INSECTA MUNDI. Vol. 9. No. 1-2. March - June. 1995 53

A Report on a Collection ofCeratopogonidae (Diptera) from Rondonia (Brazil) and Iquitos (Peru) 1. Tribes Palpomyiini and Stenoxenini

Andrea Dippolito and Gustavo R. Spinelli Instituto de Limnologia "Dr. Raul A. Ringuelet" Casilla de Correo 712, 1900 La Plata, A.rgentina

and

Willis W. Wirth FlO! ida State Collection of 1304 NW 94th St., Gainesville, Florida 32606, U.S.A.

Abstract: Examination of two large collections of from Rondonia, Brazil, and Loreto, Peru, resulted ill records of II) species ill the triBes Palpomyiini and Stenoxenini, ineluding three previously undeseriBed: schmitzorum Dippolito & Spinelli, pseudolacustris D. & S., and_Stenoxenus aductus D. & S. from Rondonia. The hItherto unknown male of PalDomyia versicolor Macfie is descrIbed and figured.

Introduction Tribe Palpomyiini This is the first paper in a series comprising a taxonomic study of ceratopogonids recently col­ Bezzia clavipennis Spinelli & Wirth lected by U. Schmitz in Rondonia, Brazil, and by J. Castner and P. Skelleyin Loreto, Peru. Datafor the Bezzia clavipennis Spinelli & Wirth, 1989: 775 (female; Brazil, collections are as foHuws. BRAZIL. Rondonia, 62 Colombia, Ecuador, venezuela). km SW Ariquemes, vic. Rancho Grande, iv-ix, 1992, DIstrIbutIOn. BraZIl, ColombIa, Ecuador, Ven­ U. Schmitz, UV light trap (abbreviated RRG in text below).PERO: Dept Loreto, 40 km NE lqmtos, ezuela. Explorama Inn on Rio Amazon, 22.viii-v.ix.1992, J. Castner & P. Skelley (ExI); and 160 km NE Iquitos, SpecimenExamined. One female from Rondonia Explorama Camp, Rio Sucasari, 2 km from Rio (RRG). Napo, 27-31.viiL1992, J. Castner &P. Skelley (ExC), The present contribution deals with the preda- ceous midges belonging to the tribes Palpomyiini Bezzia mazaruni Macfie and Stenoxenini. For general ceratopogonid termi Bezzia mazaruni Macfie, 1940a: 193 (female; Guyana); Lane, nology see Downes & Wirth (1981); for special 1958' 27 (in key); Spinelli &Wirth, 1989' 766 (redescrihed' telms dealing with the tribes Palpomyiini and figs., synonymy, distribution). Stenoxenini see Wirth et al. (1974). The holotypes Bezzia coloradensis Wirth, 1952: 238 (male, female; California); and allotypes of the new species from Brazil are Dow & Turner, 1976: 131 (redescribed; figs.; distribution); Wirth, 1933: 301 (redescribed; figs.; distribution). deposited in the lnstituto (lswaldo Cruz, RIO de Janeiro, Brazil; those from Peru are deposited in the National Museum of Natural History, Distribution. Belize, Brazil, Cayman Islands, El Smithsonian Institution, Washin~ton, D.C. Salvador, Guyana, Haiti, Honduras, Jamaica, Ven­ Paratypes and other specimens exam~ned ale in ezuela, USA (California and Florida). the Florida State Collection of Arthropods, Gainesville. 54 Volume 9, nos. 1-2, March-June, 1995, INSECTA MUNDI

Specimens Examined. Two females f:t am 2), claws shm t, slightly cULVed, each with internal Rondonia (RRG). basal tooth. Wing (Fig 3) with costal ratio 0.86 (0.84-0.88, n 2); vem M2 Just sessIle, membrane Benia megatheca Spinelli & "Hirth slightly infuscated. Halter with knob dark brown. Abdomen (Fig. 4): Dark brown. Three pairs of Bezzia megatheca Spinelli & Wirth, 1990: 22 (female; Colombia). gland rods (well visible in paratype). Two sper Distribution Brazil, Colombia unequal, measuring .067 by 0.055 mm, neck 0.01 mm, and 0051 by 0 043 mm, neck 0 016 mm SpecimensExamined. Onefemale from Rondonia whi~h r:~res7ts (RRG) the first record of B Male. Unknown. megatheca rom raZl. Distribution. Brazil, known only flom the type Benda pulchripes Kieffer locality. Bezzia pulchripes Kieffer, 1917: 330 (female; Paraguay); Lane, 191}8: 27 (in key); Mayer, 191}9: 232 (pupa; Brazil); Wirth, 1959a: 234 (male; Brazil); Spinelli & Wirth, 1989: 768 (re­ Types. Holotype female, Brazil, Rondonia, 62 km described; figs ; distribution) SW Ariguemes, vic. Rancho Grande, 20.ix.1992, U. Schmitz, UV light trap. Paratype female, same Distribution. Widely distributed, from Mexico to data except 25 ix 1992 northern Argentina

Specimens Examined. Nine females from es~ecH:Y elrdaug~er Rondonia (RRG). ran e, an ort 0 a, w 0 operated the light trap by which the '.vere Bezzia schmitzorum Dippolito & Spinelli, collected. new species (Figs. 1-4) Discussion. Bezzia sckmitzorum belongs to the Description. Female. Wing length 2.05 (1.97- venustula group of the subgenus Homobezzia. It is 2.18, n - 2) mm; breadth 0.18 (0.71 0.75, n - 8) mm. very similar to B. ca.,!oensis Spinelli & Vlilth, flOm whichitcanbe distinguishedbythe paired, rounded, Head: Dark brown. Eves sA:rated b~/ distance equal to diameter of £5 om atidial acets. An- glabrous areas of the 8tb sternite, by possessing 3 tenna dark brown, bases of flagellomeres pale; ~;~s of gland rod~ rne irc cUJ:oensis), and by the lengths of flagellomeres in proportion of 16-14-12­ 1 erent pattern 0 eg co oratIOn. 12 13 13 14 16 26 29 33 32 43; antennalratio 1.48. Palpus (Fig. 1) dark brown, narrow apex of seg- Pachyhelea pachymera (Williston) ments 3-4 whitish, lengths of segments in propor­ Ceratopogonpachymfrus WHliston, 1900: 224 (female; Mexico). tion of 5-12-15-12-12; 3rd segment with sensilla Probezziapachymera (Williston); Malloch, 1914a: 137 (combina- implanted in small, irregular pits. Mandible with tion). 8 strong teeth Thorax: Dark brown. Scutum covered by fine records); Wirth, 1962: 275 (in key); Wirth, 1974: 53 (distri- pubescence and small hairs, with minute bution); Wirth et ai, 1974; 604 (in list, key); Grogan & anteromedian sine' 3 realar setae, one ostalar. Wirth,. 1980:. 74 (redescribed; figs.;. synonymy; distribution); Ceratopogon magnus Coquillett); 1905: 61 (female; Texas). narrow pale rings subbasally on mid and hind .zohunnseniella magna (Coqlljl!ett);Malloch, 1914b' 227 (in key; femora and all tibiae, fore tibia slightly pale combination). subapically; fore femur armed with 3 ventral spines u'ohannsenomyia magna (Coquillett), Malloch, 1915. 333 (inke)', combination); Johannsen, 1943: 784 (in list N. Amer. spe­ of similar length; hind tibial comb with 8 bristles. cies) Tarsi yellowish blown, talsomeres 4 5 dark, '"en­ Sphae70mias albidiventris Kieffer, l!H7. 316 (female, Colom­ tral palisade setae absenton fore tarsus, inone row bia). ontarsomeres 1-3ofmidleg, in2 rows ontarsomeres lkmohewa albidiventris (Kieffer); Kieffer, 1917' 364 (combina- tion). 1-3 ofhind leg; hind tarsal ratio 240 (230-2 50, n - INSECTA MUNDI, Vol. 9, No. 1-2, March - June, 1995 55

Johannsenomyia latifemoris Ingram & Maefie, 1931: 231 (fe Male. Wing length 1.30 mm, breadth 0.43 mm. male; Argentina; Macfie, 1940b: 75 (male description). Similar to female with usual sexual differences. Fore femur with 2 ventral spines, mid and hind Distribution Widely distributed, from southern femora unarmed Texas to Argentina. Gemtaha(F'Ig. 6): NmthstermterIbbon-shaped, v;ith deep caudomedian excavation, membrane Specimens Examined Three females, 4 males spiculate; 9th tergite tapering to conical shape; from Rondoma (RRG). cercielongated. Gonocoxite straight, 3 times longer t.han broad, with a small ventroapical lobe; gonostylus short, taperIng dIstally, WIth a Palpomyia oliveirai Lane ventromesalcluster ofshort, blacksetae. Aedeagus Palpomywolwezrm Lane, 1947: 443 (female; Brazil); Lane, 1960: triangular, heavilysc1erotized; basalarms recurved, 383 (in key). basalarch O. 5 oftotallength, tip caplike. rammeres narrowl fused anteriorl heavil sclerotized' basal apo emes near y orizon a; is a por ion IVI , each with bulbous tip, then recurved in a long SpecimenExamined. One female from Rondonia slender proJectIon. (RRG) Distribution. Brazil, known only from the type Palpomyia pseudolacustris locahty. Dippolito & Spinelli, new species (Figs. 5-6) Types. Holotype female, allotype male, Brazil, Description. Female. Wing length 2.35 mm, Rondonia, 62 km SW Anquemes, ViC Rancho breadth 0 75 mm. Head: Reddishbrown. Eyes sepa- Grande, 25.iv.1992, U. Schmitz, UV light trap. rated by a dIstance equal to dIameter of 3 omma­ Paratypes, 9 males, as folloW's: same data as types, tidial facets Antenna dark brown, bases of 4 males; same data except 27.iv.1992, 1 male; flagellomeres pale; lengths offlagellomeres in pro- Lv.1992, 3 males; 20.ix.1992, 1 male. portionof12 12 12 11 11 12 13 14 24 26 29 29 35; antennal ratio 1.47. Palpus brown, lengths of Discussion. This species, a typical member ofthe segments in proportion of 21-35-39-29-37. Man­ Palpomyia tibialis gIOup, is very similar to P. dible with 10 t.eeth lacustris Lane, Forattini & Rabello, from which it Thorax: Scutum blackIsh, WIthout anterIor can be dIstmgUlshed by elongated gonocoxIte with spine. Legs dark brown, base and narrow apex of ventroapicallobe, and by the parameres with b"11- fore femur slightly paler; fore and mid femora with bous tip and recurved projection. 3, hind femur with 5-6 ventral spines, hind tibial comb with7 bristles. Tarsiyellowish, 5thtarsomeres Palpo1n,Jda l!ersicolor Macfie (Figs. 7-9) palpomylo "erSlc%r Martie, 1939a' 215 (lema Ie; Brazi 0; I,ane, basItarsus 0 mI leg, In two rows on tarsomeres ­ 1960· 382 (in key) 2 on hind leg; a pair ofstout, black, apieal spines on tarsomeres 1-3 of mid le~; 5th tarsomeres with Diagnosis. Female. A medium-sized species of the Palpomyia distincta group. Thorax blackish, abdomen pale brown Legs yeIlowisb brown, distal 1/3 of hind femur dark brown; base and apex of extending to 0.88 oftotal length; membrane deeply tibiae brown, these markings more extensive on infuscated, veins dark brown; venation as figured. hind leg; tarsomeres 3-5 infuscated, fOle femur Halter knob dark brown. swollen, fore tibia arcuate; all femora armed with Abdomen: Darkbrown' a n r o g an ro s. wo ovoi sper on hind leg. Second radial cell 3 times longer than neck;;~lll~fu;l, measuring a 098 by 0091 mm, and 1st. Halter knob whitish No gland rods VIsible Two spermathecae. 56 Volume 9, nos. 1-2, March-June, 1995, INSECTA MUNDI

Male. Genitalia (Figs. 7-9). Ninth stemite 3.5 Paryphoconus mayeri Wirth times broader than long, with broad, shallow Pmyphoconus mayel iWirth, 1959a. 236 (female, Brazil), Mayer, caudomedian excavatIOn; 9th tergite long, slIghtly 1959: 232 (pupa); Lane, 1961b: 454 (in key); Spinelli & tapsring, csrci slongatsd. Gonocoxits 2.5 timss Wirth, 1984: 884 (in key). longer than broad, with prominent mesoventral setoselobe; gonostyIusslendeI, cuned, withpointed Distribution Brazil tip, 0.5 as long as gonocoxite, with mesoventral Specimens Examined. Two females from Rondonia (R.EG) rotized, recurved basal apodeme; distal portion Paryphoconus nigripes Macfie stout wiLh blunt Lip. Pmyphoconus ltigl ipes Macfie, 1939b. 8 (female, Argentina), Lane, 1956: 303 (in key; Brazilrecords); Lane, 1961b: 455 (in key); Spinelli & Wirth, 1984: 88G (in key). Distribution. Brazil. DIstnbutIOn. Argentma, BraZIl, Guyana. Specimens Examined. One female, 2 males from Rondonia (RRG). Specimen Examined. One female from Rondonia (RRG). Discussion. The female here recorded agrees perfectly with Macfie's description of P. versicolor. This is the first description of the male of this Paryphoconus subflavus Macfie species. Pmyphocoltus subflavus Macfie, 1940c. 23 (female; Guyana); Lane, 1956: 300 in key); Lane, 1961b: 455 (in key); Wirth & Ratanaworabhan, 1912' I;{76 lredescnbed; hgs ; dlstIlbu- maya Spinelli & Wirth tion); Spinelli & Wirth, 1984: 887 (in key; synonymy; wing photo). Phaenobezzia maya Spinelli & Wirth, 1986: 234 (female, male; Paryphoconus travassosi Lane,1956: 304(female; Brazil); Lane, Belize). 1961b: 458 un key; record type-locality); WlIth & Ratanaworabhan, 1972: 1378 (redescribed; figs.; Brazil). Distribution. Belize, Brazil, Costa Rica, EI Salva- dor, Honduras, Mexico, Panama, Texas. Distribution. Brazil, Colombia, Guyana.

Specimens Examined. Two females from SpecimenExanrined. One female from Rondonia Rondonia (RRG), which representthe first record of (RRG). P. maya for Brazil. StenoxenUB aductuB Dippolito & Spinelli, TribeStenoxenini new species (Figs. 10-13)

Paryphoconus aemulus Macf'ie rea t 1.06 mm. ea : Dar rown, c1ypeus ye ­ Paryphoconus aemulusMacfie, 1940a: 180(male; Guyana); Lane, 1946' 206 (furoa Ie; Rrazi I); Wirtb & Ratanaworabban, 1972' 10vy'. Antenna ",vith flagellomeres in proportion of 1371 (female notes; figs.; records Brazil); Spinelli & Wirth, 26-15-15-14-14-12-12-13-45-45-43-44-49; antennal 1984: 888 (female notes' III key!. ratio 1.87; flagellomeres 3-10 blackish with numer- ous stout black hastate setae, 11-15 pale brown Distribution. Brazil, Guyana, Peru. WIth flOe scattered haIrs (FIg. 10). Palpus (FIg. 11) yellowish, external side brownish; lengths of seg- Specimen Examined. One female from Iquitos ments in proportion of 6-14-24-26. Mandible with (ExI),whiehrepresents the firstreeord ofP. aemu1us 5 teeth. for Peru. Thorax: Blackish. Legs yellowish brown, nar­ row apex ofhind tibia brown; tarsomeres 1-3 oHare and mid legs and 5th tarsomere of hind leg mfuscated. Wmg (FIg. 12) WIth membrane hyalme; INSECTA MUNDI, Vol. 9, No. 1-2, March - June, 1995 57 costal cell much broadened anteriOr to medial fork, Annals ofthe Entomological Society ofAmerica 36. rom crossvein obsolete; medial fork broadly sessile; 763·791. vem M2 sharply elbowed near base, well separated Kieffer, J. J. 1917. Chironomides d'Amerique con­ from vein M3+4; distal section of vein M3+4 1.2 serves au Musee National Hongrois de Budapest. l ...nnales Musei Nationalis Hungariei 15: 292 364. times as long as its stem; anal lobe well developed. Lane, J. 1946. New Neotropical Ceratopogonide Halter (pedicel and knob) dark brown. (Heleidae)(Diptera, Nemoeera). Revista de Abdomen: Dark brown. Five pairs of gland Entomologia 17: 202 215. rods, the one extending from segment 8 more scle­ Lane, J. 1947. Novas espeCles de Pa[pomyw do Brasil rotized Spermathecae (Fig 13) oval, without scle- (Dipt€lra, C€lratopogonida€l). R€lvista d€l Entomologia rotized neck; very large, measuring 0.230 by 0.092 18: 438-447. mm. Lane, J 1956 On "Paryphoconlls" and "Stenoxenlls" (Dlptera, Ceratopogomdae). Revlsta BraslleIra de Bi~ln~~fl· 2:-~8 Male. Unknown. Lane,. .:Oneo~oplcal Eezzw. RevlstaBrasilella de Entomologia 8' 25-36 Distribution. Brazil, known) only from the type Lane, J. 1960. Additionaldataon"Palpomyia" (Diptera, locality. Ceratopogonidae). Revista Brasileira de Biologia 20: 381-389. Lane, J. 1961a. Further notes on Neotropical Type. Holotype female, Brazil, Rondonia, 62 km Ceratopogonidae (Diptera). Revista Brasileira de SW i\riquemes, vic Rancho Grande, 1 v 1992, II Biologia 21. 37-44. Schmitz, UV light trap. Lane, J. 1961b. The genera Stenoxenus and Pmyphoconus in the Neotropics (Diptera. Discussion This species is readily distinguished Ceratopogonidae). Studia Entomologica 4: 452-458. from its Neotropical congeners by Its very large Macfie, J. VI. S. 1939a. A report on a colleetion of Brazilian Ceratopogonidae (Dipt.). Revista de spermatheca without sclerotized neck. Because of Entomologia 10: 187 219. the wing venation and many other characters it Macfie, J. W. S. 1939b. A key to the species of resembles S. johnsoni Coquillett, but the latter Ceratopogonidae akin to Macropeza Mg. (Diptera). species has only one pair ofgland rods and the vein Transactions of the Royal Entomological Society of M2 nearly reaches vem M3+4. London 89: 1-12. Macti€l, J WS 1940a. Ceratopogonidae (Diptera) from Bntlsh GUIana and Tnmdad. Proceedmgs of the References Royal Entomological Society of London (B) 9' 179- CoquIllett, D. W. 1905. New nematocerous Diptera 195. from North America. Journal of the New York Macfie, J WS 19411b I:eratopogonidae (Iliptera) Entomological Society 13: 56-69. from North-eastern Brazil. Proceedings ofthe Royal Dow, M. I., and Turner, E. C. 1976. A taxonomic Entomological SocietY of London (B) 9'73-79 revision ofthe Nearctic species ofthe genus Bezzia Macfie, J. W. S. 1940c. A report on a collection of (Diptera. Ceratopogonidae). Virginia Polytechnic Ceratopogonidae (Diptera) from British Guiana. Institute and State University Research Division Entomologist's Monthly Magazine 76: 23-32. Bulletin 103. Malloch, J. R. 1914a. Synopsis ofthe genus Probezzia, 1-162. with description of a new species (Diptera). Pro- Downes, J.A., andWirth,vt'. \\T. 1981. Geratopogonidae, ceedings of the Biological Society oHVashington 27. pp. 393·421, In McAlpine, J. F., et al., eds. Manual 137-139. of Nearctic Diptera, Vol. 1. l ...grieulture Canada Malloch, J. R. 1914b. NotesonNorthAmericanDiptera, Monograph 27, 674 pp. with descriptions of a new species in the collection Grogan, W. L., Jr., and Wirth, W. W. 1980. The ofthe Illinois State Lahoratory of Natural History. taxonomic status of the predaceous midge Bulletin ofthe Illinois State Laboratory of Natural Pachyhelea pachymera (WiIIiston)(Dlptera: History 10: 213-243, 3 plates. Ceratopogonidae). Proce€ldings of the Entomologi- Malloch, J. R. 1915. The Chironomidae, or midges, of cal SOCIety of Washmgton 82: 74-80. Illinois, with particular reference to the species Ingram, 1\, and Macfie, JWS 1931 Diptera of occurring in th€l Illinois Riv€lr. Bull€ltin of the Patagoma and South ChIle. Part II, Fasc. 4, pp. 155· IlhnOIs State Laboratory of Natural HIstory 10: 232 London 275-543, 24 plates Johannsen, O. A. 1943. A generic synopsis of the Mayer, K. 1959. DIe Puppen braslhamscher HeleIden Ceratopogonidae (Heleidae) ofthe Americas, a bib· (Diptera) Deutsche Entomologische Zeitschrift (N liography, and a list ofthe North American species. F.) 6: 230-233. 58 Volume 9, nos. 1-2, March-June, 1995, INSECTA MUNDI

Spinelli, G. R. 1983. Notas sobre Ceratopogonidae Wirth,W. W. 1959a. Newspecies andrecordsofHeleidae (Diptera: Nematocera) de la Republica Argentina. from Brazil. Deutsche Entomologische Zeitschrift III. Nuevos apoltes al conocimiento de la tIibu (N.F.} 6. 234-137. Palpomyiini. Revista de la Sociedad Entomologica Wirth,W. W. 1959b. Pachyhelea, a newgenusofAmerican :Argentina 42: 17 24. Celatopogonidae related to Palpomyia (Diptela). Spinelli, G. R., andWirth,W. W. 1984. A review ofthe Bulletin ofthe Brooklyn Entomological Society 54: Neotropical predaceous midge genus Paryphoeonus 50-52. (Diptera: Ceratopogonidae). Proceedings of the Wirth,W. W. 1962. A reclassification ofthe Palpomyia- Biological Society of Washington 97: 882-908. Bezzia-Macropezagroups, anda revisionofthe North Spinelli, G. R., and Wirth, W. W. 1986. The Neotro American Sphaeromiini(Diptera, Ceratopogonidae). pica! species of Phaenobezzw (Diptera: Annals ofthe Entomological Society ofAmerica 55: Ceratopogonidae). The Florida Entomologist 69: 272287. 231-236. Wirth, W. W. 1974. A catalogue of the Diptera of the Spinelli, G R, and Wirth, WW 1989 The Neotro- Americas south of the United States. 14. plcalpredaceousmIdges of the genusBezzta(Dlptera: Ceratopogonidae. Museumde ZooIogia, Umversidade I :eratopogomdae) Part I 'I'heglabra and bre"icornis de Sao Paulo, Fascicle 14, pp 1-89 groups. Limnobios 2: 762-778. Wirth, W. W. 1983. The North American predaceous Spinelli, G R.,and Wirth, W. W. 1990 Neotropical midges of tbe BezzlO bzcolor group (I hptera' predaceous midges of the genus Bezzia (Diptera: Ceratopogonidae). The Florida Entomologist 66: Ceratopogonidae). Part III. The gibbera group of 292-310 species. Insecta Mundi 4: 11-32. Wirth, W. W., and Ratanaworabhan, N. C. 1972. A Williston, S. W. 1900. Supplement (part), pp. 217-248. reVISIon of the tnbe StenoxenlDl (Dlptera: In: Godman, F. D., & O. Salvin, eds. Biologia Ceratopogonidae) Annals ofthe Entomological So- CentIali Americana. Zoologia - Insecta - Dipteza. ciety ofAmerica 65: 1368-1388. Vol. 1, 378 pp. London. Wirth,W.W., Ratanaworabhan, N. C., andBlanton,F. Wit th,W. W. 1952. The IIeleidae of CalifOInia. Univer­ S. 1974. Synopsis ofthe genera ofCeratopogonidae sity ofCalifornia Publications in Entomology 9: 95- (Diptera). Annaies de Parasitologie humaine et 266. comparee 49. 595-613. INSECTA MUNDI. Vol. 9. No. 1-2. March - June. 1995 59

III

Figules 1-4. Bezzia schmitzOl um, female, 5-6, Palpomyiapseudolacustl is, 5, female, 6, male. 1, palpus, 2 femola and tibiae of (top to bottom) hind, mid and fore legs; 3, 5, wing; 4, spermathecae and tip ofabdomen; 6, genitalia. 60 Volume 9, nos. 1-2, March-June, 1995, INSECTA MUNDI

/ '\ ...... -/ ~'.:'. r .~ ;r=-/\ ....., \- .="..... / :~:~-"' ';:"~-~-".: .. (/ ..,'. " T 7 I I \ , ~. '. .. " .:.,...:; " " r \. ,; . / \ 1'/11 7J 0 :;" ,."\ -. '" "'- ',: / f f , .:' \~ M''. fhi \ \ J\J , /1 / (/1,'11 f 11 y , I( ~~'PY r::\ I I \\\)'-'V," / ~ \.C'- - J -y ~LJ ~ { 11 ""'~'-./~

~ -"~ I\ ········7:·~

I .,# '~ ••.~l-· .... ;.//; . \ 7 , . l~ 13 \/

Figures '7-9. Palpomyla verslcolor, male; 10-13, Stenoxenus aductus, female: 7, genitalia, aedeagus and parameres omitted' 8 aedeagus; 9, parameres, Hl, antennal segments 10-11; 11, paJpus; 12, wing; 13, spermatheca.