Mammalia International Journal of the Systematics, Biology and Ecology of Mammals

MAMMALIA INTERNATIONAL JOURNAL OF THE SYSTEMATICS, BIOLOGY AND ECOLOGY OF MAMMALS

EDITOR-IN-CHIEF Christiane Denys, France

ASSOCIATE EDITORS EDITORIAL BOARD Stéphane Aulagnier, France Giovanni Amori, Italy Josef Bryja, Czech Republic Ernesto Capanna, Italy Patrick Gouat, France Louise H. Emmons, USA Laurent Granjon, France Steve M. Goodman, Madagascar Boris Krasnov, Israel John Gurnell, United Kingdom Boris Krysˇtufek, Slovenia Heikki Henttonen, Finland Géraldine Veron, France Herwig Leirs, Belgium Darrin Lunde, USA François Moutou, France Georgy I. Shenbrot, Israel Peter J. Taylor, South Africa Marcus Vinicius Vieira, Brazil ABSTRACTED/INDEXED IN AGRICOLA (National Agricultural Library) · Baidu Scholar · Biobase · Cabell's Directory · CABI (over 50 subsections) · CNKI Scholar (China National Knowledge Infrastructure) · CNPIEC · Dimensions · EBSCO (relevant databases) · EBSCO Discovery Service · Engineering Village · Gale/Cengage · Genamics JournalSeek · Geobase · GeoRef · Google Scholar · Japan Science and Technology Agency (JST) · J-Gate · Journal Citation Reports/Science Edition · JournalGuide · JournalTOCs · KESLI- NDSL (Korean National Discovery for Science Leaders) · Microsoft Academic · Naviga (Softweco) · Primo Central (ExLibris) · ProQuest (relevant databases) · Publons · ReadCube · Reaxys · SCImago (SJR) · SCOPUS · Sherpa/RoMEO · Summon (Serials Solutions/ProQuest) · TDNet · Ulrich's Periodicals Directory/ulrichsweb · WanFang Data · Web of Science: Biological Abstracts; BIOSIS Previews; Current Contents/Agriculture, Biology, and Environmental Sciences; Science Citation Index; Science Citation Index Expanded; Zoological Record · WorldCat (OCLC)

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ISSN 0025-1461 ∙ e-ISSN1864-1547 ∙ CODENMAMLAN

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RESPONSIBLE EDITOR Prof. Dr. Christiane Denys, Muséum National d'Histoire Naturelle, Systematics and Evolution, 55, rue Buffon, 75231 Paris cedex 05, France, Tel. : +33 (0) 1 40 79 30 85, Email: [email protected]

JOURNAL MANAGER Dr. Gunda Stöber, De Gruyter, Genthiner Straße 13, 10785 Berlin, Germany, Tel.: +49 (0)30 260 05 – 279, Fax: +49 (0)30 260 05 – 298, Email: [email protected]

RESPONSIBLE FOR ADVERTISEMENTS Claudia Neumann, De Gruyter, Genthiner Straße 13, 10785 Berlin, Germany. Tel.: +49 (0)30 260 05-226, Fax: +49 (0) 30 260 05-264, Email: [email protected]

TYPESETTING Compuscript Ltd., Shannon, Ireland

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Offenlegung der Inhaber und Beteiligungsverhältnisse gem. § 7a Abs. 1 Ziff. 1, Abs. 2 Ziff. 3 des Berliner Pressegesetzes: Die Gesellschafter der Walter de Gruyter GmbH sind: Cram, Gisela, Rentnerin, Berlin; Cram, Elsbeth, Pensionärin, Rosengarten-Alvesen; Cram, Dr. Georg-Martin, Unternehmens- Systemberater, Stadtbergen; Cram, Maike, Wien (Österreich); Cram, Jens, Mannheim; Cram, Ingrid, Betriebsleiterin, Tuxpan/Michoacan (Mexiko); Cram, Sabina, Mexico, DF (Mexiko); Cram, Silke, Wissenschaftlerin, Mexico DF (Mexiko); Cram, Björn, Aachen; Cram, Berit, Hamm; Cram- Gomez, Susana, Mexico DF (Mexiko); Cram-Heydrich, Walter, Mexico DF (Mexico); Cram-Heydrich, Kurt, Angestellter, Mexico DF (Mexico); Duvenbeck, Brigitta, Oberstudienrätin i.R., Bad Homburg; Gädeke, Gudula, M.A., Atemtherapeutin/Lehrerin, Tübingen; Gädeke, Martin, Student, Ingolstadt; Gomez Cram, Arturo Walter, Global Key Account Manager, Bonn, Gomez Cram, Ingrid Arlene, Studentin, Mexico, DF (Mexiko), Gomez Cram, Robert, Assistant Professor. Sondon UK, Lubasch, Dr. Annette, Ärztin, Berlin; Schütz, Dr. Christa, Ärztin, Mannheim; Schütz, Sonja, Diplom.- Betriebswirtin (FH), Berlin; Schütz, Juliane, Berlin; Schütz, Antje, Berlin; Schütz, Valentin, Mannheim; Seils, Dorothee, Apothekerin, Stuttgart; Seils, Gabriele, Journalistin, Berlin; Seils, Christoph, Journalist, Berlin; Siebert, John-Walter, Pfarrer, Oberstenfeld; Anh Vinh Alwin Tran, Zürich (Schweiz), Tran, Renate, Mediatorin, Zürich (Schweiz). Mammalia 2020 | Volume 84 | Issue 1

Contents

Taxonomy/phylogeny Museology/history of science

Frank E. Zachos, Les Christidis and Stephen T. Garnett Kees Rookmaaker Mammalian species and the twofold nature of taxonomy: The hornless rhinoceros (Rhinoceros sondaicus inermis a comment on Taylor et al. 2019 1 Lesson, 1836) discovered by Lamare-Picquot in the Sundarbans of Bangladesh in 1828, with notes on the Consuelo Lorenzo, Jorge E. Bolaños-Citalán and Oscar G. history of his Asian collections 74 Retana-Guiascón Rediscovery of Heteromys nelsoni in its type locality after over a century 6 Evolutionary biology

Aldo Caccavo and Marcelo Weksler Ecology Mesial hyperdontia in Sigmodontinae (Rodentia: Cricetidae), with comments on the evolution of the Govan Pahad, Claudine Montgelard and Bettine Jansen anteroconid in Myomorpha 90 van Vuuren Phylogeography and niche modelling: reciprocal enlightenment 10 Biogeography

Renan L.A. Ribeiro and Rita C. Bianchi Andrea Boscherini, Giuseppe Mazza, Mattia Menchetti, Evaluation of bait use for mammal richness 26 Alessandro Laurenzi and Emiliano Mori Time is running out! Rapid range expansion of the Jayasilan Mohd-Azlan, Sally Soo Kaicheen, Lisa Lok and invasive northern raccoon in central Italy 98 Jedediah F. Brodie Diversity and size-structured persistence of tropical Márcia M.A. Jardim, Diego Queirolo, Felipe B. Peters, carnivores in a small, isolated protected area 34 Fábio D. Mazim, Marina O. Favarini, Flávia P. Tirelli, Rhaysa A. Trindade, Sandro L. Bonatto, Júlio César Nick Ching-Min Sun, Chun-Chieh Liang, Bo-Ye Chen, Bicca-Marques and Italo Mourthe Chung-Chi Lin, Kurtis Jai-Chyi Pei and Hou-Feng Li Southern extension of the geographic range of black- Comparison of two faecal analysis techniques to assess and-gold howler monkeys (Alouatta caraya) 102 Formosan pangolin Manis pentadactyla pentadactyla diet 41 Ethology Marjorie Biffi, Pascal Laffaille and Laëtitia Buisson Local habitat preferences of a semi-aquatic mammal, the Arielli F. Machado, Cristiane F. Marks, Brisa Peres, Geruza Pyrenean desman Galemys pyrenaicus 50 L. Melo and Nilton C. Cáceres Movement and use of environmental structures, Akiko Takii, Shigeyuki Izumiyama and Takashi Mochizuki climbing supports and shelters by Akodon montensis An initial record of a long-distance dispersal route of a (Sigmodontinae, Rodentia) in the Atlantic Forest of male sika deer in central Japan 63 southern Brazil 107

Emmanuel M. Hema, Tinsgnimi Dialla, Djidama Sirima, Yaya Ouattara, Ismael Tou and Luca Luiselli Preliminary data on density and habitat of carnivores in two protected areas of Burkina Faso (West Africa) 69