Archiv für Molluskenkunde 148 (2) 161–183 Frankfurt am Main, 19 Dec. 2019

Old and new taxa of Moquin-Tandon, 1856 (: : ) in Spain and adjacent France

Hans D. Boeters

Karneidstr. 8, D-81545 München, Germany ([email protected])

Abstract. A lectotype of utriculus Paladilhe, 1874 is designated. Bythinella artiasensis Fagot, 1887 is synonymised with B. baudoni (Paladilhe, 1874). Paludinella canaliculata Paladilhe, 1869 and Palu dinella guranensis Paladilhe, 1870 are synonymised with Bythinella marginata simoniana (Moquin- Tandon, 1856). Bythinella tejedoi n. sp., B. batalleri cuenca n. subsp., B. tajoensis n. sp., and Bythinella ro lani n. sp. are described as new for Spain. Bythinella andorrensis errensis n. subsp. is new for the French Pyrenees. Key words. , new taxa, new synonyms, lectotype designation, Pyrenees. DOI. https://doi.org/10.1127/arch.moll/148/161-183

Introduction (Bourguignat, 1861), B. reyniesii (Dupuy, 1851), B. ru­ biginosa (Boubée, 1833), B. ru fes cens, B. simoniana (Mo- In Spain, of Bythinella are known from the pro- quin-Tandon, 1856), B. troyana Bernasconi, 2000, and B. vinces of Barcelona, Gerona, Lérida, and Tarragona utriculus (Paladilhe, 1874). Boe ters & Falkner (2012) in Cataluña, and from only the province of Huesca in treated B. servainiana (Pala dilhe, 1870) as distinct spe- Ara gón (Boeters [1988]), with Panticosa as the western- cies and B. rufescens as junior of B. reynie­ most locality (Fagot 1907). In the present publication, sii. They redescribed B. gibbosa (Moquin-Tandon, 1856) I update the information, as previously given by me from Haute-Garonne, and described as new B. ullaensis (Boeters [1988]), by presenting new data on this Boeters & Falkner, 2012 from the Pyrénées-Atlantiques, in regions of northern Spain outside of both Cataluña and B. syntriculus Boeters & Falkner, 2012, and B. rouchi Aragón. Earlier, I treated Bythinella rufescens (Küster, Boeters & Falkner, 2012 from the Ariège, and B. carca­ 1852) as a valid species with B. artiasensis Fagot, 1887 sonis Boeters & Falkner, 2012 from Aude; they excluded as a (Boeters [1988]). However, Boeters & B. eutrepha from the Pyrenees. The current list of 17 spe- Falkner (2012) have since showed that Paludina rufes­ cies in the French Pyrenees is presented in Appendix 1. cens is a junior synonym of Hydrobia reyniesii Dupuy, 1851, now a Bythinella. In the present publication, B. ar­ ti asensis Fagot, 1887 is treated as a junior synonym of Materials and Methods Paludinella baudoni Paladilhe, 1874. Also in this pub- lication, I consider B. espanoli Bech, 1980 to be a valid Living and empty shells were separated from species, not a junior synonym of B. batalleri Bofill, 1925 the substrate by fractionating using a set of sieves with as I earlier had believed (Boeters [1988]). mesh sizes of 2.60 and 0.56 mm. Animals for dissection In this revision of Spanish Bythinella material, I also were preserved in 75% ethanol by weight. consider species living in the French Pyrenees which bor- The counting of whorls of shells follows Gitten- der Spain. Bernasconi (2000) listed 13 species of Bythi­ berger et al. (1970), and anatomical investigations were nella from the French Pyrenees: B. baudoni, B. bertrandi made in accordance with my previous work (Boeters Bernasconi, 2000, B. dar rieuxii (Folin & Bérillon, 1999). Drawings of shells and animals were made with 1877), B. eurystoma (Paladilhe, 1870) [= B. servainiana a Leica M80 Drawing Device. Shells were photographed (Paladilhe, 1870) sensu Ber nasconi], B. eutrepha (Pal- with a Leica R8 digital system. Length and diameter adilhe, 1867), B. ginolensis Fagot, 1881, B. pyrenaica of shells were measured with a 5 mm measure plate

© E. Schweizerbart’sche Verlagsbuchhandlung (Nägele u. Obermiller) and Senckenberg Gesellschaft für Naturforschung, 2019, ISSN 1869-0963