British Columbia Marine Faunistic. Survey Report: Asteroids From the Northeast Pacific
P. Lambert
Department of Fisheries and the Environment Fisheries and Marine Service Resource Services Branch Pacific Biological Station Nanaimo, British Columbia V9R 5K6
June 1978
Fisheries and Marine Service Technical Report No. 773 Fisheries and Marine Service Technical Reports
These report s co ntain scien tific and tec hn ica l in formation tila t represents an im porta n t con t ri bu t ion to ex isting k nowlc: dge but w hi ch ror som e reaso n m ay no t be app ropriate for primary scienti f ic (i .e. J OI/III0/) publicat ion. Techn ictl Reports are direc ted primari ly towards ,\ world wi de au dience and have an in terna ti onal distri but ion. No res tri ct ion is pl aced 011 subjec t m atter and the se ries rdlec ts the broa d interes ts and policies of the Fi <; heries and Mal'inc Se r \'icc, namel\' , lI c, heries manage m ent , technology ,1I1d develop illent , ocea n sc ience s and aq uatic environ men ts relevan l to Ca nada. TechniGii Repo rts may be cited as rull publica ti ons. The correct cita ti on ap lKars ab o\'e the abstr,lct or e,lch repon Eac h rcport w ill be ,lbSl raL'lcd in Aql/a/ic Sciences ond fi l'heries Ah.I'/I'oc/s and will be indexe d Service des peches et des sciences de 1a mer Rapports techn iques Ces r • Cover design by Christine Rusk Fisheries and Marine Service Technical Report 773 June 1978 BRITISH COLUMBIA MARINE FAUNISTIC SURVEY REPORT : ASTEROIDS FROM THE NORTHEAST PACIFIC by P. LambertI Department of Fisheries and the Environment Fisheries and Marine Service Resource Services Branch Pacific Biological Station Nanaimo, British Columbia V9R 5K6 IAssociate Cur?tor Marine Biology Division B.C. Provincial Museum Victoria, B.C. - ii - (c) Minist~r of Su~pl y and Services Canada 1978 Cat. no . Fs 97-6/773 ISSN 0701-7626 - iii - This is the fourth Technical Repo~t dealing with the eastern Pacific fauna collected by the Pacific Biological Station Previous reports Bernard, F . R. 1967. Prodrome for a distributional check-list and bibliography of the recent marine mollusca of the west coast of Canada. Fish. Res. Board Can. Tech. Rep. 2: 261 p. 1971. British Columbia marine faunistic survey report on the Brachiopoda. Fish. Res. Board Can. Tech. Rep. 268: 10 p. Austin, W. C., and M. P. Haylock. 1973. British Columbia marine faunistic survey report: ophiuroids from the northeast Pacific. Fish. Res. Board Can. Tech. Rep. 426: 36 p. - iv - ABSTRACT Lambert, P. 1978. British Columbia marine faunistic survey report: asteroids from the northeast Pacific. Fish. Mar. Servo Tech. Rep. 773: 23 p. Sixty-eight living species and subspecies belonging to thirteen families, are present in British Columbia marine waters. This inventory is based on the Marine Faunistic Survey of the Pacific Biological Station that sampled the subtidal b~nthos of the continental shelf and slope to 2200 m. Key words: Asteroids, benthos, echinoderms, invertebrates, inventory. / / RESUME Lambert, P. 1978. British Columbia marine faunistic survey report: asteroids from the northeast Pacific. Fish. Mar. Servo Tech. Rep. 773: 23 p. Soixante-huit especes et sous-especes appartenant a treize fami11es ont ete denombrees dans 1es eaux marines de 1a Co1ombie-Britannique. L'inventaire se fonde sur 1e re1eve de 1a faune marine de 1a Station bio1ogique du Pacifique, qui a echanti110nne 1e benthos infratida1 du plateau et du talus continental jusqu'a 2200 m. Mots c1es: Etoi1es de mer, benthos, echinodermes, invert~br~s, inventaire. INTRODUCTION TllP asteroids listed in this report are contained in the collections o( the Pacific Biological Station at Nanaimo, made primarily during the Fisheries Research Board of Canada Faunistic Survey from 1960 to 1972. However, smaller collections from 1912 to the present are also included. Specimens from the Bering Sea to northern Washington are included, but the bulk of the specimens are from off the British Columbia coast. The material is arranged by class, family, genus, and species. All identifications were made by the author and based on the monographs of Fisher (1911, 1911-1930) and D'Yakonov (1950) and papers by Grainger (1966) and Alton (1966). To verify certain identifications, identified specimens were borrowed from the National Museum of Canada and the U.S. National Museum. Range extensions resulting from study of this collection, that of the British Columbia Provincial Museum , and the Pacific Environment Institute, are summarized in another paper (Lambert, in press). The Fisheries Research Board collection first carne into the author's hands when he was preparing a handbook for the B.C. Provincial Museum on the seas tars of British Columbia. Specimens will be deposited in the B.C. Provincial Museum, Victoria, National Museum of Canada, Ottawa, and a representative series will remain at the Pacific Biological Station. I am indebted to Dr. Frank Bernard for making the collection available and for supervising the computerization of the data and answering all my queries on station data. I am also grateful to Dr. Dan Quayle who collected most of the specimens over the years and who kindly relinquished his interest in this group. LITERATURE CITED Alton, M. S. 1966. Bathymetric distribution of sea stars (Asteroidea) off the northern Oregon coast. J. Fish. Res. Board Can. 23: 1673-1714. D'Yakonov, A. M. 1950. Sea stars (Asteroids) of the U.S.S.R. seas. Keys to the fauna of the U.S.S.R. Moscow and Leningrad 34: 202 p. Fisher, W. K. 1911-1930. Asteroidea of the North Pacific and adjacent waters. Bull. U.S. Nat. Mus. 76. Part 1, vi and 419 p. (1911); Part 2, iii and 245 p. (1928); Part 3, iii and 356 p. (1930). Grainger, E. H. 1966. Sea stars (Echinodermata: Asteroidea) of Arctic North America. Bull. Fish. Res. Board Can. 152: 70 p. - 2 - FORMAT A systematic list of the species identified is given, followed by an alphabetical listing of station data. - 3 - Species included in the Fisheries Research Board Collection Class Asteroidea ORDER PHANEROZONIA FAMILY PORCELLANASTERIDAE Ctenodiscus crispatus (Retzius, 1805) Eremicaster pacificus (Ludwig, 1905) FAMILY ASTROPECTINIDAE Leptychaster anomalus Fisher, 1906 Leptychaster arcticus (Sars, 1851) Leptychaster pacificus Fisher, 1906 Psilaster pectinatus (Fisher, 1905) Dipsacaster anoplus Fisher, 1910 Dipsacaster borealis Fisher, 1910 FAMILY LUIDIIDAE Luidia foliolata Grube, 1866 FAMILY BENTHOPECTINIDAE Pectinaster agassizi evoplus (Fisher, 1910) Luidiaster dawsoni (Verrill, 1880) Nearchaster aciculosus (Fisher, 1910) Nearchaster variabilis (Fisher, 1910) Benthopecten claviger Fisher, 1910 Benthopecten mutabilis Fisher, 1910 - 4 - FJ\MIL Y CONIASTERIDAE Gephyreaster swifti (Fisher, 1905) Pseudarchaster dissonus Fisher, 1910 Pseudarchaster parelii alascensis Fisher, 1911 Mediaster aequalis Stimpson, 1857 Ceramaster patagonicus (Sladen, 1889) Hippasteria californica Fisher, 1905 Hippasteria spinosa Verrill, 1909 Cryptopeltaster lepidonotus Fisher, 1905 FAMILY ASTEROPIDAE Dermasterias imbricata (Grube, 1857) ORDER SPINULOSA FAMILY ASTERINIDAE Patiria miniata (Brandt, 1835) FAMILY ECHINASTERIDAE Poraniopsis inf1ata (Fisher, 1906) Henricia aspera Fisher, 1906 Henricia asthenactis Fisher, 1910 Henricia leviuscula (Stimpson, 1857) Henricia leviuscula annectens Fisher, 1910 Henricia leviuscula dyscrita Fisher, 1911 Henricia leviuscula multispina Fisher, 1910 Henricia longispina Fisher, 1910 Henricia longispina aleutica Fisher, 1911 Henricia sanguino1enta (0. F. Muller, 1776) - 5 - FAMILY SOLASTERIDAE So l as t er borealis (Fisher, 1906) Solaster dawsoni Verrill, 1880 Sola ster endeca (Linnaeus, 1771) Solaster paxillatus Sladen, 1889 Solaster stimpsoni Verrill, 1880 Crossaster papposus, (Linnaeus, 1768) Lophas t er furcilliger Fisher, 1905 FAMILY PTERASTERIDAE Pter ast er jordani Fisher, 1905 Pteraster mi1itaris (0. F. Muller, 1776) Pteraster obscurus (Perrier, 1891) Pteraster tesse1atus Ives, 1888 Dip10pteraster mu1tipes (Sars, 1865) Hymenaster perissonoties Fisher, 1910 Hymenaster quadrispinosus Fisher, 1905 ORDER FORCIPULATA FAMILY BRI SINGIDAE Craterobrisinga synaptoma Fisher, 1917 Freyel1ast er fecundus (Fisher, 1905) FAMILY ZOROASTERIDAE Zoroaster evermanni Fisher, 1905 Zoroaster evermanni mordax Fisher, 1919 Zoroaster ophiurus Fisher, 1905 - 6 - FAMILY ASTERIDAE Pedicellaster magister Fisher, 1923 Tarsaster a1askanus Fisher, 1928 Amp~eraster marianu~ (Ludwig, 1905) Rathbunaster californicus Fisher, 1906 ~ty1asterias forreri (de Lorio1, 1907) Disto1asterias sp. Orthasterias koehleri (de Loriol, 1897) Pycnopodia he1ianthoides (Brandt, 1835) Asterias amurensis Lutken, 1871 Leptasterias aequa1is (Stimpson, 1862) Leptasterias arctica (Murdoch, 1885) Leptasterias hexactis (Stimpson, 1862) Leptasterias hy10des Fisher, 1928 Evasterias trosche1ii (Stimpson, 1862) - 7 - BRITISH COLUMBIA MARINE FAUNISTIC SURVEY ASTEROID COLLECTION Am~i1 _~'r_; _I:~~ _~~l~il r i_anus Year:- St. No. Hon. Latitude Longitude Depth (M) 0 0 1964 175 9 47 58.0N 125 47.2W 924 51°09.8N 130009.0W 858 1964 214 9 0 1968 29 9 48 13.0N 126.18.0W 951 Asterias amurensis Year St. No. Non. Latitude Longitude Depth (M) 0 0 11 54 18.0N 116 33.SW 220 1965 96 0 0 1965 97 11 57 16.2N 170 09.6W 38 Benthopecten c1aviger Year St. Ho. Mon. Latitude Longitude Depth (M) 0 0 1964 202 9 49 53.2N 127 22.7W 0 1830 1965 46 8 51 43.8N 131 ° 13 . 5W 1830 Benthopecten mutab1ilis Year St. No. Non. Latitude Longitude Depth (M) 0 0 1964 201 9 49 53.0N 127 17.5W 1830 Ceramaster sp. Year St. No. Mon. Latitude Longitude Depth (M) 0 0 1917 77 0 52 30.0N 123 34.0W 73 Ceramaster patagonicus Year St. No. Non. Latitude Longitude Depth (M) 0 0 1961 810 4 54 11.5N 132 54.0W 0 0 70 1963 54 8 58 07.0N 148 50.0W 0 0 241 1965 77 8 54 37.0N 0 133055.0W 229 1966 34 6 51 45.0N 128 00.JW 100 0 0 1970 25 6 51 07.3N 127 49.3W 0 95 1971 47 9 54 28.6N 133° 53. 3W 256 1972 606 2 48°24. 5N 126°04.3W 360 - 8 - Craterobrisinga synaptoma Year St. No. Mon. Latitude Longitude Depth (M) 0 48 38.5N 126°44.5W 1964 184 9 0 0 1353 1964 202 9 49 53.2N 127 22 . 7W 1830 Cros saster papposus fear St. No . ~' :on . Latitude Longitude Depth (M) 0 6 49 18.4N 124°12 . 0W 20 1914 45 0 5 49°01. 2N 123 43.0W 30 1917 7S 0 1921 103 7 49 26.8N 124 °11. OW 40 0 0 120 5 49 00.5N 125 44.3W 1934 0 35 218 4 49 10 . 5N 126°13 . 3W 102 1934 0 1959 1 9 49°13. ON 123 53.0W 130 0 0 1960 297 9 54 07.3N 132 16.1W 75 305 9 54°11.2N 131°22.4W 1960 0 20 1960 332 9 51 31. Cr~Etopeltaster 1eeidonotus Year St. Uo. Mon. Latitude Longitude Depth (M) 1971 42 3 53°01. ON 132055.')W 1061 Ctenodiscus crispatus Year St. No. Mon. Latitude Longitude Depth (M) 0 0 1944 1 11 49 00.0N 124 00 . 8W 110 1960 901 1 54°17.0N 130°17 .OW 60 0 800 8 49 30.0N 123007.0W 260 1961 0 0 1962 714 10 49 12.0N 123 54 . 0W 75 0 9 1 50°51. 4N 127 14.'JW 136 1963 0 0 1963 10 1 50 46.5N 126 09.5W 192 0 0 1963 55 8 58 07.0N 148 50.0W 209 0 0 1963 56 8 58 05.0N 149 40.0W 220 1963 124 9 56°11.0N 153°21. OW 201 0 0 1963 135 9 56 42.0N 153 19.0W 128 0 0 1963 143 9 56 55.::JN 152 40.0W 146 0 1963 144 4 58 06.8N 149002.1W 49 0 0 1964 185 9 48 44.4N 126 30.0W 461 1966 9 4 50°41. 2N 126°37 . 5W 64 0 0 1966 29 6 51 20.0N 127 42.0W 160 0 1966 30 6 51 20 . 8N 127°42.~W 160 0 0 1968 35 9 48 23 . 2N 126 05.5W 360 0 0 1968 45 9 48 44.SN 126 30.3W 436 0 0 1968 52 9 49 04.5N 126 51.5W 304 0 0 1970 21 6 50 46.5N 127 06.3W 205 Dermasterias imbricata Year St. l~o . Mon . 'Latitude Longitude Depth (M) 0 0 1964 151 4 53 43.5N 132 15.7W 46 Diplopteraster mu1tipes Year St. No. Mon . Latitude Longitude Depth (M) 0 1963 52 8 58 07.0N 148°50.0W 179 0 0 1963 57 8 57 50 . 0N 150 16.JW 174 0 1963 126 9 56°11. ON 153 40.0W 228 0 0 1963 127 9 55 53.0N 153 45.0W 228 1963 129 9 56°01.0N 154°28.0W 231 0 1965 23 6 54°35.2N 130 35.0W 102 0 0 1968 35 9 48 23.2N 126 05.5W 360 1968 45 9 48°44.5N 126°30.3W 436 0 0 1971 63 9 54 31,8N 132 39.8W 366 0 0 1972 606 2 48 24.5N 126 08.6W 360 - 10 - Dipsacaster anop1us Year St. No. Hon . Latitude Longitude Depth (M) 0 0 1964 184 9 48 38.3N 126 44.SW 1353 1964 185 9 480 44.4N 1260 30 .0W 461 1965 46 8 51 0 43.0N 1310 13.5W 1830 1965 66 8 53°01. ON 1320 55.0W ll90 1971 38 9 52 0 06.3N 1310 26.2W 1032 Dipsacaster borealis Year st. ;~o . Mon. Latitude Longitude Depth (M) 1963 109 9 57 0 12.0N 150°01. OW 275 1963 llO 9 57 0 04.5N 1510 26.0W 256 1963 121 9 56 0 18.0N 152°45.0W 256 1963 124 9 56°11.0N 1530 21.0W 201 1965 79 8 54 0 32.0N 1330 25.0W 340 Disto1asterias sp. Year St. No . Ho n. Latitude Longitude Depth (M) 1968 46 9 48 0 44 . 5N 1260 21.5W 494 Erernicaster pacificus Year St. No. Mon. Latitude Longitude Depth (M) 1964 200 9 49 0 43.5N 1280 28.5W 2200 1964 202 9 49 0 53. 2N 1270 22 . 7W 1830 1964 207 9 50 0 32.0N 129 0 01 . 5W 1830 1964 209 9 50 0 25.7N 1290 ll. OW 2013 Evasterias trosche1ii Year St. i'lo. Mon. Latitude Longitude Depth (M) 0 0 1961 868 4 53 55.0N 131 32.0W II Freye11aster fecundus Year St. No. Hon. Latitude Longitude Depth (M) 1965 72 8 54 0 03.0N 1340 00.0W llOO - 11 - GeEhyreaster swifti Year St. No. Non. Latitude Longitude Depth (M) 0 38 9 49 01.5N 123034.7W 64 1962 0 0 716 6 50 00.0N 123 57.0W 91 1962 0 0 2 1 50 36.9N 126 57.7W 26 1963 0 0 1965 6 5 54 10.3N 131 50.0W 73 Henricia aSEera Year St. No. Mon. Latitude Longitude Depth (M) 0 1934 111 5 48 39.0N 126°46.0W 128 0 1961 807 5 54 08.2N l32026.0W 4 0 0 1962 26 8 52 08.0N 131 09.0W 27 0 0 1962 718 6 49 58.0N 125 10.5W 18 0 1963 14 1 50 38.3N 126°34.0W 64 0 0 1963 36 6 49 09.5N 123 40.5W 102 0 1963 125 9 56°11. ON 153 40.0W 228 0 1968 35 9 48 23.2N 126°05.5W 360 0 0 1971 47 9 54 28.6N 133 53.3W 256 0 1972 606 2 48 24.5N 126°08.6W 360 Henricia asthenactis Year St. No. Mon. Latitude Longitude Depth (M) 0 1963 25 2 51 51.0N 128°41.0 128 Henricia 1eviuscu1a Year st. I~o . Mon. latitude Longitude Depth (M) 0 0 1917 75 5 49 01.ZN 123 43.0W 0 30 1934 ll2 6 49 51.0N 127006.0W 100 0 1934 114 5 49023.7N 126 34.0W 85 0 1934 132 7 49°31. 7N 126 34.0W 150 0 0 1934 231 6 49 22.2N 126 28.0W 20 0 1934 425 7 49 44.0N 121° 57 . OW 2 0 0 1935 7 6 52 09.0N 131 14.0W 45 1935 24 6 53°02. ON l32026.0W 110 0 1935 31 7 53 09.0N 132°21. OW 25 0 1935 37 7 52 08.0N l31007.0W 40 0 0 1950 97 5 48 53.2N 125 18.0W 5 1960 104 7 54°01. IN l32° 35 .OW 32 0 1960 287 7 54 08.3N 131°48.2W 29 0 1961 632 5 48°30.0N 125 04.0W 10 0 0 1961 811 4 48 58.0N 125 21.CJW 38 1961 818 6 54°11.5N l32054.0W 183 12 - 11('11 r i (' iii I l ' V i lJ S C \J I ;t .. - . - - - - . - _. - _.- YCil r St. ~~o . Mon . Latitude Longitude Depth eM) 1962 26 8 S2008.0N 131009.0W 27 0 1962 720 6 49 39 . 0N 124°SS.0W 2S 1963 1 1 S0036 . 4N 126°S0.9W 113 0 1963 3 1 S0037 . 7N 127 03 . 8W 60 1963 7 1 SOoSO.7N 127°20. SW 139 1963 12 1 S0044.0N 127°01. 6W SS 1963 14 1 S0038.3N 126°34.0W 64 0 1963 38 6 49 11.0N 123°41. OW 36 0 1964 lS4 4 S4002.0N 132 3S.0W 16 0 1966 19 4 49 47.0N 12SoS8.0W 20 1966 28 6 Sl020.0N 127°40. OW lS0 0 1966 61 10 SloSS.0N 131 04.2W 730 1968 12 8 S0043 . 0N 128°30.0W 90 0 1969 7 6 S4012.9N 132 S7.3W 36 1969 31 8 51 004.5N 127026.5W 110 0 0 1972 60 6 2 48 24 . 5N 126 08.6W 360 Henricia 1ev i uscu1a annec tens Yea r St. No . r;on. La t itude Longitude Depth eM) 0 0 1917 75 S 49 01 . 2N 123 43 . 0W 30 0 1934 120 S 49 00 . 3N 12S044.3W 21 0 1935 27 6 S2044.0N 132 07.0W 46 0 1960 297 9 S4007.0N 132 16.1W 7S 0 1961 632 S 48 30.0N 12S004.0W 10 1962 26 8 S2008.0N 131009 . 9W 27 1963 3 1 S0037 . 7N 127°03. OW 60 0 1963 7 1 SOoSO . 7N 127 20.SW 139 1963 14 1 S0038.3N 126°34 . 0W 64 0 1963 2S 2 51 51.0N 128041.0W 128 1963 58 8 57038 . 0N 150016.JW 156 0 1963 125 9 56°U .ON 153 40.0W 228 0 0 1963 139 9 56 50. 0N 153 19.0W 165 1965 6 5 540 10 . 3N 131050 . 0W 73 0 0 1965 77 8 54 37.0N 133 55 . 0W 229 0 0 1966 34 6 51 45. 0N 128 00.0W 100 0 1966 61 10 51°55.0N 131 04.2W 730 Henrici a 1evi us cu1a dyscrita Year St. No . Mon . La titude Longit ude Dept h eM) 0 0 1965 72 8 54 03.0N 134 00.0W UOO - 13 - Henricia 1eviuscu1a mu1tispina Year St. No. Hon. Latitude Longitude Depth (M) 0 1964 169 9 48022.8N 125 14.1W 183 0 0 1966 4 2 50 38.1N 126 34. 0W 91 Henricia 10ngispina Year St. i'40. Mon. La titude Longitude Depth (M) 0 0 1963 125 9 56 11.0N 153 40.0W 228 0 0 1970 21 6 50 46.5N 127 06.3W 205 Henricia 10ngispina a1eutica Year St. No. Mon. Latitude Longitude Depth (M) 0 0 1963 125 9 56 11.0N 153 40.0W 228 Henricia sanguino1enta Year St. No. Mon. Latitude Longitude Depth (M) 0 0 1960 308 9 54 11.1N 130 59.4W 30 0 0 1960 333 9 53 37.9N 131 15.0W 46 0 1961 511 4 48 58.0N 125°21. OW 38 0 1963 130 9 56°01.0N 154 28.0W 228 1964 23 9 48°22.8N 1250 14.0W 183 0 0 1965 106 11 57 16.9N 170 01.0W 37 0 0 1968 29 9 48 13.0N 126 18.5W 951 0 0 1972 638 2 54 09.3N 130 23.0W 80 Hippasteria ca1ifornica Year St. ,\lo. Mon. Latitude Longitude Depth (M) 1964 184 9 48°38.5N 1260 44.5W 1353 1964 200 9 49 0 43.5N 1280 28.5W 2200 1964 214 9 510 09.0N 1300 09.0W 858 0 0 1965 72 8 54 03.0N 134 00.0W 1100 0 0 1968 29 9 48 13.0N 126 18.5W 951 1968 34 9 OOoOO.ON OOOoOO.OW 0 0 0 1968 35 9 48 23.2N 126 05.5W 360 0 0 1971 39 9 52 05.5N 131 26.7W 1061 0 0 1971 42 3 48 51.2N 125 07.0W 15 1971 45 9 53002.0N 132°51. 8W 1204 0 1972 634 2 52°07.2N 131 28.0W 1230 - 14 - Hi£l£lasteria seinosa Year St. No. Mon. Latitude Longitude Depth eM) 0 0 1960 297 9 54 07.3N 132 16.1W 75 0 0 1963 5 1 50 41.8N 127 07.7W 132 0 0 1963 12 1 50 44.0N 127 01.6W 55 0 0 1963 28 2 51 44 . 5N 128 34 .0W 146 0 0 1963 53 8 58 07.0N 148 50.0W 241 0 0 1963 56 8 58 05.0N 149 40.0W 220 0 0 1963 58 8 57 38.0N 150 16.0W 156 0 1963 125 9 56 °11. ON 153 40.0W 228 0 1964 175 9 47°58.0N 125 47 . 2W 924 0 1964 214 9 51°09.0N 130 09.0W 858 0 0 1965 6 5 54 10.3N 131 50.0W 73 0 1965 77 8 54°37.JN 133 55.0W 229 0 0 1968 12 8 50 43.0N 128 30.0W 90 0 0 1968 38 9 48 46.5N 125 22.2W 95 1968 46 9 48°44.5N 126°21.5W 494 0 1971 47 9 54 28.6N 133053.3W 256 0 0 1972 605 2 48 25.2N 126 07.0W 304 0 0 1972 634 2 52 07 . 2N 131 28.0W 1230 Hymenaster perissonoties Year St. No . Ho n. Latitude Longitude Depth eM) 0 1964 184 9 48°38.5N 126 44.5W 1353 0 1964 200 9 49 43.5N 128°28.5W 2200 0 0 1968 24 9 47 58.2N 125 57.4W 1537 Hymenaster quadrispinosus Year St. No. Mon. Latitude Longitude Depth eM) 0 1965 42 8 51 ° 47 .ON 131 13.0W 1876 1965 46 8 51°43 .0N 131 ° 13 . 5W 1830 Le£ltasterias aequa1is Year St. No. Mon . Latitude Longitude Depth err) 19 35 37 7 52°08.0N 131° 07 . OW 40 - 15 - LeEtasterias arctica Year St. No. Mon. Latitude Longitude Depth (M) 0 1965 106 11 57 0 16.9N 170 01. 9W 33 LeEtasterias hexactis Year St. No. Han. Latitude Longitude Depth (t·n 1934 132 7 49 0 31. 7N 126 0 34.0W 150 1934 226 7 49 0 15.2N 126°07.0W 115 1934 424 7 49 0 51. 2N 1270 04.0W 85 1935 22 6 52 0 07.0N 131°04.0W 20 1935 24 6 530 02.0N 1320 26.0W 110 1950 97 5 48°53.2N 1250 18.0W 5 1963 125 9 56 0 11.0N 1530 40.0W 228 1963 143 9 56°55.0N 1520 40.0W 146 LeEtasterias hylodes Year St. Uo. Mon. Latitude Longitude Depth (M) 1934 444 7 49 0 57.5N 1260 54 . 0W 120 1963 80 9 580 00.5N 1500 32.0W 146 1963 104 9 570 17.8N 1510 24.0W 110 Leptychaster anornalus Year St. l'~ O • Mon. La titude Longitude Depth (M) 1963 124 9 56 0 11. ON 1530 21. OW 201 1963 143 9 56 0 55.0N 1520 40.0W 146 1964 169 9 480 22.8N 1250 14.1W 183 1964 205 9 50 0 40.0N 1280 48.5W 197 0 1968 45 9 48°44.5N 126 30.3 436 Leptychaster arcticus Year St. No. Mon. Latitude Longitude Depth (M) 1961 856 5 49 0 01.7N 1230 40.0W 40 1963 21 2 51 °21. 6N 1290 03.0W 210 1963 125 9 56 0 11. ON 1530 40.0W 228 1964 185 9 48°44.4N 126°30.0W 461 - 16 - Leptychaster pacificus Year St. Ho. Mon. Latitude Longitude Depth (M) 0 0 1917 77 9 52 30.0N 123 34.0W 73 0 0 1919 94 9 49 10.5N 129 56.0W 45 0 1962 701 6 49 59.1N 124°58.0W 115 0 0 1963 2 1 50 36.9N 126 57.7W 26 963 5 1 50° 41. 8N 127007.7W 132 0 0 1963 80 9 58 00.5N 150 32.0W 146 0 1968 35 9 48 23.2N 126°05.5W 360 Lophaster furci11iger Year St. No. Hon. Latitude Longitude Depth (M) 0 0 1964 175 9 47 58.0N 125 47. 2W 924 0 0 1964 202 9 50 00.0N 128 08.1W 1830 0 1965 72 8 54°03.0N 134 00.0W 0 noo 1965 77 8 54°37.0N 133 55.0W 229 0 0 1965 79 8 54 32.0N 133 25.0W 340 0 1968 29 9 48°13. ON 126 18.5W 951 Luidia folio lata Year St. 1\)0. Mon. Latitude Longitude Depth (M) 0 1934 113 7 49°21. ON 126 43 .0W 55 0 0 1934 133 7 49 22.3N 126 55.0W 137 0 1934 134 7 49 30.0N 125°51. OW 95 0 0 1960 32 6 48 47.9N 123 12.3W 14 0 0 1960 298 6 49 00.0N 123 30.0W 60 0 0 1961 13 6 48 42.6N 125 35.6W 75 0 0 1962 53 9 49 03.0N 123 43.0W 0 0 73 1962 720 6 49 39.'JN 124 55.0W 25 0 1963 56 8 58°05.0N 149 40 . 0W 220 0 0 1966 10 4 50 42.5N 126 36.0W 42 Luidiaster dawsoni Year St. No. Mon. Latitude Longitude Depth (M) 0 1965 6 5 54°10.3N 131 50.0W 73 0 1965 77 8 54°37.9N 133 55.0W 229 1968 45 9 48°44.5N 126°30.3W 436 1971 47 9 54°28.6N 133053.3W 256 0 0 1972 604 2 48 24.2N 126 05.0W 274 - 17 - Hediaster aegualis Year St. No . Mon. Latitude Longitude Depth (M) 0 0 1916 74 8 49 02.4N 123 45.0W 50 0 0 1917 75 5 49 01. 2N 123 43.0W 30 0 0 1934 8 2 49 10.3N 126 13.0W 56 0 0 1934 111 5 48 39.0N 126 46.0W 128 0 0 1934 124 5 49 15.7N 126 20.4W 555 0 0 1959 1 9 49 l3. ON 123 53.0W 130 0 1959 2 12 48°50. 4N 123 40.0W 55 0 1960 299 6 54 08.9N l32025.0W 66 0 0 1960 322 9 52 59.0N 129 28.QW 35 0 0 1960 331 9 51 33. 3N 128 43.5W 22 0 0 1960 333 9 53 37.9N 131 15 . 0W 40 0 1960 997 9 54007.0N 132 16 .0W 49 0 1961 850 4 50 32.0N 127°36.0W 57 0 1962 26 8 52 08.0N 131° 09 . 'JW 27 0 1963 2 1 50 36.9N 126°57 .7W 26 0 0 1963 3 1 50 37.7N 127 03.8W 60 0 1963 4 1 50 40.4N 127°11. OW 58 0 1963 7 1 50 50.7N 127020.5W l39 0 0 1963 12 1 50 44.0N 127 01.6W 55 0 0 1963 14 1 50 38.3N 126 34 .0W 64 0 0 1963 16 1 50 31.4N 126 21 .5W 112 0 1963 136 9 56 43.0N 153007.0W 128 0 0 1963 624 1 50 36.9N 126 57.0W 26 0 1964 153 4 54 02. 0N l32035. OW 16 0 0 1964 166 9 48 32.JN 125 02.0W 64 0 0 1964 169 9 48 22.8N 125 14.1W 183 0 1966 16 4 50°11. IN 127 50.2W 110 0 0 1966 28 6 51 20.0N 127 40.9W 150 0 0 1968 12 8 50 43.0N 128 30 . 0W 90 0 1968 38 9 48 46.5N 125022 . 2W 95 0 1970 27 6 51 07.8N 12S00S.6W l32 0 0 1970 53 6 50 02.0N 127 45.5W 81 0 0 1972 636 2 54 09.9N 130 26.0W 101 Nearchaster aciculosus Year St. No. i·lon. Latitude Longitude Depth (M) 0 1964 175 9 47058. 'J N 125 47.2W 924 0 1964 214 9 51 09.0N l30009 . 0W 858 1965 66 8 53°01. ON l32°55.0W 1190 0 1965 77 8 54 37.0N l33° 55. OW 229 0 0 1968 23 9 48 04.2N 125 49.5W 653 - 18 - ~earchaster aciculosus Year St. No. :Ion. Latitude Longitude Depth (M) 1968 29 9 48°13. ON 126°18.5W 951 0 1968 34 9 48 23.2N 126°05.5W 360 0 0 1971 39 9 52 05.5N 131 26.7W 1061 0 1971 42 3 48°51. 2N 125 07.0W 15 0 0 1971 45 9 53 02.0N 132 51.8W 1204 0 0 1972 634 2 52 07.2N 131 28.0W 1230 Nearchaster variabilis Year St. No. Lon. Latitude Longitude Depth (M) 0 0 1964 175 9 47 58.0N 125 47.2W 924 0 1965 81 8 54 31.0N 132037.0W 370 0 1971 47 9 54 28.6N 133053.3W 256 Orthasterias koehleri Year St. :,0 . Mon. Latitude Longitude Depth (M) 0 1934 218 4 49 10.3N 126°31. 3W 102 0 0 1934 231 6 49 22.2N 126 28.0W 20 0 0 1960 303 9 54 01.0N 132 35.1W 17 0 1961 898 4 54 °11. ON 133 55.0W 50 0 0 1962 704 6 49 45.0N 123 55.0W 80 0 0 1963 15 1 50 39.0N 126 24.0W 128 0 0 1964 453 9 54 02.0N 132 35.0W 16 0 0 1966 34 6 51 45.0N 128 00.0W 100 0 0 1966 61 10 51 55.0N 131 04.2W 730 0 0 1969 7 6 54 12.9N 132 57.3W 36 0 0 1969 31 8 51 04.5N 127 26.5W 110 0 1969 34 8 51 °51. 4N 128 31.0W 106 Patiria miniata Year St . No. Kon . Latitude Longitude Depth (M) 0 0 1934 231 6 49 22.2N 126 28.0W 20 0 0 1934 425 7 49 44.0N 121 57.0W 2 0 0 1934 444 7 49 57.5N 126 54.0W 120 0 0 1935 7 6 52 09.0N 131 14.0W 45 0 0 1959 38 1 48 59.4N 125 03.0W 8 - 19 - Pectinaster agassizi evop1us Year St. No. Mon. Latitude Longitude Depth (M) 1964 200 9 49 0 43.5N 1280 28.5W 2200 1965 72 8 54 0 03.0N 1340 00.0W 1100 Pedice11aster magister Year St. No. Mon. Latitude Longitude Depth (M) 1963 32 2 51 0 13.5N 1290 05.0W 223 1964 175 9 47 0 58.0N 1250 47.2W 924 1968 35 9 480 23.2N 1260 05.5W 360 1971 38 9 52 0 06. 3N 1310 26.2W 1032 Poraniopsis inf1ata Year St. No. Hon. Latitude Longitude Depth (H) 0 1960 928 9 54 °Il. 2N 130 58.0W 70 1962 34 8 530 56.0N 1330 08.7W 11 Pseudarchaster dissonus Year St. i'lo. Mon. Latitude Longitude Depth (M) 1964 202 9 50 0 00. 0N 1280 08.1W 1830 1965 46 8 510 43.0N 131° 13 . 5W 1830 Pseudarchaster pare1ii a1ascens Year St. No. Mon. Latitude Longitude Depth (M) 1963 33 2 510 22.0N 1290 03.5W 223 1963 53 8 580 07.0N 1480 50.0W 241 1965 66 8 530 01.0N 1320 55.0W 1190 1965 81 8 54 0 31.0N 1320 37.0W 370 1966 30 6 510 20.0N 1270 42.0W 160 1968 35 9 480 23.2N 1260 05.5W 360 1968 45 9 480 44.5N 126°30. 3W 436 1972 606 2 480 24.5N 1260 08.0W 360 - 20 - Psjlaster eectinatus Year St. No. Mon. Latitude Longitude Depth (M) 1964 202 9 50 0 00.0N 128°08.1W 1830 1965 46 8 51 0 43.0N 131 0 13.5W 1830 Pteraster jordani Year St. No. No n. Latitude Longitude Depth (M) 1963 129 9 56 0 01.0N 1540 28.0W 231 1963 136 9 56 0 43.0N 1530 07.0W 128 Pteraster mi1itaris Year St. No. Mon. Latitude Longitude Depth (M) 1963 125 9 56 0 11.0N 1530 40.0W 228 1963 129 9 56 0 01.0N 1540 28.0W 231 0 0 1963 626 9 56 11.0N 153 40.0W 228 0 1964 217 9 50 58.2N 127047.9W 105 0 0 1966 34 6 51 45.0N 128 00.0W 100 0 0 1968 12 6 50 43.0N 128 30.0W 90 Pteraster obscurus Year St. ,-1o. Mon. Latitude Longitude Depth (M) 0 1965 106 11 57 16.9N 170°01. gW 37 Pteraster Tesse1atus Year St. Uo . Mon. Latitude Longitude Depth (N) 0 0 1917 75 5 49 01.2N 123 43.0W 30 0 0 1960 305 9 54 11.2N 121 22.4W 20 0 1961 533 4 49°13. ON 125 57.0W 20 0 0 1963 5 1 50 41.8N 127 07.7W 132 0 0 1963 7 1 50 50.7N 127 20.5W 139 0 0 1963 36 6 49 09.5N 123 40.5W 102 0 1963 129 9 56°01.0N 154 28.0W 231 1964 154 4 54°02.0N 132035.0W 16 0 0 1965 81 8 54 31.0N 132 37 .0W 370 0 0 1968 12 8 50 43.0N 128 30.0W 90 0 1971 63 9 54 31.8N 132039.8W 366 - 21 - Pycnopodia he1ianthoides Year St. No. Non. Latitude Longitude Depth (M) 0 1964 154 4 54°02.ClN 132 35.'JW 16 Rathbunaster Ca1ifornicus Year St. No. Mon. Latitude Longitude Depth (M) 0 0 1968 35 9 48 23.2N 126 05.5W 360 0 1968 47 9 48 47.0N 126°31.0W 285 Solaster borealis Year St. :~o . Mon. Latitude Longitude Depth (M) 0 0 1964 175 9 47 58.0N 125 47.2W 924 0 1964 214 9 Sl009.0N 130 09.0W 8S8 0 1965 66 8 53°01. 8N 132 55.0W 1190 0 0 1968 23 9 48 04.2N 125 49.5W 653 0 0 1968 29 9 48 04.2N 126 18.5W 951 1968 46 9 48°44.5N 126°21. 5W 494 1971 39 9 52°05.5N 131 ° 26 . 7W 1061 1971 42 3 48°51. 2N 125007.0W 15 0 1972 609 2 48°24.0N 126 10.!)W 472 0 1972 634 2 52 07.2N 131°28.0W 1230 Solaster dawsoni Year St. :Jo. Mon. Latitude Longitude Depth (M) 0 0 1960 299 6 54 08.9N 132 25.0W 66 0 0 1961 897 4 49 15.0N 125 55.0W 36 0 0 1961 899 5 54 08.8N 132 02.0W 0 30 1963 12 1 50 44.0N 127°01.6W 55 0 0 1964 154 4 54 02.0N 132 35.0W 16 Solaster endeca Year St. ,'Jo. Mon. Latitude Longitude Depth (M) 0 1960 305 9 54011.2N 131 22.4W 20 0 0 1960 333 9 53 37.9N 131 05.0W 46 0 0 1960 997 9 54 07.0N 132 16.0W 49 0 1961 811 4 48 58.0N 125°21. OW 38 0 0 1962 704 6 49 45.9N 123 55.0W 80 0 0 1963 14 1 50 38.3N 126 34.0W 64 0 1963 103 9 57 22.0N 151°19.5W 128 - 22 - Solaster l2axi11atus Year St. No. Hon. Latitude Longitude Depth (M) 1963 S3 S SSo07.0N 14SoS0 .0W 241 1965 23 9 4S004. 2N 12S049 . SW 6S3 Solaster stimpsoni Year St. No. Mon. Latitude Longitude Depth (M) 0 1934 21S 4 49 10.3N 126°31. 3W 102 1935 7 6 S2009.0N l31014.0W 4S 0 S3044.0N 131 20.0W 17 1960 lS3 S 0 1960 322 9 S2°S9.0N 12'1 2S.0W 22 0 1960 332 9 Sl 31.0N 12S03S.0W 22 0 1960 333 9 S3037.9N 131 1S.0W 40 0 1961 S03 4 4S032.JN 123 1S.0W 31 0 1961 Sll 4 48 S8.0N 125°21. OW 38 1963 2 1 50036 .9N 126 Os 7 . 7W 26 0 1963 3 1 S0037.7N 127 03.3W 60 1965 6 5 S4 °10 . 3N l31050.0W 73 0 1966 30 6 51 20.0N 127°42.0W 160 1972 606 2 4S024.5N 126 °os .OW 360 Sty1asterias forreri Year St. ~~ o . Mon. Latitude Longitude Depth (M) 0 1934 120 5 49 00.5N 125044.3W 35 1934 21S 4 4'1010.3N 126°31. 3W 102 0 1935 31 7 53 09.0N l32 °21. OW 0 0 25 1935 35 7 52 20.0N 132 00.0W 0 55 1960 291 6 54 0S.1N 131°51. OW 40 1960 297 9 54 °0 7. 3N 132016 . 1W 75 1960 299 6 54 °OS. 9N l32°2S.0W 66 0 0 1960 99S 7 54 06.5N 132 27.0W 22 0 0 1962 26 52 0S.0N 131 09.9W 27 8 0 1962 710 6 49 45.0N 1230SS.0W 20 1963 113 9 56°46.0N lSl043 . 0W 220 1964 203 9 50000.0N 12S00S.1W 165 0 1964 215 9 Slo19.SN 129 4S.0W 230 0 1966 34 6 51 45 .0N 12S000.0W 100 Sty1asterias forreri Year St. No. Mon. Latitude Longitude Depth (M) 1966 61 10 51 °S5 .ON l31 °04 . 2W 730 1969 7 6 54 °12 . 9N 132057.3W 36 0 1969 31 S Sl004.SN 127 26.SW 110 1969 34 S 51 0~1. 4N 128°31. OW 106 1970 25 6 51 °07 . 3N 127049.3W 9S - 23 - Tarsaster a1askanus Year St. No. Hon. Latitude Longitude Depth (M) 1964 175 9 47 0 58.0N 1250 47. 2W 924 0 0 1966 77 10 52 58 . 5N 133 04.2W 2100 Zoroaster everrnanni Year St. t;o. Mon. Latitude Longitude Depth (M) 1964 184 9 48 0 38.5N 126°44.5W 1353 1965 66 8 530 01.0N 1320 55.0W 1190 1965 68 8 530 01.0N 1320 55.0W 1000 1968 31 9 48°16.3N 1260 13.6W 1061 1971 42 3 48°51. 2N 1250 07.0W 15 Zoroaster everrnanni rnordax Year St. 1,10. Mon. Latitude Longitude Depth (M) 1971 42 3 480 51. 2N 1250 07.0W 15 1972 634 2 52 0 07.2N 1310 28.0W 1230 Zoroaster ophiurus Year St. No. Hon. Latitude Longitude Depth (M) 1964 209 9 500 25.7N 129°11. OW 2013