植物研究雑誌 J. J. Jpn. Bo t. 76: 76: 137-150(2001)

Umbelliferae Umbelliferae in Japan 1

Takasi Takasi Y AMAZAKI

Nakano-ku ，Tokyo ，165'- JAPAN (Received (Received on January 9， 2001)

The Umbelliferae in Japan has been c1 assified into 30 genera and 90 species. of these species has been treated by several authors. However ， the scientific names of some taxa 紅 'e different by the authors ， so 1 attempt to sort out the right scien- tific tific names of species these and genera in this paper. The genus Spuriopimpinella Ki tag. is is united to the genus Pimpinella L.， and the genera Tilingia Regel and Dystaenia Ki tag. to 血e genus Ligusticum L. It is c1 ear that Angelica sec t. Acutilobi with 7 species is an en- demic demic section of 白e Japanese Islands. In this paper a key to Japanese genera is provided and and 11 genera are treated; Hydrocotyle ， Centella ，Sanicula ， To rilis ， ， ，Pleurospermum ，Bupleurum ， ， Pternopetalum and Aegopodium.

Key words: Japan ，taxonomy ，Umbelliferae

Taxonomy of U mbelliferae in Japan has Ohba H. ， Umbelliferae in Iwatsuki & al. been revised by several authors ， such as Hara eds. ， F l. Japan IIc: 268-303 (1999). (1 954) ， Hiroe and Constance (1 958) ， Ki tamura and Murata (1961) ，Ohwi (1 975) ， Key to the genera Kitagawa (1982) ，Ohba (1999). However ， A. Stems slender ， creeping; the the treatments for some genera and species axillary ， with a single umbel; fruits are are different by authors. 1 attempt to put out S住ongly flattened laterally (Subfam. I. a new taxonomy of Umbelliferae in consid- Hydrocotyloideae ). ering ering the related species in Russia and China. B. Petiole with a pair of free stipules at base; base; involucre wanting or inconspicu- Umbelliferae Umbelliferae Scopoli ous; fruits without ribs and reticula-

Hara H. ， Umbelliferae in Enum. Sperm. tions... ・H ・....・ H ・-… H ・H ・..… 1. Hydroco かle Jap. Jap. 3: 291-328 (1 954). B. Petiole sheathing at base ， without Hiroe Hiroe M. & Constance L. ， Umbelliferae in stipule; involucre conspicuous; fruits Japan Japan in Univ. Calif. Pub l. Botany 30: 1-144 with evident ribs and reticulations … (1 958). … 2. Centella Ki tamura S. & Murata G. ， Umbelliferae in A. Stems erect or ascending ， not creeping; Coloured Il lustr. Herbaceous Pl. Japan 11: 1- inflorescence terminal and axillary ， with 32 ， plates 9 (1961). a few to many umbels; fruits terete ， Ohwi J. ， Umbelliferae in Fl. Japan ed. 2: slightly appressed laterally ， or flattened 969-1003 (1975). dorsi -ventrally. Ki tagawa M. ， Umbelliferae in Satake & B. Sepals prominent ， froming a persistent l. al. eds. ，Wild Fl. Japan. Herbaceous Pl. 2: corona on fruit (Subfam. II. Saniculoi-

276-290 ， pls.249-272 (1 982). deae)... ・H ・.....・ H ・....・ H ・...・ H ・... 3. Sanicula

-137- 138 138 植物研究雑誌第76 巻第3号 平成13 年6月

B. B. Sepals inconspicuous or small ， not H. Umbellets paniculately arranged; forming forming a corona on fruit (Subfam. 11 1. i町 egular in an umbelle t... Apioideae Apioideae ). … 9. Cη 'P totaenia C. C. Oil-tubes of fruits solitary to many ， H. Umbellets umbellately arranged; intercostal ， sometimes lucking; flowers regular or irregular in an fruits fruits with or without spines ， inner umbelle t. face face of endosperm flattened or con ・ 1. Umbellets with irregular 1-3 cave. cave. flowers; cauline leaves with lin- D. D. Mericarps suborbicular or semi- ear segments ， much different orbicular orbicular in cross section; inner from radical ones. …・・ face face of endosperm concave. . 10. Pternopetalum E. E. Mericarps without or with ob- 1. Umbellets with 5 to many flow- scure scure ribs outside (Tribe 1. ers; cauline leaves nearly the Scandiceae) Scandiceae) . same with radical ones. F. F. Fruits ellipsoid ， acute at apex ， 1. Oil-tubes lacking in frui t.

dense-prickly... ・H ・H ・H ・...4. Torilis K. Stylopodium distinct ，conica l..…・ F. F. Fruits lanceolate to linear- . 11. Aegopodium cylindric ， beaked at apex ， K. Stylopodium low ，fla t...... glabrous ，hirsute or setose. .12. Ch α maele G. G. Mericarps rounded at base ， 1. Oil-tubes present in frui t. glabrous glabrous or hirsute outside..... K. Rachis of distinct ， .. .. 5. Anthriscus so the umbels paniculately ar- G. G. Mericarps long-caudate at base ， ranged; ultimate leaf-segments appressed-setose appressed-setose outside …・ ovate to linear.

...・ H ・H ・H ・..… 6. Osmorhiza L. Stems terete; involucral bracts E. E. Mericarps with semi-winged 5 ribs lacking ，rarely 1-2.

(Tribe (Tribe 2. Smyrnieae )...・ H ・....・ H ・.... M. not fleshy; ultimate seg- ..7. ..7. Pleurospermum ments of leaves ovate to D. D. Mericarps semiorbicular to lanceolate; growing in forests or compressed-semiorbicular compressed-semiorbicular or flat- dry meadows. …… 13. Pimpinella tened tened dorsi-ventrally in cross sec- M. Rh izome fleshy; ultimate segmnts tion; tion; inner face of endosperm of leaves linear-lanceolate; grow-

flattened flattened or nearly so ， or slightly ing in wet places... ・H ・. 14. Cicuta convex ， not concave. L. Stems distinctly ridged; involuc-

E. E. Fruits slightly appressed dorsi- ral bracts presen t.....・ H ・.. 15. Sium ventrally; ventrally; meric 紅 ps semiorbicul- K. Racis of inflorescences lacking ， 紅 in cross section ，ribs slender or so the umbels terminal and distinct ， not winged (Tribe 3. axill 紅 y; ultimate leaf-segments

Ammineae). linear ...... ・ H ・..…...・ H ・..… 16. Apium F. F. Ribs of meric 紅 ps slender ，linear F. Ribs of mericarps highly ridged or or obscure (distinctly ridged in or wing -like ， at least in lateral Siumsuave) Siumsuave) (Subtribe 1. Carinae). ribs (Subtribe 2. Seselinae). G. G. Leaves simpl~ ， entire. …・… G. Fruits with carpophore; dorsal ..8. ..8. Bupleurum ribs of mericarp thickly corky. G. G. Leaves lobed or dentate ， usually H. Sepals distinct; style ca. 2 mm

compound. long.. …H ・H ・-… H ・H ・... 17. Oenanthe June June 2001 Journal of Japanese Botany Vo 1. 76 No. 3 139

H. Sepals inconspicuous; style 0.2- and endocarp... ・H ・-…. 24. Glehnia 0.3 0.3 mm long... 18. Apodicarpium G. Fruits glabrous outside; lateral G. G. Fruits with a bilobed slender ribs winged ， not corky ， nearly the C 紅 pophore; ribs of mericarp same or wider than the dorsal thinly thinly corky or wing-like (thickly ones; with one to several distinct corky corky in Cnidium). gaps between mesoc 紅 p and en- H. Ribs of mericarps firm ， corky. doc 紅 p. 1. 1. Ribs of mericarps stout; calyx- H. Dorsal ribs of mericarps winged ， teeth teeth and fruit glabrous. nearly the same with the lateral

J. J. Calyx-teeth inconspicuous: ribs ones... ・H ・...・ H ・...25. Conioselinum of of meric 紅 ps thick ， obtuse and H. Dorsal ribs of meric ぽ ps thickly corky corky throughout. …. 19. Cnidium filiform ， or narrowly winged ，n 紅- J. J. Calyx-teeth prominent and persis- rower than the lateral ones. tent; tent; ribs of mericarps ending in a 1. Dorsal ribs protruding ， thickly n紅 row thin wing ， dilated and filiform ，slightly narrower than corky corky at base the lateral ones … .. .. 20. Pterygopleurum …… 26. Coelopleurum 1. 1. Ribs of mericarps thin ， not wing- 1. Dorsal ribs thinly filiform ， like; like; calyx-teeth and fruit slightly protruding. puberulent puberulent or papillate-puberulent J. Oil-tubes slender and extending … 21. Libanotis nearly to quite the length of the H. Ribs of meric 征 ps thin ， wing-like fruits; flowers with equal petals. (at (at least on the lateral ribs) ， not K. Pericarp multi-layered ， opaque;

corky... ・H ・H ・H ・..… 22. Ligusticum calyx-teeth short or inconspicu-

E. E. Fruits much appressed dorsi- ous... …...・ H ・.....・ H ・..… 27. Angelica ventrally; ventrally; c紅 pels compressedly K. Pericarp single-layered ， hyaline;

semiorbicular semiorbicular or elliptic in cross calyx-teeth distinct ...・ H ・..， section ， lateral 2 ribs winged ， .. 28. Ostericum slightly slightly or distinctly broader than 1. Oil-tubes prominent ， extending

the the dorsal ones (Tribe 4. 1/ 3-1/2 of the fruit 田 length; m 紅 m Peucedaneae ). ginal flowers in an umbellet with F. F. Dorsal ribs of mericaprs thin ， unequal petals (Subtribe 3.

filiform ， not winged; lateral ribs Tordylinae) ....・ H ・.. 29. thickly thickly membranceous ， equaling C. Oil-tubes solitary under the ribs ，2 or or shorter than a half of the body toward commissure; fruit ellipsoid ， (Subtribe (Subtribe 1. Ferulinae) … beset with long spines ，endpsperm …・ 23. Peucedanum flatted at commissure (Tribe 5.

F. F. Dorsal ribs of mericarps stout ， Dauceae).... ・H ・...・ H ・...・ H ・..30. Daucus wing-like; wing-like; lateral ribs winged ， nearly nearly the same or broader than a Subfamily 1. Hydrocotyloideae Drude half half of the body (Subtribe 2. Drude in Eng l. & Prantl ，N at. Pflanzen- An gelicinae) . fam. 3， 8: 116 (1898). G. Fruits pilose outside; lateral ribs narrowly narrowly winged ，corky ， nearly Hydrocotyle L. the the same with the dorsal ones; L. ， Sp. Pl. 234 (1753). without without gaps between mesoc 訂 P 140 140 植物研究雑誌第76 巻第3号 平成13 年6月

A. A. Creeping stems with erect or ascending B. Petioles crisped-pubescent throughout; branches ， branches not rooting; umbels fruits ca. 0.8 mm long ・… on branches. … 8. H. dichondroides B. B. Umbels (1-) 2-5 in fascicles. C. C. Stems and petioles glabrous or 1. Hydrocotyle javanica Thunb. ， Diss. sp 紅 sely depressed-pubescent; lea- Bo t. Hydroco t.: 6， t. (1798); Hiroe & ves ves 2.5-5 cm long ， 3-6 cm wide ， Constance in Univ. Cali f. Pub l. Bo t. 30: 7 ，f. petioles petioles 2-5 times longer than the 2 (1958); Ohwi ， F l. Jap. ed. 2: 971 (1 975);

lamina; lamina; fruits ca. 1. 3 mm long.... ・H ・- H.Ohba in Iwatsuki & al. eds. ， F l. Jap. IIc: 1. . 1. H. javanica 269 (1 999). C. C. Stems and petioles densely crisped ・ Hydrocotyle nepalensis auc t. non Hook.: pubescent; pubescent; leaves 0.7-3 cm long H.Hara in J. Jpn. Bo t. 17: 342 (1941). and and wide ，petioles as longa as or オオパチドメ slightly slightly longer than the lamina; Japan: C. to W. Honshu (westward from

fruits fruits ca. 1 mm long... …H ・H ・- Ib 訂紘i Pref よ Shikoku ，Kyushu (northward

.. .. 2. H. pseudoconfert α， from Amami-Ohshima). Edges of evergreen B. B. Umbels solit 訂 y; leaves 0.7-2.5 cm forests at low elevations. long long and wide. Distr.: Japan ，Taiwan ， C. to S. China ， C. C. Leaves with many punctate dots be- Indochina ，Malay Peninsula ，Java ， India and neath ，shallowly lobed ， lobes over- Ceylon. lapping; lapping; in f1 0rescences longer than Note: Hydroco η le nepalensis Hoo k. in

the the leaves. …H ・H ・..… .3. H. ram 抑ora Nepal very resembles H. javanica ， but diι C. C. Leaves without dots benath ， deeply fers from the latter by the leaves with lobed lobed into 112-3/4 of the length ， subacute lobes and acute teeth and lobes lobes opening ， if overlapping at in f1 0rescences are generally soliatry in the apex ， at least opening at base. leaf-axils. It is necessary to make a critical D. D. Leaves generally 5-1obed into study based on more materials. 1/2-2/3 1/2-2/3 of the length; in f1 ores- cences cences shorter than the leaves. 2. Hydrocotyle pseudoconferta Masam. ..4. ..4. H. maritime in J. Soc. Trop. Agric. 4: 301 (1 932); D. D. Leaves generally 3-10bed into 3/4 S.L.Liou ， F l. Reip. Popu l. Sin. 55 (1): 14 ，f. of of the length; in f1 0rescences as 2， 1-3 (1979); Yamazaki in J. Jpn. Bo t. 75: long long as or slightly longer than the 371 (2000).

leaves. …....・ H ・... 5. H. batrachium Hydroco かle nepalensis auc t. non Hook.: A. A. Stems creeping throughout ，without erect Liu & Kao ， F l. Taiwan ed. 2 ，3: 1029 (1 993) ， or or ascending branches; umbels radica l. pro p紅白. B. B. Petioles glabrous or long-pilose at タイワンチドメグサ apex; apex; fruits 1-1.3 mm long. Distr.: The Philippines ， China (Taiwan ， C. C. Stems and leaves living in winter; Zhejuang) and Kyushu. winter winter buds absent; leaves lobed in 1/ 3- 1/ 4 of the length; ripe fruits ca 1 3. Hydrocotyle ramiflora Maxim. in

mm long.... ・H ・.. 6. H. sibthorpioides Bull. Acad. Sc i. St. -Pe t. 31: 46 (1886); H. C. C. Stems and leaves whithered in win- Hara in J. Jpn. Bo t. 17: 344 (1941); Hiroe & ter; ter; winter buds present; leaves Constance in Univ. Calif. Pub l. Bo t. 30: 9 ，f.

lobed lobed in 1/ 2-2/3 of the length ...・ H ・... 3 (1958) ， pro p紅白， exc l. syn. H. maritima ..7. ..7. H. yabei Honda; S.L.Liou ， F l. Reip. Popu l. Sin. 55 (1): June June 2001 JournaI of Japanese Botany Vo I. 76 No. 3 141

26 ，f. 7，6-10 (1979); H. Ohba in Iwatsuki & K yushu et Korea. 1. a1. eds. ， F 1. Jap. IIc: 269 (1 999). タカサゴチドメ Hydroco η le wilfordi Maxim. in Bul l. Japan: S. Ryukyu (Miyako Is l. and Acad. Acad. Sci. St. -Pe t. 31: 45 (1 886); S.L.Liou ， Ishigaki Is 1.). In heavy soils on rough pitted l. Fl. Reip. Popu l. Sin. 55 (1): 23 ，f. 7， 1-5 coral plains at low altitudes. (1 979). Dsitr.: Ryukyu ，Taiwan ， C. to S. China ヤマチドメ and Vietnam. Japan: Japan: S. to E. Hokkaido ， Honshu ， Note: Hydroco かle batrachium is fre- Shikoku and Kyushu. In meadows ，more or quently treated as a variety of H. less less wet places; low altitudes to 800 m. sibthorpioides Lam. but has more close af- Distr.: Distr.: Japan ， S. Korea and C. China. finity with H. m αritima Honda.

4. 4. Hydrocotyle maritima Honda in Bo t. 6. Hydrocotyle sibthorpioides L 制 n. ， and Zoo l. 2: 1825 (1 934); H.H 紅 a in 1. Jpn. Encyc l. Meth. 3: 153 (1789); H.Hara ，Enum. Bo t. 17: 344 (1941); H.Ohba in Iwatsuki & Sperm. Jap. 3: 312 (1 954); Hiroe & 1. a1. ed. ， F l. Jap. IIc: 270 (1 999). Constance in Univ. Calif. Pub l. Bo t. 30: 11 ， Hydroco 砂le wilfordii auc t. non Maxim.: f. 4 (1958); S.L.Liou ， F l. Reip. Popu l. Sin. Makino in Bo t. Mag. Tokyo 4: (415) (1 890). 55 (1): 17 ， f. 2，4-6 (1 979); H.Ohba in Hydroco η le ramifolia Maxim. v紅. Iwatsuki & a1. eds. ， F 1. Jap. IIc: 271 (1999). maritim α(Honda) Hiroe in Ac t. Phytotax. Hydroco η le rotundifolia Roxb. ， F l. Ind. Geobo t. 14: 40 (1 950). ed. 2: 88 (1 832). Hydroco η le formosana Masam. v紅. Hydroco η le rotundifolia Roxb. s maritima maritima (Honda) Hatus. ， F 1. Ryukyus ，adi t. sibthorpioides (Lam.) Spreng. in Sys t. et et co 町 ec t. ed.: 886 (1 975). Vege t. 6: 349 (1820). ノチドメ Hydroco かle zollingri Molkenboer ex Japan: Japan: Honshu ， Shikoku ， Kyushu and Miq. ， P 1. Jungh. 1: 91 (1 85 1). Ryukyu. Lawns and roadsides on hills; low- var. sibthorpioides. lands lands to 700 m. チドメグサ Distr.: Distr.: Japan ，Korea ，Taiwan ， C. to S. Japan: Honshu ，Shikoku ，Kyushu (north- China and Indochina. ward from Amami-Ohshima). More or less wet roadsides in low altitudes. 5. 5. Hydrocotyle batrachium Hance in Dsitr.: Widespread in w 紅 m-temperate to Ann. Sc. Na t. ser. 4， 18: 220 (1862); H.Hara 佐opic regions of Asia and Africa. in in J. Jpn. Bo t. 60: 136 (1985); H.Ohba in V 訂. tuberifera (Ohwi) Yamazaki in J. Iwatsuki Iwatsuki & al. ed. ， F 1. Jap. IIc: 270 (1999). Jpn. Bo t. 69: 112 (1994). Hydroco りJle rotundifolia Roxb. var. Hydrocotyle tuberifera Ohwi in J. Jpn. batrachium batrachium (Hance) Chermez. in Bul l. Soc. Bo t. 26: 232 (1951). Bo t. Fr. 68: 508 (1921). Hydrocotyle keelungensis Liu ，Chao & Hydroco η le formosana Masam. in J. Soc. Chuang in Techn. Bull. Experim. For. Na t. Trop. Trop. Agric. 2: 51 (1930). Taiwan Univ. no. 26: 15 (1 961). Hydrocotyle Hydrocotyle sibthorpioides Lam. v訂. オキナワチドメ batrachium batrachium (Hance) Hand.-Mazz. ex Shan in Japan: Ryukyu (Southward from Okinawa Sinensia Sinensia 7: 480 (1936); S.L.Liou ， F l. Reip. Is 1.). Popu l. Sin. 55(1): 18 ，f. 2， 7 --6 (1979). Distr.: Ryukyu and Taiwan. Hydrocotyle Hydrocotyle benguetensis auc t. non Elm.: Note: This V 紅 iety has been named ‘tuberi- Hatus. ， F l. Ryukyus: 450 (1 971) ， p. p. exc l. fera' ， but the tubers forming in the type 142 142 植物研究雑誌第76 巻第3号 平成13 年6月

specimen specimen are an abnormal character. ture fruits ca. 1.3 mm long ….....

…...・ H ・-… .var. ]aponzc α Key to the varieties of of Hydrocotyle sibthorpioides 8. Hydrocotyle dichondroides Makino in A. A. Leaves glabrous on upper surface and Bo t. Mag. Tokyo 24: 242 (1 910); Hiroe &

petioles petioles ...・ H ・.....・ H ・..… var. sibthorpioides Constance in Univ. Calif. Pub l. Bo t. 30: 12 ， A. A. Leaves sparsely setulose on nerves on f. 5 (1 958); Ohwi ， F l. Jap. ed. 2: 971 (1 975); upper upper surface; petiole crisply long- H.Ohba in Iwatsuki & a l. eds. ， F l. Jap. IIc:

pubescent pubescent ne 訂 apex.... ・H ・..v 紅 . tuberifera 271 (1 999); Yonekura in J. Jpn. Bo t. 74: 367 (1999). (1999). 7. 7. Hydrocotyle yabei Makino in Bo t. Hydroco η le sibthorpioides Lam. var. Mag. Tokyo 24: 243 (1910); H.Hara ，Enum. dichondroides (Makino) Hiroe in Ac t. Sperm. Sperm. Jap. 3: 312 (1 954); H.Ohba in Phytotax. Geobo t. 14: 40 (1950). Iwatsuki Iwatsuki & al. eds. ， F l. Jap. IIc: 270 (1 999). Hydroco η le delicat αauc t. non Elmer: Hydroco η le rotundifoli αRoxb. v紅. H.Hara ，Enum. Sperm. Jap. 3: 311 (1 954). paucijlora paucijlora Yabe ， Rev. Umbe l. Jap.: 14 ケチドメグサ (1 902). Japan: S. and W. Kyushu and Ryukyu. Hydroco η le nitidula auc t. non Richard: Edges of evergreen forests at lowlands. Nakai in Bo t. Mag. Tokyo 36: 25 (1922). Distr.: Japan and Taiwan. Hydrocotyle Hydrocotyle sibthorpioides Lam. var. pauc 伊ora (Yabe) Yamazaki in J. Jpn. Bo t. 2. Centella L. 69: 69: 112 (1 994). L. ， Gen. Pl. ed. 6: 485 (1 764). var. var. yabe i. ヒメチドメ 1. (L.) Urban in M 紅t. Distr.: Distr.: Honshu ， Shikoku and Kyushu. Fl. Brasi l. 11: 287 ，t. 78 ，f. 1 (1879); H.Hara ， Edges of forests ，more or less wet places; Enum. Sperm. Jap. 3: 303 (1954); Hiroe & low altitudes to 1000 m. Constance in Univ. Calif. Pub l. Bo t. 30: 13 ， V 訂. japonica (Makino) Hiroe in Ac t. f. 6 (1 958). Phytotax. Phytotax. Geobo t. 14: 41 (1950). Hydrocotyle asiatica L. ， Sp. Pl.: 234 Hydrocotyle Hydrocotyle japonica Makino in J. Jpn. (1753). Bo t. 3: 47 (1 926); F.Maek. in J. Jpn. Bo t. Hydrocotyle lun α ta Lam. ，Encyc l. Meth. 11: 11: 377 (1936); H.Hara ，Enum. Sperm. Jap. 3: 152 (1 789). 3: 3: 311 (1954). CenteU α boninensis N akai & Tuyama in ミヤマチドメ Bo t. Mag. Tokyo 50: 31 ，f. 23 (1936). Japan: Japan: C. and W. Honshu ， Shikoku and Centella asiatica L. var. boninensis Kyushu. Kyushu. In shady places on hills or moun- (Nakai & Tuyama) H.Hara ，Enum. Sperm. tains ， lowlands to 1300 m. Jap. 3: 304 (1954). Distr.: Distr.: Japan and Korea (Is. Chejudo). ツボクサ Japan: Japan: C. to W. Honshu (westward from Key to the varieties of Hydroco かle yabei Fukushima Pref よ Shikoku ，Kyushu and A. A. Leaves 3-9 mm long ，3-10 mm wide ， Ryukyu. Sunny meadows in lowlands espe- lower lower lobes generally much divaricate; cially near seashors.

ripe ripe fruits ca. 1 mm long.... ・H ・.var. yabei Distr.: Subtropical to tropical regions of A. A. Leaves (5-) 6-12 (-15) mm long ，(6 ー) the Old World. 7-13 (-16) mm wide ， lower lobes gener- ally ally par 叫lel or narrowly divaricate; ma- June June 2001 Journal of Japanese Botany Vo l. 76 No. 3 143

Subfam. Subfam. 11. Saniculoideae Drude Calif. Pub 1. Bo t. 30: 17 ， f. 8 (1 958); Drude in Eng 1. & Prantl ，N a t. Pflanzen- S.L.Liou ， F 1. Reip. Popu 1. Sin. 55 (1): 58 fam. fam. 111 ， 8: 135 (1 898). (1 979); H.Ohba in Iwatsuki & a 1. eds. ， F 1. Jap. Jap. IIc: 272 (1 999). 3. 3. Sanucula L. Sanicul αjaponica Siebold ， Syn. Oecon. L ， Sp. P1.: 235 (1 753). Jap.: 46 (1830) ，nom. nud. Sanicula Sanicula europ αea L. var. chinensis A. A. Stems erect ，several branched; leaves al- (Bunge) Diels in Bo t. Jahrb. Sys t. 29: 491 ternate ， almost cauline ， radical leaves (1 900). few or wanting at flowering (Sec t. 1. Sα nicula elata Ham. var. japonica

Sanicula) …・...・ H ・....・ H ・...・... 1. S. chinensis (Siebold) Koidz. in Bo t. Mag. Tokyo 44: 95 A. A. Stems scape-like ， with one to several (1 930). simple simple or once branched scapes; leaves Sanicula elata Ham. v紅 . chinensis mostly mostly radica 1. (Bunge) Makino ， 11 1. F1. Nippon: 281 (1940) ， B. B. Scape or branches of scape with one comb. nud. normal normal sess i1 e leaf at apex; leaf-teeth ウマノミツノて mucronate mucronate at apex; fruits with slender Japan: Hokkaido ， Honshu ， Shikou ，

hooked pric k1 es or short curved prick 田 Kyushu and Ryukyu (Okinawa Isl ふIn more les les (Sec t. 2. Erythrosanicula). or less wet forests on hills; low altitude to C. C. Leaves 4-15 cm long ，5-20 cm 1000m. wide; wide; fruits with slender hooked Distr.: Japan ，Korea ，N. to C. China. prikles. prikles. Manchuria and Sakhalin. D. D. Leaflets broadly obovate; petals dark dark pu 中 le; involucral bracts Sec t. 2. Erythrosanicula Baill. ，His t. P1. large ，much exceeding the 7: 536 (1 880). .. .. 2. S. rubriflora Sec t. Tuberculatae Durde in Eng 1. & D. Leaflets cuneate-ovate or oblan- Prantl ，Na t. Pflanzenfam. 111 ， 8: 137 (1 898). ceolate; ceolate; petals white; involucral 2. Sanicula rubriflora F. Schmidt ex bracts bracts 1-2 times as long as the Maxim. ， Prim. F1. Amur.: 123 (1 859);

flower...... ・H ・.....・ H ・.. 3. S. kaiensis Schisch k. in F 1. URSS 16: 62 (1950); Hiroe C. C. Leaves 2-5 cm long ，3-7 cm wide; & Constance in Univ. Calif. Pub 1. Bo t. 30: fruits fruits with short hair-like prickles... 16 ，f. 7 (1 958); H.Hara ，Enum. Sperm. Jap.

.. .. 4. S. tuberculat α， 3: 322 (1954); S.L.Liou ， F 1. Reip. Popu 1. Sin. B. B. Scape with one scale-like sessile leaf 55 (1): 37 ，f. 12 (1 979); H.Ohba in Iwatsuki or or one much smaller petiolate leaf at & a1. eds. ， F 1. Jap. IIc: 272 (1999). apex; apex; leaf-teeth with an awn at apex; Sanicula vaniotiana H.Lev. in Bull. Acad. fruits fruits with short straight prickles (Sec t. Geogr. Bo t. 11: 299 (1 902). 3. 3. Pseudopetagnia) … 5. S. lamelligera クロバナウマノミツバ Japan: Japan: C. and N. Honshu (Iwate ，Nagano

Sec t. 1. Sanicula and Yamanashi Pr ，印efs. ふ Genrally on Sec t. Eusanicula Wolff in Pfl.-reich ，IV- limestones; 1000一~150∞Om. 22 ， H t. 61: 61 (1 913). Distr.: Japan ，Korea ，NE. China ，Mongolia 1. 1. Sanicula chinensis Bunge ，Enum. P1. and E. Siberia. Chin. Chin. Bor.: 106 (1833); Schischk. ， F 1. URSS 16: 16: 66 (1 950); H.Hara ，Enum. Sperm. Jap. 3. Sanicula kaiensis 勘1akino et Hisauchi 3: 3: 321 (1954); Hiroe & Constance in Univ. in J. Jpn. Bo t. 5: 366 ， photo. (1 928) ，nom. 144 144 植物研究雑誌第76 巻第3号 平成13 年6月

nud.; nud.; J. Jpn. Bo t. 10: 755 ， photo. 1 (1934); Subfam. 111. Apioideae Drude H.H 紅 a，Enum. Sperm. Jap. 3: 321 (1 954); Drude in Eng l. & Prantle ，N a t. Pflanzen- Hiroe Hiroe & Constance in Univ. Calif. Pub l. Bo t. fam. 111 ， 8: 145 (1898). 30: 30: 19 ，f. 9 (1958); H.Ohba in Iwatsuki & al eds. ， F l. Jap. IIc: 272 (1999). Tribe 1. Scandiceae DC. ヤマナシウマノミツバ DC. ，Prod r. 4: 220 (1 830). Dist r.: C. Honshu (Nagano and Yamanashi Prefs ふGenerally on limestones; 4. Torilis Adans. 1000-1700 1000-1700 m. Endemic. Adans. ，Fam. Pl. 2: 99 (1 763). A. A. Segments of leaves acute (sometimes 4. 4. Sanicula tuberculata Maxim. in Bull. acuminate) at apex; calyx-tube obtuse to Acad. Acad. Sci. St.副 Pe t. 11: 431 (1 867); H.Hara ， rounded at base ， not attenuate into

Enum. Sperm. Jap. 3: 322 (1 954); Hiroe & pedicel; umbels with 3-8 rays; fruits el 四 Constance Constance in Univ. Calif. Pub l. Bo t. 30: 21 ， lipsoid ， 3-3.5 mm long …1. T. J αrponica f. f. 10 (1 958); S.L.Liou ， F l. Reip. Popu l. Sin. A. Segments of leaves acuminate at apex;

55(1): 55(1): 39 ，f. 13 (1979); H.Ohba in Iwatsuki calyx 圃 tube acute at base ， gradually at- & a l. eds. ， F l. Jap. IIc: 273 (1999). tenuate into pedicel; umbels with 2-3 フキヤミツノ f rays; fruits oblong ，4-5 mm long... …・ Japan: Japan: C. to W. Honshu (westward from .. 2. T. scabra Gifu Gifu Pref.) ，Shikoku and N. Kyushu. In for- ests ests on mountains. 1. Torilis japonica (Hout t.) DC. ， Prodr. Dsitr.: Dsitr.: Japan ，Korea and NE. China. 4: 219 (1830); H.H 紅 a，Enum. Sperm. Jap. 3: 3: 328 (1 954); Hiroe & Constance in Univ. Sec t. 3. Pseudopetagnia Wolff. in Eng l.， Calif. Pub l. Bo t. 30: 24 ， f. 12 (1 958);

Pfl.-reich ，IV 闘 22 ， H t. 61: 52 (1913). H.Ohba in Iwatsuki & al. eds. ， F l. Jap. IIc: 5. 5. Sanicula lamelligera Hance in J. Bo t. 274 (1999). 8: 8: 11 (1 878); H.Hara ，Enum. Sperm. Jap. 3: Tordylium anthriscus L. ， Sp. Pl.: 240 321 321 (1 954); Hiroe & Constance in Univ. (1 753). Calif. Calif. Pub l. Bo t. 30: 21 ， f. 11 (1958); Caucalis japonica Hout t.， Na t. Hist r. 11 ， S.L.Liou ， F l. Reip. Popu l. Sin. 55(1): 48 ，f. 8: 42: t. 45 ，f. 1 (1777). 20 (1979); H.Ohba in Iwatsuki & al. eds. ， F l. Torilis anthriscus auc t. non Gaertner Jap. Jap. IIc: 273 (1 999). (1 788): (L.) Gmelin ， F l. Bad. 1: 613 (1 805). Sanicula Sanicula satsumana Maxim. in Bul l. ヤブジラミ Acad. Acad. Sc i. St. -Pe t. 21: 47 (1 886). J apan: Hokkaido ， Honshu ， Shikoku ， Sanicula Sanicula wakayamensis Masam. in Trans. Kyushu and Ryukyu. Roadsides and waste Na t. His t. Formos. 31: 340 (1941). places at lowlands. Sanicula Sanicula lamelligera Hance var. Distr.: Common weeds in temperate to wakayamensis wakayamensis (Masam.) Murata in Ac t. subtropical regions from N. Africa ，Europe Phytotax. Phytotax. Geobo t. 20: 195 (1962). to E. Asia ，naturalized in N. America. ヒメウマノミツノ f J apan: Honshu (Wakayama Pre f.よ Kyushu 2. Torilis scabra (Thunb.) DC. ， Prodr. 4: (Yaku-shima (Yaku-shima and Tanega-shima) and 219 (1830); H.Hara ，Enum. Sperm. Jap. 3: Ryukyu. In thickets along s佐eams; lowlands 328 (1954); Hiroe & Constance in Univ. to to 500 m. Calif. Pub l. Bo t. 30: 26 ， f. 13 (1958); DistL: DistL: Japan ，Taiwan and C. to W. China. H.Ohba in Iwatsuki & al. eds. ， F l. Jap. IIc: 274 (1999). June June 2001 Journal of Japanese Botany Vo l. 76 No. 3 145

Chaerophyllum Chaerophyllum scabrum Thunb. ， F l. Jap.: (1 979); Yamazaki in 1. Jpn. Bo t. 76: 175 119 (1 784) ， p. p. (2001).

Anthriscus Anthriscus scα br α(Thunb.)， Kozo-Polj. in Chaerophyllum nemorosum M.Bieb. ， F l. Bull. Bull. Soc. Na t. Moscou ， n. s. 29: 151 (1 916). Taur.-Cauc. 1: 232 (1 808). オヤブジラミ Anthriscus sylvestris (L.) Hoffm. var. Japan: Japan: Honshu ， Shikoku ， Kyushu and nemorosa Trautv. in Tr. Bo t. Gard. 5: 437 Ryukyu. Roadsides or waste places at low- (1 877). lands. lands. Anthriscus sylvestris (L.) Hoffm. subsp.

Dist r.: Japan ， S. Korea ，Taiwan and C. to nemorosa (M.Bieb.) Koso ・PO り. in Ac t. Hort. S. S. China. Petrop. 34: 103 (1 920); H.Hara ，Enum. Sperm. Jap. 3: 299 (954). 5. 5. Anthriscus (Pers.) Hoffm. Anthriscus sylvestris (L.) Hoffm. var. Hoffm. ，Gen. Umbell. ed ， 1: 38 (1814). aemula W om. in Fl. South East Europe ， Part URSS 5: 770 (1931). A. Fruits black or dark brown when ripe ， Anthriscus aemula (Wom) Schischk. in Fl.

smooth..... ・H ・...・ H ・-… H ・H ・... 1. A. sylvestris URSS 16: 129 (1950); Kitag. ，Neo ・Lineam. A. Fruits yellowish brown when ripe ， Fl. Manshuricae: 477 (1 979) ， cum f.

tuberuculate-hirsute.. tuberuculate-hirsute.. .・ H ・. 2. A. nemorosa hirsut 折uctus (Ohwi) Kitag. Anthriscus Anthriscus nemoros α(M.Bieb.) Spreng. 1. 1. Anthriscus sylvestris (L.) Hoffm. ， V 紅 . hirsut 折uctus Ohwi in Bull. Sci. Mus. Gen. Umbell. ed .1: 40 (1 814); S.L.Liou ， F l. Tokyo ， no. 33: 81 (1953). Reip. Reip. Popu l. Sin. 55 (1): 74 (1 979); Ohwi ， F l. Anthriscus sylvestris (L.) Hoffm. subsp. Jap. Jap. ed. Eng l.: 671 (1 965); H.H 訂 a，Enum. nemorosa (M.Bieb.) Koso-Polj. var. Sperm. Jap. 3: 299 (1 954); Hiroe & hirsut{ 斤uctus (Ohwi) H.Hara ，Enum. Sperm. Constance in Univ. Calif. Pub l. Bo t. 30: 26 ， Jap. 3: 300 (1 954). f. 14 (1 958) ， exc l. syn. A. nemorosa Anthriscus sylvestris (L.) Hoffm. f. (M.Bieb.) (M.Bieb.) Spreng.; H.Ohba in Iwatsuki & a l. hirsut{ 斤uctus (Ohwi) H.Ohba in Iwatsuki & eds. ， F l. Jap. IIc: 278 (1 999). al. eds. ， F l. Jap. IIc: 278 (1 999). Anthriscus Anthriscus sylvestris (L.) Hoffm. subsp. オニジャク，ケジャク aemula auc t. non Woron.: Ki tam. in Ac t. J apan: Sakhalin ， Hokkaido ， Honshu ， Phytotax. Phytotax. Geobo t. 20: 195 (1 962). Shikoku and Kyushh. Anthriscus Anthriscus sylvestris (L.) Hoffm. subsp. Distr.: Japan ，China ，Sakhalin ，Mongolia ， nemorosa auc t. non M.Bieb.: H.Hara ，Enum. S. Russia ，Balkan Peninsula and It aly. Sperm. Jap. 3: 299 (1954). シャク 6. Osmorhiza Rafin. Japan: Japan: S. Kuriles ， Hokkaido ， Honshu ， Rafin. ， in Joum. de Phys. 89: 257 (1 819). Shikoku and Kyushu. Edges of forests on mountains ，more or less wet places; low alti- 1. Osmorhiza aristata (Thunb.) Rydb. in tudes tudes to 1000 m. Bo t. Surv. Nebraska 3: 37 (1894); H.Ohba in Distr.: Distr.: Japan ，Korea ，NE. ，N. to C. China ， Iwatsuki & al. eds. ， F l. Jap. IIc: 275 (1999). Mongolia to Europe. Chaerophyllum αristatum Thunb. ， F l. Jap.: 119 119 (1 784). 2. 2. Anthriscus nemorosa (M.Bieb.) Osmorhiza japonica Siebold & Zucc. in Spreng. ， Pl. Umbell. Prodr.: 27 (1813); Abh ，Akad ，Muench. 4 (2): 203 (1845). Schischk. ， Fl. URSS 16: 127 (1950); Osmorhiz αα murensis F .Schmidt ex S.L.Liou ， F l. Reip. Popu l. Sin. 55(1): 75 Maxim. ， Prim. Fl. Amur.: 129 (1859). 146 146 植物研究雑誌第76 巻第3号 平成13 年6月

Osmorhiza Osmorhiza aristata (Thunb.) M 紘 ino & Schischk. ， F l. URSS 16: 232 (1950); Ohwi ， Yatabe in Bo t. Mag. Tokyo 17: 14 (1903); Fl. Jap. ed. 2: 983 (1975); K.T.Fu & Y.C.Ho ， Hiroe Hiroe & Constance in Univ. Calif. Pub l. Bo t. Reip. Popu l. Sin. 55(1): 166 ，f. 86 (1997). 30: 30: 30 ，f. 15 (1958); Schischk. ， F l. URSS Pleurospermum uralense auc t. non 16: 16: 140 (1950); H.Hara. Enum. Sperm. Jap. Hoffm.: Hulten ， F l. Kamtch. 3: 155 (1 929). 3: 3: 315 (1 954); S.L.Liou ， F l. Reipu l. Popu l. Pleurospermum austriacum (L.) Hoffm. Sin. Sin. 55 (1): 77 ，f. 34 (1997). subsp. u1 ・alense auc t. non Hoffm.: H.Hara in V 紅. aristata. J. Fac. Sci. Univ. Tokyo ，sec t. 3， 6: 93 ヤブニンジン (1952); H.Ohba in Iwatsuki & al. eds. ， F l. J apan: Hokkaido ，Honshu ， Shikoku and Jap. IIc: 284 (1 999). Kyushu. Kyushu. In forests on hills and mountains; Pleurospermum austriacum auc t. non L.: low altitude to 1800 m. Hiroe & Constance in Univ. Cali f. Pubu l. Distr.: Distr.: Japan ，Korea ，Taiwan ， C. to N. Bo t. 30: 81 ，f. 43 (1958). China ，Mongolia ， E. Siberia and Sakhalin. オオカサモチ V 紅. montana Makino in J. Jpn. Bo t. 2: 7 Japan: S. Kuriles ，Hokkaido ，N. to C. (1918). (1918). Honshu. In mountain to alpine meadows; Osmorhiza Osmorhiza montan α(Makino) Makino in 700-2600 m in Honshu ，low altitudes to J. J. Jpn. Bo t. 5: 28 (1928). 1500 m in Hokkaido. Osmorhiza Osmorhiza amurensis F.Schmidt var. Distr.: Japan ，Korea ，NE. China ，Kuriles ， mon ωna (Makino) Kitag. in Rep. Ins t. Sci. Kamchatka ， Sakhalin and E. Siberia. Res. Res. Manch. 1: 1 (1936). ミヤマヤブニンジン Tribe 3. Ammineae Koch. Japan: Japan: S. Kuriles ，Hokkaido and N. to C. Koch ， Gen. Umbe l.: 114 (1 824); Drude in Honshu. Honshu. In forests on mountains; 500-2000 Eng l. & Prantl ，Na t. Pflanzenfam. 111 ， 8: 178 m. (1 898). Dist r.: Japan ，Korea ，NE. China ，Sakhalin ， U ssuri and Amur. Subtribe 1. Carinae Drude Drude in Eng l. & Prantl ， N a t.

Key to the varieties of Osmorhiza arist α， ta Pflanzenfam. 111 ， 8: 183 (1 898). A. A. Stems and pedicels pubescent; ultimate

segments segments of leaves acute... …V 紅 . aristata 8. Bupleurum L. A. Stems and pedicels glabrescent; ultimate L. ， Sp. Pl.: 236 (1 753). segments segments of leaves acuminate. ……. ....var. ....var. montana A. Cauline leaves dilated and clasping the stem at base. Tribe Tribe 2. Smyrnieae Koch B. Stems 7-15 cm tall; bracteoles orbicu- Koch ， Gen. Umbe l.: 133 (1 824); Drude in lar-ovate 5-6 mm long ，4-5 mm wide at Eng l. & Prantl ，Na t. Pflanzenfam. 111 ， 8: 147 fruiting ， ca. 2 times as long as the frui t... (1 898). . 1. B. ajanense B. B. Stems 30-100 cm tall; bracteoles 7. 7. Pleurospermum Hoffm. ovate to lanceolate ， 1-7 mm long ，0.5- Hoffm. ， Gen. Umbe l. ed. 1: 8 (1 814). 4 mm wide at fruiting ，1-2 times as long long as the frui t. 1. 1. Pleurospermum camtschaticum C. Stems 20-60 cm tall; bracteoles Hoffm. ， Gen. Umbe l. ed. 1: 10 (1 814); ovate to oblong-lanceolate ，4-7 mm F. Schmidt ， Reis. Sachal: 140 (1868); long ，2 -4 mm wide ， 1-2 times as June June 2001 Journal of Japanese Botany Vo 1. 76 No. 3 147

long long as the frui t. Y.Li. & M.L.Sheh ， F l. Reip. Popu l. Sin. 55 D. D. Cauline leaves oblong -ovate to (1): 237 (1 979) ， pro p紅白; Hiroe &

ovate ，abruptly acute at apex ，dis 由 Constance in Univ. Calif. Pub l. Bo t. 30: 55 ， tinctly tinctly dilated at base; bracteoles f. 29 (1 958); H.Ohba in Iwatsuki & al. eds. ，

broadly broadly ovate ， acute ...・ H ・ Fl. Jap. IIc: 276 (1999)...... 2. B. nipponicum レブンサイコ D. D. Cauline leaves lanceolate ， Japan: S. Kuriles ， Hokkaido and N. attenuate-acumlnate attenuate-acumlnate at apex ， Honshu (M t. Shirakami). More or less wet slightly slightly dilated at base; bracteo- rocky places.; 1000-1700 m. les les triangular-lanceolate ，aristate- Distr.: Japan ， Kuriles ， Kamchatska ，

acuminate acuminate ...・ H ・-….. 3. B. yesoense Sakhalin and E. Siberia. C. C. Stems 40-100 cm tall; bracteoles lanceolate lanceolate to ovate ， 1-3(-5) mm 2. Bupleurum nipponicum Koso-Polj. in long ， 0.5-2.5 mm wide at fruiting ， J. Russ. Bo t. 1913: 1 ，t. 1 (1 913); in Fedch. ， shorter shorter or slightly longer than the F l. t. Asia Ross. 10: 31 ，t. 12 (1 915); H.Har 丸

fruit.... ・H ・H ・H ・-… 4. B. longiradiatum Enunl. Sperm. Jap. 3: 302 (1 954); H.Ohba in A. A. Cauline leaves gradually narrowing and Iwatsuki & al. eds. ， F l. Jap. IIc: 277 (1999). not not clasping the stem at base. Bupleurum longiradiatum Turcz. subsp. B. B. Middle cauline leaves lanceolate ，6- 10 sachalinense (F.Schmidt) Kitag. var. cm long ，0.7- 1. 5 cm wide ， abruptly pseudon 伊'P inicum Kitag. in J. Jpn. Bo t. 38: narrowing narrowing upward ， acute or short- 306 (1 963). acuminate acuminate at apex ， apical awns 0.5-1 ハクサンサイコ

mm long... ・H ・-…....・ H ・-… 5. B. chinense Distr.: C. and N. Honshu. In subalpine to B. B. Middle cauline leaves linear- alpine meadows; 1600-2400 m. Endemic. lanceolate ，7: -15 cm long ，0.3- 1. 5 cm Note. A 'form with lanceolate acuminate wide ， gradually nerrowing upward ， bractoles of umbellets has been named f. attenuate-acuminate attenuate-acuminate at apex ， apical stenolepis Ki tag. (Jap. Name: Shinano-saiko ， awns 1-2 mm long..... J. Jpn. Bo t. 44: 194 ， 1969). .. .. 6. B. scorzonel ゆ lium 3. 3. Bupleurum yesoense Nakai ex H.Hara 1. 1. Bupleurum ajanense (Regel) Kr asnob. in J. Jpn. Bo t. 9: 515 ，f. 3 (1 933). exYamazaki in J. Jpn. Bo t. 70: 341 (1995). Bupleurum yesoense N akai ，Vege t. Apoi: Bupleurum ajanense (Regel) Kr asnob. in 30 ，62 (1 930) ，nom. nud. XV Intern. Bo t. Cong. Abst r.: 255 (1 993) ， Bupleurum nipponicum Kozo-Polj. var. nom. illeg. yesoense (Nakai) H.Hara in J. Jpn. Bo t. 9: Bupleurum ranunculoides L. var. y 515 (1 933) ， pro syn.; H.Ohba in Iwatsuki & ajanense ajanense Regel in Regel & Tiling ， F l. Ajan.: al. eds. ， F l. Jap. IIc: 277 (1 999). 97 (1 858). エゾサイコ Bupleurum triradiatum Adams ex Hoffm. Distr.: Hokkaido. In alpine meadows. V 紅 .α かinum auc t. non Rupr.: F.Schmidt ， F l. Endeinic. Sacha l.: 135 (1868); H.Hara ，Enum. Sperm. Jap. Jap. 3: 303 (1954). 4. Bupleurum longiradiatum Turcz. in Bupleurum triradiatum Adams ex Hoffm. Bull. Soc. Na t. Mosc. 17: 719 (F l. Baic. V 紅 . ajanense (Regel) Kozo-Polj. in Ac t. Dahur. ， no. 505) (1 844). Hort. Hort. Peterop. 30: 200 (1914). subsp. sachalinense (F.Schmidt) Kitag. in Bupleurum triradiatum auc t. non Adams: J. Jpn. Bo t. 36: 241 (1 961). 148 148 植物研究雑誌第76 巻第3号 平成13 年6月

Bupleurum sachalinense F.Schmid t.， Reis. A. Pedicels 4- 5 (-6) mm long at fruiting; Amur. Sacha l.: 135 (1 868). petals pale yellow. supleurum longiradiatum Turcz. var. B. Bracteoles ovate ， 2.5-5 mm long ， sachalinense sachalinense (F.Schmidt) Boiss. in Bull. longer than the flower......

Soc. Soc. Bo t. Fr. 57: 413 (1910) ， in nota. ・H ・H ・..var. sachalinense Bupleurum longiradiatum Turcz. f. B. Bracteoles narrowly ovate ，1. 5-2 mm sachalinense sachalinense (F.Schmidt) Nakai in Bo t. Mag long ， as long as or shorter than the

Tokyo 50: 367 (1936). flower. …....・ H ・.....・ H ・...・ H ・-… .var. elatius Bupleurum longiradiatum Turcz. var. A. Pedicels 1. 5-2 mm long at fruiting; pet-

breviradiatum breviradiatum auc t. non F. Schmid t: Ohwi ， als light yellow...... ・H ・..var. shikotanense l. Fl. Jap. ed. Eng l.: 670 (1965); H.Ohba in Iwatsuki Iwatsuki & al. eds. ， F l. Jap. IIc: 276 (1999). 5. Bupleurum chinense DC. ， Prodr. 4: var. var. sachalinense. 128 (1830); Y.Li & M.L.Sheh ， F l. Reip. エゾホタルサイコ Popu l. Sin. 55 (1): 290 (1979). Distr.: Distr.: N. Honshu ， SE. Hokkaido and Bupleurum scorzonerifolium Willd. 五 Sakhalin. Sakhalin. latum Nakai in 1. Jpn. Bo t. 13: 486 (1 937). V 訂. elatius (Kozo ・Polj.) Ki tag. in J. Jpn. Bupleurum stenophyllum (Nakai) Ki tag. Bo t. 36: 242 (1961). var. kiusianum Ki tag. in 1. Jpn.Bo t. 21: 100 Bupleurum longiradiatum Turcz. var. (1 947). breviradiatum breviradiatum F.Schmidt f. elatius Kozo- Bupleurum falcatum L. f. kiusianum Polj. Polj. in Bull. Soc. Na t. Mosc. n. s. 29: 60 (Kitag.) Hiroe in Ac t. Phytotax. Geobo t. 14: (1 916). 145 (1 952). Bupleurum longiradiatum auc t. non ツクシサイコ，マンシュウミシマサイコ Turcz. ， Hiroe & Constance in Univ. Calif. J apan: W. Honshu (Yamaguchi Pref.) and Pub l. Bo t. 30: 55 ，f. 28 (1 958) ， pro p紅白. Kyushu. In sunny meadows on mountains; ホタルサイコ 400-500 m. J apan: HOkkaido ，Honshu ， Shikoku and Dist r.: Japan ，Korea ，NE. and C. to N. Kyushu. Kyushu. In meadows on hills and mountains; China. low altitude to 2000 m. Endemic. var. var. shikotanense (Hiroe) Ohwi in Bull. 6. Bupleurum scorzonerifolium Willd. ， Natn. Natn. Sci. Mus. Tokyo no. 33: 80 (1953). Enum. Hort. Bero l. supp l.: 300 (1 809); Bupleurum shikotanense Hiroe in Ac t. Kitag. in J. Jpn. Bo t. 21: 98 (1944); Y.Li & Phytotax. Phytotax. Geobo t. 14: 145 (1952). M.L.Sheh ， F l. Reip. Popu l. Sin. 55 (1): 267 ，f. コガネサイコ 145 (1979). Dsit r.: S. Kuriles and NE. Hokkaido Bupleurum falcatum L. var. scorzoneri- (Shiretoko (Shiretoko Peninsula). In meadows on stony folium (Willd.) Ledeb. ， F l. Ross. 2: 267 slopes. slopes. (1 844) ， ut s; Regel ，Ten t. Fl. Ussur.: 69 Note: Note: Subsp. longiradiatum in Korea ，NE. (1861); Hiroe & Constance in Univ. Calif. China and SE. Siberia differs from subsp. Pub l. Bo t. 30: 53 ，f. 27 (1958). sachalinense sachalinense by the small linear-lanceolate Bupleurum falcatum L. subsp. scorzoneri-

bracteoles bracteoles (1-2 mm long) and longer folium (Willd.) Koso 幽 Po l. in Ac t. Hort. pedicels pedicels (5-7 mm long at fruiting). Petrop. 30: 219 (1914). Bupleurum falcatum L. subsp. scorzoneri-

Key to the .v arieties of folium (Willd.) Kozo ・Polj. v紅 . komarowii Bupleurum longiradiatum subsp. Koso-Po l. in Bul l. Soc. Na t. Mosc. ， n. s. 29: sacha /i nense 59 (1916); H.H 紅 a，Enum. Sperm. Jap. 3: June June 2001 Journal of Japanese Botany Vo l. 76 No. 3 149

300 (1 954). less wet places; lowlands to 800 m. Bupleurum scorzonerifolium Willd. va r. Distr.: Japan ，Korea ，Taiwan and China. stenophyllum stenophyllum Nakai in J. Jpn. Bo t. 13: 487 Note: A form with deeply lobed leaves has (1937). (1937). been named f. dissecta (Y abe) H.Hara Bupleurum scorzonerifolium Willd. v訂. (Enum. Sperm. Jap. 3: 309 ， 1954) and with obvallatum obvallatum Nakai in J. Jpn. Bo t. 13: 488 dark pu 中le leaves as f. atropurpurea (1937). (1937). (Makino) Ohwi ex H.H 紅 a (Enum. Sper. Jap. Bupleurum stenophyllum (Nakai) Ki tag. in 3: 309 ， 1954). Cryptotaenia canadensis (L.) J. J. Jpn. Bo t. 21: 99 (1948). DC. in N. America closely resembles C. ja- Bupleurum falcatum auc t. non L.: H.Ohba ponica ， but differs from the latter by fre- in in Iwatsuki & a l. eds. ， F l. Jap. IIc: 275 ， pro quently bi-lobed or parted leaflets with p紅白. distinct petiolules ， and oblong-cylindric ミシマサイコ fruits with persistent ca. 1 mm long styles. apan.: Japan.: Honshu (westward from Fukushima Pref. and Toyama Pre f.よ Shikoku 10. PternopetaIum Fr. and Kyushu. In meadows on hills; low Franch. in Nouv. Arch. Mus. Histr. Na t. altidudes altidudes to 900 m. Paris ，ser. 2， 8: 246 (1 886). Distr.: Distr.: Japan ，Korea ，NE. ，N. to C. China ， Mongolia ， E. Siberia and Dahuria. 1. Pternopetalum tana 匙ae (Franch. & Sav.) Sav.) Hand.-Mazz. ，Symb. Sinic. 7-3: 719 9. 9. Cryptotaenia DC. (1933); H.Hara ，Enum. Sperm. Jap. 3: 320 DC. ，Mem. Ombe l.: 42 (1 829) ，nom. cons. (1954); Hiroe & Constance in Univ. Cali f. Deringa Deringa Adans. ，Fam. Pl. 11: 498 (1 763) ， Pub l. Bo t. 30: 37 ，f. 18 (1 958); F.T.Pu ， F l. nom. r句ec. Reip. .Popu 1. Sin. 55(2): 57 (1 985); H.Ohba in in Iwatsuki & al. eds. ， F l. Jap. IIc: 278 1. 1. Cryptotaenia japonica Hass k. in (1999). Retzia Retzia 1: 113 (1855); H.Hara ，Enum. Sperm. Carum tanakae Franch. & Sav. ，Enum. Pl. Jap. Jap. 3: 308 (1 954); S.L.Liou ， F l. Reip. Popu l. Jap. 2: 371 (1877). Sin. Sin. 55(2): 19 (1 985). Pimpinella tanakae (Franch. & Sav.) Diels Cryptotaenia Cryptotaenia canadensis auc t. non DC.: in Eng l. Bo t. Jahrb. 29: 494 (1 900) ，exc l. Siebold Siebold & Zucc. in Abh. Akad. Muench. 4 speclm. (2): (2): 203 (1845); Buwarda in Fl. Males. 4(2): Cryptotaeniopsis tanakae (Franch. & 133 133 (1 949); Hiroe & Constance in Univ. Sav.) Boiss. in Bull. Herb. Boiss. ser. 2， 2: Cali f. Pub l. Bo t. 30: 32 ，f. 16 (1958). 806 (1 902); Hiyama in Yaso 4: 41 (1 938) ， Cryptotaenia Cryptotaenia canadensis (L.) DC. var. ja- cum f. conformis Hiyama et f. linearis ponica ponica (Hassk.) M 紘 ino in Bo t. Mag. Tokyo Hiyama. 22: 22: 175 (1908). イワセントウソウ Deringa Deringa japonica (Hass k.) Koso-Po l. in Japan: C. to W. Honshu (west of Gunma Bull. Bull. Soc. Na t. Moscou ， n. s. 29: 136 (1916). Pref よ Shikoku and Kyushu. In forests on Cryptotaenia Cryptotaenia canadensis (L.) DC. subsp. mountains; 1000-2400 m. jα rponica (Hassk.) Hand.-Mazz. ，Symb. Sin. Dsitr.: Japan ， S. Korea and C. China. 7-3: 7-3: 713 (1933). ミツノ T 11. Aegopodium L. Japan: Japan: S. Kuriles ，Hokkaido ，Honshu ， L. ， Sp. Pl.: 265 (1753). Shikoku Shikoku and Kyushu ， probably escaped in the the Ryukyus. In secondarey forests ，more or A. Leaves bi-ternate ， pubescent beneath; 150 150 植物研究雑誌第76 巻第3号 平成13 年6月

leaflets leaflets irregularly acute-se 町 ate ，central 3: 291 (1 954); Hiroe & Constance in Univ. leaflets leaflets 3-8 cm long ，1. 5-5 cm wide.... … Cali f. Pub l. Bo t. 30: 50 ， f. 25 (1 958); 1. . 1. A. podagraria S.L.Liou ， F l. Reip. Popu l. Sin. 55(2): 138 ，f. A. A. Leaves 2-3 times ternately pinnate; 56 1-3 (1985); Schischk. ， F l. URSS 16: 457

leaflets leaflets glabrous on both surfaces ，irregu 聞 (1950); H.Ohba in Iwatsuki & al. eds. ， F l. larly larly incised-lobed ， 0.8-3 cm long ，0. 4- Jap. IIc: 281 (1 999).

1. 5 cm wide...... ・H ・-…....・ H ・.2. A. alJ フestre Cα rum alJ フestre (Ledeb.) Koso-Po l. in Bull. Bull. Soc. Na t. Moscou 29: 199 (1916). 1. 1. L. ， Sp. Pl.: Aegopodium αかestre Ledeb. f. tenera 265 (1 753); Hiroe & Constance in Univ. H.H 訂 a in Bo t. Mag. Tokyo.50: 365 (1936). Calif. Calif. Pub l. Bo t. 30: 49 (1 958). エゾボウウフウ イワミツバ Japan: S. Kuriles ， Hokkaido and N. to C. Japan: Japan: A native of Europe ，naturalized in Honshu. In forests;1000-1800 m in Honshu ， Hokkaido (Sapporo). 200-1000 m in Hokkaido. Distr.: Distr.: Japan ， Sakhalin ， Kuriles ， 2. 2. Aegopodium alpestre Ledeb. ， F l. Al t. Kamchatka ， N. Korea ，NE. China ， Dahuria 1: 1: 354 (1 829); H.H 紅 a，Enum. Sperm. Jap. and W. to E. Siberia.

山崎敬:日本のセリ科植物 I 日本のセリ科植物は約30 属90 種ほど知られて ら日本のセリ科植物を再検討した.ここでは日本 いる.これを纏めた論文の多くは，永年この科の 産の属の検索表と，チドメグサ属，ツボクサ属， 研究を続けていた北川政夫氏の結論を基礎として ウマノミツバ属，ヤブジラミ属，シャク属，ヤブ いる.しかしその見解は氏独自のもので，ロシア ニンジン属，オオカサモチ属， ミシマサイコ属， や中国の研究者とのあいだにいくつかの異なった ミツバ属，イワセントウソウ属，エゾボウフウ属 見解が見られる.これらの相違を検討して妥当な の11 属を扱い，各属内の種及び変種の検索表を掲 学名を選択しなければならない.こうした観点か げた (165_" 東京都中野区ーーーーーーー)