Records of the Indian Museum 40:265-329. Michener, C. D. 1942. Taxonomic observations on descriptions of the Maa, T.-c. 1939. Xylocopn orientalia critica (Hymen.). I. Subgenus of new genera and species (Hymenoptera: Apoidea). Journal 50:273-282. Bomhoixylocopa >wvum. Lingnan Science Journal 18:155-160. New York Entomological Society D. 1943. bees of the with Maa, T.-c. 1953. An inquiry into the systematics of the tribus Apidini or Michener, C. The American genus Anthocopa Annals of the honeybees. Treubia 21:525-640. notes on old world subgenera. Entomological Society Maa, T.-c. 1954. The xylocopine bees (Insecta) of Afghanistan. of America 36:49-86. and Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening i Michener, C. D. 1944. Comparative external morphology, phylogeny, of Natu- Kjobenhavn 116:189-231. a classification of the bees. Bulletin of the American Museum XI. ral Manning, F. 1961 . A new fossil bee from Baltic amber Verhandlungen History 82:151-326. C. D. 1947. A revision of the American of Bul- Internationaler Kongress fiir Entomologie IWien, 1960] 1:306-308, pi. Michener, species Hoptitis. letin of the American Museum of Natural 89:257-318. 5,fig.l. History the anthidiine bees. Marikovskaya, T. P. 1976. On the systematics of the tribe Anthophorini. Michener, C. D. 1948a. The generic classification of no. 1381:1-29. Entomologicheskoe Obozrenie 55:684-690 [in Russian; translation into American Museum Novitates C. D. 1948b. Notes on the American bees of the Melecta. English, Entomological Review 55:126-130]. Michener, genus Marikovskaya, T. P. 1980. A new genus of bees of the family Proceedings of the Entomological Society of Washington 50:15-18. 1951. 1043-1255 m C. F W. Anthophoridae. Entomologicheskoe Obozrenie 59:650-653 [in Rus- Michener, C. D. Superfamily Apoidea. Pp. Muesebeck and K. V. Krombein America Nortli sian; translation into English, Entomological Review 59:126-130]. (eds.), Hymenoptera of States of Marschall, A. F von. 1873. Noiiicjiclntor zoologicus cnntineus nomina of Mexico-Synoptic Catalog. Washington; United Department secundum No. 2. systematica generum animnliuni tain viventitiin quant fossiliuni, Agriculture Monograph 72 Scientific Papers, Natural History Museum, The University of Kansas

Michener, C. D. 1952. A new genus of panurgine bee from Arizona. Jour- Michener, C. D., and R. W. Brooks. 1987. The family Melittidae in Mada- nal of the Kansas Entomological Society 25:24-28. gascar Annales de la Societe Entomologique de France (n.s.)23:99- Michener, C. D. 1954. Bees of Panama. Bulletin of the American Museum 103. of Natural History 104:1-176. Michener, C. D., R. W. Brooks, and A. Pauly 1990. Little-known Michener, C. D. 1 96 1 . A new parasitic genus of Ceratinini from Australia. meganomiine bees with a key to the genera. Journal of African Zool- Journal of the Kansas Entomological Society 34:178-180. ogy 104:135-140. Michener, C. D. 1962. Observations on the classification of the bees com- Michener, C. D., and T. L. Griswold. 1994a. The classification of old world monly placed in the genus Mcgacltile. Journal of the New York Ento- Anthidiini. University of Kansas Science Bulletin 55:299-327. mological Society 70:17-29. Michener, C. D., and T. L. Griswold. 1994b. The neotropical Sft'//s-like Michener, C. D. 1963. New Ceratinini from Australia. University of Kan- cleptoparasitic bees. Proceedings of the Entomological Society of sas Science Bulletin 44:257-261 . Washington 96:674-678. C. D. 1965a. classification of the bees of the Australian E. S. Michener, A and Michener, C. D., W. LaBerge, and J. Moure. 1955. Some American South Pacific regions. Bulletin of the American Museum of Natural Eucerini bees. Dusenia 6:213-230. 1-15. C. R. B. History 130:1-362, pis. Michener, D., J. McGinley and N. Danforth. 1994. The Bee Genera Michener, C. D. 1965b. A generic review of the Dufoureinae of the west- of North and Central America. Washington: Smithsonian Institution ern hemisphere. Annals of the Entomological Society of America Press, viii -i- 209 pp. 58:321-326. C. Michener, D., and J. S. Moure. 1956. The generic positions of certain Michener, C. D. 1966a. The classification of the Diphaglossinae and North South American eucerine bees. Dusenia 7:277-290. of the of Sci- S. American species genus Caiipolicana. University Kansas Michener, C. D., and J. Moure. 1957. A study of the classification of the ence Bulletin 46:717-751. more primitive non-parasitic anthophorine bees. Bulletin of the Michener, C. D. 1966b. A new genus of ceratinine bees from the Red Sea American Museum of Natural History 112:395—452. area. Journal of the Kansas Entomological Society 39:572-576. Michener, C. D., and E. Ordway. 1964. Some anthidiine bees from Mexico. Michener, C. D. 1968. Nests of some megachilid bees, with description of Journal of the New York Entomological Society 72:70-78. a new Hoplith. Journal of the Entomological Society of Southern Af- Michener, C. D., and G. Poinar 1997. The bee fauna of the Dominican rica 31:337-359. amber. Journal of the Kansas Entomological Society 69(1996 Michener, C. D. 1969. African genera of allodapine bees. Journal of the suppl.):353-361. Kansas Entomological Society 42:289-293. Michener, C. D., and R. R. Sokal. 1957. A quantitative approach to a prob- Michener, C. D. 1970. Social parasites among African allodapine bees. lem in classification. Evolution 11:130-162.

of . Zoological Journal the Linnean Society (London) 49:199-215, pis. Milliron, H. E. 1961 Revised classification of the bumblebees-A synop- 1,2. sis. Journal of the Kansas Entomological Societv 34:49-61. Michener, C. D. 1975. Larvae of African allodapine bees. 3. The genera Mitchell, T B. 1924. New megachilid bees. Journal of the Elisha Mitchell Allodapula and Eucondylops. Journal of the Entomological Society of Scientific Society 40:154-165. Southern Africa 38:243-250. Mitchell, T B. 1934. A revision of the genus MegachUe in the nearctic re- Michener, C. D. 1 977. Allodapine bees of Madagascar American Museum gion. Part 1. Transactions of the American Entomological Societv Novitates no. 2622:1-18. 59:295-361, pis. XX-XXl. Michener, C. D. 1978a. The classification of halictine bees: Tribes and old Mitchell, T B. 1936. A revision of the genus Megachile in the nearctic re- world nonparasitic genera with strong venation. University of Kan- gion. Part IV. Transactions of the American Entomological Societv sas Science Bulletin 51:501-538. 62:117-166, pis. Vlll-Xl. Michener, C. D. 1978b. The parasitic groups of Halictidae. University of Mitchell, T B. 1943. On the classification of neotropical Megachile. Annals Kansas Science Bulletin 51:291-339. of the Entomological Society of America 36:656-671. Michener, C. D. 1980. The large species of Hoiiialictus and related Mitchell, T B. 1973. A Snhgeneric Reinsion of the Bees of the Genus Coelioxi/s Halictinae from the New Guinea area. American Museum Novitates of the Western Hemisphere. Raleigh: Department of Entomology, North no. 2693:1-21. Carolina State University, iii -^ 129 pp. Michener, C. D. 1981. Classification of the bee family Mehttidae with a Mitchell, T B. 1980. A Generic Reinsion of the Megachiline Bees of the West- review of species of Meganomiinae. Contributions of the American ern Hemisplierc. Raleigh: Department of Entomology, North Carolina Entomological Institute 18(3):i-iii -> 1-135. State University, [iij -i- 95 pp. Michener, C. D. 1982. A new interpretation of fossil social bees from the Mocsary, A. 1894. E fauna apidarum hungariae. Termeszetrajzi Fiizetek Dominican Republic. Sociohiology 7:37^5. 17:34-37. Michener, C. D. 1983. The classification of the Lithurginae. Pan-Pacific Morawitz, F. 1873a. Die Bienen Daghestans. Horae Societatis Entomologist 59:176-187. Entomologicae Rossicae 10:129-189. Michener, C. D. 1986a. Family-group names among bees. Journal of the Morawitz, F 1873b. Nachtrag zur Bienenfauna des Gouvernements von Kansas Entomological Society 59:219-234. St. Petersburg. Horae Societatis Entomologicae Rossicae 9:151-159. Michener, C. D. 1986b. A review of the tribes Diphaglossini and Morawitz, F 1875. Mellifera Latr Pp. 1-160 in A. P. Fedtschenko, Reisen Dissoglottini. University of Kansas Science Bulletin 53:183-214. in Turkestan, Zoogeogrnpthisclie Folgc. Teil V, pt. I. Apidae Genuinae. Mos- Michener, C. D. 1986c. New Peruvian genus and a generic review of cow. Andreninae. Annals of the Entomological Society of America 79:62- Morawitz, F 1876. Zur Bienenfauna der Caucasuslander Horae Societa- 72. tis Entomologicae Rossicae 12:3-69. Michener, C. D. 1989. Classification of the American CoUetinae. Univer- Morawitz, F. 1895. Beitrag zur Bienenfauna Turkmeniens. Horae Societa- sity of Kansas Science Bulletin 53:622-703. tis Entomologicae Rossicae 29:1-76. C. D. 1990. Classification of the H. first Michener, Apidae. University of Kansas Morice, F D., and J. [Durrant. 1915. The authorship and publica- Science Bulletin 54:75-153. tion of the "Jurinean" genera of Hymenoptera.... Transactions of the Michener, C. D. 1992. Nomenclatural problems in the Meganonuinae and Entomological Society of London 1914:339^36. a review of the genus Pseudophilanthus. Journal of the Kansas Ento- Motschulskv, V. 1 856. Lettres . . . a M. Menetries, 3. Etudes Entomologiques mological Society 65:146-150. IHelsinkiJ 5:3-38, 1 pi. [Fossil bee, p. 28.] C. D. 1995. S. Michener, Some genus-group names of bees. Journal of the Moure, J. 1940. Apoidea neotropica. Arquivos de Zoologia do Estado Kansas Entomological Society 67(1994):373-377. de Sao Paulo 2:39-64, pis. l-Ill (also Revista do Museu Paulista 25:39- Michener, C. D. 1996. A review of the genera of Brachynomadini and a 64). of 111. new South American genus. Journal the Kansas Entomological Moure, J. S. 1941a. Apoidea neotropica. Arquivos do Museu Society 69:87-96. Paranaense 1:41-99, 1 pi. C. D. 1997. Tetralonia. S. Michener, The identity of the type species of Jour- Moure, J. 1941b. Notas sobre abelhas do grupo Tetrapedia Klug. Revista nal of the Kansas Entomological Society, in press. de Entomologia [Rio de Janeiro] 12:515-521. Genus-Group Names of Bees 73

S. 1942. Salobra. S. 1960. Notes on the of the bees described Moure, J. Abelhas de Papeis Avulsos do Departamento Moure, J. types neotropical by de Zoologia, Sao Paulo 2:291-321. Fabricius. Studia Entomologica 3:97-160. S. Batatais. S. 1961. A classification of the old Moure, J. 1943a. Abelhas de Arquivos do Museu Paranaense Moure, J. preliminary supra-specific world bees. Studia 4:181-242. 3:145-203, pi. XI. meliponine Entomologica S. 1964a. Os de com a de dois novos Moure, J. S. 1943b. Notas sobre abelhas da cole<;ao Zikan. Revista de Moure, J. especies Zikanapis, descrigao e duas novas. Studia 7:417^58. Entomologia [Rio de Janeiro] 14:447—184. subgeneros especies Entomologica halictine S. Peru. Boletin del Moure, J.S.I 964b. Two new of bees from the Araucanian Moure, J. 1944a. Abejas del Museo de Historia Natu- genera of South America. of the Kansas ral "Javier Prado" 8(28-29):67-75. subregion Journal Entomological 37:265-275. S. 1944b. A. Barbiellini. Society Moure, ]. Apoidea da coleqao do Conde Amadeu Moure, J. S. 1964c. Um novo Paracolletini do litoral Paulista. Boletim da Revista de Entomologia [Rio de Janeiro] 15:1-18. Universidade do Parana, 2:51-56. S. 1944c. Zoologia Moure, J. Abelhas de Monte Alegre (Est. Sao Paulo). Papeis for of Paranthidiwn. Moure, J. S. 1965. New placements some species Pro- Avulsos do Departamento de Zoologia [Sao Paulo] 6:103-126. ceedings of the Entomological Society of Washington 67:29-31. Moure, J. S. 1945a. Contribugao o conhecimento dos para Diphaglossinae, bees. Moure, J. S. 1967a. A check-list of the known euglossine Pp. 395- particularmente Ptikiglossn. Arquivos do Museu Paranaense 4:137- 415 in Atas do Simpdsio sobre n Biota Anmzonica, Vol. 5, Zoologia. 178, pi. XV. S. 1967b. of eucerine bees Moure, J. A new genus and two new species S. os Revista Moure, J. 1945b. Notas sobre Epicharitina. de Entomologia from South America (Hymenoptera, Apoidea). Dusenia 8:147-152. [Rio de Janeiro) 16:293-314. Moure, J. S. 1969. Micrommation. novo genero de Halictidae do Parana. Barbiellini. Moure, J. S. 1945c. Apoidea da coleqao do Conde Amadeu A. Atas da Sociedade de Biologia do Rio de Janeiro 12:247-249. 11. Revista de de 16:394-414. Entomologia [Rio Janeiro] sul Moure, J. S. 1972. Descrii;ao de duas especies novas de Hylaeus do do Moure, J. S. 1946a. Contribuigao o conhecimento dos para Meliponinae. Brasil. Revista Brasileira de Biologia 32:275-284. Revista de Entomologia [Rio de Janeiro] 17:437^143. Moure, J. S. 1975. Notas sobre as especies de Meliponn descritas por Moure, J. S. 1946b. do Brasil. Chacaras e Quintals 74:609-612. Meliponas Lepeletier em 1836 (Hymenoptera-Apidae). Revista Brasileira de Moure, S. 1946c. Notas sobre abelhas da sub familia Chilicolinae. J. Pp. Biologia 35:615-623. 243-250, Xll in Livro de a Roniucihio Fcnvira d' Almeida. pi. Homeimgcni Moure, J. S. 1989a. Camargoia, um novo genero neotropical de Sao Paulo: Sociedade Brasileira de Entomologia. Meliponinae. Boletim do Museu Paraense Emilio Goeldi, Serie S. 1946d. Notes sobre as Boletim 4(12- Moure, J. mamangabas. Agricola Zoologia 5:71-78. 1-3. S. n. e n. 13):l-32,pls. Moure, J. 1989b. Sakagamilla affabrn gen. sp. de Meliponinae (Hy- 1947a. e Moure, J. S. Novos agrupamentos genericos algumas especies menoptera, Apidae) de Rondonia. Revista Brasileira de Zoologia novas de abelhas sulamericanas. Museu Paranaense Publica(;6es 6:681-684. no. 3:1-37. S. 1989c. Avulsas, Moure, J. Glossuropoda, novo subgenero de Eiiglossn, e duas S. 1947b. la Salta. novas da do mesmo Memorias do Moure, J. Notas sobre algunas abejas de provincia de especies Amazonia, subgenero. Revista de la Sociedad Entomologica Argentina 13:218-253. Instituto Oswaldo Cruz 84:387-389.

S. 1948. S. 1992. Melikerria e , dois novos em Moure, J. Notas sobre algumas abelhas de Tacanas, Tuciunan, Moure, J. Eomelipona subgeneros 1806. 32-38 in Eucontro Brasileiro sobre de Argentina. Revista de Entomologia [Rio de Janeiro] 19:313-346. Melipona llliger, Pp. Biologia AbeUuis e Outros Inseetos Socinis. [Naturalia, edi(;ao Sao Paulo: Moure, J. S. 1949a. Notas sobre algunas abejas de Tacanas, Tucuman, especial.] Universidade Estadual Paulista. Argentina, 11. Revista de Entomologia [Rio de Janeiro] 20:437^60, Moure, J. S. 1994a. Sobre a sistematica de latttarsis pis. X, XI. posi<;ao Exomalopsis Friese. Brasileira de S. la Revista Zoologia 9(1992):273-276. Moure, J. 1949b. Las especies chilenas de sub-familia Lithurginae. S. 1994b. n. a Moure, J. Lissopedia, gen. de Paratetrapediini para regiao Arquivos do Museu Paranaense 7:265-286, pis. 11-V. com as de tres novas. Revista S. das neotropical, descri(;6es especies Moure, J. 1950a. Contribuigao para o conhecimento especies Brasileira de 9(1992):305-317. Brasileiras de Hypotngona Cockerell. Dusenia 1:241-260. Zoologia Moure, J. S. 1995. Compsockyta fasciata, antoforideo parasita do nordeste Moure, J. S. 1950b. Contribuiqao para o conhecimento do genero Eulaema da Argentina. Acta Biologica Paranaense 21:143^148. Lepeletier. Dusenia 1:181-200. F. Moure, J. S., and J. M. Camargo. 1978. A fossil stingless bee from Co- Moure, J. S. 1950c. Alguns agrupamentos novos de abelhas neotropicais. Journal of the Kansas Society 51:560-566. Dusenia 1:385-396. pal. Entomological Moure, J. S., and P. D. Hurd, Jr. 1960. On the systematic position of three S. Moure, J. 1951a. Notas sobre Meliponinae. Dusenia 2:25-70. anthidiine bees described by Adolfo Ducke. Boletim do Museu Halictidae. Moure, J. S. 1951b. Aripimuarthra, um novo genero de Dusenia Paraense Emilio Goeldi, Nova Serie, Zoologia no. 28:1-13. 2:137-140. P. of Moure, J. S., and D. Hurd, Jr. 1982. On two new groups neotropical S. 1951c. Notas sobre abelhas do Dusenia Moure, J. antigo genero Pn.si/j/wt'. hahcHne bees. Dusenia 13:46. 2:189-198. the Halictid Moure, J. S., and R D. Hurd, Jr. 1987. An Annotated Catalog of Moure, J. S. 1953a. Notas sobre CoUetidae sul-americanos. Dusenia 4:61- Bees of the Western Hemisphere. Washington: Smithsonian Institution 78. Press, vii -i- 405 pp. Moure, J. S. 1953b. , no de da B. Nogueirapis grupo Trigonini regiao Moure, J. S., and Lucas de Oliveira. 1962. Novo genero de Panurginae Ciencia e Cultura 5:247-249. neotropical. para a America do Sul. Boletim da Universidade do Parana, Zoologia Chile. Moure, J. S. 1953c. Notas sobre Megachilidae de Bolivia, Peru y no. 15:1-14. Dusenia 4:113-124. in Moure, J. S., and C. D. Michener 1955a. The bee family Fideliidae South S. Moure, J. 1954a. Novas notas sobre abelhas do antigo genero Pasiphae. America. Dusenia 6:199-206.

Dusenia 5:165-190. Moure, J. S., and C. D. Michener. 1 955b. A contribution toward the classi- S. 1954b. sul-americanos. Dusenia 5:259- Moure, J. Notas sobre Epeolini fication of neotropical Eucerini. Dusenia 6:239-331. 286. a Moure, J. S., and S. F. Sakagami. 1984. Notes on Pseudagapostemon, S. sulamericanos. Dusenia 6:115- of a Moure, J. 1955. Notas sobre Epeolini neotropical halictine bee genus, with descriptions new subgenus 138. and two new species from southern Brazil. Dusenia 14:1-9. Paracolletini sul Brasil. D. 1975. Moure, J. S. 1956. Tres especies novas de do do Moure, J. S., and Urban. Dicranthiduim novo subgenero de Dusenia 7:305-315. Hvpanthidioides [sic] Moure, 1947. Revista Brasileira de Biologia Brasil. Moure, J. S. 1958. Augochlorodes, a new genus of Halictinae from 35:837-842. D. 1990. Carlotieola n. e Carloticola trichina Journal of the Kansas Entomological Society 31:53-55. Moure, J. S., and Urban. gen. F. Smith n. da Acta Paranaense 19:89-99. Moure, J. S. 1959. On the species o( Megalopta described by (Hy- sp. Argentina. Biologica D. Urban. 1994. n. e notas sobre menoptera, Apoidea). Journal of the New York Entomological Soci- Moure, J. S., and Rhynostelis, gen. Revista Brasileira 11:297-302. ety 66[1958]:179-190. Hoplostelis Dominique. de Zoologie 74 SciENTinc Papers, Natural History Museum, The University of Kansas

Newman, E. 1834. division of British insects into natural or- 1970. Attempted Pasteels, J. J. Especes nouvelles de Megachilidae (Hymenoptera, ders. 2:379^31. Entomological Magazine Apoidea) d'Afrique Noire, I. Generes Creightonella Cockerel] et Noskiewicz, J. 1936. Die Piilearktischeii CoUetes-Artetu Lwowie: Prace Chaiicodoma Lepeletier. Revue de Zoologie et de Botanique Africaines Naukowe. 532 pp., 27 pis. 82:205-232.

Noskiewicz, J. 1 948. sur du 1972. Revision Remarques quelques especes genre Megachile Pasteels, J. J. des Anthidiinae (Hymenoptera Apoidea) de de la Silesie et des Polskie Pismo la pays limitrophes. Entomologiczne region indo-malaise. Bulletin et Annales cie la Societe Royale Beige 18:31-59 [in Polish, summary in French). d'Entomologie 108:72-128.

Noskiewicz, 1961 . zur Kenntnis der Arten 1977. Les J. Beitrage palaarktischen der Pasteels, J. J. Megachilini parasites (Coelioxys s. 1.) d'Afrique Gattung Stelis. Polskie Pismo Entomologiczne 31:113-133. noire. Revue de Zoologie Africaine 91:161-197. W. 1848. Adnotationes in 1982. Nylander, expositionem monographicam apum Pasteels, J. J. Les Coelioxt/s Latreille (Hymenoptera Apoidea) du borealium. Notiser ur Sallskapets pro Fauna et Flora Fennica groupe afra Lepeletier d'Europe et du proche-orient. Bulletin et III. Annales de la Forhandlinger 1:165-282, pi. Societe Royale Beige d'Entomologie 118:109-120. A. A. 1948. 1984. Revision Ogloblin, Descripcion de dos generos nuevos de ParacoUetini Pasteels, J. J. des Anthidiinae (Hymenoptera, Apoidea, del argentinos. Notas Museo de la Plata 13:165-177, pis. I-IV. Megachilidae) de I'Afrique subsaharienne. Memoires de la Classe des A. A. 1953. Ogloblin, Un nuevo Halictidae (Hym. Apodidae [sic]): un Sciences, Academie Royale de Belgique (IN^1-2)19(1):1-165. nuevo subgenero de Temnosonm. Boletin Sociedad Entomologica Ar- Patton, W. H. 1879a. Generic arrangement of the bees allied to Melissodes gentina no. 2:[2-3]. and Anthophora. Bulletin of the United States Geological and Geo- Ogloblin, A. A. 1955. Descripcion de un nuevo subgenero de graphical Sur\'ey of the Territories 5:471^79. Anthophorinae. Dusenia 6:231-237. Patton, W. H. 1879b. List of a collection of aculeate Hymenoptera made Mr. S. W. in Ogloblin, A. A. 1956. Dos generos nuevos de la tribu Exomalopsini by Williston northwestern Kansas. Bulletin of the United (Apidae, Hym.) de la Republica Argentina. Dusenia 7:149-158. States Geological and Geographical Survey of the Territories 5:349- 370. Osychnyuk, A. Z. 1983. A new subgenus and new species of bees of the A. 1980. genus Andrena (Hymenoptera, Andrenidae) from the asiatic part of Pauly, Descriptions preliminaires de quelque sous-genres nouveaux dans la famille the USSR. Entomologicheskoe Obozrenie 62:794-799 [in Russian; afrotropicaux des Halictidae. Revue de Africaine 94:119-125. translation into English, Entomological Review 62:124-129]. Zoologie A. 1981. (Lahrohaliclus) nouveau Osychnyuk, A. Z. 1984a. A new palearctic subgenus and a new species of Pauly, Lasioglossum saegeri, sous-genre et nouvelle de Halictidae du Pare la genus Andrena. Vestnik Zoologii 1984(2):23-30 [in Russian[. espece National de Garamba (Zaire). Revue de Osychnyuk, A. Z. 1984b. A new east-Mediterranean subgenus and a new Zoologie Africaine 95:717-720. A. 1984a. Glossodialictus unttei, nouveau et nouvelle species oi Andrenn bees. Vestnik Zoologii 1984(l):19-24 [in Russian]. Pauly, genre espece de Halictides recolte au Pare National de (Zaire). Revue de Packer, L. 1993. Two distinctive new species of halictine bees from high I'Upemba Africaine 98:703-708. altitude in the new world tropics. Canadian Journal of Zoology Zoologie 71:1653-1662. Pauly, A. 1984b. Classification des Halictidae de Madagascar et des iles voisines I. Halictinae. der Naturforschenden Pagliano, G., and P. Scaramozzino. 1990. Elenco dei generi di Hy- Verhandlungen Gesellschaft in Basel 94:121-156. menoptera del mondo. Memoria della Societii Entomologica Italiana A. 68 (1989):1-210. Pauly, 1984c. Paradialictus, un nouveau genre clepto-parasite recolte G. au Pare National des (Zaire). Revue de Africaine Panzer, W. F 1804a. Faunae Insectorum Germanicae. Heft 85. Niirnberg: Virungas Zoologie Felssecker. The 98:689-692. actual date of publication may have been as early as A. 1984d. Contribution a I'etude des de 1801; see C. D. Sherborn, 1923, Annals and Magazine of Natural His- Pauly, genres afrotropicaux Nomiinae. Revue de Africaine 98:693-702. tory (9)11:567 and Sandhouse, 1943:532. Zoologie Pauly, A. 1990. Classification des Nomiinae Africains. Annales Musee Panzer, G. W. F. 1804b. Faunae Insectorum Germanicae. Heft 86. Niimberg: de Central [Tervuren], Sciences no. 261 :1- Felssecker. According to Commission Opinion 151 (1944) the correct Royal I'Afrique Zoologiques 206. date for is this heft [1801-1802], but Sherborn (ref. supra) gives the A. 1991 . date as 1804. Pauly, Classification des Halictidae de Madagascar II. Nomiinae. Annales de la Societe de France (NS)27:287-321. Panzer, G. W. F. 1806. Kritische Rei'iswn der Insektenfauna Deutschlands. Entomologique Perez, 1890. des melliferes du sud-ouest. la Vol. 2. + J. Catalogue Actes de Societe Nurnberg: Felssecker. [14] 271 pp., 2 pls^ Linneenne de Bordeaux 44:133-192. Panzer, G. W. E 1809. Faunae Insectorum Germanicae. Heft 107. Niirnberg: 1897a. Felssecker. actual Perez, J. Quelques especes de megachiles nouvelles ou mal The date of publication mav have been as early as connues. Rendus des Seances, in Actes de la Societe 1806; see Panzer, 1804a. Comptes Linneenne de Bordeaux 52:lviii-lxvii. F and E. Parker, D., G. Bohart. 1979. Dolichostelis, a new genus of para- Perez, J. 1897b. Sur une forme nouvelle de buccal des sitic bees. I'appareil Journal of the Kansas Entomological Society 52:138-153. hymenopteres. Comptes Rendus de I'Academie des Sciences [Paris] Pasteels, J. J. 1965. Revision des Megachilidae (Hymenoptera Apoidea) 125:259-260. de I'Afrique Noire 1., Les Genres [sic], Chaiicodoma et Creightoniella 1901. Perez, J. Contribution a I'etude des xylocopes. Actes de la Societe Megachile (s. str.). Annales Musee de Central [Ter- Royal I'Afrique Linneenne de Bordeaux (6)6:1-128. vuren]. Sciences Zoologiques No. 137. ix + 579 pp. Perkins, R. C. L. 1899. Aculeata 1966. Hymenoptera [except ants]. Pp. 1-115, Pasteels, J. J. Megachilidae (genres: Creightoniella [sic], et Megnchde II ui R. C. L. pis. 1, Perkins and A. Forel, Hymenoptera Aculeata, i)i D. Chaiicodoma) peu connues ou nouvelles des regions palearctique et Fauna Haumiensis. Vol. 1 no. 1 . Sharp (ed.). , London: Cambridge Uni- africaine. Bulletin et Annales de la Societe Royale d'Entomologie de versity Press. Belgique 102:1-19. Perkins, R. C. L. 1908. Some remarkable Austrahan Hymenoptera. Pro- Pasteels, J. J. 1968a. Statut, affinites et des Anthidiinae origines parasites. of the Hawaiian 2:27-35. Naturaliste Canadien 95:1055-1063. ceedings Entomological Society Perkins, R. C. L. 1912. Notes, with of new on ac- 1968b. descriptions species, Pasteels, J. J. Revision des Megachilidae (Hymenoptera Apoidea) uleate Hymenoptera of the Australian Region. Annals and Magazine de I'Afrique Noire II. Le genre Annales Musee de Coelioxys. Royal of Natural History (8)9:96-121. I'Afrique Central [Tervuren], Sciences No. 167. [iij -t- 139 Zoologiques Perkins, R. C. L., and L. E. Cheesman. 1928. Part V, Hymenoptera, Fasc. PP- 1, Apoidea, Sphecoidea, and Vespoidea. Pp. 1-31 in bisects of Samoa Pasteels, J. J. 1969a. La et des systematique generique subgenerique and other Samoan terrestrial Arthropoda. London: British Museum (Natu- Anthidiinae de I'ancien (Hymenoptera, Apoidea, Megachilidae) ral History). monde. Memoires de la Societe Rovale de A. Fasc. d'Entomologie Belgique Perty, J. M.'l833. 3, pp. 125-224, pi. 25-40, in Delectus Animalium 31:1-148. Articulatorum ... Brasdiam. Miinchen. Pasteels, 1969b. New Anthidiinae J. J. (Hymenoptera, Apoidea, Pesenko, Y. A. 1983. Tribe Nomioidini (in the palearctic fauna). Fauna of from the Mediterranean area and from the East. Megachilidae) Near the USSR, Insecta-Hymenoptera 17(1): 1-198 [in Russian]. Leningrad: Israel Journal of Entomology 4:409-434. Nauka. Genus-Group Names of Bees 75

Pesenko, Y. A. 1984. A subgeneric classification of bees of the genus Popov, V. B. 1951. The parasitic bee genus Ammohates Latr I. Tribes Halictui Latreille st'(;S!( stncto. Entomologicheskoe Obozrenie 63:340- Ammobatini and Pasitini, their size and taxonomic position. Trudy 357 |in Russian, English summary; English translation in Entomo- Instituta Zoologii, Akademii Nauk SSSR 9:895-949 [in Russian]. logical Review 63(3):l-20]. Popov, V. B. 1952. Bee fauna (Hymenoptera, Apoidea) of southwestern Pesenko, Y. A. 1986a. An annotated key to females of the palaearctic spe- Turkmenii and its ecological significance. Trudv Zoologicheskova cies of the genus Lashghssum sensii stricto (Hymenoptera, Halictidae), Instituta, Akademii Nauk SSSR 10:61-117 [in Russian]. with descriptions of new subgenera and species. Pp. 113-151 in Y. A. Popov, V. B. 1955a. A new subgenus of bees of the genus Heriades. Pesenko (ed.), Systematics of Hymenopterous Insects. Trudy Entomologicheskoe Obozrenie 34:280-286 [in Russian]. Zoologicheskova Instituta, Akademii Nauk SSSR 159 |in Russian]. Popov, V. B. 1955b. On the parasitic genus Radoszkozcskiana (Hymenoptera, Pesenko, Y. A. 1986b. Systematics of the bees of the genus Halictus Latreille Megachilidae) and its origins. Zoologicheskii Zhurnal 34:547-556 [in (Hymenoptera, Halictidae) with description of the 7th and 8th Russian]. metasomal sterna of males: Subgenus T\/tthalictus Pesenko. Popov, V. B. 1955c. Generic groupings of the palearctic Melectinae. Trudy Entomologicheskoe Obozrenie 65:618-632 [in Russian). Zoologicheskova Instituta, Akademii Nauk SSSR 21 :321-334 ]in Rus- Pesenko, Y. A., and 1. M. Kerzhner 1981. Nomioides Schenck, 1866: Pro- sian]. posed designation of type species. Bulletin of Zoological Nomencla- Popov, V. B. 1956. New and little-known bees from central Asia. ture 38:225-227. Entomologicheskoe Obozrenie 35:159-171 [in Russian]. Pesenko, Y. A., and A. A. Sitdikov. 1988. Classification and phylogenetic Popov, V. B. 1957. On the genera Moraiuitzella, gen. nov., and Trilia Vach. relations of the genera of the tribe Eucerini (Hymenoptera, Entomologicheskoe Obozrenie 36:916-924 [in Russian[. Anthophoridae) with two submarginal cells. Entomologicheskoe Popov, V. B. 1958. On three subgenera of the genus Andrena. Trudy Obozrenie 67:846-860 [in Russian; English translation in Entomologi- Vsesoyuznovo Entomologicheskovo Obshchestva (Horae Societatis cal Review 69:88-104 (1990)]. Entomologicae Unionis Soveticae) 46:109-161 [in Russian]. Peters, D. S. 1970. Pseudoheriades n. gen., Afroheriades n. subgen., Popov, V. B. 1962. New genera of bees from middle Asia. Trudv Instituta Pseudoheriades primus n. sp., neue Formen aus der Familie Zoologii, Akademii Nauk SSSR 30:291-309 [in Russian[. Megachilidae. Entomologische Zeitschrift 80(16):153-160. Popov, V. B. 1963. A new genus of Megachilidae (Hymenoptera) from Peters, D. C. 1972. Drei neue Arten der Megachilidae aus Mexico. central Asia. Zoologicheskii Zhurnal 42:865-869 [in Russian, English Senckenbergiana Biologica 53:373-382. summary]. Peters, D. S. 1978a. Arclwruuies gen n., eine verhaltnismassig urspriingliche Popov, V. B. 1964. On the bee genera Traclnisn Panzer and Trachusomimus, Gattung der Megachilidae. Entomologica Germanica 4:337-343. gen. n. Entomologicheskoe Obozrenie 43:405-417 [in Russian; English Peters, D. S. 1978b. Systematik und Zoogeographie der west- translation. Entomological Review 43:207-2141. palaarktischen Arten von Osmin Panzer 1806 s. str, Monosinia Tkalc Popov, V. B., and D. Guiglia. 1936. Note sopra i gen. Ctenoplectra Sm. e 1974 und Orientosmia n. subgen. Senckenbergiana Biologica 58:287- Macropis Panz. Annali del Museo Civico di Storia Naturale di Genova 346. 59:275-288. Peters, D. S. 1984. Revision der von H. Brauns beschriebenen oder Provancher, L. 1882. Faune Canadienne. Les Insectes Hvmenopteres. behandeltenafrikanischenHcrwrft's- und OsHim-Arten. Entomofauna NaturalisteCanadien 13:225-242. 5:359-387. Provancher, L. 1885-1889. Additions et Corrections an Volume II de la Faune Philippi, R. A. 1866. Einige Insekten von Chile. Stettiner Entomologische Entomologitpie du Canada Traitant des Hymenopteres. Quebec: Darveau. Zeitung 27:109-117, Taf. II. Pp. 1-475. [The parts on bees were published in 1888.] Piton, L.-E. 1940. Pnleontohgie dii Gisement Eocene de Menat (Puy-de-Dome) Quilis P., M. 1927. Los apidos de Espana. Genero Bombus Latr. Trabajos (Flore el Faiine). Clermont-Ferrand: Vallier (thesis); Paris: Lechevalier. del Laboratorio de Historia Natural de Valencia 16:1-119, 10 pi. vii + 303 pp. Radoszkowski, O. 1865. Tribu des Melectides. Horae Societatis Pittioni, B. 1939. Tanguticobombiis subg. nov. (Hymenopt., Apidae). Entomologicae Rossicae 3:53-60. Zoologischer Anzeiger 126:201-205. Radoszkowski, O. 1868. Materiaux pour servir a I'etude des insectes de Pittioni, B. 1948. Beitrage zur Kenntnis Italienischer Bienen. I. Uber einige la Russie IV. Notes sur quelques Hymenopteres de la tribu des Apides. Arten der Gattung Andrena Fabr Bollettino dell'Istituto de Horae Societatis Entomologicae Rossicae 5:73-90, pi. III. Entomologia della Universita di Bologna 17:46-61. Radoszkowski, O. 1872a. hi Seance du 3 (15) Mai. Horae Societatis Pittioni, B. 1949. Beitrage zur Kenntnis der Bienenfauna SO-Chinas. Eos Entomologicae Rossicae 8:Bulletin pp. xv-xviii. 25:241-284. Radoszkowski, 0. 1872b. Supplement indispensable a I'article publie par Pittioni, B. 1950. Die westpalaarktischen Ajten der Gattung Spatukriella M. Gerstaecker en 1869, sur quelques genres d'hymenopteres. Bulle- tin 1. Pop. Bollettino della Societa Veneziana di Storia Naturale 5:76-113. de la Societe Imperiale des Naturalistes de Moscou 45:1-40, pi. Popov, V. B. 1931a. Zur Kenntnis der palaarktischen Schmarotzer- Radoszkowski, 0. 1874a. Supplement indispensable a I'article publie par hummeln (Psitltyrus Lep.). Eos 7:131-209. M. Gerstaecker en 1869, sur quelques genres d'hymenopteres. Bulle- Popov, V. B. 1931b. To the knowledge of the genera Pasites Jurine and tin de la Societe Imperiale des Naturalistes de Moscou 46(3)(1873):133- 1 vol. Paramniobatodes gen. nov. Ezhegocinik Zoologicheskovo Muzeya 151, pi. [This issue was misnumbered 47, which otherwise does Akademii Nauk SSSR (Annuaire du Musee Zoologique de 1' Academie not exist according to D. B. Baker, in lift., 1992; he also provided the des Sciences de I'URSS) 32:453^67 [in Russian, English summary]. actual year of publication as 1874.] Popov, V. B. 1938. Notiz iiber die Gattung Chelynici Prov. und einige Radoszkowski, 0. 1874b. Supplement indispensable a I'article publie par Untergattungsgruppierungen der Gattung Stelis Panz. Konowia M. Gerstaecker in 1869, sur quelques genres d'hymenopteres. Bulle- 17:36-41. tin de la Societe Imperiale des Naturalistes de Moscou 48(1 )( 1873): 132- Popov, V. B. 1939. Subgeneric groupings of genus Prosopis F. Comptes 164. Rendus (Doklady) de I'Academie des Sciences de I'URSS 25:167-170. Radoszkowski, 0. 1884. (Quelques nouveaux Hymenopteres d'Amerique. V. B. 1 Popov, 1940. A new genus of bees from Turkestan. Trudy Horae Societatis Entomologicae Rossicae 18:17-22, pi. Zoologicheskova Instituta, Akademii Nauk SSSR 6:53-60 [in Russian|. Radoszkowski, O. 1886. Faune Hymenopterologique Transcaspienne. V. B. 1947. of the 1-11. Popov, Generic groupings subfamily Dioxynae [sic|. Horae Societatis Entomologicae Rossicae 20:3-56, pis.

Entomologicheskoe Obozrenie 29:84-92 [in Russian, English sum- Radoszkowski, 0. 1891 . Revision des armures copula trices des males des mary]. genres Cilissa et PseudocUi^sa. Horae Societatis Entomologicae Rossicae Popov, V. B. 1950a. Generic groupings of the mid-Asian bees of the sub- 25:236-243. family Anthidiinae. Doklady Akademii Nauk SSSR 70:315-318 [in Radoszkowski, O. 1893. Fauna Hymenopterologique Transcaspienne. Russian]. Horae Societatis Entomologicae Rossicae 27:38-81. Popov, V. B. 1950b. Concerning the genus Amegilla Friese. Rafinesque, C. S. 1814. Principtes Fondanientaux de Soiniologie.... Palermo. Entomologicheskoe Obozrenie 31:257-261 [in Russian]. 52 pp. 76 Scientific Papers, Natural History Museum, The University of Kansas

C. S. 1815. dc In Nature... Palermo. 224 Rafinesque, Analyse pp. Robertson, C. 1903b. Synopsis of Sphecodinae. Entomological News Rayment, T. 1930a. Microi;lossii and Melilrihus, new genera of Australian 14:103-107. bees. of the of Victoria Proceedings Royal Society (n.s.)42:211-220, pi. Robertson, C. 1903c. Synopsis of Epeolinae. Canadian Entomologist 21. 35:284-288. T. 1930b. New and remarkable bees. of the Rayment, Proceedings Royal Robertson, C. 1903d. Synopsis of Nomadinae. Canadian Entomologist Society of Victoria (NS)43:42-61 . 35:172-179. T 1931. Bees in the collections of the Western Australian Mu- Rayment, Robertson, C. 1904. Synopsis of Anthophila. Canadian Entomologist seum and the Agricultural Department, Perth. Journal of the Royal 36:37-43. of Western Australia 17:157-190. Society Robertson, C. 1905. Synopsis of Euceridae, Emphoridae and Rayment, T. 1935. A Cluster of Bees. Endeavour 752 Sydney: pp. Anthophoridae. Transactions of the American Entomological Soci- T. 1950. New bees and Part XIII. Victorian Naturalist Rayment, wasps. ety 31:365-372. 67:20-25. Robertson, C. 1918a. Some genera of bees. Entomological News 29:91- T. 1954. Remarkable bees f the rain forest. Rayment, rom Australian Zo- 92. ologist 12:46-56. Robertson, C. 1918b. Erroneous generic determinations of bees. Science T. 1959a. A new of bees in the Colletidae. Austra- Rayment, genus family (n.s.)48:368-369. lian Zoologist 12:324-329, pi. 38. Robertson, C. 1922. Synopsis of Panurgidae. Psyche 29:159-173. Rayment, T. 1959b. A new and remarkable colletid bee. Australian Zo- Rodeck, H. G. 1945. Two new subgenera of Nomflda Scopoli. 12:334-336, 11. Entomologi- ologist pi. cal News 56:179-181. Rebmann, O. 1967. Beitrag zur Kenntnis der Gattimg Megachile Latr Rodeck, H. G. 1947. Laminomada, a new subgenus of Notimda. Annals of (Hym., Apidae): Subgenus Eutriclmraea Thorns, und Neoeutriclmraen the Entomological Society of America 40:266-270. nov. subg. Entomologische Zeitschrift 77:33-38. Rohwer, S. A. 1911 . A new of nomadine bees. News Rebmann, O. 1970. 6. zur Kenntnis der Latr genus Entomological Beitrag Gattung Megnchile 22:24-27. Nachrichtenblatt der Bayerischen Entomologen 19:37-47. Roig-Alsina, A. 1987. The classification of the Journal Reed, E. C. 1892. Revision de las abejas chilenas descritas en la obra de Caenoprosopidini. of the Kansas Entomological Society 60:305-315. Gay. Actes de la Societe Scientifique du Chili (Santiago de Chile) 2:223- A. 1989. The tribe its 239. Roig-Alsina, Osirini, scope, classification, and revisions of the and Osirinus. of Kansas Sci- Reed, E. C. 1894. Entomolojia [sic] Chilena. Anales de la Universidad genera Parepeclus University ence Bulletin 54:1-23. [Santiago, Chile] 85:599-653. A. 1991. Cladistic of the Nomadinae s. str with Reinig, W. F. 1930. Untersuchungen zur Kenntnis der Hummelfauna des Roig-Alsina, analysis of a new Journal of the Kansas So- Pamir-Hochlandes. Zeitschrift fiir Morphologic und Okologie der description genus. Entomological Tiere 17:68-123. ciety 64:23-37. A. 1994. a new of and Ribble, D. W. 1968a. Revisions of two subgenera of Amireua: Micrninlreuii Roig-Alsina, Meliphilopsis, genus emphorine bees, Ashmead and Derandrcnn notes on the the of , new subgenus. Bulletin of the University relationships among genera Emphorina. of Nebraska State Museum 8:237-394. Reichenbachia 30:181-188. A., and C. D. Michener 1993. Studies of the and Ribble, D. W. 1968b. A new subgenus, Belaudreun, of the genus Andreua. Roig-Alsina, phylogeny classification of bees. of Kansas Science Journal of the Kansas Entomological Society 41:220-236. long-tongued University Richards, O. W. 1929. A revision of the humble-bees allied to Bcmbus Bulletin 55:124-162. B. de. 1840. Sur orientalis Smith, with the description of a new subgenus. Annals and Romand, I'Hymenoptere nomme Acanthopus Goryi (voy. cette Magazine of Natural History (10)3:378-386. Revue, 1840, p. 248). Revue Zoologique par la Societe Richards, O. W. 1935. Notes on the nomenclature of the aculeate Hy- Cuvierienne 1840:335-336. B. de. 1841. Notice sur divers insectes de la menoptera, with special reference to the British genera and species. Romand, Hymenopteres famille des melliferes. de 68-70. Transactions of the Royal Entomological Society of London 83:143- Magazin Zoologie (2)3:1-8, pis. 176. Romankova, T G. 1988. A new far-eastern bee of the tribe Anthidiini. Richards, O. W. 1968. The subgeneric divisions of the genus Bombus Vestnik Zoologii 1988(4):25-30. Latreille. D. A. L. and M. F. de 1997. Bulletin of the British Museum (Natural History), Entomol- Roubik, W., J. Segura, J. Camargo. New sting- ogy 22:211-276. less bee genus endemic to Central American cloud forests: Phyloge- Ritsema, C. 1873. Beschrijving van een nieuw Hymenopteren-genus uit netic and biogeographical implications. Systematic Entomology de onder-familie der Andrenidae Acutilingues. Tijdschrift voor 21:101-114. 1958. Entomologie 16:224-228, pi. 10. Rozen, J. G., ]r. Monographic study of the genus Nontadopsis Ritsema, C. 1874. [Zodion, Hymenoptera, etc.]. Tijdschrift voor Ashmead. University of California Publications in Entomology 15:i- Entomologie 17:lxviii-lxxv. iv + 1-202. R. R. 1992. Roberts, B., and W. Brooks. 1987. Agapostemonine bees of Rozen, J. G., Jr. Systematics and host relationships of the cuckoo Mesoamerica (Hymenoptera: Halictidae). University of Kansas Sci- bee genus Oreopnsites. American Museum Novitates no. 3046:1-56. ence Bulletin 53:357-392. Rozen, J. G., Jr, and L. Ruz. 1995. South American panurgine bees C. — Robertson, 1897. North American bees Descriptions and synonyms. (Andrenidae: Panurginae), Part II. Adults, immature stages, and bi- Transactions of the Academy of Science of St. Louis 7:315-356. ology of Neffnpis longiliuguu, a new genus and species with an elon- Robertson, C. 1900. Some Illinois bees. Transactions of the Academy of gate glossa. American Museum Novitates no. 3136:1-15. Science of St. Louis 10:47-55. Ruz, L. 1980. Pseiidosnrus, nuevo genero de Panurginae Chileno. Revista Robertson, C. 1 901 . Some new or little-known bees. Canadian Entomolo- Chilena de Entomologia 10:25-28. gist 33:229-231. Ruz, L. 1991. Classification and phylogenetic relationships of the Robertson, C. 1902a. Synopsis of Andreninae. Transactions of the Ameri- panurgine bees: The Calliopsini and allies. University of Kansas Sci- can Entomological Society 28:187-194. ence Bulletin 54:209-256. C. 1902b. or little-known IV. 1993. Robertson, Some new bees, Canadian Ento- Ruz, L., and J. G. Rozen, Jr. South American panurgine bees 34:321-331. mologist (Apoidea: Andrenidae: Panurginae), Part 1. Biology, mature larva, and C. 1902c. of Robertson, Synopsis Halictinae. Canadian Entomologist description of a new genus and species. American Museum Novitates 34:243-250. no. 3057:1-12. C. 1902d. — Robertson, Some new or little-known bees. II. Canadian En- Ruz, L., and H. Toro. 1983. Revision of the bee genus Liphnnthus. Univer- tomologist 34:48-49. sity of Kansas Science Bulletin 52:235-299. C. 1903a. Robertson, Synopsis of Megachilidae and Bombinae. Transac- Sakagami, S. F 1975. Stingless bees (excl. Tetnii^ouuhi) from the continen- tions of the American Entomological Society 29:163-178. tal southeast Asia in the collection of the Bernice P. Bishop Museum, Genus-Group Names of Bees 77

Honolulu, journal of the Faculty of Science, Hokkaido University, Schulz, W. A. 1911 . Zweihundert alte Hymenoptera. Zoologische Annalen, ser. VI, Zoology 20:49-76. Wiirzburg 4:1-220. Iran. Sakagami, S. F. 1978. Tetragoniila stingless bees of the continental Asia Schwammberger, K.-H. 1971. Zwei neue Bienen-Arten aus and Sri Lanka. Journal of the Faculty of Science, Hokkaido Univer- Stuttgarter Beitrage zur Naturkunde no. 255:1-4. sity ser. VI, Zoology 21:165-247. Schwarz, H. F 1938. The stingless bees (Meliponidae) of British Guiana Sakagami, S. F, and C. D. Michener. 1987. Tribes of Xylocopinae and ori- and some related forms. Bulletin of the American Museum of Natu- Annals of the of America gin of the Apidae. Entomological Society ral History 74:437-508, pi. LlI-LXIl. 80:439^30. Schwarz, H. F 1939a. The Indo-Malayan species of Trigona. Bulletin of Sakagami, S. F, and K. Yoshikawa. 1961. Bees of Xylocopinae and Apinae the American Museum of Natural History 76:83-141. collected by the Osaka City University biological expedition to south- Schwarz, H. F. 1939b. A substitute name for Patera Schwarz. Entomologi- east Asia 1957-58, with some biological notes. Nature and Life in cal News 50:23. Southeast Asia [Tokyo] 1:409-444. Schwarz, H. F. 1940. Additional species and records of stingless bees Sandhouse, G. A. 1937. The bees of the genera Augochlora, Augochloropsis (Meliponidae) from British Guiana based on specimens collected by in the United and Atigocldorella (Hymenoptera; Apoidea) occurring the Terry-Holden Expedition. American Museum Novitates no. States. Journal of the Washington Academy of Sciences 27:65-79. 1078:1-12. G. A. 1939. The North American bees of the Osmia. Sandhouse, genus Schwarz, H. F. 1948. Stingless bees (Meliponidae) of the western hemi- of the of + 1-167. Memoirs Entomological Society Washington l:[ii] sphere. Bulletin of the American Museum of Natural History 90:i- G. A. 1943. The of the and of Sandhouse, type species genera subgenera xvii -I- 1-546. bees. of the Uniteci States National Museum 92:519-619. Proceedings Schwarz, M. 1966. Beitrag zur Subfamilie Nomadinae. Polskie Pismo H. de. 1890. Histoire Naturelle ties in A. Saussure, H\/intiioptcies Grandidier, Entomologiczne 36:383-394. Histoire NaturcUc et dc Vol. 20. Paris: 1770. Anniii^ Historico Natitralis. Annus IV. Hilscher. Phi/siquc, Politique Madagnscar. ScopoH, J. A. Lipsiae: Nationale. xxi + 590 27 Imprimerie pp., pis. 150 pp., 2 pi. A. 1861. Die Nassauischen Bienen. des Vereins fiir Schenck, Jahrbiicher Shinn, A. F. 1967. A revision of the bee genus Calhopsis and the biology Naturkunde im Nassau 14:1^14 [for date, see Michener, Herzogthum and ecology of C. andreniformis. University of Kansas Science Bulle- 1986a]. tin 46:753-936. Schenck, A. 1 866. Verzeichniss der naussauischen aculeata Hymenoptera Shuckard, W. E. 1840. JBees]. Pp. 158-170 in W. Swainson and W. E. mit der deutschen Arten. Berliner Hinzufijgung iibrigen Shuckard, On the History and Natural Arrangement of Insects. London. Zeitschrift 10:317-369. in Reise der Osterreichischen Entomologische Sichel, J. 1867. Hymenoptera mellifera, Schenck, A. 1869. der nassauischen Bienen, Zweiter Beschreibung Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859..., des Nassauischen Vereins fiir Naturkunde 21- Nachtrag. Jahrbiicher Zoologischer Theil 2:143-156. 22:1[269]-114[382]. dated 1868, 1869, 1-114; [Preprint, published pp. Silveira, F. A. 1995. Phylogenetic relationships and classification of Jahrbiicher for 1867-68, in 1870, 269-382. For dates of published pp. Exomalopsini with a new tribe Teratognathini. University of Kansas Schenck's works, see Michener, 1986a.] Science Bulletin 55:425^54. Schenck, A. 1873. Ueber und zweifelhafte Bienen-Arten. einige streitige Sinha, R. N. 1958. A subgeneric revision of the genus Osmia in the west- Berliner Zeitschrift 17:243-259. Entomologische ern hemisphere. University of Kansas Science Bulletin 39:211-261. Schmiedeknecht, H. L. O. 1885-1886. (Die Bienen Apidne Eiiwpaene Sitdikov, A. A., and Y. A. Pesenko. 1988. A subgeneric classification of Vol. 2. Berlin: Friedlander. [iv] + 1-205 1867-1071] + [2, in- Europa's}. bees of the genus Eucera Scopoli (Hymenoptera, Anthophoridae) with 111-205 in dex], pis. 16, 17. [Pp. 1-110 published in 1885, 1886.] a scheme of the phylogenetic relationships between the subgenera. Schrottky, C. 1901. Biologische Notizen solitarer Bienen von S. Paulo Trudy Zoologicheskii Institut, Akademiia Nauk SSSR 175:75-101 Jin (Brasilien). Zeitschrift fiir 6:209-216. AUgemeine Entomologie Russian]. C. 1902a. Ensaio sobre as abelhas solitarias do Brazil. Revista Schrottky, Skorikov, A. S. 1914a. Les formes nouvelles des bourdons. Russkoe do Museo Paulista 5:330-613, Xll-XlV. pis. Entomologicheskoe Obozrenie 14:119-129 [in Russian]. Schrottky, C. 1902b. Hymenopteres nouveau de I'Americ^ue Meridionale. Skorikov, A. S. 1914b. Contribution a la faune des bourdons de la partie Anales del Museo Nacional de Buenos Aires 7:309-315. meridionale de la province maritime. Russkoe Entomologicheskoe C. 1906a. Neue und bekannte siidamerikanische Bienen. Schrottky, wenig Obozrenie 14:398^07 [in Russian]. Zeitschrift fiir systematische Hymenopterologie und Dipterologie Skorikov, A. S. 1922a. The bumblebees of the Petrograd district. 6:305-316. Petrogradskii Agronomicheskii Institut, Entomologicheskaya Schrottky, C. 1906b. Zur Synonymie der Apiden. Zeitschrift fiir Stantsiya. Fauna Petrogradskoi Gubernii 2(c)no. 11:1-51 lin Russian]. systematische Hymenopterologie und Dipterologie 6:115-118. Skorikov, A. S. 1922b. Les Bourdons de la faune palearctique, Partie 1, Schrottky, C. 1909a. Nuevos himenopteros sudamericanos. Revista del Severnoi Oblastnoi Stantsii Zashchity Museo de La Plata 16:137-149. Biologie generale. Izvestiya Rastnii ot Vreditelei (Bulletin de la Station Regionale Protectrice des Schrottky, C. 1909b. Synonymische Bemerkungen iiber einige Plantes a Petrograd) 4:1-160, 15 maps. siidamerikanische Halictinae. Deutsche Entomologische Zeitschrift A. S. 1933a. Zur Hummelfauna und seiner 1909:479^85. Skorikov, Japans Nachbarlandes. Mushi 6:53-65, 2 fig. Schrottky, C. 1909c. Himenopteros de Catamarca. Anales de la Sociedad Skorikov, A. S. 1933b. Zur Fauna und Zoogeographie der Hummeln des Cientifica Argentina 68:233-272. Himalaya. Doklady Akademii Nauk SSSR 5:243-248. Schrottky, C. 1909d. Hymenoptera nova. Anales de la Sociedad Cientifica Skorikov, A. S. 1937. Die Hummeln im Aspekte der Argentina 67:209-228. gronlandischen Meddelelser 20:37-64. Schrottky, C. 1910. Berichtigung (Hym.). Deutsche Entomologische Zirkumpolarfauna. Entomologiske of the fauna Zeitschrift 1910:540. Skorikov, A. S. 1938a. Zoogeographie uniformity bumblebee of the Causasus, Iran and Anatolia. Obozrenie Schrottky, C. 1911. Descripqao de abelhas novas do Brazil e de regioes Entomologicheskoe visinhas. Revista do Museo Paulista 8:71-88. 27:145-151 [in Russian, German summary]. A. S. 1938b. iiber die Hummelfauna Schrottky, C. 1913. La distribucion geografica de los Himenopteros Skorikov, Vorlaufige Mifteilung Burmas. Arkiv for 30B:l-3. argentinos. Anales de la Sociedad Cientifica Argentina 75:115-144, Zoologi 180-286. Sladen, F W. L. 1915. The bee genus Tlirinchosloma in India. Canadian 47:213-215. Schrottky, C. 1914. Einige neue Bienen aus Siid-Amerika. Deutsche Entomologist Entomologische Zeitschrift 1914:625-630. Sladen, F. W. L. 1916a. Bees of Canada-Fam. Megachilidae. Canadian Schrottky, C. 1920. Himenopteros nuevos o poco conocidos Entomologist 48:269-272. sudamericanos. Revista do Museo Paulista 12:179-227. Sladen, F. W. L. 1916b. Canadian species of the bee genus Stelis Panz. Canadian 48:312-314. Schulz, W. A. 1906. Spolia Hi/nienoptewlogicn. Paderborn: Pape. Pp. 1-356. Entomologist 78 Scientific Papers, Natural History Museum, The University of Kansas

F. 1853. Smith, Catalogue of Hymenopterous Insects in the Collection of the Strand, E. 1910. Apidologisches aus dem Naturhistorischen Museum zu British Museum. Part 1. London: British + Museum. Pp. [i] 1-198, pis. Wiesbaden. Jahrbucher des Nassauischen Vereins fiir Naturkunde i-vi. 63:37^5. Smith, F. 1854. Insects in the Collection the Catalogue of Hymenopterous of Strand, E. 1913a. Apidae von Ceylon, gesammelt 1899 von Herrn. Dr. W. British Museum. Part 2. London: British Museum. Pp. 199-465, pis. Horn. Archiv fur Naturgeschichte, Abt. A 79(2):135-150. vii-xii. E. 1913b. H. Strand, Sauter's Formosa-Ausbeute, Apidae I. Supplementa Smith, F 1858. of the insects collected at Catalogue hymenopterous Entomologica no. 2:23-67. Sarawak, Borneo; Mount Ophir, Malacca; and at A. R. Singapore by Strand, E. 1914. H. Sauter's Formosa-Ausbeute, Apidae III. Archiv fiir Wallace. Journal of the of the Linnean of Lon- Proceedings Society Naturgeschichte, Abt. A80(l):136-144. don, Zoology 2(1857):42-180, pis. 1, 2. E. 1921 . Strand, Apidologisches, Lnsbesondere uber palaarktische Amirena- Smith, F. 1861. Descriptions of new genera and species of exotic Hy- Arten, auf Grund von Material des Deutschen Entomologischen menoptera. Journal of Entomology [London] 1:146-155. Museums. Archiv fiir Naturgeschichte, Abt. A 87(3):266-304. Smith, F. 1865. of some new of in- Descriptions species hymenopterous E. 1926. Strand, Miscellanea nomenclatorica zoologica et palaeontologica. sects belonging to the families Thynnidae, Masaridae and Apidae. Archiv fiir Naturgeschichte, Abt. A 92(8):30-75. Transactions of the Society of London (3)2:389-399, Entomological E. 1932. Miscellanea Strand, nomenclatorica zoologica et palaeontologica. pi. 21. Folia Zoologica et Hydrobiologica 4:193-196. Smith, F 1868a. Descriptions of aculeate Hymenoptera from Australia. Swenk, M. H. 1907. The bees of Nebraska—III. News Transactions of the of London 1868:231-258. Entomological Entomological Society 18:293-300. Smith, F. 1968b. [A new name for Oestropisis.] Proceedings of the Entomo- Taschenberg, E. 1883. Die Gattungen der Bienen (Anthophila). Berliner logical Society of London, p. xxxix. Entomologische Zeitschrift 27:37-100. Smith, F. 1875. Descriptions of new species of Indian aculeate Hy- Thomson, C. G. 1872. Skandinaviens Hymenoptera. Vol. 2. Lund: menoptera, collected by Mr G. R. James Rothney, member of the Berling. Pp. 1-286. Entomological Society. Transactions of the Entomological Society of R. D. S. London 1875:33-50. Thorp, W., Horning, Jr., and L. L. Dunning. 1983. Bumble bees and cuckoo bumble bees of California. Bulletin of the California In- Smith, F. 1879. Descriptions of Nero Species of Hymenoptera in the Collection sect Survey 23:i-viii + 1-79 pp. of the British Museum. London: Britisn Museum, xxi + 240 pp. Timberlake, P. H. 1939. New of bees of the from Snelling, R. R. 1956. Bees of the genus Centris in California. Pan-Pacific species genus Dufourea California Annals of the Entomologist 32:1-8. (Hymenoptera, Apoidea). Entomological of America 32:395-414. Snelling, R. R. 1966. Studies on North American bees of the genus Hi/laeus Society Timberlake, P. H. 1941. Ten new of Stelis from California. 2. Description of a new subgenus and species. Proceedings of the species Journal of the New York 49:123-137. Biological Society of Washington 79:139-144. Entomological Society Timberlake, P. H. 1952a. of new of Snelling, R. R. 1967. Description of a new subgenus of Osmia. Bulletin of Descriptions species Nomadopsis from the Southern California Academy of Sciences 66:103-108. California and Texas, and of a new allied genus from South America. Annals of the of America Snelling, R. R. 1974. Notes on the distribution and taxonomy of some Entomological Society 45:104-118. P. H. 1952b. new for the bee Ruiziella. North American Centris. Contributions in Science, Natural History Timberlake, A name genus Annals Museum of Los Angeles County no. 259:1-41. of the Entomological Society of America 45:528. P. H. 1953. Erratum. Snelling, R. R. 1982. The taxonomy of some neotropical Hylaeus and de- Timberlake, Annals of the Entomological Society of America scriptions of new taxa. Bulletin of the Southern California Academy 46:598. of Sciences 81:1-25. Timberlake, P. H. 1954. A revisional study of the bees of the genus Perditn Snelling, R. R. 1984. Studies on the taxonomy and distribution of Ameri- F Smith, with special reference to the fauna of the Pacific coast. Part can centridine bees. Contributions in Science, Natural History Mu- I. University of California Publications in Entomology 9:345^32. seum of Los Angeles County no. 347:1-69. Timberlake, P. H. 1955. A new genus for two new species of dufoureine R. bees from Snelling, R. 1985. The systematics of the hylaeine bees (Hymenoptera: California. Pan-Pacific Entomologist 31:105-108. P. H. 1956. Colietidae) of the Ethiopian zoological region: The genera and sub- Timberlake, A revisional study of the bees of the genus Perdita genera with revisions of the smaller groups. Contributions in Sci- F. Smith, with special reference to the fauna of the Pacific coast. Part ence, Natural History Museum of Los Angeles County no. 361:1-33. II. University of California Publications in Entomology 11:247-350. Snelling, R. R. 1 986. Conbibutions toward a revision of the new world nomadine Timberlake, P. H. 1960. A revisional study of the bees of the genus Perdita bees: A partihoning of the genus Nomiuki. Contributions in Science, Natu- F Smith, with special reference to the fauna of the Pacific coast. Part ral History Museum of Los Angeles County no. 376:1-32. IV. University of California Publications in Entomology 17:1-156. Snelling, R. R. 1990. A review of the native North American bees of the Timberlake, P. H. 1967. New species of Pseudopanurgus from Arizona. American Novitates no. genus Chalicodoma. Contributions in Science, Natural History Mu- Museum 2298:1-23. seum of Los Angeles County no. 421:1-39. Timberlake, P. H. 1969. Metapsaem/thia, a new panurgine bee genus. En- Snelling, R. R., and R. W. Brooks. 1985. A review of the genera of tomological News 80:89-92. cleptoparasitic bees of the tribe Ericrocini. Contributions in Science, Timberlake, P. H. 1980. Review of North American Exomalopsis, Parts I- Natural History Museum of Los Angeles County no. 369:1-34. IV. University of California Publications in Entomology 86:i-vi + 1- 158. Spinola, M. 1808. Insectorum Liguriae Species Novae aut Rariores. Vol. 2. Genuae. ii -i- 262 pp. Titus, E. S. G. 1901. A new genus in the Coelixinae [sic]. Canadian Ento- Spinola, M. 1838. Compte-rendu des Hymenopteres, recueillis par M. mologist 33:256. Fischer pendant son voyage en Egypte. Annales de la Societe Titus, E. S. G. 1904a. Notes on Osmiinae with descriptions of new genera Entomologique de France 7:437-546. and species. Journal of the New York Entomological Society 12:22- M. 1 843. Sur 27. Spinola, quelques Hymenopteres peu connus, recueillis en Espagne, E. S. G. 1904b. pendant I'annee 1842, par M. Victor Ghiliani, voyageur-naturaliste. Arinales Titus, Some new Osmiinae in the United States National de la Societe Entomologique de France (2)1:111-144. Museum. Proceedings of the Entomological Society of Washington Spinola, M. 1851 . Hymenopteros. Pp. 153-569 in Gay, C, Historia Fisica y 6:98-102. Politica de Chile. Zoologia. Vol. 6. Paris: casa del autor. Titus, E. S. G. 1906. Some notes on the Provancher Megachihdae. Pro- Spinola, M. 1853. Compte rendu des hymenopteres inedits provenants ceedings of the Entomological Society of Washington 7:149-165. du voyage entomologique de M. Ghiliani dans le Para en 1846. Tkalcu, B. 1966. Metallinetla gen. n. der Familie Megachilidae. Acta Memorie della Reale Accademia delle Scienze di Torino (2)13:19-94. Entomologica Bohemoslovaca 63:200-202. F K. 1954. Fauna B. 1 Stoeckhert, Apoideorum Germaniae. Abhandlungen Tkalcii, 967. Bemerkungen zur Taxonomie einiger palaarkKscher Arten der Bayerischen Akademie der Wissenschaften (n.f.)65:l-87. (Anhang der Familie Megachilidae. Acta Entomologica Bohemoslovaca 64:91- by H. Bischoff, pp. 70-73.) 104. Genus-Group Names of Bees 79

Tkalcu, B. 19b9. zur Kenntnis der Beitrage Fauna Afghanistans- Urban, D. 1989. Dois generos novos de Eucerinae Revista Chalkodoma neotropicais. Lep., Megachilidae, Apoidea. Acta Musei Moraviae Brasileira de Zoologia 6:117-124. 54:347-384, 6 unnumbered plates. D. 1991 . Ananthidium, di Urban, um genero novo Diantliidiini neotropical. Tkalcu, B. 1971. Xcnochalicodoma, nomcn novum. Bulletin de la Societe Revista Brasileira de Zoologia 7:73-78. Entomologique de Mulhouse 1971:34. D. 1993. n. de Urban, Ctenanthidium, gen. Dianthidiini com quatro especies Tkalcu, B. 1972. centre Arguments I'interpretation traditionnelle de la novas da America do Sul. Revista Brasileira de Zoologia 8(]991)-85- phylogenie des abeilles. Bulletin de la Societe Entomologique de 93. Mulhouse, 1972:17-28. Apr. -June, Urban, D. 1994a. Tylanthidium, gen. n. de Anthidiinae da America do Sul B. 1974a. Tkalcii, Ergebnisse der Albanien-Expedition 1961 des (Hymenoptera, Megachilidae) e nota taxonomica. Revista Brasileira "Deutschen Entomologischen Institutes," Hymenoptera: Apoidea V de Zoologia 11:277-281. zur (Megachilidae). Beitrage Entomologie [Berlin] 24:323-348. Urban, D. 1994b. Gnatlmnthidiurn, gen. n. de Anthidiinae da America do B. 1 974b. Revision und Tkalcu, Klassifikation der bisher zur Untergattung Sul. Revista Brasileira de Zoologia 9(1992):337-343. Thomson D. 1995a. n. Hoptcftnia gestellten Anthocofia-Arten. Acta Entomologica Urban, Mourennihidium , gen. de Dianthidiini do Brasil. Revista Bohemoslovaca 71:114-135. Brasileira de Zoologia 12:37^5. Tkalcu, B. 1975a. Die OsHjm-Arten der D. Untergattung Hamosmin subgen. Urban, 1995b. Grnfantliidiwn, gen. n. de Dianthidiini do Brasil e uma n. Acta Entomologica Bohemoslovaca 72:34-49. especie nova de Duckeanthidnim Moure & Hurd. Revista Brasileira Tkalcij, B. 1975b. de 12:435-443. Sammelergebnisse der von RNDr. A. Hotter geleiteten Zoologia in den 1971 1972 Urban, D. 1996. Mielkeanlhidium n. Algerien-ExpediHonen Jahren und (Hymenoptera: , gen de Dianthidiini da America do Apoidea). 1. Teil: Megachilidae. Zbornik Slovenskeho Narodneho Sul. Revista Brasileira de Zoologia 13:121-125. Prirodne Rerum Naturalium Urban, D., and S. Moure. 1 993. Muzea, Vedy [Acta Musei Nationalis J. Ccblurgus longipalpiis gen. e sp. n., primeiro Slovaci Bratislava] 21:165-190. representante de Dufoureinae do Brasil. Anais da Academia Brasileira Tkalc-ij, B. 1978. zur Kenntnis der deCiencias 65:101-106. Beitrage Fauna Afghanistans:Mt'/iff»rgfl Latr., Eucera Vachal, 1 897. Eclaircissements sur Scop., Apjdae; Lilhurgc Latr., Stelis Pz., CreightoiicUn J. de genre Scrapter et description d'une Cockll., Megachilidae. Casopis Moravskeho Musea 63:153-181. espece nouvelle de Dufourea. Bulletin de la Societe Entomologique B. 1979. de France 1897:61-64. TkalciJ, Neue palaarktische Taxa der Familie Megachilidae. Acta 1899. Entomologica Bohemoslovaca 76:318-329. Vachal, J. Contributions Hymenopteriques. Annates de la Societe B. 1980. Zwei de France 68:534-539. Tkalcu, neue Arten der Gattung Waiiiin gen. n. aus Entomologique Vachal, 1903-1904. Etude sur les Vorderindian. Annotationes Zoologicae et Botanicae, Bratislava, no. J. Halictus d'Amerique. Miscellanea 135:1-20. Entomologica (Narbonne) 11:89-104, 121-136 (1903); 12:9-24, 113-128, 137-144 Tkalcu, B. 1984a. Neue palaarktische Arten der Gattungen Pseudohermdes (1904). und Archeriades Vachal, J. 1905a. Un nouveau mit Beschreibung von Hofferia gen. n. Annotationes genre d'Hymenopteres, de la familie Apidae. Zoologicae et Botanicae, Bratislava, no. 158:1-22. Bulletin de la Societe Entomologique de France, 1905:204. B. 1984b. Vachal, J. 1905b. Manuelia, un nouveau Tkalcu, Systematisches Verzeichnis der westpalaarktischen genre d'Hymenopteres melliferes. Bulletin Tetrnkvua- und £»cmj-Arten, deren Mannchen als Bliitenbesucher de la Societe Entomologique de France 1905:25-26. verschiedener Vachal, J. 1909a. nouvelles ou du haut bassin Of/in/s-Arten festgestellt wurden; mit Beschreibung Especes Htigieuses d'Apidae neuer Taxa. du Parana et des et delimitation d'une nouvelle Nova Acta Regiae Societatis Scientiarum Upsaliensis (Ser. regions contigues V:C)3:57-77. sous-famille Diphaglossinae. Revue d'Entomologie (Caen) 28:5-64. B. 1993a. Vachal, J. 1 909b. Sur le Melihmm S. F et Serv. et sur les voisins Tkalcu, Quatre nouveaux sous-genres palearctiques de la tribu genre genres de la sous-famille Annales de la des Osmiini. Bulletin de la Societe Entomologique de Mulhouse Anthophorinae. Societe 1993:55-56. Entomologique de France 78:5-14. Tkalcu, B. 1993b. Neue Taxa der Vecht, J. van der. 1952. A revision of the oriental of Bienen von den Kanarischen Inseln. Pp. preliminary species the 791-858 in H. F G. genus Cerntina. No. 16:ii -^ 1-85. Hohmann, LaRoche, Ortega, and J. Barquin, Btcncn, Zoologische Verhandelingen und Verhoeff, C. 1890. Fin zur deutschen Wespen Ameisen der Kanarischen Inseln. Veroffentlichungen aus Beitrag Hymenopteren-Fauna. Nachrichten 16:321-335. dem Ubersee-Museum Bremen, Naturwissenschaften 12:1^65, pis Entomologische H. I-XII. Viereck, L. 1904a. A bee visitor of Po»ft'rfenrt (pickerel-weed). Entomo- News 15:244-246. Tkalcu, B. 1995. Die Bienen der Tribus Osmiini der Mongolei. logical H. L. 1904b. American Entomologische Abhandlungen 57:109-147. Viereck, genera of the bee family Dufoureidae. H. 1976. News 15:261-262. Toro, Clulunalopsis, nuevo genero chileno de Exomalopsini. Entomological Viereck, H. L. 1909. of new Anales del Museo de Historia Natural de Valparaiso 9:73-76. Descriptions Hymenoptera. Proceedings of H. the of Toro, 1980. Anstrcpianurgns nuevo genero de Panurginae chileno. Anales Entomological Society Washington 11:42-51. del Viereck, H. L. 1912. Contributions to our Museo de Historia Natural de Valparaiso 13:209-212. knowledge of bees and ichneu- H. the Toro, 1989. Contribucion al conocimiento de los Panurginae chilenos. mon-flies, including descriptions of twenty-one new genera and Acta Entomologica Chilena 15:229-232. fifth-seven new species of ichneumon-flies. Proceedings of the United and V. States National Museum 42:613-648. Toro, H., Cabezas. 1977. Nuevos generos y especies de Colletini H. L. 1916. The sudamericanos, primera parte. Anales del Museo de Historia Natu- Viereck, Hymenoptera, or Wasp-like Insects of Connecti- ral de Valparaiso 10:45-64. cut. Connecticut State Geological and Natural History Survey Bulle- and V. tin I-X. Toro, H., Cabezas. 1978. Nuevos generos y especies de Colletini 22:1-824, pis. H. L. 1917. sudamericanos, segunda parte. Anales del Museo de Historia Natu- Viereck, Contributions to our knowledge of the bee genus ral de Perditn Smith. Valparaiso 11:131-148. Bulletin of the American Museum of Natural History and M. 37:241-242. Toro, H., Herrera. 1980. Las especies chilenas del genero H. L. 1922. CaUimychtum (Andrenidae-Apoidea) y descripcion de un nuevo Viereck, New bees of the genus Andrena. Occasional Papers of Anales del Museo de Historia Natural de the Boston of Natural genero. Valpariso 13:213- Society History 5:35-45, pi. 4. 225. Viereck, H. L. 1924. Prodromus of Andrena, a genus of bees. Canadian Toro, H., and A. Moldenke. 1979. Revision de los Xeromelissinae chilenos. Entomologist 56:19-24. Anales del Museo de Historia Natural de Valparaiso 12:95-182. Vitzthum, H. G. 1930. Acarologische Beobachtungen (14 Reihe). Toro, H., and L. Ruz. 1972. Revision del genero Spinolieila. Anales del Zoologische Jahrbiicher, Abteilung fiir Systematik, Okologie und Museo de Historia Natural de Valparaiso 5:137-171, 293-295. Geographic der Tiere 59:281-350. Tosi, A. 1896. Di nuovo di fossile nell'ambra di Sicilia O. 1911. Studien genere Apiaria Vogt, fiber das Artproblem. Mitteilung 2, Teil 2. - M. sicula. Meliponon/tes succini, Rivista Italiana di Paleontologia Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Ber- 2:352-356. lin 1911:31-74. 80 Scientific Papers, Natural History Museum, The University of Kansas

Walker, K. L. 1986. Revision of the Australian species of the genus Warncke, K. 1991c. Die Bienengattung Osmin Panzer, 1906, ihre Homalictus Cockerell. Memoirs of the Museum of Victoria 47:105-200. Systematik in der Westpalaarktis und ihre Verbreitung in der Tiirkei. Warncke, K. 1968. Die untergattungen der westpalaarktischen 7. Die Untergattung Foveosmia nov. Linzer Biologische Beitrage 23:267- Bienengattung Andrcna F. Memorias e Estudos do Museu Zoologico 281. da Universidade de Coimbra, no. 307:1-111. Westwood, J. O. 1835. [Characters of new genera and species of hy- Warncke, K. 1970. Beitrag zur Systematik und Verbreitung der menopterous insects.] Proceedings of the Zoological Society of Lon- Bienengattung Prosopis F. in der Westpaleiarktis. Bulletin des don 3:51-54, 68-72. Recherches Agronomiques de Gembloux (NS)5:745-768. Westwood, J.'0. 1838. Description of a new genus of exotic bees. Transac- Warncke, K. 1972. Westpalaarktische Bienen der Unterfamilie Panurginae. tions of the Entomological Society of London 2:112-113. Polskie Pismo Entomologiczne 42:53-108. Westwood, J. O. 1840a. Synopsis of the genera of British insects. Pp. 1- Warncke, K. 1974. Die Sandbienen der Tiirkei, 1 Teil. Mitteilungen der 154 ill An Introduction to tlic Modern Classification of Insects.... Vols. 1, Munchner Entomologischen Gesellschaft 64:81-116. 2, 1838-1840. London: Longman et al. [Commission Direction 63 (1957) Warncke, K. 1975a. Die Sandbienen der Tiirkei, Teil B. Mitteilungen der gives dates for the parts of this work; the parts of the Synopsis on Miinchner Entomologischen Gesellschaft 65:29-102. bees all appeared in 1840.] Warncke, K. 1975b. Beitrag zur Systematik und Verbreitung der Westwood, J. 0. 1840b. In ]. Duncan, The Natural Historii of the Bees, m W. Furchenbienen in der Tiirkei (Hymenoptera, Apoidea, HaUctus). Jardine (ed.), Tlic Naturalists' Library. Vol. 26. Entomology, Vol. 6. Polskie Pismo Entomologiczne 45:81-123. Edinburgh: Lizars. viii -^ 17-301, 30 pis. 1976. 1 875. Warncke, K. Zur Systematik und Verbreitung der Bienengattung Westwood, ]. 0. Descriptions of some new species of short-tongued Nomin Latr. in der Westpalaarktis und dem turkestanischen Becken. bees belonging to the genus Nomia. Transactions of the Entomologi- Reichenbachia 16:93-120. cal Society of London 1875:207-222, pis. IV, V. Warncke, K. 1977. Ideen zum naturlichen System der Bienen. Mitteilungen Williams, P. H. 1985. A preliminary cladistic investigation of relation- der Miinchner Entomologischen Gesellschaft 67:39-63. ships among the bumble bees. Systematic Entomology 10:239-255. Warncke, K. 1978. liber die westpalaarktischen Arten der Bienengattung Williams, P H. 1991. The bumblebees of the Kashmir Himalaya. Bulletin Colletes Latr. Polskie Pismo Entomologiczne 48:329-370. of the British Museum (Natural History), Entomology 60:1-204. Warncke, K. 1979. Beitrage zur Bienenfauna des Iran: 3. Die Gattung Williams, P. H. 1994. Phylogenetic relationships among bumblebees Rophites Spin., mit einer revision des Westpalaarktischen arten der (Bombus Latr.): A reappraisal of morphological evidence. Systematic Bienengattung Rophites Spin. BoUettino del Museo Civico di Storia Entomology 19:327-344. Naturale di Venezia 30:111-155. Wu, Y.-r. 1982. Description of a new subgenus of Nomia. Zoological Re- Warncke, K. 1980. Die bienengattungen Noniui und Systwplui im Iran mit search 3:275-280 [in Chinese, English summary]. erganzungen zu den No»n(!-Arten der Westpalaarktis. Linzer Wu, Y.-r. 1985. A study on the genus Rliopmlomelissa of China with de- Biologische Beitrage 12:363-384. scriptions of new subgenus and new species. Zoological Research Warncke, K. 1981. Beitrag zur Bienenfauna des Iran 14. Die gattung 6:57-68 ]in Chinese, English summary]. Halictiis Latr., mit Bemerkungen iiber bekannte und neue Hnlictus- Yarrow, I. H. H. 1971. The author and date of certain subgeneric names in Arten in der Westpalaarktis und Zentralasian. BoUetino del Museo Bombus. Journal of Entomology (B)40:27-29. Civico di Storia Naturale di Venezia 32:67-166. Yasumatsu, K., and Y. Hirashima. 1950. Revision of the genus Osmia of Warncke, K. 1983. Zur Kenntnis der Bienengattung Pnsjfcs Jurine, 1807, Japan and Korea. Mushi 21:1-18, pis. 1-3. in der Westpalaarktis. Entomofauna 4:261-347. Zanden, G. van der. 1988. Beitrag zur Svstematik und Nomenklature der Warncke, K. 1985. Beitrage zur Bienenfauna des Iran 19-20. Die Gattungen palaarktischen Osmiini, mit Angaben uber ihre Verbreitung. Panurgus Pz. und Melitiirgiila Fr. BoUettino del Museo Civico di Storia Zoologische Mededelingen 62:113-133. Naturale di Venezia 34(1983):221-235. Zanden, G. van der. 1989. Neue oder wenig bekannte Arten und Warncke, K. 1988a. Die Bienengattung Ofinia Panzer, 1806, ihre Systematik Unterarten der palaearktischen Megachiliden. Entomologische in der Westpalaarktis und ihre Verbreitung in der Turkei. 2 und 3: Abhandlungen (Dresden) 53:71-86. Die Untergattungen Tergosniin und Exosmia. Entcimofauna 9:389^03. Zanden, G. van der. 1 991 . Neue oder wenig bekannte Arten der Osmiini Warncke, K. 1988b. Die Bienengattimg Osmia Panzer, 1806, ihre Systematik aus dem palaarktischen Gebiet. Reichenbachia 28:163-171. in der Westpalaarktis und ihre Verbreitung in der Turkei. 1: Zanden, G. van der. 1994. Neue Arten und Unterarten, eine neue Untergattung Ht'/icosm/n Thomson, 1872. Entomofauna 9:1-48. Untergattung und einige neue Falle von Synonymic der Warncke, K. 1990. Die Bienengattung Osmia Panzer, 1806, ihre Systematik palaarktischen Bauchsammler. Reichenbachia 30:167-172. in 1972. der Westpalaarktis und ihre Verbreitung in der Tiirkei. 4. Die Zavortink, T J. A new subgenus and species of Megandrena from Untergattung Platosmia subgen. nov. Entomofauna 11:481^95. Nevada, with notes on its foraging and mating behavior. Proceed- Warncke, K. 1991a. Die Bienengattung Osmia Panzer, 1906, ihre Systematik ings of the Entomological Society of Washington 74:61-75. in ihre in Tiirkei. 5. T. 1975. of from North der Westpalaarktis und Verbreitung der Die Zavortink, J. A new genus and species eucerine bee Untergattung Pentndentosmia subg. nov. Entomofauna 12:13-32. America. Proceedings of the California Academy of Sciences Warncke, K. 1991b. Die Bienengattung Ofmin Panzer, 1906, ihre Systematik (4)40:231-242. in der Westpalaarktis und ihre Verbreitung in der Tiirkei. 9. Die Zeuner, F. E., and F.J. Manning. 1 976. A monograph on fossil bees. Bulle- Untergattung Atiuosmia subg. n. Linzer Biologische Beitrage 23:307- tin of the British Museum (Natural History), Geology 27:149-268, pis. 336. 1-i. Genus-Group Names of Bees 81

ADDENDUM

After this paper was in proof. Dr. Donald B. Baker (University Museum. Oxford) kindly sent me commentaries on certain dates of publication. Whereas I have usually used widely accepted dates, Dr. Baker has indicated that the following are correct:

Author, date Corrected Date

Agassiz, L. 1846. 1847

Guerin-Meneville, F. E. 1830. pp. 1-24, 1830: pp. 25^71. 1831

Heer, O. 1850. 1849

Lepeletier de S. Fargeau. A. L. M. 1836. 1835

Morawitz, P. 1873a. 1874

Radoszkowski. O. 1868. 1867

Schenck.A. 1866. 1867

Schenck.A. 1973. 1874

Smith, F 1858. pp. 42-88 and pis. 1, 2, 1857; pp. 89-180. 1858

Spinola. M. 1838. 1839

Vachal.J. 1899. 1900 iX DATE DUE QL566 .M43 1997 -• "IK. of i.,o .„„i Harvard[:"'" MCZ M„.,,i Library Vk'i -.,

3 2044 062 458 666 PUBLICATIONS OF THE NATURAL HISTORY MUSEUM, THE UNIVERSITY OF KANSAS

The University of Kansas Publications, Museum of Natural History, beginning with Volume 1 in 1946, was discontinued with Volume 20 in 1971. Shorter research papers formerly published in the above series were published as The University of Kansas Natural History Museum Occasional Papers until Number 180 in Decem- ber 1996. The Miscellaneous Publications of The University of Kansas Natural His- tory Museum began with Number 1 in 1946 and ended with Number 68 in Febru- ary 1996. Monographs of The University of Kansas Natural History Museum were initiated in 1970 and discontinued with Number 8 in 1992. The University of Kan- sas Science Bulletin, beginning with Volume 1 in 1902, was discontinued with Vol- ume 53 in 1996. The foregoing publication series are now combined in a new series entitled Scientific Papers, Natural History Museum, The University of Kansas, be- gun with Number 1 in 1997. Special Publications began in 1976 and continue as an outlet for longer contributions and are available by purchase only. All manuscripts are subject to critical review by intra- and extramural speciaUsts; final acceptance is at the discretion of the editor.

The publication is printed on acid-free paper Publications are composed using Microsoft Word" and Adobe PageMaker® on a Macintosh computer and are printed by The University of Kansas Printing Service.

Institutional Ubraries interested in exchanging pubUcations may obtain the Sci- entific Papers, Natural History Museum, The University of Kansas, by addressing the Exchange Librarian, The University of Kansas Libraries, Lawrence, Kansas 66045- 2800, USA. Available back issues of The University of Kansas Science Bulletin may be purchased from the Library Sales Section, Retrieval Services Department, The University of Kansas Libraries, Lawrence, Kansas 66045-2800, USA. Available issues of former publication series. Scientific Papers, and Special Publications of the Natural History Museum can be purchased from the Office of Publications, Natu- ral History Museum, The University of Kansas, Lawrence, Kansas 66045-2454, USA. Purchasing information can be obtained by calling (913) 864-4450, fax (913) 864- 5335, or e-mail ([email protected]). VISA and MasterCard accepted; include expiration date.

Editor of SciENTinc Publications: William E. Duellman

Printed BY The University of Kansas Printing Services Lawrence, Kansas