Zootaxa, a Taxonomic Catalogue of the Palaearctic Bees of the Tribe Osmiini

ZOOTAXA

1865

A taxonomic catalogue of the Palaearctic bees of the tribe Osmiini (Hymenoptera: Apoidea: Megachilidae)

STEFAN UNGRICHT, ANDREAS MÜLLER & SILVIA DORN

Magnolia Press Auckland, New Zealand STEFAN UNGRICHT, ANDREAS MÜLLER & SILVIA DORN A taxonomic catalogue of the Palaearctic bees of the tribe Osmiini (Hymenoptera: Apoidea: Megachilidae) (Zootaxa 1865) 253 pp.; 30 cm. 3 Sept. 2008 ISBN 978-1-86977-255-0 (paperback) ISBN 978-1-86977-256-7 (Online edition)

FIRST PUBLISHED IN 2008 BY Magnolia Press P.O. Box 41-383 Auckland 1346 New Zealand e-mail: [email protected] http://www.mapress.com/zootaxa/

© 2008 Magnolia Press All rights reserved. No part of this publication may be reproduced, stored, transmitted or disseminated, in any form, or by any means, without prior written permission from the publisher, to whom all requests to reproduce copyright material should be directed in writing. This authorization does not extend to any other kind of copying, by any means, in any form, and for any purpose other than private research use.

ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition)

2 · Zootaxa 1865 © 2008 Magnolia Press UNGRICHT ET AL. Zootaxa 1865: 1–253 (2008) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA Copyright © 2008 · Magnolia Press ISSN 1175-5334 (online edition)

A taxonomic catalogue of the Palaearctic bees of the tribe Osmiini (Hymenoptera: Apoidea: Megachilidae)

STEFAN UNGRICHT, ANDREAS MÜLLER & SILVIA DORN ETH Zurich, Institute of Plant Sciences, Applied Entomology, Schmelzbergstrasse 9/LFO, CH-8092 Zurich, Switzerland [email protected] [email protected] [email protected] Corresponding author: Andreas Müller, email: [email protected]

Table of contents

Abstract...... 4 Introduction ...... 4 Rationale and scope of the catalogue...... 4 Placement and classification of the Osmiini...... 9 Synopsis...... 18 Taxonomic inventory of the Palaearctic Osmiini ...... 18 Faunistic inventory of the Palaearctic Osmiini...... 20 Sources...... 21 Primary nomenclatural sources...... 21 Secondary nomenclatural sources...... 22 Conventions ...... 23 Glossary of nomenclatural terms...... 23 Format of the catalogue ...... 24 Abbreviations in the catalogue ...... 25 Family-group names ...... 27 Genus-group names...... 28 Species-group names...... 39 Chelostoma LATREILLE ...... 39 Haetosmia POPOV ...... 50 Heriades SPINOLA...... 51 Hofferia TKALCU ...... 57 Hoplitis KLUG ...... 58 Hoplosmia THOMSON ...... 131 Ochreriades MAVROMOUSTAKIS ...... 136 Osmia PANZER ...... 137 Protosmia DUCKE ...... 200 Pseudoheriades PETERS...... 206 Stenoheriades TKALCU ...... 206 Stenosmia MICHENER...... 207 Wainia TKALCU...... 209

Accepted by L. Packer: 26 May 2008; published: 3 Sept. 2008 3 Fossil taxa...... 210 Incertae sedis ...... 210 Appendix ...... 212 Nomina dubia...... 212 Nomina nuda...... 214 Osmiini erroneously attributed to the Palaearctic region ...... 215 Palaearctic taxa erroneously attributed to the Osmiini...... 215 Acknowledgements...... 216 References ...... 217 Index ...... 236

Abstract

This taxonomic catalogue covers all family-, genus- and species-group names of the Palaearctic bees of the tribe Osmiini as published by the end of 2006. As the taxonomy of the Palaearctic Osmiini is currently in a poor state, the primary goal of this catalogue is to provide a complete coverage of the widely dispersed and often not easily accessible taxonomic literature, thus laying the basis for further taxonomic work. We therefore refrain from proposing new synonyms, new names, new ranks or new combinations, and the taxa accepted here are based on a literature survey, generally adopting the most recent published opinion. The generic and subgeneric system proposed by Michener (2000) serves as a general backbone for this catalogue. We list six available family-group names, 93 available genus-group names and 935 available species-group names that currently split up into the valid names of two subtribes, 13 genera, 43 non-nominotypical subgenera, 604 species and 76 non-nominotypical subspecies. Starting with Linnaeus (1758), a total of 99 mainly Euro- pean taxonomists contributed to the available species-group names by the end of 2006. Taxon accounts provide the refer- ence to the original description, the name-bearing type(s), distribution, and literature sources for species identification. Apart from the extant taxa the catalogue also treats the extinct representatives of the osmiine bees following the tribal classification of Engel (2005).

Key words: Apiformes, classification, distribution, fossils, mason bees, nomenclature, osmiine bees, pollinators, synon- ymy, taxonomic history, types

Introduction

“The Palearctic [bee] fauna is the most difficult in the world to study because of the lack of catalogues […] combined with the great number of species, many of them named long ago […].” —Charles D. Michener (2000: 113)

Rationale and scope of the catalogue

Objects without names cannot be communicated about in an unambiguous manner. Knowledge on taxa that has accumulated for centuries in particular will remain inaccessible without correct scientific names (Winston, 1999). The most important function of taxonomy (i.e., the assignment of organisms to taxa) and in particular of nomenclature (i.e., the naming of the recognized taxa) is simply the provision of a universal framework for information retrieval. As a vehicle for information exchange, nomenclature is one of the most basic tasks of systematic biology to ensure an up-to-date directory of valid names. In fact, taxonomy is rather unique in biology in that it is fundamentally retrospective, evaluating all previous works with nomenclatural bearing since Linnaeus’s Systema Naturae (1758), published 250 years ago.