l*l 331:!y"
Checklist of
of Canada and Alaska
Checklist of BEETLES of Canada and Alaska
Edited by
Yves Bousquet Biosystematics Research Centre Ottawa, Ontario
Research Branch Agriculture Canada Publication 186UE t99l oMinister of Supply anrl Services Canada 1991
Available in Canada through
Assoeiated Bookstores and other borksellers or by mail fr,rm
Canada Com-nunication Group Publishing Ottawa, Canada KlA0S9 -
Cat. no. A43-1861/1991E ISBN 0-660-13?67-4
Canadian Cataloguing in Publication Data
Checklist c f beetles of Canada and Alaska
(Publication ; 18'61/E)
Includes bibliographical references. Cat. no. A43. 1861i1991E ISBN 0-660- r3767-4
1. Beetles-(lanada-Nomenclature. 2. Beetles- Alaska-Nomr:nclature. I. Bousquet, Yves. II. Canada. Agriculture Canada. Research Branch. III. Series: .?ublication (Canada. Agriculture Canada). English ; 1861/E. QI581.C4 1991 595.76 C91-099101-4
Stafr Editor Frances Smith CONTENTS
List ofcontributors v SuperfamilyScarabaeoidea 143 Acknowledgments v Family Lucanidae (J. McNamara) 143 Summary vi Family Passalidae (J. McNamara) 145 R6sum6 vi Family Scarabaeidae (J. McNamara) 145 Synoptic table 1 SuperfamilyByrrhoidea 158 Introduction 4 Family Byrrhidae (P.J. Johnson) 158 Format 4 SuperfamilyBuprestoidea 160 Subsequent revisions ofthe checklist 5 Family Buprestidae (A. Davies) 160 Abbreviations and symbols 5 SuperfamilyDryopoidea 169 ORDERCOLEOPTERA 6 Family Ptilodactylidae (L. LeSage) 169 Family Heteroceridae (L. LeSage) 169 SuborderArchostemata 6 Family Limnichidae (L. LeSage) 171 Family Cupedidae (J.M. Campbell) 6 Family Dryopidae (L. LeSage) 771 Family Micromalthidae (J.M. Campbell) 6 Family Elmidae (L. LeSage) 172 SuborderAdephaga 7 Family Psephenidae (L. LeSage) 174 Family Rhysodidae (Y. Bousquet) 7 L74 Family Trachypachidae (Y. Bousquet) 7 SuperfamilyElateroidea Family Artematopidae (Y. Bousquet) 174 Family Carabidae (Y. Bousquet) 8 Family Elateridae (Y. Bousquet) 175 Family Haliplidae (R.E. Roughley) 60 Family Throscidae (Y. Bousquet) 186 Family Amphizoidae (Y. Bousquet) 61 Family Eucnemidae (Y. Bousquet) 186 Family Noteridae (R.E. Roughley) 61 Family Dy'tiscidae (D.J. Larson and R.E. Superfamily Cantharoidea 188 Roughley) 62 Family Lycidae (J. McNamara) 188 Family Gyrinidae (R.E. Roughley) 72 Family Phengodidae (J. McNamara) 190 (J. 190 Suborder Polyphaga 74 Family Lampyridae McNamara) Family Cantharidae (J. McNamara) 192 SuperfamilyStaphylinoidea 74 195 Family Hydraenidae (R.E. Roughley) 74 SuperfamilyDermestoidea Family Derodontidae (D.E. 195 Family Ptiliidae (J.M. Campbell) 75 Bright) Family Nosodendridae (J.M. Campbell) 196 Family Limulodidae (J.M. Campbell) 76 Dermestidae (Y. Bousquet) 196 Family Agyrtidae (J.M. Campbell) 77 Family Family Leiodidae (S.B. Peck) 77 SuperfamilyBostrichoidea 199 Family Scydmaenidae (A. Davies) 82 Family Bostrichidae (J. McNamara) 199 Family Micropeplidae (J.M. Campbell) 84 Family Lyctidae (J. McNamara) 20L Family Silphidae (J.M. Campbell) 84 Family Anobiidae (J. McNamara) 201 Family Staphylinidae (J.M. Campbell and A. Family Ptinidae (Y. Bousquet) 205 Davies) 86 Superfamily Lymexyloidea 206 Family Scaphidiidae (J.M. Campbell) 124 Family Lymexylidae (J.M. Campbell) 206 Family Pselaphidae (A. Davies) 125 SuperfamilyCleroidea 206 SuperfamilyHydrophiloidea 130 Family Trogossitidae (Y. Bousquet) 206 Family Hydrophilidae (R.E. Roughley) 130 Family Cleridae (J. McNamara) 208 Family Georyssidae (R.E. Roughley) 135 Family Melyridae (D.E. Bright) 2LI Family Sphaeritidae (R.E. Roughley) 135 Superfamily Cucujoidea 213 Family Histeridae (A. Davies) 135 Family Sphindidae (J.M. Campbell) 213 SuperfamilyEucinetoidea I4I Family Nitidulidae (J. McNamara) 2I4 Family Eucinetidae (J.M. Campbell) I4I Family Rhizophagidae (Y. Bousquet) 2I8 Family Clambidae (J.M. Campbell) I4l Family Cucujidae (Y. Bousquet) 219 Family Scirtidae (J.M. Campbell) 142 Family Cryptophagidae (Y. Bousquet) 22I SuperfamilyDascilloidea 143 Family Languriidae (J.M. Campbell) 223 Family Dascillidae (J.M. Campbell) 143 Family Erotylidae (J.M. Campbell) 225 Family Rhipiceridae (J.M. Campbell) I43 Family Phalacridae (J.M. Campbell) 226
Ill Family Cerylonidae (J.M. Campbell) 227 Family Anthicidae (Y. Bousquet) 27t Family Corylophidae (J.M. Campbell) 228 Family Pedilidae (Y. Bousquet) 274 Family Coccinellidae (J. McNamara) 229 Family Aderidae (Y. Bousquet) 275 Family Endomychidae (J.M. Campbell) 287 Family Scraptiidae (L. LeSage) 275 Family Lathridiidae (Y. Bousquet) 239 SuperfamilyChrysomeloidea 277 Family Biphyllidae (J.M. Campbell) 247 Family Cerambycidae (J. McNamara) 277 Family Byturidae (J.M. Campbell) 24I Family Bruchidae (J. McNamara) 300 SuperfamilyTenebrionoidea 242 Family Chrysomelidae (L. LeSage) 3C,1 Family Mycetophagidae (Y. Bousquet) 242 SuperfamilyCurculionoidea 323 Family Tetratomidae (L. LeSage) 243 Family Nemonychidae (J. McNamara) 323 Family Ciidae (J. McNamara) 244 Family Anthribidae (J. McNamara) 324 (L. Family Melandryidae LeSage) 245 Family Ithyceridae (J. McNamara) 32:,5 Family Mordellidae (J. McNamara) 248 Family Attelabidae (J. McNamara) 3tl6 Family Rhipiphoridae (J.M. Campbell) 250 Family Rhynchitidae (J. McNamara) 326 Family Colydiidae (Y. Bousquet) 251 Family Brentidae (J. McNamara) 327 Family Zopheridae (J.M. Campbell) 252 Family Apionidae (J. McNamara) 328 Family Tenebrionidae (Y. Bousquet and J.M. Family Curculionidae (J. McNamara) 329 Campbell) 253 Family Scolytidae (J. McNamara) 35€; Familr' Lagriidae (J.M. Campbell) 261 Family Platypodidae (J. McNamara) :]65 Famils Cephaioidae (J.M. Campbell) 262 Famil r Meloidae (J.M. Campbell) 269 ORDERSTREPSIPTERA 366 Famil r Oedemeridae (J.M. Campbell) 266 SuborderStylopidia 366 Famil r' Prostomidae (J.M. Campbell) 267 Family Corioxenidae (S.B. Peck) 366 Famil Synchroidae (L. LeSage) 267 Family Halictophagidae (S.B. Peck) 336 'z Familr Mycteridae (J.M. Campbell) 267 Family Stylopidae (S.B. Peck) 367 Famil,r Boridae (J.M. Campbell) 268 Famil r Pythidae (J.M. Campbell) 268 References 368 Famil r Pyrochroidae (J.M. Campbell) 269 Index 397 Famil,r Salpingidae (J.M. Campbell) 270
iv LIST OF CONTRIBUTORS
Y. Bousquet, Biosystematics Research Centre, D.J. Larson, Department of Biology, Memorial Agriculture Canada, Ottawa, Ontario K1A 0C6 University of Newfoundland, St. John's, D.E. Bright, Biosystematics Research Centre, Newfoundland AlB 3X9 Agriculture Canada, Ottawa, Ontario K1A 0C6 L. LeSage, Biosystematics Research Centre, J.M. Campbell, Biosystematics Research Centre, Agriculture Canada, Ottawa, Ontario K1A 0C6 Agriculture Canada, Ottawa, Ontario K1A 0C6 J. McNamara, Biosystematics Research Centre, A. Davies, Biosystematics Research Centre, Agriculture Canada, Ottawa, Ontario KlA 0C6 Agriculture Canada, Ottawa, Ontario K1A 0C6 S.B. Peck, Department of Biology, Carleton P.J. Johnson, Department of Entomology, University, Ottawa, Ontario KlS 586 University of Wisconsin, Madison, Wisconsin R.E. Roughley, Department of Entomology, 53706 University of Manitoba, Winnipeg, Manitoba R3T 2N2
ACKNOWLEDGMENTS
We wish to thank the following persons for Moldenke (Clytrinae); H.G. Nelson (Dryopoidea); reviewing sections of the checklist: R.L. Aalbu A.F. Newton Jr. (Leiodidae, Staphylinidae); S. (Tenebrionidae); R.S. Anderson (Silphidae, Oygur (Gyrinidae); J. Pakaluk (Corylophidae); J.D. Curculionidae); F.G. Andrews (Lathridiidae); LS. Pinto (Meloidae); D. Pollock (Mycteridae, Boridae, Askevold (Chrysomelidae); F.M. Atton (Gyrinidae); Pythidae, Salpingidae); V. Puthz (Steninae); E.G. E.U. Balsbaugh (Chrysomelidae); R. Baranowski Riley (Cassidinae); W.T. Schultz (Eumolpinae); T.N. (Leiodidae); W.F. Barr (Cleridae); E.C. Becker Seeno (Chrysomelidae); R.B. Selander (Meloidae); (Elateroidea); H.P. Brown (Dryopoidea); D. S.A. Slipinski (Cerylonidae); C.L. Staines Chandler (Anthicidae, Pselaphidae, Melandryidae, (Hispinae); W.E. Steiner Jr. (Phalacridae); J.B. Tetratomidae, Synchroidae, Scraptiidae); J. Cicero Stribling (Ptilodactylidae); M. Thayer (Lampyridae); W. Connell (Nitidulidae); M. (Staphylinidae); M.C. Thomas (Cucujidae, Daccordi (Chrysomelinae); J.G. Edwards Prostomidae); C.A. Triplehorn (Tenebrionidae); R.E. (Synetinae); D.G. Furth (Chrysomelidae); R.E. White (Anobiidae, Chrysomelidae); D.P. Wooldridge Gordon (Coccinellidae): L.H. Herman (Limnichidae); D.K. Young (Scirtidae, (Staphylinidae); A.T. Howden (Curculionidae); H. Pyrochroidae); P. Zwick (Leiodidae). Howden (Scarabaeidae); C. Johnson (Ptiliidae); J.B. Thanks are also extended to B.F. Carr, J.L. Karren (Chlamisinae); J.M. Kingsolver (Bruchidae); Carr, and D.W. Langor, for providing several J.L. Krysan (Chrysomelidae); S. Laplante (Cleridae, provincial records; to D. Beauchamp, B. Walter, L. Cerambycidae); J.F. Lawrence (Melandryidae, Robillard, and R. Hutchinson, for typing the check- Tetratomidae, Synchroidae, Scraptiidae); I. Lobl list; and to A. Smetana, for checking the termina- (Scaphiidae); W.V. Miller (Heteroceridae); A.R. tion of all species-group names. SUMMARY
All species and subspecies of beetles known to presence or absence of the taxa in each province occur in C.anada and Alaska are listed along with and territory, in Labrador, and in Alaska. their synonyms; these names are arranged alpha- Introduced and holarctic species are highlighted. betically t,nder the respective genus-group names. References to pertinent identifrcation keys are indi- Valid genus-group and family-group names are cated under the corresponding supraspecific taxon. listed in p aylogenetic order. No new taxa, combina- The work also includes a brief introduction for tions, or synonyms are proposed. Species distribu- every family treated and an index to all supraspe- tions are presented in tabular form, indicating the cifrc names.
RESUME
L'ouvrage r6pertorie toutes les espdces et sous- tabulaire, indiquant la pr6sence ou l'absl:nce des espdces dt,: Col6optdres connues du Canada et de taxa dans chaque province et territoire, le l'Alaska arnsi que leurs synonymes; ces noms sont Labrador et l'Alaska. On met en relief les espbces 6num6r6s par ordre alphab6tique d la suite du nom introduites ou holarctiques et on mentionne, sous du groupe-genre correspondant. Les noms valides le taxon suprasp6cifique correspondra.nt, les des group':-genre et groupe-famille sont pr6sent6s r6f6rences contenant des tableaux de d61;ermina- par ordre phylog6n6tique. Cette liste ne propose tion utiles. L'ouvrage comprend aussi une courte aucun nouveau taxon, combinaison ou synonyme. introduction sur chaque famille consid6r'6e ainsi La r6partibion des espdces est pr6sent6e sous forme qu'un index des noms suprasp6cifrques.
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More than 250 000 available for Canada and Alaska. Leng's catalogue species have been described, and the actual number (1920) and its supplements (Leng and Mutchler may easily exceed a million. Given such staggering 1927, 1933; Blackwelder 1939a; Blackwelder and frgures, it is not surprising that Coleoptera can be Blackwelder 1948) as well as the world-based found neally everywhere, from the tropics to the Coleopterorum Catalogus list most of the species Arctic and from caves to the summit of high moun- that occur in Canada and Alaska. However, these tains. Coleoptera species are particularly abundant publications are out of date for many groups, and on vegetation, inside rotten wood, under the bark of their distribution data are incomplete. Beaulne dead trees, and in soil, ground litter, fungi, dung, (1913-1929)started a checklist of the beetles of carrion, rodent burrows, and nests. Several groups Canada but never completed it, and as pointed out are adapted to life in fresh or brackish water. by Lindroth (1966) for the ground beetles, many of Although some dytiscids can venture into salt water Beaulne's distributional records are inaccurate. and some species do complete all stages of their life This checklist is intended to provide the correct cycle in the intertidal zone where they are periodi- names of the beetles that occur in Canada and cally immersed, no beetles have become completely Alaska, give a brief survey of the distribution of the adapted to marine life. Beetles are important to taxa in the area covered, and lead the user's to the humans because many species are beneficial in the proper publications for the identification of the biological control of pest arthropods and weeds or taxa the checklist of Strepsiptera has been added are major pests of stored products, fabrics, forests, at the end simply because the group has often been wood prodructs, and crops. included in the Coleoptera. FORMAT J.C. Fabriciusr respec- With some exceptionsreflecting the author's J.L. LeConte, G. Horn, and personal perspective, the concept of the families tively. An asterisk [*] placed after a species-group used in this checklist follows that of Lawrence name indicates that the taxon is Holarctic, a dagger (1982a). N,najor changes concern the trachypachids, ttl that it is introduced in North Americra, and a limulodids, scaphidiids, lyctids, lagriids, pedilids, double dagger t{l that it is probably introduced in rhynchitids, scolytids, and platypodids, which are North America. treated as distinct families, and the leptinids, AII synonyms based on material collected in trogids, and geotrupids, which are included respec- North America north of Mexico should be included tively within the Leiodidae and the Scarabaeidae and are listed in italics. Synonyms of Hola:nctic and for the last two groups. The superfamily concepts introduced species based on material frorri outside and the listing of the families follow Lawrence North America have been included occasionally, (1982a). particularly when the name has been in use in The classification and information for each North American literature. Misidentifications, family are the responsibility of the respective spelling errors, and invalid emendations are not authors. No new taxa, combinations, or synonyms llited. The ending of all species-group names has are proporied. Variety and form names published been checked to conform with the gender of their before 1961 are given subspecifrc rank unless they respective genera. were deemed to be synonyms or their status has Valid genus-group names are in capital letters, been changed (article +5 (g) of the International their synonyms in italics. All synonyms crf genus- Code of Zoological Nomenclature, 3rd ed.). group names whose type species are indigengu-s to Varieties and forms published after 1960 are North America north of Mexico should be included. omitted. New distributional records are listed but Some of the synonyms whose type species are not not identified as such. indigenous to the continent are included also when The arLthors of species-group names are listed in the name has been used in North American litera- parentheses when the original combination differs ture. from the one used in the checklist. To avoid confu- References to keys are indicated under the cor- sion, initials are included for several authors, such responding supraspecific taxon. It should be noted as C.R. Sahlberg, R.F. Sahlberg. J.R. Sahlberg, J. that only references including keys useful for the MiiIIer, O.F. Miiller, W. Horn, J.E. LeConte, and O. determination of all or most Canadian taxa of the Fabricius. LeConte, Horn, and Fabricius stand for group treated are considered. Keys published before the 1930s are usually out of date, and most of from other collections have been included. Doubtful them are therefore omitted. records from the literature that apparently are An abbreviation of two letters is used to indi- based on misidentifications or mislabeled specimens cate Canadian provinces and Alaskan records (see are rejected. section entitled "Abbreviations and symbols"). A question mark (?) inserted in a geographic Labrador is considered to be a distinct geographic column indicates that the taxon was recorded in the entity. The distributional citation in the checklist is area, but the record is questionable. The absence of from west to east. a geographic entry after a species-group name indi- The distributional records are based on pub- cates that the taxon was recorded in Canada. but lished records and on specimens in the Canadian the province was not specified. National Collection. For some families. records SUBSEQUENT REVISIONS OF THE CHECKLIST This publication is set as a data base that can sions to their attention. We would also welcome be easily updated to maintain an accurate summary receiving specimens that represent new provincial of our current knowledge of the beetles of Canada or Alaskan records. Specialists and other users and Alaska. The authors intend to maintain a cor- may send material to the Biosystematics Research rected version of the checklist and to republish it Centre for confirmation, and with the sender's per- whenever we have accumulated enough changes. mission, for incorporation as voucher specimens in The authors would appreciate the assistance of all the Canadian National Collection. users of this checklist in calling errors and omis- *TI AB B REV r.rl' O N S AND SYMB O L S AB Alberta NT NorthwestTerritories AK Alaska ON Ontario BC British Columbia PE Prince Edward Island LB Labrador PQ Quebec MB Manitoba SK Saskatchewan NB New Brunswick YK Yukon Territorv NF Newfoundland H Holarctic NS Nova Scotia I Introduced in North America * Holarctic f Probably introduced in North America t Introduced in North America $ No subgeneric name available Micromalthidae Order Coleoptera Suborder Archostemata FamilyCUPEDIDAE reticulatedbeetles J.M. Campbell Three species of Cupedidae occur in Canada. Neboiss (1984) the species of Cupes. Ponornarenko This small, primitive family of beetles contains only (1969) discussed the fossil origins of the Coleoptera 30 extant species around the world, but it is well with particular reference to the srrborder represented by fossils dating back to the early Archostemata and the family Cupedidae, rerviewing Triassic. l,ittle is known about the biology of the the fossil and extant species of the family. Atkins species-the larvae apparently bore into wood, and (1979) cataloged the North American taxa of the the adults iue collected from foliage or under bark. family. The farnily Cupedidae is well known taxonomi- cally. Atk.ns (1963) revised the world species, and BC (1); oN (2);PQ (1) Subfamily CUPEDINAE Tribe Priacmini Genus PRIAC MA LeConte P. serrata (LeConte) BC Tribe Cupedini Genus CUPEII Fabricius C. capitatus tr abricius ON Genus TENO T{ERGA Neboiss T. concolor (Si estwood) ON PQ cinerea (flay) oculata (Oasey) t r iI ineat c; (Melsheimer) Family MICROMAL'THIDAE telephone-pole beetles J.M. Campbell This fz: mily contains only one species, which known to occur in Coleoptera. The larvae are pae- was descri ced from the eastern United States but dogenetic, producing either living larvae or a single has subserluently become established in western egg that develops through a different larval type to Canada, C uba, Brazil, Hawaii, Hong Kong, and produce an adult male (Pringle 1938). South Afrir:a. The larvae are wood borers and are Atkins (1982) cataloged the species for North found in decaying oak logs and in moist, creosote- America. treated timbers and poles. The life cycle of Micromalthus debilis is one of the most complex BC (1) Genus MICROMALTHUS LeConte M. debilis Le(jonte BC Trachvoachidae Suborder Adephaga Family RIIYSODIDAE wrinkled bark beetles Y. Bousquet The family Rhysodidae includes only two and larvae live under the bark of dead trees and in species in Canada. The adults are cylindrical, elon- rotting wood. gate beetles resembling in habitus some species of Bell (1970) revised the North American species. the families Colydiidae or Cucujidae. Both adults eC (1);ON (1) Genus OMOGLYMMIUS Ganglbauer Key to species: Bell and Bell (1982) (Subgenus BOREOGLYMMIUS Bell & Bell) O. americanus (Castelnau) - ON orolas (Newman) e ra r at u s (Audinet-Serville) Genus CLINIDIUM Kirby Key to North American species: Bell and Bell (1975) (Subgenus ARCTOCLINIDIUM Bell) C. calcaratum LeConte - BC Family TRAC IIYPAC HIDAE Y. Bousquet This family includes two genera: Trachypachus, decades, however, some authors have highlighted which has three species in western North America many character states that suggest that and one species in Eurasia, and Systolosorno Solier, Trachypachus and Sysfolosoma are more closely which has two species in Chile. Members of related to Hydradephaga than to Carabidae (see Trachypachus are usually found on dry, more or Kavanaugh (1986) for a different point of view). Iess shaded soil, often under dead leaves. Nothing Lindroth (1961) has redescribed the two species is known about their feeding habits. that inhabit Canada. This group has been included within the Carabidae for a long time. During the last two aK (1);YK (1); NT (1); BC (2); aB (1); SK (1) Genus TRACHYPACHUS Motschulsky T. holmbergi Mannerheim AK YK .\T BC AB SK inermis Motschulsky oregonus Casey specularis Casey T. gibbsii LeConte _BC alticola Casey c alifor nicu s Motschulsky Carabidae Family CARABIDAE ground beetles Y. Bousquet With 9,t6 species and subspecies in Canada and in the family. In recent years, some authc,rs haYe Alaska, thr,: family Carabidae is an important ele- included the rhysodids within Carabidae, but based ment of thr: Coleoptera fauna in the northern part on larval characters I prefer to exclude thenL; on the of North A:nerica. In Canada and Alaska members other hand, some others consider the cicinclelids to of these species occur in most terrestrial habitats as be a distinct family, a view with which I do not well as in .marshes and bogs; a few species live on agree. Thanks to the work of Wallis (1961) lor tiger trees. Adults of most species are carnivorous or beetles and Lindroth (1961-1969), the carabid fauna omnivorous,; only those of a few groups seem to be of Canada and Alaska is well known taxonomically. primarily phytophagous; larvae of most species are Bell (1990) recently published a key to the North probably ca.rnivorous, but those of a few groups are American genera. known to be phytophagous or parasitic. The Carabidae of North America are here AK(237); YK (164); NT (207); BC (a83); AB (386); grouped in 36 tribes. The classification used is basi- SK (314); MB (350); ON (503); PQ (451); NB (2a3); cally that of Lindroth (1961-1969), with a few modi- NS (24a); PE (137); LB (.94); NF (175) ' II (84); I fications re,flecting recent taxonomic developments (46) Tribe Cicindelini Genus OMUS Eschscholtz Leptontut Casey Megomus Casey O. audouini audouini Reiche BC aequicorr,.is Casey audens Casey borealis ()asey delicatuhn Casey distans Casey oregonentiis Casey paruulus Casey rugipenn,.s Casey tacomae ()asey thoracicus Casey uandykei W. Horn O. dejeani Rei:he BC foueatus l)asey robustus Casey Genus CICIN.)ELA Linn6 (Subgenus CICINDELA s. str.) (Pachydetn Rivalier) (Tribonia Rivalier) C. ancocisconr nsis Harris PQ carolinae Casey dowiana Casey eriensis (iasey C. decemnotata Say YK BC SK albertina Casey lantzi lJ.a.rcts C. depressula depressula Casey AK BC scapularis Casey C. duodecimgrrttata Dejean BC AB SK MB ON PQ NB NS PE LB NF bucolica tlasey edmontor'.ensis Can:r hudsonic,z Casey proteus Birby C. formosa folmosa Say AB SK fletcheri Criddle luxuriosa Casey manitoba Lelrg Carabidae C. formosa generosa Dejean SK MB ON PQ latecincta Godd C. formosa gibsoni Brown SK C. fulgida westbournei Calder AB SK lln- : elegans Calder C. hirticollis athabascensis Graves AB bt\ C. hirbicollis couleensis Graves BC AB C. hirticollis rhodensis Calder _ONPQNBNS_LB NF nigrita Davis shermani Casey C. hirticollis shelfordi Graves AB SK MB C. lengi versuta Casey _BCABSKMB C. limbalis limbalis KIug NT BC AB SK MB oN pe -NS-- NF arnoena LeConte awemeana Casey eldorensis Casey m il ita r is Varas-Arangua spreta LeConte C. limbata hyperborea LeConte NT_ABSKMB LB C. lirnbata nympha Casey AB SK MB C. longilabris longilabris Say AK YK NT BC AB SK MB oN pe NB NS PE LB NF albilabris Kirby nouaterrae Leng terracensis Casey C. nebraskana nebraskana Casey BC ABSK MB calgaryana Casey canadensis Casey chamberlaini Knars montdna LeConte splssllorsis Casey uteana Casey C. oregona guttifera LeConte AK YK NT BC audax Casey oregonella Casey sterope Casey C. oregona oregona LeConte BC oualipennis Casey quadripennis Casey sonoma Casey C. parowana wallisi Calder BC azurea Cald,er C. patruela patruela Dejean ON PQ C. pugetana Casey BC nigerrimoides Hatch C. purpurea audubonii LeConte BC AB SK MB auguralis Casey graminea Schaupp nigerrina Leng C. purpurea purpurea Olivier BC ON PQ NS marginalis Fabricius ramosa Gistel C. repanda novascotiae Vaurie PQ NB NS PE C. repanda repanda Dejean NT BC AB SK MB ON PQ NB NS PE LB NF baltimorensis Hetbst d.uncaniKnats hoosieri Mares maehleri Robinson unijuncta Casey C. scutelaris criddlei Casey _MB C. scutelaris lecontei Haldeman -MBONPQ C. scutelaris scutelaris Say _ABSK C. sexguttata denikei Brown _MBON C. sexguttata sexguttata Fabricius oN PQ NB NS guttata Emmons harrisii Leng illinoensis Mates kansanal(nans leuettei Casey quadriguttata Davis tridens Casey uarians Ljungh uiolacea Fabricius Carabidae C. tranqueba'ica borealis Harris NT BC C. tranqueba:ica kirbyi LeOonte NT_ABSK nae C. tranquebartca roguensis Harris -BCABsx C. tranqueba ica tranquebarica MB ON pe Ns NS PE NF Herbst crinifron.s Casey horiconezsis Leng minor Lt:ng obliquat':.t Dejean turbulenta Casey u ir idula Varas-Arangua uulgaris Say uulgaris;,ninor Haris wichitan.a Casey C. tranquebar:ica vibex Horn BC (Subgenus CI CINDELIDIA Rivalier) C. punctulatri punctulata Olivier AB SK MB ON PQ NB boulderensis Casey chihuah:ne BaLes fontinar,ra Casey jenisoni llistel prolixa (.)asey (Subgenus C' T,INDERA Westwood) (Plectogratr. ha Rivalier) C. tericola cinctipennis LeConte YK _AB SK cyanella LeOonte C. terricola irrperfecta LeConte Bcy C. terricola tcrricola Say SK MB pusilla E ay sayanellt Casey (Subgenus EI,LIPSOPTERA Dohktourow) C. cuprascenr, LeConte MB amnicohL Casey mundukr Casey C. lepida Dejean AB SK MB ON PQ insomnil: Casey C. nevadica k.nausi Leng AB SK MB Tribe Carabini Genus CALOSOMA Weber Key to Norbh American species: Gidaspow (1959) (Subgenus CitLOSOMA s. str.) (Callipa'o Motschulsky) (S y nc alc so ma Breuning) C. lrigidum l.jrby BC SK MB ON PQ NB NS PE Ieuettei Casey (Subgenus CTTLODREPA Motschulsky) C. scmlator (Fabricius.t _ONPQ C. wilcoxi Letlonte _ONPQ (Subgenus C;T LLITROPA Motschulsky) (Paratrooa Vacher de Lapouge) C. exlernum Sayt ON longipenne Dejean (Subgenus CI{RYSOSTIGMA Kirby) (Lyperostenia Vacher de Lapouge) (Tapinosthenes Kolbe) C. calidum (Fabricius) NT SK MB ON PQ NB NS NF comes Casey concretum Casey expansu,n Casey laticolle Casey stellatur,z Casey 10 Carabidae C. cancellatum Eschscholtz BC aenescens LeConte esuriens Casey praestans Casey rectilaterum Casey sagax Casey transuersum Casey C. lepidum LeConte AB SK MB C. obsoletum Say AB SK MB microstictum Casey C. tepidum LeConte BC caelator Casey cogitans Casey indigens Casey irregulare Walker pellar Casey semicupreum Casey (Subgenus CAILISTHENES Fischer von Waldheim) (Callistenia Vacher de Lapouge) (Isostenia Vacher de Lapouge) C. luxatum Say BC SK debile (Casey) diffractum (Casey) etcaratum (Casey) opacum (G6hin) parowanunr (Casey) pimelioides (Walker) pustulosum (Casey) reflexum (Casey) semotum (Casey) striatulum LeConte striatum Breuning tahoense Van Dyke tegulatum (Casey) utense (Casey) uiator (Casey) (?) zimtnermanl (LeConte) C. moniliatum ( Leconte) BC AB SK bicolor (Walker') concinnum (Casey) laqueatum LeConte uancouuericum (Csjkl) C. monticola I Casey) neuadense (Casey) C. subaeneum Chaudoir C. wilkesii (Leconre) Genus CARABUS Linn6 (Subgenus CARABUS s. str.) (Cryocarabus Vacher de Lapouge) (H omoeocar abus Reitter) (.Lic hnocarab us Reitter) (Neocarabus Hatchl C. chamissonis Fischer AK YK NT_AB SK MB ON PQ LB von Waldheim brachyderus Wiedemann groenlandicus Dejean washingtoni Casey C.goryt Dejean ON clarkei Blwnenthal limbatus Say C. granulatus granulatus Linn6t _BCAB MB ON PQ C. granulatus hibernicus Lindrotht _NBNSNF C. maeander maeander Fischer AK NT BC AB SK MB ON PQ NB NS PE LB NF von Waldheimx atlanticus Vacher de Lapouge excatenatus Kraatz excostatus Kraatz h udsonicus Motschulsky 11 Carabidae lapilayi Castelnau lecontei G6hin simoni I{eyden C. nemoralis O.F. Mullert BC AB SK oN PQ NB NS PE NF canadensis Vacher de Lapouge C. vinctus W':ber ON carinatu s Dejean georgiae Cslki interruptus Say ligatus Getrnat (Subgenus DIOCARABUS Reitter) C. taedatus a-gassii LeConte AK tiK BC SK MB ON PQ canadic,ts Roescke coloradensis Breuning gladiatcr Motschulsky montant:.cus Casey oregonetrcis LeConte patulicotlis Casey stocktonensis Casey C. taedatus taedatus Fabricius bacciuorus Fischer von Waldheim bicanaliceps Casey francisutnus Casey rainieri Van Dyke seriatus Wiedemann C. truncatico llis truncaticollis AK tT( NT Eschscholtz ala s ken :,:i s Basilewsky lutshniA ianus Basilewsky (Subgenus H 0MICARABUS G6hin) C. serratus Say BC AB SK MB ON PQ NB NS PE NF canadensis LeConte lineato p unct atzs Dej ean lolumi l4otschulsky uegasensis Casey (Subgenus T,\NAOCARABUS Reitter) C. sylvosus s.ylvosus Say ON PQ lhermin:eri Dejean (Subgenus MEGODONTUS Solier) C. vietinghol hi vietinghoffii NTYKAK Adarr:s* Tlibe Cychrini Genus SPHAIRODERUS Dejean S. canadensiri canadensis Chaudoir oN PQ NS blancha,di Leng palpalis Motschulsky S. Iecontei Ie:ontei Dejean MB ON PQ NB NF bicarinatus LeConte diffractt.ts Casey niagarensis Castelnau S. nitidicoliis brevoorti LeConte SK MB ON PQ NS granulosus Chaudoir S. nitidicollis nitidicollis PQ NF Chevlolat Genus SCAI'HINOTUS Dejean (Subgenus STIAPHINOTUS s. str.) (Anabol'rs Gishel') (Irichroa Newman) (Megaliridia Casey) S. elevatus coloradensis Van Dyke yu_ S. viduus (Dejean) pe NB ir r eg ula r i s Beutenmuller leonard:'.i (Harris) t2 Carabidae (Subgenus NOMARETUS LeConte) Key to species: Gidaspow (1973) S. bilobus (Sayt MB ON PQ NB NS (Subgenus PSEUDONOMARETUS Roeschke) Key to species: Gidaspow (1973) S. merkelii (Horn) - BC idahoensis (Webb) S. regularis (LeContet - BC S. relictus (Horn) - BC (Subgenus STENOCANTHARUS Gistel) (Pemphus Motschulsky) Key to species: Gidaspow (1973) S. angusticollis (Fischer AK BC von Waldheim) longipes Casey nigripennis Roeschke opacus Casey uelutinus M6n6tries (Subgenus NEOCYCHRUS Roeschke) Key to species: Gidaspow (1973) S. angulatus (Harris) - BC rnaritimus Van Dyke (Subgenus BRENNUS Motschulsky) Key to species: Gidaspow (1968) S. johnsoni Van Dyke - BC hlahowyae Perrault S. marginatus (Fischer AK BC AB von Waldheim) columbianus (Casey) confusus (Casey) cupripennis (Casey) /ollor (Roeschke) fulleri (Horn) gracilis (G6hin) insularis (Casey) montanicus (Casey) oregonus (Casey) wrangelli (Casey) Genus CYCHRUS Fabricius Key to North American species: Gidaspow (1973) C. rickseckeri LeConte - BC C. tuberculatus Harris - BC pustulosus Casey Tlibe Nebriini Genus LEISTUS Friilich Key to North American species: Erw.in (1970a) (Subgenus LEISTUS s. str.) L. fernrgineus (Linn6)f - NF (Subgenus NEOLEISTUS Erwin) L. ferruginosus Mannerheim AK BC AB ferrugineus Dejean nigropiceus Casey Genus PELOPHILA Dejean P. borealis (Paykull)* AK YK NT BC AB SK MB ON PQ - LB NF californica Motschulsky e s c hs c holtzi (Mannerheim) shermani Casey ulkei Horn P. rudis (LeConte) NT BC AB - MB ON - NF 13 Carabidae Genus NEBRIA Latreille N. acuta acuta Lindroth AK BC N. arkansana edwardsi Kavanaugh BC N. brevicollis (Fabricius)t _v v_ PQ N. campbelli Kavanaugh BC N. charlottae Lindroth BC N. crassicorn is crassicornis BC Van fryke N. crassicornis intermedia Van Dvke BC AB N. diversa LeConte BC liuidaLeConte townsendi Casey N. frigida R.I'. Sahlberg* BC paruula J. Sahlberg reducta'Casey uiridis E:lorn N. gebleri cai;cadensis Kavanaugh _BC N. gebleri gebleri Dejean AK BC AB melanariaHatch N. gouleti Kavanaugh ? N. gregaria Fischer von Waldheim AK cuneata casey N. gyllenhali castanipes (Kirby)* AK YK NT BC AB SK MB ON PQ LB NF curtulata Casey (?) elias Motschulsky labradorica Casey moesta leCorrte prominens Casey N. haida Karanaugh _BC N. hudsonica LeConte _YKNTBCABSK yu_ N. kincaidi k ncaidi Schwarz AK BC columbi'ma Casey N. lacustris I lcustris Casev MB ON PQ NB expansa Casey N. lituyae Kavanaugh AK N. Iouiseae Kavanaugh -BC N. mannerheimii Fischer AK BC von S'aldheim coruallis Casey hippisleyi Casey oregona Casey N. meanyi meanyi Van Dyke AKYK_BC N. meanyi sylvatica Kavanaugh -BC N. metallica Fischer von Waldheim AK BC pacifica Chaudoir N. nivalis gar;pesiana Kavanaugh _PQ LB N. nivalis nir alis (Paykull)* AK YK I.TT _ _PQ v bifaria llIannerheim mollis l!:lotschulsky N. obliqua obliqua LeOonte AKYK_BC SK incerta Casey obtusa I.,eConte testaceipes Casey texana Casey N. pallipes Say PQ NB NS PE N. piperi Van Dyke AK--eC 9* N. sahlbergii sahlbergii Fischer AKYK-BCAB von Waldheim aleuta\anDyke uiolacea Motschulsky N. schwarzi schwarzi Van Dvke AB N. suturalis l.eConte :" ON PQ LB longula LeConte nimbosa Casey N. virescens .florn BC breuis Casey T4 Carabidae Tlibe Notiophilini Genus NOTIOPHILUS Dum6ril N. aeneus (Herbst) oN PQ NB NS porrectus Say N. aquaticus (Linn6)x BC SK MB ON PQ NB NS LB NF hardyi Putzeys N. biguttatus (Fabricius)t BC NB NS NF N. borealis Harris* AK YK NT BC AB MB ON PQ _LB NF N. directus Casey BC AB lanei lJatc]l N. intermedius Lindroth AK BC MB PQ LB NF N. nitens LeConte BC N. semistriatus Say AK YK- BC AB MB ON PQ NS LB americanus }{atris coloradensis Casey confusus LeConte punctatus LeConte N. simulator FalI BC AB euanescens Casey N. sylvaticus Eschscholtz AK BC Tribe Loricerini Genus LORICERA Latreille Key to species: Ball and Erwin (1969) (Subgenus LORICERA s. str.) L. decempunctata Eschscholtz AK _BC L. pilicornis congesta Mannerheimx AK L. pilicornis pilicornis AK YK NT BC AB SK MB ON PQ NB NS is NT (Fabricius)* californica LeConte neoscotica LeConte semip unct ata Eschscholtz sierraeYanDyke uteana Casey Tribe Opisthiini Genus OPISTHIUS Kirby O. richardsoni Kirby AK YK NT BC SKAB TYibe Elaphrini Genus DIACHEILA Motschulsky Arctobia C.G. Thomson D. arctica amoena (Faldermann)* LB americana Motschulskv subpolaris LeOonte D. polita (Faldermann)* NTYKAK Genus BLETHISA Bonelli HelobiumLeach Rhaphiona Fischer von Waldheim Key to species: Goulet and Smetana ( 1983) B. catenaria W.J. Brown* AKYKNT_MB B. inexspectata Goulet and Smetana AK B. julii LeConte NT_ABSKMBONPQNBNSNF B. multipunctata aurata AK YK NT BC AB SK MB ON PQ NB NS PE - NF Fischer von Waldheimx hudsonica Casey B. oregonenis LeConte BC acutangula Chaudoir columbica Casey B. quadricollis Haldeman NT BC AB SK MB ON PQ NB NS PE LB NF arnericana Hatris 15 Carabidae Genus ELAI'HRUS Fabricius Key to species: Goulet (1983) (Subgenus ARCTELAPHRUS Semenov) E. lapponicus lapponicus Gyllenhal* AK YK NT BC AB - MB ON PQ - LB obscurit.tr I{trby E. lapponicus obliteratus AK ManrLerheim (Subgenus NEOELAPHRUS Hatch) E. cicatricosrrs LeConte PQ rhodear; us Casey E. clairuillei KJrby AK YK NT BC AB SK MB ON PQ NB NS PE LB NF /rosll H.ppisley lynni Pierce politus ,eCorrte torreyer,rsis Tanner E. fuliginosus Say AB - MB ON PQ E. olivaceus LeOonte NT BC AB SK MB ON PQ NB NS PE - NF (Subgenus ELAPHRUS s. str.) (.Trichel tphrus Semenov) E. americanrrs americanus Dejean AK YK NT BC AB SK MB ON PQ NB NS PE LB NF bitubercsus Casey gratiost^s Mannerheim interme lizs Kirby punctat ssjmzs LeConte sinuatu i LeConte E. americanrrs syivanus Goulet BC AB E. californicrLs Mannerheim AK - NT BC AB SK MB ON PQ NB NS PE hesperh'.s Casey similis LeCorrte E. lecontei Crotch - NT BC AB SK MB ON deuinctt:s Casey spisslco''nls Casey E. parviceps Van Dyke* AK - NT - MB ON E. ruscarius Say ON PQ texanus Casey E. tubercular us Maklin* AK YK NT (Subgenus E LAPHROTERUS Semenov) (Elaphriatus Semenov) E. angustico lis angusticollis AX YK NT R.F. liahlberg* E. purpuranr' Hausen AK YK NT BC AB pallipes Hotr' Tribe Omophronini Genus OMO.?HRON Latreille Key to North American species: Benschoter and Cook (1956) O. americanus Dejean AB SK MB ON PQ NB NS PE - NF fontinal;.s Casey illustris Casey iridescens Casey locustri'; Casey lengi Casey proximus Chandler soil Kirby tanneri Chandler texanus Casey O. Iabiatus (Fabricius) NS O. ovalis Horn - BC AB SK concinnits Casey frater Casey gemma'Jasey O. robustus llorn AB ON breuipen.nis Casey decoloratus Fall 16 Carabidae O. tesselatus Say AB SK MB ON PQ NB NS PE ellipticus Casey lecontei Dejean Tribe Promecognathini Genus PROMECOGNATHUS Chaudoir P. crassus LeOonte BC contractus Casey corpulentus Casey grandiceps Casey Tribe Scaritini Genus SCARITES Fabricius S. subterraneus Fabricius ON beckwithii Stephens denticollis Chaudoir interruptus (Herbst) spinipes (Sulzer) subtemeus Bonelli Genus PASIMACHUS Bonelli P. elongatus LeConte AB SK MB angustulus Casey euanescens Casey pirnalis Casey uegasensis Casey uernicatus Casey Tribe Clivinini Genus DYSCHIRIUS Bonelli Key to North American species: Bousquet (1g88) (Subgenus AKEPHORUS LeConte) D. obesus LeConte - BC (Subgenus DYSCHIRIUS s. str.) D. affinis Fall ON PQ duplicatus Fall D. alticola Lindroth -BC D. aratus LeConte _BCAB MB D. brevispinus LeConte -ONPQ D. campicola Lindroth AB MB D. compactus Lindroth D. criddlei FaIl _:rf MB D. er5rthrocerus LeConte -oNPQ D. foigidus Mannerheim AK YK NT BC _ONPQ LB aeneolus LeConte D. globulosus (Say) AK YK -BCAB SK MB ON PQ NB NS NF paruus LeConte D. haemorrhoidalis (Dejean) -oN D. hiemalis Bousquet AK NT BC AB MB ON PQ _LB D. integer LeOonte AK NT BC AB SK MB ON PQ NB NS PE NF apicalis LeOonte basalis LeConte nigripes LeConte tr an s m arin u s Mannerheim D. interior Fall _AB SK MB arizonicus Van Dyke D. laevifasciatus Horn BC AB D. larochellei Bousquet NB NS _NF D. longulus LeConte BC AB SK MB_PQ D. montanus LeConte BC AB MB ON PQ _v thompsoni Hatcll L7 Carabidae D. nigricornis Motschulsky* AK YK NT secretus Fall D. pacificus .l,indroth _BC D. pallipenn:s (Say) AB SK ON PQ D. perversus Fall AB MB desertut Fall D. pilosus LrConte MB ON PQ hispidu,; LeConte D. planatus ,indroth AB SK MB D. politus (Dejean)* AK YK _BCABSKMB ON PQ NF aureolu.; Notman D. quadrima :ulatus Lindroth AB SK MB D. sellatus LeConte PQ NB NS PE NF D. setosus LcConte AK nc AB SK MB ON PQ NB NS PE alternatus Hatch D. sextonj Bousquet ON po D. sphaerico lis tSayr BC AB sr me ON PQ NB NS NF subpunt:tatus Hatch D. subarcticus Lindroth AK YK NT D. tenuispin rs Lindroth AB D. timidus Lindroth -me D. truncatus LeConte AK Nr sc AB SK MB Genus CLIVINA Latreille (Subgenus S 0MICLN'INA Kult) C. dentipes.l)ejean ON coruina Putzeys confusa LeConte fissipes Putzeys georgiana LeConte (Subgenus PARACLIVINA Kult) C. bipustula' a (Fabrici us) ON quadrintaculafo (Palisot de Beauvois) (Subgenus CLIVINA s. str.) C. americana Dejean ON PQ NS acuducta Haldeman cordata Putzeys ludouiciana Putzeys C. collaris tIlerbst)t BC MB ON PQ elongatt Randall randalli LeConte C. fossor il,inn6)t _BCAB ON PQ NB NS PE NF C. impressifrons LeConte C. oregona Fal) -BC _-ONPQ Genus SCHIZOGENIUS Putzeys Key to North American species: Whitehead (1972) (Subgenus SCHIZOGENIUS s. str.) S. amphibius (Haldeman) PQ frontalis (LeConte) S. depressus LeConte BC S. ferrugineus Putzeys sx ON pe sulcatu;; (LeConte) S. lineolatus {Sayr ON PQ S. litigiosus Fall ec S. sulcifrons Putzeys ON PQ NB TYibe Brachininr Genus BRA(IHINUS Weber Key to North American species: Erwin (19706) (Subgenus NEOBRACHINUS Erwin) B. cordicollis Dejean - ON PQ NB gracilis Blatchley leptocerus Chaudoir uelox LeConte 18 Carabidae B. cyanipennis Say MB ON PQ NB cephalotes Dejean rejectus LeConte B. cyanochroaticus Erwin - BC - SK MB ON PQ B. fumans (Fabriciust - BC AB - MB ON PQ affinis LeConte amplipennis Bates atbarae Stehr cyanopterus LeConte suffians LeConte tabasconus Bates B. janthinipennis (Dejean t - ON PQ pumilio LeConte B. medius Harris - BC - ON PQ minutus }J.arris B. ovipennis LeConte - ON PQ B. quadripennis Dejean - BC AB SK MB ON stygicornis Say I schern ihhii Mannerheim B. tenuicollis LeConte - ON PQ ballistarius LeConte similis LeConte B. texanus Chaudoir AB Tribe Paussini Genus METRIUS Eschscholtz M. contractus contractus - BC Eschscholtz Tribe Gehringiini Genus GEHRINGIA Darlinston G. olympica Darlington NT BC AB Tribe Broscini Genus BROSCUS Panzer B. cephalotes (Linn6tf NS PE Genus MISCODERA Eschscholtz Leiochiton Curtis M. arctica (Paykull)x AK YK NT BC AB SK MB ON PQ - NS PE LB NF americana Mannerheim hardyi Chaudoir Genus BROSCODERA Lindroth B. insignis tMannerheim) AK BC Genus ZACOTUS LeConte Z. matthewsii LeConte AK BC angustus Casey fre deric ki Nunenmacher subopacus Hopping Tlibe Trechini Genus BLEMUS Dejean Las iotrec hu s Ganglbauer B. discus (Fabriciustf - ON PQ NB Genus TRECHUS Clairville (Subgenus TRECHUS s. str.) 19 Carabidae T. apicalis Motschulskyx AK YK NT BC AB SK MB ON PQ NB NS LB NF borealis Schaeffer brumalts Casey canader'.sis Putzeys fuluus (",eCorte) micans .LeConte) pallescens Casey puritan,ts Casey rhodensis Casey T. chalybeus Dejean* AK BC AB brachycierus Jeannel califor n ic us Motschulsky tahoensi.s Casey T. crassiscapus Lindroth NS T. obtusus Erichsont ---ec-- _:uIu T. oregonensis Hatch -BCAB _ :.T _ T. ovipennis Motschulsky y_ BC_ T. quadristriatus (Schrank)f _ _ _ONPQ T. rubens (Fabricius)t PQN eNs pe- y T. tenuiscapr.rs Lindroth _YK_BCAB Tribe Bembidiini Genus ASAI HIDION des Gozis Tachypi:.ts Dejean A. alaskanurn Wickham AK YK NT A. yukonens'r Wickham AKYK_BCAB Genus BEM.IIIDION Latreille (Subgenus CHRYSOBRACTEON Netolitzky) (Parabrzcteon Notman) B. alaskense Lindroth AK B. balli Lindroth ---ABix B. carinula ( jhaudoir YK NT BC AB SK ff,fS On PQ NB NF tubercu o/um Notman B. co1villenstr Lindroth .AK B. foveum Motschulsky* AKYKNTBC_SK MB bryanti Carr (.?) glabr iusculum Motschulsky grahamiHatch B. hesperium Fall BC binariu;n Casey B. inaequale inaequale Say MB ON PQ NB NS PE arenari,r,m Dejean lacustre LeConte B. inaequale opaciceps Casey AK BC AB SK B. lapponicum Zetterstedt* AK_NT BC AB SK pugetanumFall B. levettei Casey AK YK NT BC SK MB oN PQ NB NS LB NF carrianlm Casey B. lorquinii (lhaudoir BC tacomar Clssy B. punctatostriatum Say BC SKAB oN PQ NB (?) sigiliare Say stigmat:icum Dejean B. zephyrurr Fall BC marginosum Casey tristicuium Casey (Subgenus BRACTEON Bedel) B. bowditchii LeConte BC (Subgenus ODONTIUM LeConte) B. confusum Hay'ward AB SK MB ON PQ aeneorubrum Casey marquettense Casey nitidult:m Dejean 20 Carabidae B. coxendix Say NT_AB SK MB snowi Casey unicum Casey uenator Casey uigilans Casey ueesi Hatch & Ortenburger (Subgenus PSEUDOPERYPHUS Hatch) (B r a cteomimus Lindroth) B. chalceum Dejean BC AB SK MB ON PQ NB NS NF antiquum Dejean dilatatum (LeConte) B. honestum Say ON PQ NB basale (LeConte) platyderum Chaudoir (Subgenus OCHTHEDROMUS LeConte) B. americanum Dejean -oN PQ illini Casey B. bifossulatum (LeOonte) BC AB SK MB ON PQ cheyennense Casey ferreum Casey nuperum Casey regestum Casey suffiatum Casey (Subgenus EURYTRACHELUS Motschulsky) (Eudromus l\trby) (P la ty t r a c hel us Motschulsky) B. interventor Lindroth AK YK NT BC AB SK MB ON PQ B. nitidum (Kirby) AK YK NT BC AB SK MB ON PQ edolatum Casey (Subgenus HYDRIUM LeConte) B. levigatum Say AB delawarense Casey (Subgenus METAILINA Motschulsky) B. dyschirinum LeOonte AK YK BC AB agitabile Casey aleneanum Casey atrolucens Casey heechelum }Iatch perturbatum Casey speculinum Casey B. lampros tHerbsilf BC NF B. properans ( Stephens tt PQ NB NS PE NF (Subgenus PI{YLA Motschulsky) B. obtusum Audinet-Servillet oN PQ (Subgenus LIONEPFIA Casey) Key to species: Erwin and Kavanaugh (1981) B. castum Casey AK BC brumale Casey nescium Casey serenum Casey uaciuum Casey B. disjunctum Lindroth BC B. erasum LeConte BC AB lasciuum Casey lubricum Casey probatum Casey B. lindrothellum Erwin & Kavanaugh AK BC B. lummi Erwin & Kavanaugh BC B. sequoiae Lindroth BC (Subgenus ACTEDIUM Motschulsky) B. lachnophoroides Darlington AB SK (Subgenus TRECHONEPHA Casey) B. iridescens I LeConte) BC AB 2I Carabidae atnicum Casey deceptor Casey fabrum Oasey impium Casey obliuiost:.m Casey p ar allel c'colle (Motschulsky) repens Casey uolatile t)asey JR B. simplex Hayrvard PQ essexense Casey $ B. kuprianov:.i Mannerheim AK YK NT BC AB adultum Casey b iimp r es s u m Mannerheim bucolicum Casey dilutum Casey funereurz LeConte maehlint Hayward ouipennt (Motschulsky) (Subgenus PI,ATAPHODES Ganglbauer) B. arcticum Lindroth* AK YK B. brachytho:'ax Lindrothx AK YK NT BC B. complanul om (Mannerheim) AK BC AB paruulutn Notman B. compressum Lindroth* AK YK NT B. farrarae Ilatch AK BC B. haruspex l.)asey _BC B. incertum ( Motschulsky) AKYK?BC AB ampliatum Casey blanditum Casey (?') breue (Motschulsky) improuit,um Casey liuidulu n Casey saturatum Casey testatun Casey tetr ag ly 1 t u m (Mannerheim) B. manninge:rse Lindroth AK YK NT BC AB ochropu:; Lindroth B. occultator Notman AK _AB SK PQ NB NS LB NF B. quadrifoveolatum Mannerheim AK YK BC AB 9* iller Casey B. rosslandicrm Lindroth BC AB B. viator Casey BC (Subgenus P.I,ATAPHUS Motschulsky) (.Melomt'lus Casey) (M icromelomalus Casey) (Trachelonepho Casey) B. basicorne Notman _PQ_NS B. carolinense Casey _PQNBNS NF heeneanzm Casey B. curtuiaturn Casey AK YK BC AB decrepitu.m Casey effetum Casey flebile Casey timefactum Casey B. gebleri turbatum Casey YK BC AB conflictt:m Casey B. gordoni Lindroth BC B. gratiosum Casey AK YK BC AB pallidumHatch B. hyperboraeorum Munster* AK YK NT B. kalumae Lindroth BC B. planatum (LeConte) AK YK NT BC AB--oNpeNnNs--Nr adjutor Casey aequalis: (Walker) solutum Casey 22 Carabidae B. planiusculum Mannerheim AK BC B. rufrnum Lindroth AK YK - BC .AB B. rusticum lenensoides Lindroth AK YK - BC B. nrsticum rusticum Casey - ON PQ NB NS - LB NF notmani Casey B. siemicola Casey - BC B. stillaguamish Hatch - BC B. sulcipenne hyperboroides AK YK - BC Lindroth B. sulcipenne prasinoides Lindroth - BC .AB - MB - PQ NB LB NF (Subgenus BLEPHAROPLATAPHUS Netolitzky) B. hastii C.R. Sahlbergx AK YK NT BC MB ON PQ - LB (Subgenus TRICHOPLATAPHUS Netolitzky) B. planum (Haldeman) - ON PQ NB NS filicorne Casey guexii Chatdo\r uulsum Casey B. rolandi Fall - ON PQ NB NS (Subgenus HIRMOPLATAPHUS Netolitzky) B. concolor (Kirby) AK YK NT BC .AB SK MB ON PQ NB NS NF longulum (LeConte) subaeneum (LeConte) B. nigrum Say - ON PQ NB NS PE facile Casey morosum Casey B. quadrulum LeConte - YK - BC .AB aegrotum Casey callidum Casey porrectum Casey tartareurn Casey tritum Casey uiridinigrum Casey B. recticolle LeConte - BC AB oblatum Casey pertinax Casey tetr ag onoderum Chaudoir umbraticola Casey B. salebratum (LeOonte) NT BC .AB SK MB ON PQ NB NS NF consessor Casey inopinum Casey mackinacense }latc}r purpurascens (LeConte) (Subgenus HYDRIOMICRUS Casey) B. quadratulum Notman PQ NB NS NF proximum Notman B. semistriatum (Haldeman) - ON PQ - NS (Subgenus BEMBIDIONETOLITZKYA Strand) (Daniela Netolitzky) B. mckinleyi carneum Lindroth - BC AB B. mckinleyi mckinleyi FalI AK YK $ B. lenae Csiki* AK YK NT intermedium Poppius $ B. commotum Casey _BCAB seclusum Casey B. nebraskense LeConte _BCAB denueranum Casey gouanicum Casey tractabile Casey $ B. actuosum Casey - BC AB debilicolle Casey urgens Casey 23 Carabidae B. nevadense Illke BC uiaticun.: Casey B. plagiatum (Zimmermann) ON B. striola (LeConte) BC angustit:r Casey modulatum Casey shastani.cum Casey uancouu zri Casey (Subgenus PiIRYPHANES Jeannel) B. dauricum Motschulskyx AK YK NT MB B. grapii Gyl enhal* AK YK NT BC AB SK MB ON PQ NB NS is NF nitens Casey picipes (t{irby) scrutatum Casey seducturn Casey B. lacunariurn (Zimmerrnann) ON PQ histricurn Casey militare Casey B. platynoides Hayward _BC insopita:zs Casey merens li)asey optatum Casey sedulutr, Casey B. stephensi lrotch'l -oN PQ NB PENS NF canadense Hayward B. texanum ( lhaudoir) MB cogitans Casey inquietum Casey B. yukonum lall* AK YK NT BC MB PQ $ B. bimaculat rm (Kirby) AK YK NT BC AB SK MB ON B. postremur:r Say -PQ NB B. sordidum rKirby) AI< YK NT BC AB SK MB oN PQ LB B. umiatenser Lindroth AK (Subgenus PIIRYPHUS Stephens) B. bruxellense Wesmaelt -PQNBNSPE NF B. femoratunr Sturm* NB NS B. obscurellu m (Motschulsky)x AK YK NT BC AB sx lle ONPQNB_PE NF albidipezne Casey caducunt Casey fuscicrus (Motschulsky) lucidum (Mannerheim) mixtum LeConte parowar.um Casey petulans Casey (?) subinflatum (Motschulsky) B. petrosum ilttuense Lindroth AK B. petrosum'letrosum Gebler* AK YK Nr ec AB SX ON PQ Nn lls pr LB NF castaliu;n Casey exiguiceps Casey lepusculum Casey lucidum (LeConte) substric,um (LeConte) wenatchee Hatch B. poppii sch,rlleri Lindroth AK B. rupicola (lGrby) AK NT BC AB sr l.rn oN- auersan:; Casey B. sejunctum sejunctum Casey NT AB SK LB NF B. sejuncturn semiaureum Fall _BC _:a YNS fenderi llatch B. tetracolum Sayt _BC oN PQ NB NS NF nactum Casey $ B. transverszrle Dejean YK_BC SK MB ON PQ NB NS LB NF acoman,tm casey animatu.m Casey 24 Carabidae (?) erosum (Motschulsky) ercursum Casey haplogonum Chaudoir ma nner heimii (LeConte) marinicum Casey perspicuum (LeConte) sarpedon Casey tuolumnae Casey $ B. scopulinum (Kirby)x AK YK NT BC AB SK MB ON PQ NB PENS NFLB bellulum Casey gelidum (LeConte) (Subgenus LIOCOSMIUS Casey) B. mundum (LeConte) BC b ifa s c iat um (Motschulsky) hilare Casey (Subgenus PERYPHODES Casey) B. ephippigerum (LeConte) _BC B. salinarium Casey _BCAB SK MB $ B. constricticolle Hayward _AB B. nudipenne Lindroth _AB SK MB (Subgenus EUPETEDROMUS Netolitzky) B. graciliforme Hayward -oNPQ B. immaturum Lindroth _ONPQNBNSNF B. incrematum LeConte AK YK Nt EC AB sr MB ON PQ NB NS PE LB NF arcuatum LeConte fortunatum Casey gulosum Casey /ezgi Notman mobile Casey nubiferum Casey oblectans Casey sernotum Casey (Subgenus LEUCIIYDRIUM Casey) B. tigrinum LeConte -BC $ B. consimile Hayward ,AB SK MB B. flohri Bates BC .AB SK MB henshawi Hayward B. obtusidens Fall ,AB SK MB B. scudderi LeConte --nc ,AB SK MB spissicorne Casey $ B. dejectum Casey ,AB SK MB aberti}].atch fidele Casey tencenti Hatch B. mormon Hayward _BC B. obtusangulum LeConte _ BC ,{B SK MB cornix Casey (Subgenus NOTAPHUS Stephens) B. aeneicolle (LeConte) NT BC .q,B SK MB B. castor Lindroth _ ,A,B SK MB ON PQ NB NS NF B. coloradense Hayward NT BC .{B SK MB albertanum Casey amplipenne Casey imperitum Casey prociduum Casey B. cordatum (LeConte) MB ON placabile Casey B. debiliceps Casey BC 25 Carabidae B. dorsale Say _BCAB SK MB B, graphicum Casey NT BC AB SK MB ON lassulutn. Casey B. indistinctum Dejean AK BC barbarae Casey caudex (;asey conse nt a neum (LeConte) curiosun,, Casey derisor C.iasey deuinctum Casey erpositurn Casey extricatum Casey formale l}asey franciscc:,num Casey ornatellt'.m Casey reconditum Casey B. insulatum (LeConte) NT BC AB SK MB caliginosum Casey B. intermeditLm (Kirby) AK BC AB SK MB ON PQ B. nigripes (IrJrby)x AK YK NT BC AB SK MB ON PQNB_PELB NF concitatt.Lm Casey quadrah colle (Mannerheim) retracturn Casey B. patruele Dejean NT BC AB SK MB ON PQ NB NS PE NF contristans Casey editum Oasey exclusunt Casey fenisex (.asey fraternun LeOonte imitator Casey marcidum Casey mediocn: Casey monstra,'.um Casey plectile Oasey pos ticun,'. (Haldeman) subexiguum Casey B. rapidum (LeConte) SK MB ON PQ NB docile Ctsey fugitans Casey negligeni Casey sociale ( asey B. semipunct etum (Donovan)* AK YK BC SKAB ON accuratt'.m Casey prouoanitm Casey B. umbratum (LeConte) NT BC AB SK u ar iolos u m ( Motschulsky) B. variegaturn Say ON PQ postfascntum Hamilton B. versutum LeConte oN PQ NB $ B. constricturn (LeConte) NB NS PE ciuile Ca.sey festinans Casey uernula Casey B. contracturr Say _PQ NB NS PE NF B. viridicolle (LaFert6-S6nectdre) NT AB SK MB ON hamiferum Chaudoir particeps Casey R B. oberthueri Hayward AB SK MB ON PQ NB (Subgenus FIJRCACAMPA Netolitzky) B. affine Say ON decipiens Dejean fallax Dejearr thespis Casey JR B. impotens Casey BC SK MB ON NB 26 Carabidae aestiuum Casey frugale Casey gratuitum Casey imbelle Casey ind,igens Casey pictum (LeConte) pullulum Casey uirgatulum Casey B. mimus Hay'ward - SK MB ON PQ NB NS PE - NF pellax Casey rotundiceps Casey B. timidum (LeConte) NT BC AB SK MB ON oregonum Casey sord.idulum Chaudoir B. versicolor (LeConte) AK - NT BC AB SK MB ON PQ NB NS PE LB NF terracense Casey tolerans Casey wisconsinium Casey (Subgenus EMPHANES Motschulsky) (Omala Motschulsky) B. diligens Casey - BC AB SK MB parabile Casey uilescans Casey B. vile (LeConte.) - BC (Subgenus BEMBIDION s. str.) (Lopfto Stephenst B. mutatum Gemminger & Harold AK YK NT BC AB SK MB ON PQ - NS NF arillare (LeConte) uegetum Casey B. praecinctum LeConte AB SK MB alutaceum Hatch ueridicum Casey B. quadrimaculatum dubitans AK YK NT BC AB SK MB (LeConte) caseyi Letg crurale (LeConte) gregale Casey pugetanurn Casey sapphicum Casey tenax Casey B. quadrimaculatum oppositum Say AB SK MB ON PQ NB NS PE - NF (Subgenus SEMICAMPA Netolitzky) B. convexulum Hayrvard AK YK - BC AB nouellum Casey B. morulum LeConte AK YK NT BC AB SK MB NF browni Lindroth B. muscicola Hayward - SK MB ON PQ NB NS PE B. praticola Lindroth - BC - SK - ON PQ NB B. roosevelti Pic AB SK MB blaisdelli Casey concinnum Blaisdell p er conc innunt Blaisdell B. semicinctum Notman - ON PQ NB NS (Subgenus DIPLOCAMPA Bedel) (Paralopha Casey) B. transparens (Gebler)* AK YK NT BC AB SK MB ON pQ NB NS pE LB NF edmontonense Casey sulcatum (LeConte) trepidum (LeConte) (Subgenus TREPANEDORIS Netolitzky) B. acutifrons LeConte AK - NT BC AB SK MB m i c r o r e t ic u la t u m Hatch B. canadianum Casey NT - AB SK MB ON PQ B. concretum Casey AK YK NT BC AB SK MB ON PQ NB NS PE - NF congruens Casey habile Casey 27 Carabidae B. connivens (LeConte) BC digressum Casey elizabetl:ae Hatch B. fortestriatrm (Motschulsky) AK YK NT BC SK MB ON PQ NB NS PE NF cdutum .LeConte) concurre'.ns Fall peregrinum Casey utnbrati:um Casey B. frontale (l.eOonte) _BC_SKMBONPQ NS PE B. pseudocaubum Lindroth _BCABSKMBONPQ )u B. siticum Cz.sey _BC adolescew Casey (Subgenus AIvIERIZUS Chaudoir) B. oblonguloiJes Lindroth _BC B. oblongulurn (Mannerheim) AK BC crassicorne (Casey) heeni (C*ey') B. spectabile (Mannerheim) AK BC longicor.rc (Casey) B. wingatei Filand oN PQ NB NS LB NF Genus PHR\PEUS Casey P. ricksecker (Hayward) BC rutilinut Casey Genus ELAP HROPUS Motschulsky Barytac,rys Chaudoir Tachyur a Motschulsky E. anceps (Lt.Conte) NT AB SK MB ON PQ E. capax (Le0onte) -oN YNS E. dolosus (LeConte) -oN apacheanus { Casey) audax (l,eConte) rapar (I eConte) E. granarius (Dejean) -oN PQ _NS glossem'r (Casey) E. incurwus ( Jay) BC SK MB ON PQ NB NS PE parallel 1s (Casey) rhodens'.s (Cslki) E. parvulus (Dejean)t BC E. saturatus (Casey) ---oNFeNeNs-- E. tripunctatus (Say) _ONPQNBNS barnesi (Stehr) laredoanus (Casey) seruzs ({lasey) E. vernicatus, (Casey) SK MB ON PQ NB laetificus (Casey) unionis (Csiki) E. vivax (LeConte) oN PQ mendar (LeConte) E. xanthopur (Dejean) ON NS ancilla rCasey) famelia.Ls (Casey) loaipennis (Casey) Genus TACFiYTA Kirby Key to species: Erwin (1975) T. angulata Casey YK BC AB SK MB ON PQ NS T. falli (Hayrvard) BC fu T. nana inornata (Say) BC ON PQ arizonica Casey californ;;,ca Casey collaris Casey debilicol,lis Casey picipes lktby riu ulari s (Motschulsky) T. nana kirb ri Casey BC AB ON PQ 28 Carabidae Genus POROTACHYS Netolitzky P. bisulcatus tNicolaitt - ON PQ NB frontalis (Hayward) Genus POLYDERIS Motschulsky Microtachys Casey P. laevis {Sayt - ON PQ - NS congesta (Casey) flumenalis (Casey) recta (Casey) troglodytes (Dejean) unistriata (Casey) unistriolata (Cslki) Genus TACHYS Stephens Isotachys Casey T. bryanti Lindroth ,\B T. halophilus Lindroth ,\B SK MB T. vittiger LeConte - BC marginellus LeConte Genus PARATACHYS Casey P. edax (LeConte) - BC P. oblitus {Casey) - ON PQ appalachius (Casey) cuneatus (Casey) gentilis (Casey) iowensis (Casey) obliquus (Casey) P. proximus tSayt - ON PQ nubifer {Casey) P. rhodeanus (Caseyr - ON PQ P. scitulus (LeConte) - ON PQ NB pallescens (Casey) p a I I i d i u s c u I u s (C sikl) Genus MIOPTACIfYS Bates T ac hy menis Motschulsky M. flavicauda (Say) - BC r\B ON PQ ma rg inicollis (Motschulsky) r eflexicoll is (Motschulsky) Tlibe Nomriini Genus NOMIUS Castelnau Haplochile LeConte N. pygmaeus (Dejeant - BC AB - MB ON PQ Genus PSYDRUS LeConte Monillopatrobzs Hatch P. piceus LeConte - BC AB SK MB ON PQ punctdtus (Hatch) Tribe Patrobini Genus PLATYPATROBUS Darlington P. lacustris Darlington NT - l\B SK MB ON PQ NB Genus PATROBUS Dejean Geo p a t r ob u s Darlington N eo pat r ob u s Darlington P. lossifrons (Eschscholtz)x AK BC l\B dimorp hic us Darlington /uluus Mannerheim latiusculus Chaudoir long i uen I ri s Mannerheim 29 Carabidae P. foveocollis (Eschscholtz)* AK YK NT BC AB SK MB ON PQ - LB NF ang usticollis Mannerheim insularis Casey Iaeuiceps Casey obt usiu sc u I us Chaudoir tenuis (l'eConte) NF P. lecontei Chaudoir - BC AB SK MB - PQ NB - canadensis Casey grauidus Darlington ruftpes LeConte p. tongicbrnis (Say) NT BC AB SK MB ON PQ NB NS PE - NF americanus Dejean P. septentrionis Dejlan* AK YK NT BC AB SK MB ON PQ - LB NF hyperboreus Dejean Iabradorinus Casey minuen.; Casey tritus Casey P. stygicus Chaudoir AK YK NT BC AB SK MB ON PQ - LB NF Genus DIPLOUS Motschulskv Platidiu.s Chaudoir D. aterrimus, (Dejean) AK YK NT BC AB breuiceps (Casey) coloradensis (Casey) fulcratus (LeConte) reflexus (Casey) tenuitarsis (Casey) D. californic rs (Motschulsky) - BC breuius;ylas (Casey) lnciszs (Casey) Iatipenris (Casey) rectus (Jasey) sietan es (Casey) strenuus (Casey) troc han te ricus (LeConte) D. frlicornis iCasey) - BC D. rugicollis (Randall) PQ NB NS longipalpis (Notman) Genus PLAI]IDIOLUS Chaudoir Patroboidea Van Dyke P. vandykei Kurnakow AK YK - BC AB rufus (Van Dyke) Tlibe Pterostichini Genus LOK\NDRUS LeConte Key to Ncrth American species: Allen (1972) L. minor (Chaudoir) - ON inquietrc Casey L. velocipes Casey - ON Genus STOMIS Clairville S. pumicatus (Panzer)t NS Genus LOP}{OGLOSSUS LeConte L. scrutator (LeConte) - ON PQ canadensis (Chaudoir) Genus POECILUS Bonelli Feronio Latreille Leconteus Lutshnik (Subgenus POECILUS s. str.) P. chalcites (Say) - ON PQ micans Chaudoir soyi (BruII6) 30 Carabidae P. corvus (LeContet AB SK MB P. lucublandus (Say) NT BC AB SK MB ON pe NB NS pE castanipes Kirby conue*icollis (Say) dilatatus LeConte elucens Casey fraternus Say manhattanis (Casey) planifer Casey P. scitulus LeConte - BC AB SK MB (Subgenus DERUS Motschulsky) (D er ulus Tschitsch6rine) P. nearcticus (Lindroth.t* NT Genus STEREOCERUS Kirby Boreobi a Tschitscherine S. haematopus (Dejean)* AK YK NT BC AB SK MB ON pe LB NF similis I\trby - S. rubripes (Motschulsky)* AK hry z hanou s kii (Lindroth) Genus MYAS Dejean Key to North American species: Allen (1980) M. coracinus (Say) - ON M. cyanescens Dejean - ON pe NB foueatus LeConte Genus PTEROSTICHUS Bonelli (Subgenus CYLINDROCHARIS Casey) Key to species: Barr (1971) P. rostratus (Newman) - ON pe NB grandiceps (LeConte) piceatus (Casey) sulcatulus (Casey) (Subgenus HOLCIOPHORUS LeConte) P. lama (M6netri6st - BC o/er (Dejean) at e r r imu s (Motschulsky) cephalus (Casey) domitor (Casey) pollens (Casey) uancouueri (Casey) (Subgenus FIYPHERPES Chaudoir) (B r ac hy s tilu s Chaudoir) (Gonoderus Motschulsky) (H ap locoelus Chaudoir) P. adoxus(Say) - ON pe NB NS rejectus LeConte sufflatus (Casey) P. a)gidus LeConLe AK BC anthrat (Casey) innatus (Casey) responsor (Casey) ualidus (Dejean) P. amethystinus Mannerheim AK BC jejunus Casey metlakatlae Casey nouellus Casey stoicus (Casey) P. castaneus (Dejean) AK BC brunneus (Dejean) terracensis (Casey) wrangelli Casey P. crenicollis LeConte AK BC pugetanus Casey rectilatus Casey 31 Carabidae P. ecarinatur, Hatch BC AB P. herculane,rs Mannerheim BC scenicus Casey P. neobrunneus Lindroth BC P. nigrocoerrleus Van Dyke BC P. protractus LeConte BC AB fontinalis Casey inornah.ts Bland prouenss (Casey) utensis t.Casey) uiuax (Casey) P. tristis (De"iean) ON PQ NB NS interfector (Newman) subarcuztus LeConte sustentu s LeConte tetriculc Casey zephyru; Casey (Subgenus L.DPTOFERONIA Casey) Key to speries: Hacker (1968) P. pumilus Casey BC longicol'.is (Casey) longicol'.is LeConte oregonui (Chaudoir) oregonis Cslki P. sphodrinus LeConte BC AB idahoanus (Casey) (Subgenus M ONOFERONIA Casey) P. diligendus (Chaudoir) PQ apalachius Horn osculan:, Casey (Subgenus G A.STRELLARIUS Casey) P. honestus (Say) ON PQ NB NS america ous (Castelnau) atronite * (Casey) deficien.t (Casey) fastiditu s (Dejean) scolopateus (Casey) (Subgenus S'IEROPUS Dejean) (Refonic: Casey) (.Sterode rus Motschulsky) P. circulosus Lindroth AK YK (Subgenus B')THRIOPTERUS Chaudoir) (Dysidit.s Chaudoir) (.Pararg;ttor Casey') P. adstrictus Eschscholtz+ AK YK NT BC AB SK MB ON PQ NB NS PE LB NF angusticollis (Casey) commixius Chaudoir latebricola (Casey) latescans (Casey) laxicolli.; (Casey) luczotii .Dejean) maehlini LeConte oblong ius c ulus (Motschul sky) oblongotzotatus (Say) ob t u sa n,7 ulu s (Motschulsky) saxatilis (Casey) sericeus (Casey) seriepun ctatus Mannerheim sex,p unc :atus (Mannerheim) shastan.rs (Casey) P. lustrans L,eConte BC atrolucens (Casey) p uncticollis (Motschulsky) P. mutus (Say) BC MB ON PQ NB NS PE NF carbonarius (Dejean) egens (Casey) 32 Carabidae rnlorosus (Dejean) picicornis (Kirby) politus (Harris) puluinatus Hausen stenopus Hausen P. oregonus LeConte BC colligatus (Walker) P. pensylvanicus LeConte NT BC AB SK MB ON PQ NB NS PE NF (Subgenus MORPHNOSOMA Lutshnik) (Euferonia Casey) (Omaseidius Jeannel) P. coracinus (Newman) oN PQ NB NS PE LB NF adjunctus LeConte erebeus Casey flebilis LeConte lacustris (Casey) ludibundus (Casey) moerens (Newrnan) (?) monedulus (Newman) roanicus (Casey) rufitarsis (Nicolay & Weiss) strigosulus (Casey) uenator (Casey) uashingtonensis (Nicolay & Weiss) P. Iachrymosus (Newman) ON PQ NB P. melanarius (IlligerX _BCAB SKMBONPQNBNSPE-NF P. novus Straneo -MBONPQ P. relictus (Newman) ON protensus LeConte P. stygicus (Say) oN PQ bisigillatus (Harris) (?) picipes (Newman) probus Casey quadrifer (Casey) rugicollis (Haldeman) subaequalis (Casey) umbonatus (Casey) uapidus Casey (Subgenus ABACIDUS LeConte) (Peristethus LeConte) P. atratus (Newman) ON P. permundus (Say) ON planifer (Casey) (Subgenus MELANIUS Bonelli) (Ly p ero som us Motschulsky) (Lyperus Chaudoir) (M e tame I ani us Tschitsch6rine ) (Omaseus Dejean) P. castor Goulet & Bousquet ON PQ P. caudicalis (Say) NT BC AB SK MB ON PQ NF agrestis Bland breuibasis (Casey) P. corvinus (Dejean) NT BC AB SK MB ON PQ NB aequalis (Casey) subpunctatus (Harris) tenebrosus (Chaudoir) P. ebeninus (Dejean) ON acutangulus (Chaudoir) (Subgenus PSEUDOMASEUS Chaudoir) P. luctuosus (Dejean) BC AB SK MB ON PQ NB NS PE NF abjectus LeConte confluens (Casey) hamatus (Harris) P. tenuis (Casey) AB MB ON PQ NB NS PE NF testaceus (Casey) (Subgenus LAGARUS Chaudoir) (Pseudargutor Casey) (P se udol ag ar us Lutshnik) 33 Carabidae P. Ieconteianus (Lutshnik) BC SK MB ON PQ NB NS erythropus (Dejean) nitidus iKirby) (Subgenus ARGUTOR Dejean) (Americ cmasezzs Csiki) (Micronnseus Casey) (.O ma se : rlu s Lutshnik) P. corruscultLs LeConte ON PQ P. femoralis iKirby) AB SK MB oN PQ aequicoi.Iis (Casey) desidiosus LeConte P. patruelis (Dejean) AK BC AB SK MB ON PQ NB NS PE LB NF bicolor (Kitby) linearis (Mannerheim) P. strenuus (Panzer)f BC NF (Subgenus CRYOBIUS Chaudoir) Key to No th American species: Ball (1966) P. arcticola (Chaudoir) AK YK NT MB PQ LB I ab r ad.a rens is ( Chaudoir) P. auriga Ball AK P. barryorun Ball AK NT AB P. biocryus liall AK P. brevicornis (Kirby)* AK YK NT BC nfg ON P6-LB NT delicatus (Casey) e p ipl e ur alis (J. Sahlberg) fa s t idiosus (Mannerheim) P. bryanti (\ian Dyke) NT P. bryantoid:s Ball AK P. cacumenir; Ball AI{ P. caribou Brrll AK YK NT _MB P. chipewyarr Ball NT BC AB _MBON P. empetricola (Dejean)* AKYK_BC globicollis Csikt r ot undi :olli s ( Mannerheim) r ufi coll,:. s (Mannerheim) P. gerstlensis Ball AK P. haftorni I.,indroth AK P. hudsonicr:.s LeConte AK YK NT BC P. kotzebuei Ball AK P. mandibulrryoides Ball AK YK NT BC P. nivalis {R F. Sahlbergt+ AK P. parasimil:s BaIl* AK YK pl:) P. pinguedineus (Eschscholtz)x AK YK I'ff - - rg alaskensis (Poppius) beringi (Casey) diplogrnus (Chaudoir) holmbergi (Poppius) incogni;lus (Poppius) mandibularis (Kirby) montanellus Poppius splendidus (J. Sahlberg) s ub nititlul u s (Poppius) s ub s inuos us (Chaudoir) washingtoni (Casey) P. planus (J. Sahlberg) AK blaisdeili Van Dyke P. riparius ( Dejean) AK YK NT BC AB breuiusculus (Casey) caseyi Csiki /ofuus (Mannerheim) fus coae,zeus ( Chaudoir) laeuilatus Notman patulus (Casey) P. similis Mannerheim* AK q uadr icollis ( Mannerheim) P. soperi BalI AK NT P. stantonersis BaIl NT 34 Carabidae P. surgens LeConte AB P. tareumiut Ballx AK YK NT P. tiliaceoradix Ball AK P. ventricosus tEschscholtz)x AI{ YK NT aleutorum (Lutshnik) b reu icoll is (Tschitsch6rine) hy perboreus (Mannerheim) otariidinus (Casey) r ufi sca p us (Mannerheim) r ug ifer us ( Tschitsch6rine) s ubcaudatus ( Mannerheim) s ubexarat us (Mannerheim) u indic atus (Mannerheim) (Subgenus LYPEROPHERUS Motschulsky) Key to species: Budarin (1976) P. agonus Horn* AK YK NT P. costatus (Men6tri6s)* AK YK NT P. punctatissimus (Randall) - YK NT BC AB SK MB ON PQ NB NS - LB NF P. sublaevis (J. Sahlberg)* AI{ YK NT rufofemoralis Van Dyke P. vermiculosus (M6n6tri6s)x AK YK NT innuitorum (W.J. Brown) Genus CYCLOTRACHELUS Chaudoir Key to species: Freitag (1969) (Subgenus EVARTHRUS LeConte) C. sodalis sodalis (LeConte) - ON corax (LeConte) fatuus (LeConte) inflatipennis (Casey) I icinoide s ( Motschul sky) retractus (Casey) sulcatus (Casey) uagans (.LeConte') Genus ABAX Bonelli A. parallelepipedus NS NF (Piller & Mitterpacher)f aler (Villers) s/riolo (Fabricius) Tribe Zabrini Genus DISAMARA Lindroth Pseud,amara Lindrotn D. arenaria (LeConte) _ONPQNBNSPE Genus AMARA Bonelli (Subgenus CURTONOTUS Stephens) (Leirus Zimmermann) A. alpina (Paykull)* AKYKNTBCAB-MBONPQ-LBNF alaskana Cslki angustata J. Sahlberg arguta (Casey) borealis (Chaudoir) brunnipennis Dejean deficiens (Casey) inanis (Casey) obtusa LeConte rubripennis (Casey) subtilis (Casey) A. aulica (Panzer)t A. blanchardi Hay'ward ___sc____PQIUNS__NF A. bokori Csiki* AK YK NT BC MB sahlbergi Poppius A. carinata (LeOonte) _BCABSKMBONPQNB 35 Carabidae adstricta (Putzeys) catenulata (Casey) giluipes (Casey) laticollis (LeConte) A. hyperborea Dejean* AK YK NT BC AB SK MB ON PQ NFLB canadensis (Putzeys) elongatr.r (LeConte) imperfe':ta (W.J. Brown) tristis (Prtzeys) A. lacustris l,eConte AK YK NT BC AB SK MB ON NS manitobensis (Casey) A. pennsylva-nica Hayward -oN PQ NS fuluipes (Putzeys) A. thoracica Hayward BC AB SK A. torrida (Panzer)* AK YK NT BC AB SK MB ON PQ Ne NS pn LB NF albertana (Casey) biarcuala (Casey) breuilat'ris (Kirby) breuipennis (Casey) consueta Fall cylindrica LeConte dura (Casey) holnrbergi (Putzeys) hudsonica Hayward labradarensis (Casey) melanollastrica Dejean reflexa rPrtzeys) rufimat a (Kirby) scrutat( (Casey) somnoh nta (Putzeys) (Subgenus B RADYTUS Stephens) (P seuda brady tzs Csiki) A. apricaria Paykull)t BC SKAB MB ON PQ NB NS LB NF putzeystana Csiki putzeysti Hotn A. avida (Sa d BC SK MB ON PQ NB NS PE NF confinis Dejean A. browni Li rdroth -YK NT A. colvillensi s Lindroth AK NT -ms-- A. exarata Dejean ON curta ((:asey') furtiua;Jay grauida (Casey) stygiali:; (Casey) A. fulva (O.F. Mriller)i _PQ NB NS PE LB NF A. glacialis ( Mannerheim)* AK YK Nr sc SK oN PQ _LB nainensis Casey A. Iatior (Kirby) _BC SK MB oN PQ NB NS NF decepta (Casey) humphreysi (Casey) laeuistri.ata (Putzeys) Iibera LeConte oregona LeConte relicta (Casey) A. schwarzi.{ayward YK NT AB MB ON PQ septentrionalis (LeConte) (Subgenus PERCOSIA Zimmermann) Key to species: Hieke (1978) A. obesa (Say) YK NT BC SK MB ON PQ NB NS PE NF dffinis (LeConte) extensa (Casey) latissima (Casey) sulcatuta (Casey) uentricosa (Casey) (Subgenus CELIA Zimmermann) (.Isopleurus l{rtby) A. aurata De;jean euanida (Casey) 36 Carabidae fragilis (Casey) gouanensis (Casey) irnitatrix Horn proba (Casey) A. bifrons (Gyllenhal)t PQ NB NS PE LB NF A. brunnea (Gyllenhal)* AK am pl ico ll is (Mannerheim) A. californica Dejean BC mericana Dejean perspecta Casey pugetana (Casey) A. chalcea Dejean ON coruina (Casey) maneei (Casey) pinorunr (Casey) schotti (Casey) sodalis (Casey) A. discors Kirby AK YK NT BC AB SK MB ON PQ NS LB NF gibba (LeConte) lugubris (Casey) merula (Casey) neuadica (Casey) obsolescens (Casey) paganica (Casey) paroudna (Casey) scolopax (Casey) spaldingi (Casey) sphaerops (Casey) wahelandi Hatcft A. hicksi Lindroth* NT A. idahoana (Casey) _BC AB exlineae Minsk & Hatch A. musculis (Say) _BC AB SK MB ON PQ I\D breuitarsis (Casey) contempta (LeConte) crenulata (Casey) curticeps (Casey) fluminea (Casey) limbalis (Casey) lyncea (Casey) minnesotana (Casey) nugator (Casey) paulula (Casey) scintilla (Casey) shoemaheri (Casey) uegrandis (Casey) A. pseudobrunnea Lindroth AKl'K-BCABSKMB-PQ LB A. quenseli (Schbnherr)x AK YK NT BC AB SK MB ON PQ NB NS pe LB NF brumalis (Casey) brunnescens (Casey) ceruicalis (Casey) defecta (Casey) definita (Casey) docilis (Casey) eldorensis (Casey) erplanatula (Casey) exposita (Casey) femoralis Hotn fontinalis (Casey) hornis Cslki i ndi s t inc ta (Mannerheim) laxicollis (Casey) mimica (Casey) obligata (Casey) piperi (.Casey) relucens (Mannerheim) remota (Zimmermann) remotestriata Dejean tahomae (Casey) washoeana (Casey) A. rubrica Haldeman ON PQ 37 Carabidae ferruginea Casey liquida (Casey) lubrica (Casey) lucina (Oasey') nigripennis (Casey) pallida (Casey) piceonitt:ns (Casey) politissitna (Casey) proditor (Casey) A. sinuosa (Casey) AK YK NT BC AB SK MB ON PQ NB PE NFLB elusa (Casey) hospes (Casey) nitida (Yitby) nupta (Casey) subaenett (LeConte) subaenescens Csiki thoracicu (Casey) A. spuria Lindroth AK NT (Subgenus AIIARA s. str.) A. aenea (DeGeer)f oN PQ NB NS PE NF deuincta Casey A. aeneopolit.r Casey NT_ABSKMB oN PQ NB_ LB NF A. coelebs Hayward _BCABSKMB leuiceps Jasey nebrask<,.na Casey oblonguta Casey rustica ()asey A. communis (Panzer)t A. conflata Lt:Conte BC Iu_ diffidenr Casey leydeni l.)asey semota (Jasey) A. confusa Lt Conte AK YK BC SK MB PQ acomant'. Casey ebenina Casey impeditu Casey oblongifrrmis Casey protenso Putzeys subarctica Casey subpunctata LeConte uiridula Casey A. convexa L+:Conte BC AB SK MB ON PQ breuiforrnis Casey cockerell,i Casey frugalis (Casey) lecontei,Jsiki ouiformis Casey piceola (.Jasey polita LeConte wingatet Casey A. cupreolata Putzeys NT BC SK MB ON PQ NB NS PE eneruis Oasey haywarc!.i Csiki inflaticol.lis Casey neoscotica Casey paruiceps Hayward tenax Ctrsey A. ellipsis (C;rsey) BC AB SK MB ON PQ w innipe;1en sis (Casey) A. emancipata Lindroth BC A. erratica (Duftschmid)x AK NT BC -s^ Ite - pe -iuNr A. eurynota (Panzer)t NF A. familiaris (Duftschmid)t AK BC AB -oN PQ NB NS PE_NF humilis Casey A. farcta LeConte YK BC AB SK MB albertae (Casey) formalis (Casey) funebris (Casey) margino.tella (Casey) olympia (Casey) 38 Carabidae parallela (Casey) sho.stanica (Casey) solita (casey) subdepressa (Casey) subsimilis (Casey) uancouueri (Casey) A. impuncticollis (Say) SK MB ON PQ NB NS PE NF (? ) a nt hracina Haldeman arcuata Casey dfficilis LeConte edor Notman otiosa Casey wadei Casey A. interstitialis Dejean* AK YK NT A. Iaevipennis Kirby AK_NTBCABSK ms oN PQ NB NS PE NF aeneolucens Casey A. littoralis Mannerheim AK YK NT BC AB SK MB ON PQ NB NS PE NF conuergens Casey fallaxLeConte hesperia Casey inepta LeConte heeni Casey lacustrina Casey laurana Casey microcephala Casey mystica Casey oodiformis Casey pullmani Casey recticollis Casey (?) teres Notman turbata Casey A. iunicollis Schiddtex AK YK BC AB SK MB ON PQ NB NS LB NF carriana Casey marquettensis Casey A. patruelis Dejean* AK YK NT BC AB SK MB ON PQ NB NS PE NF columbiana (Casey) inaequalis Kirby reducta (Casey) splendida Haldeman A. sanjuanensis Hatch BC (Subgenus ZEZEA Csiki) (Pseudotriaena Minsk & Hatch) (Triaena LeConte) A. angustata (Say) SK MB ON PQ flebilis (Casey) indistincta Haldeman A. Iongula LeConte BC afoueolata Hayward alaxnoguia Minsk & Hatch atrichata Minsk & Hatch glabrata Minsk & Hatch profuga (Casey) A. pallipes Kirby BC AB SK MB ON PQ NB NS depressa (LeConte) h,incaidi Minsk & Hatch lawrenciana (Casey) parallela (Casey) sherrnani (Casey) A. scitula Zimmermann BC AB SK MB irregularis (Casey) obliqua Casey prouana Casey uinta (Casey) uapida (.Casey) Tribe Harpalini Genus CRATACANTHUS Dejean C. dubius (Palisot de Beauvois) AB SK bisectus Casey 39 Carabidae litoreus Jasey pensylucinicus Dejean Genus PIOS0MA LeConte P. setosum L-'Conte - BC AB SK breuipenne Casey Genus EURYDERUS LeConte Nothopus LeConte Key to species: Bail (1960) E. grossus (S,ryt - BC AB SK MB ON arizonic,ts (Casey) obtusus .Casey) priuatus (Casey) ualens ((.)asey) zabroides LeConte Genus HARF ALOBRACIIYS Tschitsch6rine H. leiroides t vlotschulskyrx AK - NT - A-B Genus HARPAI-US Latreille (Subgenus OPHONUS Dejean) H. puncticeps (Stephensti - ON PQ - NS H. rufibarbis t Fabricius rt PQ (Subgenus PSiEUDOPHONUS Motschulsky) Key to Norbh American species: BaIl and Anderson (1962) H. bicolor tFz bricius t MB ON PQ - NS admissu; Casey compar .,eConte excubant: Casey feroculut Casey nactus Casey H. ery.thropu r Dejean MB ON PQ - NS abstrusus Casey cupiens t)asey deludent Casey effetus Casey fenisex Casey rufopicet.ts Casey H. faunus Sa.r MB ON PQ badius Ltejean conuiuus LeConte H. longicollis LeConte - ON PQ dolosus ()asey haldemoni Casey latescanr Casey uagans L,eConte H. paralus Cusey AB SK spaldingi Casey H. pennsylva:ricus (DeGeer) - BC - SK MB ON PQ NB NS PE inmixtus Casey longior \:Jtby mormoni.cus Casey H. rufrpes (DeGeer)t PQ NB NS PE - NF (Subgenus M IGAPANGUS Casey) H. caliginosu; tFabricius) MB ON PQ dux Casey (Subgenus PI,ECTRAI-IDUS Casey) Key to species: El-Moursy (1958) H. erraticus Siay MB ON PQ caudalis Casey (Subgenus PIIARAJ-US Casey) H. indianus C siki - ON testaceus LeConte 40 Carabidae (Subgenus IIARPAI-US s. str.) H. affinis (Schranktt -BCABONPQNBNSPELBNF aenescens Casey oenezs (Fabricius) assimilis Dejean canonicus Casey conuictor Casey lustralis Casey uiridiaeneus Palisot de Beauvois uiridis Say (Subgenus HARPALOMERUS Casey) H. amputatus Say AK YK NT BC AB SK MB ancillaris Casey bracatus Casey cuculus Casey cupreolatus Casey papagonalis Casey rotundicollis I\ttby stephensi l\ttby transuersus Casey (Subgenus EUHARPALOPS Casey) H. animosus Casey _BCABSK montanus LeConte montuosus Csiki H. egregius Casey AK YK NT BC AB SK MB ON PQ _ NS PE LB NF alienus LeConte baergi Csiki cascadiensis Hatch instructus Casey rufimanus LeConte H. fraternus LeConte -BCAB nimius Casey occidenlalis Chaudoir praestans Casey wadei (Casey) H. funerarius Csiki AB SK MB ON PQ funestus LeConte reuersus Casey H. hoppingi Lindroth _BC H. laticeps LeConte AK YK NT BC AB SK MB ON PQ NB NS - LB NF solutus Casey H. lewisii LeConte NT-ABSKMBONPQNBNS aesopus Casey H. viduus LeConte _ONPQ prouidens Casey $ H. indigens Casey - ON PQ NB beatulus Casey piperi (Casey) H. ventralis LeConte AB SK MB electus Casey $ H. fulvilabris Mannerheim AK YK NT BC AB SK MB ON PQ NB NS PE LB NF H. megacephalus LeConte MB ON (Subgenus CORDOHARPALUS Hatch) H. cordifer Notman AK BC w ashingtoniensls Van Dyke $ H. carbonatus LeConte NT BC AB SK H. fallax LeConte - ON PQ NB NS amiculus Casey recisus Casey H. herbivagus Say - BC AB SK MB ON PQ NB NS PE ossenszs Casey 4L Carabidae foueicollis LeConte mutabilis (Haldeman) H. intactus Casey _BC uintanus Casey H. nigritarsis C.R. Sahlbergx AK YK NT BC AB MB ON PQ NFLB curtatus Mannerhein proximus LeConte recensus Casey H. pleuriticus Kirby AK BC SKAB MB ON PQ NB NS PE LB NF aequabilis Casey lasciuus tJasey liuidulus Casey niualis Casey perspicuus Casey placidus Casey pumilio Casey H. seclusus Casey -BCAB parowanus Casey H. solitaris D,:jean NT BC AB SK MB ON PQ NFLB fuliginos zs (Duftschmid) opicus Ctl.sey H. somnulentrs Dejean BC blanditu; Casey contactut'Casey pacificus FalI H. uteanus Cr,Lsey AB SK MB oppositut Casey xA H. alaskensis Lindroth AK H. innocuus LeConte NT sc AB oN pe fugitans Casey intactus Jasey persolus Casey $ H. cautus Dej:an BC aduena I eConte (? ) alb ior, icus Mannerheim crenatell n Casey columbianus Casey (?) defix.us Walker oregonensis Casey H. ellipsis Le(lonte H. obnixus CzLsey ---scTsK------H. ochropus FJrby AK BC AB SK MB ON PQ H. opacipennis (Haldeman) AK YK NT BC AB SK MB ON PQ lacustris Casey leuiceps Oasey oodioides Chaudoir H. plenalis Casey NT BC MB ON PQ NB NS NF latebricoi,a Casey modulatus Casey $ H. desertus LeConte BC AB SK captiosu:t Casey couleensts Hatch dulciculus Casey lucidus LeConte lustrans Casey metuens Casey uaciuus Oasey R Y H. fulgens Csiki ON nitidulus Chaudoir Genus IIARP ALELLUS Lindroth H. basilaris (Kirby) AK YK NT BC AB SK MB ON PQ NS durescans (Casey) 42 Carabidae er.tensus (Walker) furuiculus (Casey) obesulus (LeConte) oblongus (Casey) renoicus (Casey) sejunctus (Casey) stocktonensis (Casey) subenormis (Casey) uaricornis (LeConte) uentricosus (Casey) Genus TRICHOTICHNUS Morawitz (Subgenus TRICHOTICHNUS s. str.) (Episcopellus Casey) (Pteropalus Casey) T. autumnalis (Say) ON nitescens (Casey) T. dichrous (Dejean) ON fluuialis (Casey) uersutulus (Casey) T. vulpeculus (Say) ON PQ nigripennis (Dejean) Genus SELENOPHORUS Dejean (Subgenus SELENOPHORUS s. str.) (Gynandropus Dejean) (Hemisopalus Casey) S. gagatinus Dejean ON PQ NB m(turus Haldeman uiridescens LeConte S. hylacis (Say) ON americanus (Dejean) S. opalinus (LeConte) oN PQ iripennis LeConte S. pedicularius Dejean ON aereus LeConte puellus Putzeys troglodjttes Dejean S. planipennis LeConte BC AB SK MB ON ( Subgenus CELIAMORPHUS Casey) S. ellipticus Dejean ON currens (Casey) opaculus (Casey) oualis Dejean Genus DISCODERUS LeConte Selenalius Casey D. parallelus (Haldeman) BC ON amer icanus (Motschulsky) gener Casey hesperus Casey paruuliceps Casey Genus GEOPINUS LeConte G. incrassatus (Dejean) AB SK MB ON PQ fluuiaticus Casey Genus ANISODACTYLUS Dejean (Subgenus ANADAPTUS Casey) A. discoideus Dejean sK yu oN PQ A. nivalis Horn BC paruulus (Casey) A. pitychrous LeConte _ABSKMB A. sanctaecrucis (Fabricius) BC AB SK MB ON PQ NB lls PE baltimoriensis (Say) uteanus (Casey) 43 Carabidae A. viridescenr LeConte chalceus LeConte idahoensis (Hatch) Iecontei Gemminger & Harold (Subgenus G\T{ANDROTARSUS LaFert6-S6nectbre) (Triplect:us LeConte) Key to Norl;h American species: Noonan (1973) A. dulcicollis LaFert6-S6nectdre) ON ellipticu: LeConte elongatu.; Chaudoir modicus (Casey) A. merula Ge'mar MB ON PQ aethiops (Casey) crassus LeConte grauidus LeConte hetnpi (Casey) margina.lus (Casey) pinguis l.,eConte sulcipennis (Casey) uolcotti iCasey) A. ovularis {Casey} -oN semirubilus (Casey) A. rusticus t S ry r MB ON PQ NB NS PE oblongus (Casey) lrjslls Dr jean (Subgenus AIIISODACTYLUS s. str.) (.Cephaloqyna Casey) A. binotatus ( Fabricius tt _BC A. californicu r Dejean _BC aleneanus Casey angustus Casey comes Ct.sey confusus LeConte humeral,s Casey maestus Casey obsolescens Casey oregonus Casey paganict s Casey sinuatus Casey A. carbonariu s t Say t oN PQ breuior (t laseY) luctuosu,:; Dejean rufipenn,'.s LeConte A. furvus LeC onte -oN A. harrisii Le'lonte BC AB SK MB ON PQ NS- Iacertosus Casey A. kirbyi Lincroth BC MB ON PQ NB NS (?) Iecontei Chaudoir A. melanopus (Haldeman) ON A. nigerrimus {Dejean) oN PQ NN NS NF Iaticollis (Kirby) opacus (()asey) punctulatus LeOonte A. nigrita Dejean BC AB SK MB ON PQ NB NS PE inte r p un.:t at u s (Kirby) A. similis Le(lonte BC incertus Jasey incisus C. asey punclico,lis Chaudoir semipunctdtus LeConte sericatus Casey solidus Casey (Subgenus AI'LOCENTRUS LeConte) A. amaroides LeConte BC (Subgenus PS,EUDAPLOCENTRUS Noonan) A. laetus Dejr an ON gemmeus (Casey) 44 Carabidae (Subgenus SPONGOPUS LeConte) A. verticalis (LeConte) SK MB ON PQ Genus AMPHASIA Newman (Subgenus AMPI{ASIA s. str.) A. interstitialis { Say) ON PQ fuluicollis Newman mollis Casey obscuripennis (Dejean) (Subgenus PSEUDAMPHASIA Casey) A. sericea (Harris) SK oN PQ femorata (Dejean) Genus DICHEIRUS Mannerheim Key to species: Noonan (1968) D. piceus (Menetriest BC alutaceus Casey angustulus Casey australinus Casey decoloratus Casey facilis Casey incidens Casey iteg ul ar is Motschulsky insularis Casey parallelus LeConte rupimontis Casey sodalis Casey uillosus Motschulsky Genus XESTONOTUS LeConte X. lugubris tDejeant oN PQ NS manhattanis (Casey) tioganus (Casey) Genus NOTIOBIA Perty (Subgenus ANISOTARSUS Chaudoir) (Diaphoromerus Chaudoir) (Eurytrichus LeConte) (Stilbolidus Casey) Key to North American species: Noonan (1973) N. nitidipenni s (LeConte) oN PQ agitabilis (Casey) cephala (Casey) conspectd (Casey) delicata (Casey) tenuitarsis (Casey) N. picea tLeConte) oN PQ soyi (Blatchley) N. terminata tSayl ON PQ NB NS agilis (Dejean) angusticollis (Casey) conniuens (Casey) florid,ana (Casey) foueicollis (Bates) fuscipennis (Casey) hebes (Casey) inerrans (Casey) ocreata (Say) parallela (Casey) similis (Say) suboualis (Casey) subuirens (Casey) testacea (Haldeman) uernicata (Casey) 45 Carabidae Genus TRIC JOCELLUS Ganglbauer (Subgenus T RICHOCELLUS s. str.) T. cognatus (Gyllenhal)x AK YX NT BC AB SK MB ON PQ NB NS PE LB NF axillar i s (Mannerheim) boreellus Casey conflag r at us ( Mannerheim) lateralis Casey longius<. ul us (Mannerheim) montico.ta Casey nitens (l,eConte) punctipr,:nnis Casey ruficrus (Kirby) (Subgenus O REOXENUS Tschitsch6rine) T. mannerheimii (R.F. Sahlberg)* AK YK NT BC MB - PQ porsildi W.J. Brown Genus AMEIIINUS Casey A. linearis (I.eConte) - ON Genus BRAITYCELLUS Erichson (Subgenus L OCELLUS Motschulsky) (Glyceri n Casey) B. politus tF:rllt BC (Subgenus B RADYCELLUS s. str.) B. harpalinu; tAudinet-Serviller* - BC (Subgenus C \THARELLUS Casey) B. lecontei C;iki AK YK NT BC AB SK MB ON PQ NB NS PE LB NF cordicol'is (LeConte) (Subgenus S' IENOCELLUS Casey) B. congener tl,eContet - BC AB SK MB ON PQ alutacet.s (Casey) B. nigerrimu; Lindroth AB SK B. nubifer Lt Conte - BC AB nubicollis (Casey) uentrali; LeConte B. rupestris r Say) - ON PQ NB NS antennolis (Casey) (.?) cinct;rs (Say) (?) debilipes (Say) elong att":lus (Dej ean) flauipes (Kirby) parallel,^rs Chaudoir A B. insulsus (Oasey) -oN B. neglectus il,eConte) _ BC AB SK MB ON PQ NB NS PE _ NF B. nigriceps LeConte MB ON PQ occultus (Casey) $ B. californicr.s (LeContet - BC B. tantillus (Dejean) - ON dfficilis (Dejean) (Subgenus LIOCELLUS Tschitsch6rine) B. nigrinus (Dejean) AK BC AB SK MB ON PQ NB NS PE - NF quadricollis (LeConte) tibi,alis (Kirby) B. semipubescens Lindroth AB ON PQ NB (Subgenus TRILIARTHRUS Casey) B. atrimedeus (Say) MB ON PQ NB B. badipennis (Haldeman) - ON PQ curticoliis (Casey) properus (Casey) 46 Carabidae B. conformis (Fall) BC tetricus (Casey) B. georgei Lindroth AB B. kirbyi (Horn) oN PQ B. lugubris (LeConte) AB _MBONPQ NB PENS NF frosti (Fall) protractus (Casey) Genus STENOLOPHUS Stephens (Subgenus STENOLOPHUS s. str.) S. anceps LeConte BC fidelis Casey rolu nd i colli s Motschul sky S. carbonarius (Dejean) ON NS S. fuliginosus Dejean BC AB SK MB ON PQ NU NS PE N" fuscipennis (LeConte) uersicolor Kirby S. fuscatus Dejean ON PQ S. humidus Harnilton ON PQ_ S. incultus Casey BC fr_ _ _ consors Casey debiliceps Casey S. limbalis LeConte BC indistinctus Motschulsky longitarsis Casey S. megacephalus Lindroth oN PQ S. ochropezus (Say) _MBONPQ NB NS conuexicollis LeConte floridanus Casey (?) gracilis Casey testaceicollis Casey S. plebejus Dejean ON PQ (?) lugubris (Haldeman) riuularis Casey (Subgenus AGONODERUS Dej ean) S. comma (Fabricius) BC SK MB ON PQ NB NS PE NF dorsalis (LeConte) gracilitarsis Casey obliqulus Casey similis (I\trby) S. lecontei (Chaudoir) ON PQ plagiatus Casey S. lineola (Fabricius) AB SK MB ON PQ furcatus (Fabricius) s S. conjunctus (Say) BC SK MB ON PQ NB NS PE captiosus Casey humilis (Dejean) immunis (Kirby) misellus (Dejean) moquinus Casey thoracicus Casey S. rotundatus LeConte -oN S. rotundicollis (Haldeman) -oN pe- incitatus Casey scitulus Casey Genus ACUPAJ-PUS Latreille (Subgenus PHILODES LeConte) (Goniolophus Casey) A. alternans (LeConte) oN PQ testaceus (LeConte) A. rectangulus Chaudoir oN PQ (Subgenus ACUPALPUS s. str.) A. canadensis Casey __ A. carus (LeConte) _ NC_:"MB3Ni3NBN3Ff;_ Nil 47 Carabidae curlipet nis Casey expertus Casey subrectu,.s Casey triuialis Casey A. meridiant s ll-inn6t+ - BC AB - PQ A. nanellus (lasey - ON PQ NB NS A. pumilus I indroth - ON PQ - NS (Subgenus T A.CHISTODES Casey) A. indistinct rs Dejean - ON obscurus (Casey) A. partiariu: tSayt ON PQ conuerg tns (Casey) Iyratus Casey') A. pauperculus Dejean ON PQ NB NS PE consimi'.is Dejean A. testaceus Dejean - ON PQ fusciceps (Casey) micros LeConte Tlibe Licinini Key to North American species: Ball (1959) Genus DIPL ]CHEILA Brull6 Rembus Dejean (Subgenus If IOREMBUS Jeannel) D. assimilis rl-eConter ON PQ modesta Casey D. impressic',llis rDejeanr ON PQ cliens C*ey foueata Jasey laticollis (LeConte) planulala Casey D. major ma. or (Leconte ) - ON expansa Casey oblonga Casey procera Casey D. obtusa (LeConte) NT BC AB SK - ON PQ NB NS PE parallelt Casey D. oregona tllatcht BC AB SK MB D. striatopurLctata (LeConte) NT BC AB SK MB ON PQ alternar.s Casey amplipennis Casey breuicoli,is Casey D. undulata larr AB - MB Genus DICAELUS Bonelh (Subgenus P.ARADICA-ELUS Ball) D. dilatatus lilatatus Say PQ carolinensis Casey dejeanii Dejean & Boisduval planicollis LeConte D. elongatus Bonelli - ON PQ osieuill rnsis Casey debilice;ts Casey obscuru.; LeConte simplex Dejean D. politus Dt jean - ON PQ angustus Casey leonardli Hatis D. sculptilis ntricatus LeConte - ON ocellatus Blatchley D. sculptilis rpioides Ball - SK MB ON D. teter Bonelli ON PQ ouipenn'.s Casey (Subgenus DICAELUS s. str.) D. purpuratus purpuratus Bonelli * ON uiolaceus Bonelli 48 Carabidae (Subgenus LIODICAELUS Casey) D. laevipennis laevipennis LeOonte AB eudnescens (Casey) Genus BADISTER Clairville (Subgenus BADISTER s. str.) B. lernrgineus Dejean - BC anthracinus LeConte B. neopulchellus Lindroth NT BC AB SK MB ON PQ - NS PE B. notatus Haldeman - ON PQ angustus Casey giluip.es Casey terminalis LeConte B. obtusus LeConte NT - AB SK MB ON PQ - NS B. pulchellus LeConte - ON (Subgenus BAUDIA Ragusa) B. grandiceps Casey - BC AB - MB ON PQ - NS B. transversus Casey MB ON PQ B. ocularis Casey - ON PQ NB NS PE B. parviceps Ball - BC MB ON PQ B. reflexus LeConte - ON seclusus Blatchley Tribe Oodini Genus OODES Bonelli O. amaroides Dejean - ON O. americanus Dejean - ON PQ fluuialis LeConte O. brevis Lindroth - ON Genus LACHNOCREPIS LeConte L. parallela tSayt MB ON PQ Genus STENOCREPIS Chaudoir (Subgenus STENOUS Chaudoir) S. cuprea tChaudoirt - ON leucodacty Ia (LaFert6-S6nectdre) Tribe Panagaeini Genus PANAGAEUS Latreille P. fasciatus Say -oN Tlibe Callistini Genus CHLAXNIUS Bonelli Key to North American species: Bell (1960) (Subgenus EURYDACTYLUS LaFert6-S6nectdre) (Gly ptoder us LaFert6-S6nectdre) C. tomentosus tomentosus (Say) AB SK MB ON PQ amplus LeConte lacustrinus Casey luctuosus Germar (Subgenus ANOMOGLOSSUS Chaudoir) C. emarginatus Say - ON PQ NB NS delectans (Casey) grauis (Casey) semotus (Casey) 49 Carabidae C. pusillus Say - ON elegant,ilus Dejean nanulu,; (Casey) (Subgenus CHLAENIUS s. str.) C. aestivus liay - ON cobaltir'.us Dejean congener LeConte C. cordicollir, Kirby MB ON PQ NB C. erlrthropr.. s Germar - ON oblongi;cennis Casey rufilabris Dejean umbritursis Casey C. laticollis :iay ON brachyclerus Chaudoir breuicol.lis LeConte rufipes Dejean C. sericeus sericeus (Forster) - BC AB SK MB ON PQ NB NS PE - NF peruirict.is LeConte uteanus Casey C. solitarius Say AB chlorophanus Dejean (Subgenus C HLAXNIELLUS Reitter) C. brevilabr-s LeCon[e - ON (.?) cons tnilis LeConte texaneltus Casey C. impunctilrons Say MB ON PQ C. nebraske.'rsis LeConte AB SK sedulus Casey C. nemoralir Say - ON Iongico,lis Chaudoir C. pensylvarLicus pensylvanicus Say BC AB SK MB ON PQ NB NS PE - NF (?) fulgtceps Newman impunctifrons l\ttby pubescc'ns l{arlis uicinus Dejean C. tricolor Dejean - BC AB SK MB ON PQ NB NS NF atripen,'tis LeConte chaly be ipennis Chevrolat jacinto Casey quadricollis Kirby uigilant Say (Subgenus F,RACFM,OBUS Chaudoir) C. Iithophilus lithophilus Say NT BC AB SK MB ON PQ NB NS PE - NF sma ra g' dig e r Motschulsky uiridanus Dejean (Subgenus A GOSTENUS Motschulsky) C. alternatus Horn AK - NT BC AB SK MB ON PQ NB NS NF alberta,tus Casey C. harpalinr s Eschscholtz - BC AB crestonensis W.J. Browrr C. interrupt rs Horn - BC C. niger Rardall NT BC AB SK MB ON PQ NB NS PE - NF ludouici.anus Leng $ C. purpuric< llis Randall NT BC AB SK MB ON PQ frostii Carr punctipennis Casey Tribe Pentagonicini Genus PENIIAGONICA Schmidt-Goebel Didetut LeConte Rhomb tdera Reiche P. picticornis Bates -ONPQ 50 Carabidae Tribe Perigonini Genus PERIGONA Castelnau Trechicus LeConte P. nigriceps tDejeant - ON PQ umbripennis (LeConte) P. pallipennis (LeConte) - ON Tribe Platynini Genus CALATHUS Bonelli Key to North American species: Ball and Ndgre (1972) (Subgenus CALATHUS s. str.) (Fuscocalathus Ndgre) C. fuscipes (Goeze)* - BC (Subgenus NEOCAJ-ATHUS BalI & Nbgre) C. gregarius (Sayt - ON PQ NB NS distinguendus LeConte C. ingratus Dejean AK YK NT BC AB SK MB ON PQ NB NS PE - NF acomanus Casey aquilus Casey beringi Casey calator Casey coloradensis Casey confusus LeConte incommodus Mannerheim labrad,orinus Casey nanulus Casey planifer Casey reductus Casey C. opaculus LeConte - ON PQ alutaceus Casey appalachius Casey obesulus Casey sonoricus Casey uentricosus Casey C. ruficollis grandicollis Casey - BC (Subgenus PROCALATHUS Jedlicka) C. advena(LeConte) AK BC AB SK - ON PQ - LB NF binarius (Casey) b reu ius cul u s (Mannerheim) b runne s cen s (Mannerheim) dzlcis (Mannerheim) lenis (Mannerheim) mollis (Eschscholtz) scolopax (Casey) Genus LAEMOSTENUS Bonelli Key to species: Casale (1988) (Subgenus LAEMOSTENUS s. str.) L. complanatus (Dejean )* - BC (Subgenus PRISTOI.IYCHUS Dej ean) L. terricola (Herbst)f - BC PQ NB NS PE - NF Genus SYNUCHUS Gyllenhal Pristodactyla Dejean Key to species: Lindroth (1956) S. irnpunctatus (Say) - BC AB SK MB ON PQ NB NS PE - NF americanus (Dejean) americanus (LeConte) conuexu.s (Casey) coruinus (LeConte) Genus OLISTHOPUS Dejean Odontonyx Stephens 51 Carabidae O. micans LcConte -oNPQ O. parmatus (Say) _ONPQ cinctus )ay iterans t)asey pictus C'asey Genus SERI{IODA Kirby Agonodromius Reitter S. bembidioides Kirby AK YK NT BC AB SK MB cicatricosa Motschulsky colonica Casey montico,!,a Casey S. bogemannii (Gyllenhal)* AK YK _ BC .AB inuidiosa Casey s t r ig icolle (Mannerheim) tacomae Casey S. obsoleta ([iay) NT BC AB SK MB ON PQ NB NS _ LB NF consimi.e (LeConte) insulina Casey luctuosc (Dejean) uicina (()emminger & Harold) S. quadripun ctata (DeGeer)* AK YK NT BC AB SK MB ON PQ NB NS _ LB NF octocola (Mannerheim) stigmost.t, (LeConte) Genus ANCIIOMENUS Bonelli Tetraleucus Casey A. picticornis, Newman -oN Genus PSEL DANCHUS Casey P. aeneolus (LeConte) -BCAB bjorkmcnae Gray & Hatch P. quadratus (LeConte) -BC dilleyanus (Casey) metuen: (Casey) Genus ATRANUS LeConte A. pubescens (Dejean) _ONPQNB obconia.rs (Haldeman) pallesce.zs Casey Genus RHADINE LeConte Comstockia Van Dyke R. Iindrothi llan AB SK Genus PARANCHUS Lindroth P. albipes (F rbricius)'l _NBNSNF clemens (LeConte) Genus OXYI'SELAPHUS Chaudoir Anchus LeConte O. puncticeps (Casey) -BC_SKMBONPQ_NSPE pusillus (LeConte) Genus AGOI{UM Boneili (Subgenus E UROPHILUS Chaudoir) A. anchomenoides Randall AK _ NT BC AB SK MB ON PQ NB NS PE - NF lanellum Gray A. canadenser Goulet MB ON PQ NB NS A. consimile (GyllenhaI)x AK YK NT BC AB _ MB ON PQ - NS - LB NF fragile (Mannerheim) inualidum Casey A. darlingtorri Lindroth A. exaratum (Mannerheim)* AK5*rec r,rn:x:a]"]'_ A. galvestonicum (Casey) _ _ _ _ 52 Carabidae A. gratiosum (Mannerheim) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF antiquum (Notman) carri (.Casey) properans (Casey) ruficorne (LeConte) symmetricum (Casey) A. lutulentum (LeConte) NT BC AB SK MB ON PQ NB NS PE adustum (Casey) atriceps (Casey) elegantulum (Casey) A. palustre Goulet A. picicornoides Lindroth - - - scABsxrunBNiSNsNspr- A. retractum LeConte NT BC AB SK MB ON PQ NB NS PE collusor (Casey) dil uti p e nne (Motschulsky) elongatulum Haldeman facile (Casey) serenum (Casey) A. simile Kirby AKYK_BCABSKMB A. sordens Kirby AK YK NT BC AB SK MB oN pe Ns Ns pr LB NF ineptum (Casey) frosti (Casey) picicorne (LeConte) A. superioris Lindroth AK-NT-AB-MBONPQNB A. thoreyi Dejeanx AK YK NT BC AB SK MB ON PQ NB NS PE _ NF gemellum (LeConte) (?) lenum Dejean picipenne Kirby (Subgenus PLAT\' IOMICRUS Casey) A. bicoior (Dejean)* AK YK NT BC AB ma r g inell un'L (LeConte) A. fer"ruginosum (Dejean) AK _BC AB SK erasutn (LeConte) marcidum (Casey) uadei (.Casey) A. nigriceps LeConte* AK NT BC AB SK MB ON PQ NB PE NF fragilissimum (Casey) A. piceolum (LeOonte) _BC AB SK MB ON PQ NF d.issensum (Casey) fractum (Casey) frigidulum (Casey) lasciuum (Casey) tritum (Casey) (Subgenus AGONUM s. str.) (Melanagonum Casey) (Paragonum Casey) (Punctagonum Casey) (Taphranchus Casey) A. affrne Kirby AK YK NT BC AB SK MB ON PQ NSNB PE NF cor6o (LeConte) A. albicrus Dejean -oN A. belleri (Hatch) BC A. brevicolle Dejean BC frater (LeConte) A. corvus (LeConte) BC AB SK MB ON PQ debiliceps Casey hyslopi Casey A. cupreum Dejean NT BC AB SK MB ON PQ NB borealinum Casey chalceum LeConte crassicolle (LeConte) cupreolucens Casey esuriale Casey Iongulum Casey marquettense Casey oualicauda Casey parallelum Casey protractum (LeConte) seminitidum Kirby 53 Carabidae A. cupripenne (Say) NT BC AB SK MB ON PQ NB NS PE - NF gemmeum Casey nitidulunt Dejean A. deceptivurn (LeContet NS A. excavatum Dejean - ON PQ NB ontarioni; (Casey) trinariunt (Casey) A. fidele Case, - ON PQ NB NS PE laeue (Let)onte) molestwn (LeConte) subinflat,tm Casey A. harrisii LeConte - BC AB SK MB ON PQ NB NS PE aethiops l)asey mordax Casey A. melanariun Dejean - BC AB SK MB ON PQ NB NS PE maurum Haldeman militare l)asey (?) scuteliiare (.Say) A. metallescer:rs (LeConte) NT BC AB SK MB ON PQ NB NS NF lacustre Oasey terracens: Casey A. moerens Dr jean MB ON PQ A. muelleri (Eerbst)f - BC - ON PQ NB NS PE - NF hardyi (LeConte) A. mutatum ((lemminger & Harold) AK BC - SK MB ON PQ NB NS PE LB NF dtratum t LeConte) A. octopunctat.um (Fabricius) - ON PQ NB NS PE A. placidum (fiay) - BC AB SK MB ON PQ NB NS PE LB NF amplior (.)asey aztecanu;n Casey citatum (iasey morosum Dejean rhodeant.m Casey A. propinquurr (Gemminger & Harold) AK YK NT BC AB SK MB ON PQ NB NS PE - NF amens Ci:tsey fratercuhLm (LeConte) humile Crcey insuetum Casey piceum LeConte A. quinquepu:rctatum Motschuisky* AK YK NT - AB SK MB - PQ - LB perforatum (LeConte) A. subsericeurn (Leconte) - BC AB SK latiusculum Casey suffusum Casey uteanum Casey uirid.issintum Casey A. tenue (Lec')nte) - ON PQ NB NS PE A. trigeminunr Lindroth MB ON PQ NB NS PE (Subgenus MICRAGONUM Casey) (Circinal;:dia Casey) A. aeruginosum Dejean - ON PQ NB NS undulatum (Casey) A. nutans (Sayr - ON PQ femoratun Dejean A. striatopunr.tatum Dejean - ON breuiceps (Casey) decipiens LeConte luculentum (Casey) A. variolatum (LeConte) - BC coronadiizum (Casey) (?) limba:um Motschulsky shastanicum (Casey) (Subgenus CIRCINALIA Casey) A. crenistriatum (LeConte) - oN PQ - NS PE liticola (Casey) politissimum (Casey) roticolle (Casey) statenense (Casey) A. punctiforme t Sayt 2 - 54 Carabidae foueicolle Chaudoir ludouicianum (Casey) A. rufrpes Dejean ON rubripes (Zimmermann) (Subgenus STEREAGONUM Casey) A. errans (Say) BC SK MB ON PQ erythropus Kirby subcordatum (LeConte) A. ferreum Haldeman ON ocreatum Haldeman quadrulum (Casey) solid.ulum (Casey) (Subgenus STICTANCHUS Casey) Key to species: Liebherr (1986) A. decorum (Say) BC SK MB ON PQ arenarium (Casey) californicum (Dejean) charmis (Bates) ftornii (Hausen) itnpictum (Casey) irruptum (Casey) luxatum (Casey) obscurum (LeConte) sitnplex (LeConte) solutum (Casey) syrdcusense (Hatch) tepidum (Casey) t e stac eonot um (Hausen) thoracicum (Dejean) uteanum (Casey) uinnulum (Casey) A. extensicolle (Say) SK MB ON PQ NB NS clientulum (Casey) (? ) cy ane sce ns (Motschulsky) gaudens (Casey) lecontei LeConte ob s c urat utn (Chaudoir) proximum (Harris) uigilans (Casey) uiride (LeConte) (Subgenus DERATANCHUS Casey) A. quadrimaculatum (Horn) AB Genus PLATYNUS Bonelli (Subgenus PLATYNUS s. str.) P. brunneomarginatus (Mannerheim) BC bicolor LeConte bicoloratus Gemminger & Harold cinctellus LeConte ma r ginat u s (M6n6tries) rugiceps (Mannerheim) tenebricosus Gemminger & Harold P. cincticollis (Say) oN PQ NS (?) blandus Germar coruinus (Dejean) deplanatus (Chaudoir) (?) maculifrons (Say) marginalis (Haldeman) P. decentis (Say) AK NT BC AB SK MB ON PQ NB NS LB NF albertanus (Casey) aleneanus (Casey) coracinus (LeConte) d.epressus (Haldeman) gagates (Dejean) iowanus (Casey) missourianus (Casey) oblongipennis (Casey) DD Carabidae pacatus ( Casey) sinuatus (Dejean) turbidus (Casey) P. mannerhei:nii (Dejean)* AK YK NT BC AB SK MB ON PQ NB NS PE LB NF inter s t itiali s (Motschulsky) octofoueo t atus (Maklin) stygicus ,-eConte P. opaculus L':Conte - ON PQ 6oopls (Casey) inquisitor (Casey) P. tenuicollis LeConte) - ON PQ NB NS distinguendus (Casey) margina;:us (LeOonte) pennsylu znicus (Casey) reflexus -l,eConte (Subgenus PI ATYNIDIUS Casey) P. hypolithos Sayr - ON erythrop'.ts Dejean ontarien::is (Casey) P. ovipennis (Mannerheim) - BC arachnoides (Casey) maurus (Motschulsky) rotundip znnis (Motschulsky) similatut: (Casey) Tribe Ctenodactylini Genus LEPT( )TRACHELUS Latreille Sphaera':ra Say L. dorsalis (F.rbriciusj - ON Tribe Odacanthini Genus COLL. URIS DeGeer (Subgenus C()SNANIA Dejean) (.O dacan,, hella Liebke) C. pensylvanica (Linn6r ON PQ Tlibe Masoreini Genus TETR,,\GONODERUS Dejean T. lasciatus (llaldemanj ON undulatt'.s LeConte Tribe Lebiini Genus LEBIA. Latreille Key to Nor:,h American species: Madge (1967) (Subgenus LOXOPEZA Chaudoir) L. atriceps LeConte AB SK MB nanulinc: (Casey) L. atriventris Say - SK MB ON PQ enormis .Casey) L. grandis Hrntz - ON PQ magister (Casey) majuscui.a (Chaudoir) riuularis (Casey) L. tricolor Sa., - ON PQ (Subgenus LIMPRIAS Bonelli) L. divisa LeC rnte AB SK MB concinna LeConte (Subgenus LI)BIA s. str.) (Ap helog enia Chaudoir) 56 Carabidae (D ianchomena Chaudoir) (Metabola Chaudoir) L. analis Dejean ON anchora Chevrolat ap p endic ulata Chaudoir bonellii Putzeys (? ) s ubfr.g ur ata Motschulsky L. cyanipennis Dejean BC AB SK barbarae Casey montanaHorn melaenaHatch ruficollis LeConte L. fuscata Dejean BC MB ON PQ NB NS canonica Casey L. guttula LeConte BC metuens Casey pacifica Casey L. lobulata LeConte -oN (? ) b r unnicol lis Motschulsky L. moesta LeConte BC SK MB ON PQ NB NS NFLB cynica Casey L. ornata Say -oN PQ (?) apicalis (Haldeman) asheuillensis Casey axillaris Dejean brunnea Haldeman erythrocephala Dejean fl au iu entris Motschulsky fluuiatilis Casey frigida Chaudoir marginella Dejean nigripennis Dejean reperta Casey uirginica Casey L. perita Casey _BC L. pleuritica LeConte _SK ON L. pulchella Dejean AB SK MB ON rufopyga (Chaudoir) tahoensis Casey uiuida (Bates) L. pumila Dejean BC AB SK MB ON PQ NB NS frugalis Casey illini Casey ludouiciana Casey maculicornis LeConte quadrata Casey rhodopa Schwarz tertiaria Casey L. solea Hentz SK MB ON PQ NS (.? ) fl au olineata Motschulsky scapularis Dejean websteri Casey L. viridis Say YK NT BC SK MB ON PQ NB NS PE adolescens Casey bracata Casey castigata Casey cobaltina Casey (?) cyanella (Motschulsky) duluthiana Casey euoluta Casey histrica Casey incitata Casey magica Casey papago Casey planifera Casey prominens Casey smaragd.ula Dejean subaffinis Casey subopaca Schaeffer trajecta Casey 57 Carabidae truckeens s Casey uermiculi za Casey uiridis D< jean L. vittata (Fabricius) - BC AB SK MB ON PQ - NS amnicola Casey conjunget:s LeConte connecta tlChaudoir) debiliceps Casey depicta Horn flauouittata Chevrolat furcata LeConte sonomae l)asey spraguei (}{orlr) tempeana Casey Genus DROM. US Bonelli (Subgenus DRIMIUS s. str.) (Dinodrortius Casey) D. piceus Dejean - BC AB - MB ON PQ NB NS quadricollis LeConte Genus APRIS''IUS Chaudorr A. constrictus Jasey BC AB A. latens (LeCrntet - SK MB ON PQ - NS (.?) fuscipt nnis Motschulsky A. pugetanus l)asey - BC A. iubsulcatus (Dejean) - ON PQ NB NS cordicolli, ; (LeConte) Genus MICRC LESTES Schmidt-Goebel Blechrus Motschulskv Bomius L eConte M. brevilobus Lindroth - ON M. curtipennis (Casey) AB SK M. linearis (Lt,Conte) - BC AB SK MB ON PQ - NS anEustus (LeConte) prominul rs (Casey) M. major Lindroth - BC M. nigrinus (N'[annerheim) - BC fretus (.Ca,sey) obispinus (Casey) M. pusio (Leconle) - ON Genus SYNTC,MUS Hope M etable tt.. s Schmidt-Goebel s. americanus (Dejean) AK \T( NT BC AB sK MB ON PQ NB NS PE LB NF borealis ( 4immermann) Genus AXINOPAIPUS LeConte Variopalpis Solier A. biplagiatus tDejeant - BC AB SK - ON PQ Genus CALLtr.IDA Dejean (Subgenus CAI,LEIDA s. str.) (Spongoloba Chaudoir) C. punctata LeConte MB ON PQ C. purpurea tsiayt IIB cyanipennis Chaudoir smaragdina Dejean (Subgenus PHILOPHUGA Motschulsky) Key to species: Larson (1969) C. viridis amorna (LeConte) - BC AB SK MB canora (.Casey) obscura ( lasey) puella (C*ey) 58 Carabidae Genus TECNOPHILUS Chaudoir Key to species: Larson (1969) T. croceicollis peigani Larson AB SK Genus CYMINDIS Latreille (Subgenus PINACODERA Schaum) (Pla nes us Motschuisky) C. limbata Dejean - ON PQ - NS (?) fuscicollis (Motschulsky) (?) laeuigata (Motschulsky) C. platicollis tSayt - ON PQ fuscata Dejean uirescens Drrty (Subgenus CYMINDIS s. str.) C. americana Dejean - ON PQ continens Casey uenator Dejean C. borealis LeConte - BC AB SK MB ON PQ NB NS NF C. cribricollis Dejean - YK NT BC AB SK MB ON PQ NB NS - LB NF abstrusa LeConte acomana Casey alternans Casey alticola Casey coseyi Liebke hirbyi Casey marginata Kirby obliqua Casey planifera Casey reflexa LeConte rupimontis Casey sinuata Casey tardaLiebke C. interior Lindroth AB SK MB C. neglecta Haldeman AB SK MB ON PQ NB C. pilosa Say - ON PQ cribrata LeConte pubescens Dejean C. planipennis LeConte NT BC AB SK MB - PQ bipartita Casey b reu i p enni s Zimmermann directa Casey gouanica Casey C. unicolor Kirby AK YK NT BC AB - MB ON PQ - LB NF hudsonica LeConte parowana Casey Genus APENES LeConte (Subgenus APENES s. str.) (Nominus Motschulsky) A. sinuatus (Say) - ON pustulatus (Dejean) Tlibe Zuphiini Genus ZUPHIUM Latreille Z. americanum Dejean -oN Tfibe Galeritinr Genus GAJ-ERITA Fabricrus Galeritina Jeannel Galeritula Strand, Key to New World species: Reichardt (1967) (Subgenus PROGAIERITINA Jeannel) Key to species: BalI & Nimmo (1983) 59 Haliplidae G. janus (Fablicius) MB ON PQ ahasueru; Liebke americana (Fabricius) borealis Oastelnau caseyi Lel'g caseyi Lit:lske cordicollis Chaudoir cyanipeniis Dejean intermed o Casey occipitali.; Casey thoracica Casey Family HALIPLIDAE crawling water beetles R.E. Roughley Haliplidae is a family of aquatic beetles that dent invasion of the water (for alternative views, includes 3 7 species in Canada and Alaska. The see Kavanaugh 1986). There is no recetrt taxo- adults are distinguished among aquatic beetles by nomic revision of the North American fipecies. their enlarged hind coxal plates suspended below Matheson (1912) and Roberts (1913) provided the the abdominal sterna. Adult haliplids swim slowly last comprehensive treatments. Proper species or crawl o r submerged vegetation. Members of identification often requires examination of male Brychius are found in erosional portions of streams. genitalia. Therefore, previously widespread species Those of other genera use a wide variety of freshwa- are found to represent species complexes. A num- ter habitats but are most often found in permanent, ber of regional taxonomic works, such as Gordon clean, nutrient-rich ponds and lakes or in slow and Post (1965), Matta (1976), Brigham (1982), backwaters of streams. Larvae feed on periphyton Hilsenhoff and Brigham (1978), and Gunderson or algae. Adults are primarily algophagous but may and Otremba (1988), are useful for interpreting the be facultative scavengers. Canadian fauna. The hr'liplids have often been placed near Dytiscidae in classifications. This position is main- AK (3); YK (4); NT (9); BC (23); AB (15); liK (10); tained by lleutel and Roughley (1988), who, how- MB (13); ON (17); PQ (18); NB (3); NS (3); LB (2); ever, sugg(, st that haliplids represent an indepen- NF(4) . H(1) Genus BRYCI'IIUS C.G. Thomson Key to North American species: Carr (1928) B. albertanus Carr _AB SK MB B. hornii Crotr:h BC B. hungerfordi Spangler -oN B. pacificus Carr BC Genus HAJ-IPLUS Latreille Key to North Arnerican species: Wallis (1933) (Subgenus IL.aI-IPLUS s. str.) H. allisonae Brigham BC H. apicalis C.(i. Thomson* YI< Nr BC AB SK MB strigatus Roberts ^K H. blanchardi Roberts H. distinctus ''Vallis BC _-PQ H. dorsomacu, atus Zimmermann BC -NF H. falli Mank A]{ YK NT MB_PQ H. immaculicollis Harris AK }'K NT BC AB SK MB ON PQ NB \Tq DF LB NF americanus Aub6. impressus Kirby H. longulus LeConte NT BC AB MB ON PQ hoppingiWallis H. robertsi Zirnmermann BC pallidus loberts H. stagninus lL,eech BC AB SK MB (Subgenus PARAIIAPHLUS Wallis) 60 Noteridae H. borealis LeConte _ABSKMBONPQ H. connexus Matheson -MBONPQ NB NS H. fasciatus Aub6 ON PQ H. leopardus Roberts -PQ H. pantherinus Aub6 oN PQ H. triopsis Say ON PQ (Subgenus LIAPHLUS Guignot) H. canadensis Wallis _BCAB SK MB ON PQ H. columbiensis Wallis NT BC H. cribrarius LeConte -YKNTBCAB yu oN PQ NB _LB NF H. leechi carteri Leech _BC :" H. leechi leechi Wailis _BCAB H. nitens LeConte NT ON H. salinarius Wallis _BC H. salmo Wallis _BCAB H. subguttatus Crotch NT BC AB SK MB ON PQ _NS NF H. ungularis Wallis _BC H. vancouverensis Matheson _BCAB Genus PELTODYTES R6gimbart Cnemidotus Erichson P. callosus (LeConte) BC AB P. duodecimpunctatus (Say) oN PQ P. edentulus (LeConte) BC AB SK MB ON PQ P. lengi Roberts ON PQ P. muticus (LeConte) oN PQ P. torbulosus Roberts BC AB SK MB oN PQ Family AMPHIZOIDAE trout-stream beetles Y. Bousquet Amphizoidae is a small monogeneric family con- waterfalls. They apparently feed on living stonefly taining the genus Amphizoa, which has four nymphs. species: three occur in western North America and Kavanaugh (1986) recently revised the species one in the People's Republic of China, wesrern of this family. Szechwan Province. Both larvae and adults are found crawling over submerged rocks, debris, or wood in rivers, fast-flowing mountain streams, or aK (1); YK (2); BC (3); AB (2) Genus AMPHIZOA LeOonte Dysmalhes Mannerheim A. insolens LeConte AKYK_BC josephi Matthews sah I ber g ii (Mannerheim) A. lecontei Matthews YK_BC carinata Edwards planata Van Dyke A. striata Van Dyke BC Family NOTERIDAE burrowing water beetles R.E. Roughley Noteridae is a family of aquatic beetles that larvae of a European species burrow in the mud and includes only two species in Canada. Adults and take air from submerged plant tissue. Most North larvae are found in standing water and are associ- American larvae crawl on the substrate or pond ated with dense masses of aquatic vegetation. The vegetation. Adults are predaceous, and larvae may 61 Dytiscidae be omnivorous. A detailed study of the life history known (Beutel and Roughley 1987), and ther system- of any North American species is needed. atic position of the family near Dytiscidae is well Taxonomically, Nearctic noterids are well supported (Beutel and Roughley 1988, Kavanaugh known bet:ause all genera have been revised. No 1986). additional species are expected for the Canadian fauna. TlLe classification of noterids is fairly well oN (2);PQ (1) Genus HYDFi,OCANTHUS Say Key to Nevr World species: Young (1985) (Subgenus IIIDROCANTHUS s. str.) H. iricolor Say ON PQ similato- Zimmermann Genus SUPIJISELLUS Crotch Key to North American species: Young (1979a) S. puncticolli; (Crotch r - ON Family DYTISCIDAE predaceous diving beetles D.J. Larson and. R.E. Roughley In Car'ada and Alaska 262 species of Dytiscidae especially within the larger genera. Identifrcation are known, making this the largest family of of species may be difficult, as differences between aquatic beetles in the region. These beetles occur in species are often subtle. There is therefore a need a wide valiety of shallow-water habitats. The rich- for additional analysis to provide better species defi- est fauna is in the emergent vegetation zone along nitions and characters for their recognition. the edges of pools, small ponds, and small, slow Reviews and revisions of many genera hirrve been streams. .t\. number of species are characteristic of prepared or are in progress, but no single publica- cold-wate: habitats, such as small seepages and iion treats the entire fauna. The regional works of springs, as weII as bog, forest, alpine, and arctic Hatch (1953) and Larson (1975) together ccver most pools. Clear, gravelly streams and saline ponds of the western and boreal species. Wallis (1973) each contain distinctive groups ofspecies. and Larson and Colbo (1983) have publisled fairly Adults are predators or scavengers, and all lar- complete faunal lists for Manito ba and vae are apparently predaceous. The range of prey Newfoundland, respectively. Leech and Chandler taken probably is determined by what prey of suit- (1956) and D.S. White et al. (1984) provide good able size , s available, but some hydroporines feed keys to adults and larvae of most North l\merican largely olL microcrustacea, whereas some colym- genera. A number of works treating the fauna of betines prey mainly on culicid larvae. As far as is various states have been published in the United known or can be inferred, aII species in the fauna States. Some of these works are especially valuable are univo itine or semivoltine. Most species over- in interpreting the Canadian fauna: California winter as adults, but time of breeding and occur- (Leech and Chandler 1956), Florida (Young 1954), rence of larval stages vary greatly among species. Maine (Malcolm 1971), Nevada (La Rivers 1951), Represenuative larvae of most genera have been North Dakota (Gordon and Post 1965), North and described, but larvae are known for only a small South Carolina (Brigham 1982), and Utah (R.D. percentagr: of species. Anderson 7962). Zaitzev (1953) outlijned the The feLmily is relatively well known taxonomi- Palaearctic distribution of Holarctic species. cally. The principal taxonomic problems include relating the northern Nearctic fauna with that of the Palaezrrctic; analyzing and interpreting patterns AK (92); YK (111); NT (118); BC (167); AB {153); SK of geogral,hical variation, which in many species is (r24); MB (145); ON (155); pQ (147); NB (82); NS complex; aind developing suprasp ecifrc classifrcation, (77):PE (31);LB (7a); NF (78) ' H (45) Subfamily COPELATINAE TYibe Copelatini Genus COPELATUS Erichson Key to No:r'th American species: Young (1963o) 62 Dytiscidae C. chewolati Aub6 - ON renouatus Guiglot C. glyphicus (Say) - oN pe - NS pE - NF Subfamily AGABETINAE Tlibe Agabetini Genus AGABETES Crotch A. acuductus (Harris) - ON pe Subfamily LACCOPHILINAE T?ibe Laccophilini Genus LACCOPHILUS Leach Key to North American species: Zimmerman (1970) L. biguttatus Kirbyx AK YK NT BC AB SK MB ON pe NB LB inconspicuus Fall stroehmi C.G. Thomson L. maculosus Say - BC AB SK MB ON pe NB NS pE cal ifornic u s Motschulsky descipiens LeConte fusculus Sharp shermaniLeech trunco tus Mannerheim L. proximus Say - ON americanus Au.b6 confusus Sharp L. undatus Aub6 - ON Subfamily FfYDROPORINAE Tlibe Hydrovatini Genus HYDROVATUS Motschulsky Key to North American species: Young (1g56, 1g63b) H. pustulatus (Melsheimer) - ON pe compressus Shatp cuspidatus auct. amer. Tribe Methlini Genus CELINA Aub6 Key to North American species: Young (1979b) C. hubbelli Young - ON pe Tlibe Hyphydrini Genus DESMOPACHRIA tsabinston D. convexa (Aub6) NT BC AB SK MB ON PQ NB NS PE Tlibe Bidessini Key to genera: Young (1967) Genus BIDESSONOTUS R6gimbart Key to North American species: Young (1g54) (LeConte] B. inconspicuus - ON pe - NS B. pulicarius (Aub6) - ON Genus LIODESSUS Guignot Key to North American species: Larson and Roughley (1990) L. affinis (Say) AK yK NT BC AB SK MB ON pe NB NS pE NF charlotti (Clark) - emilianus (Clark) e ry t h r ostotn us (Mannerheim) 63 Dytiscidae macularis (LeOonte) microrei.iculatzs (Hatch) nanus (l\tb6) nigrinut (Casey) obscurellus (LeConte) L. cantralli (''loung) .AB SK MB L. fl avicollis r LeConte) _MBONPQ L. fuscatus (Crotch) SKMBONPQ_NS_LB Genus NEOCLYPEODYTES Young N. ornatellus (Fali) Genus IJVAIiiUS Guignot U. granarius (Aub6) MB ON PQ NB NS U. lacustris (Say) oN______U. subplicatt.s (Hatch) Tribe Hydroporini Genus HYGIIOTUS Stephens Coelamitus C.G. Thomson Key to North American species: Anderson (1971, 1976, 1983) H. acaroides lLeconte) MB winnipe,g (Wallis) H. bruesi (Fzll) AB H. compar (tr all) AK _BC AB SK MB ON PQ NB LB H. dissimilis Gemminger & Haroid _BC AB_MBONPQ dispar (. -eConte) H. falli tWallis ) - _ONPQ NF H. farctus (L rConte) ABSKMB_PQNB H. impressol unctatus (Schaller)x AK \-I( NT BC AB SK MB ON PQ NB NS PE LB NF similis (.Kjtby) H. infuscatur (Sharp) - YK NT BC AB SK MB dentiger (FalI) oregonu.; (FalI) H. intermedi -rs iFall,1 _BC H. laccophili rus t LeContet NT_ABSKMBONPQNBNS H. lutescens ileConte) AK _BC impress /rons Motschulsky infacetus (Clark) mediali:: (LeConte) H. marklini (Gyllenhal)x AK YK NT AB SK MB PQ NF canadensis (Fa11) uirgo (F:rll) H. masculinr s (Crotchl BC AB SK MB fastidiotus (Fa11') H. nigrescen r tFallt BC -pe H. novemlinc,atus hudsonicus (Fa11)* AK YK NT MB H. nubilus tI eConte) oN PQ NN NS H. obscurepl rgiatus t Fall t sc columbi znus (W.J. Brown) H. patruelis ileConte) AK YK NT BC SK MB ON PQ LB NF borealis (Fall) discoideus (LeConte) quebecensis (W.J. Brown) H. picatus (Iriirby) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF H. punctilintatus tFall t YK BC AB SK MB ON H. salinarius lWallist _NT _ABSKMB H. sal Balfour-Browne YK NT BC AB SK MB ON PQ NB NS PE LB NF punctatus (Say) H. sellatus (LeConteJ NT _ABSKMBON H. semivittarus tFall) BCAB_MB H. suturalis rl,eConte) AK YK NT BC AB SK MB ON PQ in lre ouoideur,; (LeConte) H. sylvanus ,Fallt ON PQ H. tumidiver tris (Fallt _NT BC AB SK MB ON H. turbidus (LeOonte) AK YK NT BC AB SK MB ON PQ NSpn-- H. unguicularis (Crotch)* AX YK NT BCABSKMB-PQ LB NF mongoli:us (Jakovlev) 64 Dytiscidae Genus FIYDROPORUS Clairville (Subgenus HETEROSTERNUTA Strand) (H eter o ster nu s Zimmermann) (H etero s teth u s Falkenstrom) Key to species: Matta and Wolfe (1981) H. alleghenianus Matta & Wolfe PQ NB H. cocheconis Fall PQ H. pulcher LeConte _ONPQNBNS-LBNF H. wickhami Zaitzev -oNPQNB concinnus LeConte (Subgenus SANFILIPPODYTES Franciscolo) H. browni Wallis BC H. brumalis W.J. Brown PQ H. compertus W.J. Brown BC AB H. edwardsi Wallis BC AB H. falsificus W.J. Brown BC H. pacil-rcus Fall BC H. planiusculus Fall BC AB sx y" oN pe LB NF H. terminalis Sharp BC s H. acadianus Balfour-Browne - H. persimilis Crotch - pq- N3 LB NF aequus Fall - - H. paugus Fall AK YK NT BC AB SK MB ON PQ NB NS LB NF H. rubyae Larson YK NT BC AB SK MB ON PQ NB LB NF DD H. stagnalis Gemminger & Harold _NT_AB SK MB ON PQ NB NS rD -NF collaris LeConte (Subgenus NEOPORUS Guignot) (C ir cinopo ru s Guignot) Key to species: Fa]] (1923) and Wolfe (1984) H. blanchardi Sherman _ONPQ H. carolinus Fall -oNPQNBNS-LBNF H. clypealis Sharp _ONPQNBNSNF anticus Shato H. dilatatus Fall NB H. dimidiatus Gemminger & Harold _AB semirufus LeConte H. mellitus LeConte ON PQ NB H. mixtus LeConte ON PQ H. solitarius Sharp SK MB ON PQ NB NS pn ie NF H. spurius LeConte ON PQ NB peltatus Shatp H. sulcipennis Fall _ONPQ NB H. superioris Balfour-Browne - Nr ec AB SK MB ON sericeus LeConte H. tennetum Wolfe -oN H. undulatus Say NT BC AB SK MB ON PQ NB NS PE LB NF anticus Sharp consimilis LeConte fasciatus Haris placatus Fall pubipennis Aub6 p un ctati s sirn u s Attb6. scrutator Sharp uelutinus Aub6 H. vittatus LeConte AB SK MB ON H. vitiosus LeConte oN PQ (Subgenus IIIaDROPORUS s. str.) Key to North American species: Fall (1923) H. appalachius Sherman YK NT BC AB SK MB ON PQ signatus Sharp H. badiellus Fall AK YK NT BC AB SK MB ON PQNBNS_LBNF H. brevicornis Fall PQNBNS_LBNF H. carri Larson AB H. columbianus Fall YK NT BC AB s* ms ON PQ NB 65 Dy'tiscidae H. dentellus Fall AK YK NT BC AB SK MB ON PQ NB NS PE LB H. despectus Sharp - YK NT BC AB - MB ON PQ - NS NF rusticut Sharp H. dichrous Melsheimer - ON PQ discicol',is Say H. fatigus G rrdon - BC H. fortis Le('onte - BC H. funestus Fall - BC - SK H. fuscipennis Schaumx AK YK NT BC AB SK MB ON PQ NB LB NF criniticrtxis Larson piceus ,cittrrn puberut'us Mannerheim H. lapponunL Gyllenhal AK YK NT BC MB ON PQ - LB NF labradc rensis Fall [L larsoni NLlsson - YK NT BC AB SK MB - PQ - LB lL longiusculus Gemminger & Harold AK BC oblongus Aub6 H. mannerhr:imi Balfour-Browne AK YK NT BC AB humerolis Alb6 H. melsheimeri Fall ON PQ H. morio Aur6* AK YK NT BC AB SK MB ON PQ - LB NF caligintsus LeConte me lanor:ep ha I us Gyllenhal H. nigellus l{annerheim* AK YK NT BC AB SK MB ON PQ NB LB NF H. niger Say - ON PQ NB NS PE - NF latifrons Sharp modestus Attb6 H. notabilis LeConte AK YK NT BC AB SK MB ON PQ NB NS PE LB NF arcticut C.G. Thomson H. obscurus Sturm* AK YK NT BC AB SK MB ON PQ NB LB NF hockin1i Larson H. occidentalis Sharp - YK - BC AB SK H. pervicinus Fall - BC AB SK MB H. polaris Fr,Ll1 AK YK NT MB - PQ subuiret;cens Fall H. puberulur LeContex AK YK NT BC AB - MB ON PQ NB LB NF leuanderi J. Sahlberg H. rectus Fall AK YK NT BC AB SK MB ON PQ NB LB NF H. rufinasus Mannerheim AK YK NT BC AB SK MB ON PQ - NS H. sibiricus,,I. Sahlbergx AK YK NT boreus llordon H. signatus \{annerheim AK YK NT BC AB SK MB ON PQ NB NS PE LB NF inornat'"ts Sharp youngi k,rdorl H. similaris Fall - BC H. sinuatipe* Fall - BC H. striola (Gyllenhal)x AK YK NT BC AB SK MB ON PQ NB NS - LB NF ambiguus Aub6 subtonsus LeConte uittula .Eic]l.son H. tademus Leech - BC AB H. tartaricus LeConte* AK YK NT BC AB - MB ON PQ cam ina., iu s Motschulsky lugubri: Motschulsky mel an c nolicu s Motschulsky H. tenebrosus LeConte AK YK NT BC AB SK MB ON PQ - NS NF luridiptnnis LeConte lutulen: us LeConte H. transpun-'tatus H.P. Chandler - BC AB H. tristis (Pr,rykull)* AK YK NT BC AB SK MB ON PQ NB NS - LB NF elongat:ilus Schiodte ru fi co p, ll u s Mannerheim uarians LeOonte Genus POTITIVIONECTES Zimmermann Deronectes auct. amer. P otarno dy t e s Zimmermann Key to North American species: Zimmerman and Smith (1975a) P. aequinoctialis (Clarkt AB P. deceptus Fallr - BC 66 Dy'tiscidae P. depressus t Fabricius)* NT_AB_MB P. elegans (Panzer)8 _BCABSKMBON P. griseostriatus (DeGeer)x AK YK NT BC AB SK MB ON PQ NB NS pn in Nr catascopium (Say) coloradensis (FalIt mathiasi (Hatch) parallelus (Say) suffusus (Sharp) P. rotundatus ( LeConte t ON PQ NB NS PE LB NF P. spenceri (Leech) - YK _BCABSKMB P. striatellus (LeConte) - YK NT BC AB SK MB infaustus (Clark) pulcher (Motschulsky) Genus OREODYTES Seidlitz Key to North American species: Zimmerman (1985) O. alaskanus (Fall) AK YK BC AB p r o d u c t o t r u n c at u s (Hatch) rainieri (Hatch) recticollis (Fall) O. angustior (Hatcht _BC O. congruus (LeContet _BCAB O. crassulus iFall; _BCAB O. laevis (Kirby) AK YK NT BC AB MB ON PQ duodecimlinealzs (LeConte) -iuNu hortensis (Hatch) semiclarus (.Fall) yukonensis (Fall) O. leechi Zimmerman AK YK O. obesus cordillerensis Larson - YK _BCAB O. picturatus (Horn) _BC O. quadrimaculatus (Horn) _BCAB brodei (.Gellerrnan) O. sanmarkii (C.R. Sahlberg)* - YK NT riuolis (Gyllenhal) O. scitulus {LeConte) NT BC AB SK MB ON PQ NB NS NF bisulcatus (Fa],\) O. snoqualmie (Hatchl BC AB Genus LACCORNIS des Gozis Agaporus Zimmermann Key to species: Wolfe and Roughley (1990) L. conoideus (LeConte) - YK NT BC AB SK MB ON PQ NB _NF L. deltoides (Fall) ON L. latens (Fall) ON PQ NB L. oblongus t Stephens tx NT L. pacificus Leech BC AB Subfamily COLYMBETINAE Tribe Agabini Key to North American genera: Leech (1942) Genus AGABINUS Crotcl' Key to species: Leech (1941) A. glabrellus (Motschulsky) _BC A. sculpturellus Zimmermann _BC Genus AGABUS Leach Allonychus Zaitzev Anisornera LeConte Apator Semenov Arctodytes C.G. Thomson Gaurodytes C.G. Thomson Heteronychus Seidlitz Ilybiosoma C'rotch Key to North American species: Fa\l (1922b), Larson (1989) A. adpressus Aub6* AK YK NT BC solas Leech 67 Dytiscidae A. aeruginosrs Aub6 -oN disposit'ts Guignot A. ajax Fal1 YK NT BC AB SK MB ON PQ LB fuscipennis auct. amer. A. ambiguus (Say) YK _BC AB SK MB ON PQ NB NS PE LB NF fimbriatus LeConte reticulaLus A,e.b6 A. antennatus Leech AK YK NT BC AB SK MB ON PQ LB clauatus LeConte A. anthracin:rs Mannerherm AK YK NT BC AB SK MB ON PQ NB NS LB NF scapuloris Mannerheim A. approximatus FaIl NT BC AB SK A. arcticus (['aykull)x AK YK NT BC AB SK ms oN pe LB NF reticulaius (Kirby) subfasctttus LeConte A. audeni Wl1lis AK YK NT BC AB SK MB A. austinii S.rarp AK YK NT BC AB A. bicolor (K:rby) AK YK NT BC AB SK MB oN- NS- mutus S.harp A. bifarius (F.irby)* AK NTYK BC AB SK MB oN PQ NB NS LB hessleri Hochhuth A. bjorkmanare Hatch BC AB rectus (LeConte) A. canadensir Fall YK_BCABSK MB A. clavicorniu Sharp* AK YKNTBC_SK MB uerzs W,J. Brown A. col)rynbus :eech YK NT MB A. confertus ,eOonte BC A. confrnis (( iyllenhal)* AK YK NT BC AB SK MB oN pe - NS- LD NE longulu: (LeConte) ouoideut (Crotch) A. coxaiis Shrrpx AK YK NT BC A. discolor (F.arris) AK YK NT BC AB SK MB ON PQ NB NS PE ie- A. discors Lelonte _BC A. disintegrabus (Crotch) -oN PQ A. elongatus (Gyllenhal)* AK YK NT BC AB SX MB bryanti Jan A. erichsoni tlemminger & Haroldx AK YK NT BC AB SK MB ON PQ NB NS NF lutosus t Crotch) nigroaer,,eus Erichson sobrinus Motschulsky A. erythropterus (Say) ON PQ NB NS NF A. falli (Zimrnermann) AB SK MB oN PQ semtpunctatus auct. amer. sharpi F all A. fuscipenni.s (Paykull)* YKAK NT BC AB SK MB ON ontarior,,is FalI A. gagates Arrb6 -oN PQ NB NS A. hypomelar; Mannerheim AK BC AB i rregula 'is Mannerheim A. infuscatus Aub6 AKYKNTBCAB_MBONPQ _LBNF A. inscriptus (Crotch) AK YK NT BC AB SK MB ON PQ _NS _LBNF A. leptapsis (LeConte) _YK_BCABSKMBONPQ NB NS _LBNF A. lugens LeConte -BC perplexus Sharp suturalis (Crotch) A. lutosus LeConte BC AB SK discolor LeConte lecontei lCrotch) mimusLeech A. margaretae Larson AB SK A. moestus (Ourtis)* AK YK NT _MBONPQ borealis Sharp nigripalcis (J. Sahlberg) A. nectris Leech -BC AB SK A. oblongulus Fall _BC A. obtusatus (Say) oN PQ f.fn NS nitidus lSay) A. opacus Aub6* AK \'K NT BC AB SK MB ON PQ NB LB NF gelidus )!a71 pseudoc' tn fertus Wallis sharpi Jacobson 68 Dytiscidae A. phaeopterus (Kirby) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF A. pisobius Leech _BCAB A. punctulatus Aub6 _YK AB SK MB ON PQ NB LB aeneolus (Crotch) A. semipunctatus (Kirby) AK YK NT BC AB SK MB ON PQ NB NS LB NF stridulator Shatp A. semivittatus LeConte ON PQ spilotus LeConte texanus Sharp A. seriatus (Say) YK NT BC AB SK MB ON PQ NB NS NF arcfzs Melsheimer intersectus Ctotch parallelus LeConte striatus Aub6, A. smithi W.J. Brown _BCAB A. strigulosus (Crotch) AK YK NT BC AB SK MB nanus (.LeConte) A. subfuscatus Sharp MB ON PQ NB NS PE A. thomsoni (J. Sahlberg)* AKYKNTBCABSKMB-PQ LB NF A. tristis Aub6x AK YK NT BC AB SK PQ LB NF atrdtus Mannerheim crotchi Zaitzev dubizs Mannerheim piceus Zaitzev piceolus Zaitzev subopocus Mannerheim A. triton Fall SK MB A. vancouverensis Leech AK_-SCT A. velox Leech AK YK NT _MR A. verisimilis W.J. Brown _BCAB A. wasastjernae (C.R. Sahlberg)* AK YK NT BC AB SK MB ON pe- _LBNF henaiensis Fall palustris Wallis A. zattzewi Poppius* AK YK NT MB hudsonicus Leech A. zetterstedti C.G. Thomson* AK YK NT MB browni Leech Genus CARRI{YDRUS Fall C. crassipes Fall YK NT BC SK MB ON Genus ILYBIUS Erichson Key to North American species: Larson (1987) I. angustior (Gylienhal)* AK NB NS PE LB NF I. biguttulus (Germar) YNTBCYSKMB3NiS NB NS NF laramaeus LeConte L churchillensis Wallis _MB I. confusus Aub6 f_ I'_ ON NS deniheiWallis I. discedens Sharp AK YK NT BC AB SK MB ON PQ NB NS _LBNF L fraterculus (LeConte) I. ignarus (LeConte) --NTBCY:"YuoNpe NS __I" I. incarinatus Zimmermann -oNPQ I. oblitus Sharp I. picipes (Kirby)* AK Nr sc AB !x nrn 3N pe -isNr L pleuriticus (LeConte) _ \'K"K NT BC AB SK MB ON PQ NB NS PE LB NF inuersus Sharp I. quadrimaculatus Aub6 AK BC AB I. subaeneus (Erichson)* AK YK NT BC AB SK MB ON PQ LB NF uiridianeus Crctch I. vittiger (Gyllenhal)* AK YK NT BC MB Tribe Colymbetini Genus COLYMBETES Clairville Cymatopterus Dejean Key to North American species: Zimmerman (1981) C. dahuricus Aub6x AK YK NT BC AB SK MB ON PQ obscuratus Mannerheim seminiger LeConte 69 Dytiscidae C. densus LeConte AK YK - BC AB SK MB inaequalLs Horn C. doiabratus (Paykull)* AK YK NT BC MB - PQ - LB NF drewsonl. LeConte groenlanCicus At:.b6 margina'.is (Fabricius) thomsont. Sharp triseriatu.s I{trby C. exaratus LeConte - YK NT BC AB SK MB i ncog nitr"r s Zimmerman rugipenn.is Sharp C. paykulli Erichsonx - YK NT BC AB SK MB ON PQ NB NS NF longulus LeConte C. sculptilis Ftarris AK YK NT BC AB SK MB ON PQ NB NS PE LB NF Genus NEOS'IUTOPTERUS Balfour-Browne P seudo sc utop te r u s Hatch Key to species: Larson (1975) N. angustus (LeConte) - YK - BC AB SK MB ON PQ NB NS - LB NF N. hornii (Crc,tch) AK YK NT BC AB SK MB ON PQ NB LB coriaceut: Hotn Genus RHANTUS Dejean Key to Nor',h American species: Zimmerrnan and Smith (1975b) (Subgenus NI,\RTUS Zaitzev) (Rantogilon des Gozis) R. sinuatus (l,eConte) NT BC AB SK MB ON PQ - NS (Subgenus RIIANTUS s. str.) R. binotatus (Harris) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF aequalis ll.atc]r assimilit (Kirby) diuiszs ( {ub6) flauogriseus Crotch longipes Sharp obscurus, Shatp plebeius Shatp R. consimilis Motschulsky NT BC AB SK MB ON PQ disceden; Shatp loslzs (LeConte) R. frontalis (l{arsham)* AK YK NT BC AB SK MB ON PQ fla u icoll t s E schscholtz gyllenhat i (Castelnau) notatus (Fabricius) punctatus (Hope) sericozs Sharp suturalk (Boisduval & Lacordaire) R. gutticollis Sayt - BC AB dominguensis Gemminger & Harold hoppingt.Wallis hubbelliHatch maculiccllis (Arb6) mexican, t s (Castelnau) R. suturellus (Harris) AK YK NT BC AB SK MB ON PQ - NS - LB NF subopacus (Mannerheim) zimmermanni WalTis R. wallisi Hatch AK YK NT BC AB SK MB ON PQ NB NS - LB NF suturellus anct. Tribe Coptotomini Genus COPTJTOMUS Say Key to eastern North American species: Hilsenhoff(1980) C. longulus LeConte - BC AB SK MB ON PQ NB NS PE lenticus Hilsenhoff C. loticus Hil;enhoff - ON PQ 70 Dytiscidae Tlibe Matini Genus MATUS Aub6 Key to species: Young (1953) M. bicarinatus (Say) ON M. ovatus ovatus Leech ON PQ Subfamily DYTISCINAE TYibe Dytiscini Genus DYTISCUS Linn6 Leionotus l\trby Macrodytes C.G. Thomson Key to species: Roughley (1990) D. alaskanus Balfour-Browne AK YK NT BC AB SK MB ON PQ LB NF paruulus Motschulsky D. circumcinctus Ahrens* AK YK NT BC AB SK MB ON PQ arrrizs Mannerheim ooligbukiilirby D. cordieri Aub6 AK NT BC AB SK MB ON PQ NB sublimbatus LeConte D. dauricus Gebler* AK YK NT BC AB SK MB ON PQ NB NS NFLBPE confluens Say franklinii l{trby frontalis Motschulsky uentralis Motschulsky uexatus Sharp D. fasciventris Say _YKNTBCABSK MB ONPQNBNS_LB D. harrisii Kirby AK YK NT BC AB SK MB ONPQNBNS_LB NF D. hatchi WaIIis -BCABsx D. hybridus Aub6 MB ON PQ NB compar (Melsheimer) D. marginicollis LeConte NT BC SKAB MB alb ioni c u s Motschulsky fu scostr iat us Motschulsky D. verticalis Say MB ON PQ NB NS PE Tlibe Hydaticini Genus HYDATICUS Leach Key to North American species: Roughley and Pengelly (1982) (Subgenus I{YDATICUS s. str.) H. aruspex Clark* AK YK NT BC AB SK MB ON PQ NB NS PE NFLB americanus Sharp laeuipennis C.G. Thomson modestus S}:'atp H. oiceus LeConte AB SK MB ON PQ NS Tribe Aciliini Genus ACILIUS Leach Key to North American species: Hilsenhoff(1975) A. abbreviatus Mannerheim AK _BCAB oregonensis Ctotch A. athabascae Larson MB ON PQ A. mediatus (Say) _ ICT:" -oN PQ NB NS maccullochii (Kirby) A. semisulcatus Aub6 AK YK y:'T:" MB ON PQ NB NS PE LB NF A. sylvanus Hilsenhoff MB ON PQ Genus GRAPHODERUS Dejean Key to North American species: Wallis (1939) G. fascicollis (Harris) oN PQ 7T Gytininae G. liberus (Sr,Ly) NT BC SK MB ON PQ NB NS NF brunnipznnis Aub6 rugicolli.s I{ttby thoracic ^ts Haris G. manitober.sis Wallis G. occidentalis Horn _YKN,scABs^MBoNpe G. perplexus Sharp AK YK NT BC AB SK MB ON PQ NB Ns po ie Nr elatus S tarp Genus THERMONECTUS Dejean T. basilaris (]{arris) -oN Tribe Cybistrini Genus CYBIfITER Curtis C. frmbriolatrrs (Say) MB ON NS crotchi \{ilke oliuieri (.)totc}' Family GYRINIDAE whirligig beetles R.E. Roughley Adult lyrinids form a group of beetles wonder- Dineutus; few changes have been required since fully adalrted for life on the water surface; 36 that publication. Fall's work (1922a) is the lastest species ocsur in Canada and Alaska in streams, revision of the North American species of G'yriruus; a rivers, pords, and lakes. They can be found quite taxonomic revision of Nearctic Gyrin zs is in far from s rore but are more common closer to the progress and should help ease the diffrculty of iden- shore. Thc adults are readily distinguished by their tifying specimens in this genus. Morrisette (1979) highly mo<1ifred, short and flat middle and hind legs provides good regional works on gyrinids for and their horizontally separated eyes. Adults glide Quebec, Gordon and Post (1965) for North Dakota, on the surlace of water, cluster together, and when Sanderson (1982) for North and South Carolina, disturbed lnove rapidly in circles and scatter widely and Ferkinhoff and Gundersen (1983) for Minnesota or dive. Such aggregations often include a number and environs. The general classification of ryrinids of species, Adults produce defensive secretions, is now well understood (Folkerts 1979). Beutel and which discourage predation by aquatic vertebrates, Roughley (1988) suggest that gyrinids are l;he most and feed on dead and dying insects trapped on the primitive family of the suborder Adephaga end that water surf;lce. Larvae are aquatic and respire cuta- they represent a very early and independent use of neously wibh the aid of lateral abdominal filaments. aquatic habitats by beetles (see also Kavanaugh Pupation occurs on emergent plants along the 1986). shoreline. There is no recent taxonomic treatment of the aK (7); YK (10); NT (14); BC (18);AB (21); SK (23); North American species of gyrinids. Roberts (1895) MB (22); ON (27); PQ (30); NB (14);xS (101; pE (5); prepared the latest revision of the species of LB (8);NF (10) . H (4) Subfamily GYRININAE Key to North American genera: Folkerts (1979) Tlibe Enhvdrini Genus DINEIJTUS Macleay Cyclinus Kirby D. assimilis (.iGrby) BC SKAB MB ON PQ PE americanus (Say) D. discolor ArLb6 -oN PQ NS labratus (Melsheimer) D. horni Roberts _SK MB ON D. nigrior Rolrerts MB ON PQ 72 Gyrininae Tribe Gyrinini Genus GYRINUS O.F. Miiller (Subgenus GYRINULUS Zaitsev) G. minutus Fabricius* AK YK NT BC SK MB ON PQ NB NS (Subgenus GYRINUS s. str.) G. aeneolus LeConte SKAB MB ON PQ NB G. aeratus Stephensx AX YK NT BC AB -oN PQ instabilis Fall G. affrnis Aub6 NT BC AB SK MB ON PQ NB NS LB NF canadensis R6gimbart G. analis Say MB ON PQ _NS G. aquiris LeConte AB_ MB ON PQ NB PE NT G. bifarius Fall NT BC AB SK MB ON PQ NB NS LB NF G. cavatus Atton -YK NT BC AB MB -NS G. confinis LeConte NT BC AB SK MB ON PQ NB NS LB NF G. consobrinus LeConte BC AB SK G. dichrous LeConte _SK MB ON PQ G. dubius Wallis _SK MB PQ NF G. fraternus Couper MB ON PQ -NS PE :u G. frosti Fall _:* PQ G. gehringi Chamberlain -oN PQ NF G. hoppingi Leech NT BC AB SK G. impressicollis Kirby mn on PQ G. latilimbus Fall _YK BC AB SK -oN PQ NN NS LB NF G. lecontei Fall MB ON PQ pe G. maculiventris LeConte NT BC AB ;K yu oN PQ Tu_ G. marinus Gyllenhalx AK YK G. marginellus FalI PQ NB G. opacus C.R. Sahlberg* AK YK NT BC AB SK MB ON PQ LB NF hoppingiLeech G. pectoralis LeConte _YK NT BC AB SK MB ON PQ NB G. piceolus Blatchley PQ G. picipes Aub6 AK YK NT BC AB SK MB ON PQ G. pleuralis Fall AK YK NT BC ABSK_ON PQ lln - G. plicifer LeConte BC AB G. pugionis Fall _SKMBON PQ NB NS G. punctellus Ochs BC AB SK G. sayi Aub6 _YK NT BC AB SK MB ON PQ NB NS pt LB NF lugens LeConte G. ventralis Kirby NT ABSK_ON PQ NB- G. wallisi Fail AK YK NT BC AB SK MB ON PQ 73 Hydraenidae Suborder Polyphaga Superfamily Staphylinoidea Family FIYDRAENIDAE minute moss beetles R.E. Roughley Twenty.ssyen species of Hydraenidae are found such as wing venation and the absence of tire basal in Canada and Alaska. The family includes minute piece on the aedeagus, leave little doubt about its aquatic beeitles, most of which are under 2 mm long. place in the superfamily Staphylinoidea. Perkins Adults are found along the edges of aquatic habi- (1980) recently revised the Canadian and .Alaskan tats, particularly streams; some also live in semi- species in his monograph of the Hydraenidae of the aquatic habitats. Adults and larvae feed on peri- Western Hemisphere. phy'ton. The farnily is sometimes assigned to the super- NT (a); BC (21);AB (10); SK (4); UB (5); ON (5);PQ family Hyclrophiloidea; however, several features, (7);NB (3);NS (1);NF (1) Subfamily IIYDRAENINAE Tribe Hydraenini Genus HYDR\XNA Kugelann H. anguiicollil Notman NT_AB-MB ON PQ NB NF H. atlantica F erkins PQ NS H. circulata Perkins _BCAB__ IU H. nigra Hatcr -BC H. occidentali r Perkins _BC H. pacifica Pe.:kins _BCAB H. pennsylvar.ica Kiesenwetter ON PQ_ H. petila Perk ins _BC H. vandykei d'Orchynont _BC Genus LIMNEBIUS Leach Limnocharis Horn L. alutaceus (Casey) _BCAB L. borealis Pelkins _BC Subfamily OCHTHEBIINAE Genus GYMNOCHTHEBIUS d'Orchwnont G. nitidus (Le,lonte) NT oN PQ Genus OCHTII{EBIUS Leach (Subgenus OCHTHEBIUS s. str.) (Enicocerus Stephens) O. benefossus LeConte oN PQ O. bisinuatus Perkins BC AB__ O. borealis Pe:rkins BC O. intermptus LeOonte BC aberti }Ia.tc}r O. kaszabi Janssens NT BC AB SK MB ON PQ NB O. lecontei Perkins BC O. lineatus LeConte BC AB SK MB milleriHatch O. marinus PnLykull BC AB SK MB holmbergi Maklin O. rectus LeConte BC (Subgenus ASIOBATES C.G. Thomson) (H omaloc ht hebizs Kuwert) .(Trymoch thebizs Kuwert) O. brevipennir; Perkins BC 74 Ptiliidae O. cribricollis LeConte _BCABSK MB_ PQ O. discretus LeConte _BC O. hibernus Perkins NT O. mimicus W.J. Brown _BCAB Genus NEOCHTHEBIUS d'Orchymont N. vandykei (Knisch) lapid.icola (Van Dyke) Family PTILIIDAE feather-winged beetles J.M. Campbell Members of Ptiliidae, a family of very small bee- will be found, particularly in eastern Canada. tles, have fringed wings with long filaments, which Hatch (1957) studied the western species. Bistrom give them a feather-like appearance. The family and Silfverberg (1979) examined many of the comprises about 560 species around the world, of generic type species and frxed the generic names. which 32 are known to occur in Canada and Alaska. Dybas and Dybas (1990) recently published a key to Most feather-winged beetles are found in associa- the North American genera. Many Canadian tion with moist organic matter, such as leaf litter, records require confrrmation, as the identification of decaying wood, and fungi. most species has not been checked by specialists. Ptiliidae is probably the most poorly known family of beetles in North America. Undoubtedly, aK (5);BC (16);MB (1); oN (2); PQ (10); NB (5); NS when the family is revised, many additional species (1) . H(1);I(11) Subfamily PTILIINAE Genus MOTSCHULSKIUM Matthews M. sinuatocolle Matthews BC Genus PTENIDIUM Erichson P. nitidum (Heer)t NB P. pullum Maklin _BC P. pusillum (Gyllenhal)t _BCPQ _NS apicale Erichson terminale Holdeman Genus ACTIDIUM Matthews A. crotchianum Matthews BC Genus PTILIOLA Haldeman P. brevicollis (Matthews)t PQ P. collani Mannerheim -oN canadensis LeConte Genus PTILIOLUM Flach P. columbianum (Matthews) _BC P. obscurum (Matthews) _BC P. sharpi (Matthews) _BC P. spencei (A]libert)t PQ Genus PTILIUM Gyllenhal Millidium Motschulsky P. minutissimum (Ljungh)t PQ 75 Limuiodidae Subfamily ACROTRICHIINAE Genus NEPHANES C.G. Thomson N. titan (Nev'man)f PQ Genus PTILI IPYCNA Casey P. moerens (ldatthews) MB ON PQ quadratr.t (Casey) Genus ACROTRICHIS Motschulsky Macdonuldium Abdullah & Abdullah Trichopteryx Kirby & Spence A. castanea (lvlatthews) BC A. cognata (Niatthews)f BC Ne- platonoffi Renkonen pruinosc Hatdn A. dilfinis (Mrtthews) BC A. fasciculari ; (Herbst)i NB A. fungi (Abdullah & Abdullah) PQ A. grandicolli s (Mannerheim)f PQ A. henrici (M,rtthews) BC fratercuia (Matthews) A. insularis (.v{aklin) AK BC A. intermediz (Gillmeister) AK A. josephi (M,rtthews) BC A. laticollis N aklin AK A. parallelopi peda (Matthews) BC A. sericans (Iteer)i PQ NB A. sitkaensis (Motschulsky)* AK A. thoracica ( Waltl)t PQ anthracina (Matthews) A. vicina (Ma bthews) BC A. volans (Mctschulsky)i NB fennica l:lenkonen A. xanthocer:. (Matthews) BC Family LIMULODIDAE horse-shoe crab beetles J.M. Campbell Members of this small family are obligate reported in Canada, but one species of Litnulodes, guests of ants. The family comprises about 30 which I have tentatively identifred as L. pa,radoxus, species distributed in North and South America and occurs in southern Ontario. Seevers antl Dybas Australia, lnly one of which is found in Canada. (1943) revised the classification of the farnily and Recent workers have placed the limulodids in included a key to all genera and most North the family Ptiliidae (e.g. Lawrence 1982o), but I American species. prefer to nraintain the group as a separate family, on basis of papers by Seevers and Dybas (1943) and Dybas (19?6). This family has not been previously oN (1) Genus LIMU -ODES Matthews (Subgenus LIMULODES s. str.) L. paradoxus Matthews ON 76 Leiodidae Family AGYRTIDAE agyrtid beetles J.M. Campbell Eleven species of agyrtids are known in However, these studies have shown that agyrtids America north of Mexico, of which eight are found are more closely related to species of the family in Canada and Alaska. Adults are scavengers of Leiodidae than to Silphidae and therefore must be decaying organic material. Larval feeding habits treated as a separate family. The North American are probably similar to those of the adults species are well known taxonomically. The species (Anderson and Peck 1985). of Canada and Alaska were treated bv Anderson Until recent papers by Lawrence (1982o) and and Peck (1985). Lawrence and Newton (1982), species of this family had been included in the family Silphidae. AK (4); NT (1); BC (6);AB (3); ON (1) Genus NECROPHILUS Latreille N. hydrophiloides Gu6rin-M6neville AK BC a/er Motschulsky N. pettiti Horn ON Genus IPELATES Reitter Pelates }Iotn Pelatines Cockerell I. latus (Mannerheim) AK BC Genus AGYRTES Frrtlich (Subgenus AGYRTECANUS Reitter) A. longulus (LeConte) AK BC politus (Casey) Genus LYROSOMA Mannerheim L. opacum Mannerheim AK Genus PTEROLOMA Gyllenhal P. nebrioides W.J. Brown Genus APTEROLOMA Hatch A. caraboides (Fall) _BC A. tenuicorne (LeConte) _BCAB Family LEIODIDAE small scavenger beetles S.B. Peclz In Canada and Alaska 128 species of Leiodidae various decaying organic materials in usually moist are known. Those grouped here have been placed forest or freld habitats. The two biggest subfamilies under many family names, but all share the derived are Leiodinae and Cholevinae. Members of the (but diffrcult to see) character of a nearly enclosed Leiodinae subfamily (= Anisotomidae, Agathidiidae), gutter that circles the antennal articulation on the or round fungus beetles, are often globose and com- dorsal face of segments 7,9, and 10 or 8-10 in 11- pact in body shape, and many species can contract segmented antennae and segments 7-9 in 10-seg- into a ball-like form. Some species are specialized mented antennae, with internal sensory vesicles in for digging in soil, where they feed on subterranean some segments. The higher classification here lists fungi; others are specialized predators of slime five subfamilies and follows recent advances molds; and still others feed on soft fungi or are more reported by Lawrence (1982o), Lawrence and generally saprophagous (Wheeler 1984). The feed- Newton (1982), and Newton (1984). ing habits of both adult and larval Coloninae (= Members of Leiodidae are, for the most part, Colonidae) are not known. Members of the scavengers, both as larvae and adults, feeding on Cholevinae subfamily (= Catopidae, Leptodiridae), 77 Leiodidae or small carrion beetles, are more elliptical in body The leiodid fauna of Canada and Alaska is only shape and feed on carrion or dung. Some species are partly known. Several genera have been reviewed specializecl occupants of ant nests, caves (but not in and revised recently, and others are being prepared. Canada), rr the nests and burrows of vertebrates. No single publication treats the entire fauna, but a Members irf the Leptininae family (= Leptinidae), or generic review of the North American fauna is mammal rrest beetles and beaver beetles, are nest available (Peck 1990). The largest faunal works scavengers and ectoparasites on small mammal (Hatch 1933, 1957; Jeannel 1936) are no longer hosts or on beavers. completely reliable. Distributions and bionomics of Collec;ing Leiodidae usually requires a concen- the species are usually incompletely known, and tration on the particular habits or habitats of the many species in the northern United States may genera or species sought. They can be collected in eventually be found in southern Canada. the following ways: in baited pitfall traps; by sifting and Berlese extraction of litter or fungi; by sweeping AK (36);YK (18); NT (16); BC (84);AB (39)t SK (18); forest vegc,tation on still, warm, and humid evenings; MB (26); oN (57); PQ (53); NB (1e);NS (15); t B (7); and by use,, of large-area flight intercept traps. NF(10) . H(6);I(1) Subfamily LEIODINAE Tribe Hydnobiini Genus HYDIIOBIUS Schmidt H. contortus Hatch -BC H. crestonensis Hatch _BCAB H. femoratus Hatch BC H. longulus l,eConte _BC longidets LeConte H. luggeri H rtch AK H. matthewsii Crotch --ec-- H. obtusus L,eConte SK H. pumilus L,eConte ON PQ hiseri Hatch latidens LeConte H. simulator W.J. Brown _BC H. stacesmit ri Hatch _BC H. strigilatu r Horn _BC H. substriatrLs LeOonte AB pe NS curuid.ens LeConte H. validus \[.J. Brown PQ Genus TRIAIRTHRON Markel T. lecontei Horn _BC cedonuli'.i Schaufuss Tribe Leiodini Genus ANO(]DUS LeConte Neocyrtlsa W.J. Brown Key to species: Daffner (1988) A. fusciclavus (Fall t BC sternitu; (Hatch) A. insolitus tW.J. Brownt BC SK A. obsoletus Melsheimer) oN-- blandis :; ima (Zimmermann) A. potens tW.J. Brownt BC _ONPQ A. puritanus {Fallt _ONPQ A. secretus (W.J. Brown) -oNPQ A. superans Fall) AB_MBON Genus LIONOTHUS W.J. Brown Key to No;th American species: Daffner (1988) L. forticornis Daffner -oN L. ulkei W.J Brown -oN PQ 78 Leiodidae Genus CYRTUSA Erichson Caenocyrta W.J. Brown Key to North American species: Daffner (1988) C. subtestacea (Gyllenhal)* BC oN PQ picipennis (LeConte) Genus LIOCYRTUSA Daffner Key to North American species: Daffner (1988) L. luggeri (Hatch) _BCABSKMBONPQNB L. nigriclavis (Hlisnikovsky)* _BC_ONPQ Genus ECARINOSPHAERULA Hatch E. ecarinata (Horn) BC Genus ISOPLASTUS Horn I. fossor Horn oN PQ uncus Wheeler Genus LEIODES Latreille L. assimilis (LeConte) AB SK MB ON PQ NB NS LB NF L. bradorata W.J. Brown yyNrBC PQ L. canadensis (W.J. Brown) SK MB L. collaris (LeConte) AKYK_BC AB MB ON PQ NB LB NF L. conferta (LeConte) L. curvata (Mannerheim) AK BC AB _ 9*PQ morula LeConte L. diffrcilis (Horn) L. dilatata Hatch - - - ncy L. grossa Hatch -BCAB L. impersonata W.J. Brown -oNpe NB NS L. lateritia (Mannerheim) AKBC_ L. lostine (Hatch) _BCAB L. merkeliana (Horn) NT BC L. mirabilis (Hatch) _BC L. paludicola (Crotch) -BC L. puncticollis (C.G. Thomson) AKYK_BCAB SK nfn ON Pq L. punctostriata Kirby AK YK NT BC AB MB ON PQ NB NS LB NF indistincta (LeConte) laefo (Mannerheim) punctatostriato Horn punctistriata LeConte L. punctulata (GyIlenhal) AKYKNTBCAB_MBONPQ L. rhaetica (Erichson) AK L. rufrpes (Gebler) AK YK NT BC AB SK MB ON L. serripes Hatch _BCABSK L. strigata (LeConte) AKBCABSK-ONPQ NB L. tenuis Hatch _BC L. triepkei (Schmidt) -YKNT-AB-MBONPQ NB L. valida (Horn) NTBCAB_MBONPQ NB Genus ZEADOLOPUS Broun Apheloplastus W.J. Brown Key to North American species: Daffer (1988) Z. egenus (LeConte) oN PQ impubis (Zimmermann) Tlibe Scotocryptini Genus AGLYPTINUS Cockerell A. laevis (LeConte) oN PQ Tribe Pseudoliodini Genus COLENIS Erichson C. impunctata LeConte BC MB ON PQ NB NS 79 Leiodidae Tribe Agathidini Genus ANIS()TOMA Panzer Key to North American species Wheeler (1979) A. amicaW.J. Brown BC oN PQ A. basalis (Lc,Conte) _ONPQ_NS dichroa .LeConte) A. blanchard' (Horn) PQ A. confusa (Horn) BC AB _9* interstrialis Hatch A. discolor (l\'lelsheimer) BC -oN PQ piceo (Melsheimer) A. errans W..1. Brown AK BC AB SK -oN PQ NS A. expolita S .J. Brown -oN polita (LeConte) A. geminata r.Horn) -oN PQ A. globososa {atch BC AB_ MB ON PQ globosa LeConte) A. horni Whe,eler -oN PQ NB A. humeralis (Fabricius)i BC A. inops W.J. Brown -oN PQ NB A. nevadensi:i W.J. Brown BC MB bicolor (..Jorn) Genus AGAT'HIDIUM P anzer Key to North American species: FaIl (1933) A. alutaceum Fa11 AK A. athabasca:rum Fali AB_ A. angulare Idannerheim AK _BC A. californicu.m Horn _BC A. cavisternum Fa1l -BC A. columbianum Fall _BC A. concinnuni Mannerheim AK -BC A. conjuncturn W.J. Brown _BC A. contiguun Fall _BC A. depressun. Fall AK _BC AB PQ A. difforrnis (LeConte) -oN canadense W.J. Brown A. effluens Mannerheim AK A. jasperanurn Fall YK A. maculosurr W.J. Brown A. obtusum llatch A. pulchrurn LeConte AK ma ndib rilatum Mannerheim A. rotundulu.n Mannerheim AK hincaidi Hatch Subfamily CATOPOCERINAE Genus CATOPOCERUS Motschulsky Homoeosoma ArsLin Pinodytr,'s Hotn Typhloleiodes Hatch C. cryptophagoides (Mannerheim) AK BC nolilzs l[otschulskv Subfamily COLONINAE Genus COLCN Herbst (Subgenus CiJRVIMANON Fleischer) C. bidentaturn (C.R. Sahlberg)* -BCABONPQ NF C. mannerheimi Szl'rnczakowski ON Iu_ (Subgenus COLON s. str.) C. femorale I'Iatch _BC C. serripoides Hatch _BC C. tibiale Har,ch _BC 80 Leiodidae (Subgenus MYLOECHUS Latreille) C. aedeagosum Hatch _BC C. celatum Horn _BC C. clavatum Mannerheim AK _BC C. cornplicatum Hatch -BC C. dentatum LeConte -oN PQ C. discretum Hatch -BC C. elongatum Notman -oN C. inerme Mannerheim AK _BC C. kincaidi Hatch -BC C. magnicolle Mannerheim AK NT BC AB MB ON PQ NB LB NF C. nevadense Horn _BC C. pribilof Hatch AK C. pusillum Horn _YK pe Subfamily CHOLEVINAE Tribe Ptomaphagini Genus PTOMAPHAGUS Illiger Key to North American species: Peck (1973) P. breviorJeannel ON PQ P. nevadicus Horn BCAB_MB densus Hatch piperi Hatc}r thomomysiHatch Tribe Nemadini Genus NEMADUS C.G. Thomson N. brachyderus (LeConte) PQ NS N. decipiens (Horn) --ec--- 9* N. gracilicornis Fall _MB N. parasitus (LeConte) _MB ON PQ N. pusio (LeConte) BC T?ibe Cholevini Genus CATOPS Paykuli C. alpinus sitkanus Jeannel AK YK NT BC C. alsiosus (Horn)x AK YK NT BC AB MB ON PQ NB NS LB C. americanus Hatch AK YK NT BC AB SK MB ON PQ NB NS clauicornis LeConte C. basilaris Say AK YK NT BC AB SK MB ON PQ NB NS LB NF co d o ueri n u s Mannerheim spencianus (Kirby) C. egenus (Horn) AK YK NT BC AB SK ON C. gratiosus (Blanchard) _BCAB oN PQ NS C. luridipennis Mannerheimx AK YK NT BC AB SK MB ON PQ b r unnip e nni s Mannerheim C. mathersi Hatch -YK_BCAB C. simplex Say AK BC AB SK_ONPQ NB NS_LBNF Genus SCIODREPOIDES Hatch Sciodrepa C.G. Thomson S. fumatus terminans (LeConte) AK _NTBCABSKMBONPQ S. watsoni hornianus (Blanchard) BC AB SK MB ON PQ YNS NF Genus CATOPTRICHUS Murray C. frankenhaeuseri (Mannerheim) AK BC Genus PRIONOCHAXTA Horn P. opaca (Say) ON PQ NS 81 Scydmaenidr,Le Subfamily LEPTININAE Genus LEPTINILLUS Horn L. validus (Horn) AK l.K NT BC AB SK MB ON PQ NB NF Genus PLAI'YPSYLLUS Ritsema Platy psitllus Westwood P. castoris Ritsema* AK BC MB ON PQ castorinus Westwood Genus LEPTINUS P.W.J. Muller Key to North American species: Peck (1982) L. occidentarnericanus Peck AK YK BC Family SCYDMAENIDAE antlike stone beetles A. Dauies Forty-eight species of Scydmaenidae have been Scydmaenidae, but these works included rlany syn- recorded in Canada and Alaska. Improvements in onyms, and the higher classification was confused. the current taxonomic status and in the methods Marsh (1957) made major improvements for the and frequency of collection of the species should western North America fauna, and recently Franz greatly in:rease this number. Scydmaenidae tend (1985) viewed Casey's North American gemera in a to be common locally, possibly following distribu- worldwide perspective. The last publlication, tions of the mites on which both the adults and lar- together with recent work by Suter and Newton rrae feed. They are found chiefly in wet mosses, (personal communication), forms the basis of the deep forer;t litter, decaying logs and tree stumps, present arrangement. tree holes, sawdust piles, and compost; some species are associirted with ants and caves. LeCorrte (1852) and Casey (1897) described AK (5); BC (12);AB (1); SK (1); MB (3); ON (2e); PQ most of the North American species of (14);NB (8);NS (6);LB (1) Subfamily SCYDMAENINAE Tlibe Lophioderini Genus LOPF:IIODERUS Casey L. biformis (.iweklin) AK BC L. insignis Ni.arsh BC L. myops Ca;ey _BC L. similis Mr.rsh -BC Tribe Euconnini Genus DRASITOPHUS Casey D. laevicollis Casey BC Genus EUCONNUS C.G. Thomson (Subgenus EUCONNUS s. str.) E. clavipes (Say) MB ON PQ pilosico Jlis (LeConte) E. lecontei (Sichaufuss) -oN E. semiruber Casey AB SK MB E. ventralis ilasey _NS (Subgenus CONNOPHRON Casey) E. ater (Casr:y) ON E. brevicornis (Say) ON E. comptus (Casey) ON E. debilitans (Casey) oN PQ NB E. elongatus (Casey) ON E. flavitarsis (LeConte) oN PQ NB E. gaudens (Casey) ON NB 82 Scydmaenidae E. illustris (Casey) ON E. novellus (Casey) -PQ E. optatus (Casey) ON PQ E. oreophilus (Casey) ON E. pertinax (Casey) ON E. pumilus (Casey) ON E. tenebrosus (Casey) ON E. testaceipes (Casey) ON PQ E. trifrdus (Casey) ON (Subgenus SCOPOPHUS Casey) E. affrnis Casey ON (Subgenus PSOMOPHUS Casey) (Xestophus Casey) E. debilis Casey ON E. fatuus (LeConte) ON PQ NB NS E. salinator (LeConte) _MBON_NB (Subgenus PYCNOPHUS Casey) (Nudatoconnus Franz) E. rasus (LeConte) ON PQ $ E. longiceps Fall AK BC Tribe Stenichnini Genus MICROSCYDMUS Saulcy & Croissandeau Opresus Casey M. misellus (LeConter oN PQ NB Genus STENICHNUS C.G. Thomson (Subgenus CYRTOSCYDMUS Motschulsky) S. badius (Casey) ON _NS S. ovipennis {Casey) _BC S. perforatus (Schaumr -oN S. turbatus tCaseyt PQ NB NS (Subgenus BRACHYCEPSIS Brendel) S. adjutor Fall AK S. pacificus (Casey) S. rothi (Marsh) S. subpunctatus (LeOonte) ON PQ _NS_LB (Subgenus PARASCYDMUS Casey) S. corpusculus (Casey) oN PQ NB (Subgenus TAPHROSCYDMUS Casey) S. californicus Motschulskv AK Tfibe Eutheiini Genus EUTHEIA Stephens E. morae Marsh BC E. scitula Meklin AK Genus \IERAPHIS Casey V. lacuna Marsh BC Tribe Leptoscydmini Genus LEPTOSCYDMUS Casey L. caseyi (Brendel) ON PQ Tlibe Scydmaenini Genus SCYDMAENUS Latreille Acholerops Casey 83 Silphidae S. retrusus (Casey) PQ S. zimmermz:.nni Schaum oll punctatus Casey Family MICROPEPLIDAE micropeplid beetles J.M. Campbell Micropeplidae is a small family of beetles with the feeding habits of the species; however, both lar- 10 species known in Canada and Alaska. They are vae and adults are assumed to feed ott fungal found in a variety of habitats, including deciduous spores. and conifr,,rous leaf litter, inside beaver lodges, in The family is well known in North .t\merica. Carex clurnps growing in shallow water, on freshly Campbell (1968) revised the New World spr;cies and cut stumps of fir and cedar and possibly associated published a catalog of the North Americatr species with the fungus Pyrobema, which is found in forests (1984c). after a firr:. Matthews (1970) described two species, M. hoogervdorni and M. hopkinsi, from Pliocene AK (a); YK (1); NT (1); BC (8); AB (a); SK. (2); MB deposits irr western Alaska. Little is known about (1); ON (5);PQ (a) ' H(1) Tribe Micropeplini Genus KALI|jSUS LeConte K. nitidus Lc Conte BC Genus MICROPEPLUS Latreille Anhenooeplus Koch Key to North American species: Campbell (1973b) M. browni Cr'Lmpbell -oNPQ M. brunneus Maklin AK BC M. cribratus LeConte AB -oN M. laticollis li{eklin AK YK BC AB SK _ONPQ M. punctatus LeConte AK BC costatus Maklin oregonensis Hatch' M. robustus ()ampbell BC M. sculptus l,eConte BC AB sx ON PQ obliquus LeConte M. smetanai Campbell BC M. tesserula Curtis* AK NT BC AB MB ON PQ costatus LeConte costipenzis Makiin Family SILPHIDAE carrion beetles J.M. Campbell The ca.rrion beetles belong to a family of about The family name Silphidae is userl in the 175 species found primarily in the Northern restricted sense of Lawrence (I982a), Lawrence and Hemisphere, with a few species having ranges Newton (1982), and Anderson and Peck (198{5). These extending into South America, southeast Asia, and authors exclude the agyrtids, treating them es a sepa- Australia. Twenty-five species are known to occur in rate family, Agyrtidae. The Canadian and Alaskan Canada and Alaska. Carrion beetles are common, species are well known taxonomically, based on a and because oftheir large size and often bright colors handbook ofthe species by Anderson and Peck (1985). they are frequently collected. Both adults and Iarvae ofmost species feed on carcasses ofvertebrates; other species are scavengers, and some species of Aclypea AK (8); YK (6); NT (9); BC (13); AB (17); SK (18); are phy'tophagous and may be considered to be pests MB (18); ON (18); PQ (15); NB (10); NS (11); PE (7); ofcultivated crops such as spinach. LB (1); NF (6) . H (4) 84 Silphidae Subfamily SILPHINAE Genus NECRODES Leach Proton ec rod es Portevin N. surinamensis (Fabricius) - BC AB SK MB ON PQ NB NS PE - NF Genus THANATOPHILUS Leach Philas Portevin T. coloradensis (Wickham) AK BC obalskii (Portevin) T. lapponicus (Herbst)* AK YK NT BC AB SK MB ON PQ NB NS - LB NF cal ifor ni cus (Mannerheim) caudatus (Say) tuberculatus (Germar) T. sagax (Mannerheim) AK YK NT BC AB SK MB ON PQ T. trituberculatus (Kirby) AK - NT - AB SK MB ON Genus OICEOPTOMA Leach O. inaequale tFabricius) - ON PQ O. noveboracense (Forster) AB SK MB ON PQ NB NS PE marginale (Fabricius) marginata Kirby Genus NECROPHILA Kirby & Spence N. americana {Linn6} MB ON PQ NB NS affinis (I{ttby) canadensis (Kirby) peltata (Catesby) terminata (KJrby) Genus HETEROSILPHA Portevin H. ramosa {Say) NT BC AB SK MB ON ceruaria (Mannerheim) Genus ACLYPEA Reitter Blitophaga Reitter A. bituberosa {Leconte) NT BC AB SK MB A. opaca tl-innet* AX YK NT Subfamily NICROPHORINAE Genus NICROPHORUS Fabricius Neonicrophorus Hatch N. americanus Olivier - ON pe grandis Fabricius uirginicus Fr6lich N. carolinus (Linn6J AB med.iatus Fabricius mysticallis Angell N. defodiens Mannerheim AK YK NT BC AB sK MB oN PQ NB NS pE - NF binotatus Portevin conuersator Walker hadjakensis Portevin lateralis Porte.rin mannerheimi Portevin plagiatus Motschulsky pollinctor LeConte N. guttula Motschulsky - BC AB SK hecate Bland immaculosis }{atch punctostriatus Pietce N. hybridus Hatch & Angell - BC AB SK MB ON N. investigator Zetterstedt* AK YK NT BC AB SK MB oN Pe - NS pE - NF aleuticus Gistel confossor LeConte 85 Staphylinidae infodiens Mannerheim mar it imu,s Gu6rin-M6neville melsheimeri lltrby particeps Fischer von Waldheim pollinctor Mannerheim N. marginatus, Fabricius BC AB SK MB ON PQ cordiger Portevin labreae Pterce latifrons Pierce mchittrictei Pierce m ontez u r,' ta e Matthews obtusiscu lellum Pier ce requiescalor Gistel N. nigrita Marrnerheim BC alphaPierce ruficornit Motschulsky N. obscurus K.rby AB SK MB melsheimeri LeOonte N. orbicollis S,ry SK MB ON PQ NB NS PE hallii \{trby quadrisignatus Castelnau N. pustulatus Herschel AB SK MB ON PQ NB NS bicolon N ewman fasciatus Portevin lordzs Mrnnerheim unicolor l)ortevin N. sayi Castel rau AB SK MB ON PQ NB NS NF lunatus I eOonte luniger Harold lunulatus Gistel N. tomentosur Weber SK MB ON PQ NB NS PE a ngustifosciolzs Portevin aurigaster Portevin uelutinus Fabricius N. vespilloider, Herbst* AK 1T< NT BC AB SK MB ON PQ NB NS PE NF hebes I\trby pygmaeu.r; I{trby Family STAPFIfl INIDAE rove beetles J.M. Campbell and A. Dauies The role beetles form one of the largest families they can also be found in a variety of habitats of insects in the world, with over 32 000 described around human dwellings. species. ,[n Canada, ll29 species have been Several authors are currently investigating the reported, b,ut hundreds more remain to be either higher classifrcation of the family. The system fol- described or recorded. Most rove beetles are preda- Iowed in this checklist has been extensively modi- tors of oth,:r insects and live on or in the soil, in fied from the conventional system used in earlier ground litt*:r, moss, or in decomposing organic mat- papers. The system is based primarily on a paper ter. Some r:ove beetles prey on aphids, bark beetles, by Lawrence and Newton (1982), but will uindoubt- or other ir.sects on trees and herbaceous plants, edly be modifred further as more phylogene bic stud- whereas others are specialized feeders on pollen or ies are completed. Adults of most genera of rove are myceto rhagous. Rove beetles prefer moist habi- beetles (except the genera of Aleocharinae) can be tats along t,he margins of streams, lakes, and snow- identifred with the use of keys by Arnett (1968) and frelds, and in decomposing vegetation, manure, and Moore and Legner (1974 and 1979) or with keys carcasses. They are common in all biomes, ranging based on the European fauna such as those ofLohse from tropical forests to tundra well above arctic and (1964) and Lohse et al. (1974). Most larvae have alpine timberlines. Some species are associated not yet been described. Newton (1990) provides keys with mammal burrows, ant and termite nests, to the North American genera of larvae and adults intertidal and coastal habitats. and saline flats: that inhabit soil litter. Other Nearctic genera can 86 Staohvlinidae be identified using the keys by Kasule (1966), based type specimens. In recent years a few genera of on the rove beetle larvae of Britain. North American Aleocharinae have been revised, Taxonomically, the North American fauna of and their species can now be identified. These many tribes and subfamilies is now well known: include the genera Aleochara, Myllaena, Gymnusa, Quediini and Xantholininae (Smetana Lg7l, 1982); and Deinopsls by Klimaszewski (1979, 1982, 1984) most Tachyporinae (various generic revisions by and the genus Falagriini by Hoebeke (1985). Ashe Campbell between 1973 and 1988); Oxyporinae (1984,1986) revised adults and larvae ofthe genera (Campbell f969); and Pseudopsinae (Herman 1975, of the aleocharine subtribe Gyrophaenina. Other Newton 1982). The large subfamilies Steninae and subfamilies with most species poorly known in Staphylininae are becoming well known among the Canada include Paederinae and Oxytelinae, except Canadian fauna, although comprehensive revisions for the genus Bledius, which was revised by of most genera are not yet available. However, Herman (7972a-f986). except for a few genera, Aleocharinae, the largest Moore and Legner (1975a) published a catalog subfamily of Canadian rove beetles, still remains of the North American Staphylinidae. almost totally unknown. Although Seevers (1978) provides a key and a redescription ofthe genera and AK (308);YK(r72); NT (155);BC (581);AB (265); SK tribes of the subfamily, identification even to genus (126); MB (278); ON (aa1); PQ (a60); NB (167); NS remains difficult or impossible without access to (195);PE (19); LB (63);NF (171) ' H (88);I (99) Subfamily EMPELINAE Genus EMPELUS LeConte E. brunnipennis (Mannerheim) BC Subfamily PROTEININAE Tribe Proteinini Genus MEGARTHRUS Curtis M. americanus Sachse MB ON PQ NS PE M. arcuatus Hatch AI{ BC IU M. atratus Maklin AK BC AB M. excisus LeConte -oN M. pictus Motschulsky AK BC M. sinuaticollis Boisduval & AK YK NT BC AB Lacordaire* angulicollis Maklin Genus PROTEINUS Latreille P. atomarius Erichsonf BC ON paruulus LeConte P. basalis Makiin AK BC AB P. brachypterus (Fabricius)* AK BC P. limbatus Maklin* AK BC -oN maekliniFavel Subfamily OMALIINAE Tribe Eusphalerini Genus EUSPHALERUM Kraatz Anthobium auct. (Subgenus EUSPHALERUM s. str.) E. convexum (Fauvel) ON PQ NB NS NF E. fenyesi (Bernhauer) SC AB _MBONPQ NB NS NF E. pothos (Mannerheim) AI{ BC AB ON PQ NB NS NF dirn idiat um (Melsheimer) E. rugulosum (Meklin) AK E. segmentarium (Meklin) AK E. torquatum ( Marsham)t NF (Subgenus ONIBATHUM Tottenham) 87 Staphylinida I E. californicum (Fauvel) _BC E. farrarae Hatch _BC E. grayae Ha:ch AK BC AB E. swauki Ha.tch _BC Tribe Omaliini Genus ACRLTLIA C.G. Thomson A. crenulata lHatch) AK _BC A. tumidula (Maklin) AK _BC Genus PYCNOGLYPTA C.G. Thomson Key to Norbh American species: Campbell (19836) P. aptera Carnpbell MB_PQ NB NS_LB NF P. lurida (Gy lenhal)* AK YK Nr--ix MB ON PQ NB NS Genus ELONIUM Leach Acrolocha C.G. Thomson E. diffusum ( :'auvel,) -oN PQ NB NS NF E. exsculptun tMiiklin r BC E. fractum {Fauvel) v_ MB ON E. helferi (St.elt RC E. leechi tHa chj BC E. minutum r Olivierl'i BC striatum (Gravenhorst) E. rugosum t . {atch r BC Genus HAPALARAEA C.G. Thomson Phyllodr epa C.G. Thomson H. angustior Hatcht BC H. caseyi (Be'nhauer & Schubert) BC pacifica .Casey) H. floralis tPr.ykull)+ - BC AB_MBONPQ NS NF H. hamata (Fauvel) BC MB ON PQ NS H. humerosa tFauvel) H. humilis (Ntaklin) AK BC ___9*PQ_ H. longula (Nl,iklin) AK BC H. megarthrcides (Fauvel) BC -AB H. strigipennLs (Maklin) AK Genus OMALIUM Gravenhorst Ochthen'nus Motschulsky O. algarum Casey BC fucicola Jasey O. callosum llaklin AK O. excavaturr Stephenst PQ O. flavidum Ilamilton -o^ PQ O. foraminosum Meklinx AK YK NT BC AB SK MB ON PQ O. oxyacanth;le Gravenhorstt - ?- O. plagiatum Mannerheim AK O. repandum Erichson pe- O. rivulare (Faykull)t AK _BC -o^ PQ NS NF Genus PHLOEONOMUS Heer (Subgenus PIILOEOSTIBA C.G. Thomson) P. iapponicus (Zetterstedt)E AK YK NT BC AB SK MB ON PQ NB NS argus Lt.Conte (Subgenus PHLOEONOMUS s. str.) (Distemrnus LeConte) P. flavipennis (Maklin) AK P. pusillus (Gravenhorst)* AK BC AB -oN PQ Ns-:Ne laesicollis (M€iklin) P. suffusus (Casey) AK Genus MICRI\J-YMMA Westwood Key to species: Bdcher (1988) 88 Staphylinidae M. brevilingue Schipdte* AK YK NT dicksoni Maklin M. marinum (Strctm)* PQ NB - NF stimpsoni LeConte Tribe Coryphiini Key to North American species: Campbell (1978c) Genus CORYPHIUM Stephens Occlephelinus Hatch C. arizonense tBernhauer, - BC C. brunneum (Hatch) - BC C. hyperboreum (Maklint* AK YK C. nigrum Campbell PQ - NS Genus SUBHAIDA Hatch S. ingratatHatcht AI( BC AB S. monticola Campbell AB Genus HOLOBOREAPHILUS Campbell H. nordenskioeldi (Maklin)x AK YK NT - MB - PQ nearcticus (B\ait) Genus BOREAPHILUS C.R. Sahlberg B. henningianus C.R. Sahlbergx AK YK AB SK MB ON PQ NB americanus Notman Tribe Anthophagini Genus BRATHINUS LeConte Key to species: Frank et al. (1987) B. nitidus LeConte PQ NB NS NF B. varicornis LeConte - ON PQ NB Genus ACIDOTA Stephens Key to North American species: Campbell (7982a) A. crenata (Fabricius)* AK BC AB SK MB ON PQ NB NS PE LB NF seriata LeConte A. quadrata (Zetterstedt)x AK YK NT BC AB - MB ON PQ - LB fr a nkenh ae u s eri Miiklin patruelis Leconte A. subcarinata Erichson - ON PQ NB NS NF Genus OLOPHRUM Erichson Lathrium LeConte Key to North American species: Campbell (1983o) O. boreale (Paykull)x AK YK NT BC AB - MB - PQ helleni Scheerpeltz nig r op ic eum Motschulsky quesneliHatch O. consimile (Gyllenhal)* AK YK NT BC AB SK MB ON PQ NB NS NF Iimbatum Maklin marginatum (Kirby) paruulum Meklin O. latum Maklin* AK YK NT O. obtectum Erichson - ON PQ emarginatum Erichson rotundicolle (Say) O. rotundicoile (C.R. Sahlberg)* AK YK NT BC AB SK MB ON PQ - NS NF conuexicolle LeConte conuercum Meklin Genus ARPEDIIIM Erichson Key to North American species: Campbell (19840) A. cribratum Fauvel - BC MB ON PQ NB NS - LB NF 89 Staphylinidae angulare Fanvel columbier,.se Hatch Genus EUCNIICOSUM Reitter Reuelstoh,:a Hatch Key to Nort.r American species: Campbell (19840) E. brachyptenLm (Gravenhorst)* AK YK NT MB beringense (Van Dyke) E. brunnescens (J. Sahlberg)* AK YK NT BC AB MB ON PQ LB NF hoppingi (Hatch) E. tenue (LeConte)* AK YK NT BC AB SK MB oN PQ .B NF Genus AMPHICHROUM Kraatz Staphygrophis Horn A. floribundum LeConte BC flauicorne'Casey A. macuiatum (Horn) BC immaculctum Hatch A. maculicolle (Mannerheim) AK Genus PELEC OMALIUM Casey Heterops -Vlannerheim P. laevicolle (LeConte) P. puberulum tFauvel) _BCAB _ :a_ I'_ . _ P. testaceum (:Vlannerheim) AK BC AB binotatun',. Casey nigrwn Hatch Genus LESTE VA Latreille Zeslo Bla:kweider Paralestetn Casey Pseudolesteua Casey Teuales C*ey L. pallipes Le( jonte ON PQ NB NS NF biguttata LeConte picescens LeConte Genus UNAMIS Casey U. columbiensis Hatch _BC U. fulvipes Fall _BC Genus PHLAEOPTERUS Motschulsky Phloeopte,.us LeConte Tilea Fatvel P. castaneus (Oasey) BC AB cascadien;is Hatc}' P. cavicollis (Fauvel) AK BC AB breuipennis Casey longipalp;rs Hatch P. frosti Hatch AK BC P. fusconiger trlotschulsky AK BC P. houkae Hat,:h AK BC AB P. lagrandeuri Hatch _BC P. loganensis llatch _BCAB Genus VELLICA Casey V. longipennis Casey BC Genus OROBANUS LeConte O. falli Mank _BC O. rufipes Cas,:y _BC O. simulator LeConte _BC O. tarsalis Hatch _BC Genus ARTOCHIA Casey A. productifrorrs Casey AK BC 90 Staphylinidae Genus XYLODROMUS Heer X. concinnus (Marsham)t AK BC SKAB Iacustris Casey X. depressus (Gravenhorst)t AK BC Genus ANTHOBIUM Leach Lathrimaeum Erichson A. atrocephalum (Gyllenhal)* AK A. clarki Hatch _BC A. fimetarium (Mannerheim) AK _BC A. occiduum (Casey) _BC AB A. reflexicolle Casey _BC A. sinuosum Hatch _BC Genus DEINOPTEROLOMA Jansson Mathrilaeum Moore Key to species: Smetana (19856) D. pictum tFauvel) -BC D. subcostatum (Maklin) AK _BC humerale (Casey) Genus TANYRHINUS Mannerheim T. singularis Mannerheim AK BC Genus TRIGONODEMUS LeConte T. fasciatus Leech BC T. striatus LeConte ON PQ NS Genus PORRHODITES Kraatz Key to North American species: Campbell (1984b) P. fenestralis (Zetterstedt)x AK YK NT BC AB MB PQ LB breuicollis Maklin P. inflatus tHatcht _BCAB -oN PQ Genus OROCHARES Kraatz Paradeliphrum Hatc}: Key to North American species: Campbell (1984b) O. tumidus (Hatch) BC Genus GEODROMICUS Redtenbacher G. brunneus (Say) PQ coeszs Erichson G. fauveli Casey _BC G. integer Casey _BCAB G. rufrpennis Van Dyke AK G. schultzi Fauvel -BC G. strictus Fauvel o" pe NF nigrita Farel G. temporalis Casey BC G. verticalis (Say) AK YK AB MB ON PQ NB NF ouipennis LeConte Genus MICROEDUS LeConte M. austinianus LeConte AK YK _BCAB_PQ M. fenderi Hatch _BC M. laticollis (Mannerheim) AK _BC M. porterae Hatch _BCAB M. rogersi Hatch _BC Genus HAIDA Keen Key to species: Campbell (19786) H. keeni Keen AK BC 91 Staphylinidar,, Subfamily OLISTHAERINAE Genus OLIST HAXRUS Dejean O. megacephz,.lus (Zetterstedt)* AX YK NT AB _PQ laticeps i,eConte O. substriatur; (Gyllenhal)x YK NT AB oN PQ nitidus I eConte Subfamily PIESTINAE Tribe Piestini Genus SIAGCTNIUM Kirby & Spence Prognatl, us Berthold Key to Nort,h American species: Moore (1975) S. americanurn Melsheimer AB NF conuergeizs Sachse S. punctatum LeConte AB SK MBONPQ_NS NF S. stacesmith: Hatch _BC AB SK MBONPQ_LB Tlibe Trigonurini .Genus TRIG(,NURUS Mulsant Key to Nor h American species Blackwelder (1941) T. crotchii Lellonte BC lecontei flharp T. dilaticollis Van Dyke BC T. sharpi Blat.kwelder BC T. subcostatu,' Meklin AK Subfamily OSORIINAE Tribe Osoriini Genus HOLO IROCHUS Erichson Neotroch,^ts Blackwelder H. laevicaudus (LeConte) MB Tlibe Lispinini Genus CLAVILISPINUS Bernhauer C. prolixus (L,-.Conte) MB PQ C. rufescens (,:Iatch) BC AB Genus THOR,\COPHORUS Motschulsky Glyptomtt Erichson T. costalis (Erichson) MB ON PQ Tlibe Eleusinini Genus RENAIDIA Motschulsky Eumalus Sharp R. canadensis Horn R. nigrella (Lt:Conte) ---ecAB- _ 9*______jubilaea Motschulsky Subfamily OXYTELINAE Key to genera: Herman (1970) Tribe Coprophilini Genus DELEi\STER Erichson D. concolor LcConte BC D. dichrous (Ciravenhorst)t oN PQ LB NF 92 Staphylinidae Genus COPROPHILUS Latreille Zonoptilus Solsky Key to North American species: Campbell (1979a) C. castoris Campbell NB C. sexualis Leech - 9*PQ Genus SYNTOMIUM Curtis Eunonia Casey S. confragosum Meklin AKBCAB_PQ S. grahami Hatch AK_PQ S. keenianum (Casey) _BC )" S. malkini Hatch AK BC Genus MITOSYNUM Campbell M. vockerothi Campbell NB Genus APLODERUS Stephens Phloeonaeus Erichson A. linearis LeConte Genus BLEDIUS Leach Microbledius Herman Psamathobledizs Herman Key to North American species: Herman (I972a,1976, 1983, 1986) B. albonotatus Miiklin AK BC alb idi p ennis Bernhauer ornatus LeConte B. analis LeConte AB_MB ON PQ B. annularis LeConte AK YK NT BC AB SK MB ON PQ NB LB NF Ianguidus Casey B. aquilonarius Herman AK YK NT BC AB oN PQ B. assimilis Casey _MB ON PQ B. basalis LeConte -NS--- ignauus Casey misellus Casey B. confusus LeConte AK YK NT BC AB SK _ONPQ B. coulteri Hatch BC SK MB B. emarginatus (Say) -oNPQ troglodytes Erichson B. flavipennis LeConte BC SK MB agonus Casey cuspidatus LeConte nelsoniHatch tenuis Casey B. foraminosus Casey BC deceptiuus FalI relictus Fall B. fumatus LeConte . MB ON B. gravidus Casey NT BC AB SK MB ON missionensis Hatch B. habrus Herman AK YK BC B. ineptus Casey AB SK MB ON B. longipennis M,iklin AK B. mandibularis Erichson SK MB breuidens LeConte B. melanocephalus (Say) MB confinis Fall B. monstratus Casey BC B. mysticus Fall BC B. nardus Herman BC B. neglectus Casey NB NS NF B. nitidicoilis LeConte _MB B. omega Herrnan AB _MB oN PQ B. opacifrons LeConte nc _MB condonensis Hatc}' lectus Casey piceus Fall regularis Fall specularis Fall 93 Staphylinidr,Le B. opaculus LeConte _NS PE _NF B. philadelphicus Fall -oNPQ NB _NF dickersoni Notman dissimit'.is Fall folli Wendeler mirlas.Notman B. politus Erichson NB nigriceJts Notman B. rotundicollis LeConte MB B. rubiginosus Erichson MB B. ruficornis LeConte B. semiferrugineus LeConte _:c_:"yuo^rF8_ _ _ _ _ canaliailatus Notman B. strenuus tlasey BC AB SK MB ON arizonensis Fall furtiuus Casey B. suturalis LeConte YKAK NT BC AB SK ON luteiper nis LeConte medialis Fall oregonensis Hatch pleuralt.s LeConte B. tarandus Herman AK YK NT BC AB SK MB ON PQ NF diuisus LeConte B. tau LeCorrte MB ON PQ NB NS NF B. turgidus t.)asey AK YK NT BC AB SK MB ON PQ NF borealis Blatchley bowronensis Hatch B. viriosus lr.erman AK YK NT BC AB SK B. washingtonensis Hatch _BC _9*PQ B. zophus H rrman _YK_BC Genus OCH'|HEPHILUS Mulsant & Rey Ancyrol horus Kraatz O. biimpressus (Meklin)* AK BC O. columbierLsis (Hatch) AKYK_BC _v O. planus (LeOonte) AKBC_ONPQ Genus THIN ODROMUS Kraatz T. corvinus (Casey) g*PQ -_ T. diffusus (tlasey) _ nc ______ Genus CARI'ELIMUS Leach P a rat r a,go p hlo e us Hatch Trogophloeus Mannerheim C. bilineatus Stephenst BC C. congener Jasey BC C. conjunctus (Casey) BC C. diffusus (tlasey) BC C. harneyi H.atch BC C. nanuloides Hatch BC C. obesus (Kiesenwetter)f -PQ fo s s ula t u s Motschulsky mancus (Casey) spectatl;.s (Casey) C. vancouvelensis Hatch -BC C. volans (Notman) _BC Genus THIN OBIUS Kiesenwetter Torrentornus Bietig T. alaskanus (Fall) .AK T. frizzelli Hatch _BC T. teres (Fall) AI{ Tlibe Oxytelini Genus PLA1 YSTETHUS Mannerheim P. americamrs Erichson MB ON PQ P. degener Mulsant & Reyt _:'T:" -oN 94 Staphylinidae Genus OXYTELUS Gravenhorst O. fuscipennis Mannerheim AKYK-BC ABSKMBONPQ_NS O. Iaqueatus (Marsham)t _BC O. montanus Casey BC AB ON PQ NB O. nimius Casey T_ _ _ONPQ O. pennsylvanicus Erichson _ONPQ O. sculptus Gravenhorstf _BC _ONPQ Genus APOCELLUS Erichson A. sphaericollis (Say) AB MB ON PQ amabilis Sachse Genus ANOTYLUS C.G.Thomson Key to species: Hammond (1976) A. insecatus (Gravenhorst)t -oN A. insignitus (Gravenhorst) A. niger (LeConte) _BC MB A. nitidulus (Gravenhorst)f _BC b or eal is (Motschulsky) punctatus (LeOonte) rugulosus (Say) A. rugosus (Fabricius)t _BC MB ON PQ NS A. sobrinus (LeConte) _BC AB SK MB IU __Iu A. suspectus (Casey) MB A. tetracarinatus (Block)t _BC PQ Subfamily PSELJDOPSINAE Key to genera: Newton (1982) Genus PSEUDOPSIS Newman Key to species: Herman (1975) P. montoraria Herman _BC P. sagitta Herman P. subulata Herman T NF Genus ZALOBIUS LeConte Key to species: Herman (1977) Z. spinicollis LeConte BC Genus NANOBIUS Herman Key to species: Herman (1977) N. serricollis (LeConte) BC Genus ASEMOBIUS Horn Key to species: Herman (1977) A. caelatus Horn BC Subfamily PHLOE OCIIARINAE Key to genera: Smetana and Campbell (1980) Genus CHARIfYPHUS Sharp Chapmania Bernhauer P seude le u s is Bernhauer Siberia Blackwelder Triga Farel Trigites Handlirsch Key to species: Herman (I972b) C. picipennis (LeConte) oN PQ NS Genus VICELVA Moore & Leqner V. vandykei (Hatch) AK YK BC 95 Staphylinidu e Subfamily TACI{YPORINAE Tribe Tachyporini Genus TACFIINUS Gravenhorst Key to North American species: Campbell (1973o), (1983) (Subgenus T.ACHINUS s. str.) T. addendus Horn _MB ON PQ _NS T. angustatus Horn - YK _BC AB T. apterus M:iklin AK T. basalis Erichson* AK YK _BC AB ix me ON PQ NN NS circumcinctus Maklin T. beckeri Ci,Lmpbell - YK _BC T. brevipennis J. Sahlberg* AK YK NT T. canadensis Horn oN pe T. contortus Hatch _BC AB uarians Hatch T. corticinus Gravenhorstt PQ T. crotchii H.rrn BC T. dilatatus l{atch BC AB T. elongatus Gyllenhal* AK YK NT BC AB SK MB ON PQ LB NF parallelus Hot:n T. fi mbriatus, Gravenhorst ON PQ_NS T. frigidus E ichson AI( YK NT BC AB SK MB ON PQ_NS TII NF propinq tus Mannerherm T. fumipennis tSayt AB _MB ON PQ NB NS T. horni Can pbell ON PQ_NS T. instabilis \{tiklin* AK YK BC T. jacuticus j rcuticus Poppiusx AK l-I( NT nearctic'ts Campbell T. jacuticus r Lllrichi Campbell PQ LB NF T. lanei Hatch BC AB T. limbatus itlelsheimer ON PQ NB NS T. luridus Er ichson _MB ON PQ NB NS T. maculicoll s Maklin ec- T. memnonil s Gravenhorst _MB ON PQ NB NS batychrous Gravenhorst rulzzs Sz.chse T. minimus (lampbell ON PQ T. nigricornil; Mannerheim AK BC AB T. picipes Er chson _ONPQNBNSPE T. quebecensis Robert AK YK NT BC AB SK MB ON PQ NB NS NT acutus Ilatch T. rainieri H rtch BC T. rufipes (D rCeert+ BC -PQNBNS NF T. schwarzi Ilorn _PQ-NS T. scrutator tlemminger & Harold _AB MB ON PQ NB NS discoideus Melsheimer repandu:.s Hotn T. semirufus Horn AK BC T. smeLanai tlampbell BC AB T. stacesmitl i Campbell BC T. subterranlus t Linne rt BC T. tachyporoides Horn NT BC AB SX ff,fS oN pe Nn Ns - ie lrr T. thruppi H;rtch - YK BC AB _MB ON PQ NB NS _LBNF T. vergatus ( ampbell oN PQ (Subgenus T.,\CHINODERUS Motschulsky) T. debilis Ho'n BC Genus COPROPORUS Kraatz Erchom'ts Motschulsky Key to North American species: Campbell (1975) C. ventriculus (Say) - NT BC AB SK oN PQ NB NS acuducttrs (Kirby) affinis (I{trby) flauidus (Casey) gibbulus (Erichson) maneei liicheerpeltz punctul 96 Staphylinidae Genus CILEA Jacquelin du Val Key to North American species: Campbell (1975) C. silphoides ( Linn6tt BC AB oN PQ NB Genus TACHYPORUS Gravenhorst Key to North American species: Campbell (1979b) (Subgenus PAI-PORUS Campbell) T. nitidulus (Fabricius)* AK YK NT BC SK MB ON PQ LB NF faber Say (Subgenus TACHYPORUS s. str.) T. abdominalis (Fabricius)* AK YK NT BC AB MB ON PQ NB NF T. borealis Campbell AK YK NT BC AB SK MB ON PQ NS T. browni Campbeil PQ NB T. californicus Horn _BC tehamoe Blackwelder T. canadensis Campbeli -\'K-BC SK MB ON PQ NB NF T. dimorphus Campbell _BC T. dispar (Paykull)t _BC _ONPQ NB NS NF chrysomelinus arct. T. elegans Horn -ONPQ T. flavipennis Campbell AK YK NT BC AB SK MB ON PQ NB NF T. inornatus Campbell AK YK NT BC SK MB ON PQ NB T. jocosus Say v MB ON acaudus Say arduus Erichson infuscatus Bernhauer maculipennis LeConte snyderi Blackwelder T. lecontei Campbell BC AB SK MB ON PQ T. maculicollis LeConte BCABSKMBONPQ_NS T. mexicanus Sharp YK_ BCABSKMB_PQ_NSNF alleni Blackwelder oregonus Blackwelder temacus Blackwelder T. nanus Erichson -oNPQ T. nimbicola Campbell AK YK NT BC AB PQ NB NS -LB NF T. ornatus Campbell SK MB_PQ T. pacifrcus Campbell BC T T. pulchrus Blatchley MB ON PQ T. rulomoides Campbell MB ON PQ NB T. rulomus Blackwelder NT BC AB MB ON PQ NT T. stacesmithi Campbell TY BC _:u T. transversalis Gravenhorstf _ONPQ Genus SEPEDOPHILUS Gistel Key to North American species: Campbell (1976) S. basalis (Erichson) ON pulicarius (Sachse) setifer (Casey) S. bisigratus (Horn) BC S. cinctulus (Erichson) ON PQ corticola (Casey) S. crassus (Gravenhorst) ON PQ moestus (Say) S. ctenidialis Campbell ON S. Iittoreus (Linn6)t BC AB MB ON PQ NN NS NF carissimus Flatch hnoxii (LeConte) S. marshami (Stephens)t -PQ _NS S. opicus (Say) -PQ S. scriptus (Horn) oN PQ S. testaceus (Fabricius)t -ABSK ON PQ NB NS I\TF elongatus (Blatchley) imbricatus (Casey) limuloides (Casey) S. velocipes (Casey) NS uirginiae (Casey) S. wickhami Campbeil ON PQ 97 Staphylinidr.e Tlibe Bolitobiini Genus LORDITHON C.G. Thomson Key to No:th American species: Campbell (1982b) (Subgenus LORDITHON s. str.) L. anticus (Forn) ON PQ NB NS L. appalachi,rnus Campbell MB ON PQ NB NS L. axillaris {,lravenhorst) PQ L. cascadensis iMalkin) BC hatchi (llalkin) L. diffrcilis Campbell BC L. facilis {Caseyt -oN PQ NB NS_ _NF L. niger (Grevenhorstt -oN PQ L. poecilus (J,{annerheim t BC AB L. scutellarii Campbeli -oN PQ NB NS _NF L. thoracicus thoracicus AK YK NT BC AB MB ON PQ NB NS LB NF (Fabricius)x cincticoilis (Say) occiduus (Casey) (Subgenus B OLITOBUS Tottenham) L. bimaculat rs (Coupen BC AB MB ON PQ NB NS kelleyi l[.alkin L. cinctus (Gravenhorsl) -oN PQ atricautt.atus (.Say) gentilis iLeConte) L. fungicola ilampbell AK YK NT BC AB SK MB ON PQ NB NS PE L. idahoae Hatch AB :uy L. longiceps lleConte) AK _BCAB -oN PQ NS elelas (Elernhauer) L. notabilis (:ampbell PQ L. obsoletus Sayr NS sellolzs Sachse L. oregonus l)ampbell BC L. quaesitor Hornt -oN PQ NS rostratus (LeConte) Genus CARI'HACIS des Gozis Key to North American species: Campbell (1980) C. intrusus ( lornJ -MBONPQ C. nepigoner:sis (Bernhauer) - BC MB ON PQ NB NS _NF rz6er (1\lalkin) wallisi (Malkin) Genus BOLI'IOBIUS Leach B. analis tPaykullt* NT AB SK MB ON PQ NS cingulai.us arct. B. kremeri tldalkin] _BC Genus BRYCITPORUS Kraatz B. rubescens (Hatch) _BC B. rufescens LeConte _BC -oN peNe- LB paruulu.s Casey rubidus Leconte testaceui LeConte B. varians (F atch) _BC Genus MYCIITOPORUS Mannerheim (Subgenus ISCHNOSOMA Stephens) M. pictus Horn _BCABSK MB ON PQ NB NS NF M. splendidus (Gravenhorst)* AK YK NT BC AB SK MB ON PQ NS NF elegans Matthews inquisiti^ts Casey pallidus Mannerheim (Subgenus MYCETOPORUS s. str.) M. american rs Erichson MB ON PQ NB lucidulus LeConte 98 Staphylinidae M. brunneus (Marsham) _BC M. consors LeConte MB ON PQ NB NS M. discalis Hatch -BC M. inquisitus Casey M. insignitus Maklin AK___f:"yu9*PQIU___ M. neotomae Fall _BCAB M. punctatissimus Hatch _BC M. punctulatus Hatch _BC M. rusosus Hatch AK YK NT BC AB SK MB ON PQ - NS Subfamily TRICHOPIIYINAE Genus TRICHOPHYA Mannerheim Eumitocerus Casey T. piiicornis (Gyllenhal)f BC ONPQ_NS Subfamily HABROCERINAE Genus HABROCERUS Erichson H. capillaricornis (Gravenhorst)t BC-ONPQ _NF H. magnus LeConte ON PQ _NF H. schwarzi Horn _AB_MBONPQ Subfamily ALE O CHARINAE T?ibe Oxypodini Key to North American genera: Seevers (1978) Genus DEVIA Blackwelder Dasyglossa Kraatz D. congruens (Casey) AK_NT MB Genus GNATHUSA Fenyes G. caribou Lohse AK YK NT G. tenuicornis Fenyes _BC Genus OCYUSAl{raatz O. canadensis Lohse AK YK Genus ILYOBATES Kraalz Gennadota Casey L canadensis (Casey) -oN Genus MOLUCIBA Casey M. g:randipennis Casey _BC Genus OCAJ-EA Erichson lsoglosso Casey O. vancouveri Casey _BC Genus OXYPODA Mannerheim Hylota Casey O. bradorensis Lohse PQ O. chillcotti Lohse YTNT_ _ _ MB O. demissa Casey O. egestosa Casey AK BC _ 9N_ _ O. famula Casey _BC O. frigida Bernhauer _BC O. glenorae Casey _BC O. hemingi Lohse AK O. hiemalis Casey -oN-- O. impressa Casey AK BC O. irrasa Meklin AK O. lacustris Casey -oN-- O. lassula Casey AK O. lata Lohse MB 99 Staphylinida e O. leechi Lohse _YK O. madgei Lc,hse AK O. manitobae Casey MB O. nelsoni Lchse AK__- O. optiva Casey _BC O. parafunebris Lohse AK YK NT MB O. regressa Clasey _BC O. subpolaris Casey AK Genus PENT ANOTA Bernhauer P. meuseli Bernhauer* AK Genus BRACIHYUSA Mulsant & Rey Tetralinz Casey B. americana (Fenyes) _BC B. helenae (C.asey) AK .\T Genus GNYI:ETA C.G. Thomson Euliusa Casey G. brevicornis Casey BC G. caerula (C.R. Sahlberg)'l NF G. manitobae Casey _MB Genus MER()NERA Sharp M. venustula (Erichson) -oNPQ Tribe Athetini Genus ACROTONA C.G. Thomson A. adjuvans Oasey A. onthophik Lohse AKYK_ _ _ _ 9"______ Genus HYDTrOSMECTA C.G. Thomson H. pseudodior;ica Lohse YK Genus CANASTOTA Casey C. canadensis; (Casey) -oN Genus DATOMICRA Mulsant & Rey Hilarina:. Casey Micromt:ta Casey Oligomia Casey Taxicere.tla Casey D. surgens (Casey) BC D. wrangeli (tlasey) AK Genus DIME'IROTA Mulsant & Rey Dalotia ()asey Dimetrotina Casey Engamotla Casey D. alaskana ( Oasey) AK BC D. altaica (Berrnhauer)* LB - _ pa:) D. bradorensrs Lohse yv_ D. burwelli L,rhse DN!q D. campbelli -ohse AK \'K D. caribou Lohse _YK oat D. dempsterensis Lohse -YK !q D. moesta Miiklin AK maeklini (Fenyes) D. munsteri (Bernhauer)* AK YK NT MB D. nearctica Lohse AK YK NT D. omissa Casey _BC D. picipennis (Mannerheim)x AK BC carlottae (Casey) D. prudhoensis Lohse AK D. regissalmonis Lohse AK 100 Staphylinidae D. resplendens Casey _BC D. retrusa Casey -BC D. smetanai Lohse AK YK NT MB pe LB D. stercoris (Fenyes) _BC columbica (Bernhauer) D. venti Lohse YK D. vicaria Casey BC Genus PSEUDOTA Casey P. cornicula Casey _BC P. formalis Casey _BC P. irrupta Casey _BC P. nanulina Casey _BC P. nescia Casey _BC P. vana Casey _BC Genus AMISCHA C.G. Thomson Colposura Casey A. analis (Gravenhorst)t NF Genus GEOSTIBA C.G. Thomson Sibiota Casey G. circellaris (Gravenhorst)t NF impressula Casey G. parvipennis (Bernhauer) AB Genus BOREOSTIBA Lohse B. campbelliana Lohse AK YK B. hudsonica Lohse B. lagunae Lohse -YK----PQ----- B. lamellifera Lohse _YKPQ B. sibirica (Maklin)* AKYKNT_PQ_LB Genus ADOTA Casey Panalota Casey A. comparabilis (Maklin) -AK A. maritima (Mannerheim) AK BC massettensis Casey Genus ANEPSIOTA Casey A. quadricollis Casey _BC A. terminalis Casey _BC renominata Bernhauer & Scheerpeltz Genus ATHETA C.G. Thomson Elytrusa Casey Stonalota Mannerheim A. circulicollis Lohse PQ A. granulata (Mannerheim) AK YK NT MB-PQ-LB nomadica Casey A. martini Lohse - I't( Genus ATHETALIA Casey A. metlakatlana (Bernhauer) _BC A. nimia (Casey) _BC Genus ATHETOTA Casey A. atriventris Casey BC A. wickhami (Casey) BC Genus LAMIOTA Casey L. achromata (Casey) BC L. concessa (Casey) BC L. keeni (Casey) BC L. profecta (Casey) BC 101 Staphylinidae Genus LIOGLUTA C.G. Thomson P s e udon; eg ist a Bernhauer L. aloconotoic.es Lohse LB L. granulosa Lohse AK L. insolens (Casey) _BC L. nigropolita (Bernhauer) _YK PQ LB L. nitens (Maklin) AKYK-BC L. trapezicollis Lohse _YK L. vasta (Maklin) AK \.K Genus MOCITA Mulsant & Rey M. amblystegii (Brundin tx AK YK NT MB: M. breviuscula (Maklin) AK BC AB prudens (Casey) Genus AI-AO.BIA C.G. Thomson A. sparreschr eideri (Munster)+ YKAK Genus SCHISiTOGLOSSA Kraatz S. aubeiodes .llrundin -AK S. holmbergi Brundin .AK Genus PSEUIfOUSIPAJ-IA Lohse P. microptera. Lohse YKAK Genus BORE fPHILIA Benick B. caseyi Loh;le AK YK NT B. caseyiana..,ohse _YK B. chillcotti Lrhse MB B. eremita (R re)* AK-MB LB B. fusca (C.R. Sahlberg)x AK_NT B. gelida (J. Sahlberg) AKYKNT_MB PQ B. hyperborei,. (Brundin)* AK_NT R. insecuta (F ppelsheim)* AK YK MB B. islandica (.iftaatz)x AK YK NT B. manitobensis Lohse AK_MB B. nearctica l,ohse AK YK MB PQ- LB B. nomensis (Casey) AK B. subplana tJ. Sahlbergt* AK_NT Genus CLUSIOTA Casey C. claviventris Casey BC Genus DINA-IiA-EA C.G. Thomson D. borealis Lchse D. planaris (I\Ieklin) AKYKPQ__-__ Genus BESSOBIA C.G. Thomson B. cryptica Lchse PQ Genus HOMIIOTUSA Casey H. manitobae (Casey) MB Genus MICRODOTA Mulsant & Rey Hilara Mrlsant & Rey Sonomota Casey M. globicollis (Bernhauer) ON M. holmbergi (Bernhauer) AK Genus PARAI)ILACRA Bernhauer P. glenorica ({)asey) BC Genus PHILI{YGRA Mulsant & Rey P. awemeana (Casey) MB P. botanicaru.m Muonaf N" L02 Staphylinidae P. criddlei (Casey) P. erudita (Casey) P. falcifera Lohse P. junii Lohse _YK P. laevicollis (Meklin) AK P. leechi Lohse NT P. malleoides Lohse AK YK NT P. polaris (Bernhauer)x AK YK NT P. prognata (Casey) BC P. pseudoboreostiba Lohse _YK P. ripicoloides Lohse P. rostrifera Lohse _ 5NT P. satanas (Bernhauer) P. stylifera Lohse -T P. surrufa (Casey) P. varula (Casey) Genus XENOTA Mulsant & Rey Delphota Casey Donesia Casey H alob r e cthina Bernhauer Nemota Casey Roualida Casey X. apposita (Casey) _BC X. brumalis (Casey) _BC X. cursor (Meklin) AK X. districta (Casey) _BC X. fanatica (Casey) _BC X. fungi (Gravenhorst)t X. mollicula (Casey) _BO-_ v X. mordax (Casey) _BC X. pallitarsis (Kirby) MB X. postulans (Casey) _BC X. pratensis (Maklin) AK X. pugnans (Fenyes) AK militaris (Casey) X. relicta (Casey) _BC X. remulsa (Casey) -BC X. resplendens (Casey) _BC X. rurigena (Casey) _BC X. subintima (Casey) _BC X. sumpta (Casey) X. tenuicula (Casey) _:r_ _ MB X. wrangelica (Casey) AK Genus LYPOGLOSSA Fenyes M eg acrotona Scheerpeltz L. angularis (Maklin) AK YK -BC-ONPQ NF L. fenyesi (Bernhauer) _BC Genus PONTOMALOTA Casey P. Iuctuosa Casey BC Genus STROPHOGASTRA Fenyes S. penicillata Fenyes MB Genus TARPHIOTA Casey T. debilicollis Casey BC T. fucicola (Maklin) AK T. geniculata (Maklin) AK BC insolita Casey T. seditiosa Casey BC Genus TRICHIUSA Casey T. columbica Casey BC T. postica Casey ON 103 Staphylinidae Tribe Aleocharini Genus AJ-ECiCHARA Gravenhorst Key to North American species: Klimaszewski (1984) (Subgenus CI)PROCHARA Mulsant & Rey) (.Eucharina Casey) (Funda Blackwelder) A. bilineata Gyllenhalt - BC AB - MB ON PQ - NS PE - NF anthomviae Sprague ontarionis (Casey) A. bimaculata Gravenhorst NT BC AB SK MB ON PQ - LB deserticc'la (Casey) innocua (Casey) obsolescens (Casey) recta (Casey) A. suffusa (Casey)* AK BC AB - PQ acumincta (Casey) perturbc,ns Fenyes A. sulcicollis Mannerheim AK BC cylindre']a (Casey) debilicornis (Casey) rugosa (Jasey) tibialis ( Casey) A. verna Say': AK YK - BC AB SK MB ON PQ - NS PE LB NF (Subgenus XINOCHARA Mulsant & Rey) A. castaneipcnnis Mannerheim AK YK - BC AB ON PQ acomantl (Casey) concurr(,ns (Casey) eludens .Casey) glenorar.a (Casey) insulant. (Casey) mannertteimi (Casey) oregona Hatch rotundicollis (Casey) A. fumata Gravenhorsti - YK - BC AB - MB ON PQ NB NS PE affiuens (Casey) defecta (Casey) languidu Sachse A. inexspectata Klimaszewski - - PQ - NS A. lacertina Sharp - BC AB SK MB ON PQ NB NS den s iu er, t r is Bernhauer idonea ('t)asey) imbricata (Casey) salicola lCasey) A. lanuginosaL Gravenhorstf - BC AB ON PQ NF A. sculptivenbris (Casey) - ON PQ NB - NF A. tristis Gravenhorstf PQ (Subgenus Cr\I-OCHARA Casey) (.OreochcLra Casey) A. laramiens s (Casey) AB SK A. rubricalis Caseyt BC - ON A. rubripenn s (Casey) MB ON PQ rubripes Blatchley wallawallae Hatc}r A. speculicollis Bernhauer AB ON PQ A. villosa Mannerheimf AK BC AB (Subgenus AI,EOCHARA s. str.) A. curtula (Greze)t - BC - ON PQ NB NS NF bugnioni Fa:uel fuscipes (Linn6) A. gracilicornis Bernhauer* NT BC - SK MB ON PQ - NS americata Casey ellipsicoi.lis Casey kansana Casey pleuralis Casey A. Iata Gravr nhorst+ - ON PQ A. assiniboin Klimaszewski - YK - BC - SK MB ON - r ufo nig r z Klimaszewski t04 Staphylinidae A. tahoensis Casey YK NT BC AB SK MB ON NS montanica Casey postpicta Casey A. thoracica Casey -oN PQ collusor Casey A.' sekanai Klimaszewski AK YK AB MB ON unicolor Klimaszewskr (Subgenus ECHOCHARA Casey) A. ocularis Kiimaszewski MB ON PQ (Subgenus EMPLENOTA Casey) (Polistoma Casey) (Polystoma Stephens) A. curtidens Klimaszewskr _BC A. litoralis (Miiklin) AK BC PQ NB NS NF arenaria (Casey) longiceps (Casey) maritima (Casey) quadrifer (Casey) trilimbata (Casey) A. pacifica (Casey) BC Genus PHLOEOPORA Erichson P. arctica Lohse NT Genus PAROCAI-EA Bernhauer P. nearctica Lohse AK YK NT P. pseudobaicalica Lohse AKYKNT_PQ Tribe Hoplandriini Genus HOPLANDRIA Kraatz Key to species: G6nier (1989) (Subgenus HOPLANDRIA s. srr.r H. lateralis (Melsheimer) ON PQ arizonica Casey carinata Casey ochracea Ktaatz texana Casey (Subgenus LOPHOMUCTER Notman) H. klimaszewskii G6nier _PQ H. Iaevicollis (Notman) -PQ Tlibe Falagriini Key to North American species: Hoebeke (1985) Genus CORDALIA Jacobs C. obscura (Gravenhorst)f ON PQ Genus MYRMECOCEPHALUS Macleay Lorinota Casey Stenagria Sharp M. arizonicus (Casey) BC acomanus (Casey) fontinalis (Casey) M. cingulatus (LeConte) oN PQ NS sinuosus Casey tenuicornis Casey M. gatineauensis Hoebeke PQ NB Genus FAI-AGRIA Leach F. dissecta Erichson _BCABSKMBONPQNB NS angulata Casey ery t hr optera Melsheimer 105 Staphylinidae globosa l{elsheimer iowana (iasey ithacana Casey sterilis Casey subsimilis Casey tetana Casey F. sulcata (Paykull)t AB ON PQ sicula Jekel Genus ALEODORUS Say Chitalia Sh.ary A. bilobatus (llay) ON PQ canadent:is (Casey) nigresceris (Casey) turbatus (Casey) A. intricatus (Casey) AB SK A. scutellaris iLeConte) ON granulos ts (Casey) illustris (Casey) Tribe Myrmedoniini Genus DRUS;LLA Leach D. cavicollis ((lasey) AK PQ Genus XENO DUSA Wasmann Pseudolomechuso Mann Key to Nonh American species: Hoebeke (1976) X. cava (LeCo nte) ON lobata Cr,sey probata l)asey X. reflexa (W:,Llker) BC SK MB ON PQ Genus ZYRAfi Stephens Platyusa Casey Z. criddlei (Ce.sey) MB Z. loricatus Casey -oN Tlibe Gyrophaenini Subtribe Gyrophaenina Key to North American genera: Ashe (1984) Key to North American species: Seevers (1951) Genus GYRO PIIAENA Mannerheim (Subgenus G\ROPHAENA s. str.) G. affinis C.R. Sahlbergf BC MB PQ NB lacustris Casey inconspir.:ua Casey subpunctata Casey G. antennalis Casey NB_ G. californica Casey ec----- G. caseyi Seevers -PQ G. criddlei Casey _MB G. egena Casey ON PQ exilis Ca,:ey G. flavicornis Melsheimer oN PQ G. gaudens C,rsey oN PQ G. geniculata Meklin AK G. gilvicollis Casey ; --- G. insolens Casey ON-NB G. keeni Casey YK BC AB ON PQ NB G. modesta Casey NB G. nana (Paykull)* AK YK BC AB MB ON perpolita Casey G. nanoides Seevers ON PQ G. sculptipennis Casey _PQ 106 Staphylinidae G. uteana Casey PQ pacifica Casey G. vitrina Casey - ON PQ dttonsa Casey (Subgenus PHAENOGYRA Seevers) G. gracilis Seevers - 2 G. subnitens Casey MB ON Genus EUMICROTA Casey E. socia lErichsont PQ Subtribe Bolitocharina Genus BOLITOCHARA Mannerheim Ditropalia Casey Pleurotobia Casey Silusid.a Casey Stictalia Casey Venusa Casey B. arcuata (Casey) _BC B. carlottae (Casey) _BC B. densicollis (Casey) -BC B. notata M:iklin AK Genus LEPTUSA Kraatz Dianusa Casey Eucryptusa Casey Ulitusa Casey L. fontana (Casey) -oN Subtribe Homalotina Genus HOMALOTA Mannerheim H. depressiuscula Mannerheim AK Genus PLACUSA Erichson P. turbata Casey - BC Subtribe Cyphina Genus CYPHEA Fauvel C. wallisi Fenyes MB Tlibe Autaliini Genus AUTAI-IA Leach A. brevicornis Casey _BC A. truncatula Casey _BC TYibe Phytosini Key to western Norbh American species: Moore (1956) Genus AMBLOPUSA Casey A. borealis Casey AK BC A. brevipes Casey AK BC pallida Casey Genus DIAULOTA Casey D. densissima Casey AK BC insolita Casey 107 Staphyiinida: Genus LIPAIiOCEPHALUS Maklin L. brevipennis Miiklin AK L. cordicollis LeConte BC Genus THIN JSA Casey T. fletcheri Casey AK BC diuergens Casey nigra Ct.sey robustulz Casey Tribe Gymnusini Key to North American species: Klimaszewski (1979) Genus GYMIIUSA Gravenhorst G. atra Casel'* AK YK NT BC MB ON PQ NB NS _LBNF G. brevicollis (Paykull)t PQ _LB G. campbelli Klimaszewski AK YK NT -oN PQ NB NF G. grandicepr; Casey MB ON PQ lrs NF G. konopacki. Klimaszewski AK YK NT MB G. pseudovariegata Klimaszewski AKYKNT_AB MB ON PQ NB NS _ I,B NF G. smetanai .l(] imaszewski AK YK NT MB ON NF Tribe Deinopsini Kev to North American species: Klimaszewski (1979) Genus DEIN,JPSIS Matthews D. canadensi,i Klimaszewski -oN D. harringtorLi Casey -oNPQ NB NS NF D. rhadina K limaszewski -oN Tribe Myllaenini Kev to North American species: Klimaszewski (1982) Genus MYLI.AENA Erichson M. arcana Casey AB ON PQ NB intmunci.a Casey M. audax Car;ey NT BC oN PQ NB abdita Casey dissimuLans Casey molesta Casey uegeta Casey M. decreta Casey BC AB impellen.s Casey scobinel't.a Casey umbra Qasey M. fenyesi Bcrnhauer BC esuriens Casey M. insomnis ilasey AK PQ NB NS M. Iudifrcans Casey yl'BCysKyuSN PQ breuiues;lis Casey M. procidua Casey PQ M. vulpina B:rnhauer NS Tlibe Oligotini Genus CYPHA Leach Hypocyphtus Gyllenhal C. crotchi Horn BC AB 108 Staphylinidae Subfamily OXYPORINAE Key to North American species: Campbeil (1969), (19780) Genus OXYPORUS Fabricius (Subgenus PSEUDOXYPORUS Nakane & Sawada) O. lateraiis Gravenhorst MB ON PQ NB NS breuis Melsheimer O. occipitalis Fauvel oN PQ O. quinquemaculatus LeConte _ T_:"ysKy" oNPQ-NS (Subgenus OXYPORUS s. str.) O. femoralis femoralis Gravenhorst ON PQ pulcher Ziegler O. kiteleyi Campbell PQ O. rufipennis LeConte -oN PQ_NS O. stygicus Say PQ_NS O. vittatus vittatus Gravenhorst MB ON PQ NB NS cinclzs Gravenhorst dimidiatus Melsheimer fasciatus Melsheimer Subfamily STENINAE Key to North American species: Casey (1884) Genus DIANOUS Samouelle D. chalybaeus LeConte AK NT oN PQ NB NF c oer u les cen s Blatchley D. nitidulus LeConte AK YK BC AB SK -PQ _NS NF zephyrus Casey Genus STENUS Latreille Laestris Melsheimer (Subgenus "HEMISTENUS" Motschulsky) S. croceatus (Casey) oN PQ S. kryzhanovskii Puthzx - - Nr ec AB ix Yu - - - - - S. niveus Fauvels AK I'K NT _ MB uislo Sanderson S. pubescens fraternus (Casey) AK YK NT BC AB SK MB ON S. pubescens sandersonianus Puthz AK YK _v S. retrusus (Casey) AK _BC S. umbratilis (Casey)x AK YK NTBCAB-MB -PQ LB NF (Subgenus IIYPOSTENUS Rey) (Arezs Casey) S. advena (Casey) NT-MBONPQ NB NS S. aiexanderi Puthz* -BCAB-MBONPQ S. annularis Erichson NB NS _T" S. arculus Erichson _ _ _:"TSKyuSNPQ NB S. cubensis Bernhauer -oN S. dissentiens (Casey) AK BC AB SK MB ON PQ NB lls - hubbardi (Casey) -Ne leuiceps (Casey) lucidus (Casey) politulus (Casey) politus Sanderson simiolus (Casey) S. flavicornis Erichson BC SK MB ON PQ NB NS geniculatus Say S. fulvicornis Stephensf BC S. insperatus Puthz lrn oN PQ NB S. lugens (Casey) NB perbreuis L. Benick S. pinguis (Casey) BC AB SK MB trajectus (Casey) utensis (Casey) S. plicipennis (Casey) AK YK NT BC AB MB agnatus (Casey) /orzs L. Benick 109 Staphylinidae S. pollens (Casey) AK NT BC AB SK MB patens (Casey) S. punctatus Erichson -oN PQ artus ((.asey) dilutus lCasey) obsoletus (Casey) S. quebecensis Puthz AK_NTBCABSK MB ON PQ NB NS NF S. reconditus breviusculus Puthz AX BC S. reconditus, reconditus (Casey) AK_NTBCAB MB ON PQ NB canader.sis (Casey) propinquus (Casey) S. rossi Sanc erson AX YK NT BC AB SK MB ON PQ NB PE NF S. varipes (Casey) -MBONPQ ft (Subgenus P ARASTENUS Heyden) S. pterobrachys Gemminger & Harold AK _BC breuipermis Maklin S. sibiricus J. Sahlberg* AK NT BC AB MB ON PQ NS LB NF alpicolo Sanderson confusus Casey deducto" (Casey) exaspentlus L. Benick montan'"ts Casey noctiua1lus Casey wenzeli Prt]az (Subgenus SIENUS s. str.) S. ageus Caseyx AK YK NT AB MB ON PQ NB NS LB NF rigidus Casey S. alveolatus Casey _BC AB S. angustus'Jasey ONMB PQ NB NS NF indigen.; Casey mendan Casey pettiti C.asey placidu.; Casey tutnicol,'is Casey S. assequens Rey* AK YK NT BC MB ON PQ NF nanulu:, Casey pusio Clsey S. austini Cr.sey AK YK NT BC SK MB ON PQ NS LB NF amplifir::atus L. Benick S. bilentigat rs Casey -oN PQ murphyanus Bernhauer S. bilineatus J. Sahlberg* AK NT BC AB SK MB ON PQ coruus Casey tacomar: Casey S. brivioi Pu,hz AK YK NT AB SK MB oN PQ NB NS NF S. callosifror.s Puthz ON :u S. canaliculatus Gylienhal* AK YK NT BC AB MB oN PQ NB NS NF congener Maklin S. cariniceps Miiklin AK BC S. carinicollis Casey ON PQ -NS-- NF S. caseyi Puthz ON PQ NB NS S. clarostigma Puthz AB S. clavicornis (Scopoli)t _PQ NB S. colon Say ON carolintts Bernhauer S. colonus E:ichson SK MB ON PQ NB NS PE arizonae Casey championi Sharp chapini Blackwelder floridan,us Casey inermis Sharp integer Dasey interpres L. Benick militaris Casey obstrusus Casey suspectus Blatchley weiseri Betnhaner S. columbianus Sanderson BC 110 Staphylinidae S. comrna LeConte* YK NT BC AB SK MB ON PQ punctiger Casey S. coriaceus Puthz* AK-NT S. dolosus Casey AKYKNT_ABSKMBONPQNB jejunus Casey uillosus Casey S. egenulus Puthz NT BC AB SK MB ON PQ LB NF S. egenus Erichson TY -MBONPQ S. eilipticus Casey BC S. erythropus Melsheimer ON PQ NSNS_ S. exploratus Fall* AK YK BC AB incanoides Prt}:rz S. fasciculatus J. Sahlberg* AK YK NT BC AB SK MB ON PQ NB NF S. femoratus Say MB ON PQ S. fulvoguttatus Notman -oNPQ S. geminorum Puthz BC S. hyperboreus J. Sahlberg* AKYK NT BC AB sr MB ON PQ LB falli Scheetpeltz frigid,us Fall S. insularis Casey AK BC S. intrusus Casey* AK YK NT SK MB ON PQ NB NS S. jacuticus Poppius* NT S. juno Paykull* AK YK NT BC AB SK MB ON PQ NB NS PE LB NF S. kamtschaticus Motschulskyx AK YK aquilonius L. Benick S. koebelei Puthz BC raniermontis Ptt]az S. Iabilis Erichsonx AK YK NT S. laccophilus Cdsey AK_NTBCAB SK MB ON PQ NB LB anastamozans Casey edar Notman rugifer Casey uerticosus Casey S. latipennis J. Sahlberg* AK YK NT S. latissimus hatchi Sanderson AKYK_BCAB -pe S. lustrator Erichsont S. mammops Casey* AK \'K NT BC AB - ftfS oN PQ Ne Ns Fe is Nr humilis Blatchley rulomus Blackwelder S. maritimus Motschulsky AK BC AB S. melanarius Stephens* _MB oNpeNs-- NF S. melanopus (Marsham)l vvv_ _ ON S. mendosus Puthz ON S. morio Gravenhorst* AK YK NT BC AB s^ me ON PQ NB diues Casey enod.is Casey haplus Casey indistinctus Casey luculentus Casey subgriseus Casey terricola Casey S. mundulus Casey _BC S. neglectus Casey MB ON PQ_ S. occidentalis Casey AK-NTBCABSKMB S. parallelopipedus Mdklin AK BC AB adspector Maklin animatus Casey conuictor Casey S. paululus L. Benick AK YK campbelliPrthz S. pauperculus Casey MB ON PQ brumalis Casey S. planatus Puthz BC -oN S. pluto Casey MB ON PQ S. pudicus Casey -oN enodis Casey juuencus Casey minor Casey sibyllinus L. Benick S. pumilio Erichson* AK YK NT MB ON PQ atomarius Casey 111 Staphylinid re S. pusillimus Puthz MB ON PQ NB NS S. scabiosus Casey AK YK rVr ec AB ix MB ON PQ NB NS NF S. schwarzi Oasey MB ON PQ NB NS:u_ S. scrupeus Casey AK _NTBCABSK MB ON PQ S. sculptilis Casey BC S. sectator Oasey BC S. semicolon LeConte AK _NTBCABSK lo*peNn S. sordidus .l)uthz *t_ y MB ON PQ S. sphaeroprir Casey _ _ONPQNB NS v S. strangula.tus Casey _ONPQNB S. stygicus Say AK \'K NT BC AB SK MB ON PQ NB NS NF S. tenuis Casey _ABSK _ONPQ dfficilis Casey eriensis Casey ingratus Casey inornatus Casey paralle,-us Casey pauper Casey subtilis Casey S. venustus Casey _BC S. vespertinrrs Casey _BC incultu:; Casey luctuos;ts Casey S. vexatus C.asey _BCAB MB S. vicinoides Puthz _BCAB SK MB S. vicinus Crrsey ON simplet Casey tricuspidatus L. Benick S. vinnulus ilasey* AK YK NT AB MB ON PQ NB NS LB confusoLdes Renkonen (Subgenus lESNUS Rey) (MutintLs Casey) S. californicrrs Casey BC S. curtus Casey _NT_AB_MBONPQ S. formicetorurn Mannerheim+ YKAB_MB_PQ borealis M,iklin S. gratiosus Casey -oN PQ NS hirsutu; Casey S. immargin atus Miiklin* AK YK NT BC AB MB PQ NB NS monticrla Casey uariabi,;.is Prthz S. tahoensis Casey BC pugetensis Casey uacuus Casey Subfamily EUAESTHETINAE Genus NOR.lf ENSKIOELDIA J. Sahlberg Key to species: Puthz (197h) N. columbiana Puthz _BC Genus STIC'IOCRANIUS LeConte Key to species: Puthz (7974b) S. puncticeps LeConte ON Genus EUAIISTHETUS Gravenhorst E. americanus Erichson MB ON PQ NB NS NF E. brevipennis Casey T_ I'- T_ ON PQ NB E. floridae Casey _NS E. ganglbaueri Bernhauer ____ oN PQ E. iripennis Casey oNPQ-NS E. laeviusculus Mannerheim AK l'K NT MB oNPQ-NS E. pacificus Jlall AK BC T:" E. pugetensis Hatch _BC oNPQNBNS-- t12 Staphylinidae Subfamily LEPTOTYPHLINAE Genus CHIONOTYPHLUS Smetana C. alaskensis Smetana 4K Subfamily PAEDERINAE Key to genera: Blackwelder (1939b) Tlibe Pinophilini Genus PAJ-AMINUS Erichson P. luteus Casey -oN Tlibe Paederini Subtribe Medonina Genus LITHOCHARIS Deiean Metaxyodonta Casey (Subgenus PSEUDOMEDON Mulsant & Rey) (Ramona Casey) L. capitula {Casey) - BC L. obsoleta tNordmann)t - BC L. thoracica {Casey) MB ON PQ NB clarescens (Casey) ruficollis (Casey) (Subgenus LITHOCHARIS s. str.) L. ochracea (Gravenhorst)'l - BC - ON PQ NB alutacea Casey quadricollis Casey Genus ACHENOMORPHUS Motschulsky (Subgenus ADEROCHARIS Sharp) A. corticinus tGravenhorst) MB ON PQ millepunctatus (Say) rufescens Nordmann Genus SUNIUS Curtis Medonella Casey Micromedon Casey Oligopterus Casey Xenocharis Bierig (Subgenus CAI-ODERMA Casey) S. rugithorax Hatch - BC (Subgenus IIYPOMEDON Mulsant & Rey) (Asteria Fauvel) (Hemimedon Casey) (Lena Casey) S. melanocephalus (Fabriciustf PQ S. peregrinus (Casey) - BC discolor Casey S. quadripennis Casey - BC (Subgenus TRACHYSECTUS Casey) S. confluentus rSay) MB ON PQ NB NS confluens Erichson Genus MEDON Stephens Oxymedon Casey (Subgenus MEDON s. str.) M. americanus Casey PQ M. fusculus (Mannerheimt+ - - PQ M. guppyi Hatch - BC 113 Staphylinidr.e (Subgenus PLATYMEDON Casey) (Paramzdon Casey) M. capitalis Flatch _BC M. convergens (Casey) AK M. insularis Casey -BC M. laticollis iCasey) MB M. mcleodi llatch _BCAB M. pugetensis Hatch _BC M. shastanic,us (Casey) _BC M. truncatu'r Hatch BC M. vancouve,ri (Casey) _BC borealil Casey (Subgenus X'IEDONODONTA Casey) M. ctenopho:um Hatch _BC Subtribe Scopaeina Genus ORUII Casey (Subgenus FYCNORUS Casey) Key to North American species: Herrnan (1965) O. dentiger (LeConte) AK - MB ON PQ humilis (Notman) iowanus Casey (Subgenus C RUS s. str.) Key to North American species: Herman (1965) O. distinctu, Casey BC O. punctatu,r Casey BC filius C*ey pallidus Casey sonomae Casey Genus SCOI'AEUS Erichson Leptorus Casey Pseudo"us Casey (Subgenus SCOPAXOMA Casey) S. brunniper LeConte - BC AB (Subgenus SCOPAXUS s. str.) S. notangulrLs Casey PQ (Subgenus -cCOPAEOPSIS Casey) S. ventralis Casey) - ON Subtribe Stilicina Genus PACI{YSTILICUS Casey P. hanhami (Wickham) _BCAB_MBNB Genus RUG.[LUS Leach Stilicosoma Casey R. biarrnatus (LeConte) MB ON PQ NB abbreuiellus Casey R. dentatus Say -oN luculentus Casey minusculus Casey nigrolucens Casey R. fragilis (Gravenhorst)t _ONPQ_NS dngustatus (Fourcroy) s cutella t us Motschulsky R. latiusculus (Casey) R. oregonus (Casey) AK- sc- R. rufrpes Germarf - - - il------Genus MEGASTILICUS Casey M. formicar:us Casey -oN rt4 Staphylinidae Subtribe Lathrobiina Genus LATHROBIUM Gravenhorst (Subgenus APTERAI-IUM Casey) L. sibiricum Fauvelx AK (Subgenus ABLETOBIUM Casey) L. shermani Fall ON (Subgenus LATHROBIUM s. str.) (Litolathra Casey) L. amplipenne Casey ON deceptiuum Casey L. arrnatum Say oN PQ procerum Casey L. confusum LeConte ON PQ L. divisum LeConte BC L. fauveli Duvivier -ABSKMB oN PQ NB NS PE NF simplex LeConte L. fulvipenne (Gravenhorst)t - BC AB NF p unct ulat u m Mannerheim L. hesperum (Casey) BC L. leconteanum Scheerpeltz NF concolor LeConte L. nigrolineum (Casey) _MB L. obtusum Casey _MB L. shoshonicum (Caseyr -BC L. simile LeConte -MBONPQ-NS-- L. sparsellum Casey AB _MBONPQNBNS L. spissicorne Casey ON PQ PE L. vancouveri Casey AK _BC L. washingtoni Casey AK NT BC AB SKMBONPQ_NS NT (Subgenus LATHROBIOMA Casey) L. othioides LeConte ON PQ L. scolopaceum (Caseyt L. tenue LeConte nc - - MnoNFS f"NS: (Subgenus LATHROLEPTA Casey) L. debile LeConte ON PQ (Subgenus TETARTOPEUS Czwalina) (Deratopeus Casey) Key to North American species: Watrous (1980) L. angulare LeConte PQ NB NS nigrescens (Casey) semirubrum (Casey) L. convolutum Watrous -ONPQ L. furvulum {Casey) NT AB SK MB ON PQ NB NS PE LB N" L. Iacustre (Caseyt MB ON PQ NB NS L. nigrum LeConte AK YK NT BC AB SK MB ON PQ NB NS PE NF finitimum LeConte stibium (Casey) L. nitidulum LeConte ON_NBNS NF bicolor LeConte bicoloratutn Bernhauer & Schubert paruipenne (Casey) L. punctulatum LeConte AK NT BC AB SK MB ON PQ capitosum (Casey) hebes Casey L. rubripenne tCasey) ON PQ agitans (Casey) callidwn (Casey) L. sollicitum Fall AK YK AB L. tetricum (Caseyt ON PQ canoaensis (Bierig) nigriceps Notman $ L. puncticolle Kirby MB 115 Staphylinidr:e Genus LOBIiATHIUM Mulsant & Rey (Subgenus EULATHROBIUM Casey) (Lathro.iropis Casey) L. grande (LeConte) L. subseriatum (LeOonte) ___sc___g*PQ_fr___ ustulatum Casey (Subgenus LOBRATHIUM s. str.) (Lathro,Siella Casey) (Lathro;laxis Casey) L. caseyianum ( Scheerpeltz) _BC bipartitum Casey L. collare (Erichson) MB ON PQ NS L. coloraden,;e Casey AK NT BC L. impressinLum Hatch _BC L. longiuscu um (Gravenhorst)f -ox L. multipunc,tum (Gravenhorst)t -BO L. rubricolle (Casey) _BC L. sewardi (Fall) AK (Subgenus PSEUDOLATHRA Casey) (Linolathra Casey) (Microluthra Casey) (.Parala::hra Casey) L. ambiguunr (LeConte) _ONPQ L. dimidiatum (Say) -oN horni Duvivier rufoparlitum Fairmaire & Germain L. famelicunr (Casey) _PQ L. lineiformt, (Casey) L. oregonense (Casey) BC 9*_ Subtribe Paederina Genus PAEI)ERUS Fabricius Leucopt.ederus Casey N eo p ae t:.le r us Blackwelder Paederillus Casey P. littorarius Gravenhorst NT BC SK MB ON PQ NB NS NF pugetensis Casey Subtribe Cryptobiina Genus HOMAEOTARSUS Hochhuth (Subgenus GASTROLOBIUM Casey) H. badius (Gravenhorst) ON H. bicolor (Gravenhorst) oN PQ Dr\ H. carolinus (Erichson) - rq (Subgenus HESPEROBIUM Casey) H. californicus (LeConte) BC pacificus Casey uancout'eri Casey H. cinctus (Say) BC AB oN PQ NS latebricola Nordmann H. cribratus (LeConte) -oNPQ H. pallipes (Gravenhorst) -oNPQ grauenhorsti I\trby H. sellatus (]-eConte) oN PQ Genus OCHI|HEPHILUM Stephens (Subgenus OCHTHEPHILUM s. str.) (C ry p to b ium Mannerheim) O. fracticorne (Paykull)+ BC ON PQ NS brachelytratum Korge furcaticcrne Grimmer 116 Staphylinidae Subtribe Astenina Genus ASTENUS Dejean A. americanus (Casey) -oN A. binotatus (Say) -oN A. brevipennis (Austin) MB ON A. cinctus (Say) _BCAB PQ A. discopunctatus (Say) AB -oNPQ NB centralis Ars.stin A. linearis (Erichson) A. longiusculus (Mannerheim) BC (?') t risig natu s Boheman Subfamily STAPFIYLININAE Tribe Philonthini Genus NEOBISNIUS Ganglbauer Key to New World species: Frank (1981) N. jucundus (Horn t - BC AB ON N. lathrobioides (Baudi )* PQ N. occidentoides Frank AB - MB N. senilis tHornt - BC N. sobrinus (Erichsont ON PQ NB I\TF arduus (Sharp) ocreatus (Hotn) simulator Smetana N. terminalis ( LeConte) PQ NS adustus (Sharp) delicatulus (Sharp) N. viliosulus t Stephenslt ON PQ fuluicornis (Notman) Genus ERICHSONIUS Fauvel Actobius Fatvel Key to North American species: Frank (192b) E. alumnus Frank _ONPQ E. civicus Frank -oN E. fraterculus (Horn) -oNPQ E. inutilis tHorn.l -oNPQ E. nanus (Horn) AK - NT BC _ONPQ NB NS NF E. patella {Horn r -oNPQ E. pusio tHorn ) Ilrr *oNPQ E. rosellus Frank -oNPQ Genus PHILONTHUS Curtis P. aequalis Horn P. agilis {Gravenhorsilt - BC nlger Melsheimer P. albionicus Mannerheim AK BC P. asper Horn NF P. aurulentus Horn AK BC P. blandus ( Gravenhorst) ON PQ-NS paederinus Sachse pulchellus Melsheimer P. brunneus (Gravenhorst) oN PQ breuis Melsheimer dimidiatus (Say) picatus l\ttby P. bucephalus Horn BC P. carbonarius (Gravenhorst)t AK BC Of,l PQ NB NS NF uorlzs (GyllenhaI) P. caurinus Horn BC P. cephalicus Casey _AB - l,ff - P. (Gravenhorst)t cephalotes - BC yuoN PQ NS NF P. chlorinus Tottenham BC _ NS uiridanus Hotn P. cognatus Stephensf BC ON PQ NS NF fu sci p enni s (Mannerheim) r17 Staphylinidar,r P. concinnus t Gravenhorst)f BC AB SK MB ON PQ NS PE NF oc hro pus ( Gravenhorst) P. confertoides Hatch BC P. confertus f,eOonte P. couleensis Hatch NS__Nr P. crestonensrs Hatch BC P. crotchi Horn BC P. cruentatus (Gmelin)t BC AB _MB ON PQ -NF P. cunctans F.orn NS NF P. cyanipennis (Fabricius)* BC AB _ IYID ON PQ NB NS coe r ule i7:, ennis (Mannerheim) P. debilis (Gravenhorst)t BC ON PQ NF P. dimidiatus C.R. Sahlbergf AB P. distans Horn BC P. duplicatus Bernhauer & Schubert AK YK BC AB basalis LIotn P. ferreipenn:s Horn _BC_ P. fi metarius (Gravenhorst)t NF P. finitimus Cjasey PQ P. flavibasis Oasey ---ec- P. forcipatus .Flatch _BC P. fraternus Fall -YK P. furvus Nordmann NT BC ON pe NB NS -LB NT ca li forn i : us Mannerheim comptus Haldeman P. fusiformis Melsheimer P. gaudens Tlrttenham P. grandicollir Horn P. hepaticus ltrrichson PQ cinctulu:: Melsheimer (?) nanu;; Melsheimer uaricolor Boheman P. hudsonicurr Horn AK P. hyperboreus alaskensis Smetana AK_NT v P. instabilis }Iorn _YK ecABi*Ms-- P. lacustrinuir Scheerpeltz NT _ABSKMBONPQ lacustris Casey P. Iecontei Hr.rrn BC P. leechensis Hatch AK BC ie NF P. lindrothi Smetana -NS NF P. lomatus Erichson BC_ONPQ _NS NF georgianus Sachse P. Iongicornis Stephenst NF P. micans (Glavenhorst)t P. neonatus Simetana -NS IU P. nigritoides Smetana AK P. occidentalis Horn P. opacipennis Notman -:"y -NS NF P. palliatus (Gravenhorst) _NS P. perforatus Fauvel fu P. perversus Horn -sc- P. pettiti Horn _BC P. picicornis lJorn _BC P. politus (Lirrn6)t AK_ _BCAB SK MB oN PQ we Ns NI' horrisi Melsheimer mandibularis Kirby P. protervus Casey -MB P. puberulus Horn BC P. pubes Hor:r BC P. pumilio Ct;.sey _MB P. quadricollis Horn BC AB _Fe P. quediinus Horn -AB _PQ P. rainieri Hatch BC P. rectangulus Sharpt AK_ BC SK MB ON PQ NF tetr ag onoce p halus Notman P. sanguinolentus (Gravenhorst)t BCi P. septentrionis Fall _YK P. sericinus [Iorn -"e P. siegwaldii Mannerheim AK BC ON PQ _LBNF P. sordidus (Gravenhorst)t BC AB SK MB ON PQ NB NS \TET long i pen nis Provancher pachycephalus Nordman 118 Staphylinidae P. spiniformis Hatch _BC P. stacesmithi Hatch _BC P. stictus Hausen PQ P. subvirescens C.G. Thomson* AI( YK NT BC AB PQ P. theveneti Horn AK BC P. triangulum Horn _BC P. umbratilis (Gravenhorst)t -BC NB NS P. umbrinus (Gravenhorst) NS P. varians (Paykull)f NT BC ON PQ NB NF P. vulgatus Casey AK NS Genus GABRONTHUS Tottenham G. thermarum (Aub6)t oN PQ Genus GABRIUS Stephens G. bidentatus (Horn) BC T\TF G. brevipennis (Horn) AK_N' BC AB SK MB ON PQ NB NS G. decipiens (Horn) BC G. downiei (Hatch) BC G. linearis (Casey) BC G. malkini (Hatch) AK__ NF microphthalmus (Horn) BC G. pe G. nigritulus (Gravenhorst)t BC AB - lle oN pumilus Mannerheim G. oceanus (Hatch) BC G. ottawensis (Casey) ON G. picipennis (Maklin) AK YK NT BC AB SK MB ON PQ NE NS -inNr G. seattlensis (Hatch) BC NF G. subnigritulus (Reitter)+ BC _PQ NF G. virilis (Horn) BC G. wawawai (Hatch) BC _-__ Genus CAFIUS Stephens (Subgenus EUREMUS Bierig) C. bistriatus Erichson NB NF bilineatus Erichson rufifrons Bierig C. luteipennis Horn BC (Subgenus BRYONOMUS Casey) C. canescens Miiklin AK _BC C. seminitens Horn _BC (Subgenus REMUS Holme) C. femoralis Meklin AK BC mutatus Gemminger & Harold (Subgenus PSEUDOREMUS Koch) C. nudusjohnsoni Fall BC Genus BELONUCHUS Nordmann B. rufrpennis (Fabricius) ON PQ for mos us ( Gravenhorst) pallipes Melsheimer Genus LAETULONTHUS Moore & Legrer L. Iaetulus (Say) oN PQ Genus HESPERUS Fauvel H. apicialis (Say) ON PQ haernaturus (Erichson) 119 Staphylinide:e Tribe Staphylinini Genus CREOPHILUS Samouelle Saprophilus Streubel C. maxillosuii (Linn6)t AK YK NT BC AB SK MB ON PQ NB NS LBPE NF arcticus Erichson uillosus (Gravenhorst) Genus ONTIIOLESTES Ganglbauer O. cingulatus (Gravenhorst) NT BC AB SK MB ON PQ NB NS _LBNF O. murinus (Linn6)t NF Genus HADIIOTES Meklin H. crassus (I{annerheim) AK BC extensu! LeConte Genus THINOPINUS LeConte Tri c hon r n t h us Motschulsky T. variegatus (Motschulsky) BC Genus PLATYDRACUS C.G. Thomson P. cinnamopterus (Gravenhorst) -oNPQ P. comes (Letlonte) ? P. exulans (Lrichson) -oN P. fossator ((|ravenhorst) MB ON PQ NB immaculatus (Horn) P. maculosui (Gravenhorst) ON PQ ery thr otr ennis (Mannerheim) uiduatu r (Fabricius) P. mysticus ( 0richson) -ONPQ-____ P. rutilicaudrr (Horn) _BC P. tarsalis (N lannerheim) AK _BC sub meta: llicus (Mannerheim) P. tomentosus (Gravenhorst) ON P. violaceus ( Gravenhorst) ON PQ NS cupriper',nis (Melsheimer) P. viridanus Horn) ON PQ NS P. vulpinus ( \ordmann) ON PQ Genus STAP.FIYLINUS Linn6 S. aeneocephllus DeGeerf BC S. ater Gravenhorstt BC Io*pe NS-- NT S. badipes LcConte ix MB ON PQ NB NS carbona',us LeConte ^B S. capitatus 131and BC AB _ONPQ NB NF S. erythropterus Linn6f MB ON PQ NS S. melanarius Heert _ONPQ S. nigrellus Ilorn BC S. pleuralis L,eConte BC S. saphyrinus LeConte BC v- S. winkleri Bernhauert BC Tlibe Xanthopygini Genus TYMF'ANOPHORUS Nordmann Diaphoetes Waterhouse T. puncticollis (Erichson) BC AB oN PQ NB NF borealis (Hatch) Tribe Quediini Key to North American species: Smetana (1971) Genus BEERIA Hatch B. nematocera (Casey) BC t20 Staphylinidae Genus HETEROTHOPS Stephens H. asperatus Smetana _BC H. carbonatus Fall _BC H. conformis Smetana _BC H. fraternus Smetana _BC H. fusculus LeConte _BC AB SK MB ON PQ _NS californicus LeConte fumigatus LeConte H. marmotae Smetana H. minor Smetana N,ecAB-tu3NPe-ns--wr H. pusio LeConte _BC_ONPQ H. sordidus Smetana AK_ NT BC Genus QUEDIUS Stephens (Subgenus MEGAQUEDIUS Casey) Q. explanatus LeConte BC Q. manitobensis (Casey) AB -ll"- (Subgenus MICROSAURUS Stephens) (Anastictodera Casey) (Quediochrus Casey) Q. campbelli Smetana SK oN PQ Q. canadensis (Casey) oN PQ Ne- Q. caseyi caseyi Scheerpeltz --Nt- AB SK MB oN PQ curtipennis (Casey) Q. caseyi divergens (Casey) _BC AB paruipennis Bernhauer uteanus (Casey) Q. charactus Hatch -BC Q. crescenti Hatch _BC Q. criddlei (Casey) AK YK NT BC AB SK MB hoodiHatch Q. erythrogaster Mannerheim _BC AB SK ON PQ neotomae Hatc}' Q. fulgidus (Fabricius)t _BC _MB iracundus (Say) Q. griffinae Hatch BC Q. incisus Hatch BC AB Q. Iimbifer Horn BC _NS Q. mesomelinus (Marsham)t A"- BC AB oN PQ LB NF _YK BC Q. mordax Smetana AB -y" Q. pellax Smetana BC Q. peregrinus (Gravenhorst) ON PQ -NS siluicola Casey Q. pullmani Hatch BC AB SK MB _ Q. rainieri Hatch v_ BC _NS Q. spelaeus spelaeus Horn BC AB SK quadriceps Casey Q. tahomae Hatch BC Q. terminatus Melsheimer Q. velox Smetana AK BC AB (Subgenus QUEDIUS s. str.) Q. curtipennis Bernhauert _BC parallelus Hatch Q. horni Hatch _BC Q. labradorensis insularis Smetana AK. Q. labradorensis labradorensis AK NT BC AB SK MB ON pe---ie NF Smetana Q. lanei Hatch _BC Q. laticollis (Gravenhorst) ON marginatus Korge neomolochinus Korge Q. molochinus (Gravenhorst)t _PQ NF nigribasis Kotge Q. molochinoides molochinoides AKYK_BC oN PQ NF Smetana (Subgenus QUEDIONUCHUS Sharp) Q. plagiatus Mannerheimx AI{ YK NT BC AB MB ON PQ LB NF r2l Staphyiinida: laeuigati ts (Gyllenhal) I ongipen n is Mannerheim ruftpennis Maklin (Subgenus Dl STICHALIUS Casey) Q. capucinus (Gravenhorst) ON PQ ater (Zie1let) bordzs Melsheimer inuersus (.Say) seeuersi Korge Q. frater Smetana _BCAB Q. montivagus Smetana _BC Q. nevadensis (Casey) _YK _BCAB Q. pediculus (Nordmann) AK _BCAB Q. tetricus Smetana _BC Q. transparerLs Motschulsky AK _BCAB marginalis Mdklin pugetensis Hatc}l (Subgenus RI\PHIRUS Stephens) (.Quediel'.us Casey) Q. adnexus Snetana _NF Q. aenescens Maklin AK BC AB agnatus iCasey) Q. breviceps (Casey) AK BC Q. brunnipen nis Mannerheim AKYKNTBCABONPQ_LBNF Q. densiventris (Casey) AKYK-BCAB-PQNF alutaceu; Hatch bonapari:ei }{atc}i' Q. elevatus Hatch _BC Q. felimani (Zetterstedt) AK YK NT BC AB SK MB PQ LB NF hyperboreus Erichson Q. ftigidus Srretana AK-NT_AB ON NF Q. fulvicollis r Stephens)* AKYK_BCAB _MBONPQ_LB NF pugetant.rs (Casey) raphiroides Hatch Q. impar Smctana BC AB Q. nanulus (C asey) _BCAB helenae ( Casey) humilis rtCasey) Q. oculeus (Crrsey) AK BC sparsus (Casey) Q. prostans Horn BC rupimonlis (Casey) Q. rusticus Snetana _BCAB_MB_PQ Q. seriatus H,rrn _BC v_ Q. simulator iJmetana AKYKNT_ABSKMBONPQ - in- Q. sublimbatr.rs Maklinx AKYKNTBCABSKMB_PQ _LBNF orbiceps ICasey) solitarius (Casey) Q. vilis SmetzLna BC (Subgenus P/.RAQUEDIUS Casey) Q. puncticeps Horn BC Genus HEMIiIUEDIUS Horn H. ferox (LeC,rnte) ON PQ NS Genus ANAQIIEDIUS Casey A. vernix (LeConte) ON PQ NB NS Genus ACYL()PHORUS Nordmann (Subgenus AI{ACYLOPHORUS Smetana) A. pratensis l,eConte ON PQ NF (Subgenus ACYLOPHORUS s. str.) A. alternans Smetana -oN A. caseyi Leni; -oNPQ _NS A. flavicollis Sachse -oN A. pronus Eri,chson AK NT BC MB ON PQ NB NS NF r22 Staphylinidae Subfamily XANTHOLININAE Key to North American species: Smetana (1982) Tribe Diochini Genus DIOCHUS Erichson D. schaumi Kraatz _ONPQ breuipennis Casey pallidiceps Casey thoracicus Casey T?ibe Othiini Genus ATRECUS Jacquelin du Val A. americanus (Casey) - ON PQ - NS A. macrocephalus (Nordmann) AK - NT BC AB ON PQ NB NS NF A. quadripennis (Caseyt - BC AB Genus PAROTHIUS Casey P. californicus (Mannerheim.t - BC P. punctatus Smetana - BC Tribe Xantholinini Genus GAUROPTERUS C.G. Thomson G. fulgidus {Fabricius)* - ON Genus NIIDOBIUS C.G. Thomson N. cephalus (Say) AK - NT BC AB SK MB ON PQ NB NS NF assimilis (Kirby) consentane us (Nordmann) flauipennis (Nordmann) giluipennis (Casey) hiesenwetteri (Sachse) nubipennis Casey p alliat us (Melsheimer) N. pugetanus Casey - BC corticallis Casey debilis Casey limbalis Casey Genus LEPTACINUS Erichson Leptacinodes Casey Xa nt ho p hius Motschul sky L. batychrus tGyllenhal)+ - BC MB ON PQ seriatus LeConte L. intermedius Donisthorpet - BC AB SK MB ON PQ NF L. pusillus (Stephens)f - ON PQ nigritulus LeConte Genus PHACOPHAILUS Coiffait P. parumpunctatus tGyllenhal)* - ON PQ cephalicus (LeConte) Genus STENISTODERUS Jacquelin du Val S. rubripennis tleConte) - ON Genus LITHOCIIARODES Sharo Nematolinus Casey Oligolinus Casey L. longicollis (LeContet - ON PQ Genus LINOHESPERUS Smetana L. borealis (Casey) - BC L. cuspifer Smetana - BC AB L. subhamaLus Smetana AB t23 Scaphidiidae Genus STIC'IOLINUS Casey S. flavipes (leConte) oN PQ NS scolopaceus Casey Genus HESF;EROLINUS Casey Leiolinus Casey H. parcus (LeConte) tarsalis (Casey) uteonus (Casey) Genus GYRCTIIYPNUS Samouelle G. angustatus Stephenst _BC_ONPQ NB NS NF G. campbelli Smetana MB ON PQ G. fracticorn:.s (O.F. Mulier)t _BCAB_MBONPQ NB NS NF' obsidiar us (Melsheimer) Genus NEOI|YPNUS Coiffait & Saiz N. beckeri Srretana ON PQ N. emmesus,'Gravenhorst) oN PQ luteiuen; ris (Casey) sang uinolent us (Melsheimer) N. fragilis (C,rsey) _BC ON N. hamatus ( Say) NT BC AB SK IYI D ON PQ NS macilenius (Casey) N. lineatus (lVendeler) ON linearis Blatchley) N. melanops .Casey) oN PQ N. monstrosr- s Smetana ON PQ N. obscurus (Orichson) ---ecABix MB oN PQ N" NS NF oregonul: (Casey) protractils (Casey) N. picipennis (LeConte) _BC Genus OXYBLEPTES Smetana O. kiteieyi Slretana ON PQ NB Genus XANTHOLINUS Dejean X. linearis (Olivier)t R11 NS Genus HYPNOGYRA Casey H. gularis (Lr,rConte) ON uernicaht Casey Genus XEST()LINUS Casey X. abdominalis Casey SK MB ON PQ frontalis Hatc}: indianeri sis Blatchley ouiceps Casey Family SCAPHIDIIDAE shining fungus beetles J.M. Campbell Scaphidiidae is a small family of fungus-feeding consensus among researchers on their proper place- beetles that comprises over 1200 species around the ment. The family Scaphidiidae is poorly known in world, but; only about 50 species occur in North North America, and species recognition of most America: cif these. 17 are recorded in Canada and Canadian taxa is not possible. Casey (1893, 1900) Alaska. IVIost species are found in fungi, but some has completed the most recent revision of the North live in rottrng wood, leaf litter, and under bark. American shining fungus beetles. Althorrgh many recent authors (Lawrence 1982a; Lawrence and Newton 1982) have placed the aK (1); YK (1); NT (1); BC (4); AB (1); SK (2); MB scaphidiids in the Staphylinidae family, there is no (3); oN (13);PQ (10);NF (1) t24 Pselaphidae Tribe Scaphidiini Genus SCAPHIUM Kirby S. castanipes Kirby AK YK NT BC AB MB ON PQ Genus SCAPHIDIUM Olivrer S. obliteratum LeConte -oN S. piceum Melsheimer -oN S. quadriguttatum Say _ONPQ q uadr inota t u m Castelnau quadripustulatum Say T?ibe Scaphisomini Genus SCAPHISOMA Leach S. castaneum Motschulsky pe_ S. convexum Say | - yuBN S. lacustris Casey -:'- S. repandum Casey oN S. rubens Casey _i PQ Genus TOXIDIUM LeOonte T. compressum Zimmermann ON T. gammaroides LeConte ON pe- Genus BAEOCERA Erichson Eubaeocera Cornell Key to Norbh American species: Cornell (1967) B. apicalis LeConte -oN PQ distincta (Blatchley) B. congenera Casey BC SK MB ON PQ speculifera Casey D'\ B. deflexa Casey - rq B. lalsata Achard SK_ONPQ NF concolor auct. youngi (Corr]ell) B. humeralis FaIl B. nana Casey __BC___ oN Fe rubriuentris Casey Family PSELAPHIDAE short-winged mold beetles A. Dauies Eighty-frve species of Pselaphidae are recorded J.L. LeConte, E. Brendel, and T.L. Casey described in Canada and Alaska: however. the members of most North American species in the mid to late this family have been poorly collected because of nineteenth century; their work was subsequently their minute size and the obscurity of their habi- brought together by Bowman (1934). The present tats. They are found chiefly in moist habitats, arrangement of the supraspecific taxa is based on including the edges of bogs and marshes, under the recent world catalog of genera by Newton and bark of dead trees, and especially in leaf litter and Chandler (1989). Chandler (1990o) also published a rotten wood; a few are myrmecophilous. Adults of key to the North American genera. most Pselaphidae are predatory on small inverte- brates such as nematodes, mites, and collembolans; the larvae are seldom seen, and their habits are equally unknown. AK (10); YK (1); NT (3); BC (23); AB (16); SK (9); Jeannel (1955) divided Pselaphidae into six sub- MB (16); ON (56); PQ (38);NB (15);NS (11); pE (2); families, all of which are represented in Canada. LB (1);NF (3) ' l(2) 125 Pselaphidae Subfamilv FARONINAE Tribe Faronini Genus MEGARAFONUS Casey Key to Ncrrth American species: Schuster and Marsh (1958) (Subgenus IIEGARAI'ONUS s. str.) M. ventralis Casey AK BC Genus SONTfMA Casey Rafonus Casey Key to North American species: Marsh and Schuster (1962) S. margemina Park & Wagner AK BC S. parviceps (Maklin) AK BC Subfamilv EUPLECTINAE Trib; Tlogastrini Genus CON,CPLECTUS Brendel Hexirherius Grigarick & Schuster Prorhet:ius Raffray C. canaliculttus (LeConte) PQ syluatit.us (Raffray) Genus ORO ?US Casey Key to Nc'rth American species: Schuster & Grigarick (1960) O. striatus Oasey BC Genus TETIIASCAPIIA Schuster & Marsh T. dentata Park & Wagner BC Genus FOVIIOSCAPHA Park & Wagner F. terracola Park & Wagner BC Tlibe Euplectini Genus THE|IIUM Casey Apothinus Shatp T. cavifrons (LeConte) ON Genus EUP],ECTUS Leach Key to North American species: Wagner (1975) E. acomanus Casey PQ E. confluens LeConte PQ E. duryi Casey PQ NB NS barri P;trk & Wagner E. elongatus Brendel ON E. karsteni (Reichenbach)t --s^- ON PQ PE iowensis Casey E. longicollis Casey ON E. signatus tReichenbach)t SK MB ON pa:) E. silvicolus Chandler NS Genus PYCI{OPLECTUS Casey Key to Ncrrth American species: Wagner (1975) P. congener (Casey) ON tenellus Casey P. interruptrrs (LeConte) ON hudsonicus (Casey) P. linearis (LeOonte) ON irnpressiceps (Casey) Genus RAMECIA Casey Liniolis Grigarick & Schuster R. crinitus (tsrendel) PQ NS 126 Pselaphidae Genus TRIMIOPLECTUS Brendel Key to North American species: Chandler (1990b) T. obsoletus Brendel - ON PQ Genus BIBLOPLECTUS Reitter Key to North American species: Chandler (19906) B. integer (LeConte) MB ON PQ NB NS B. ruficeps tleContet - ON Genus ACTIUM Casey Proplectus Raffray Key to North American species: Grigarick and Schuster ( 1971) A. barri Park & Wagner - BC AB A. retractum Casey AK BC hatchiPatk & Wagner A. wawonaensis Grigarick & Schuster - BC Genus ACTIASTES Casey Key to North American species: Grigarick and Schuster (1971) A. foveicollis (LeConte.l - BC - ON PQ costalis (Brendel) pennsy lu anicus (Bowman) A. globiferum (LeConte) PQ blandurn (Casey) Genus ACTIZONA Chandler A. borealis Chandler - BC AB Genus CUPILA Casey Key to North American species: Grigarick and Schuster (1968) (Subgenus CUPILA s. str.) C. clavicornis (Maklinl AK BC Genus MELBA Casey Key to eastern North American species: Chandler ( 1990b) (Subgenus MELBA s. str.) M. parvr:la tl,eConte) - ON Subfamily BATRISINAE Tlibe Batrisini Genus BATRISODES Reitter Key to western North American species: Grigarick and Schuster (1962) Key to eastern North American species: Park (1947) (Subgenus BABNORMODES Park) B. riparius (Sayt - ON PQ (Subgenus EMPINODES Park) B. albionicus (Aub6) AK BC B. denticauda (Caseyt - BC (Subgenus EXCAVODES Park) B. frontalis tl,eConte) AB - MB ON PQ B. globosus (LeConte) AB ON PQ NB NS B. scabriceps (LeConte) - ON PQ harringtoni (Casey) B. sinuatifrons (Brendelt - ON (Subgenus PUBIMODES Park) B. striatus tl,eConte) - ON PQ aterrimus (Casey) cephalotes (Casey) psotai Park sirnplex (LeConte) I27 Pselaphidae Subfamily GONIACERINAE Tribe Bythinini Genus MAC HAXRODES Brendel M. carinatus (Brendel) ON Genus TYCI{OBYTHINUS Ganslbauer Bythinopsis Raffray T. bythinioides (Brendel) ON PQ NS Tlibe Brachyglutini Genus RYB,I\XIS Saulcy Bryaxis Leach R. clavata tlirendel) ON PQ NB brendeii (LeConte) R. conjunctr tLeContet ON PQ NB NS R. mystica ( asey AB -MBONPQ NB NS R. transversa Fall AK - NT -AB SK MB ON PQ NB NF R. varicornir, t Brendel) ON PQ NB Genus BRAI.)HYGLUTA Thomson Nisa Ci,Lsey B. arguta (Casey) ON B. terebrata (Casey, SK ON PQ intricatt (Casey) perfora o (Brendel) Genus REIC HENBACHIA Leach Key to western North American species: Grigarick and Schuster (1967) R. albionica Morschulskyr AK americt:..na Raffray compar (LeConte) francistana Casey fundata Casey R. arcilera C asey _PQ R. binodiferi,L Casey AK BC AB R. borealis Casey AB - MB ONPQNB_PE R. canadens.s (Brendel) ON R. congener Brendelr _PQ R. corporalis Casey oN PQ R. cyiindrana tBrendel r ON R. distans C,rsey ON R. facilis Ca rey ON R. fusLicorni s Casey BC AB R. kapones (lrigarick & Schuster - BC AB R. peregrinaLor Casey MB oN---- R. propinqu:. rleconte) MB ON PQ NB NS R. rubicundr. (Aubet ON R. scabra (B'endelt ON R. spatulifer Casey - \aK - BC AB SK MB ON PQ NB LB NF Genus NISAXIS Casey N. tomentos..r tAub6, ON cincinnLtta Casey minuta (Brendel) Genus EUTIiICHITES LeOonte E. zonatus (llrendel t ON PQ zimmer,nanni LeOonte Genus DECTTiRTHRON Brendel Key to eastern North American species: Park (1958) (Subgenus DECARTHRON s. str.) D. abnorme (LeConte) ON PQ D. formiceti il,eConte) _ yl"T:"yu ON D. inusitaturn Park _PQ I28 Pselaphidae D. laurenticum Casey ON D. Iongulurn (LeConte) oN PQ D. scarifrcatum Brendel D. stigrnosum Brendel UN Tribe Tychini Genus LUCIFOTYCHUS Park & Wagner Key to western North American species: Park and Wagner (1962) Key to eastern North American species: Park (1956) L. cogrratus (LeConte) L. testaceus (Casey) T---BC--nneoNpeNs--- NF Subfamily PSELAPHINAE Tribe Pselaphini Genus PSELAPHUS Herbst (Subgenus PSELAPHUS s. str.) P. bellax Casey BC AB SK MB ON PQ NB NS P. erichsoni LeConte -oN P. fustifer Casey _AB MB ON PQ NB NS Tribe Tyrini Genus TYRUS Aub6 Pytna Casey T. corticinus (Casey) BC AB T. humeralis tAub6t NT nfn OW Pe Ne-- compdr LeOonte Genus CEDIUS LeConte Key to North American species: Park (1949) (Subgenus SINISTROCEDIUS Park) C. ziegleri LeConte ON Tlibe Ceophyllini Genus CEOPHYLLUS LeConte C. monilis LeConte MB ON Tlibe Tmesiphorini Genus TMESIPHORUS LeConte T. costalis LeConte ON Tribe Ctenistini Genus PILOPIUS Casey P. piceus (LeConte) ON PQ lacustris Casey saginatus Casey Subfamily CLAVIGERINAE Tribe Adranini Genus ADRANES LeConte A. lecontei Brendel _ABSK MB ON A. taylori Wickharn BC_SK t29 Hydrophilicae Superfamily Hydrophiloidea FamilyIIYDROPHILIDAE waterscavengerbeetles R.E. Roughley Hydr'tphilidae forms a large family of mostly species, both aquatic and terrestrial, are scav- aquatic t'eetles; 149 species are known within the engers. Canadiar' and Alaskan borders. The family can be Smetana (1988) has recently revised 1;he taxon- divided into two biologically different groups: one omy of the Canadian species, and his contribution aquatic lnd the other terrestrial. The aquatic was used as the basis for this checklist. species or:cur in a wide range of habitats, but most prefer shallow standing water. The terrestrial AJ{Q0; YK (20); NT (36); BC (81);AB (65); SK (52); species (subfamily Sphaeridiinae) live in most MB (6a); oN (e2); PQ (86);NB (38);NS (33); PE (2); kinds of lecaying organic matter. Adults of most LB (5);NF (24) ' H (11);I (18) Subfamily IfYDROC HINAE Genus HYjI'ROCHUS Leach H. brevitars is Knisch -oN PQ H. currani IV.J. Brown _BCABSX MB ON PQ H. granulatns Blatchley _BC MB ON PQ ouellet, Leech H. neosquarnifer Hellman -oN PQ H. pseudoscuamifer D.C. Miller BC_ MB ON PQ H. scabratus Mulsant :" PQ gibbost.ts Melsheimer H. squamife r LeConte BC AB SK MB ON PQ NB laticollis Notman H. subcupreus Randall PQ Subfamily HELOPHORINAE Key to North American species: Smetana ( 19854) Genus HEL OPHORUS Fabricius (Subgenus ]IELOPHORUS s. str.) H. g'randis I lliger* ON PQ NB NS (Subgenus ITHOPALELOPHORUS Kuwert) H. angustic,rllis d'Orchymont SK MB ON PQ NB NS NF H. auricollisi Eschscholtzx AK YK BC PQ NF angust tIus Mannerheim consimilis Mannerheim inquinttus Mannerheim H. browni Angusx YK NT H. columbia.nus McCorkle _NTBCABSK MB H. electus d'Orch).rnont _NT_ABSK MB corri d'Orchymont H. frosti Srr.etana PQ NB H. furius Srnetana NT BC AB SK MB ON PQ frater ,t\rr:.etana [I. inflectus McCorkle BC MB ON PQ [I. lacustris LeConte AK YK NT BC AB SK MB ON PQ NU lls _LBNF lL ledatus rl'Orchymont BC .AB H. leechi M,:Corkle BC AB SK MB H. linearis ..-eConte _ABON oblitus Shatp pallent Casey H. linearoioes d'Orchlnnont H. lineatus Say _ _ _ :cy:"yuoN pe NB o b solet e s ulcat us Motschulsky uentratis Motschulsky H. marginicollis Smetana -oN H. nitiduloicles d'Orchymont AK YK NT BC AB s^ MB ON PQ NB NF 130 Hydrophilidae H. nitidulus LeConte BC AB SK MB gregolis d'Orchymont H. oblongus LeConte* AK YK NT BC AB SK MB ON PQ H. orchymonti Smetana -oN H. orientalis Motschulskyx _NTBCAB SK MB ON PQ NB NS H. parasplendidus parasplendidus YK NT MB Angus H. sempervarians Angus AK YK NT BC AB SK MB ON PQ NB NS LB NF H. splendidus J. Sahlberg* YK NT (Subgenus GEPHELOPHORUS Sharp) H. sibiricus (Motschulsky)x AK YK NT (Subgenus CYPHELOPHORUS Kuwert) H. tuberculatus Gyllenhal* AK YK BC AB SK MB ON PQ NS scaber LeConte (Subgenus ORPHELOPHORUS d'Orchyrnont) ,, H. arcticus W.J. Brown* NT MB ON Subfamily BEROSINAE Genus BEROSUS Leach Enoplurus Hope B. fraternus LeConte _BC AB SK MB ON PQ cal ifor nicus Motschulsky B. hatchi D.C. Miller NT BC AB SK MB ON PQ B. maculosus Mannerheim AK te sselatu s Motschulsky B. oregonensis D.C. Miller AB SK B. peregrinus (Herbst) ON PQ auritus Melsheimer imma c uI at u s Zimmermann subsignatus LeConte B. striatus (Say) NT BC AB SK MB ON PQ B. stylifer Horn _AB -oN falcatus Shatp Subfamily CHAETARTHRIINAE Genus CHA.ETARTHRIA Stephens Key to northwestern species: Miller (1974) C. atra (LeContet _PQ_NS C. nigrella (LeConte) - BC ,,2 C. pallida tl-eContet - SK nigriceps (LeConte) Subfamily IIYDROBIINAE Tlibe Hydrobiini Genus LACCOBIUS Erichson Key to Canadian species: Gentili ( 1985) L. agilis (RandaII) NT BC AB_MB ON PQ NF L. borealis Cheary AK YK NT BC AB ON v_ L. carri d'Orchymont AB SK ON pacificus D.C. Miller L. cinereus columbianus D.C. Miller AK YK NT BC AB SK MB -PQ L. minutoides d'Orchymont ON PQ NB L. reflexipenis Cheary NT MB ON PQ NB NS reflexipenis Malcolm L. spangleri Cheary SK MB ON PQ NB \TIT spangleri Malcolm L. teneralis Cheary -oN L. truncatipenis D.C. Miller NT AB SK MB ON Genus IfYDROBIUS Leach 131 Hydrophili H. fuscipes llinn6)* AK NT BC AB SK MB ON PQ NB NS _ LB NF H. melaenus (Germar) 5 -oNPQNBNS globostts (Say) Genus SPERCHOPSIS LeConte S. tessellata (Zie gler) ON PQ NS Genus AMFITOR Semenov A. latus (Hcrn) _BC A. scabrosus (Horn) .4K BC AB Genus PAR.{CYMUS C.G. Thomson P. confusus Wooldridge _BC P. despectur; (LeConte) AB SK MB ON PQ P. subcupreus (Say) _YKNTBCAB SK MB ON PQ NB P. tarsalis t).C. Miller _BC Genus CRENITIS Bedel Crenip.lilus Ilorn C. alticola (.lall) BC C. digesta (n,eConte) _AB sK y" PQ NB NS C. maculifrc ns W.J. Brown YK_BCAB 9* C. malkini l).C. Miller BC AB C. monticol,L (Horn) -PQ NS C. morata (l{orn) AK YK NT BC AB -PQ _NF C. paradign.a ( d'Orchymont) BC AB paracy noides (d'Orchymont) C. rufiventris (Horn) BC C. snoqualnLie D.C. Miller BC Genus ANACAENA C.G. Thomson Crenitu.lus Winters A. limbata ( Fabricius)t BC AB SK MB ON PQ NS NF Tribe Helocharini Genus ENOCHRUS C.G. Thomson Key to Ncrth American species: Gundersen (1978) (Subgenus I'INOCHRUS s. str.) E. piceus pir:eus D.C. Miller BC carinalus (Horn) (Subgenus IIETHYDRUS Rey) E. blatchleyr tFallr MB E. californicus (Horn) _BC E. cinctus tSay) _SK MB ON PQ NB limbalis (Melsheimer) E. consors {l,econte) PQ E. consortus Green MB ON E. ochraceus (Melsheimer) Nr ec AB sr MB ON PQ NB NS NF lacustris (LeConte) simplet: (LeConte) E. perplexurr t LeContet MB ON PQ NS fimb r iatu s (Melsheimer) E. pygrnaeus nebulosus (Say) MB ON PQ E. sayi Gunrlersen -oN (Subgenus L,UMETUS Zaitsev) E. diffusus (LeConte) NT BC AB SK MB ON E. hamiltoni (Horn) NT BC AB MB ON PQ NS NF collinus W.J. Brown conjunr:tus (Fall) horniLeech liuidus (Walker) pacificus Leech pyretus Leech r32 Hydrophilidae E. reflexipennis (Zimmermann) PQ NB NS angustulus (Casey) Genus CYMBIODYTA Bedel Key to species: Smetana (1974) C. acuminata Fall - YK NT BC AB SK MB ON PQ - NS C. blanchardi Horn - ON PQ C. chamberlaini Smetana - ON C. columbiana Leech AK BC C. minima Notman - BC MB ON PQ C. pacifica Leech - BC hatchi D.C. Millet C. semistriata (Zimmermann) - ON PQ NB C. toddi Spangler MB ON C. vindicata Fall NT BC AB SK MB ON PQ NB NS - LB NF Genus HELOCOMBUS Horn H. bifidus {LeContet - ON PQ NB Subfamilv IIYDROPHILINAE Genus TROPISTERNUS Solier T. columbianus W.J. Brown - BC AB SK - ON T. ellipticus (Leconte) - BC ofizis Motschulsky concolor Sharp T. glaber lHerbst) - ON PQ - NS T. lateralis humeralis Motschulsky - BC T. lateralis marginatus Motschulsky - BC AB binotatus Walker limbalis (LeConte) T. lateralis nimbatus (Say) AB - MB ON Pe - NS T. mixtus (Leconte) MB ON pQ - NS T. natalor d'Orchyrnont - ON PQ Genus HYDROPHILUS O.F. Miiller H. triangularis Say - BC AB SK - ON pe subsulcatus LeConte Genus DIBOLOCELUS Bedel D. ovatus (Gemminger & Haro1d) - ON pQ oualis (Ziegler) Genus HYDROCHARA Berthold Key to species: Smetana (1980) H. brevipalpis Smetana - ON H. obtusata {Say) NT BC AB SK MB ON pQ H. simula Hilsenhoff& Tracy - ON Subfamily SPHAERIDIINAE Tlibe Sphaeridiini Key to North American species: Smetana (1978) Genus PHAENONOTUM Sharp P. exstriatum (Say) -oN dubium S}ratp sem ig I obos um (Zimmermann) Genus SPHAERIDIUM Fabricius S. bipustulatum Fabriciust _BCABSK_ONPQNBNS S. lunatum Fabriciust _BCABSKMBONPQNBNSNF S. scarabaeoides (Linn6)t _BCABSKMBONPQNBNSNF crendtum Palisot de Beauvois 133 Hydrophilidae Tribe Cercyonini Genus CERCYON Leach (Subgenus PROSTERCYON Smetana) C. roseni Knlsch NT BC AB SK MB ON PQ NB NS opacellus Fall (Subgenus CERCYON s. str.) C. adumbrat us Mannerheim AX BC unicolor' (Motschulsky) C. assecla Slnetana AB -oN PQ NB NS C. atricapilh.Ls (Marsham)t AB PQ NS mundut Melsheimer C. cinctus Slretana NT BC AB SK MB ON PQ C. connivens Fall -oN C. erraticus i]metana _BC MB ON PQ C. fimbriatus Mannerheim AK _BC C. haemorrhoidalis (Fabricius)t -BC -oN PQ NB NS PE NF basillaris Notman nigricol,t.is (Say) C. herceus frigidus Smetana AK YK NT BC AB SK MB ON PQ C. herceus h':rceus Smetana BC AB SK MB ON PQ C. impressus (Sturm)t BC C. indistinct,rs Horn _PQNB C. lateralis ( Vlarsham)-i NT BC AB ON PQ NB NS NT incrematus Notman C. limbatus . vlannerheim AK YK NT BC AB ful uiper n i s Mannerhei m C. litoralis (( iyllenhal)x PQ NB NS C. luniger Ma.nnerheim AK BC __I" C. marinus (j.G. Thomson* AK 1T( NT BC AB SK MB ON PQ C. matthewsi Smetana AK_AB C. mendax Smetana -oN C. nevadanus Knisch C. praetextat;us (Say) MB ON PQ NS testaceus Blatchley C. pygmaeus (Illiger)+ BC AB SK MB ON PQ NB NS NF apicalis (Say) C. quisquilius (Linn6)t BC AB SK MB ON PQ NB NS NF C. terminatus { Marsham;f -oN PQ nozzs llelsheimer C. tolfrno Habch AK BC C. unipunctu tus (Linn6)l BC AB SK MB ON PQ NB NS (Subgenus DICYRTOCERCYON Ganglbauer) C. ustulatus ( Preyssler)* PQ (Subgenus P.ARACERCYON Seidlitz) C. analis tPaykullt* NT BC SKAB MB ON PQ NB NS NF falliWi:fterc maculalus Melsheimer C. minusculus Melsheimer BC SK MB ON PQ NB NF Genus TECT OSTERNUM Balfour-Browne Genyon Smetana T. naviculare (Zimmermann) - ON PQ Genus DELI'OSTETHUS Sharo Ictinosternum Smetana ,, D. columbier.sis ( Hatcht Tribe Megasternini Genus MEGASTERNUM Mulsant M. obscurum (Marsham)t _BC M. posticaturn (Mannerheim) AK BC pugetense Halch 134 Histeridae Genus CRYPTOPLEURUM Mulsant C. minutum (Fabricius)t _BCAB MB ON PQ NB NS PE -NF C. subtile Sharpf _BCAB SK MB ON PQ _NF Family GEORYSSIDAE minute mud-loving beetles R.E. Roughley Georyssidae includes minute beetles, usually mud attached to the body; they are apparently algal under 2 mm long; two species are known for the feeders. Nearctic region, one of which occurs in Canada. Adults are found in mud or sand at the edges of streams and are often concealed by sand grains or BC (1);AB (1);PQ (rl Genus GEORYSSUS Latreille G. ousillus LeConte BC AB PQ Family SPHAERITIDAE false clown beetles R.E. Roughley Sphaeritidae contains only one genus, organic material (e.g. fungi) and on trees when the Sphaerites, with two species in the Holarctic sap runs. region; one of them occurs in Canada and Alaska. Adults are found in dung and other decaying eK (1); BC (1);AB (1) Genus SPHAERITES Duftschmid S. politus Mannerheim AK BC Family HISTERIDAE hister beetles A. Dauies In Canada and Alaska 118 species of Histeridae treatments of the North American Histeridae. The have been recorded. The adults and larvae are chiefly classification used here is based on Arnett (1968), predators of other insects and their larvae. Most whereas the status of the species is based on Mazur species are associated with decaying organic material, (1984) and on work by Helava (personal communica- dung, and carrion; some are found in sandy areas, in tion). mammal burrows, under bark, and in bark-beetle galleries; a few are myrrnecophilous. Although some species are associated with stored products, they most likely feed on other insects that occur there. AK (1); YK (2); NT (2); BC (57); AB (42); SK (25); LeConte (1845), Marseul (1853-1857, 1860-1862), MB (43); oN (65);PQ (58);NB (13);NS (12);PE (3); and Horn (1873) have conducted major taxonomic NF(7) . H(1);I(10) Subfamily ABRAEINAE T?ibe Abraeini Genus ABRAEUS Leach A. bolteri LeConte BC 135 Histeridae Tribe Plegaderini Genus PLEGADERUS Erichson P. comonfor.i Marseul - BC uergran.dis Casey P. consors Horn - BC egenus Casey P. fraternus Horn - BC P. molestus Casey - BC obesus Casey P. nitidus Horn - BC P. sayi Mars,eul - BC - ON PQ NB NS P. setulosus Ross BC P. transversus (Say) ON PQ NF Tribe Acritini Genus ACR.ITUS LeConte A. exiguus I lrichson I - ON PQ acicula ius (J.E. LeConte) cribriptnnis Marseul elliptictrc Casey natchez Marseul obliquu s (J.E. LeConte) subdep,'essus (Blatchley) A. nigricorn s lHoffman r-l' - ON PQ Genus AELIITES Horn A. politus tleContet - ON PQ A. simplex t,l.E. LeConter - ON PQ acupict ts (Marseul) Tlibe Teretriini Genus TERIITRIUS Erichson T. american rs J.E. LeConte -oNPQ dissonans Marseul picipes J.E. LeConte T. montanus Horn _BC Subfamily SAPRININAE Genus SAPIIINUS Erichson S. discoidalir LeConte - BC amplus Casey latiforn,.is Casey mayhewi Papp S. impressusr J.E. LeConte - BC AB ON infaustus LeConte piceus ., .8. LeConte S. lugens Errchson NT BC AB SK MB ON PQ co I iforn icus Mannerheim consimilis Waiker disting uendoides Hatch oregonensoides Hatch p se udoclete r sus Dahl gren sejunctoides }Jatch S. oregonensis J.E. LeConte - BC AB SK MB ON PQ distinguendus Marseul lecontecides Hatch p s e u do cl i s t ing u e ndu s Hatch p se udo s ej unc toide s Hatch pseudosejunctzs Hatch sejunctus Marseul spurcus LeConte S. pensylvar-"icus tPaykull) MB ON PQ profusoi.des Hatch 136 Histeridae S. sedakovii Motschulsky* YK Genus EUSPILOTUS Lewis (Subgenus HESPEROSAPRINUS Wenzel) E. assimilis (Paykull) MB ON PQ NB extraneus (Illiger) sardous (Flach) semisulcus (Hatch) simulatus (Blatchley) E. conformis (J.E. LeOonte) -oN ouiformis (Blatchley) E. cribrum (Casey) BC AB SK MB carri (Hatch) E. insertus (LeConte) BC AB SK obductus (LeOonte) opacellus (Casey) uinctus (LeConte) (Subgenus NEOSAPRINUS Bickhardt) E. scrupularis (J.E. LeConte) BC Genus XEROSAPRINUS Wenzel X. fimbriatus (LeOonte) NT BC AB SK desertorum (Marseul) X. lubricus (LeConte) _BC AB SK MB rotundiftons (Marseul) Genus GEOMYSAPRINUS Ross (Subgenus PRISCOSAPRINUS Wenzel) G. castanipennis (Fall) AB SK yu_ G. cheyennensis (Casey) _BCAB G. copei (Horn) _BCAB G. lanei (Hatch & McGrath) _BC G. laramiensis (Casey) -BCAB G. obscurus (LeOonte) pr]) G. obsidianus (Casey) _ _ T_ -oN moniliatus (Casey) (Subgenus GEOMYSAPRINUS s. str.) G. rugosifrons (FaIl) MB Genus HYPOCACCUS C.G. Thomson H. aeneipunctatus (Horn) -BC H. bigemmeus (LeOonte) H. bigener (J.E. LeConte) - - :'T 3* MBor.pe NS: eriensis (Hatch) - illinoensis Wolcott impunctellus ( Casey) lakensis Blatchley michiganensis (Hatch) ohioensis (Hatch) ontarioensis (Hatch) sphaeroides (J.E. LeConte) H. estriatus (LeConte) BC AB H. frtchi (Marseul) PQ H. fraternus (Say) BC AB SK MB ON PQ NS PE NF H. iris (Fall) AB SK MB ON PQ alutiger Wenzel H. mancus (Say) MB H. omissus (Casey) AB SK MB H. patruelis (J.E. LeConte) AB_MBONPQ jaueti (Marceul) H. seminitens (LeConte) MB ON PQ Genus BAECKMANNIOLUS Reichardt B. dimidiatipennis (J.E. LeConte) PQ NB NS NF deserticola (Marseul) desertorum (Marseul) palmatus (Say) L37 Histeridae Genus NEOI'ACHYLOPUS Reichardt N. sulcilrons tMannerheim.l - BC Genus GNA'IIHONCUS Jacquelin du Val G. rossi Hat< h - BC G. rotundatus (Kugelann)f - BC AB SK MB ON PQ NF communis (Marseul) deletus (J.E. LeConte) nanus (Scriba) Subfamilv DENDROPHILINAE Tri6e Dendrophilini Genus DENI )ROPHILUS Leach D. punctatus (Herbsu+ - BC MB ON PQ punctulatus (Say) D. xavieri M rrseult - BC - ON sexstria;lus Hatc}a Tlibe Bacaniini Genus BACFNIUS LeOonte B. punctifon:ris (LeConte) _ONPQ Tribe Paromalini Genus CARC INOPS Marseul Carcinus Marseul C. pumilio (Etrichson)t - BC AB SK MB ON PQ NF nanus Ql.E. LeConte) q uatuordecimsfrjolzs (Stephens) Genus XESTIPYGE Marseul X. conjunctu n tSayt - ON X. geminaturn tJ.E. LeConte) - ON Genus PLATYLOMAIUS Cooman P. aequalis (iiayt - BC AB - MB ON PQ affinis (,1.8. LeConte) estriatu.t (J.E. LeConte) Genus PARC,MALUS Erichson P. bistriatus Erichson AB ON PQ NB - NF P. mancus C,lsey - BC AB P. seeversi (\Venzel; - ON P. teres LeConte - ON PQ NB - NF Subfamilv TRIBALINAE Genus EPIEIRUS Erichson E. pulicarius Erichson - ON PQ minor J,8. LeOonte E. regularis iPalisot de Beauvois) - ON nigrellus (Say) uicinus LeConte Genus ONTIIOPHILUS Leach O. deflectus .l{elava - ON Genus CAEF|OSTERNUS LeConte C. americanus U.E. LeConte) - ON 138 Histeridae Subfamily HISTERINAE Tribe Histerini Genus ATHOLUS C.G. Thomson Peranus Lewis A. americanus {Paykull ) AB SK MB ON PQ diffractus (Casey) exard.tus (J.E. LeConte) perplexus (LeConte) A. bimaculatus ( Linn6)t BC AB -oN PQ obliquus (.Say) A. falli tBickhardtt BC AB SK MB ON PQ gagates Fatl A. sedecimstriatus { Sayl MB ON PQ Genus MARGARINOTUS Marseul (Subgenus MARGARINOTUS s. str.) M. guttifer Horn _AB (Subgenus PTOMISTER Houlbert & Monnot) M. brunneus (Fabricius;f ON PQ NB NS PE cadau e r inu s (Ho{frnann) impressus (Fabricius) obtusatus (Harris) uirginiae (Casey) M. egregius {Casey) -MBONPQ M. foedatus (J.E. LeConte) NB NS M. fractifrons {Caseyt -:nc- _yuoNPQ fidelis (Casey) planifrons (Lewis) M. harrisii (Kirbyt BC AB SK MB ON PQ repletus (J.E. LeConte) M. hudsonicus (Caseyt MB ON PQ NB M. immunis (Erichson) BC MB ON PQ ols--- M. intermptus (Palisot de Beauvois) -oNPQ NB M. merdarius (Hoffmann)t nc _ONPQ NS memnonius (Say) M. pluto (Casey) BC AB MB albertensis (Hatch) carri (}{atc}r) M. rectus (Caseyt BC M. umbrosus (Casey) BC umbratilis (Casey) (Subgenus STENISTER Reichardt) M. obscurus (Kugelann)f BC semisculptus (LeConte) stercorarius (Hoffmann) (Subgenus PARAIISTER Bickhardt) M. cognatus (J.E. LeConte) MB ON PQ unicus (Casey) M. lecontei Wenzel SK MB ON PQ NB NS PE M. purpurascens (Herbst)t --nc- (Subgenus PROMETHISTER Kryzhanovsky) M. confusus Wenzel -oN PQ M. marginicollis (J.E. LeConte) MB ON PQ Genus HISTER Linn6 H. abbreviatus Fabricius AK BC AB SK MB ON PQ NB bifidus Say coloradensis Casey crenicollis Flach subhernisphaericus Palisot de Beauvois H. albertanus Casey AB H. curtatus J.E. LeConte PQ NS haldemanni Marseul 139 Histeridae H. defectus .f.E. LeConte SK -oN PQ permixlus Zimmermann H. depuratol Say BC SKAB MB ON PQ NB NS paykulliKirby perbreu.'.s Casey spretus J.E. LeConte Genus SPILODISCUS Lewis S. arcuatus (Say) NS S. militaris (Horn) -BC electus (Casey) oregonus (Casey) simpliupes (.Fa\l) S. sellatus (LeOonte) BC senneui lei (Marseul) S. semiruber (Casey) BC AB SK MB soloris t Carnochan) Genus PSILr)SCELIS Marseul P. abnormal s Mann SK MB P. carri Hatch _AB P. corrosa Casey _AB P. planipes (LeConte) MB ON PQ_NS harrisi:J.8. LeConte) P. subopaca LeConte) YK BC AB MB Tribe Platysomatini Genus PLA1YSOMA Leach (Subgenus PLATYSOMA s. str.) P. aequum (l,eConte) ON PQ P. aurelianu n (Horn) PQ P. lecontei Nlarseul BC AB _MBON PQ d.epress tm J.E. LeConte tabellurt Casey (Subgenus EURYLISTER Bickhardt) P. carolinum (Paykull) oN PQ cinnamomeum (A. White) sordidum (Say) (Subgenus CYLISTER Cooman) P. coarctaturn J.E. LeConte -BCAB yuoNPQ P. punctigemm (LeConte) _BCAB _ Genus CYLISTIX Marseu. C. cylindrica (Palisot de Beauvois) C. deficiens Casey C. gracilis (J.E. LeConte) _ _ rsxy"giliH _ _ _ _ _ Tribe Hololeptini Genus HOLOLEPTA Paykull H. aequalis Say BC AB MB ON PQ fossularis Say Tribe Exosternini Genus PHEL,ISTER Marseul P. subrotunius (Say) oN PQ P. vernus (Say) ON saunieri Marseul I40 Clambidae Subfamily HETAERIINAE Tribe Hetaeriini Genus HETAERIUS Erichson H. brunneipennis (Randall) -oN H. exiguus Mannerheim BC nitidus Martin H. tristriatus Horn BC AB SK MB Superfamily Eucinetoidea Family EUCINETIDAE plate-thigh beetles J.M. Campbell Eucinetidae is a small family that contains Horn (1880) last reviewed the species of this about 30 species throughout the world; seven of family, as part of Dascillidae; they are poorly known them, placed in one genus, are known to occur in taxonomically. Canada. These beetles are associated with fungal decay and are found in leaf litter or under bark of vK (1); NT (2); BC (5); eB (3); SK (3); MB (4); ON fungus-infested logs. (a);PQ (a);NB (2);NS (2) . I (1) Genus EUCINETUS Germar E. haemorrhoidalis (Gerrnar)t BC AB SK MB oN PQ NB NS E. infumatus LeConte _v BC E. morio LeConte ON PQ _NS E. oviformis LeConte MB E. punctulatus LeConte YK_ SC AB sx MB ON PQ E. terminalis LeConte BC E. testaceus LeConte _NT BC AB SK MB oN PQ NB Family CLAMBIDAE minute beetles J.M. Campbell Clambidae forms a family of small beetles that decaying vegetation, where they probably feed on are rarely collected because of their size and habi- spores of fungi and slime molds. tat. The family contains five genera and about 50 The American species were revised by Endrridy- species in the world. Only two genera comprising Younga (1981) and are well-known taxonomically. seven species are known to occur in Canada and Alaska, but more are likely to be found. Adults are usually collected during their evening flight. They AK (1); BC (3); AB (3); oN (1); PQ (6) o I1[ can also be collected by sifting leaflitter and piles of (1); I (1) Genus CALYPTOMERUS Redtenbacher C. oblongulus Mannerheim AK BC Genus CLAMBUS Fischer von Waldheim Sternuchus LeConte C. armadillo armadillo (DeGeer)t PQ niger Hatch C. gibbulus (LeContetf, BC AB PQ r adul a Endr }dy-Younga C. howdeni Endrddy-Younga ON PQ r47 Scirtidae C. pubescens Redtenbacher* BC AB PQ puberu.lus LeConte C. smetanai Endrcidy-Younga PQ C. vulneratus LeConte PQ Family SCIRTIDAE marsh beetles J.M. Campbell The lamily Scirtidae contains about 600 species sion. Horn (1880) last reviewed the species of this throughout the world, but most of them are found in family, as part of Dascillidae. Klausnitzer (1976), temperai;e zones. Six genera and 26 species are however, reviewed several species occurring in known tc occur in Canada and Alaska. Adults are Canada; he (1980) also segregated some species for- collected, often in large numbers, on vegetation in merly assigned to Elodes to form the new genus wet habitats, particularly marshes, bogs, and Flauohelodes. swamps; they are attracted to light. Larvae are aquatic and feed on fungi, algae, diatoms, and other organic nratter. Most Canadian species are well known except AK (1); YK (1); NT (2); BC (10); AB (a); SK (3); MB those of the genus Cyphon, which is in need of revi- (3); oN (15);PQ (15);NB (7);NS (3);NF (5) Genus ELCTDES Latreille Hemic;tphon LeConte Sacodt:s LeConte E. angusta Hatch _BC E. apicalis 1,eConte _BC E. emargin;r.ta Hatch _BC E. maculicc'llis Horn PQ_NS NF Genus FLAVOHELODES Klausnitzer F. fuscipennis (Gu6rin-M6neville) ON F. pulchella (Gu6rin-M6neville) ON F. thoracica. (Melsheimer) ON Genus MICROCARA C.G. Thomson M. explanat;a LeConte BC SK MB ON PQ NB Genus CYPHON Paykull C. brevicoll:.s (LeConte) BC C. collaris (rGu6rin-M6neville) NB C. concinnus (LeOonte) BC AB- - - :a C. confusus W.J. Brown _ONPQ NF horiont Klausnitzer C. exiguus :1orn BC C. modestu'; (LeConte) --pe NF' C. nebulosus (LeConte) -oNPQ C. neovaria bilis Klausnitzer _ONPQ NS- C. obscurus (Gu6rin-M6neville) _ONPQ Ns NF C. ochreatus Klausnitzer yu oN PQ fu C. punctatr:.s (LeConte) BC C. pusillus,.LeConte) _NT BC AB SK C. rufrcollis (Say) _ONPQ C. variabilis (Thunberg) AK YK NT BC AB SK MB ON PQ lle Ns NF Genus PRIONOCYPHON Redtenbacher P. discoideus (Say) oN PQ P. limbatus LeConte oN PQ NB Genus SCIIITES Illiger S. orbiculatus (Fabricius) oN PQ S. tibialis Gu6rin-M6neville AB ON PQ NB t42 Lucanidae Superfamily Dascilloidea Family DASCILLIDAE soft-bodied plant beetles J.M. Campbell The status of Dascillidae has changed consider- occur throughout the world. The adults are usually ably since Horn reviewed it (1880). Its size has found on foliage; they feed on plants or are carnivo- been greatly reduced by the r_emoval of species now rous. The larvae live in soil or are aquatic; they feed included in the families Scirtidae, Eucinetidae, on vegetative matter such as roots or leaf mold. Artematopidae, and Ptilodactylidae, so that only one species is now known to occur in Canada. Fifteen genera and about 80 species are known to BC (1) Subfamily DASCILLINAE T?ibe Cneoglossini Genus ARAIOPIDIUS Cockerell Araeopus LeConte A. monachus (LeConte) BC Family RHIPICERIDAE cedar beetles J.M. Campbell Rhipiceridae contains five genera and about 50 The adults are often found in mating congregations species throughout the world. Only one genus with on the trunks of trees. with many males grouped one species is known to occur in southern Ontario. around one female. The larvae of this species live on the roots of trees, where they are parasitic on the nymphs of cicadas. oN (1) Genus SANDAI-US Knoch S. niger Knoch ON hnochi (Gt6in-M6neville) rubidus Melsheimer r uftpennis (Latreille) scabricollis Haldeman Superfamily Scarabaeoidea Family LUCANIDAE stag beetles J. McNq.mara Lucanidae includes 14 species in Canada. The Howden and Lawrence (1974) published a key larvae of all species breed in or beneath the decay- of the North American genera, and since the genera ing wood of logs or stumps and therefore are of little occurring in Canada include few species, specific economic importance. The adults are believed to identifrcation is relativelv easv. feed on honeydew or sap. They lay their eggs in bark crevices, especially near the roots. The adults are attracted to light and are most commonly found vK (1); NT (2); gC (7); AB (2); SK (2); MB (3); ON in wooded areas. (8);PQ (5); NB (2);NS (3);NF (1) r43 Lucanidae Subfamilv SYNDESINAE Tribe Sinodendrini Genus SINODENDRON Hellwig S. rugosum Mannerheim - BC Tlibe Ceruchini Genus CERUCHUS Macleay C. piceus (V/eber) MBONPQ-NS bolbi (t.lastelnau) uirginiznsis Casey C. punctatus LeConte BC C. striatus LeConte _BC Subfamily NICAGINAE T?ibe Nicagini Genus NIC,\GUS LeConte N. obscurus tleConter ON PQ Subfamily LUCANINAE Tribe Lucanini Genus LUC ANUS Scopoli (Subgenus LUCANUS s. str.) L. elaphus I'abricius - ON carleng:i Angell (Subgenus I'SEUDOLUCANUS Hope & Westwood) L. capreolus tl.inndt - ON domo l:'abricius muticu.; Thunberg nigricephalus (Benesh) trigonu:.s Thunberg L. placidus iay - ON lentus \)astelnat Tribe Dorcini Genus DORCUS Macleay D. parallelu* {Sayt - ON PQ carnocl,,ani Angell costatu,s LeConte nanus Casey oblongus (Charpentier) uoeti (Schonherr) TYibe Platycerini Genus PLA',YCEROPSIS Benesh P. keeni (Casey) BC pedicelt aris (Moilenk) Genus PLAIIYCERUS Geoffroy Key to species: Benesh (1946) P. marginalis Casey NT BC AB SK MB NB NS P. oregonensis Westwood - BC chalybo:eus Casey coerulescens LeConte P. piceus (K,rby) - \'K NT BC AB SK MB ON PQ NB - NF depressus LeConte 144 Scarabaeidae virescens (Fabricius) oN PQ NS angustus casey iowanus Casey peregrinus Casey quercus (Weber) securidens (.Say) Family PASSALIDAE peg and b,ess beetles J. McNamara Only one species of Passalidae occurs in Reyes-Castillo (1970) published key to the Canada. These beetles, which live in rotten logs American genera of this family. and stumps, are not economically important. The adults and larvae communicate with each other by stridulation. The adults prepare food for the larvae by chewing the wood and mixing it with saliva. MB (1);ON (1) Tribe Proculini Genus ODONTOTAINIUS Kuwert Passalotaenius Kuwert O. disjunctus (Illiger) NtB ON cornutus (Fabricius) Family SCARABAEIDAE scarErb beetles J. McNamara Scarabaeidae is one of the largest families of Mating taker; place during the summer. The eggs beetles in the world; 248 species are known to occur are laid in the soil in late July and early August; in Canada and Alaska. Beetles of this family are they hatch immediately, and the larvae begin to popular with collectors because of their bright colors feed. When bhe weather turns cold the larvae bur- and the large horns of some groups. The size of row deeper into the soil, below the frost line, where these beetles varies greatly, from 1.9 mm for they hibernate. In the spring when the weather is Pleurophorus longulus to 15 cm for Goliathus, warmer, they move up near the surface of the which is probably the largest insect known based on ground and continue to feed until they pupate in mass. The scarabs contain some of the most June. The adults emerge about the first of July, destructive beetles, such as the Japanese beetle depending on the area. Many variations occur in (Popillia japonica), the Asiatic garden beetle the life cycle of scarabs. The Phyllophoga species (Maladera castanea), and the European chafer cycle ranges firom 2 to 4 years, depending largely on (Amphirnallon majalis). These belong to a group of the latitude. scarabs that feed primarily on Iive plant matter as John Cooper is currently revising the scarabs of adults, and usually on plant roots or rotting wood as Canada and Alaska, and his work is expected to be larvae. The other group of scarabs are mainly dung published as part of The Insects and Arachnids of feeders or scavengers. They feed primarily on dung Canada, a serries published by Agriculture Canada. or decaying plant and animal matter as adults, and The classifrcation used here is taken almost entirely in many cases the larvae feed on the same material from his manuscript. as it is supplied to them by the adults. Because of their economic importance, the life cycle of many AK (13); YK (9); NT (17); BC (88); AB (96); SK (58); species is fairly well known. That of the Japanese MB (85); ON (136); PQ (109); NB (a1); NS (a7); PE beetle is probably typical of the family in general. (11);LB (7);ltIF (20) . H (1); I (21) 145 Scarabaeidae Subfamilv SCARABAEINAE Tribl Scarabaeini Genus CAIITHON Hoffmansegg Key to North American species: Robinson (1948) (Subgenus CANTHON s. str.) C. chalcites (Haldemanr ON C. pilularir:s {Linn6) - AB hudsonias (Forster) laeuis i.Dtury) obtusi'fens (Ziegler) uiridetcens Horn uoluen; (Fabricius) C. vigilans LeConte - ON (Subgenus BOREOCANTHON Halflter) C. praticok LeConte BC A-B SK MB C. simplex Leconte - BC AB (Subgenus ILAPHYROCANTHON Martinez) C. viridis 1l)alisot de Beauvois) - ON metallicus Stutrn obsoletus (.Say'1 uiridicatus (Say) Tribe Coprini Genus COF'RIS O.F. Miiller Key to New World species: Matthews (1961) C. lricator lricator rFabricius) - ON PQ anagllpticus Say tullius (O\ivter) C. minutus (Drury) PQ ammo:l, Fabricius lar Fa.:ricius reflexu:.s Panzer silenzs Fabricius Tlibe Onthophagini Genus ONTHOPHAGUS Latreille Key to Nrrth American species: Howden and Cartwright (1963) O. hecate hecate (Panzer) - BC AB SK MB ON PQ NB NS furcicollis Dejean hastator (Fabricius) lamal)ejean latebrosus (Fabricius) obtectt:s (Palisot de Beauvois) soyi C;rstelnau scabricollis l{rtby O. nuchicornis (Linn6)t - BC AB - MB ON PQ NB NS NF r hinoc'zro s Mel sheimer xiphias LeConte O. orpheus canadensis lFabriciusr - ON PQ O. orpheus pseudorpheus MB Hou'den & Cartwright O. pennsyl' anicus Harold - ON PQ Subfamily APHODIINAE Tribe Aegialiini Key to North American species: Gordon and Cartwright (1988) Genus MICRAIGIAJ-IA W.J. Brown M. pusilla (Horn) - BC AB - MB I46 Scarabaeidae Genus AXGIALIA Latreille (Subgenus LEPTAIGIALIA W.J. Brown) A. humeralis W.J. Brown -oN PQ A. montana W.J. Brown _BCAB A. rufescens Horn _BCABSK MB ON PQ NB (Subgenus PSAMMOPORUS Thomson) A. criddlei W.J. Brown AK BC AB SK MB ON PQ NF A. cylindrica (Eschscholtz) AK BC A. lacustris LeConte AK YK NT BC AB SK MB ON PQ NB NS A. nana W.J. Brown PQIU___ )u A. opaca W.J. Brown _BCAB A. terminalis W.J. Brown AK YK NT BC AB SK MB LB NF (Subgenus RHYSOTHORAX Bedel) (Anomalaeg ial ia W . J . Brown) A. spissipes LeContef oN PQ NS NF rzlo (Fabricius) rufina Sllfvergerg (Subgenus AXGIAJ-IA s. str.) A. arenaria (Fabricius)t NS A. blanchardi Horn _BC PQ A. carri Gordon & Cartwrieht AB A. conferta Horn NtrD r\I\T PQ A. crassa LeConte _ _ _ ecf:" insularis W.J. Brown A. opifex Horn oN PQ NS PE Tribe Aphodiini Genus APHODIUS illiger Key to eastern North American species: Gordon (1983) A. aenictus Cooper & Gor'lon ON A. albertanus W.J. Brown NT BC AB A. aleutus Eschscholtz _BCAB ursinus Motschulsky A. alternatus Horn BC SK MB inermis Robinson moclipsiHaLch A. anthracinus LeConte _ ^AB A. aquilonarius W.J. Brown BCTSKIo*pe----- A. badipes Melsheimer oblongus Say A. bicolor Say PQ A. bidentatus Schmidt BC AB __9* bidens LeConte A. borealis Gyllenhalx BC AB _MBON PQ NB NS LB NF errons W.J. Brown A. brevicollis LeConte AB y"_ A. browni Hinton -BC _ smithiW.J.Brcwn A. bryanti W.J. Brown AB SK A. carri W.J. Brown AB SK A. coloradensis Horn AB A. congregatus Mannerherm AK ec AB arcticus Harold A. consentaneus LeConte NT AB SK MB A. cormptor W.J. Brown AK \'K BC AB SK MB A. cribratulus Schmidt BC cribratus LeConte A. criddlei W.J. Brown AB yu_ A. decipiens Horn ___sc AB :" A. denticulatus Haldeman AB A. distinctus (O.F. Miiller)t _BC AB SK MB ON pe NS inquinatus (Herbst) m a eu I i pe n n i s Mel sheimer A. duplex LeConte BC AB SK MB A. edmontonus W.J. Brown AB I47 Scarabaeic ae A. erraticus (Linn6)t MB ON PQ NB NS NF pensuullensis Melsheimer A. fimetarius (Linn6)-f BC AB SK MB ON PQ NB NS LB NF nodifrons Randall A. fossor (Linn6)t AK BC AB SK MB ON PQ NB NS PE LB NF A. granarirLs (Linn6)t _BCAB _MB ON PQ NB NS aterr i;' n u s Melsheimer inutilt:s Horn metalticus Haldemarr spretu.s Haldeman A. guttatus Eschscholtz AKYK_BCAB _MB PQ _NS A. haemorrhoidalis (Linn6)i _BCAB SK MB ON PQ NB NS _LB NF A. hamatus hamatus Say _BCAB SK MB PQ occidentalis Horn sagittarius Harold A. hirsutus W.J. Brown BC A. hyperboreus LeConte NT BC AB SK MB ON PQ NB NS NF anguloris LeConte A. iowensis Wickham AB SK MB A. lentus Eorn ON PQ A. leopardr.Ls Horn AK YK NT BC AB SK MB ON PQ NN NS A. leptotarr;is W.J. Brown AB TU A. manitobensis W.J. Brown AB _MB ON PQ _NS A. omissus omissus LeConte YK BC AB SK MB ON A. omissus torpidus Horn YK f' BC AB SK MB A. opacus l,eConte BC AB colum,tiensis Wickham A. pardaiis LeConte BC A. pectoral:s LeConte BC AB A. peculios rs Schmidt _MB awemeanus W.J. Brown longitursis Fall tenuihr,rsis Reitter A. phaeopterus LeConte BC AB canadensis Garnett (?) rus;elli Hatch A. piceatus Robinson AK BC AB rainer', Hatch A. pinguell rs W.J. Brown BC AB SK MB A. pinguis1:Ialdeman NT ? AB SK MB oN pe NF ualidus Hotn A. prodromus (Brahm)t _BC SK MB oN PQ NB NS A. richards'tnae Hatch _BC A. rogersi Ilatch A. rubeolus (Palisot de Beauvois) _:r_ ON c op r on.y tn u s Mel sheimer A. rubripennis Horn oN PQ NB NS A. rufrpes (.Linn6)t ON PQ A. ruricola Melsheimer AB -MB oN PQ _NS NF aureliunus Harold curtus Haldeman A. scabriceps LeConte -AB SK A. scrofa (Fabricius)t -PQ NB NF A. sigmoideus Van Dyke BC A. socialis \V.J. Brown _AB _MB A. stercoros,us Melsheimer oN pe A. subaeneits LeConte BC? gentne.i }{atch perpoli,tus Gordon A. subterraneus (Linn6)t ON PQ NB NS A. talpoides,i W.J. Brown -ms A. tenellus Say AK I'K NT BC AB _MB ON PQ A. thomomJrsi W.J. Brown _MB A. vittatus Say _BCAB SK MB oN PQ NB NS _LB A. walshii }Iorn AB SK MB Genus OXYOMUS Stephens O. silvestris (Scopoli)'l BC PQ opacifrons }Iorn porcatus (Fabricius) I48 Scarabaeidae Tlibe Eupariini Genus ATAENIUS Harold Key to North American species: Cartwright (1974) A. abditus {Haldeman.t ON PQ NB attenuator Harold A. gracilis ( Melsheimer t ON PQ A. imbricatus (Melsheimer) ON sordidus Harold A. spretulus (Haldemant ON cozsors Fall falliHinton A. strigatus (Sayt oN PQ Genus DIAJ-YTES Harold D. criddlei W.J. Brown MB D. striatulus tSayt MB ON PQ NB NS cribrosus (LeConte) D. truncatus (Melsheimer) ON PQ coruinus (Haldeman) D. ulkei Horn ON PQ Genus DIAI-YTELLUS W.J. Brown Key to species: Brown (1929) D. dialytoides tFall r -oNPQ D. humeralis (Leconte) -oN Tribe Psammodiini Genus PSAMMODIUS Fallen Key to New World species: Cartwright (1955) P. caelatus (LeContet BC P. laevipennis Costa+ -oN aegialioides Haldeman P. oregonensis Cartwright BC P. sulcicollis (Illigertt * Dr'lrq Genus RHYSSEMUS Mulsant Key to New World species: Gordon and Cartwright (1980) R. californicus Horn AB R. germanus tLinnet* -oN pa:) puncticollis W.J. Brown R. neglectus W.J. Brown AB R. sonatus LeConte AB SK MB Genus PLEUROPHORUS Mulsant Key to New World species: Cartwright (1948) P. atlanticus Cartwright ON P. caesus (Creutzer)t - BC ON PQ Subfamily OCHODAEINAE Genus OCHODAXUS Audinet-Serville Cadocera Eschscholtz Psephus I\ttby Stornphax Fischer von Waldheim O. luscinus Howden _BCAB O. musculus (Say) -oN americanus Westwood O. opacus LeConte MB O. simplex LeOonte AB MBSK t49 Scarabaeidae Subfamily GEOTRUPINAE Key to North American species: Howden (1955) Tribe Bolboceratini Genus BOL,IOCEROSOMA Schaeffer B. bruneri f.tawson & McColloch MB Genus EUC.ANTHUS Westwood Key to Ncrth American species: Howden (1964) E. greeni Rcbinson AB - MB E. lazarus iltabricius; - ON melibot us (Fabricius) Genus BOLTIOCERAS Kirby Odontaeus Fischer von Waldheim Key to Ncrth American species: Wallis ( 1928) B. falli (wallis) - SK MB ON PQ B. filicornis Sayt - SK MB B. liebecki l ilallisr - ON PQ - NS B. obesus (L.econte) - BC AB Tlibe Geotrupini Genus GEOIRUPES Latreille (Subgenus (IEOTRUPES s. str.) G. stercorarius (Linn6)t PQ NB - PE - NF (Subgenus INOPLOTRUPES Jekel) (M ela notr u p e s Blanchard) G. ball Jek:l MB ON PQ NB NS similis Jekel G. hornii Bl rnchard - ON PQ NF (Subgenus ()NYCHOTRUPES Jekel) G. semiopacus Jekel MB ON PQ b la chb ur nii Melsheimer melsheimeri Jekel oualipennis Jekel G. splendidus miarophagus Say - ' G. splendidrrs splendidus - ON PQ - NS (Fab:ricius) gilnichi Jekel mixtus Ilorn starki .tekel Subfamily GLAPFIYRINAE Genus LICI INANTHE Burmeister Dasyde'ra LeConte Key to sp':cies: Carlson (1980) L. rathvoni LeConte) - BC canina Ilorn edwara'siHorn Subfamily TROGINAE Genus GLARESIS Erichson Key to Nt,rth American species: Gordon (1970c) G. canadens,is W.J. Brown AB - MB G. medialis Gordon - BC Genus TRO.K Fabricius Omorgus Erichson Pseudotrox Robinson Key to North American species: Vaurie (1955) T. aequalis Say MB ON PQ NB NS T. atrox LeConte - BC AB SK - ON 150 Scarabaeidae T. capillaris Say ON T. fascifer LeConte _BC T. foveicollis Harold ON aciculeatus Robinson T. hamatus Robinson ON T. robinsoni Vaurie AB SK MB _MBON T. scaber (Linn6) -ec AB PQ NB NS- T. scabrosus Palisot de Beauvois ON T. sonorae LeConte NT AB SK alternans LeConte lecontei Harold T. sordidus LeConte ON PQ T. spinulosus spinulosus Robinson ON T. striatus Melsheimer ON T. unistriatus Palisot de Beauvois AB SK MB ON pe NB NS ID NF porcatus Say T. variolatus Melsheimer ON PQ NB NS erinaceus LeOonte monticola Robinson Subfamily MELOLONTHINAE T?ibe Sericini Genus SERICA Macleay Camptorhina Kirby Stilbolemma Harris S. anthracina LeConte BC AB SK crdssata Walker frontalis LeConte robusta LeConte ualidaHarcld. S. atracapilla (Kirby) _BC MB ON PQ NB NS cucullata Dawson S. curvata LeConte NT BC MB S. elusa Dawson T:^ PQ S. georgiana Leng -oN PQ NB NS PE lecontei Dawson S. intermixta Blatchley NT BC AB SK MB ON PQ S. parallela Casey -oN PQ S. sericea (Illiger) NT BC _SK MB ON PQ S. sponsa Dawson -oN S. tristis LeConte MB ON PQ NB NS PE Tribe Melolonthini Key to North American genera: Hardy (1974) Genus DIPLOTAXIS Kirby Alobus LeConte Diazus LeConte Orsonyx LeConte Key to species: Vaurie (1958; 1960) D. atlantis Fall ON nigra Caftwight D. brevicollis LeConte BC AB nigropicea (Walker) D. haydenii LeConte AB SK impressifrons Caziet innoxia LeConte leuicoxa FalI D. obscura LeOonte AK BC AB SK MB cornpacta Fall consequens (Walker) protninens Fall D. sordida (Say) ON PQ carbonaria Burmeister rugosioides Schaeffer D. subangulata LeConte BC californica Schaeffer D. tenebrosa Fall BC 151 Scarabaeid,re D. tristis K rby MB ON pe NB NS pE c or p ul e n t a Burrneis ter Genus PHYLLOPHAGA Harris Ancylonycha Blanchard Endrosa LeConte Gymnis LeConte Lachnr>sterna Hope Triche:;thes Erichson Key to North American species: Luginbilt and Painter (1g53) P. anxia (LeConte) NT BC AB SK MB ON pe NB NS pE - NF alpina (Linell) breuiccllis (Blanchard) cephal,.ca (LeConte) dubia:.Smittr) inspert:ta (Smith) p u nc t i r::ol li s (Blanchard) uninot:tta (Walker) P. balia , Sa/r ON comola (Burmeister) P. crenulatit tFrrilichr ON georgicana (Gyllenhal) P. drakii (Kirby) AB SK MB ON pe NB NS pE consimilis (LeConte) grandi.; (Smith) P. ephilida 'Sayt burmasteri (LeConte) P. fervidatl'abricius) arcuatut (Smith) quercir a (Knoch) P. lraterna Jarris ON pQ cognaflt (Burmeister) P. fusca (Fr tlich) - BC - oN pe - NS NF feruens (Gyllenhal) P. futilis ( Lt.Conte) - ON PQ NB decidutt (LeConte) gibbosc,. (Burmeister) serricornis (LeConte) P. gracilis tllurmeisterr - ON inana (LeConte) uoluulo. (LeConte) P. hirsuta tl(nocht - ON pe P. hirticula Knochr - ON PQ P.ilicistKn,rcht - ON ciliata LeConte) fimb r ia ta (Burmeister) ybnesi [ianderson porcina (Hentz) P. inversa tllorn r ON P. lanceolat.t tSayt MB ON arizoncte von Bloeker cazieri von Bloeker grisian,z von Bloeker P. longispin,r tSmithr - ON PQ P. marginal s (LeConter ON PQ P. nitida t Lt Conte) AB - MB ON PQ innominata (Smith) limula lHorn) P. rugosa tl\lelsheimert AB - MB ON pQ P. tristis tFa,,bricius) - BC - ON pilosicctllis (Knoch) P. vilifrons tl.eConter ON pe hirticep,s (LeConte) Genus POL'IPHYLLA Harris MacrorLoria CroLch Key to Ncrth American species: Young (1988) P. crinita Le Conte - BC incolum.is Casey mystica Casey L52 Scarabaeidae ona von Bloeker pacifica Casey peruersa Casey santarosae von Bloeker P. decemlineata (Say) BC SKAB castanea Casey com stockiana von Bloeker laticauda Casey matrona Casey oregona Casey parilis Casey peruersa Casey potsiana Casey reducta Casey ruficollis Casey squamotecta Casey P. hammondi LeConte AB SK MB acomana Casey bisinuata Casey diffusa Casey impigra Casey latifrons Casey molesta Casey oblita Casey oklahomensis }].atc}r pimalis Casey proba Casey rufescenta Tanner sejuncta Casey speciosa Casey squatnicauda Casey subuittata LeConte uerecunda Casey P. variolosa (Hentz) ON PQ Genus AMPHIMAILON Berthold A. majalis (Razoumowsky)f ON PQ Tlibe Pachydeniini Genus PHOBETUS LeConte Key to species: Hardy (1978) P. comatus LeConte BC leachi Banett Tribe Macrodactylini Genus DICHELONYX Harris Dichelonycha lltrby Key to North American species: Saylor (1945) Key to eastern North American species: Brown (1946) D. albicollis (Burmeister) ON PQ NB NS D. backii (Kirby) _NTBCABSKMB oN PQ arizonica Barrett D. canadensis (Horn) oN PQ lrD D. diluta (FalI) ON PQ NB NS- D. elongatula (Schtinherr) _PE D. fulgida (LeConte) _BCAB _:a)u crotchi (Horn) mormona (Fall) oregona (Van Dyke) D. kirbyi W.J. Brown AB SK MB D. linearis (Gyllenhal) MB ON PQ hexagona (Germar) testacea (Kirby) D. subvittata (LeConte) MB ON PQ NB NS D. truncata (LeConte) sc AB SK 153 Scarabaeida': D. vicina (Fall) - BC columbinna Hopping deserta Hopping Genus MAC I,ODACTYLUS Latreille Stenothyax Haris M. subspinor,us (Fabricius) oN PQ barbatu:,s Fitct' elongatus (Herbst) poly p ha. g us (Burmeister) Tribe Hopliini Genus HOPLIA Illiger Echyra Erichson Key to No:th American species: Hardy (1977) H. callipyge LeConte BC cazieri \loyer conuexul,a LeConte desertic'tla Boye'r humbol,.ltensis Boyet irrorata LeConte lecontei Dalla Torre mutata Gemminger & Harold oregona LeConte pubicollis LeConte utahensis Boyer H. Iaticollis LeConte _MB?_ H. modesta llaldeman ON barbata Blatchley heluola Melsheimer monticota Melsheimer singularis Burmeister H. sackenii LeConte BC AB H. trifasciate. Say MB ON PQ NB NS ferrisi E:,oyet primarit Burmeister lrlsfis N.,elsheimer Subfamily RUTELINAE Tribe Anomalini Genus ANOUIALA Samouelle Anomalepta Casey Anomalopus Casey Hemispilota Casey Oliganomala Casey Paranornala Casey Rhombanalia Casey Spilota Burmeister Key to North American species: Potts (1977) A. binotata {'}yllenhal t ON compacia Casey maculata (Castelnau) marginella LeConte unifascifia Say A. lucicola iF abricius) ON olrolo (llabricius) cinctello Burmeister moerens (Fabricius) nigritula (Castelnau) p hy I lope' r thoides Nonfried pinicola Melsheimer q uadr im.ac ulat a Dr apiez A. undulata .Vlelsheimer ON canadensis Casey dubia BlatchTey r54 Scarabaeidae floridana Casey nigropicta Casey saginatula Casey uarians Burmeister Genus STRIGODERMA Burmeister Alamona Casey Strigodermella Casey S. arboricola (Fabricius) ON Genus POPILLIA Audinet-Serville P. japonica Newman-i ON PQ NS Tribe Rutelini Genus PELIDNOTA Macleay Key to North American species: Hardy (1975) (Subgenus PELIDNOTA s. str.) (Pelidnotidia Casey) P. punctata (Linn6) MB ON PQ breuicollis Casey breuis Casey debiliceps Casey hudsonica Casey lutea (Olivier) oblonga Casey pallidipes Casey ponderella Casey strenua Casey tarsalis Casey texensis Casey Genus COTAI-PA Burmeister Ciocotalpa Saylot C. lanigera (Linn6) SK MB ON molaris Casey fou Wickham uernicata Casey Subfamily DYNASTINAE Key to species: Endrodi (1985) Tribe Cyclocephalini Genus CYCLOCEPHALA Latreille Dichromina Casey Ochrosidia Casey Spilosota Casey C. borealis Ar-rolv NS uillosa Burmeister C. longula LeConte BC abrupta (Casey) ambiens (Casey) californica Atrow marcida (Casey) modulata (Casey) obesula (Casey) oblongula (Casey) phasma (Casey) prona (Casey) reflexa (Casey) regulifrons (Casey) rustica (Casey) 155 Scarabaeidae Tribe Pentodontinr Genus LIGf RUS Burrneister (Subgenus L,IGYRODES Casey) L. relictus (Say) MB ON PQ NB clypealis (Casey) dawsoni (Casey) paruice,:s (Casey) quadriT ennis (Casey) uernicicollis (Casey) (Subgenus L,IGYRUS s. str.) (Anagrylius Casey) (.Ligyretilus Casey) L. gibbosus 1,;ibbosus (DeGeer) AB ON bicorniculatus Casey breuiusculus Casey curtiper'.nis Casey farctus Jasey juuencus (Fabricius) lacustri; Casey laetulu:, Casey Iaticaut:ta Casey Iongulu; Casey lucubla tdus Casey morio (LeConte') parallelus Casey punctiutuda Casey remotut: Casey rubidus Casey texanus Casey uariolos us (Burmeister) uirginic'"ts Casey Genus APHONUS LeConte Key to No:th American species: Gill and Howden (1985) A. castaneus (Melsheimer) PQ cubiforr zis Casey obesus (Burmeister) saginat,"rc Casey trapezictllis Casey A. tridentatts (Say) ON PQ aterrim;ts Casey congestt,ts Casey elongatus Casey frater L.:Conte modula,lus Casey politus l)asey scutella:is Casey Tribe Oryctini Genus XYLC.,RYCTES Hope X. jamaicensis (Drury) ON PQ americanus (Palisot de Beauvois) faunus \)asey hebes Casey lacustri:; Casey obsolescens Casey sotyrus (Fabricius) tenuicornutus Casey 156 Scarabaeidae Subfamily CETONIINAE Tlibe Cetoniini Key to New World genera: Hardy (1988) Genus EUPHORIA Burrneister Erirhipidia Casey Erirhipis Burmeister Euphorhipis Casey Euphoriaspis Casey Haplophoria Casey Isorhipina Casey Rhipiphorina Casey E. fulgida (Fabricius) _ONPQ E. inda (Linn6) AB SK MB ON PQ barbata (Say) brunnea (Gory & Percheron) marilandica (Frdlich) nigripennis l{lages E. rufobrunnea Casey BC Tribe Cremastocheilini Genus CREMASTOCHEILUS Knoch Key to North American species: Potts (1945) (Subgenus CREMASTOCHEILUS s. str.) C. armatus Walker BC cribripennis Casey densicollis Casey maritimus Casey montanus Casey obliquus Casey pilosicollis Horn C. canaliculatus Kirby -oNPQ hentzi Haris C. castaneus Knoch AB SK MB NS breuisetosus Casey lecontei Westwood pocularis Casey C. crinitus LeConte BC AB bifoueatus Van Dyke pugetanus Casey C. knockii LeConte SK MB areolatus Casey crenicollis Westwood gracilipes Casey (Subgenus ANATRINODIA Casey) C. wheeleri LeConte SK MB Subfamily TRICHIINAE Key to North American species: Howden (1968) Tlibe Osmodermini Genus OSMODERMA LePeletier & Audinet-Serville Gymnodus Kirby O. eremicola (Knoch) -oNPQ NS O. scabra (Palisot de Beauvois) _ONPQ NB NS beauuoisii Bainbridge coseyi Schenkling cauiceps Casey delicatula Casey foueatus (Kirby) gracilipes Casey lacustrina Casey rugosa (Kirby) O. subplanata Casey AB MB ON montana Wickham t57 Byrrhidae Tribe Trichiini Genus TRITIHIOTINUS Casey T. affinis (C'ory & Percheron) - ON PQ paruuius Casey uentricosus Casey T. assimilis, (Kirby) NT BC AB SK MB ON PQ NB NS bistriga (Newman) uariabilis (Burmeister & Schaum) T. piger (Fabriciust ON drumrzondi (Gory & Percheron) reductus Casey r ot unc:,'.icol I is (Kirby) T. viridans tKirby) - ON Genus GNORIMELLA Casey G. maculosr tKnocht - ON PQ bigs6i, (Kirby) dissiniilis (Gory & Percheron) Superfamily Byrrhoidea Family BYRRHIDAE pill beetles P.J. Johnson The lamily Byrrhidae is represented in Canada Taxonomically, the family is reasonably well and Alaska by 26 of the 34 North American species, known in North America. Johnson (-t987) has compare(l with about 280 species around the world. treated the bionomics, larval taxonomy, and bio- The greatest diversity of Canadian byrrhids is geography of the North American generur, and has found in the montane provinces. All species in provided a key to the North American taxa (1991b). Canada irre bryophagous and express microhabitat specifrcity. Many species are associated with the moss Cerotodon purpureus on disturbed sandy- gravelly soils. Species of the coastal forests of British C.iolumbia and Alaska are closely associated aK (14); yK (9); NT (13); BC (15); AB (1:2); SK (5); with certain forest communities and are more MB (7); ON (10); PQ (12);NB (2);NS (a); l,B (5);NF polyphagous in their host mosses. (4) . H (3); I (2) Subfamilv SYNCALYPTINAE Genus CHIiETOPHORA Stephens C. spinosa Rossi t+ - BC - ON PQ Genus CURIMOPSIS Ganglbauer Key to North American species: Johnson (1986) C. albonotata (LeConter - YK AB grisea LeConte C. echinata (LeOonte) AK - NT BC AB SK PQ breuicr'tllis Casey C. moosilar.ike Johnson AK - NT - AB - PQ - LB C. seLulosa (Mannerheim) AK - NT Subfamilv BYRRHINAE Tribl Byrrhini Genus BYRRHUS Linn6 Key to North American species: El-Moursy (1970) B. americanus LeConte MB ON PQ - NS - LB angustulus Casey brunnescens Casey 158 B1a-rhidae criddlei Casey imperitus Casey manitobae casey neglectus Casey B. concolor Kirby AK YK NT SK laramiensis Casey rigens Casey B. cyclophorus Kirby YK NT AB SK MB ON PQ LB canterius Casey dfficilis Casey fasciatus atct. fuluouestitus Casey stolidus Casey torpidus Casey B. eximius LeConte AK YK NT BC callidus Casey centralis Casey explicatus Casey B. geminatus LeContex I'K PQ pacificus Casey pettiti }{otn B. kirbyi LeConte AK NT BC AB MB ON PQ NB NS LB NF consuetus Casey egenus Casey perditus Casey picipes Casey uafer Casey wickhami Casey B. pilula (Linn6)x NT Genus PORCINOLUS Mulsant & Rey P. undatus (Melsheimer) AB SK MB ON PQ crescentifer Casey glabellus Melsheimer hystris Casey murinus Fabricius Tlibe Simplocariini Genus SIMPLOCARIA Stephens S. elongata J.Sahlberg* - YK NT MB ON remotaW.J.Btown S. metallica (Sturm) NT BC AB PQ NF colurnbica Casey subnuda Casey tesselata LeConte S. semistriata ( Fabricius)t BC ON PQ NS NF Genus LIOLIGUS Casey L. nitidus (Motschulsky) AK BC aequabilis Casey heeni Casey striolatus Casey Genus LIOON Casey L. simplicipes (Mannerheim) AK BC puncticeps Casey inflatum LeConte speculare Casey Genus EXOMELLA Casey Key to North American species: Johnson (1985) E. pleuralis (Casey) AK BC 159 Buprestida: Tlibe Pedilophorini Genus EUSOMALIA Casey E. lecontei tWickham) BC Genus LISIIEMUS Casey L. acuminabus (Mannerheim) AK BC formosus Casey satelles Casey Genus MOIIYCHUS Erichson M. aeneolur; (LeConte) AK NT BC AB MB ON lateralis (.Fall1 albertt:. nus (W.J. Brown) szbseloszs (Fall) hesper rc Wickham M. oblongur; (LeConte) AK BC AB subcup,reus (Fall) insulsus Casey Tribe Tylicini Genus TYLICUS Casey T. subcanur, (LeConte) \T( NT BC AB SK ON PQ Genus CYT I LUS Erichsor C. alternatrLs (Say) MB ON PQ NB NS LB NF nigrans Casey C. mimicus Casey NT BC AB longultLs Casey Superfamily Buprestoidea Family BUPRESTIDAE metallic wood-boringbeetles A. Dauies One hundred and fifty-two species of his handbook has been used as the basis for this Buprestidae have been recorded in Canada and checklist, with updated distributiorLal data. Alaska. The adults are usually found resting or Further revision of the species in such groups as ovipositi eg on the bark and foliage of trees and Anthaxiinae is required before a proper subgeneric shrubs, and feed chiefly on pollen-rich flowers; they arrangement can be established. It should be noted are also ;active fliers. The larvae develop mainly that the number of species recorded in the northern under ther bark of the host plants, which are usually regions does not reflect the number of species pre- fairly specific, although a few are leafminers, gall- sent but rather only the lack of collecting in those makers, and coniferous cone borers. regions. Major taxonomic treatments of the North America.n buprestids include those of LeConte AK (11);YK (13); NT (18); BC (75);AB (57); SK (a1); (1860) and Chamberlin (1926). Bright (1987) pro- MB (63); ON (99);PQ (80);NB (36);NS (22);pE (7); vides keys and descriptions ofthe Canadian species; LB (4);NF (5) . H (1); I (2) 160 Buprestidae Subfamily POLYCESTINAE Tribe Polycestini Genus CHRYSOPHANA LeConte C. placida (LeConte) BC AB cupreola Van Dyke strandi Obenberger Subfamily ACMAEODERINAE Tlibe Acmaeoderini Genus ACMAEODERA Eschscholtz A. idahoensis Barr BC A. pulchella (Herbst) AB -llsoN- arizonaeHotn flauosignata Gory imtnaculata Horn mixtaLeConte ornata Olivier uoluulus Castelnau A. tubulus (Fabricius) ON culta (Weber) geranii (Harris) q uattuor de cimg uttata (F abicirs) A. vandykei Fall A. variegata LeConte distincta Kerremans Iucia FalI Genus ANAMBODERA Barr A. gemina (Horn) BC nebulosa (}lorn) Subfamily CHALCOPHORINAE T?ibe Chalcophorini Genus CHALCOPHORA Solier C. flortis LeConte MB ON PQ NB cupreola Casey Iaurentica Casey C. liberta (Germar) MB ON PQ borealis (Gory & Castelnau) paruiceps Casey C. virginiensis (Drury) - BC AB SK MB ON PQ NB NS ang ulicollis (LeConte) antennalis Casey breuicollis Casey c upr eomac ulata (Goeze) ingens Casey lacustris LeConte melanotum Muttkowski montana Casey nouaebo racensis (Fitch) obliterata Casey obscurata (FiLch) oregonensis (Filc}r) pallida Kerremans prominens Casey Genus TEXANIA Casey T. campestris (Sayt ON substrigosa (Gory & Castelnau) 161 Buprestidae Subfamily BUPRESTINAE Tlibe Dicercini Genus SPE CTRAJ-IA Casey S. gracilipes t Melsheimer) ON Genus DICERCA Eschscholtz Argante Gistel Stenuris I{ttby Key to North American species: Nelson (1975) D. callosa callosa Casey AX YK NT BC AB SK MB ON PQ pertirutx Casey rigida Casey tetrica Casey D. callosa f rosti Nelson _BCAB D. caudata LeOonte NT_ABSKMBONPQ NB NS abrup;a Casey biangulata Casey cupreola Casey filiola Casey inflatula Casey longicu.ud.a Casey pisciformis Casey D. crassico.llis LeConte BC AB californica Crotch hesperica Casey D. divarica;a (Say) MB ON PQ NB NS aestiuo Casey angus icauda Casey aurich alcea Melsheimer dabio Melsheimer incisa Casey lirnulo Casey nigra l)asey parumpunctata Melshermer rustict:,, Casey subaet,ualis Casey uancotlueri Casey D. dumolini(Castelnau & Gory) SK MB ON PQ consobrina Melsheimer D. hesperoboreaiis Hatch & Beer _BCAB D. horni Crotch _BC ampliuta Casey D. lugubris LeConte NT SK MB ON PQ NB auster't Casey lacustt'is LeConte morio Casey D. lurida (tr'abricius) oN PQ erosa Casey floridae Casey gaudens Casey gracilis Casey indis tinc ta Mel sheimer indura,ta Casey innocua Casey leuette,;. Casey porcat,'tla Casey pruinasa Gory regula,is Casey sagor lasey seriato. Casey truncata Casey D. pectorosa LeConte _BC_SK D. pugionata (Germar) ON D. punctulata (Schtinherr) oN PQ pinorum Casey transuersa Say D. sexualis Crotch _BC D. tenebrica (Kirby) AK YK NT BC AB SK MB ON PQ NB NS longipennis Casey prolongata LeConte 162 Buprestidae seuera casey subargentea Casey subcuprea Casey sulcatula Casey D. tenebrosa (Kirby) AK YK NT BC AB SK MB ON PQ NB NS PE LB acerba Casey bifoueata LeConte chrysea Melsheimer distinguenda (Castelnau & Go.y) montana Casey mystica Casey stolida Casey wickhami Casey D. tuberculata (Castelnau & Go.y) oN PQ NB NS asperata Castelnau hilaris LeConte mancaLeConte molitor Melsheimer scobina Chewolat Genus DESCARPENTRIESINA Leraut Key to North American species: Evans (1957) D. cyanipes (Say) NT BC AB SK MB ON PQ NB apicilla (Obenberger) collaris (Obenberger) cupripes (Casey) debilis (LeConte) erecta (Gory) paruiceps (Casey) D. ferrea (Melsheimer) _BCABSK MB ON PQ D. fraseri (Chamberlin) -BC D. montana (Chamberlin) _BC MB D. thureura (Say) MB costicollis (Gory) Tribe Buprestini Genus TRACHYKELE Marseul T. blondeli Marseul _BC T. nimbosa Fall -BC Genus BUPRESTIS Linn6 Ancy lo c h ir a Eschscholtz Anoplis l{trby Gymnota Gistel Key to North American species: Helfer (1941) B. adjecta (LeConte) BC AB breuis Casey B. aurulenla Linn6 BC AB MB ON adulans Casey aemula Casey affinis Casey bicostata Dejean chry sochlora (Philippi) fabulosa Casey lauta (.LeConLe) nupta Casey prospera Casey radians (LeConte) tacornae Casey uenusta Casey uillosa (LeConte) B. confluenta Say YK_BC AB SK MB ON PQ tesselata Casey B. fasciata Fabricius SK MB ON PQ NB NS fortunata Casey fulgens Casey lherminieri Chevrolat saturata Casey 163 Buprestidae sexmaculata Hausmann sexplagiata (LeConte) B. intricateL Casey BC AB con,tortae Hopping murraydnae Hopping B. laeviventris (LeConte) BC pugeta,na Casey B. langi Mannerheim AK BC AB SK MB angusta Casey bistrinotata Casey callida Casey crenata Casey d.epressa Casey fastidt.osa Casey gramitea Casey incolumis Casey Iaeuicirps Casey medio:ris Casey obliqu.a Casey oregon.a Casey ornata (Walker) patruelis Casey sediticsa Casey uiridirnicans Casey B. lineata ltabricius BC MB ON PQ NS douisi Nicolay & Weiss B. lyrata Casey BC AB SK MB ON NS acomana Casey adducta Casey caligitnsa Casey fuscat)asey lecontt::i Saunders morosd Casey nigricuns Casey B. maculat.ventris Say NT SKAB MB ON PQ NB NS PE pagancrum l{trby rusticorum Kirby sexnotzta Gory & Castelnau B. maculipennis Gory SK MB PQ deficiens Casey fusiformis Casey inconstans Melsheimer leporirua Casey reductt Casey scriptt:. Casey B. nuttalli Kirby AK YK NT BC AB SK MB ON PQ altern':tns (LeConte) boulderensis Casey conici<,auda Casey consulcris Goty contor;la Casey diruptzns Casey flauopicta Casey grauidula Casey torua Oasey B. salisbu4rensis Herbst ON cazieri Helfer ultran,rarina Say B. striata I abricius oN PQ NB canad,ensis Casey imped.it.ta Say obscura Casey B. subornal;a (LeConte) BC adoneu Casey histrio Casey punctiuentris Casey rubrorrctans Casey uiolescens Casey B. sulcicollis (LeConte) MB ON PQ NS NF lateralis Casey t64 Buprestidae Tribe Melanophilini Genus MELANOPHILA Eschscholtz Phaenops Dejean Key to North American species: Sloop (1937) M. abies Champlain & Knull ON PQ NB o6iells Obenberger M. aeneola Melsheimer ON PQ metallica Melsheimer M. californica Van Dyke BC M. drummondi drummondi (Kirby) AK YK NT BC AB SK MB ON PQ NB LB guttulata Mannerheim umbellatarum (Kirby) M. fulvoguttata (Harris) BC SK MB ON PQ NB NS PE NF croceosignata (Castelnau & Gory) decolorata (Castelnau & Gory) ftarrisi Obenberger octospilota (Castelnau & Gory) pacifica Obenberger M. gentilis LeConle BC prasina LeConte M. intrusa Horn BC Genus OXYPTERIS Kirby Apatura Castelnau & Gory O. acuminata (DeGeer)* AK YK NT BC AB SK MB ON PQ NB NS PE NF ap pendiculata (Fabricius) i mma c u I ata ( Mannerheim) longipes (Say) morio (Fabricius) O. occidentalis Obenberger BC Tribe Anthaxiini Genus ANTHAXIA Eschschoitz (Subgenus MELANTHAXIA Richter) A. aeneogaster Castelnau & Gory BC aerica Ctotch grosso Obenberger strigata LeConte A. aenescens Casey _BC A. inornata (Randall) _YKNTBCABSKMBONPQNBLB expdnsa LeConte foueicollis LeConte imperfecta LeConte A. porella Barr BC A. prasina Horn BC falsula Obenberger A. retifera LeConte AK YK BC MB ( Subgenus IIAPLANTHAXIA Reitter) A. deleta (LeConte) _BC A. pseudotsugae (Chamberlin) _BC A. quercata (Fabricius) ON PQ N" cuneiformis (Gory) A. viridicornis (Say) SK MB ON PQ subaenea (LeConte) uiridifrons (Gory) Genus AGRILAXIA Kerremans A. flavimana (Gory) ON gracilis (Melsheimer) 165 Buprestidae Subfamily CHRYSOBOTHRINAE Tribe Chrysobothrini Key to North American species: Fisher ( 1942) Genus CHRYSOBOTHRIS Eschscholtz Knowltonia Fisher Odontomus l\ttby C. adelpha Gemminger & Harold ON PQ soror LeConte C. aeneola l,eConte AB SK (?) purpurata Bland C. azttea LeConte BC AB SK ON PQ lecontet Leng C. beeri Barr _BC C. blancharrli Horn AB SK ms oll PQ NB C. breviloba Fall _BCAB C. carinipernis LeConte _BCAB C. caurina IIorn _BC C. columbiana Barr _BC C. cribraria Mannerheim NT_ABSK lln Of,l pe calcarota Melsheimer f'Loricola Gory lalo Kerremans C. dentipes iGermar) BC SK MB ON PQ NB c har acr e r is tica ( Harris) planattt Gory & Castelnau posticatis Gory & Castelnau rotund,collis Gory & Castelnau C. femorata (Olivier) BC AB SK MB ON PQ NB NS alaban,ae Gory dissim,,.Iis Dejean fastidit,sa Gory insculF'ta (Herbst) lesueuri Gory & Castelnau misello LeConte nigrituta Gory & Castelnau obscurr.t LeConte quadri;',mpresso Gory & Castelnau C. monticoler Fall BC AB grandis Chamberlin C. harrisi (Flentz) BC AB-MBON PQ NB NS_ C. laricis Van Dyke BC AB C. leechi Barr BC AB C. mali Horn BC AB SK MB C. neopusilla Fisher PQ NS_ C. nixa Horr BC fu C. oregona Ohamberlin BC _ _ T:'. planon .arginafo Chamberlin C. orono Frost MB ON C. pseudots rgae Van Dyke sc C. pusilla Gory & Castelnau MB ON PQ NB bigutta,ta Gory & Castelnau str ang ulata Melsheimer C. rugosicelrs Melsheimer C. scabripennis Gory & Castelnau xr nc AB ix MB 3N i3 NB PE proxima (Kirby) scabra Goty C. sexsignata (Say) oN PQ NB PE germari Gory & Casteinau ignipes Gory & Castelnau serguttata (Say) C. sylvania Fall _BC C. trinervia (Kirby) AK YK NT BC AB SX MB ON PQ NB NS LB NE cicat r i cos a Motschulsky C. verdigripennis Frost AB SK MB ON PQ NB NS C. viridiceps Melsheimer MB ON C. vulcanica LeConte nc AB canadensis Chamberlin 166 Buprestidae Subfamily AGRILINAE Tribe Coraebini Genus EUPRISTOCERUS Dey'r'olle (? ) R haeb oscelis LeConte E. cogitans (Weber) ON PQ NB ignarus (Fabricius) ruficollis (Herbst) Tlibe furilini Genus AGRILUS Curtis A. acutipennis Mannerheim _MBONPQ latebrus Castelnau & Gorv A. anxius Gory AK BC AB SK MB ON PQ NB NS NF grauis LeConte torpidus LeConte A. arbuti Fisher _BC manzanitae Chamberlin A. arcuatus (Say) MB ON PQ NB obliquus LeConte A. atricornis Fisher -ONPQ A. aurichalceus Redtenbacherf _ONPQ A. bilineatus bilineatus (Weber) MB ON PQ aurolineatus Gory biuittatus l{trby flau olineatus Mannerheim triuittatus Horn A. bilineatus carpini Knull A. burkei Fisher -ncAB--9*PQ----- A. celti Knull -oN A. cephalicus LeConte -BCAB-MBONPQ A. crataegi Frost AB ON PQ A. criddlei Frost A. crinicornis Horn _ _ _:"yu3Ni3Iu_ _ _ _ A. cyanescens (Ratzeburg)t -ONPQ A. defectus LeConte -ONPQ A. egenus Gory -oN A. fallax Say -oNPQ impressipenni.s Uhler zemes Gory A. frosti Knull _MBONPQ A. geminatus (Say) oN PQ A. granulatus populi Fisher BC AB tr ic hocar p ae Chamberlin A. horni Kerremans A. imbellis Crotch A. juglandis Knull A. lacustris LeConte - - -"gN;:- cuneus LeConte - pubiuentris Ctotch A. lecontei Saunders -oN subfasciatus LeOonte A. liragus Barter & W.J. Brown A. malvastri Fisher ---:"ffisKYuoNPQNB--- A. masculinus Horn -SKMBONPQ A. nigricans Gory -oN auricomus Frost A. obsoletoguttatus Gory _ONPQ interruptus LeConte subaeratus Hatris A. otiosus Say MB ON PQ uirens Goty A. parvus Saunders MB pusillus Say A. pensus Horn AB SK MB ON PQ NB NS PE betulae Fisher A. politus (Say) AK YK NT BC AB SK MB ON PQ NB NS canadensis Obenberger cupreolus LeConte 167 Buprestidar: desertus LeConte plumbcus LeConte solitar,t,us Harold A. pseudoccryli Fisher -MBONPQ A. putilius l5ay oN PQ A. rufrcollis (Fabricius) -MBONPQ cupricollis Goty A. sayi Saurders _PQ NB NS browni Catlson & Knight lateralis (Say) A. subcinctus Gory ON A. vittaticollis (Randall) BC oN PQ frenatus Gory A. walsinghami Crotch BC Subfamily TRACFTYINAE Tribe Pachyschelini Genus PACHYSCHELUS Solier P. confusus Wellso & Manley -oN P. laevigatts (Say) _ONPQ oblong zs (Motschulsky) ouatus (Say) politus Kertemans punctatus (Goty) P. purpurer.,s (Say) AB ON biimpr essus Motschulsky Tribe Brachyini Genus BRACHYS Solier B. aerosus l1{elsheimer AB SK MB ON PQ rufescens Nicolay & Weiss B. aeruginosus Gory -oNPQ B. ovatus (\Veber) _ONPQ auruleztus (Kirby) bellporli Nicolay & Weiss horni }(erremans laeuicauda LeConte molestus Gory termin,zns Castelnau & Gory Genus TAPFIROCERUS Solier T. gracilis (iiay) AB SK MB ON PQ albog u tt at us (Mannerheim) grossu:; Obenberger texanus Kerremans T. nicolayi Obenberger T. schaefferi Nicolay & Weiss - - ABsr- 3N------ 168 Heteroceridae Superfamily Dryopoidea Family PTILODACTYLIDAE toed-winged beetles L. LeSage The family Ptilodactylidae, which is primarily plant matter; those of Anchytarsus are truly tropical in distribution, comprises about 500 species aquatic, burrowing in submerged decaying wood in around the world, but only three occur in Canada. creeks (LeSage and Harper 1976a). Adults hide in the riparian vegetation or in the leaf Stribling's recent key (1986) is useful for identi- litter during the day; they are easily attracted to fying the genera. light. Larvae of Ptilodactyla prefer moist or wet habitats, such as beaver houses or wet, decaying BC (1);MB (1);oN (2);PQ (3);PE (1) Genus FTILODACTYI,A Illiger Key to North American species: Johnson & Frey'tag (1982) P. serricollis (Say) - BC MB ON PQ PE elate r ina Gu6rin-M6neville /usca Melsheimer P. carinata Johnson & Freytag PQ Genus ANCIIYTARSUS Gu6rin-M6neville Key to North American species: Stribling (1986) A. bicolor (Melsheimer) ON PQ ofer Gu6rin-M6neville debilis Ziegler Family HETEROCERIDAE variegated mud-lovingbeetles L. LeSage Heterocerids are represented in Canada and has been followed for this checklist except for the Alaska by 25 species. The adults are nocturnal and genus Dampfius, which was recently placed in syn- are readily attracted to light. During the day they onymy witlr' Heterocerus by Miller (1988b). The hide and make tunnels in sandy areas near water Canadian fauna is relatively well known, but new courses or standing water, a behavior shared by the species are still being discovered (Miller 1988o, Iarvae. Adults and larvae feed on zooplankton, 1988b). Both the arctic and the maritime fauna diatoms, and algae. have been little surveyed. Pacheco (1978) has Pacheco (1964) proposed a drastic change in the recently cataloged the North American species. traditional treatment of the family when he described 15 new genera and restricted the genus Heterocerus to the OId World. Although several of aKQ); YK (a);NT (5); BC (e);AB (16); SK (14); MB his new genera are questionable, his classifrcation (1e);ON (16);PQ (16);NB (5);PE (1) Tlibe Auryliini Genus EXPLORATOR Pacheco E. canadensis (Fall) AK YK NT AB SK MB Genus CENTURIATUS Pacheco C. auromicans (Kiesenwetter) -_ C. compactus (Fall) _ _ :"ffi:"MBg"i8 _ _ _ _ _ r69 Heteroceridae Genus MICIIOAUGYLES Pacheco M. moleculus (Fall) _ABSKMB_PQ M. mundulus (Fall) -AB Tlibe Heterocerini Genus LAN'IERNARIUS Pacheco L. brunneus (Melsheimer) _NTBCABSKMBONPQPE L. gemmatur tHornt BC AB SK L. mollinus (Kiesenwetter) - _ABSKMBONPQ L. parrotus llacheco _NTBCABSKMBONPQNB L. sinuosus .Pacheco YK NT BC AB SK MB Genus NEO F{ETEROCERUS Pacheco N. fatuus lKiesenwetter ) - MB ON NB N. gnatho {I,econte) YK _BCABSK MB ON pe labratus (LeConte) N. longilobu lus Pacheco PQ N. pallidus tSayt _ABSK MB ON PQ NB uentrahs (Melsheimer) N. sandersoni Pacheco MB Genus HETEROCERUS Fabricius Dampfius Pacheco Key to North American species: Miller (1988b) H. cof laris (liesenwetter) - SK MB ON PQ H. inciertus Pacheco.) _BCAB MB ON PQ H. mexicanus Sharp -oN H. subtilis Vt.V. Miller MB ON PQ H. undatus {Melsheimer; AB SK MB ON PQ NB H. unicus W V. Miller _BCABSK lvlD Genus PEDI TATUS Pacheco P. schwarzi Horn) _ONPQ P. tenuis tW V. Miller) MB ON Genus LAPSUS Pacheco L. tristis (Mannerheim) AK YK NT BC AB SK MB ON PQ NB Tribe Tropicini Genus TROI'ICUS Pacheco T. pusillus {flayt - ON PQ americanus (Sharp) Iimbatus (Kiesenwetter) luteolus (LeConte) 170 Dryopidae FamilyLIMNICHIDAE minute marsh-lovingbeetles L. LeSage Of the three subfamilies and 15 genera of This group of beetles was first considered as a Limnichidae described in the Western Hemisphere, tribe of Byrrhidae, but Hinton (1939o, 1939b) ele- only two genera and four species occur in Canada. vated it to family rank and placed it in the super- Adults of limnichids are usually active during the family Dryopoidea, after a study of the internal day near water in leaf litter; they are readily anatomy. The North American limnichid fauna is attracted to light. The larvae are rarely collected now well known after a series of recent revisions by and are not represented in most collections. Paulus Wooldridge, who also published a catalog of the (1970) frrst described the immature stages of two North American species in 1986. European species: they are terrestrial, Iive in moist soil, and tunnel under layers of mosses and algae, which thev use as food. eC (1);MB (1);oN (4);PQ (2) Subfamily LIMNICHINAE Tribe Limnichini Key to North American genera: Wooldridge (1975) Genus LIMNICHITES Casey Key to North American species: Wooldridge (1977) L. huronicus Casey ON L. olivaceus (LeConte) ____ ON uirginicus Casey L. punctatus (LeConte) BC MB oN PQ austinianus Casey Genus LUTROCHUS Erichson Key to North American species: Brown (1972) L. laticeps Casey ON PQ Family DRYOPIDAE long-toed water beetles L. LeSage Dryopidae includes over 200 species around the The family has not been revised since Brown world, but only six occur in Canada. Adults are (1972) put together all the taxonomic information. essentially phytophagous, feeding on diatoms, The Canadian fauna is reasonably well known for algae, and decaying wood. The truly aquatic adults Quebec and Ontario (LeSage and Harper 1975). of Helichus inhabit riflles of water courses, using a Brown (1983o) recently published a catalog of the hydrofuge plastron for breathing, as do elmids. North American species. Adults of Dryops are typically riparian. They can take flight directly from water surface when dis- turbed. Dryopid larvae are rarely collected; the few specimens collected were found in moist soil on sC (2); AB (1); SK (1); MB (1); ON (2); PQ (5); NB banks of creeks where the adults live. (2);NS (2);PE (1);NF (1) ' I (1) Genus DRYOPS Olivier (Subgmus YRDOPS Steffan) D. viennensis (Heer)t PQ Genus HELICHUS Erichson Key to North American species: Musgrave (1935); Brown (1972) L77 Elmidae H. basalis L,:Conte PQ NB NS H. columbia.rus W.J. Brown BC H. fastigiatts (Say) PQ H. lithophih.Ls (Germar) ON PQ H. striatus LeConte --ecAB SK MB ON PQ NS NS PE_NF foueatus LeConte Familv ELMIDAE riffle beetles L. LeSage Elmidae comprises about 1100 species around The family Elmidae is usually split into two sub- the world: 32 of them occur in Canada. Elmids are families (Brown 1972). The North American fauna is small beetles, usually less than 3 mm long, and most relatively well known as a result of Brown's generic are aquatic in both the adult and the larval stages. treatment (1972), his catalog (1983c), and the recent Adults breathe by means of a hydrofuge ventral plas- revisions of the genera Dubiraphia and Optioseruus. tron, the arvae with retractile cloacal gills. Adults Stenelrnis, the largest genus, is now under revision. and larver e inhabit the substratum of creeks and A systematic survey of the eastern Canadian fauna rivers, feeding on diatoms, encrusted algae detritus, undertaken by LeSage and Harper (1975) focuses on or submer:ged decaying wood (LeSage and Harper Quebec. The fauna of the Prairie Provinces, western t976b); they are good indicators of water quality Canada, and the Arctic is still poorly known. The (Hilsenhoff I977). Pupation occurs near water, present list adds many new provincial records based under ro Subfamily LARINAE Tlibe Larini Key to genera: Brown (1981) Genus LARI. LeConte Key to Noth American species: Brown (1972) L. avara amlrlipennis Darlington BC L. avara avara LeConte BC Subfamily ELMINAE Tribe Ancyronychini Genus ANC-IRONYX Erichson A. variegata (Germar) oN PQ cincta (.Say) parumctculata (Hardy) Tribe Elmini Genus CLEPTELMIS Sanderson C. ornata (Schaeffer) BC Genus DUBIRAPHIA Sanderson Key to No"th American species: Hilsenhoff(1973) D. bivittata tLeConte) MB ON PQ D. minima E.ilsenhoff _ _ T_ MB ON PQ D. quadrinotata (Say) MB ON PQ NB NSpn-- D. vittata (Nl.elsheimer) AB SK MB ON PQ NS 172 Elmidae Genus HETERLIMNIUS Hinton Ceralimnius Hinton Key to North American species: Brown (1972) H. corpulentus tl,eContet - BC AB antennatus (Fall') H. koebelei (Martinj - BC AB Genus MICROCYLLOEPUS Hinton Key to North American species: La Rivers (1949) M. pusillus pusillus (LeConte) - BC - ON PQ NB browni (Hatch) Genus NARPUS Casey N. concolor (LeConter - BC AB solutus (W.J. Brown) Genus OPTIOSERVUS Sanderson Key to North American species: White (1978) O. divergens (LeConter - BC AB SK pecosensis (Fall) O. fastiditus (LeConte) NT BC AB SK MB ON PQ NB NS PE - NF O. quadrimaculatus tHorn) - BC AB O. ovalis tl-eContet - ON PQ NB NS PE LB ampliatus (Fa'r\) O. seriatus (Leconte) - BC AB O. trivittatus (W.J. Brown) - ON PQ NB NS NF Genus OULIMNIUS des Gozis O. latiusculus (LeOonte) - ON PQ NB NS PE - NF Genus ORDOBREVIA Sanderson O. nubifera (Fall) - BC Genus PROMORESIA Sanderson Key to North American species: Brown (1972) P. elegans ll-eContet PQ P. tardella (Fall) PQ NB NS - LB NF subarctica (W.J. Brown) Genus STENELMIS Dufour Key to North American species: Brown (1972) S. bicarinata LeConte - ON PQ - NS S. concinna Sanderson PQ S. crenata (Say) MB ON PQ NB NS PE - NF sordida Motschulsky S. mera Sanderson PQ NB NS S. musgravei Sanderson PQ S. quadrimaculata Horn - ON PQ blatchleyi Musgrave sulcata Blatchley S. sandersoni Musgrave - ON PQ tarsalis Sanderson S. vittipennis Zimmermann - SK MB ON PQ T?ibe Macronychini Genus MACRONYCHUS P.W.J. Miiller Key to North American species: Hinton (1940) M. glabratus Say - ON PQ NB NS lateralis Melsheimer Genus ZAITZE\rIA Champion Z. milleri H.P. Brown - BC Z. parvula (Horn t - \'K - BC AB columbiensis (Angell) 173 Artematopidae Family PSEPHENIDAE water-penny beetles L. LeSage Psephenids form a small family of aquatic bee- The taxonomy of the North American tles; onl.' three species have been recorded so far in psephenids is relatively well known, tha.nks to the Canada. Their vernacular name refers to the works of Brown (1972, 1983b), Brown and Murvosh unique round and flat shape of the body of the lar- (I974), and Brigham (1981). vae, which live in riffles of creeks and rivers. The adults o1' Ectopria spend the daytime in the foliage of marginal vegetation; they are readily attracted to light. Tlrose of Psephenus scurry over exposed rocks in riflles during the day and hide at night. ON (3); PQ (2);NB (1); NS (2) subfamily EUBRIINAE Genus EC,fopRrALeconte Eurea LeConLe Key to Irforth American species: Brigham (1981) E. leechi Brigham - ON E. nervosa tMelsheimerr - ON PQ - NS tarsalis LeConte thorar:'ica (Ziegler) tibiali)s LeConte subfamily PSEPHENTNAE Genus ps[]pHENUS Hardeman Eurytr:.alpus LeConte Fluuit:'ola DeKay Key to liorth American species: Brown & Murvosh (1974) P. herricki lDeKayr - ON PQ NB NS lecontei (LeConte) t re nto nen si s Zimmermann Superfamily Elateroidea Family ARTEMA*TOPIDAE Y. Bousquet Thi-" family includes eight North American one found in North America. Many authors, includ- species, four of which are currently known to occur ing Arnett (1968), have placed the macropogonines in Canarla. Little is known about the bionomics of within Dascillidae. the Near:ctic species. The adults are usually found on foliage; the larvae of some species have been reared or.r mosses (Lawrence 1982q.). Lawrence (1982a) recognized three subfamilies NT (1); BC (3); AB (1); MB (1); ON (2); PQ (2); NB of Artematopidae, Macropogoninae being the only (1) Subfamily MACROPO GONINAE Genus MACROPOC'ON Motschulsky Key to North American species: Hopping (1936) M. piceus l,eConte NT BC AB - MB ON PQ NB dubiusW.J. Btown rufipes l{orn M. testaceipennis Motschulsky - BC 174 Elateridae cribricollis W.J. Brown rubricollis Pic Genus EURYPOGON Motschulsky Dasypogon LeConte E. californicus Horn BC E. niger (Melsheimer) oN pe horrisi (Westwood) Family ELATERIDAE click beetles Y. Bousquet This family is well known for the ability of the The classifrcation of the family followed here is adults to click when lying on their back. It includes Hyslop's (1917), with slight modifrcation. His classifr- 369 species and subspecies in Canada and Alaska. cation is preferred Stibick's (1979), as it is used more The adults occur on foliage and flowers, under bark, widely. There is no recent taxonomic revision of the and in rotting wood; the larvae are found in leaf litter, North American species. Most species that occur in soil, moss, and rotting wood. Larvae of most species the areas covered in this checklist could be identifred living in leaf litter and rotting wood are predaceous; using Dietrich (1945), Brooks (1961), Lane and those occurring in the soil feed on roots, and some are Lancaster (1971), and the taxonomic reYisions listed. very injurious to crops. Adults of some species appar- ently do not feed, whereas those ofother species prob- AK (53); YK (32); NT (49); BC (19a); AB (131); SK ably feed on tender plant tissues, such as leaves and (88);MB (99); ON (175);PQ (165);NB (81); NS (87); flower buds. PE (21);LB (25);NF (46) ' H (6);I (5) Subfamily PYROPHORINAE Tlibe Pyrophorini Genus LACON Castelnau Aulacon Arnett Diphyaulon Arr:.ett Lepidotus Stephens Zalepia Arnett Key to North American species: Arnett (1952) L. auroratus (Say) -_ yu oN PQ NB L. avitus (Say) _ v_ oN PQ L. discoideus (Weber) ON pennatus (Fabricius) L. impressicollis (Say) ON lepturus (Say) senilis (Germar) L. maculatus (LeConte) ON L. marmoratus (Fabricius) oN PQ L. rorulentus (LeConte) -ec-- L. sparsus (Candbze) -BC Genus DANOSOMA Thompson Key to North American species: Arnett (1952) D. brevicorne (LeConte) AK YK NT BC AB SK MB ON PQ NB cauicolle (LeConte) profusutn (Candbze) uetusturn (Walker) D. obtectum (Say) NTYK BC AB SK MB ON PQ NS Genus ALAUS Eschscholtz A. melanops LeConte BC naja Canddze A. myops (Fabriciust oN PQ canadensis Canddze A. oculatus (Linn6) oN PQ luscus (Fabricius) ]-75 Elateridae Genus DR.A.STERIUS Eschscholtz D. debilis L,eConte AK YK BC SKAB MB ON PQ NB NS incongruus Fall Genus AEC'LUS Eschscholtz A. amabilis (LeConte) -oN A. mellillus comis (LeConte) NT BC AB A. mellillus marginicollis (Horn) sx MB A. mellillus mellillus (Say) -oN PQ dorsali.s (Say) Genus COIT|ODERUS Eschscholtz C. auritus (Herbst) AB SK MB ON PQ NS caseyi'.Leng) crassicollis (Melsheimer) dilectus (Say) C. bellus (Say) ON piclzs lSteinheil) uariegutus (Steinheil) Tribe Pityobini Genus PIT)|OBIUS LeConte P. anguinusi LeConte AR SK MB ON PQ NB NS billing.si Bland Tlibe Athoini Genus LIM,INIUS Eschscholtz L. aeger Leilonte BC MB ON PQ NB NS PE LB NF hnulli ?all L. agonus (fiay) ON PQ L. anceps L:Conte SK MB ON PQ NB L. aurifer LeConte _MBONPQ L. auripilis (Say) ON PQ pubico;Iis (LeConte) L. basillaris (Say) ON PQ semiaeneus LeConte L. bicolor Van Dyke _BC L. cal ifornicus t Mannerheimj _BCABSK MB hispidus LeConte occider,.talis Candbze L. canus LeConte BC SKAB discoiaeus LeConte subcosi atus Motschulsky L. confusus LeConte oN PQ NS aenesc(:ns LeConte L. crotchi Horn BC L. ectypus (Say) SK mn- L. griseus (l']alisot de Beauvois) -oN hirticot lis Melsheimer propexits Canddze L. humidus Lane _BC L. infuscatus Motschulsky _BCAB L. lanei Van Dyke _BCABSKMB L. lecontei Lane _BCABSK consimilis Walker nitidicollis LeConte L. maculicollis Motschulsky _BC discicollis Canddze L. nitidulus Horn _BC AB L. pectoralir; LeConte AK YK NT BC MB ON PQNB_ LB NT L. pictus Van Dyke _BC y:" L. pilosulus Canddze -BC pilosus LeConte L. plebejus {Say) oN PQ m e ta I I e scens Melsheimer 176 Elateridae L. quercinus (Sayt L. rudis W.J. Brown L. rufihumeralis Lane ECAB_ L. seminudus Van Dyke BC L. snakensis Lane BC_- L. stigma tHerbst) armus (Say) L. subauratus LeConte BC L. ursinus Van Dyke AB SK MB L. venabelsi Wickham BC Genus ELATHOUS Reitter E. californicus Van Dyke BC E. discalceatus (Say) ON PQ NS ober ndor ffe r i ( Schwarz ) E. nebulosus (Van Dyke) BC Genus ATHOUS Eschscholtz Crepidophorus Mulsant & Guillebean Key to eastern North American species: Becker ( 1974) Key to North American species: Becker (1979) A. acanthus (Say) ON PQ NS carolinus Van Dyke flauangularis Hotrr maculicollis LeConte A. brightwelli {Kirby} ON PQ NB NS arcticollis Melsheimer oblong icoll is Melsheimer A. campyloides Newmant -PQ NB NS PE affinis Coupet A. cucullatus {Say) ON PQ _NS hypoleucus Melsheimer p rocer icoll is Melsheimer stripatus Melsheimer A. lossularis (LeConte) -oNPQ A. imitans Fall BC A. neacanthus Becker PQ A. nigropilis Motschulsky BC AB montanus LeConte nugalis Fall orophilus Harold speculifer FalI A. orvus Becker _ONPQ NB NST_ _ A. posticus (Melsheimert -oNPQ erebusYanDyke A. productus (Randall) SK MB ON PQ NF bipunctatus Provancher A. rufrfrons (Randallt MB ON PQ NB NS NF lengiDtry reflexus LeConte A. rufrventris rectithorax Reitter BC recticollis Fall A. rufrventris rufiventris AK BC AB (Eschscholtz) fer r ug inos us Eschscholtz maritimus Van Dyke A. scapularis (Say) A. scissus LeConte BC ___9*PQ_____ ingens Fall A. sierrae varius Lane AK BC A. vittiger LeConte BC Genus HARMINIUS Fairmaire H. triundulatus (Mannerheim) AK YK BC SK ON PQ Genus MICRATHOUS Lane M. brevis (Van Dyke) BC Genus DENTICOLLIS Piller & Mitterpacher Key to North American species: Becker (1952) 177 Elateridae D. denticor nis { Kirby t NT BC AB SK MB ON PQ NB NS NF flauinrcus (Melsheimer) D. varians (Germar) AK YK NT BC uariabilis (Eschscholtz) Genus BERNINELSONIUS Leseigneur Key to s;cecies: Stibick (1976) B. hyperboreus (Gyllenhal)* AK NT plona,:us (Eschscholtz) Genus LIGMARGUS Stibick Key to N.orth American species: Stibick (1976) L. funebris (Canddze) AK YK NT BC AB L. lecontei (Leng) -oN PQ NS planaius (LeConte) Genus ASCOLIOCERUS M6quignon Scolio :erus Motschulsky Key to North American species: Stibick (1978) A. sanborni(Horn)x AK YK NT AB MB PQ barba:tus (J.R. Sahlberg) Genus MAI,GAIOSTUS Stibick Key to North American species: Stibick (1978) (Subgenus MARGAIOSTUS s. str.) M. glacialirr (Van Dyke) AK BC AB M. grandic'rllis (Lecontel oN PQ Genus HYI'OLITHUS Eschscholtz C ry pto hy pnus Eschscholtz Key to n*orth American species: Stibick (1978) H. littoralir; Eschscholtzx AK Genus HYPNOIDUS Dillwyn Key to North American species: Stibick (1978) H. abbreviatus (Say) AK YK BC AB SK MB ON PQ NB NS PE NF silaceipes (Germar) H. bicolor ( Eschscholtz)* AK YK NT BC AB SK MB ON PQ NB NS NF fallax (Mamer}rein) lacustris (LeConte) limbatus (Mannerheim) picescens (LeConte) pumilits (Dejean) scar ifi c at u s (Mannerheim) uestitus (Mannerheim) H. impress collis (Mannerheim) AK YK NT BC AB MB ON PQ LB NF H. leei Stibick _AB H. nocturnus (Eschscholtz) AK YK_BCAB H. rivularius (Gyllenhal)* AK YKNTBC_SK MB PQ LB NF H. squalidr.s tLeContet BC AB Genus OEDOSTETHUS LeConte O. femoralis LeConte AB SK MB ON PQ NB Genus NEGASTRIUS Thomson N. caurinus lHorn; BC N. choris (Sayt _ABSK MB ON PQ N. delumbi s tHorn) PE (Horn) yI' N. dispersus BC N. dubius ( Horn) _ABSK MB N. exiguus (Randallt guttatulus (Melsheimer) -oN N. extricatus (Fall) AK YK NT BC N. gentilis rLeConte) BC AB SK N. manki (Fall) BC AB N. melsheirneri tHorn) _PQ NB NS N. musculus (Eschscholtz) AK BC 178 Elateridae N. obliquatulus (Melsheimer) ON NS N. ornatus (LeConte) BC colon (Horn) moerens (Hotrr) N. pectoralis (Say) BC AB SK MB ON PQ NS futilis (LeCofte) inops (LeConte) N. restrictul us (Mannerheim.) AK YK NT PQ N. striatulus (LeConte) AK BC AB _ONPQNBNS N. tumescens LeConte AK YK NT BC AB SK MB ON PQ NB NS Genus CTENICERA Latreille Corymbites Latreille C. aeripennis (Kirby) AK YK NT BC AB SK tincta (LeConte) C. angularis (LeConte) BC C. angusticollis (Mannerheim) AK BC fraterna (LeConte) spectabilis (Mannerheim) C. appressa (Randall) ON PQ NB NF mirifica LeConte C. appropinquans (Randall) SK MB ON PQ NB NS PE NF C. arata (LeConte) -YK NT BC AB SK MB ON PQ NB NS NF C. atropurpurea (Melsheimer) PQ NF C. bipunctata (W.J, Brown) _BCAB C. bombycina (Germar) _BCAB C. callida (W.J. Brown) _BC C. carbo LeConte _BC C. caricina (Germar) AK _BC C. comes (W.J. Brown) -BC C. confluens (Gebler) AK C. conjungens (LeOonte) _BC C. consors (W.J. Brown) AB SK MB C. crestonensis (W.J. Brown) _BCAB C. cylindriformis (Herbst) ON PQ NB NS PE appressifrons (Say) breuicornis (Say) p ar al lel ip ip eda ( Germar) submetallica (Germar) C. destructor (W.J. Brown) AB SK MB C. diversicolor (Eschscholtz) C. dorothyae Knull PQ C. elongaticollis (Hamilton) PQ NB NS-- C. fallax (Say) ON PQ NS C. falsifrca (LeConte) ON PQ NB NS NF C. fulvipes (Bland) ON PQ NS-_ C. funerea (W.J. Brown) C. furtiva (LeConte) monticola Horn C. glauca (Germar) SK simili s sima ( Motschulsky) C. hamata (Say) ON PQ NB NS C. hieroglyphica (Say) _MB ON PQ NB NS C. hoppingi (Van Dyke) AI{ YK Nr sc AB SK MB ON PQ -? C. inflata (Say) ON PQ C. insidiosa (LeConte) ON PQ NB NS C. kendalli Kirby AK YK NT BC AB SK MB ON PQ NB _NF anchorago (Randall) C. laricis (W.J.Brown) _BC AB C. lateralis (LeConte) _BC q uadr iu ittat a (Walker) uittata (Walker) C. limoniiformis (Horn) MB C. lobata (Eschscholtz) AK NT TSK telum (LeConLe) C. lutescens (FaIl) _BC AB SK C. maura (LeConte) _BC C. mediana (Germar) NT AB SK ftfS ON pe NB NS r ubid.ipennis (LeConte) p(l C. mendax (LeConte) AK YK BC AB SK -oN NF 179 Elateridae C. moerens. tLeConte) _BC C. monLanrr (W.J. Brownl _BCAB C. morula rl,eConte) AK YK NT BC AB SK MB ON PQ LB brunnipes (Bland) C. nebraskensis t Bland) _BCAB C. nigricans tFall t _BC C. nigricollis (Bland) AK _BCAB -nneoNpe- C. nitidula (LeConte) AK NT BC AB SK MB ON PQ NB NTQ DF LB NF C. ochreipennis (LeOonte) AK YK NT BC AB LB NF C. opacula {Leconte) _BC :"yuoNPQ_ serricornis (Mannerheim) C. propola columbiana (W.J. Brown) AK YK BC AB C. propola propola (LeConte) NT BC AB SK MB ON PQ NB NS PE furcifer (LeConte) C. protracta { Leconte) BC C. pruininir tHornt BC no:rlo iHyslop) pruinosa (Schwarz) pruinosula (Schwarz) ( C. pudica W.J. Brown ) _BCAB C. pulchra tLeconte) NT BC AB SK MB ON PQ NB NS PE festiua (LeConte) C. pygmaert tVan Dyket NT BC AB SK MB ON PQ NB NS NF C. pyrrhos {Herbst} -oN aequa,',is (Melsheimer) pyrrhica (Haldeman) uagralLs (Melsheimer) C. respleniens aeraria (Randail) ? NT? AB SK MB ON PQ NB NS LB NF junAi , Schenkling) racinei (Chevrolal) uiridi.. (Germar) C. resplencens resplendens AK NT BC AB SK (Esr:hscholtz) C. rotundirollis r Sayt ON russicollis (Germar) stictico (Germar) C. rufopleuralis tFall t NB NS PE C. rupestri r {Germarr _BC _yuoNPQ C. sagitticc,llis (Eschscholtz) _BC C. semimel allica (Walker) _BCAB C. sexualis tW.J. Brown.) AB ix- - C. silvatica Van Dyke _BCAB - C. sjaelandica (O.F. MtiLiler)t oN PQ NU uiridis (Say') C. spinosa ' LeContet NF C. spf enderrs lZieglert _ _ _ ABSXMBSNiSNBNS_ _ C. stricklar:Ldi (W.J. Brown) AK _NTBCABSKMBON C. suckleyi suckleyi (LeConte) BC--laeoNpo---- C. sulcicoll:s tSay) parallela (Say) C. tarsalis Melsheimer; -oNPQ NB NS C. tristis tCandbzeJ BC C. triundulata (Randall) AK _NT BC AB SK MB ON PQ NB NS PE LB NF C. uliginostt tVan Dyket BC C. umbrico a (Eschscholtz) AK BC AB rzdis (Motschulsky) C. umbripennis (LeConte) BC AB gracilior (LeConte) n ub il ipennis (LeConte) C. vernalis (Hentz) ON PQ NS C. vidua tS'.J. Brown) BC AB C. volitans (Eschscholtz) AK BC uaria (W .J. Brown) C. vulnerata (LeConte) _PQ NS C. watsoni W.J. Brown) _PQ LB C. weidtii t.\ngellr nc-- Genus OXI'GONUS LeConte Key to North American species: Roache (1963) O. montanus Schaeffer PQ NB NS 180 Elateridae O. obesus (Say) MB ON PQ acutipennis (Randall) Genus EANUS LeConte P ar anom u s Kiesenwetter E. albertanus W.J. Brown _BCAB E. decoratus (Mannerheim) AK YK NT BC AB SK MB ON PQ LB NF paruicollis (Mannerheim) uagus (LeConte) E. estriatus (LeConte) AK YK NT BC AB MB ON PQ NB NS LB NF subarcticus W.J. Brown E. granicollis (Van Dyke) _BC E. maculipennis LeConte ON PQ NB LB NF pictus (Canddze) E. striatipennis W.J. Brown AK BC Genus HEMICREPIDIUS Germar Asaphes l\trby H. bilobatus (Say) ON c au ifron s (Melsheimer) c on s ent ane u s (LeOonte) mel ano p ht hahn us (Melsheimer) planatus (LeOonte) tener (LeConte) H. brevicollis (Candbze) MB ON PQ NB NS H. carbonatus (LeConte) BC AB SK MB coracinus Candbze H. dilaticollis Motschulsky BC H. hemipodus (Say) ON PQ aeneolus (Melsheimer) oereus (Melsheimer) corporosus (Germar) decoloratus (Say) H. memnonius (Herbst) AB SK MB ON PQ NS PE baridius (Say) ruficornis (Kirby) thomasi Germar H. montanus Lane BC H. morio (LeConte) BC uernus Canddze H. pallidipennis (Mannerheim) AK BC oregonus (LeConte) H. simplex (LeConte) _BC AB H. tumescens (LeConte) _BC Tlibe Oestodini Genus OESTODES LeConte O. puncticollis Horn AB SK MB O. tenuicollis (Randall) ON PQ NB NS PE g r ac ilifor mis (RandalI) obscuripes Candbze Subfamily EI"ATERINAE Tribe Elaterini Key to North American species: Roache (1961) Genus ELATER Linn6 Ludius Berthold Steatoderus Dejean E. abruptus Say oN PQ NS coracinus (Germar) Genus PARALLELOSTETHUS Schwarz P. attenuatus (Say) ON /uscus Castelnau 181 Elateridae Genus SERICUS Eschcholtz Atractooterus LeConte Dolerosomus Motschulsky Sericosomus Dejean S. blaisdelli (Van Dyke) BC S. honestus (Randall) -oN PQ NB fusiforrzis (LeConte) S. incongrurLs (LeConte) AK YK NT BC SK MB ON PQ NB NS LB NF behren::i (Canddze) S. silaceus (tjay) -oN PQ debilis (LeConte) flauipennis (Motschulsky) umbraticus (LeConte) S. viridanus (Say) ON PQ NS sublucens (Randall) Tlibe Agriotini Genus GLY?HONYX Canddze Key to Ncrth American species: Smith and Balsbaugh (1984) G. testaceus (Melsheimer) ON Genus AGRTOTES Eschscholtz Pedetes l{trby Key to N< rth American species: Becker (1956) A. apicalis l,eConte BC A. arcanus \V.J. Brown -o* A. avulsus (.LeConte) -oN A. collaris (LeConte) -oN PQ NSpe-- A. criddlei Van Dyke SC AB ix MB v A. fer"rugineipennis (LeConte) BC AB A. fucosus (],eConte) _AB SK MB ON pe NB NS--Nr A. hoodi Bec:ker BC A. isabellinus (Melsheimer) -on PQ A. limosus ( LeOonte) _NT BC AB SK MB ON PQ NE Ns pn is Nr A. Iineatus tLinn6X BC NS NF A. mancus (Say) _AB SK MB ON PQ NB NS PE LB striatutus Melsheimer A. oblongicollis (Melsheimer) PQ A. obscurus (Linn6)t BC _9^ NF A. opaculus (LeConte) BC AB -lt montartus LeConte _MBONPQ NB A. pubescens Melsheimer _NS A. quebecensis W.J. Brown yuoNPQ A. sparsus LeConte --sc- _ A. sputator il-inn6)f NB NS A. stabilis (l,eConte) BC SK MB ON PQ A. tardus W.J. Brown BC AB v A. thevenetl,i Horn BC ____ Genus AGRIOTELLA W.J. Brown A. bigeminata (Randall) yu PQ A. columbiana W.J. Brown BC 9* YNS A. occidentzrlis W.J. Brown BC A3 Genus DALOPIUS Eschscholtz D. agnellus W.J. Brown D. asellus W.J. Brown - scTsx- Iu_ D. brevicornis W.J. Brown - a*;: D. cognatus W.J. Brown oN PQ NB NS D. corvinus W.J. Brown BC D. fucatus W.J. Brown BC AB D. fuscipes W.J. Brown _PQ D. gartrelli W.J. Brown BC AB -v D. gentilis W.J. Brown D. inordinar;us W.J. Brown --scABSf-9*PQ D. insolens W.J. Brown D. insolitus W.J. Brown -_l"T--oNpe r82 Elateridae D. insulanus W.J. Brown _BC D. maritimus W.J. Brown _BC D. mirabilis W.J. Brown ABSKMB_PQ D. pallidus W.J. Brown _BCABSKMBONPQ NB \Tq DF _NF D. parvulus W.J. Brown -- MB- D. spretus W.J. Brown - - ec - :" - D. suspectus W.J. Brown _BC D. tristis W.J. Brown D. vagus W.J. Brown NS D. vernus W.J. Brown _ - -:"-:"ilB3Nia v Tribe Ampedini Genus AMPEDUS Dejean A. antracinus (LeConte) -BC A. apicatus (Say) AK_ NT BC AB SK nfS O* PQ NB NS PE - lrr melanopygus Germar A. areolatus (Say) -oN PQ obliquus (Say) scitulus Germar A. atripennis (Horn) _BC u iolaceipennis (Candbze) A. behrensi (Horn) _BC A. brevis (Van Dyke) T_ _BC AB ___ A. collaris (Say) MB_PQ NS- A. deletus (LeConte) AK NT BC AB SK MB ON PQ NB NS is N" A. dimidiatus (LeConte) _BC A. evansi W.J. Brown NT_ -s* MB ON PQ NB NS A. fuscatus (Melsheimer) -oNPQ A. fusculus (LeConte) AB SK MB ON PQ NB iey A. impolitus (Melsheimer) _ONPQ A. laesus (LeConte) -oN A. laurentinus W.J. Brown AK- Nr sc AB_ MB ON PQ NS NS- _NF A. linteus (Say) _ONPQ lugubris Germar A. luctuosus (LeConte) NT MB ON PQ NB NS_ LB NF A. melanotoides W.J. Brown YSK -oN PQ A. melinus (LeConte) -BC A. melsheimeri (Leng) _SK MB ON PQ NS hepaticus (Melsheimer) A. militaris (Harris) (LeConte) ie Ne A. miniipennis Nr- AB- - SNFBNn- - _NF A. mixtus (Herbst) _v AB SK MB ON PQ NB NS PE A. moerens (LeConte) NT BC AB A. molestus (LeConte) -BCMBONPQNBNS A. nigricans Germar AB SK MB ON PQ NB NS A. nigricollis (Herbst) -oNPQ A. nigrinus (Herbst)* AK YK NT BC AB SK MB ON PQ NB A. oblessus (Say) AB SK MB ON PQ NB :"y sellatus (Leng) discoideus (Fabricius) A. occidentalis Lane BC AB bimaculatus (Van Dyke) A. pedalis Germar AB SK MB ON PQ rufipes (Palisot de Beauvois) A. phelpsi (Horn) BC AB hoppingi (Van Dyke) A. phoenicopterus Germar BC A. protervus (LeConte) ---o*pe NS A. pullus Germar AK YK NT BC AB SK MB ON PQ NB NS -N" Iacustris (LeConte) A. pusio Germar -oN luteolus (LeConte) A. quebecensis W.J. Brown NT BC AB SK MB PQ LB NF A. rhodopus (LeConte) T BC A. rubricollis (Herbst) -oN PQ socer (LeConte) uerticinus (Palisot de Beauvois) A. rubricus (Say) MB3N|8- A. sanguinipennis (Say) - - - - N3- - - 183 Elateridae A. sayi (Let)onter - ON A. semicinrtus rRandall) - ON PQ - NS A. subtilis tleOonte) - BC AB SK MB ON PQ - NS A. uteanus (Van Dyker - SK A. varipilis (Van Dyket - \'K NT BC AB columoianus W.J. Brown A. vitiosus LeContet - ON PQ NB Genus ME(IAPENTHES Kiesenwetter Dolopt osomus Motschulsky Key to Nrrth American species: Becker (19?1) M. caprella tl-eContet AK BC califor nicus W.J. Brown M. gentner Lane - BC M. nigriventris LeConte - BC M. rogersi }{orn ON PQ NB M. solitariL.s Fall PQ M. stigmosus (LeConte) BC AB SK MB ON PQ NB M. tartareus (Lecontej - BC Tribe Physorhini Genus ANC HASTUS LeConte B r ac h2,' cre p i s LeConte Crepiaotritus LeConte Elatrh,.us Horn A. anthrax Horn.) - BC coruzs Van Dyke militaris Candbze A. cinereipt nnis tEschscholtzr - BC AB hir s utul us Motschulsky recedens LeConte p ilifer t.: s Motschulsky puberu.lus (Mannerheim) regularis Motschulsky ta nt il I t.t s ( Mannerheim) Tribe Melanotini Genus MEL,ANOTUS Eschscholtz Cratonychus Dejean Ctenon,ychus Stephens Perimecus Stephens Key to North American species: Quate and Thompson (1967) M. americanus (Herbst) -oN ang u st.a tu s (Erichson) M. castanipes (Paykull)t MBONPQNBNSPE_NF inaequ,zlis (LeConte) paradaxus Melsheimer scrobiccllis (LeConte) texanu:; Canddze M. communis (Gyllenhal) _ONPQ pag(tnt,ts Canddze M. corticinus (Say) rlNT g I andieo lor Melsheimer uetulus (Erichson) M. decumar:Lus (Erichson) _ONPQ_NS canadensis Candbze cuneo.tus (LeConte) incertus (LeConte) M. depressus (Melsheimer) .rNT diu ar c arinus Blatchley lixus Blatchley p ar um p unc tatas (Melsheimer) M. hyslopi \/an Zwaluwenburg - ON PQ M. ignobilis Melsheimer -oN secretu.s (LeConte) M. leonardi (LeConte) MB ON PQ 184 Elateridae M. longulus oregonensis (LeConte) BC franciscanus Van Dyke uariolatus LeConte M. miscellus Quate -oN M. morosus Canddze -oN PQ debilis Blatc}:.ley longulus (LeConte) simulans Blatchley M. prasinus Blatchiey M. sagittarius (LeConte) _9* PQ carinus Blatchley M. similis (Kirby) _SK MB ON PQ NB NS flssilis (Say) exuberans (LeConte) oc hr acei p ennis (Melsheimer) s p henoidal is (Melsheimer) M. trapezoideus (LeConte) ON Subfamily CARDIOPHORINAE Genus CARDIOPHORUS Eschscholtz C. acutus Lanchester _AB C. alpestris Lanchester BC C. brunnipennis Lanchester BC C. cardisce (Say) _AB SK MB ON PQ NB NS dejeanii LeConte C. communius Lanchester BC C. convexulus LeOonte --oNpeNe---- C. convexus (Say) ON PQ amictus Melsheimer insulsus Canddze C. edwardsi Horn _BC C. fenestratus LeConte NT BC AB larquini Candbze C. gagates Erichson MB ON PQ NB NS laeuicollis Erichson C. kooskooskiensis Lanchester BC AB C. Iatiusculus Eschscholtz BC C. montanus Bland _AB C. nevadensis Blanchard BC C. plebejus Lanchester BC C. plexus Lanchester BC C. pratensus Lanchester _AB C. propinquus Lanchester BC C. pubescens Blanchard BC AB SK MB C. silvanus Lanchester _AB C. sodalis Lanchester _AB C. tenebrosus LeConte BC AB SK MB fuluipes LeConte C. tumidicollis LeConte BC AB C. tutus Lanchester BC AB Genus HORISTONOTUS Canddze H. curiatus (Say) ON areolatus (Erichson) 185 Eucnemidae: FamilyTHROSCIDAE throscidbeetles Y. Bousquet Throscidae includes about 30 North American (1979) treated the lissomines as a subfamily of species, of which nine have been reported in Eucnemidae; Lawrence (1982a) listed them in Canada. The adults, which look like small, stout Elateridae. I have followed Yensen (1975) in con- elaterids, are usually found on foliage and flowers. sidering the lissomines as a subfamily of The larvae occur in litter, in decaying wood, and Throscidae. Yensen (1975) provided a key to the under bark. extant genera of the family and revised t;he genus Few authors agree on the position of the lis- Trixagus. somines. Crowson (1961) included Drapetes in Elateridae and Lissomus in Throscidae; Burakowski NT (2); BC (6); AB (1); MB (1); ON (5); PQ (5); NB (1975) el:vated the group to family level; Lohse (2);NS (2);NF (1) Subfamily LISSOMINAE Genus DRAIIETES Dejean D. gerninatus (Say) oN PQ ame r icc,nus ( Gerstaecker) basalis (Randal1) exstriatus (Say) Subfamily THROSCINAE Genus TRD(AGUS Kugelann Throsct.Ls Latreille Key to North American species: Yensen (1975) T. carinicoll s tSchaeffer.t _BC AB ON PQ NB NF T. chevrolat {Bonvouloirj _BC ON PQ NS paruulu.s (LeConte) sejunctus (Hotn) T. mendax (.lorn) NT BC T. sericeus ( LeConte; NT BC debilis t}{orn) Genus AULONOTHROSCUS Horn A. constrictor (Say) MB ON PQ NB NS A. punctatus (Bonvouloirt PQ A. validus (LeConte) _BC _9* Genus PACI]IOPUS LeConte P. hornii (LeConte) BC fuchsi Casey Family EUCNEMIDAE false click beetles Y. Bousquet Thirty-two species of eucnemids have been many changes and additions to the Canad.i.an fauna reported in Canada and Alaska. The adults occur can be expected. Horn's revision (1886) is the latest mainly on vegetation and under bark but are rather one for the North American species. infrequently collected. The larvae bore into various hardwoods that are usually rotten. Like elaterids, adults have the clicking mechanism, but few species have been actually observed to click. The taxonomy of the North American species of AK (1); BC (13); AB (2); SK (1); MB (3); ON (18); PQ Eucnemidae has been poorly studied, and therefore (20);NB (5);NS (3);NF (1) 186 Eucnemidae Subfamily EUCNEMINAE Genus STETHON LeOonte S. pectorosus LeConte - ON PQ Genus DELTOMETOPUS Bonvouloir D. amoenicornis tSay) - ON PQ NB NS clypeatus (Say) ereptus Bonvouloir D. rufipes (Melsheimert Genus DROMAEOLUS Kiesenwetter D. basalis (LeContet - BC D. californicus Bonvouloir - BC D. cylindricollis (Sayt - ON PQ D. harringtoni Horn MB ON PQ NB D. nitens Horn - BC Genus FORNAX Castelnau Isarl hrus LeConte Onichodon Newman F. badius (Melsheimer.t - ON PQ basalis Hotn nouitius (Bonvouloir) F. bicolor (Melsheimer) PQ luridus Chevrolat F. calcealus tSay) - ON PQ spretus (LeConte) F. canadensis W.J. Brown - ON PQ NB NS F. hornii Bonvouloir - ON PQ F. orchesides (Newman) - ON PQ Genus PHAENOCERUS Bonvouloir P. americanus Horn - BC Subfamily DIRIIAGINAE Key to genera: Cobos (1964) Genus DIRHAGUS Latreille Microrhagus Dejean D. pectinatus (LeConte) - BC - ON PQ NB D. subsinuatus (LeConte) MB ON PQ imperfectus (LeConte) oblilus (Bonvouloir) D. triangularis tSayt - ON PQ Genus ENTOMOPHTTIALMUS Bonvouloir E. rufiolus (LeConte) - ON PQ pallens Bonvouloir Genus SARPEDON Bonvouloir S. scabrosus Bonvouloir - BC - ON PQ Genus RHAGOMICRUS Fleutiaux R. humeralis { Say) PQ meticulos us (Bonvouloir) Subfamily MELASINAE Genus ISORHIPIS Boisduval & Lacordairei I. obliqua (Say) - BC - ON PQ NB NS Genus MELASIS Olivier M. rufipennis Horn - BC M. tsugae Hopping - BC 187 Lycidae Genus EPIPHANIS Eschscholtz E. cornutus Eschscholtz AK BC - SK - ON PQ NB - NF cristatus LeConte Genus SCHI ZOPHILUS Bonvouloir S. subrufus Randall,l sirnplet (LeConte) trilobatus Bonvouloir Genus HYLI.S des Gozis H. frontosus tSayt canaliculatus (LeConte) H. terminalrs (Leconte, - BC AB SK - ON PQ Genus XYLOPHILUS Mannerheim Xylobius Latreille Xyloecus Dejean X. cylindriformis lHornt - BC Genus NEMATODES Berthold Hypocozlus Dejean N. atropos {l;ay} MB N. penetran; (Leconte) - ON PQ Genus ANE,,ASTES Kirby A. druryi Kirby - BC AB brunne.rc (Latreille) aroszs (Say) latreille'i LeConte Superfamily C antharoidea Family LYCIDAE net-winged beetles J. McNamara Twent;y-nine species of Lycidae are recorded in tions; his paper of 1949 includes a key to the Canada and Alaska. The adults occur on leaves, on Nearctic tribes. flowers, or under bark and are thought to be preda- ceous. The larvae are predaceous, and those of most species are found under bark. Greer, has revised the taxonomy of the North AK (3); yK (1); NT (1); BC (8); AB (5); SK (a); MB American species of Lycidae in a series of publica- (8); ON (25); PQ (19);NB (5);NS (5); LB (1); NF (1) Subfamily LYCINAE Tlibe Lygistopterini Key to North American species: Green (1950) Genus CAI-OCHROMUS Gu6rin-M6neville C. dimidiatr,is (LeConte) -BC C. fervens LeConte C. perfacetus (Say) ---:"Tsx-o*pe----- substritttus (LeConte) Tlibe Platerodini Key to North American species: Green (1951, 1953) Genus DICTYOPTERUS Mulsant D. aurora (Flerbst) AK YK NT BC AB _ MB ON PQ NB NS _ LB NF coccinatus (Say) nigripes (Schaeffer) 188 Lycidae D. hamatus Mannerheim AK BC D. simplicipes Mannerheim AK BC AB loelzs Motschulsky D. thoracicus (Randall) BC AB SK MB ON PQ praefectus (Newman) Genus EROS Newman E. humeralis (Fabricius) ON PQ incestus LeConte obliquus (.Say) Genus PLATYCIS Thomson Erotid,es Waterhouse P. sculptilis (Say) ON PQ axill aris (Melsheimer) oblira (Newman) Genus LOPHEROS LeConte L. crenatus (Germar) MB ON PQ cruciatus (Randall) L. fraternus (Randall) -oNPQ Genus EROPTERUS Green E. arculus Green _ONPQ E. trilineatus (Melsheimer) _ONPQ Genus PLATEROS Bourgeois Neoceletes Leng & Shoemaker P. batillifer Green -oN P. bispiculatus Green -oNPQ P. borealis Green -oN P. centralis Green -oN P. fl avoscutellatus Blatchley -oNPQ NS P. floralis (Melsheimer) P. lictor (Newman) ecABsxmn3Ni3 NB NS californicus Van Dyke - columbiensis W.J. Brown crateracollis (Leng & Shoemaker) P. peregrinus Green _BC oN PQ P. subfurcatus Green ON P. timidus (LeConte) ON P. volatus Green ON PQ NB Tlibe Calopterini Key to North American species: Green (1952) Genus CAI-OPTERON Gu6rin-M6neville Diagrapha Newman C. discrepans (Newman) yuSN PQ C. reticulatum (Fabricius) affinis (LeConte) - - - retiferum LeConte typicum (Newman) C. terminale (Say) MB ON PQ apicale (LeConte) diuisum (Newman) dorsale (Newman) duplicatum (Haldeman) Genus CAXNIA Newman Caeniella Cockerell C. dimidiata (Fabricius) MB ON PQ NB NS scapularis Newman Genus CELETES Newman C. basalis LeConte SK MB ON PQ NB NS mystacina LeConte tabida LeConte 189 Lampyrrida e FamiIyPHENGODIDAE glowworrns J. McNq.mara Only one species of Phengodidae occurs in debris on the ground. The males are al;tracted to Canada. The larvae of most species are lumines- light. cent, hertce the vernacular name glowworms. The Wittmer .1975) revised the North American larvae fe,,ed on soft-bodied insects and other small species of the genus Phengodes, the only genus organisrls. The females are larviform but have occurring in Canada. compound, instead of simple, eyes. Adults and lar- vae occur in leaf litter and under logs, bark, and oN (1) Genus PHIINGODES Illiger Key to North American species: Wittmer (1975) (Subgenus PHENGODES s. str.) P. plumosa tOliviert ON Familv LAMPYRIDAE fireflies J. McNamara Twenty-nine species of Lampyridae are slugs. The feeding habits of the adults are not well recorded in Canada. Members of this family are known. fairly well known because of their ability to produce Many genera of lampyrids have been revised in flashing light. Species may be recognized in flight the past few decades, but a thorough revision ofthe by the color of the light as well as by the number family would be very useful. Green (1948) pub- and the length of flashes. This flashing behavior lished a revision of the Nearctic genera. enables l;he sexes to frnd each other. All larvae and most adults are luminescent. The larvae are preda- xT (3); BC (9);AB (3); SK (5); MB (12); ON (19); PQ ceous and feed on other insects, larvae. snails, and (14);NB (8);NS (10);PE (1);NF (3) ' I (1) Subfamily LAMPYRINAE Tribe Cratomorphini Genus PYI|ACTOMENA Melsheimer Lecontea E. Olivier Key to North American species: Green (1957) P. angulata (Say) NT SK MB ON PQ NB NS flauoc:,.ncta LeConte P. borealis (Randall) AB SK MB ON PQ NB NS P. dispersa Green NT SK MB P. linearis LeConte MB ON PQ P. lucifera Melsheimer ON P. sinuata Green MB T?ibe Photinini Genus PHOTINUS Castelnau Key to N'orth American species Green (1956) P. aquilonius Lloyd MB ON PQ_NS P. ardens LeConte MB ON PQ NB NS NF taedif'zr LeConte P. consimi]is Green -oN P. ignitus llall -oN P. indictus (LeConte) -oN ablucens Fall P. marginellus LeConte -oN PQ castus LeCor,:te 190 Lampyridae P. obscurellus LeConte MB ON PQ frigidus Olivier P. punctulatus LeConte ON P. scintillans (Sayt ON Genus PHAUSIS LeConte Key to North American species: Fender (1966) P. nigra Hopping BC P. rhombica Fender BC AB- Genus ELLYCHNIA LeConte Key to western North American species: Fender (1970) (Subgenus ELLYCHNIA s. str.) E. cornrsca (Linn6) -BC y" oN PQ NB NS NF E. facula LeConle _BC E. greeni Fender _BC E. hatchi Fender _BC E. lacustris LeConte _BC Genus PYROPYGA Motschul sky Keyto species: Green (1961) P. decipiens (Har-ris) -- P. nigricans (Say) NrscABixnrnSNiSNENS - - californica Motschulsky californica (Gorham) fene stralis (Melsheimer) reuersus (Gemminger) Genus LUCIDOTA Castelnau RileyaE. Oliviet L. atra (Olivier) MB ON PQ NB NS laticor nis (Fabricius) morio (Melsheimer) tardaLeConte Genus PHOSPHAENUS Castelnau P. hemipterus (Fourcroy)f PQ NS Subfamily ANIYDETINAE Genus POLLACLASIS Newman Polyclasis LeConte P. bifaria (Say) ON PQ oualus Newman Subfamily PHOTURINAE Genus PHOTURIS Dejean Py r ec tosoma Motschulsky Key to North American species: Barber (1951) P. fairchildi Barber NS P. pennsylvanica (DeGeer) BC_SKMBON PQ NB NS PE NF marginata (Panzer) u ittiger a (Motschulsky) 191 Cantharid,re Family CANTHARIDAE soldier beetles J. McNamqra Cantharidae includes 126 species and subspecies Some of the genera of Cantharidae are poorly in Canarla and Alaska. The adults are common on known taxonomically, including the large genus goldenrc,d, milkweed, and hydrangea, where they Podabrus, and are in need ofrevision. Fender (1962) feed on nectar and pollen. Most larvae are carnivo- provides keys to the species of the Pacifrc Northwest. rous, gererally preying on other soft-bodied insects; a few are cmnivorous, feeding on plants such as wheat, aK (15); vK (8); NT (13); BC (57); AB (20); SK (9); potatoes. celery, and other plants. They occur mostly MB (22); ON (a8); PQ (48); NB (22); NS (rl5); PE (1); beneath bark and debris. LB (5); NF (16) . I(2) Subfamily Cantharinae Tlibe Podabrini Genus PODABRUS Westwood Brachynotus I\irl:y Malthacus l\trby Key to Irorth American species Fa]l (1927) P. alexand,rri Fender _AB P. brevicol,is Fall ON P. cavicolli; cavicollis LeConte BC P. conspiratus Fall AK BC P. deceptus W.J. Brown AK BC MB P. diadema (Fabricius) vy oN PQ NB NS l*" P. excursur; excursus Fall BC AB P. extremus LeConte AK \.K NT BCAB-MB ON LB P. falli Hop,ping BC AB P. fissilis Fall AK BC AB P. flavicollis LeConte ON P. frater LeConte ON q uadr,.collis ( Motschul sky) P. frosti Fender ON _NB P. fumiganus Green PQ NB P. furtivus Fall _BC P. heteronl'chus Fall NT BC AB SK MB ON PQ P. instabilis Fall TY _ IJU P. intrusus Green ON PQ NB NS _NF P. laevicoil s (Kirby) AK YK Nt sc AB SK MB ON PQ NB LB NF P. limbellur LeConte PQ NB P. lutosus LeConte _BC P. lygarius Fender _BC P. melvillei Fender _BC P. modestus (Say) ON pe-Ns -Ne P. nothoides LeConte MB ON PQ P. obscurevittatus Fall _YK NT BC AB ix MB P. pattoni LeConte PQ NB P. perplexus W.J. Brown AK NT MB P. piniphilus (Eschscholtz) AK NT BC AB MB PQ P. planulus Green MB ON PQ_NS P. probus FaIl PQ-_ _Iu P. protensu s LeConte ON P. pruinosts atrocervicus Fender _BC P. pruinosr;.s diversipes Fall _BC AB P. puberulus LeOonte MB ON PQ NB NS LB NF P. punctatuLs LeConte PQ NB NS P. puncticollis (Kirby) PQ NB marginellus LeConte P. punctula.tus LeConte ON PQ P. pustulatus Fender _BC P. pygmaeus Green PQ P. rossi Fender _BC P. rugosulus LeConte ON PQ NB NS p ar u icol lis (Motschulsky) P. scaber LeConte 192 Cantharidae P. secretus W.J. Brown PQ P. sericatus (Mannerheim) AK P. simplex Couper na" o* PQ NS P. smithi Fender _BC IU P. tetragonoderus Fall AK_ _BCAB P. tomentosus (Say) -oN cinereipennis Motschulsky rufiolus (Melsheimer) P. tricostatus (Say) ON PQ atripes (Motschulsky) benneti (Kirby) Tlibe Cantharini Genus ANCISTROI.JYCHA Markel Key to North American species: McKey-Fender (1950) A. bilineata tSayt oNPQ-NS A. dentigera (LeContet oN PQ A. neglecta tFall r oN PQ Genus CANTIIARIS Linn6 C y r tomo p tila Motschulsky Telephorus Schaeffer C. alticola (LeConte) BC C. aneba borealis McKey-Fender NT BC AB SK MB C. beta Fender BC C. consors (LeConte) BC tibialis (LeConte) tibiella (Gemminger) C. curtisi (Kirby) - YK NT BC SK MB ON PQ NB LB NF samouellei (Kirby) C. impressa (LeConte) -oNPQ-NS C. oregona tLeConte) BC C. rotundicollis Say -oNPQNBNS C. rufa Linn6t NF C. transmarina transmarina ec -_-9*PQIUNS:' (Motschulsky) C. transmarina scopa (LeConte) BC C. tubercu lata ( LeConte) PQ armigera (Couper) breuicollis (LeConte) C. westwoodi tKirby) Genus CYRTOMOPTERA Motschulsky C. divisa (LeConter - BC latiu sc ula Motschulsky Genus RHAXOI.IYCHA Motschul sky Key to North American species: McKey-Fender (1950) R. bilobata (McKey-Fender) MB oN PQ R. carolina t Fabricius) oN PQ jactata (Say) Genus RHAGONYCHA Eschscholtz Key to North American species: Green (1940) R. alaskensis Fender AK R. anthracina Mannerheim AK R. costipennis (LeConte) -oN------R. excavata (LeConte) BC MBONPQ-NS R. fraxini (Say) AK BC AB MBONPQ_NSNF atra (l\trby) binodula Mannerheim nigrita (LeConte) R. fulva (Scopoli)t -BCPQ R. gilvipes (Gemminger) NT_? dichroa (LeOonte) flauipes (LeConte) 193 Cantharidaer R. imbecillis (LeConte) ON NF albolineat a (Blatchley) R. luteicollis (Germar) ON cinctellt (LeConte) R. mandibuiaris (Kirby) AK YK NT BC AB SK MB -PQ LB NF nigritu.la (LeConte) R. mollis mc,llis Fall SK ON NS NF nigriceps (LeConte) pusilla (LeConte) R. nanula (LeConte) NS R. oriflava ( LeConte) ON R. recta (Melsheimer) ---ecAB-- oN PQ NS NF pusilla (LeConte) rufipes (Say) R. scitula (Say) _PQ NS R. septentri,rnis (Green) MB ON R. syivatica (Green) ON R. tantilla (l,eConte) -SK pzsio (l,eConte) R. vilis (LeConte) _ONPQ R. walshi (LeConte) MB Tfibe Silini Genus POLItrMIUS LeConte P. canadensis W.J. Brown P. laticornis (Say) _ONPQ fuft_ _ _ dubjzs (Melsheimer) incisus LeConte inualid,.rs (Say) undulatus (LeConte) Genus SILII) Charpentier Key to Ncrth American species: Green (1966) (Subgenus SIILIS s. str.) S. atra LeC<.rnte NT S. difficilis ciffrcilis LeConte _YK NT BC y:" MB PQ NB S. insperata. Green _BC S. lecontei Cireen _BC S. lutea LeConte _BC pallens LeOonte S. paliida Mannerheim AK _BC S. parallela Green _BC S. percomis iSay) -o* PQ NB NS curta (7,eConte) longico"nis LeConte S. spinigera LeConte BC minuta LeConte S. striatella Green BC (Subgenus DITEMNUS LeConte) S. bidentata (Say) PQ S. latiloba Blatchley MB Tlibe Malthinini Genus MAJ.THINUS Latreille Key to Ncrth American species: Fender (1972) M. occipitalis LeConte oN PQ Genus MALTHODES Kiesenwetter Key to North American species: Fender (1951) M. abruptus Fender _BC M. acuticauda Fender _BC M. alexanderi Fender _BC M. bakeri Fender _BCAB M. canadensis Fender _BC 194 Derodontidae M. columbiensis Fender BC M. fragilis (LeConte) ON PQ NB NS trdnsuersus (LeConte) M. fuliginosus fuliginosus LeConte _PQ NB NS M. fuliginosus flavicollis Fender NS M. intricatus Fender M. macnabi Fender M. medioccidens Fender ON M. niger (LeConte) _PQ NN T\TIT M. oregonus Fender M. parvulus (LeConte) oN PQ NB NS_- NT M. piceolus Fall M. piperi Fender M. similis Fender ON NS M. stacesmithi Fender M. terraenovae Fender NF Subfamily CIIAULIOGNATHINAE Tlibe Chauliognathini Genus CHAULIOGNATHUS Hentz Key to North American species: Fender (1964) C. pensylvanicus (Deceer) MB ON PQ bimac u la t us (Fabricius) Tlibe Ichthyurini Genus TRYPHERUS LeConte Lygerus Kiesenwetter Key to North American species: Fender (1960) T. frisoni Fender ON Superfamily Dermestoidea FamilyDERODONTIDAE tooth-neckedfungus beetles D.E. Bright The family Derodontidae includes four genera documented the feeding habits of Lqricobius erich- and 19 species of small beetles that occur in the sonii quite well. Adults and larvae feed on the pine temperate parts of both the Northern and Southern and spruce aphids of the family Adelgidae. The pre- Hemispheres. They are rarely encountered in the ferred host of L. erichsonii is the balsam woolly freld and are uncommon in collections. Three gen- aphid; the derodontid was introduced into eastern era and eight species are found in Canada and Canada in the 1950s for the control of this pest. Alaska. With the exception of Laricobius species, The other species of Laricobius also feed on other both larvae and adults of this family feed on various species of Adelgidae. kinds of fungi or on the products of fungal Information on adult ahd larval morphology, metabolism. Peltastica species appear to be feeding behavior, life cycle, and evolution, as well as restricted to fermenting sap flows or to areas under keys and diagnoses ofall genera and species, can be bark, where bacterial or fungal fermentation takes found in Lawrence and Hlavac (1979). Lawrence place. Derodontus species feed on the fruiting bod- (1989) recently published a catalog of the North ies of a wide range of higher Basidiomycetes. The American species. beetles prefer fresh fruiting bodies, although they have been recorded in rotting mushrooms, "slimy AK (2); BC (5); aB (3); ON (2); PQ (1); NB (2); NS fungus," and "watery fungus." Franz (1958) has (3);NF(1) . I(1) 195 Dermestida,: Subfamily PELTASTICINAE Genus PELi|ASTICA Mannerheim P. tubercula.ta Mannerheim AK BC Subfamily DERODONTINAE Genus LARI.COBIUS Rosenhauer L. erichsoni Rosenhauert _BC NF L. laticollis Fall _BCAB YNS L. nigrinus Fender L. rubidus l,eConte ---:cT-,oNpe NB Genus DERCDONTUS LeConte D. esotericus Lawrence AB D. maculatu,s (Melsheimer) __oN__N3__ D. trisig:natus (Mannerheim) AK ec Family NOSODENDRIDAE wounded'tree beetles J.M. Campbell Noso Genus NOSODENDRON Latreille N. californir:um Horn BC N. unicolor Sav ON Family DERMESTIDAE dermestid beetles Y. Bousquet DerrrLestidae is represented in Canada and The taxonomy of the North American der- Alaska b;y 48 species and subspecies. Dermestids mestids is relatively well known, thanks to the are usually found on flowers, on dried animal car- work of Beal, who revised most genera occurring in casses, in social Hymenoptera, bird and mammal the area. Mroczkowski (1968) published a catalog of nests, where they feed on pollen, feathers, hair, the world fauna. fur, or the remains of insects. Many species, how- eyer, are household and museum pests; they cause damage to a wide variety of products, such as car- pets, silk, fur, feathers, wool, leather, seeds, eK (6); rlK (3); NT (6); BC (31); AB (16); SK (10); grain, and cereal products as well as dried-insect MB (14); ON (2a); PQ (1e); NB (a); NS (5); PE (4); collection s. NF(4) . J(17) Tlibe Derrnestini Genus DERMESTES Linn6 (Subgenus DERMESTINUS Zhantiev) D. caninus {}ermar BC AB SK ON 196 Dermestidae murinus LeConte nubilus Say tessellatus Hetbst D. carnivorus Fabriciust BC rnucoreu,s LeConte sobrinus LeConte uersicolor Castelnau D. fasciatus LeConte SK MB D. frischii Kugelannt _ _ _ ncT ONPQ_NS D. maculatus DeGeert _BC _MB oN PQ elongatus Hope lupinus Mannerheim truncatus Casey uulpinus Fabricius D. marmoratus Say _BCAB_MB D. talpinus Mannerheim AK YK NT BC AB SK MB oNpeI"_ D. tristis Fall _BCAB (Subgenus DERMESTES s. str.) D. ater DeGeert _MBONPQ D. lardarius Linn6f --NrecAB SK MB ON PQ NB NSPE_NF D. pulcher LeConte _MB_PQ D. reductus Kalik _BC D. signatus LeConte _BCAB SK MB T[ibe Attagenini Key to North American species: Beal (1970) Genus ATTAGENUS Latreille A. brunneus Faldermannt elongatulus Casey extricatus Casey A. pellio (Linn6)t -BC NS A. schaefferi hypar Beal _BC A. unicolor japonicus Reittert -BC-SKMBONPQ PE NF canadensis Casey A. unicolor unicolor (Brahm)t megatoma (Fabricius) piceus (Olivier) Genus NO\IELSIS Casey Genattus Sharp (Subgenus PARANOVELSIS Casey) N. perplexa (Jayne) BC Tribe Megatomini Key to North American genera: BeaI (1967) Genus TROGODERMA Dejean Key to North American species: Beal (1954) T. glabrum (Herbst)t AB -oN PQ boronBeal T. inclusum LeConte BC -SK MB ON PQ aduena Casey brunnescens Casey frosti Casey nigrescans Casey obsolescens Casey T. ornatum (Say) ON pallipes Ziegler paruulum Casey procerum Casey pusillum LeConte serriferum Dalla Torre serrigerum Casey lorsole Melsheimer T. simplex Jayne BC ajar Casey pollens Casey 197 Dermestidae T. sinistrurn FaIl AK YK NT BC AB SK )IB ON PQ T. sternale sternale Jayne BC _ONPQ caseyi Arrow simuh.tns Casey uariipes Casey T. variabilr'Balliont _AB SK ON PE parab: Ie Beal scabricennis Casey Genus REESA Beal R. vespulac (Milliront NT BC AB SK MB ON PQ PE NF Genus MEGATOMA Herbst Key to North American species: Beal (1967) (Subgenus MEGATOMA s. str.) M. pubescens (Zetterstedt) BC prolira BeaI (Subgenus PERIMEGATOMA Horn) M. ampla ('lasey) BC M. angularis Mannerheim AK M. cylindri:a (Kirby) AK YK NT BC AB ON impre:;sa (Casey) montit:ola (Casey) M. giffardi (Blaisdel]) AK M. leucochlidon Beal ^-ec-- M. variegar.a tHornt BC AB guadalupensis (Casey) jaynei (Casey) neuad;',ca (Casey) Genus PStr UDOHADROTOMA Kalik Key to North American species: Beal (1967) P. kaliki B,ral BC P. perversa (Fall) AK BC AB Genus DETjIRTHRUS LeConte Key to North American species: Beal (1967) D. stebbinsi Beal BC Genus CR\?TORHOPALUM Gu6rin-M6neville Key to North American species: Beal (1985o) C. haemonhoidale (J.E. LeConte) -oN C. triste Le Conte -oN fuscicl,aue Casey modestum Casey nigriatrne LeConte picicorne LeConte Tribe Anthrenini Genus AN|IIHRENUS I.C. Schaeffer Key to errstern Nonh American species: Hoebeke et a]. (1985) (Subgenus ANTHRENUS s. str.) A. pimpinellae Fabriciust BC conspersus Casey lepidus LeConte neuad,icus Casey obtectus Casey occidens Casey pictus Casey suffusus Casey A. scrophu ariae (Linn6)f BC ON PQ NB NS thoracicus Melsheimer (Subgenus NATHRENUS Casey) 198 Bostrichidae A. verbasci (Linn6)t BC AB MB ON PQ caseyi Hinton destructor Melsheimer pistor Casey substriatus Casey uorax Casey (Subgenus FLORILINUS Mulsant & Rey) A. castaneae Melsheimer ON PQ NB carolinae Casey A. museorum (Linn6)t ON PQ NF (Subgenus HELOCERUS Mulsant & Rey) A. fuscus Oliviert oN PQ NS Tribe Orphilini Genus ORPHILUS Erichson Key to North American species: Beal (1985b) O. ater Erichson oN PQ O. subnitidus LeConte BC aequalis Casey chalybeus Casey Tribe Thylodriini Genus THYLODRIAS Motschulsky T. contractus Motschulskyt NT AB MB ON PQ Superfamily Bostrichoidea FamilyBOSTRICHIDAE bostrichidpowder'postbeetles J. McNamara Bostrichidae comprises 18 species in Canada Members of Bostricidae have been widely dis- and Alaska. It includes some of the most destruc- tributed through commerce because of their ten- tive beetles known. Some species cause consider- dency to bore into wood products, such as crating able damage to such stored products as dried roots and bamboo. and grains. In the tropics these insects are espe- Fisher (1950) revised the North American cially destructive, injuring felled timber and bam- species of the family. boo. The larvae do most of the damage. Some species can kill weakened trees; others have the AK (1); vK (1); NT (3); BC (8); AB (3); SK (2); MB habit of tunneling into the gTeen shoots of plants. (6); ON (13); PQ (6);NB (1);NS (1) ' I (1); H (1) Subfamily PSOINAE Tlibe Polycaonini Genus MELA.LGUS Dejean E ro pioide s Gu6rin-M6n6ville Heterarthron Dejean M. confertus (LeConte) BC inciszs (LeConte) Genus POLYCAON Castelnau Alloeocnemis LeConte P. stoutii (LeConte) BC ouicollis (LeConte) 199 Bostrichidac Tribe Psoini Genus PSOI, Herbst Acrepis LeConte P. quadrisig:rata tHornr - BC sexguttt.,.ta Lesne Subfamily ENDECATOMINAE Genus END.IICATOMUS Me11i6 E. rugosus (Randall) MB ON PQ Subfamily DINODERINAE Genus RHYZIOPERTIIA Stephens R. dominica ,Fabriciustt BC MB ON pe pusilla (Fabricius) - Genus PROSTEPHANUS Lesne P. punctatus {Sayr - ON Genus STEPHANOPACIIYS Waterhouse S. cribratus t LeConter NT BC AB SK MB ON pe S. rugosus (Ctlivier) - ON NB opacus (Casey') porcatu! (LeConte) S. sobrinus lr)aseyr - BC AB S. substriatus (Paykull)* AK YK NT BC AB MB ON pe NS pacificu:; (Casey) - - Subfamily BOSTRICHINAE Tribe Bostrichini Genus AMPI IICERUS LeConte (Subgenus AI\.{PHICERUS s. str.) A. bicaudatu,; (Sayr NT SK MB ON asperico,t lis (Germar) gracilis'Jasey serricollt's (Germar) Genus LICHIINOPHANES Lesne L. armiger {Leconte) MB ON L. bicornis (M/ebert - ON pe L. mutchieri Belkin - ON pe angustatus (Casey) L. truncaticollis tl-eConrer - ON Tribe Xyloperthini Genus SCOBICIA Lesne S. bidentata tHorn r - ON S. declivis (LeContet - BC Genus XYLOBIOPS Casey X. basilaris tSayt - ON basalis ( Dejean) h ume ral is (Melsheimer) 200 Anobiidae Family LYCTIDAE lyctid powder'post beetles J. McNarnara Only six species of lyctids are known to occur in cause further damage to wood by producing exit Canada. Members of this family are commonly holes when they emerge. The family is worldwide called powder-post beetles because of the tendency in distribution, each region having an indigenous of the larvae to reduce sapwood, especially of hard- fauna plus introduced species. woods, into a powdery frass. They are found more Gerberg (1957) has revised the New World often in recently dried wood rather than in old species. wood. They attack lumber used for furniture, crat- ing, hardwood floors, and other purposes. Second only to termites, Lyctidae cause the most destruc- BC (4);uB (1); oN (4);PQ (3);NB (1);NS (2) o I tion to wood and its products. The adult beetles (2) Tribe Lyctini Genus LYCTUS Fabricius L. brunneus (Stephens)f BC PQ NB NS carolinae Casey L. cavicollis LeConte BC L. linearis (Goeze)f BC oN PQ arillaris Melsheimer striatus Melsheimer L. opaculus LeConte -oN breuipennis Casey L. planicollis LeConte BC MB ON PQ NS Genus TROGOXYLON LeConte T. parallelopipedum (Melsheimer) ON Family ANOBIIDAE deathwatch beetles J. McNaruara Anobiids include 79 species in Canada and (cigarette beetle), are serious pests of stored prod- Alaska. Larvae of most species bore into dead hard- ucts, including drugs, tobacco, seeds, spices, cereal woods and softwoods; two species of Ernobius attack products, leather, and museum specimens. and destroy pine cones; one species, Anobium punc- North American anobiids are relatively well tq.tum (furniture beetle), damages woodwork, furni- known taxonomically, thanks to the work of White, ture, bookbindings, and similar products. The bee- who published a key to the genera (1971), a number tles often produce a tapping sound in their burrows, of generic revisions, and a catalog of the North which gives rise to the ancient superstition that they American species ( 1982). are a warning of death, hence the common name. Some species feed in the young stems or shoots of AK (4);YK (6);NT (2); BC (35);AB (18); SK (9); MB growing trees. Two species, Stegobium paniceum (21); oN @2);PQ (40);NB (8);NS (7);NF (7) o I (drugstore beetle) and Le.sioderma serricorne (5) Subfamily DRYOPHILINAE Tlibe Eucradini Genus EUCRADA LeOonte E. humeralis (Melsheimer) -oN PQ E. robusta Van Dyke _BC 201 Anobiidae Subfamily ERNOBIINAE Tribe Xestobiini Genus XESiTOBIUM Motschulsky Hyperisus Mulsant & Rey Key to North American species: White (1975o) X. abietis Fisher BC X. alfine LeConte BC X. gaspensrs R.E. White -oN PQ X. marginicolle (LeConter BC Genus UTOBIUM Fall Key to North American species: White (1966, 1976) U. elegans iHorn) BCAB_MBONPQ U. granulal,um R.E. White _ v_ BCAB_PQ U. marmorrtum Fisher BC Tlibe Ernobiini Genus EPISERNUS Thomson E. trapezoideus (Fall) BC Genus ERITIOBIUS Thomson Philoxylon LeConte E. bicolor R.E.White NF E. gentilis Jlall BC E. luteipen.ris LeConte PQ E. mollis (Iinn6)+ ON PQ_NS NF conu et ifrons (Melsheimer) E. nigrans ,lall _BC E. pallitarsis Fall _BC E. punctulatus (LeConte) _BC E. schedli ![.J. Brown ON PQ NF Subfamily ANOBIINAE Tribe Euceratocerini Genus CTENOBIUM LeConte C. antennal;um LeConte oN PQ Tlibe Gastrallini Genus GASTRALLUS Jacquelin du Val G. marginipennis LeConte BC Tlibe Nicobiini Genus TRICHODESMA LeConte Ptinodzs LeConte T. cristata Oasey T. gibbosa (Say) _ _ _:c_ _ _ ON T. klagesi Fall ON TYibe Stegobiini Genus STETIOBIUM Motschulsky S. paniceum (Linn6)t AK BC SK MB ON PQ NB obesurn (Melsheimer) tenuistriatum (Say) Genus OLIGOMERUS Redtenbacher Key to North American species: White (1976) O. alternan; LeConte ON PQ 202 Anobiidae O. crestonensis Hatch BC O. obtusus LeConte lo*pe-- O. sericans (Melsheimer) _ONPQ Tribe Anobiini Genus PLATYBREGMUS Fisher P. canadensis Fisher ON NS Genus ANOBIUM Fabricius A. punctatum (DeGeer)f BC MB_PQ NS pumilum LeConte Genus HEMICOELUS LeConte Cacotemnus LeConte Key to North American species White (1976) H. carinatus (Say) BC SKAB MB ON PQ NB errons (Melsheimer) linearis (LeConte) H. defectus (Fall) _BC yu oN PQ H. gibbicollis (LeConte) AK BC destructor (Fisher) roguensis (Hatch) H. pusillus (Fall) -oNPQ H. umbrosus (Fall) AK YK AB MB ON PQ NB Genus MICROBREGMA Seidlitz M. emarginatum emarginatum BC AB SK MB ON PQ (Duftschmid)t foueatutn (Kirby) M. emarginatum granicolle Fall BC Tribe Hadrobregmini Genus HADROBREGMUS C.G. Thomson Coelostethus LeConte H. americanus (Fall) BC AB H. notatus t Sayt oN PQ NS__- H. quadrulus (Leconte) BC H. truncatus (Fall) BC Genus PRIOBIUM Motschulsky Key to North American species: White (1975b) P. sericeum (Sayt MB ON PQ NB NS Genus DESMATOGASTER Knutson D. subconnata (FaIlt YK AB MB PQ Subfamily PTILININAE Tribe Ptilinini Genus PTILINUS O.F. Mriller P. basalis LeConte BC P. Iobatus Casey YK_BCAB_MBON serricollis Say P. pruinosus Casey -MBONPQ NB P. rufrcornis Say -ABONPQ _NS bicolor Melsheimer Subfamily XYLETININAE Tribe Xyletinini Genus VRILLETTA LeConte Pseudoxyletinus Pic V. decorata Van Dyke BC V. laurentina Fall ON PQ 203 Anobiidae Genus EIIIRILLETTA Fall Oligor;wodes Fall Key to North American species: White (1985) E. brevis R E. White AB -oN E. harrisii lFall) -oNPQ E. peltata (Harris) MB ON PQ Genus XYI ETINUS Latreille Key to N crth American species: White (1973, 1977) X. bicolor E.E. White -YK AB SK MB X. confusus R.E. White NT BC AB PQ X. fucatus i,eConte _BC_SK MB ON PQ NB NF X. lugubris LeConte AB SK yu_:a X. rotundic'rllis R.E. White -BCABSK Tribe Lasiodermini Genus LASIODERMA Stephens Key to North American species: White (1990) L. serricorne (Fabricius)t AK MB ON PQ castanzum Melsheimer Subfamily DORCATOMINAE Tlibe Calymmaderini Genus CALYMMADERUS Solier Eupac.lus LeConte C. nitidus ( LeConte) TYibe Petaliini Genus PETALIUM LeConte Key to N'rrth American species: Ford (1973) P. bistriatum (Say) ON P. seriaturr Fall ON PQ Tribe Prothecini Genus SCULPTOTHECA Schilsky S. puberula tLeConte) ON PQ Genus STAGETUS Wollaston Key to New Worid species: White (19756) S. prolundus (Leconte) ON PQ s triat orc u n ctatzs (LeConte) TYibe Dorcatomini Genus DOIi,CATOMA Herbst Key to North American species: White (1966) D. falli R.E, White -oN D. integra ( Fall) _BC D. moderata R.E. White MB ON PQ D. pallicornis LeConte -SKMBONPQ NB NF foueata R.E. White D. setulosa LeConte oN PQ Genus CAENOCARA C.G. Thomson Tylistus LeOonte C. bicoior (Germar) MB ON C. blanchardi Fall _BC C. oculaturrL (Say) _ONPQ NE NS C. scymnoides LeConte _YKNTBCABSK MB ON PQ C. tenuipalpum Fall _,, 204 Ptinidae Genus BYRRHODES LeConte B. intermedius (Leconte) PQ Subfamily TRICORYNINAE Tribe TYicorynini Genus TRICORYNUS Waterhouse C atorama Gu6rin-M6neville Hemiptychus LeConte Key to North American species: White (1965) T. borealis (LeContet MB ON T. confusus (Fall) PQ T. densus (Fall) AB rogersi (Hatch) T. nigritulus (Lecontet - ON T. nubilus tFallt schuhi (I].atc]e], Family PTINIDAE spider beetles Y. Bousquet Only 16 ptinid species are found in Canada and Ptinidae, the common name of which refers to Alaska, and all but three are introduced on this con- the fact that the adults of many species look like tinent. Many of these species are found in mills, small spiders, is closely related to Anobiidae. The warehouses, granaries, stores, museums, and hospi- species are divided into two groups that are consid- tals where they feed, in both the adult and larval ered to be subfamilies or tribes. Papp has reviewed stages, on a wide variety of products including the North American fauna (1962). Many species grain, seeds, and drugs. These species are usually that occur in Canada can be identifred with the key minor pests, since they rarely build up large popula- provided by Brown (1940). tions. Spider beetles also occur in nests of mam- mals, birds, and bees; they are probably scayengers AK (1); BC (13);AB (5); SK (6); IVIB (7); ON (11); PQ that feed on dry animal and vegetable matter. (11);NB (9);NS (10); PE (1); NF (5) . I (13) Subfamily GIBBIINAE Genus GIBBIUM Scopoli Scotias Czenpinski G. aequinoctiale Boieldieuf - ON PQ NB NS psylloides auct. Genus MEZIUM Curtis M. affrne Boieldieut - BC - SK MB ON PQ NB NS Subfamily PTININAE Genus SPHAERICUS Wollaston S. gibboides tBoieldieut+ - BC Genus TIPNUS C.G. Thomson Epauloecus Mulsant & Rey T. unicolor (Piller & - NB NS Mitterpacher)t Genus TRIGONOGENIUS Solier T. gJobulus Soliert - BC farctus LeConte Genus NIPTUS Boieldieu N. hololeucus (Faldermann)f - BC AB SK MB ON PQ NB NS PE - NF 205 Trogossitida e Genus PSEI.IDEUROSTUS Heyden P. hilleri (Reitter)t BC AB ON PQ NB alienus (W.J. Brown) Genus PTINUS Linn6 (Subgenus I'TINUS s. str.) P. bicinctus Sturmf _BCABSK MB oN PQ NB NS P. clavipes ['anzert _BC ON PQ NS b r unne;t. ts Duftschmid hirtellus Sturm testaceus Olivier P. fur (Linn6)t BC ON PQ NB NS NF humeralis Say lofro Fabricius P. raptor Sturmt BC _SKMBONPQ NB NS NF P. villiger (Reitter)-f BC AB SK MB ON PQ NS NF (Subgenus CYNOPTERUS Mulsant & Rey) P. californicus Pic BC MB_PQ P. fallax Fall BC P. falli Pic -oN P. ocellus W J. Brownf AK BC _SKMBONPQ NS NS _NF tectus aoct. Superfamily Lymexyloidea FamilyLYMD(YLIDAE ship-timberbeetles J.M. Campbell Thirt;'ssvsn species of Lymexylidae are known that Lymexylidae is closely related to Cleroidea and around the world. Only one rare species occurs in Cucujoidea. Arnett (1968) places the farnily near Canada. .\dults are found in decaying wood, on tree Lampyridae, based on wing venation. Wheeler trunks, or under bark. The Canadian species is (1986) revised the world genera and higher classifi- probably .introduced, since it also occurs in Europe cations, including a study of the phylogerny, mor- and was firrmerly very destructive to wooden ships. phology, and known biology of the genera. The classifrcation of the family is now fairly sta- ble. Crorvson (1955) and Wheeler (1986) believe aB (1);ON (1);PQ (1);NB (1); NS (1);NF (1) Subfamily FIYI ECOETINAE Genus IIYLItrCOETUS Latreille Elatero;;.des J.C. Schaeffer Hyloceotus Melsheimer H. Iugubris lJayf AB ON PQ NB NS NF americc:nus }{aris Superfamily Cleroidea Family TROGOSSITIDAE bark-gnawing beetles Y. Bousquet Trogossitidae includes 22 species in Canada and Barron (1971) has revised the trogositids of Alaska. Most North American species are found North America. The species are divided into two under the bark of dead trees, in logs, and in stumps; subfamilies, Peltinae and Trogossitinae. a few species occur in fungi. The adults and larvae of many species feed on wood-eating insects; others feed on dercaying matter or fungi. Tenebroides mau- AK (a);YK (a); NT (5); BC (17); aB (6); SK (6); MB ritanicus is a well-known stored-product pest that (6); oN (12); PQ (e); NB (2); NS (2); PE (1); LB (1); feeds on grains and cereal products. NF(3) . H(2) 206 Trogossitidae Subfamily PELTINAE Tribe Peltini Genus CAI-ITYS C.G. Thomson Nosodes LeOonte Peltidea Motschulsky C. minor Hatch _YK_BCABSK_ONPQ C. scabra (Thunberg)* AK_NTBCAB_MBONPQ sercata (LeConte) silphides (Newman) Genus OSTOMA Laicharting Gaurambe C.G. Thomson O. columbiana Casey AK YK NT BC AB SK MB ON PQ NB LB O. ferruginea (Linn6)* AKYKNTBCABSK_ONPQ NS NF fraterna (Randall) nigrina Casey se ptentr io nalis ( Randall) O. pippingskoeldi (Mannerheim) BC Genus GRYNOCHARIS C.G. Thomson G. oregonensis (Schaeffer) BC G. quadrilineata (Melsheimer) MB ON PQ marg inata (Melsheimer) Genus THYMALUS Latreille T. marginicollis Chevrolat BC AB SK MB ON PQ NS NF aubei L6veill6 fulgidus Erichson Genus ERONYXA Reitter Ostomodes Reitter E. paliida (Motschulsky) BC dohrni (Reitter) lagrioides Reitter pilosula (Crotch) Subfamily TROGOSSITINAE Tribe Trogossitini Genus NEMOSOMA Latreille Pseudalindria Fall N. punctulatum Van Dyke BC punctatum Van Dyke Genus CORTICOTOMUS Sharp Parafilumis Casey C. caviceps (Fall) BC laeuiuentris Casey Genus AIRORA Reitter A. aequalis Reitter BC bicolor Casey polita Casey punctiuentris Casey A. cylindrica (Audinet-Serville) ON nlgello (Melsheimer) nigra (Melsheimer) teres (Melsheimer) Genus TEMNOCHILA Westwood T. chlorodia (Mannerheim) NT BC melanica lJ.atc}a prosternalis Schaeffer u ir idicy anea (Mannerheim) 207 Cleridae Genus TEI'IEBROIDES Piller & Mitterpacher Trogossita Olivier T. americanus tKirby) ON PQ castat teus (Melsheimer) (Horrr) nigrit ^rs T. collaris Sturm) -- T. corticalis (Melsheimer) AK YK NT ec- sxusBNpe - - - - dubius (Melsheimer) internrcdius (.Horn) limbalis (Melsheimer) T. crassico'nis (Horn) BC califo.ni"u" 11fornt pleurulis (.Horn) T. laticollir, (Horn) -oN obscu:us (Hotn) T. mauritanicus t Linn6rf BC SK MB ON PQ NB PE NF nitidt s (Horn) T. occidenlalis Fall BC T. sinuatu,r (LeConte) BC Family CLERIDAE checkered beetles J. McNarnara Cleridae includes 50 species and subspecies in In North America the taxonomy of clerids is rea- Canada and Alaska. Most species are predaceous, sonably well known, but the literature is scattered. the adul':s feeding on adult bark beetles and the lar- A revision of the family would be helpful. Barr vae on eggs, larvae, and adults of wood-boring bee- (I962a) has published a key to the genera. There is tles in galleries. Several species of Enoclerus and no comprehensive work for the Canadian fauna, but Thanasimus are believed to be imoortant in the most eastern species could be identifred by referring natural r:ontrol ofbark beetles in coniferous forests. to Knull (1951) and most western species by refer- Adults of Cleridae are found mostly on plants, tree ring to Barr 0962b). trunks, or under bark. Some are found on flowers, feeding on pollen. Some species of Necrobia are AK (5); vK (4); NT (5); BC (22); AB (12); SK (10); scavengers and are economically important as pests MB (14); ON (32); PQ Q4; NB (11); NS [4); NF (4) of stored food, carcasses, and skins. . I(5) Subfamily CLERINAE Tlibe Tillini Genus MONOPHYLLA Spinola Elasn ocerus LeConte Macrttelus lllug M. terminata (Say) ON megatoma Spinola Genus CYIyIATODERA Gray C. bicolor (Say) oN PQ C. decipiens Fall BC _-n'___ C. inornatiL (Say) ON PQ tenera LeConLe C. ovipennis LeConte BC pilosei.Ia LeConte C. undulata (Say) ON boscil (Chevrolat) longicollis Spinola uauth.ieri Spinola Tribe Phyllobaenini Genus PH'{LLOBAENUS Deiean Hydnocera Newman 208 Cleridae P. dubius (Wolcott) P. humeralis (Say) - - - ecABSfMBoNFeNs- - - - cyanescens (LeConte) cydneus (Spinola) dfficilis (LeConte) P. lecontei (Wolcott) P. pallipennis (Say) T- I'BC-:"MBBNiSNBws- - - serratus (Newman) P. scaber (LeConte) _BC P. subfasciatus (LeConte) _BCABSKMB P. unifasciatus (Say) -oN punctatus (Spinola) P. verticalis (Say) -oNPQNB brachypterus (I0.ug) I ine a toc oll is ( Spinola) tenellus (.Say') Genus ISOHYDNOCERA E.A. Chapin I. curtipennis (Newman) MB ON PQ NB NS longicollis (Ziegler) L tabida (LeConte) _ONPQ Tribe Thaneroclerini Genus ZENODOSUS Wolcott Z. sanguineus (Say) -ONPQ Tribe Clerini Genus PRIOCERA Kirby P. castanea (Newman) - ON maculata Ziegler rufescens Spinoia Genus OPILO Latreille O. domesticus (Sturmt* Genus THANASIMUS Latreille T. dubius (Fabricius) AK YK NT BC AB SK MB ON PQ NB - NF erythrocep halus (Spinola) ruficeps (Spinola) T. trilasciatus (Say) - ON PQ T. undatulus (Say) AK \T( NT BC AB SK MB ON PQ NB NS NF abdominalis I{rtby mel anocep halus Chevrolat nubilus (l\lug) Genus PLACOPTERUS Wolcott P oeci loc hroa Chevrolat P. thoracicus tOliviert MB ON PQ monilis (Melsheimer) pallipes (Wolcott) Genus ENOCLIIRUS Gahan E. barri Knull - BC E. coccineus coccineus (Schenkling) AB SK corallinus (Fall\ E. eximius (Mannerheim) - BC holosericeus (A. White) E. ichneumoneus (Fabricius.t - ON rufus (Olivier) E. Iaetus tKlugt - BC abruptus (LeConte) E. lecontei (Wolcott) - BC nigriuentris (LeConte) E. muttkowskii (Wolcott) - ON PQ NB liljebladi Wolcott 209 Cleridae E. nigrifrons tSayt * ON PQ gerhardi Wolcott testaceipes Wolcott E. nigripes nigripes {Say) MB ON PQ NB dublzs (Spinola) incertus (LeConte) E. nigripes rufiventris (Spinola) - ON PQ NB E. rosmarus (Say) - ON oculatus (Spinola) E. schaefferi Barr - BC AB humer,zlis Schaeffer E. sphegeus (Fabriciust - BC AB sobrius (Walker) Genus TRIOHODES Herbst Key to Norbh American species: Foster (1976) T. apivorus borealis Wolcott & E.A. Chapin interruptus LeConte nuttalii Say trifasaatus Strnn T. nutalli tliirby) AB SK MB ON PQ T. ornatus lrartwegianus A. White AK YK NT BC AB Subfamilv KORYNETINAE Tribi Epiphloeini Genus PHI OGISTOSTERNUS Wolcott Phyllo nenus Spinola P. dislocatus {Sayt MB ON PQ NB distrop.hus (IArg) transuersalis (Spinola) Genus NEI(IHNEAWolcott & E.A. Chapin N. laticorni ; (Sayt - ON Tribe Enopliini Genus NECTRTHOPLEURA Barr Key to sprecies: Barr (1976) N. thoracicr'L ( Say r - ON PQ b imac t.iata (Melsheimer) nig ript: n ni s ( Spinola) p ennsy lu anica (Chevrolat) p uncto tis s im a (Chevrolat) Genus CHARIESSA Perty B r achy mor p hus Chevrolat C. elegans Horn - BC lemberti La Rivers C. pilosa (F)rster) - BC AB SK MB ON PQ cinclo lSpinola) margirLata (Say) onusta (Say) Genus BOSCHELLA Barr B. fasciata tl-eConte) - BC trilobata (Van Dyke) Genus CREGYA LeConte Galeruclerus Gahan C. oculata (Say) - ON marginipennis ( Spinola) 2to Melyridae Tribe Korynetini Genus NECROBIA Olivier N. ruficoilis (Fabricius)t -BC-oNPQ N. rufrpes (DeGeer)t _BCABONPQNF N. violacea (Linn6)t AK YK NT BC AB SK MB ON PQ NF errons Melsheimer Subfamily TARSOSTENINAE Genus TARSOSTENUS Spinola T. univittatus (Rossi)f BC alb ofa sc iat us ( Melsheimer) Family MELYRIDAE soft-winged flower beetles D.E. Bright Melyridae includes 49 species known to occur in careful study. There are no recent revisions or taxo- Canada. The adults commonly occur on flowers and nomic treatments of the Nearctic species; the infor- evidently feed on other insects as well as on pollen. mation below therefore reflects our best effort at Larvae are primarily predators, occurring in vari- achieving a list based on available literature. ous habitats but most commonly in soil, leaf litter, Arnett (1968) provides a key to the genera and or under bark. One species, Malachius deneus,has Hatch (1962) a key to the species of the Pacific been introduced in North America and causes con- Northwest. siderable damage to developing wheat in the Prairie Provinces. The taxonomy of the North American melyrids YK (2); NT (3); BC (35);aB (16); SK (5); MB (6); ON is in poor condition, and the family is in need of (10);PQ (10);NB (1);NS (1);NF (1) ' I(2) Subfamily MALACHIINAE Genus COLLOPS Erichson C. bipunctatus (Say) AB SK C. cribrosus LeConte BC cy anip en nis Motschulsky C. discretus Fall BC C. hirtellus LeConte C. quadrimaculatus (Fabricius) __ftBCy:"_ oNpe----- C. tricolor (Say) SK MB oN PQ C. versatilis FalI Bc C. vittatus (Say) - - lrr AB ir nrs ON PQ hauorthi Westwood Genus ENDEODES LeConte Key to North American species Moore & Legner (1975b) E. collaris LeConte BC Genus MALACHIUS Fabricius Anthocomus Erichson Hapalorhinus LeConte Microlipus LeConte M. aeneus (Linn6)l -BCAB ON PQ NB NS NF M. auritus LeConte _BC M. bipunctatus (Harrer)t ON PQ M. blaisdelli Hopping -Bcms M. erichsonii (LeConte) criddlei W.J. Brown lateralis LeConte M. flavilabrus (Say) ON PQ coeruleus Randall 2tl Melyridae M. floricola Marshall - BC M. francisc,rnus Fall - BC M. horni FrLll - BC AB antenruatus Hopping M. macer fl.orn - BC M. mixtus l{orn - BC M. mirandrrs (LeConte) - BC M. moerens. LeConte - BC u n ifor in is Motschulsky M. nigrinui Fall - BC M. thevene,ii Horn - BC AB contorl.us Fall M. ulkei H<'rn AB - MB ON PQ Genus TAN'AOPS LeConte C ap halistes Motschulsky T. ignitus Fall BC bosalis W.J. Brown blaisdelli (Malkin) n u ne n;n ac her i Marshall sierroe Marshall T. malkini l{arshall - BC Genus HYF'EBAXUS Kiesenwetter Pseude'baeus Horn H. apicalis Sayt oblitus (.LeConte) H. bicolor (',eContet - BC AB Genus ATTAI-US Erichson (Subgenus .\CLETUS LeConte) A. nigrellus (LeConte) - YK - BC AB ON PQ falli (I:lopping) (Subgenus ,\TTALUS s. str) ( S calolt ter u s Motschulsky) A. morulus smithi HoppinC - BC A. pallifrons tMotschulskyt - ON A. humeral .s LeConte pettiti Hotn Subfamilv RHADALINAE Key to geriera: Peacock (1987) Genus SEM.IJULISTUS Schilskv Eurelymis Casey S. ater ( Le('ontel - BC AB Subfamily DASYTINAE Key to North American genera: Blaisdell (1938o) Genus DOL,ICHOSOMA Stephens D. foveicollc tKirbyt - BC AB - MB ON PQ Genus DASYTES Paykull D. cruralis LeConte) - BC Genus AMPHIVECTURA Blaisdell A. monticol r tBlaisdellr - BC AB Genus HOF PINGIANA Blaisdell H. hudsonir:a (LeConte) - \'K NT BC AB SK MB - PQ breuilabris Blaisdell kingiW.J.Brown 212 Sphindidae Genus TRICHOCHROUS Motschulsky T. alberbensis Blaisdell AB Genus AMECOCERUS Solier Listrus Motschulsky A. confusus Casey _BC A. glabratus Hatch _BC A. medicatus Blaisdell AB A. olSrmpianus Blaisdell _BC A. plenus Casey _BC A. provincialis Blaisdell -BC A. quadricollis Hatch _BC A. robustus Blaisdell _BC A. senilis (LeConte) AB Genus DASYRHADUS FalI D. impressicollis Fall BC Superfamily Cucuj oidea FamilySPHINDIDAE dry-fungusbeetles J.M. Campbell Sphindidae is a small family comprising 35 (1979) revised the higher classification of the fam- species around the world, of which five are known to ily, which is in need of a taxonomic revision; most occur in Canada. Adults and larvae feed on spores current records are unreliable. of slime molds. Casey (1898) prepared the last revision of the North American sphindids. Sen Gupta and Crowson BC (2);AB (1); oN (3);PQ (4) Subfamily SPHINDINAE Tribe Odontosphindini Genus ODONTOSPHINDUS LeConte O. clavicornis Casey BC O. denticollis LeConte ON pc) Tribe Sphindini Genus SPHINDUS Chewolat Coniophagus Mink S. americanus LeConte -BCABONPQ S. trinifer Casey _ONPQ Subfamily EURYSPHINDINAE Genus EURYSPHINDUS LeConte E. hirtus LeConte PQ 213 Nitidulidaer Family NITIDULIDAE sap beetles J. McNamara The lamily Nitidulidae comprises 101 species in the pollen and petals of the same plantli or some- Canada and Alaska. Members of this family are times of others. mostly saprophagous and mycetophagous. The Parsons (1943) published a revision of the majority live in decaying fruit, fermenting plant Nearctic Nitidulidae, and most western species can juice, an Subfamily CATERETINAE Genus HEI'ERHELUS Jacquelin du Val H. pennatus (Murray) BC ON PQ NF crinitt..s (Murray) H. sericans (LeConte) BC Genus CAI'ERETES Herbst (Subgenus CATERETES s. str.) C. scissus Parsons AK_NT AB SK ON PQ Genus BOIiEADES Parsons B. abdomiralis (Erichson) MB ON PQ Genus BRI'CHYPTERUS Kugelann B. globularius Murray _BCAB B. troglodytes Murray _BC B. urticae (Fabricius)f NT BC AB SK MB ON PQ NF pusill Lrs (Melsheimer) Genus BRACHYPTEROLUS Grouvelle B. pulicarius (Linn6)t AB SK MB ON PQ NB NS NF mordelloides Notman Subfamily CARPOPHILINAE Genus COI,IOTELUS Erichson C. obscuru,r Erichson MB ON PQ NB NS Genus COLOPTERUS Erichson Colastus Erichson CYlloP'liss MurraY Key to North American species: Parry & Howden (1975) C. maculatus (Erichson) C. semitect.us (Sayt - - - - 3Npe- C. truncatus (Randall) YK BC AB SK MB ON PQ NS infimus (Erichson) limba,tus (LeConte) obliquus (LeConte) Genus CAIIPOPHILUS Stephens (Subgenus CARPOPHILUS s. str.) (Tribrachys LeConte) C. antiquus Melsheimer PQ C. brachypterus (Say) mn on PQ carbonatus LeConte 214 Nitidulidae C. dimidiatus (Fabricius)* MB oN PQ C. discoideus LeConte BC ON apicalis LeConte caudalis (LeConte) C. hemipterus (Linn6)f AK BC AB ON PQ b imac ulatu s Melsheimer C. ligneus Murray AK decipiens l{orn AB C. lugubris Murray -oN pe C. marginellus Motschulskyf MB ON nitens Fall C. mutilatus Erichsonf BC luridus Murray C. pallipennis (Say) AB SK pollens Sharp C. sayi Parsons MB PQ Subfamily NITIDULINAE Genus EPURA-EA Erichson Epuraeanella Ctotch E. adumbrata Mannerheim AK-NTBCAB PQ E. aestiva (Linn6)t AK BC AB ms oN PQ NF conu exi u sc ula Mannerheim E. alternans Grouvelle _BC alticola Fall E. alternata Parsons -oNPQ E. ambigua Mannerheim AK_- BC E. avara (Randall) AK YK BC MB ON PQ NB NS NF nubila LeCotte E. boreades Parsons BC _ONPQ NB E. corticina Erichson PQ E. erichsoni Reitter NT BC-SK MB ON PQ E. flavomaculata Miiklin AK _AB MB ON PQ NS E. fulvescens Horn -oNPQ E. helvola Erichson SK MB ON PQ castanea (Melsheimer) rufa Reitter E. horni Crotch ON PQ duryiBlatchley E. labilis Erichson BC MB ON PQ NS__ E. linearis Maklin AK BC AB PQ E. obliquus Hatch _I' BC AB PQ NB E. obtusicollis Reitter BC AB SK MB_PQ E. ovata Horn BC MB_PQ E. parsonsi Connell BC AB SK PQ Ns__ NF E. peltoides Horn I"_ _AB MB ON PQ E. planulata Erichson AX BC AB MB ON PQ NB placida Meklin E. populi Dodge BC AB MB_PQ E. rufa (Say) _ONPQ bodio (Melsheimer) rotundicollis Reitter E. rufida (Melsheimer) _ONPQ E. terminalis Mannerheim* AK YK _BC AB MB ON PQ NB infuscata M€iklin E. truncatella Mannerheim AK YK NT BC AB SK MB ON PQ LB NF nigraMeklin ornatula Notman E. umbrosa Horn PQ NB Genus APHENOLIA Reitter A. monogama (Crotch) BC Genus STELIDOTA Erichson S. octomaculata (Say) ON PQ biseriata Reitter 2t5 Nitidulidae Genus OMOSITA Erichson (Subgenus IJAPROBIA Ganglbauer) O. colon (Linn6)t - BC AB SK MB ON pe NB NS NF O. discoidezL (Fabricius){ AK BC AB - MB ON pe - NS NF inuersc: LeConte Genus NITIDULA Fabricius N. bipunctata (Linn6)t AK YK - BC AB - MB ON pe N. carnaria (Schaller)i BC - ON pe - NS N. nigra Sc.iraeffer AK yK - BC AB - MB N. rufipes (,inn6)t - BC AB - MB ON pe NB NS NF ossium l{ttby N. ziczac Say NT BC AB - MB humerr:tlis LeConte inorna;a Horn unig u t i:a t a Melsheimer Genus PRO METOPIA Erichson P. sexmaculata tSayr ON pe Genus LOB [OPA Erichson Cerophorus Castelnau L. setosa Hi,rold BC substriata (Hamilton) L. undulata tSayr AB - MB ON pO Genus SOR')NIA Erichson S. grisea (Linn6rt NF S. guttulata tl-eConter - BC Genus PHE.\OLIA Erichson P. grossa tF.rbricius) ON pO Genus THA-YCRA Erichst-rn Perthaiycra Horn Key to New World species: Howden (1961) T. concolor (LeConte) - YK - BC AB - MB ON pe T. dentata I l. Howden - BC T. keltoni H Howden - BC AB T. leechi H. Howden BC T. mixta H. Howden BC AB T. monticola H. Howden - BC T. murraf lHornt - BC AB T. parsonsi .r{. Howden - BC T. sinuata H. Howden - BC AB Genus POC,,\DIUS Erichson P. fulvipenn s Erichson BC dorsali:; Horn P. helvolus llrichson NT BC - SK MB breuiusculus Reitter infusca;',us Reitter limbatt.s Reitter Genus AMP.] ICROSSUS Erichson A. ciliatus t()liviert - ON unilineatus (.Say) Genus CYCIIRAMUS Kugelann Quadrifrons Blatchley C. adustus !.richson * ON pO Genus PALL,ODES Erichson P. pallidus tl)alisot de Beauvois) - ON silaceus Erichson 216 Nitidulidae Genus CYLLODES Erichson C. biplagiatus LeConte MB Subfamily MELIGETHINAE Genus MELIGETHES Stephens Key to North American species: Easton (1955) M. atratus (Olivier)l _BC M. canadensis Easton AKYK_BCAB_MBONPQ M. cleominis Easton _BCABSKMB M. dauricus Motschulsky* _BCABSKMB californicus Reitter moerens LeConte rufimanus LeConte M. nigrescens Stephens* _YK_BCAB_MBONPQ_NSPE seminulum LeOonte M. pinguis Horn M. saevus LeConte AB-nrn------NF M. simplipes Easton _ONPQ_NSPE Subfamily CRYPTARCHINAE Genus PITYOPHAGUS Shuckard P. rufipennis Horn - BC Genus CRYPTARCHA Shuckard (Subgenus CR\?TARCHA s. str.) C. ampla Erichson - BC AB - MB ON PQ (Subgenus LEPIARCHA Sharp) C. concinna Melsheimer - BC AB - MB ON PQ bella Reitter liturata LeConte piclo Melsheimer Genus GLISCHROCHILUS Reitter (Subgenus LIBRODOR Reitter) G. fasciatus (Olivier) - BC MB ON PQ - NS g eminat us (Melsheimer) q uadr imaculoso (Melsheimer) G. quadrisignatus (Say) - BC AB - MB ON PQ - NS NF b ip unctatu s (Melsheimer) bipustulatus (Melsheimer) canadensis W.J. Brown s exp u st ulatu s (Reitter) sirnilis (Melsheimer) G. sanguinolentus rubromaculatus PQ NB NS (Reitter) G. sanguinolentus sanguinolentus - BC AB ON PQ (Olivier) G. siepmanni W.J. Brown - BC AB SK MB ON PQ NB (Subgenus GLISCHROCHILUS s. str.) G. confluentus tsayt - ON PQ G. lecontei W.J. Brown - BC IVIB G. moratus W.J. Brown - BC AB SK MB ON PQ G. vittatus (Say) AK BC AB - MB ON PQ dejeanii (Kirby) sepulcralis (Randall) Subfamily CYB OCEPHALINAE Genus CYBOCEPHALUS Erichson C. californicus Horn _BC 217 Rhizophagi lae Family RHIZOPIIAGIDAE rhizophagid beetles Y. Bousquet Twerrty-four species and subspecies of rhi- The taxonomy of the Nearctic species has been zophagids are known to occur in Canada and Alaska. poorly studied, and most genera are in need of mod- They are collected most commonly under the bark of ern revision. Crowson (1955) recognized four sub- dead trees, in rotten logs, and in decaying vegetation. families, two of which (Rhizophaginae and A few species live in ant nests, and some species of Monotominae) are found within the borders of Monotomit are occasionally found in buildings. Very Canada and Alaska. Sen Gupta (1988) recently little is known about their feeding habits. Members reviewed the world genera of the family. of Rhizophaginae are mostly predaceous, feeding on wood-boring beetles, especially scolytids; those of AK (3); vK (1); sC (15);AB (7); SK (3); MB (a); ON Monotominae are apparently fungus feeders. (15);PQ (11);NB (a);NS (3);NF (5) . I (3) Subfamilv RHIZOPHAGINAE Genus RHIZOPHAGUS Herbst Key to North American species: Bousquet (1990) (Subgenus ;\NOMOPHAGUS Reitter) R. brunneuri brunneus Horn AB ON PQ R. galbus Bousquet _BC v_ R. pseudobrunneus Bousquet AKBCAB_MB (Subgenus ITURHIZOPHAGUS M6quignon) R. grouvellei M6quignon BC (Subgenus RHIZOPHAGUS s. str.) R. dimidiatus Mannerheirn AKYK_BCABON PQ NS NF testaceits M6quignon R. minutus minutus Mannerheim BC R. minutus rotundicollis Bousquet -oN pe NF R. paralleloi:ollis Gyllenhalf _BC_ON PQ NS NF robustt,ts Schaeffer R. remotus .LeConte AK BC AB MB ON PQ NS luteus ,VI6qrignon R. sayi Sche,effer -oN bipunc,tatus (Say) R. sculpturu.tus Mannerheim BC Subfamily MONOTOMINAE Genus PYCI\OTOMINA Casey P. cavicollis (Horn) ON PQ Genus MACREUROPS Casey M. longicoll:s (Horn) BC Genus BAC'IRIDIUM LeConte Crine Pascoe B. ephippigerum (Gu6rin-M6neville) BC -oN B. striolaturn (Reitter) _ONPQ Genus HES PEROBAXNUS LeConte H. abbrevia':us (Motschulsky) BC rufipennis (LeConte) Genus MOITIOTOMA Herbst M. americana Aub6 ON PQ M. arida Casey ON M. fulvipes .Melsheimer --SCAB-nfe opaca iJiegler p ar allelocollis Motschulsky subnigra Motschulsky 218 Cucujidae M. longicollis (Gyllenhal)f _BCABSK _ONPQ NB NF M. parallela LeConte _BC_SK _ONPQ M. picipes Herbstt _BCAB MB ON PQ NB NF foueata LeConte M. producta LeConte NB M. quadraria Casey SK ON Family CUCUJIDAE flat bark beetles Y. Bousquet The cucujids currently include 31 species and sub- Lawrence's concept (1982a) of the family is fol- species in Canada and Alaska. The adults are usu- lowed here. Many authors rank the silvanids as a ally found under the bark ofdead trees, in logs, and in distinct family. There is no modern taxonomic revi- decaying plant matter. Some species apparently feed sion of Cucujidae, but some genera have recently on fungi and decaying matter, others prey on insects. been revised. Six of the species occurring in Canada, Cryptolestes ferrugineus, C. pusillus, C. turcicus, Ahasuerus aduena, Oryzaephilus surinqmensls, and O. mercator, AK (3);vK (1);NT (2); BC (15);AB (10); SK (9); MB are associated with stored products; four ofthem are (17); oN (26); PQ (24); NB (5); NS (a); PE (1); LB among the most serious pests in the country. (1);NF (5) . H (1);I (10) Subfamily CUCUJINAE Genus CUCUJUS Fabricius C. clavipes clavipes Fabricius _MB oN PQ _NS NF C. clavipes puniceus Mannerheim AK BC AB SK MB subnitens Schaeffer Genus PEDIACUS Shuckard P. depressus (Herbst)f _BC subglaber LeConte P. fuscus Erichsonx AK YK NT BC AB SK MB ON PQ NB LB planus LeConte s ub car inatu s Mannerheim Subfamily PASSANDRINAE Genus CATOGENUS Westwood C. rufus (Fabricius) oN PQ Subfamily LAEMOPHLOEINAE Genus PLACONOTUS Macleay Siluanophloeus Sharp Key to New World species: Thomas (1984) P. zimmermanni (LeConte) MB ON PQ apertus (Casey) illustris (Casey) sobrinus (Casey) Genus LAXMOPHLOEUS Deiean L. biguttatus (Say) - BC - SK MB ON PQ NB NS PE b isig nat u s Gu6rin-M6neville L. fasciatus Melsheimer MB ON PQ Genus CHARAPHLOEUS Casey C. adustus (LeContet MB oN PQ -NS C. convexul us (LeConte t ON 2r9 Cucujidae Genus CRY FIOLESTES Ganglbauer Key to New World species: Thomas (1988) C. ferrugine,us (Stephens)t - BC AB SK MB ON PQ NF C. punctatus (Lecontet - ON adumbratus Casey extrica:lus (Casey) geminctus (LeConte) C. pusillus Sch Genus LEPTOPHLOEUS Casey L. alternans (Erichsontf - BC PQ Subfamilv SILVANINAE Tribl Uleiotini Genus ULEIOTA Latreille Bronte:i Fabricius U. debilis rl,econter MB ON PQ U. dubia duoia tFabriciust - ON PQ denticulata Srrrith U. dubia trr ncata Motschulsky BC Genus DENDROPHAGUS Schtinherr D. cygnaei ltlannerheim AK BC AB - MB ON PQ NF americ:t nus Mannerherm germat i Mannerheim glaber LeConLe Genus TEL.IIPHANUS Erichson H et erotlr om ia Haldeman T. velox Haldeman - ON PQ Tlibe Silvanini Genus SILVANUS Latreille Key to spr:cies: Halstead (1975) S. bidentatus (Fabricius)t - BC - ON PQ NB S. muticus Siharp - BC - ON PQ S. nitidulus LeConLe AB - MB S. planatus Germar AB SK MB ON PQ cognatt.ts LeConte zimmer ma nni Gu6rin-M6neville Genus CAT.I:[ARTOSILVANUS Grouve]le KeY to sp,:cies: Halstead (1975) C. imbellis (Leconte) NT ON PQ C. fagi tGuerin-M6nevillet* - ON Genus AHASVERUS des Gozis A. advena (\Valtltf - BC AB SK MB ON PQ musaeorum (Ziegler) A. longulus'Blatchleyt - ON PQ NB paruiceos (Casey) Genus ORYZAXPHILUS Ganglbauer Key to species: Halstead (1980) O. mercator (Fauvel)t - BC AB SK MB ON PQ NB NS NF O. surinamensis (Linnett - BC AB SK MB ON PQ Genus NAU SIBIUS Redtenbacher N. clavicorn.s (Kugelanntt - ON PQ 220 Cryptophagidae Family CRYPTOPIIAGIDAE silken fungus beetles Y. Bousquet Sixty-six species of cryptophagids are recorded fauna, only the genus Cryptophagus has been in Canada and Alaska. These beetles are usually revised (Woodroffe and Coombs 1961). Many gen- found on mold, on fungi, under bark, as well as in era previously included within the cryptophagids decaying vegetation and nests of social have been transferred recently to Languriidae. The Hymenoptera, birds, and mammals; one species, reader can expect many nomenclatural changes at Myrmedophila americana, lives in ant nests. the specifrc level, as well as many new distribution Adults and larvae of most species feed on mold and records, when the cryptophagid groups are revised. decaying vegetation; those of Telmophilus species The generic classification of the Cryptophaginae apparently feed on flower heads of certain aquatic used here follows Bousquet (1989). plants. Several species of Cryptophagus are found in stored products, where they feed on mold. AK (26); YK (1); NT (3); BC (47); AB (15); SK (7); This group is poorly studied taxonomically. MB (12); oN (1e); PQ (17); NB (2); NS (3); LB (2); Since Casey's (1900) revision of the North American NF (3) . H (11);I (8) Subfamily TELMATOPHILINAE Genus TELMATOPHILUS Heer T. americanus LeConte BC AB SK MB ON PQ Subfamily CRYPTOPHAGINAE Key to North American genera: Bousquet (1989) Tlibe Cryptophagini Genus ANTHEROPHAGUS Latreille A. convexulus LeConte PQ NB NS A. ochraceus Melsheimer - - - ec AB !r< r,re 3N PQNBNSPE_NF oregon us Casey A. pallidivestis Casey _BC A. suturalis Meklin AK BC Genus HENOTIDERUS Reitter Crosimus Casey Henoticoides Hatch H. lorna (Hatch) AK BC H. obesulus (Casey) AK_NT AB ON PQ NS Genus HENOTICUS C.G. Thomson Coniophthalmo Kangas Glyptophorus Patk H. californicus (Mannerheim) BC inconspicuus (LeConte) H. serratus (Gyllenhal) AK BC MB ON PQ denticulatus (LeConte) Genus PTERYNGIUM Reitter P. crenatum (Gyilenhal)* BC PQ malacum Casey Genus SALEBIUS Casey S. octodentatus (Mriklin) AK BC sexdentatus Casey Genus CRYPTOPHAGUS Herbst C r y pto p hag i stes Crotch Key to North American species: Woodroffe and Coombs (1961) C. acutangulus (Gyllenhal)* AK BC AB MB ON PQ NF prominens Casey restrictus Casey 227 Cryptophag idae C. bidentatus Maklin AK BC puncta tis sim us MEikIin C. cellaris (llcopoli)+ BC AB SK MB ON angustztus Casey rupimc,ntis Casey C. confertus Casey* AK - NT BC AB - PQ arc han. gelicus J.R. Sahlberg C. corticinus C.G. Thomson* AK BC - ON PQ depressulus Casey C. dentatus (Herbst)* AK BC fumatus (Marsham) q uad.ride ntat us Mannerheim C. difficilis tlasey - BC AB - PQ contractus Casey C. distinguendus Sturm* - BC heeni Casey C. hebes Casey AK BC lyraticollis Casey o/ioszs Casey C. histricus Casey - BC LB C. lapponicLrs Gyllenhalx - BC ualidus Kraatz C. laticollis Lucast - BC AB af,finic Sfrrm cribricollis Casey inscitu.s Casey C. maximus Blake - BC C. obsoletur Reittert - ON C. peregrin rs Woodroffe & Coombs - BC C. pilosus C yllenhalf - BC - - MB ON cicatrir.osus Casey infuscatus Casey paruinodus Casey plectrum Casey C. porrectur' Casey AK BC C. quadrihamatus Mtiklinx AK plagiatus Poppius C. saginatu; Sturml AK BC - ON C. scanicus tlinn6)t - NF C. scutellat rs Newmant - ON C. setulosus, Sturm* - BC - ON PQ - LB antennatus Casey grossutus Casey C. stromus Woodroffe & Coombs - SK MB C. subluma us Kraatzl - BC AB SK - ON PQ C. tubercult,sus Meklinx AK BC AB bouldetanus Casey breuipilis Casey debilis LeConLe instabtlis Bnrce lepidut Casey C. varus Woodroffe & Coombs - BC AB SK Genus MYF,MEDOPHILA Bousquet M. americana (LeConte) AK l'K - BC AB - MB - PQ blzmi iBlaisdell) Tribe Caenoscelini Genus CAENOSCELIS C.G. Thomson Macrodea Casey C. anlenna]is (Caseyt AB - MB C. basalis C asey PQ C. ferruginea tC.R. Sahlbergt* AK BC C. parallela Casey AK BC 222 Languriidae Subfamily ATOMARIINAE Tribe Atomariini Genus EPHISTEMUS Stephens (Subgenus EPHISTEMUS s. str.) E. globulus (Paykull)* BC ON apicalis LeOonte Genus ATOMARIA Stephens (Subgenus ANCHICERA C.G. Thomson) A. aleutica Casey A. apicalis Erichsonf 1"_ _ _ _ _MB A. curtula Casey ON pumilio Casey pl:) A. ephippiata Zimmermann NT BC AB SK MB ON hesperica Casey A. fulvipennis Mannerheim AK A. kamtschatica Motschulsky* AK BC fallax Casey A. laetuia LeConte BC A. nubipennis Casey BC A. ochracea Zimmermann - - lr,fe ON Pe lacustris Casey A. ovalis Casey A. pusilla (Paykull)* AK sc - -- A. saginata Casey - - - a*:a - - - (Subgenus ATOMARIA s. str.) (Agathengis des Gozis) A. capitata (Casey) BC A. fuscicollis Mannerheim* AK A. lepidula Maklin AK A. longipennis (Casey) BC nigricollis (Casey) A. melanica Hatch BC A. planulata Maklin AK BC A. quadricollis (Casey) BC A. stricticollis (Casey) BC A. subdentata (Casey) BC A. vespertina Miiklin AK BC Subfamily IfYPOC OPRINAE Genus HYPOCOPRUS Motschulsky H. tenuis Casey AB Family LANGURIIDAE lizard beetles J.M. Campbell The family Languriidae, as recently defined, is larvae and adults of the other subfamilies are widely distributed in all faunal regions but is best poorly known, but they probably feed on decaying represented in tropical areas. OnIy seven species vegetation and fungal hyphae rather than on living are known to occur in Canada, all but one of which plants (Lawrence and Vaurie 1983). are found east of the Rocky Mountains. Larvae of Until recent papers by Sen Gupta and Crowson, the subfamily Languriinae (Languriidae of previous this family was restricted to those genera now authors) are stem borers of which one, the clover placed in Languriinae. Sen Gupta (1967) trans- stern borer, Languria mozardi, is an economic pest ferred Toramus and allied genera from the family in Canada. The feeding habits and biology of the Cryptophagidae to the subfamily Toraminae. 223 Languriidze Subsequi:ntly (1968), he transferred Loberus to the of the family are reasonably well known, with most tribe Lolrerini, both in the family Languriidae. The subfamilies having been revised since 19,[8 and the family vl,as redefined, and the higher classification species cataloged by Lawrence and Vaurie (1983). and genr:ra of the world were revised bv Sen Gupta and Crorvson (1921). The North Ameiican BC (1); SK (1); MB (2); ON (6); PQ (5) . I (1) "o".i"" Subfamily XENOSCELINAE Tribe Loberini Genus LOI}ERUS LeConte L. impressr.rs LeConte ON PQ Subfamily TORAMINAE Key to species: Sen Gupta (1967) Tribe Toramini Genus TOF:AMUS Grouvelle Tomarus LeConte T. pulchellr.Ls (LeConte) oN PQ Subfamily CRYPTOPHILINAE Tribe Cryptophilini Genus CR'IIPTOPHILUS Reitter C. integer ( F{eer)f PQON Subfamily LANGURIINAE Tlibe Languriini Key to North American species: Vaurie (1948) Genus LAI\ GURIA Latreille L. angustata (Paiisot de Beauvois) ON pulchr t LeConte uhlerit Horn L. convexicollis Horn BC in t er s t,t}ialis C asey L. mozardi mozardi Latreille MB ON PQ apicalis Motschulsky Genus ACROPTEROXYS Gorham A. gracilis (Newman) SK MB ON PQ aztecana Casey diuisa (}]orn) inornata (Randall) latreillei (LeConte) nigrice o s (Motschulsky) obscuro (Motschulsky) pertenuis Casey texana Schaeffer thoracina Casey 224 Erotylidae Family EROTYLIDAE pleasing fungus beetles J.M. Campbell The family Erotylidae is found in all faunal The family is well known taxonomically; Boyle regions of the world but primarily in moist, tropical (1956) revised the North American species, and Sen areas. Only 20 species are known to occur in Gupta (1969) worked out the higher classifrcation. Canada. Adults are frequently found under bark or under logs. Both larvae and adults feed on the eC (5);AB (6); SK (4); MB (5); ON (1a); PQ (1a); NB fruiting bodies of higher fungr. (2); NS (1) Subfamily DACNINAE Genus DACNE Latreille (Subgenus DACNE s. str.) (Ezgis Paykull) D. californica (Horn) _BC cephalotes Casey elongata Casey Iaticollis Casey uteana Casey D. quadrimaculata (Say) MB ON PQ (Subgenus XENODACNE Boyle) D. picea LeConte _BC Genus MEGAI-ODACNE Crotch M. fasciata (Fabricius) -oNPQ bifasciatus (Olivier) M. heros (Say) -oNPQ Subfamily TRIPLACINAE Genus TRIPLAX Herbst T. antica LeConte - BC AB SK mormonalis Casey T. californica LeConle - BC AB T. cuneata Boyle - BC AB T. dissimulator (Crotcht AB SK MB ON PQ carri Casey T. flavicollis Lacordaire - ON PQ confinis LeConte T. frosti Casey AB SK MB ON PQ NB T. macra LeConte MB ON PQ T. thoracica Say AB SK MB ON PQ NB antennata Casey breuicollis Casey conuergens Casey latiuscula Casey melano p t era Lacordarre obliqua Casey Genus TRITOMA Fabricius Cyrtotriplar Ctotch T. biguttata tSay) PQ basalis Lacordaire T. humeralis Fabricius - ON PQ ornata Casey ruficeps LeConte taeniata LeConte uittata LeConLe T. mimetica (Crotch) - ON PQ T. pulchra Say - ON PQ - NS basalis Melsheimer 225 Phalacrida,r cincta Lacordafte dimidiata Lacordaire sellata Ktthnt T. sanguinipennis (Sayl oN PQ T. unicolor Say ON unicolor Lacordaire Genus ISCIIYRUS Lacordaire Key to Ncrth American species: Skellay and Goodrich (1989) I. quadripunctatus quadripunctatus oN PQ (OIi rier) alabartae Schaeffer antediLisa Mader Family PHALACRIDAE shining flower beetles J.M. Campbell The flamily Phalacridae comprises about 600 to use. Phalacridae and Ptiliidae are probably the species around the world; only five have been most poorly known families of beetles in Canada, reported to occur in Canada, but many other species and based on existing literature, a reliable specific certainly occur. Adults are usually found on flow- identifrcation for most genera is not possitrle. ers; larvae feed in flower heads of Compositae or on spores aILd stomata of rusts and smuts. Casey (1916) revised the North American species o[ phalacrids, but this work is very difficult BC (3);MB (1); ON (3);PQ (2) T?ibe Phalacrini Genus PHIJ-ACRUS Paykull P. penicillabus (Say) BC Tribe Olibrini Genus OLLBRUS Erichson O. rufipes l.,eConte BC tritus ,Jasey O. semistri rtus LeConte ON striatulus LeConte Genus STII,BUS Seidlitz Eustilbus Shatp S. apicalis (Melsheimer) BC ON Genus LITOCHROPUS Erichson L. scalptus Casey PQ Genus ACI-I-AMUS Sharp A. pugetan-rs Casey MB ON PQ 226 Cerylonidae Family CERil ONIDAE cerylonid beetles J.M. Campbell OnIy about 400 species of Cerylonidae, a family feed on a plasmodium of slime molds (Slipinski that consists of small to very small beetles, are 1988). known around the world, of which seven have been Adults of this family are reasonably well known recorded in Canada and Alaska. Most adults are taxonomically. Sen Gupta and Crowson (1973) collected in leaf litter, in rotten wood, and under redefined the family and revised the higher classifi- bark. Hypodacne punctata is known in the galleries cation. Lawrence and Stephan (1975) revised the of Camponotus ants, and Murmidius oualis occurs North American species, and Lawrence (1982b) cat- on stored products (Lawrence and Stephan 1975). aloged them. Little is known about the feeding habits of the lar- vae and adults, although some larvae are known to AK (1); NT (1); eC (3); AB (2); tt{B (1); ON (6); PQ feed on fungal hyphae or spores, and some larvae (3);NB (1);NS (2) . I (1) Subfamily ELIXESTINAE Genus HYPODACNE LeConte H. ounctata LeConte ON Subfamily MURMIDIINAE Genus MURMIDIUS Leach M. ovalis (Beck)t AK ON Genus MYCOCERINUS Slipinski Mychocerus a:uct. M. depressus (LeConte) ON Subfamily CERYLONINAE Genus PHILOTHERMUS Aub6 P. g)abriculus LeConte ON PQ Genus MYCHOCERUS Erichson Lapethus Casey M. discretus (Casey) BC Genus CERYLON Latreille C. castaneum Say BC AB MB ON PQ NB NS clypeale Casey stricticutn Casey C. unicolor (Zieglet) NT BC AB ON PQ NS angustulum LeConte simplex LeConte syluaticum Casey 227 Corylophidae Family CORYLOPHIDAE minute firngus beetles J.M. Campbell Members of Corylophidae, a small family of species have yet to be discovered. Casey (1900) pre- about 4315 species, are found in all major faunal pared the last revision covering the North American regions, but only 11 species are known to occur in species of Corylophidae. The classificatic,n adopted Canada. Adults are usually collected in moldy leaf here is based on Paulian's (1950) reclassifrcation of litter ancl piles of grass cuttings. Adults and larvae the Old World taxa. are thought to feed on fungal spores. The taxonomy of the Canadian corylophid species is poorly known; reliable species identifrca- YK (1); NT (2); BC (7); AB (4); SK (2); MB (1); ON tion is not possible at this time, and a number of (a); PQ (3); NB (1) . I (2) Subfamily C ORYLOPHINAE Tribe Orthoperini Genus ORTHOPERUS Stephens O. atomus (Gyllenhal)t _BC O. scutellaris LeConte NT BC AB SK lucidus Casey O. suturalis LeConte ON Tlibe Rypobiini Genus GRC)NE\IUS Casey G. fuscicorrLis Casey oN PQ Subfamily SERICODERINAE Tribe Sericoderini Genus SERICODERUS Steohens S. lateralis (Gyllenhal)t BC PQ Subfamily SACIINAE Tlibe Saciini Genus MOLAMBA Casey M. biguttat,a (LeConte) BC AB M. fasciata (Say) oN pe N" s u b fa s : iata (Gu6rin-M6nevilIe) M. obesa Casey BC Genus SACIIUM LeConte Parmu.lus Harold S. lugubre l,eConte YK NT BC AB SK yuoN_ S. montanum Casey _BC Genus ARTHROLIPS Wollaston A. decolor (LeConte) califor nicus (Matthews) AB 228 Coccinellidae Family COCCINELLIDAE ladybird beetles J. McNamara Coccinellidae comprises 162 species and sub- monograph by Gordon (1985). Only some of the species in Canada and Alaska. Many species are most recent references on coccinellids have been introduced in North America, some of these pur- listed in the checklist. Good regional works on posely for biological control. Most coccinellid adults Coccinellidae include those of Chapin (1974) for and larvae are beneficial, because they are preda- Louisiana, Belicek (1976) for western Canada and tors of aphids, scale insects, and mites. However, in Alaska, and Larochelle (1979) for Quebec. the absence of preferred food, many species feed on the immature stages of other insects. The species of Psyllobora feed on fungi, and the only species of Epilachna that occurs in Canada feeds on plants of the family Leguminaceae. /J{(26); YK (31); NT (35); BC (9a);aB (83); SK (70); The taxonomy of the North American MB (56); ON (83); PQ (77);NB (37); NS (25);pE (6); Coccinellidae is well known. thanks to the recent LB (9);NF (13) . H (2); I (8) Subfamily STICHOLOTIDINAE Tribe Microweisini Genus MICROWEISEA Cockerell Epismilia Cockerell Pseudoweisea Schwarz SmiliaWeise Key to species: Gordon (1970o) M. misella (LeContet BC AB SK MB ON PQ NB NF Genus COCCIDOPHILUS Brethes Cryptoweisea Gordon Key to North American species: Gordon 0970a) C. marginatus { Leconte) BC AB SK MB ON PQ NB Tlibe Serangiini Genus DELPHASTUS Casey Key to species: Gordon (19706) D. pusillus (LeConte) ON PQ puncticollis (LeConte) sonoricus Casey Subfamily SCYMNINAE Tlibe Stethorini Genus STETHORUS Weise Key to New World species: Gordon and Chapin (1983) S. punctillum Weiset BC -oN S. punctum punctum (LeConte) ABSK-ONPQ-NS S. punctum picipes Casey BC 6reuis Casev Tlibe Scymnini Key to North American species: Gordon (1976) Genus DIDION Casey D. longulum Casey NTYK BC AB SK PQ occiduum (Casey) paruiceps Casey D. nanum (LeConte) ON PQ D. punctatum (Meisheimer) _NTBCABSKMB ON PQ NB_PE NF 229 Coccinellidae Genus SCYMNUS Kugelann (Subgenus {ICYMNUS s. str.) S. americar..us Mulsant oN PQ S. apicanus pseudapicanus Gordon AB SK MB boreal;s Gordon S. caurinus Horn BC AB aluticctllis Casey S. fenderi Ilalkin BC S. indianensis Weise ON rustict;s Casey S. nebulosus LeConte BC harneli Hatch infuscc:tus Boheman phelps;'.i Crotch S. opaculus Horn S. paracanus linearis Gordon - - - :t ffi 3[ nre------(Subgenus PULLUS Mulsant) S. abbreviai;us LeConte ON PQ S. aquilona::ius Gordon _ABSK S. ardelio Eorn BC AB apacheanus Casey decipeins Casey sanctu; Casey S. aridus Cr,rsey BC S. brullei Mulsant SK MB ON PQ ivs--- diuisut Casey laureniicus Casey lodi Stt}rr subaen:.eus Casey S. calaveras Casey BC saginalus Casey strenuus Casey stygicu:.s Casey tenuiue stis Casey S. carri Gordon S. caudalis LeConte _ _ _ rSf nre oN pe agricola Casey indutus Casey natche zianus Casey S. cervicalir Mulsant -oN S. compar ( asey -oN impun:tusWingo uicksbt.rgicus Casey S. coniferarum Crotch BC S. fraternus LeConte ON PQ dentipes FaIl haemo;rhous LeOonte S. humboldt,i Casey _BC S. impexus Mulsant'l _BC lln NS_- S. impletus Gordon _BC S. iowensis Casey _BC_SKMB oN PQ NB collarit Melsheimer melsheimeri Weise S. kansanus Casey -ONPQ S. lacustris LeConte YI( NT BC AB SX MB ON PQ NB NS cultratus Wingo S. marginic, >ll is Mannerheim BC AB ON PQ borealis Hatch californicus Boheman desertcrum Casey S. postpictus Casey BC AB S. puncticollis LeConte :" -oNpe-- S. rubricaudus Casey _ONPQ canteri.us Casey chromopygus Casey texanus Casey S. securus J. Chapin oN_ S. solidus Casey nc blaisde'lli Casey 230 Coccineilidae S. suturalis Thunbergt PQ D/r S. tahoensis Casey S. tenebrosus Mulsant NT AB_ -oN PQ S. uncus Wingo AB SK MB ON S. utahensis Gordon AB Genus NEPHUS Mulsant (Subgenus NEPHUS s. str.) N. ornatus naviculatus (Casey) AK BC AB SK MB ON PQ NF h.incaidi (Hatch) N. ornatus ornatus (LeConte) _ONPQ NB NS frosti (Casey) sanguinifer (Casey) (Subgenus TURBOSCYMNUS Gordon) N. georgei (Weise) AK YK NT BC AB ON PQ NF bisignatus (Horlr) (Subgenus SCYMNOBIUS Casey) N. flavifrons (Melsheimer) ON PQ bioculatus (Mulsant) guttiger (Mulsant) Iudouictanus (Casey) marginellus (Mulsant) N. intrusus (Horn) ON inops (Casey) N. sordidus (Horn) SK MB ON intrusoides (Hatch) Genus DIOMUS Mulsant D. amabilis (LeConte) PQ dulcis (Casey) emertoni (Casey) D. debilis (LeConte) AB SK ON PQ aeger (Casey) minutissimus (Casey) molliculus (Casey) pusio (Casey) D. liebecki (Horn) MB ON adulans (Casey) ohioensis (Stehr) D. terminatus (Say) ON femoralis (LeConte) partitus (Casey) Tlibe Hyperaspini Genus HYPERASPIDIUS Crotch H. comparatus Casey BC juniperus Nunenmacher H. hercules Belicek _AB H. insignis Casey -AB H. mimus Casey _AB SK carri Nunenmacher c oloradens is Nunenmacher H. vittigerus (LeConte) YK_BC AB SK Genus HYPERASPIS Redtenbacher Oxynychus LeConte H. bigeminata (Randall) -oN PQ NB gueri Mulsant H. binotata (Say) SK MB ON PQ NB affinis (Randall) Leucopsis Melsheimer normata (Say) paspalis Watson H. borealis Dobzhansky YK BC AB eloli Nutting obscura Malkin 237 Coccinellidae schuhi }{atch simuloides Hatch H. consimilis LeConte NT BC AB SK MB PQ NB canadensis Dobzhansky H. conviva rlasey SK MB ON PQ congressis Watson insolen.s Casey H. disconotata Mulsant AB _MBON PQ NB_ H. dissolutzL dissoluta Crotch BC coloraclana Casey nupta,iasey H. fastidiosr Casey _BCAB s e pte nt rion is Dobzhansky H. inflexa Casey _BCAB SK MB PQ serena Casey H. jasperentis Belicek NT_AB H. lateralis Mulsant _BCAB SK flammula Nunenmacher idoe N rnenmacher laeuipennis Casey montartica Casey nig rocc. uda Dobzhansky omissa Casey wellmani Nunen.macher H. moerens (LeConte) H. octavia C asey _ _ sx_ 3NF3 NB H. oregona )obzhansky BC AB lanei E atch H. postica I eConte BC AB SK angust tla Casey elliptict Casey essigl n{alkin H. pratensis LeConte PQ H. proba (Say) SK MB ON PQ trinifer Casey H. quadrivittata LeConte BC AB SK tetrane;tra Casey H. signata signata (Olivier) SK taedato. LeConte H. simulatrix Dobzhansky BC AB SK H. troglodyt:s Mulsant _MB pe lls discretc: LeConte nouasct'liae E.A. Chapin H. undulata (Say) BC SK MB ON PQ NB elegans Mulsant guttifer a Weise maculilbra Melsheimer Genus BRA(IHIACANTHA Dejean B. albifrons iSay) AB SK MB pacifica Casey B. decempustulata (Melsheimer) -oN PQ NB NS stellata Casey troglod^rtes Mulsant B. dentipes (Fabricius) ON separatiz Leng socialis Casey B. quadripunctata quadripunctata ON PQ Melsheimer basalis Melsheimer confusa Mulsant diuersa Mulsant B. rotunda Ci'ordon ON PQ B. ursina (Fabricius) AB SK MB ON PQ NB NS congruens Casey B. uteella CzLsey BC fenyesi Leng lezgl Nunenmacher sonorana Casey 232 Coccinellidae Subfamily CHILOCORINAE Tribe Chilocorini Key to genera: Chapin (1965) Genus BRUMOIDES E.A. Chapin B. histrio (Fall) - BC paruicollis (Casey) B. septentrionis septentrionis AK YK NT BC AB SK MB ON (Weise) desertorutn (Casey) neuadensis (Leng) ouoideus (Casey) B. septentrionis davisi (Leng) - ON PQ Genus EXOCHOMUS Redtenbacher E. aethiops rBlandi AB SK mormonicus Casey E. californicus Casey - BC Genus CHILOCORUS Leach C. hexacyclus Smith AB SK C. stigma (Say) AB SK MB ON PQ NB NS biuulnerus Mulsant C. tricyclus Smith - BC Subfamily COCCIDULINAE Tlibe Coccidulini Genus COCCIDULA Kugelann C. lepida LeConte AK YK NT BC AB SK MB ON PQ NB occidentalis Horn reitteri Dodge suturalis Reitter szlzrolis Weise Genus RHYZOBIUS Stephens Lindorus Casey R. lophanthae (Blaisdell)t - BC Subfamily COCCINELLINAE Tribe Coccinellini Genus ANISOSTICTA Dejean A. bitriangularis (Say) AK YK NT BC AB SK MB ON PQ NB NS _ LB NF irregularis Weise multig uttata (Randall) A. borealis Timberlake AKYKNT_MB_PQ Genus MACRONAEMIA Casey M. episcopalis (Kirby) AK YK NT BC AB SK MB ON PQ Genus PARANAEMIA Casey P. vittigera (Mannerheim) _rn_ similis Casey Genus COLEOMEGILLA Timberlake Megilla Mulsant C. maculata lengi Timberlake -oNPQ Genus CERATOMEGILLA Crotch C. ulkei Crotch AK YK NT BC AB ON poruo (Watson) 233 Coccinellidz,.e Genus HIPI'ODAMIA Dejean H. americana Crotch NT BC AB SK MB ON PQ H. apicalis Casey _BC lezgl Johnson nig r om a c ula ta (Nunenmacher) tricolor Nunenmacher H. arctica (lichneider)x AK YK NT BC _PQ LB H. caseyi Jc,hnson _BCAB H. convergens Gu6rin-M6neville _BCABSK MB ON PQ NB juncta Jasey modestz Melsheimer obsoleta Crotch pratico!.a Mulsant H. expurgata Casey _YK_BCABSK H. falcigera Crotch AK YK NT BC AB SK MB PQ albertana Ca.sey cottlei (.Nwenmacher) H. glacialis glacialis (Fabricius) NT BC AB SK MB ON PQ H. glacialis lecontei Muisant NT_ABSKMB abduce.zs Casey defecta Johnson h op p in,;1 i Nunenmacher macher.zie E.A. Chapin p se u do;1 lacialis Johnson H. lunatomi rculata dobzhanskyi YK BC E.A. Chapin H. moesta nLoesta LeOonte _BC H. moesta b':wditchi Johnson _BCABSK MB H. oregonensis Crotch _BCAB cockere.'li Johnson d,ispar'Jasey liliputo.na Casey puncticollis Casey H. parenthesis (Say) BC AB SK MB ON PQ NB NS albomacula Fitch appror,:mata Fitch confluezta Fitch connatu Fitch discopttnctata Fitc}' insulata Fitch linearis Fitch liturico!.lis Fitch nimia I'itc}r permac"ifrons Fitch triangu.laris Fitch tridens (l\ttby) tridentifrons Fitch H. quindecirnmaculata Mulsant _SK_ON H. quinques ignata quinquesignata AK YK NT BC AB SK MB ON PQ NB_PE (KirLY) coccineu Casey leporina Mulsant mulsanti LeConte pugetar'.a Casey quadra,"ia Casey subsiru.lis Casey uteana Casey uernis Oasey H. quinquesignata ambigua LeConte BC obliqua Casey punctulata LeConte H. sinuata crotchi Casey NT BC AB SK MB disjunc;la Timberlake fontinaiis Casey straminea E.A. Chapin H. sinuata spuria LeConte AK YK _BC cornpler Casey H. tredecimpunctata tibialis (Say) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF 234 Coccineliidae H. variegata (Goeze)t H. washingtoni Timberlake BC 9*PQ_ Genus ANATIS Mulsant Key to species: Watson (1976) A. Iabiculata (Say) MB ON PQ NB canadensis Provancher coseyl Westcott q uinde c i m p u n c t a ta (Oliv iet) A. lecontei Casey BC AB A. mali (Say) AKYKNT BC AB SK MB ON PQ NB NS PE NF borealis Belicek signaticollis Mulsant A. rathvoni (LeConte) BC AB SK Genus MYZIA Mulsant Neomysia Casey Paramysia Reitter M. interrupta (Casey) BC AB coseyi (Timberlake) M. pullata (Say) YK AB SK MB ON PQ NB NS LB montana (Casey) nolons (Randall) randalli (Casey) M. subvittata (Mulsant) BC AB hornii Crotch oregona (Casey) Genus CAI-VIA Mulsant Anisocaluia Crotch C. quatuordecimguttata (Linn6)* AK YK NT BC SK MB ON PQ NB NS LB NF bicordifera (Casey) cardisce (Randall) comple*a Nunenmacher elliptica (Casey) hesperica (Crotch) incarnata (Kirby) lacustris (Casey) obliqua (Randall) postplagiata (Casey) quadrisignata (Casey) sicordi (Nunenmacher) simllis (Randall) uniformis (Casey) uancouueri (Casey) uictoriana (Casey) Genus ADAI,IA Mulsant A. bipunctata ilinn6)* AK YK NT BC AB SK MB ON PQ NB NS NFLB annectans (Crotch) bioculata (Say) coloradensis Casey disjuncta (Randall) duplicata Leng frigido (Schneider) humeralis (Say) ludouicae Mulsant mel anop I eur a (LeConte) ocellata Johnson opthalmica Mulsant ornatella Casey ouipennis Casey postica Johnson sexpustulata Johnson transuersalis Casey Genus COCCINELLA Linn6 Key to North American species: Brown (7962b) 235 Coccinellidi,'e C. alta W.J. Brown BC AB sutura'.is Casey C. californica Mannerheim AK BC AB melanc'collis Johnson C. fulgida V/atson NT BC ON PQ C. hieroglyphica humboldtiensis TY BC Nunenmacher C. hieroglyphica kirbyi Crotch _YK NT BC SKAB MB ON PQ NB NS tricuspis Kitby C. hieroglyphica mannerheimi AK YK NT MB Muklant C.johnsoni Casey AX BC C. monticola Mulsant AK_NTBCAB SK MB ON PQ Nn f,IS - alutacea Casey bigutta.ta Johnson confluenta Johnson impres';a Casey lacustris LeConte neuadica Casey postica Johnson sellica Johnson C. novemno:ata Herbst BC AB SK MB ON PQ confluenta Fitch conjunt:ta Fitch degenet'Casey diuisict'llis Fitch francis,:ana Mulsant inaequulis Fitc}:' oregontt Casey paruan,,aculato Fitch C. prolongata prolongata Crotch BC A.B C. septempr..nctata Linn6J _BCABSKMBONPQ NB NS C. transvers oguttata richardsoni AK YK NT BC AB SK MB ON PQ NB NS in Ne W.J. Brown interrulta Fitdn quinquznotata I{ttby zetterstzdti Madet C. trifasciat,r perplexa Mulsant AK YK NT BC AB SK MB ON PQ NB NS _LBNF C. trifasciatl subversa LeConte _BC barda T,eConte euginii Mtisant juliana Mulsant praedicta Mad.et C. undecimp,unctata undecimpunctata BC ON PQ NB NS PE NF Linn5t Genus PRO )YLAEA Mulsant P. quatuord,:cimpunctata (Linn6)t ON PQ NB Genus CYC.i,ONEDA Crotch C. munda (Say) MB ON PQ C. polita Casey _BCABSK flauaT'lr'berlake Genus OLLr\ Casey O. v-nigrum (Mulsant) BC A3 SK MB ON abdominalis (Say) arizonae Casey fenestra'.lis Casey minuta Casey plagiaht Casey soyi (Clotch) semiluri.aris Johnson sobrina Casey Genus NEO.FIARMONIA Crotch Agrabiu Casey Harmottiaspis Casey Neoharnonia Casey 236 Endomychidae N. venusta venusta (Melsheimer) ON centralis Casey dissimila Blatchley fattigi B\atc}:.ley notulata (Mulsant) Genus MULSANTTNA Weise Pseudocleis Casey Key to species: Chapin (1985) M. hudsonica (Casey) AK YK NT BC AB SK MB ON PQ NB NS PE LB M. picta (Randall) _YK _BCABSKMBONPQNBNS blanchardi (Johnson) c on cinnata (Melsheimer) contert(r (Mulsant) impictipennis (Weise) minor (Casey) nubilata (Casey) Tribe Psylloborini Genus PSYLLOBORA Dejean P. borealis Casey BC deficiens Casey P. vigintimaculata (Say) AK YK NT BC AB SK MB ON PQ NB NS NF obsoleta Casey separata Casey taedata LeConte Subfamily EPILACHNINAE Key to New World species: Gordon (1975) Tribe Epilachnini Genus EPILACHNA Dejean E. varivestis Mulsant oN PQ ceruina Mulsant corrupta Mulsant cuprea Cockerell genuina Mulsant juncta Johnson maculiuentris Bland toweri Johnson Family ENDOMYCHIDAE handsome ftrngus beetles J.M. Campbell The family Endomychidae is found in most taxonomically well known. However, the family has regions of the world, but the majority of species never been revised for North America. Strohecker occur in tropical areas. The range of 15 species (1953) reviewed the genera of the world and pub- extends northward into Canada. Adults and larvae lished a catalog of the North American species may be collected on soft fungi, in leaf litter, or (1986). Hatch (1962) treated the genera and species under bark with fungal growth. One species, that occur in the Pacific Northwest. A modern revi- Mycetaea subterranea (M. hirta in most publica- sion of the North American species of this family is tions), is a pest of stored products. Most species needed. feed on fungr. Because of their moderately large size and often NT (1); BC (7); AB (2); SK (2); MB (5); ON (10); PQ bright coloration, many North American species are (8);NB (2);NS (6);PE (1);NF (2) ' I(2) 237 Endomychidae Subfamily MYCETAEINAE Tribe Mycetaeini Genus SYMBIOTES Redtenbacher S. duryi Blatchley -oN Iacustris Casey oblong,ts Casey pilosus Casey waltoni Dury S. gibberosr;.s .(Lucas)t -oN duryiWalton montanus Casey Genus MYCETAXA Stephens M. subtenanea (Fabricius)f _BC_ONPQ_NSPE_NF fumata Stephens hirta (Ivlarcham) Tribe Leiestini Key to North American species: Blaisdell (1931) Genus RHANIDEA Strohecker Rhanis LeConte R. unicolor Zieglerr - ON PQ apicalis (Melsheimer) haemo,"rhoidalis (Gu6rin-M6nevi11e) Genus STE'IHORHANIS Blaisdell S. borealis ]]laisdell - BC Genus PHYMAPHORA Newman P. californica Horn - BC P. pulchella Newman MB ON PQ - NS cr a s s icor nis ( Melsheimer) p u nc t i r:' o I I i s (Zi e gl er') Subfamily STENOTARSINAE TYibe Stenotarsini Genus DA]riAI Reiche D. testacea iZiegler) _ONPQ_NS fer r ug i ne a (Gu6rin-M6neville ) lzleo (lvlelsheimer) piloso i.Melsheimer) Subfamily ELIMORPHINAE Tribe Lycoperdinini Genus LYC'OPERDINA Latreille (Subgenus (]OLGIA Mulsant) L. fermgine a LeConte - BC AB SK MB ON PQ - NS angulata (Say) g lab rala Gu6rin-M6neville TYibe Mycetinini Genus APHORISTA Gorham A. laeta (LeConte) A. vittata (Fabricius) ---:"--meo*pe----- distincta (Say) lineata (Olivier) ouipenizis Casey 238 Lathridiidae Genus MYCETINA Mulsant M. idahoensis Fall BC M. perpulchra (Newman) MB ON PQ NB NS q uadrig uttata (Melsheimer) Subfamily ENDOMYCHINAE Tribe Endomychini Genus ENDOMYCHUS Panzer E. biguttatus Say _NTSKMB ON PQ NBNS_ NF E. limbatus (Horn) BC AB endomychoides (Fall) Family LATHRIDIIDAE minute brown scavenger beetles Y. Bousquet In Canada and Alaska 55 species of lathridiids Most of the genera, particularly those of the sub- have been recorded. These species are usually family Corticariinae, are in need of modern revision. found on foliage, in rotten wood, under logs, and The generic classifrcation of the Lathridiinae used under the bark ofdead trees, as well as in fungi and here follows that of Walkley (1952). nests of social Hymenoptera, birds, and mammals. A number of species, including Cartodere constricta and Aridius nodifer, are regularly found in stored products, especially those that are damp and moldy. Adults and larvae feed on fungr, particularly molds. AK (19);YK (5);NT (6); BC (3a); AB (9); SK (8); MB Lathridiidae is another family of Cucujoidea (17); ON (22); PQ (22); NB (6); NS (8); PE (1); LB that has been poorly studied in North America. (1);NF (a) . H (6);I (15) Subfamily I"ATHRIDIINAE Genus DIENERELLA Reitter Microgramme Walkley D. arga (Reitter)t oN PQ D. cordicollis (Mannerheim) AI{ D. costulata (Reitter)t _MBONPQ D. frliformis ( Gyllenhal)f SK MB NB PE D. filum (Aub6)t ---ec- _MBONPQ D. rufrcollis (Marsham)t _BC ON PQ _NS NF p ulicar ia (Melsheimer) Genus ENICMUS C.G. Thomson E. aterrimus Motschulsky oN PQ ferrugineus Belon opaculus LeConte E. cordatus Belon -BC E. fictus FaIl AK NT BC AB ix ms o* Pe E. mimus Fail AK NT BC AB SK MB E. nitens FalI y_ yuoNpe_ E. tenuicornis LeConte BC _ _ _ _ laticollis LeConte Genus THES Semenov T. bergrothi (Reitter)f BC SK MB NB NS tricarinatus (W.J. Brown) Genus LATHRIDIUS Herbst Conithassa C.G. Thomson L. consimilis Mannerheim* AK par allelocollis Mannerheim L. crenatus LeConte BC 239 Lathridiidae L. hirtus Gyllenhalf _BC MB ON PQ L. minutus (Linn6)+ _BCAB SK MB porcatus Herbst reflextls LeConte L. nigritus (Fall) _BC L. protensi :oliis Mannerheim AK BC quadr :collis Mannerheim sobrinus Mannerheim suspectus (Fall) Genus STEPHOSTETHUS LeOonte S. breviciavis (FalI) -oNPQ NB NS S. cinnamopterus (Mannerheim) AK _BC S. costicollis (LeConte) AK _BC armat:zlus (Fall) S. curtulus (Mannerheim) AK S. lardariuir (DeGeer)f NF S. liratus (LeConte) AK BC AB SK MB ON PQ wS carinil'er (Fa\\') trilobatus (Motschulsky) S. montants (Fall) AK YK NT BC MB PQ laeuior (W .J. Brown) Genus ADISTEMIA Fall A. watsoni lWollaston)t ON PQ Genus CAIi,TODERE C.G. Thomson Coninomus C.G. Thomson C. constrict,a (Gyllenhal)t BC AB SK MB ON PQ NB NF sculpti.lis (LeConte) Genus ARI )IUS Motschulsky A. nodifer ( Westwood)f BC MB ON PQ Subfamily CORTICARIINAE Genus COI,,TICARIA Marsham C. canalicu ata Mannerheim AK C. columbia. FalI _BC C. dentigera LeConte _BC ON PQ C. dentiventris Poppius* AK YK C. fenestralis (Linn6)* AX BC deleta Mannetlr'eim denticulata I{trby kirbyi LeConte obtusa LeConte rugulo.;a LeConte C. ferrrrginea Marshamx AK NT MB PQ C. Iinearis (Paykull)x AK NT MB PQ- _ _ luIu C. orbicollis Mannerheimx AK YK _BC munsteri Stand C. parallela Fall YK AB C. pubescers Gyllenhal-i _BC grosso LeConte -on C. salpingoides Motschulsky _BC C. serrata (Paykull)f _BC octodentata (Say) C. serricollis LeConte ON C. spinulosa Mannerheim AK C. valida Fall -nc C. varicolor Fall _BC Genus MEL,ANOPHTHALMA Motschulsky M. americana (Mannerheim) BC ON PQ distinguenda auct. amer. morsa',LeConte) pusilla (Melsheimer) rufula (.LeConte) 240 Byturidae M. pumila (LeConte) _BC_ON M. villosa (Zimmermann) _BC Genus CORTICARINA Reitter C. alberta (Fall) AB C. cavicollis (Mannerheim) -BC-ONPQ-NS C. exigua (Mannerheim) AK C. fuscula (Gyllenhal) AK l'K NT BC MB ON PQ americana auct. amer. compt(r (LeConte) expansa (LeConte) grata (LeConte) C. herbivagans (LeConte) _BC leuis (LeConte) similata a:uct. Genus CORTINICARA Johnson C. gibbosa (Herbst)t _BCABSKMBONPQNBNS Genus CORTILENA Motschulsky C. picta (LeConte) -oNPQNBNS Family BIPI{YLLIDAE false skin beetles J.M. Campbell Only four species of Biphyllidae are known to This family is poorly known taxonomically, and occur in North America, one of which, Diplocoelus the North American species have not been revised. brunneus, is found in eastern Canada. Little is known about the habits of this species; adults are ON (1);PQ (1);NB (1) found in fungus or under bark. Tlibe Biphyllini Genus DIPLOCOELUS Gu6rin-M6neville Marginus LeConte D. brunneus (LeConte) - oN PQ NB Family BYTURIDAE fouitworm beetles J.M. Campbell Two species of Byturidae are found in North sion by Springer and Goodrich (1983) of the North America of which one, Byturus unicolor, the rasp- American species of the family recognized only one, berry fruitworm, is common and widely distributed somewhat variable species. across Canada. Adults ofthis species feed on leaves and flowers, and the larvae feed on the receptacle of the red raspberry, Rubus idaeus. Byturus unicolor has been known under a vari- aK (1); yK (1); NT (1); BC (1); AB (1); SK (1); MB ety of Latin and common names, but a recent revi- (1); ON (1); PQ (1); NB (1);NS (1);NF (1) Tribe Byturini Genus BYTURUS Latreille Horticola Lindemann B. unicolor Say AK YK NT BC AB SK MB ON PQ NB NS NF americanus Gemminger & Harold 247 Mycetophagidae baker;'.Barber breuicollis Casey punctatus Casey rubi Eiarber sordictus Barber Superfamily Tenebrionoidea Family MYCETOPIHGIDAE hairy fungus beetles Y. Bousquet Of the 26 species of mycetophagids recorded Parsons (I975b) has revised the mycetophagids from Nc,rth America, 15 are known to occur in of theNearcticfauna. Canada. These beetles are found mainly in decay- ing vegetable matter, in fungi, and under the bark of rotten logs. A few species, such as Typhaea ster- corea, are occasionally found in buildings. As the Latin name suggests, members of this family are BC (6);aB (4); SK (2); MB (7); ON (10); PQ (12); NB mycetophagous. (2);NS (1); PE (1) . I (1) Genus MY,IETOPHAGUS Hellwig (Subgenus MYCETOPHAGUS s. str.) M. californicus Horn - BC prouer,,sis Casey M. flexuosrrs Say MB ON PQ M. punctat us Say MB ON PQ M. serrulat,us (Caseyt MB ON PQ pictus (.Casey) quadralius Casey subderzressus (Casey) tribaheatus Casey (Subgenus ILENDUS Casey) M. distinctus Hatch - BC AB SK MB M. pluripunctatus LeConte AB - MB ON PQ (Subgenus PARILENDUS Casey) M. quadriguttatus P.W.J. Miillerf - BC AB SK MB ON PQ NB bimac ulatus Melsheimer bipust.ulatus Melsheimer b isig n at us Melsheirner uariegatus C.R. Sahlberg (Subgenus GRATUSUS Casey) M. plurigu,tatus LeConte - BC franciscanus Van Dyke M. tenuilasciatus Horn - BC AB ON PQ notalulus ( Casey, Genus LITARGUS Erichson (Subgenus AJ-ITARGUS Casey) L. balteatus LeConte - ON PQ infulatus LeConte tr(l,nsuersus LeConte (Subgenus PARALITARGUS Casey) L. didesmus (Sayt PQ asperulus Casey (Subgenus LITARGUS s. str.) L. sexpuncbatus tSay) - ON PQ obsolescens Casev 242 Tetratomidae (Subgenus TILARGUS Casey) L. tetraspilotus LeConte MB ON PQ Genus TYPHAEA Stephens T. stercorea (Linn6)f - BC - ON PQ NB NS PE fumata (Linn6) Genus THRIMOLUS Casey T. minutus Casey PQ duryi Casey FamilyTETRATOMIDAE tetratomidbeetles L. LeSage The family Tetratomidae includes nine species on beetles, such as Blatchley (1910), Chagnon and in Canada and Alaska. Both adults and larvae live Robert (1962), Hatch (1965), and Arnett (1968). in the fruiting bodies of higher fungi and are myce- Young (1976) recently transferred the genus tophagous. Pectotoma, which had been previously placed in this Previously considered a subfamily of family, to Scraptiidae. The survey of the Melandryidae, the tetratomids were raised to family Tetratomidae of Quebec by Chantal (1985) includes status by Crowson (1955). This ranking is accepted genera placed here in Melandryidae. by most recent authors, including Miyatake (1960) and Lawrence (1982a). There is no recent revision ofthefamilyinNorthAmerica. Generaandspecies aK(1); YK(1); nC(2); AB(1);SK(1); ON(5);PQ can be identifred tentatively by using general books (7); NB (1);NF (1) . I (1) Subfamily TETRATOMINAE Genus TETRATOMA Fabricius T. concolorleConte - YK - BC AB SK T. truncorum LeConte - ON PQ Genus ABSTRULIA Casey A. tesselata (Melsheimer) - ON PQ maculata Casey A. variegata Casey PQ NF Genus EUPISENUS Casey E. elongatus (LeCon[e) AK BC Genus PISENUS Casey P. humeralis (Kirbyt - ON PQ concolor (Kirby) ruficornis (LeConte) Subfamily PENTHINAE Genus PENTHE Newman Pytocis Castelnau P. obliquata (Fabricius) -oNPQ P. pimelia (Fabricius) -oNPQNB anglica (Gmelin) funerea Newman lesueuri (Castelnau) morio (Fabricius) Genus MALLODRYA Horn M. subaenea Horn prl 243 Ciidae Family CIIDAE minute tree-fungus beetles J. McNamarq, The family Ciidae, which consists of small bee- Lawrence (1971) revised the North American tles, comprises 29 species in Canada and Alaska. species and also published a catalog of the species The larvae and adults of nearly all species feed on (1982c). fungal h'7phae comprising the mycelia and especially the fruiting bodies of wood rotting Basidiomycetes. Several l;pecies are found under bark, in dead wood, eK (6); xT (6); BC (19); AB (10); SK (5); MB (14); or in the galleries ofbark beetles. ON (16);PQ (17);NB (5);NS (6);NF (6) ' I (1) Subfamily CIINAE Tlibe Ciini Genus CIS Latreille Xestocis Casey C. americanus Mannerheim AK BC AB MB ON PQ NB NS NF forsteri Abdullah frosti Dury hatch, Lawtence insolerc (Casey) minot (Hatch) nitidt s (Hatch) oweni (.Hatch) serricollis Drty strigulosus (Hatch) C. angustus Hatch _BC C. biarmatus Mannerheim AK _BC C. castlei (Dury) -o* C. creberrimus Melli6 PQ C. ephippiatus Mannerheim AK -nc _:* PQ moznt'ttei (Dtry) C. fuscipes Melli6 NT BC AB SK MB ON PQ _NS NF atripennis Me1li6, caroli:zae Casey cheurolatii Melli6 dubius Melli6 impressus Casey pallens Casey C. horriduius Casey NT BC MB ON NS mormonicus Casey C. hystriculus Casey -BC C. levettei (Casey) _BCABSKMBONPQ NB NS NF C. maritimus (Hatch) _BCMB C. pistorius Casey NT-ABSKMBONPQ C. striolatus Casey NTBCAB_MBONPQ NS-- fraternus Casey macilentus Casey C. subtilis Mel1i6 PQ confusus Blatchley julichiDrry wenzeli Dtry C. tridentatus Mannerheim AK BC ednae (I].atch) reflexus (Hatch) Genus ENNEARTHRON Melli6 E. spencen (Hatch)t -BC Genus DOLICHOCIS Dury D. indistinctus Hatch -BC PQ D. manitoba Dury NT BC AB MB ON PQ NB- 244 Melandryidae Genus ORTHOCIS Casey O. punctatus (Mel1i6) AK NT BC AB MB ON PQ NF aterrimus Casey Genus STRIGOCIS Dury S. opacicollis Dury ON Genus HADRAULE C.G. Thomson Maphoca Casey H. blaisdelli (Casey) BC H. elongatula (Gyllenhal)t NB Genus PLESIOCIS Casey P. cribrum Casey BC AB MB PQ Genus CERACIS Me11i6 Key to North American species: Lawrence (1967) C. sallei (Melliel oN PQ C. singularis t Duryt C. thoracicornis t Zieglert r,rs3Npe- - bifoueata Dtry ------laminifrons (Casey) mellyi (.Mel1i6) oblonga (Blatchley) picea (Casey) pumicdta (Meili6) unicolor (Casey) Genus SULCACIS Dury S. curtula tCasey) BC AB SK MB PQ criddlei (Drry) cylindrica (Dury) montana (Casey) nigra Dury sorror (Casey) Genus MALACOCIS Gorham Brachycis Casey M. brevicollis (Casey) MB ON PQ NF Subfamily OROPHIINAE Tlibe Orophiini Genus OCTOTEMNUS Melli6 O. laevis Casey AK BC AB SK MB ON PQ NB NS NF denudatus Casey Family MELANDRYIDAE false darkling beetles L. LeSage Melandryids are common throughout the world beetles work inconsistently. Up-to-date revisions at and include about 100 genera and 1200 species, of the species level are badly needed. Chantal (1985) which at least 55 occur in Canada and Alaska. keyed out the species of Hallomeninae and Adults live under bark, in fungi, or in flowers. Eustrophinae from Quebec. At this time, no expert is Larvae are carnivorous or feed on decaying plant working on the family in North America. The classifi- material; they live under bark, in dead logi, or in cation used here corresponds to the arrangement sug- stumps; a few are found in leaf litter, but detailed gested by J.F. Lawrence (personal communication). life-history studies, such as that of Hoebeke and McCabe (1977), are seldom published. aK (4);YK (2);NT (3); BC (27); AB (10); SK (5); MB The taxonomy of the family is in a state of disar- (e); ON (27);PQ (42); NB (11); NS (10); LB (3); NF ray at all levels. Generic keys from general books on (13) . H(3);I(2) 245 Melandryidre Subfamily IIALLOMENINAE Genus HALLOMENUS Panzer H. binotatu,; (Quenselti PQ H. debilis L,:Conte PQ ziger Fialdeman H. punctulatus LeConte AK BC - ON PQ basalis Mannerheim H. scapulars Melsheimer - BC - ON PQ Iuridus Haldeman ruficor,zis Melsheimer H. serricornis LeConte - BC AB ON PQ Subfamily EUSTROPHINAE Genus EUS'IROPHINUS Seidlitz E.bicolor(Fabriciusr - ON PQ indistittctu s ( LeConte) E. confinis t LeContet - BC - SK MB ON PQ Genus EUSTROPHUS ll I iger E. tomentosus Say - BC - ON PQ niger Nlelsheimer Genus SYNSTROPHUS Seidlitz S. repandus t Horn t - BC MB ON PQ Genus HOL,OSTROPHUS Horn H. bifasciat rs (Sayt - ON PQ q uadr i:nac ulalzs (Melsheimer) H. discolor l{orn PQ H. impressi':ollis tl-eConter - BC Subfamilv MELANDRYINAE Trile Orchesiini Genus ORCHESIA Latreille Key to North American species: Lalibert6 (1966) O. castanea (Melsheimer) - BC MB ON PQ - NS NF O. cultrifonnis Laiibert6 - NT SK MB - PQ - NS O. ornata Horn - BC O. ovata La ibertti ON PQ Genus MIC ROSCAPHA LeConte M. clavicon.is LeConte PQ Genus LEDERIA Reitter Crioscctpha Flotn L. arctica (llorn) AK BC Tlibe Melandryini Genus EMNIESA Newman E. connectens Newman - ON PQ - NS - LB NF maculcrta LeConte E. labiata (,(iay) - ON PQ NB - NF E. stacesmi-,hi Hatch - BC E. testacea leeperi Malkin - BC Genus MET,ANDRYA Fabrici us NeoemtnesaHatch M. blackmani Hatch PQ M. striata Say - BC AB SK - ON PQ NB bicolor Melsheimer 246 Melandryidae diuersa Mulsant excauata Haldeman thoracica Melsheimer Genus PHRYGANOPHILUS C.R. Sahlberg P. collaris LeConte NT BC AB - MB ON PQ NB Tlibe Hypulini Genus ZILORA Mulsant Key to North American species: Mank (1938) Z. canadensis Hausen PQ Z. hispida LeConte - YK - BC AB ON PQ NF Z.nudaProvancher PQ Z. occidentalis Mank - BC Genus HYPTILUS Paykul. H. concolor (LeConte) PQ Genus MICROTONUS LeConte M. sericans LeConte - '' ? - Genus SYMPHORA LeConte Trotomma LeConte S. flavicollis (Haldeman) MB ON PQ - NS S. rugosa (Haldeman) - ON PQ - NS Genus PROTHALPIA LeOonte P. holmbergi (Mannerheim)* AK BC fulrninans (LeConte) P. undata LeConte - ON PQ - NS NF Tlibe Serropalpini Genus AMBLYCTIS LeConte A. praeses LeConte PQ Genus XYLITA Paykull X. livida (C.R. Sahlberg)* _YK_BCAB_PQNBNS_LBNF X. laevigata (Hellenius)* -BCAB-MBONPQNB_NF decolorata (Randall) Genus SPILOTUS LeConte S. quadripustulatus (Melsheimer) PQ Genus RUSHIA Forel Carebara LeConte R. californica FaIl _BC californica Schaeffer schaefferi Csiki Genus SCOTOCHROIDES Mank S. antennatus Mank PQ Genus SCOTOCHROA LeConte Key to species: Mank (1939b) S. atra LeConte S. basalis LeConte S. buprestoides (Kirby) _-_:"AB__;.::il___Ni Genus ENCHODES LeConte E. sericea (Haldeman) _BCABSKMBONPQ 247 Mordellida: Genus SEF,ROPAIPUS Hellenius Key to North American species: Mank (1939a) S. coxalis Nlank _BC ON PQ NB NS NF S. obsoletur; Haldeman T_ _BC -PQ perfoueatus Mank S. substriatus Haldeman NT BC MB ON PQ NB LB NF Genus MYSTAXUS Kugelann M. simulator (Newman) ON PQ NB NS NF trifasc iatus (Melsheimer) Genus DIR,IA-EA Fabricius D. liturata '.LeConte) AB SK ON PQ NB quadrtmacularo (Say) D. horiei Hatch BC Genus PHLOEOTRYA Stephens P. vaudoueri Mulsantt PQ NS fusca (.LeConte) Genus ABDERA Stephens A. frrma Ha.tch BC Genus ANIIIOXYA Mulsant A. glaucula LeConte ON Subfamily OSPFIYINAE Tlibe Osphyini Genus OSPFfYA Illiger Nothut Olivier O. varians (LeConte) oN PQ Family MORDELLIDAE tumbling flower beetles J. McNamara Severrty species and subspecies of mordellids genera of the world; and Bright (1986) pr:Lblished a have beer recorded in Canada. The adults are small, catalog of the Nearctic species. Neverlheless, a humpbacked beetles, and their most distinctive fea- modern revision would be useful. Members of ture is their long, pointed abdomen. They are com- Anaspini, which used to be included within monly found on umbelliferous flowers, on which they Mordellidae, are now in the family Scraptirdae. feed. Larvae are carnivorous or feed on plant mate- rial. Sorne are found boring in rotten wood, others are leaf and stem miners, but very few are known. Liljetrlad (1945) revised the North American YK (1); NT (6); BC (18); AB (10); SK (12); MB (37); species oI'Mordellidae; Ermisch (1951) treated the ON (49);PQ (29);NB (14);NS (7);PE (1) Tribe Mordellini Genus TOMOXIA Costa T. borealis ( LeConte) T. inclusa Lr:Conte T. lineella LeConte T. serval (Sz,y) =_:"Y_;;lNifrilrs_ - T. triloba (Say) _ _ T. undulata (Melsheimer) Genus MORDELLA Linne Sphalera LeConte 248 Mordellidae M. albosuturalis Liljeblad BC M. atrata atrata Melsheimer NT BC AB SK MB ON PQ NB M. atrata lecontei Csiki -ONPQNB NS itorata LeConte M. brevistylis Liljeblad BC M. grandis Liljeblad BC M. hubbsi Liljeblad BC M. marginata lineata Melsheimer -o*--- M. marginata marginata Melsheimer _NT BC MB ON PQ NB NS M. melaena Germar _NT AB MB ON PQ M. quadripunctata (Say) BC AB SK MB ON Genus GLIPA LeConte G. bidentata (Say) _ONPQ G. octopunctata (Fabricius) -oN G. oculata (Say) -oN jouialis (LeConte) Genus CONALIA Mulsant & Rey C. melanops Ray ON PQ Tribe Mordellistenrnr Genus MORDELLISTENA Costa M. aemula LeConte _MB M. aethiops Smith sK MBoN -- M. andreae ancilla LeConte - Y - - - - - uarians LeConte M. andreae andreae LeConte ON grammica LeOonte M. andreae ustulata LeConte NT MB M. arida Leconte ON rutila Liljeblad M. aspersa (Melsheimer) M. bicinctella LeConte M. cervicalis LeConte _:'I_ illgN;: NB auricorna Helmuth '_ erilis Liljeblad picicornis LeConte M. convicta LeConte yuSNPQ NS M. discolor (Melsheimer) _ _ _ cinereofasciata Smith wolcotti Liljeblad M. divisa LeConte MB M. downesi Hatch BC M. errans Fall PQ M. frosti Liljeblad MB-PQ NB M. fulvicollis (Melsheimer) -oN M. fuscata (Melsheimer) _ONPQ M. fuscipennis (Melsheimer) -oNPQ NS PE pityptera LeConte M. gigas Liljeblad BC _ONPQ M. guttulata Helmuth -oN M. hebraica LeConte MB ON M. impatiens LeConte MB ON M. incommunis Liljeblad AB MB M. infima LeConte SK MB ON NB tiara Ray M. liljebladi Ermisch BC egregia Liljeblad M. limbalis (Melsheimer) SK MB ON PQ M. liturata (Melsheimer) ON _ft M. lutea (Melsheimer) ON M. marginalis (Say) _NT -MBONPQ NB M. morula LeConte BC SK M. nigricans Melsheimer _MBONPQ nigenima Helmuth M. nunenmacheri Liljeblad BC 249 Rhipiphoridae M. ornata (trfelsheimer) SK MB ON PQ decoreLa LeConte pectora,tis (LeConte) M. pallipes fimith MB ON M. picilabris Heimuth MB M. pustulatr,r (Melsheimer) -:-ecAB i* MB ON PQ M. quadrinobata Liljeblad -oNPQ M. scapulais (Say) _BCAB SK MB ON PQ NB NS biplagiuta Helmuth M. semiusta LeConte MB M. sericans .Fall SK MB M. sexnotata Dury -oN falli Cslki festiua Tall M. smithi Dury _MBON M. suturella Helmuth _MB M. syntaenia Liljeblad _MBNB M. testacea.Blatchley ON M. tosta LeConte _MBONPQNB M. trifasciata (Say) SK MB ON PQ NB amica L,eConte lepidula LeConte nigricollis Helmuth M. unicolor LeConte NT MB M. vilis (LeClonte) BC MB ON Genus GLIPOSTENODA Ermisch G. ambusta iLeConte) BC ON PQ diuersa (Ray) Family RHIPIPHORIDAE wedge-shaped beetles J.M. Campbell The family Rhipiphoridae contains about 400 Rhipiphoru.s, the largest North American genus of species, of which 10 are known to occur in Canada. the family, are poorly known and cannot be reliably Larvae of the subfamily Rhipiphorinae are parasitic determined. The records cited below, based on spec- on the larvae of bees and wasps; the frrst instars are imens identifred with the aid of Rivnay's revision, endoparasites and the last ones ectoparasites. are not all reliable. When revised, a number of addi- Adults are often found on flowers, where females tional species and provincial records will be lay their eggs. recorded. Rivnay (L929) revised the North American species o.l this family. However, the species of eC (3);AB (2); SK (2); ON (5);PQ (2) Subfamily PELECOTOMINAE Genus PELECOTOMA Fischer von Waldheim P. flavipes Melsheimer ON PQ Subfamily RHIPIPHORINAE Genus MACIIOSIAGON Hentz M. cruenturr (Germar) SK M. limbatun. (Fabricius) -o*- Genus RHIPIPHORUS Bosc d'Antic Dorthesia Say Myodes Latreille Myodites Latreille R. californicus (LeConte) R. columbiar:,us W.J. Brown R. fasciatus (Say) 250 Colydiidae R. mutchieri Rivnay R. nevadicus (LeConte) R. walshii (LeConte) R. zeschii (LeConte) Family COLYDIIDAE cylindrical bark beetles Y. Bousquet Only 21 species of colydiids are known to occur The North American species of colydiids are in Canada. They are usually found under bark, in usually divided into two subfamilies: Colydiinae, galleries of wood-boring larvae, and in fungi. which includes most of the species, and Adults and larvae of many species are predators of Bothriderinae. Stephan (1989) has recently revised wood-boring insects; others feed on rotten wood or the North American species of the family. fungr; larvae of some species are parasitic on vari- ous groups such as Scolytidae, Platypodidae, NT (2);sC (10); AB (3); SK (1); MB (1); ON (10);PQ Cerambycidae, and Buprestidae. (5); NB (1); NF (1) ' I (1) Subfamily COLYDIINAE T?ibe Synchitini Genus SYNCHITA Hellwig S. fuliginosa Melsheimer oN PQ nigripennis LeConte Genus ENDEITOMA Sharp E. granulata (Say) PQ floridana Casey Genus EUCICONES Sharp E. marginalis (Melsheimer) ON latus Casey Genus BITOMA Herbst Synchytodes Ctotch B. crenata (Fabricius)t B. ornata (LeConte) _ B. quadriguttata (Say) _y__:x;:__ __ trinotata Casey zimmer manni Gu6rin-M6neville Genus NAMUNARIA Reitter Coxelus Horn N. guttulatus (LeConte) ON N. pacificus (Horn) BC Genus LASCONOTUS LeConte L. borealis Horn NT _SK MB ON PQ NB_ NF L. complex LeConte -BC AB L. intricatus Kraus NT BC AB h.rausiHatc}r L. planipennis Kraus -BC L. tuberculatus Kraus _BC L. vegrandis Horn -BC apicalis Casey schwarzi lkans Tlibe Colydiini Genus AULONIIIM Erichson Key to North American species: Kingsolver & White (1967) A. longum LeOonte - BC 25t Zopheridae Genus COLYDIUM Fabricrus C. lineola Sry - BC - ON longiut,culum Say Tribe Pycnomerini Genus PYC NOMERUS Erichson Endeclus LeConte Penthetispa Pascoe P. reflexus tSay) - ON nitidus (LeConte) Subfamily BOTHRIDERINAE Tlibe Deretaphrini Genus DERETAPHRUS Newman D. oregonensis Horn - BC Tribe Teredini Genus OXYLAEMUS Erichson Redistes Casey O. californir us Crotch - BC Tribe Bothriderini Genus BOT :IRIDERES Erichson B. cryptus Stephan -oN B. geminatus (Say) -oN Family ZOPHERIDAE J.M. Campbell The f amily Zopheridae comprises about 125 Lawrence (1979). However, a revision of t,he North species, Subfamily ZOPHERINAE Genus PHEI,LOPSIS LeConte P. obcordata (Kirby) - BC AB ON PQ NB - NF porcata (LeConte) 252 Tenebrionidae Family TENEBRIONIDAE darkling beetles Y. Bousquet and J.M. Campbell The family Tenebrionidae comprises over 1200 lagrioid lineage, which is probably the sister group species in North America, but more than 8O7o of of the tenebrionids, has been given family status; them are restricted to the southwestern United these species lack defensive glands. Although the States. Only 135 species occur within the Canadian limits of the family are now fairly stable, its higher and Alaskan borders. Many tenebrionids live in classifrcation is still poorly understood. Numerous semiarid areas and deserts, but in Canada most changes have been proposed recently in the classifi- species are found in rotten wood, in fungr, under logs cation of the family (see Doyen and Tschinkel 1982), and stones, and under the bark of dead trees. A and more are expected; the alleculids, for example, number of species, such as those of Tenebrio, are probably polyphyletic in origin. Rolf L. Aalbu Gnatocerus, Tribolium, and Polorus, are stored- (personal communication) has provided the classifr- product pests, some of which are the most important cation used here. There is no recent revision of the in the country. Adults and larvae feed on plants or Nearctic Tenebrionidae, and most genera that occur plant material, including decaying vegetation, seeds, in Canada have not been revised. Identifrcation is roots, cereals and their products, and fungr. often difficult, especially in large groups like The concept of the family used for the checklist Eleodes. is similar to that of the tenebrionoid branch of Doyen and Tschinkel (1982). Adults of all species AK (3); YK (3); NT (8); BC (69); AB (32); SK (22); have paired defensive glands, with reservoirs open- MB (32); oN (70); PQ (63); NB (16);NS (1a); PE (5); ing between the seventh and eighth sternites. The LB (2); NF (7) . H (1);I (11) Subfamily PHRENAPATINAE Tribe Penetini Genus CLAMORIS des Gozis Phthora Mulsant C. americana (Horn) BC Subfamily TENTYRIINAE Tlibe Eurymetopini Genus STERIPHANUS Casey S. subopacus (Horn) alutaceus Casey Tribe Anepsiini Genus ANEPSIUS LeConte A. montanus Casey AB Tribe Coniontini Genus CONIONTIS Eschscholtz Coelotaxis Hotn Coniontellus Casey Coniontides Casey C. ovalis LeConte BC acerba Casey alutacea Casey anita Casey arida Casey breuiuscula Casey coruina Casey inepta Casey oblita Casey okanagani Boddy parilis Casey 253 Tenebrionldae punchtta Casey regulc,ris Casey sculptipennis Casey uteanu Casey uanco.zueri Casey weidtt Casey Tribe Coelini Genus COItrLUS Eschscholtz Coelotnorpha Casey Pseudocoelus Casey Keyto species: Doyen (1976) C. ciliatus Eschscholtz _BC ampli,:ollis Casey arenarius Casey curtulus Casey debilis Casey latus ()asey longuLus Casey obscurus Casey scolopax Casey sternalis Casey Tlibe Asidini Genus ASIDOPSIS Casey A. opaca tSayt AB A. polita (Sayt AB SK Genus MICjROSCHATIA Solier Acrosthatia WlTke Pycno;'totida Casey P. inaequa.is tl,eConte) - , diuersa (Casey) Genus GL\?IASIDA Casey G. sordida Leconre) AB Genus PEL,ECYPHORUS Solier P. densicollis Horn - BC corrosus Casey Subfamily DIAPERINAE Tribe Phaleriini Genus PHALEROMELA Reitter Nelite:t LeConte P. globosa (LeConte) _BC P. picta (Mrnnerheim) AK P. variegat r Triplehorn _BCAB picta (Horn) TYibe Nitionini Genus SCAPHIDEMA Redtenbacher Key to North American species: Triplehorn (1965) S. aeneolum (LeConte) - YK - BC AB - MB ON PQ NB LB Tlibe Rhipidandrini Genus RHIPIDANDRUS LeConte R. flabellicornis (Sturm) - ON PQ paradorus (Palisot de Beauvois) 254 Tenebrionidae Tlibe Bolitophagini Genus BOLITOTHERUS Candbze Phellidius LeConte B. cornutus (Panzer) _SKMBONPQNB bifurcus (Fabricius) Genus ELEATES Casey E. depressus (Randall) MB ON PQ NB tetraopes (Newman) E. explanatus Casey _BC Genus BOLITOPHAGUS Illiger B. corticola Say _ONPQ Tribe Diaperini Key to North American genera: Doyen (1984) Genus ADELINA Dejean Doliema Pascoe Schedarosus Reitter D. plana rFabricius) - ON depressa (Erichson) lecontei (I].orn) scidaria (Reitter) Genus CYNAEUS LeConte C. angustus tleConte) - BC AB SK MB ON PQ opoczs Champion Genus GNATOCERUS Thunberg CerandriaLrcas Echocerus Hotn Sicinus Champion (Subgenus GNATOCERUS s. str.) G. cornutus (Fabricius)f - BC MB ON PQ - NS maxillaris (PaIisot de Beauvois) Genus ALPHITOPHAGUS Stephens Phyletes Redtenbacher P hy lethus Redtenbacher Key to North American species: Triplehorn (1965) A. bifasciatus {Sayr - SK MB ON PQ Genus DIAPERIS O.F. Mriller Key to North American species: Triplehorn (1965) D. maculata Olivier MB ON PQ NB floridana Blatchley hydactina Fabricius hydni Fabricitts suturalis Chevrolat Genus NEOMIDA Latreille Arrhenoplita IGrby Euoplus LeConte Hoplocephala Castelnau & Brull6 Key to North American species: Triplehorn (1965) N. bicornis (Fabricius) - ON PQ - NS cornigera (Fabricius) g r acilis (Motschulsky) uirescens (Castelnau & Brull6) u ir idipennis (Fabricius) metallica (PaIisot de Beauvois) Genus PLATYDEMA Castelnau & Brull6 Histeropsis Chevrolat Key to North American species: Triplehorn (1965) 255 Tenebrionidae P. americanum Castelnau & Brull6 - BC AB - MB ON PQ NB P. excavatLm {Say) - t - ON PQ fraten;.utn Chevrolat nigrih".m Motschulsky paruulum Casey tubercllatum Castelnau & Brull6 P. neglectun Triplehorn - BC P. oregonense LeConte - BC P. ruficorne (Sturm) - ON PQ analeHald.eman opaculum Casey rufum (Melsheimer) rufiuen.tre Castelnau & Bru116 P. subcosta:um Castelnau & Brulle - ON PQ clypea,um Haldeman o b lo ng ulum Motschulsky P. teleops 'l riplehorn - ON PQ Genus PENTAPHYLLUS Dejean Key to North American species: Triplehorn (1965) P. pallidus LeConte - ON PQ Subfamilv HIPOPHLOEINAE Trib'e Hypophloeini Genus CORTICEUS Piller & Mitterpacher Hy pop,llaeus Fabricius Key to North American species: Triplehorn (1990) C. parallelr- s tMelsheimert - ON PQ C. praetern.issus (Fa11) AK YK NT BC AB SK MB ON PQ C. subopacr.s (Wallis) NT BC AB - MB ON - NF C. substrialus (LeConteJ - BC C. tenuis (t.eOonte) - BC AB ON PQ NB minor'Wallis) o c c ider: talis (Wallis ) C. thoracicr.s tMelsheimerJ - ON piliger (LeConte) Subfamilv OPATRINAE Tribe Eleodini Genus ELEODES Eschscholtz (Subgenus Iv{ELANELEODES Blaisdell) E. humeral.s LeConte - BC latiusczlus Walker E. obsoletus obsoletus (Say) NT - AB SK MB ON E. tricostatus (Say) NT - AB SK MB ? - alternatus Kitby planetus Solier: robustus LeOonte (Subgenus LITHELEODES Blaisdell) E. extricatus convexicollis AB SK Blaisdell E. extricatus extricatus (Say) NT BC AB SK E. parvulus Blaisdell - BC E. vandykei modificatus Blaisdell - BC (Subgenus PROMUS LeConte) E. opacus (Slayt NT - AB SK MB (Subgenus IILEODES s. str.) E. hispilabris connexus LeConte - BC binotatus Walker imitabilis Blaisdell immundus Blaisdell loeuis Illaisdell 256 Tenebrionidae E. hispilabris hispilabris (Say) - BC AB SK MB lecontei Gemminger sulcatus LeConte E. obscurus sulcipennis Mannerheim - BC aratus LeConte conjunctus Walker conuexicollis Walker (Subgenus BLAPYLIS Horn) (Am p hido ra E schscholtz ) (Eleodopsis Blaisdell) E. indentatus Blaisdell - BC E. novovermculus Boddy - BC AB E. nunenmacheri nunenmacheri - BC Blaisdell E. pimelioides Mannerheim - BC oregonus Blaisdell patruelis Blaisdell subligatus LeConte uerruculus Blaisdell uiator LeConte E. productus Mannerheim - BC uariolosus Blaisdell E. rolundipennis LeConte - BC strictus LeConte (Subgenus METABLAPYLIS Blaisdell) E. nigrinus LeConte - BC (Subgenus CRATIDUS LeConte) E. osculans (LeConte) - BC Genus EMBAPHION Say E. muricatum Say AB concduum LeConte Tlibe Opatrini Genus EPHALUS LeConte E. latimanus (LeConte) NS Genus BLAPSTINUS Dejean Aspidius Mulsant & Rey Heteropus Castelnau Lodinus Mulsant & Rey B. discolor Horn _BC oregonensis Casey B. intermixtus Casey _BC B. metallicus (Fabricius) -BCABSKMBONPQ_NS interruptus (Say) luridus Mulsant & Rev B. pratensis LeConte -oN B. pulverulentus Mannerheim - BC c alifor nicu s Motschulsky B. substriatus Charnpion NT BC AB SK MB ON PQ Genus GONOCEPHAIUM Chevrolat G. bilineatum Walkert _BC latifrons (LeConte) Tribe Lllomini Genus ULOMA Latreille U. impressa Melsheimer -oN U. longula LeOonte -BC 257 Tenebrionidae T?ibe Helopini Genus TAIiPELA Bates Larnperos AIIard T. micans (Fabricius) - ON PQ Genus HELOPS Fabricius H. californicus Mannerheim - BC H. convexulus LeConte - BC montanus LeConte H. perniter.rs LeConte - BC H. punctipennis LeConte - BC punctatus Gemminger H. regulus Blaisdell - BC AB Subfamilv TENEBRIONINAE rriue Tlibotiini Genus TRIBOLIUM Macleay Margu,s Dejean Key to species: Hinton (1948) (Subgenus TRIBOLIUM s. str.) T. audax Halstead - BC AB SK MB ON PQ T. castaneum (Herbst)* - BC AB SK MB ON PQ PE ferrugineum (Dejean) T. confusu:n Jacquelin du Val* - BC - SK MB ON PQ NF T. destruct:r Uyttenboogaart* - BC AB SK MB ON PQ - NS PE - NF T. madens Charpentiertt - NB NS (Subgenus APIIANOTUS LeConte) T. brevicor;re (LeConte.t - BC Genus BIL'S Mulsant B. estriatu,; (LeContet - BC AB - PQ Genus PAI,ORUS Mulsant Key to species: Halstead (1967) (Subgenus PALORUS s. str.) P. ratzeburgii (Wissman)t - BC - ON PQ - NS (Subgenus CIRCOMUS Fleischer) P. subdeprr:ssus (Wollastont* MB Genus LAT'HETICUS Waterhouse L. oryzae Waterhouse* MB - PQ NB Tribe Tenebrionini Genus ALF'HITOBIUS Stephens A. diaperinus (Panzer)f - BC AB SK MB ON PQ NB NS PE - NF A. laevigat.Ls Fabriciusi - BC - ON plcezs (Olivier) picipes (Panzer) Genus MEI|ACLISA Jacquelin du Val M. marginalis Horn - BC Genus CENTRONOPUS Solier M ene chide s Motschul sky Pyres Champion Scotobates Fiye Key to New World species: Spilman (1962) C. calcarat:s tFabricius) - ON PQ - NS aeneus (DeGeer) carolirt ensis (Palisot de Beauvois) reflexus (Say) 258 Tenebrionidae Genus TENEBRIO Linn6 T. molitor Linn6t BC AB SK MB ON PQ NB NS NF laticollis Stephens T. obscurus Fabriciust BC AB -oNPQ PE NF poliens Casey Genus NEATUS LeConte N. tenebrioides (Palisot de BC MB ON PQ Beauvois) badius (Say) picipes auct. amer. Genus IDIOBATES Casey D. castaneus (Knoch) ON PQ interstitialis (Say) Tlibe Amarygmini Genus MERACANTHAKiTbv Falacer Castelnau M. contracta (Palisot de Beauvois) ON canadensis Kirby cuprea (Castelnatt) inflata (Solier) (?) rugosa (Castelnau) (?) turnida (Melsheimer) Subfamily ALLE CLILINAE Tribe Alleculini Genus HYMENORUS Mulsant Hymenophorus Mulsant Key to North American species: Fall (1931) H. caurinus Fall BC H. discretus Casey -oN- H. megops Hatch BC H. molestus Fall _ONPQ NB H. niger (Melsheimer) -oNPQ NB H. obesus Casey _ONPQ H. picipennis Casey -oNPQ H. pilosus (Melsheimer) H. sinuatus Fall BC - :*PQ Tiibe Gonoderinr Genus ISOMIRA Mulsant Tedinus Casey Key to eastern North American species: Marshall (1970) I. comstocki Papp BC I. iowensis Casey - ow: I. oblongula Casey _ONPQ I. pulla (Melsheimer) -oNPQ ignora Blatc}:.ley I. quadristriata (Couper) MB ON PQ NB NS ueluntina LeConte I. sericea (Say) -oNPQ _NS tenebrosa Casey I. valida Schwarz -oN similis Blatchley Genus PSEUDOCISTELA Crotch Chromatia LeConte Cisfelo Fabricius Telacis Poey P. amoena (Say) -ONPQ P. brevis (Say) -ONPQ erythroptera (Ziegler) 259 Tenebrionilae P. pectinat.r Hopping - BC P. pinguis l.eConle - BC pacific:a Hopping Genus CAI'NOCHROA LeConte C. fuliginosa (Melsheimer) - ON PQ NB Genus ANI)ROCHIRUS LeConte A. erythropus (Kirbyt - ON PQ - NS fuscipes (Melsheimer) luteipes LeConte Tlibe Mycetocharini Genus MY(IETOCHARA Berthold M y ceto chare s Latreille M y cetocharis Gyllenhal M y cetct p h il a Gyllenhal Pterna Seidlitz Stigmotoma LeConte Key to North American species: Campbell (1978€) M. analis (LeConte) -BC-SKMBONPQ dauisi Hatch horni ))rry lugubris (LeConte) M. bicolor (Couper) -ONPQ M. binotata (Say) -oNPQ longula (LeConte) margiltata (LeConte) M. foveata .LeConte) -ONPQ gilipes Casey gracilis (LeConte) lecontti Borchmann tenuis (LeConle) M. fraterna (Say) MB ON PQ laticol is (LeConte) megahlps Casey nigerrtma Casey M. procera Casey _BC angusta Hatch caseyi Hatch crassu]ipes Casey downe'.Hatch malkh;iHatch neuadensis Casey Genus HYI\{ENOCHARA Campbell Key to species: Campbell (I978d) H. rufipes (LeConte) -oNPQ Subfamilv COELOMETOPINAE Tri6e Coelometopini Genus MEIIINUS LeConte M. laevis tCtliviert - ON PQ Genus XYLOPINUS LeConte T aenoltate s Motschulsky X. aenescens LeConte PQ X. saperdioides tOlivier) - ON PQ anthracinus (Knoch) calcaratus (Palisot de Beauvois) ruftpes (Say) spinipes (Fabricius) Genus HAILANDRUS LeConte H. concolor LeConte MB ON PQ 260 Lagriidae H. fulvipes (Herbst) oN PQ femoratus (Fabricius) Genus COELOCNEMIS Mannerheim Key to species: Doyen (1973) C. californica Mannerheim _BC AB barretti BlaisdelI bosolis Casey columbiana Casey dilat icoll is Mannerherm idahoensis Casey ouipennis Casey rauca Casey rugosa Linne\l spaldingi Casey utensis Casey Genus IPHTHIMINUS Spilman I. opacus (LeConte) _SX yu oN PQ NB NS I. salebrosus (Casey) -BC L serratus (Mannerheim) _BC AB serudtor (Walker) seruilis (Walker) subligatus (Walker) Genus AI-OBATES Motschulsky A. pensylvanicus (DeGeer) MB ON PQ chrysops (Herbst) inermis (Mannerheim) sublaeuis (Palisot de Beauvois) Genus UPIS Fabricius U. ceramboides (Linn6)x NTYKAK BC AB SK MB ON PQ NB PE LB NF reticulata (Say) uariolosa (DeGeer) Tribe Strongyliini Genus STRONGYLIUM Kirbv Reminius Casey Stenochia I{rtby Key to North American species: Triplehorn and Spilman (1973) S. tenuicolle (Say) ON PQ Family LAGRIIDAE lagriid beetles J.M. Campbell The family Lagriidae is primarily tropical in dis- lagriids as a distinct family until their relationship tribution and comprises over 1000 species around the with respect to other tenebrionoid groups has been world. Only six species have been recorded in Canada. better established. The classification of the North Adults of Statird and, Arthromacra are usually col- American lagriids was reviewed by Parsons (19750), lected by beating the foliage of trees; larvae are who includedAnaedus andParatenetus in the subfam- thought to breed in decaying wood or leaf litter. Little ily Lupropinae (tribe Heterotarsini of Tenebrionidae, is known about the biolory of the species. in most publications); this subfamily more properly The status of this family is uncertain. In recent should be named Goniaderinae. Doyen and Tschinkel studies of the higher classification of the (1982) have suggested that the genus Paratenetus be Tenebrionidae, Watt (t974), Doyen and Lawrence placed in or near the tenebrionid tribe Belopini. Until (1979), and Doyen and Tschinkel (1982) placed the the classification of the Lagriidae is reviewed, I have lagriids in Tenebrionidae, usually as a subfamily. chosen to follow the arrangement of Parsons (1975o). However, Doyen and Tschinkel (1982) suggested that the lagriids and related groups form the sister group to all other tenebrionids and possibly should be ttlT (1);sC (1);AB (1);MB (3); oN (5); PQ (5);NB (1); restored to family status. I prefer to recognize the NS (1) 26t Cephaloidae Subfamily GONIADERINAE Genus PAR.A.TENETUS Spinola Storthe,phora Mdklin P. crinitus lrall P. fuscus LeConte - - NrncAB- MB3N|8- - - - - P. punctatus Spinola -oNPQ Genus ANAEDUS Blanchard A. brunneur; (.Zieglet) ON Subfamily LAGRIINAE Genus ART.FIROMACRA Kirby Key to N,.'rth American species: Parsons (1975a) A. aenea (Say) MB ON PQ NB NS donacioides I\trby Subfamily STATIRINAE Genus STA IIRA Audinet-Serville Key to Ntrrth American species: Parsons (I975a) S. gagatina Melsheimer PQ respler,.dens Melsheimer Family CEPIIALOIDAE false long-hornedbeetles J.M. Campbell Cephaloidae is a small family of about 20 the genus Cephaloon. Subsequently, Arnett (1968) species rerstricted to the cooler parts of the Holarctic transferred the subfamilies Stenotrachelinae and region. l)nly nine species are known to occur in Nematoplinae from the melandryiids and pediliids, Canada ernd Alaska. Larvae are found in old, rot- respectively, to Cephaloidae. Both subfarrLilies have ting logs; adults of Cephaloon are often taken on been considered by some authors to be disbinct fam- flowers. ilies (Mamayev 1973), but I have followed Lawrence Species of this family are reasonably well (1982a) in accepting Arnett's treatment. known taxonomically. Hopping and Hopping (1934) and Arnett (1953) revised and reviewed, respec- AK (1); YK (1); NT (2); BC (5); AB (3); MB (1); ON tively, the species of the family, but included only (a); PQ (5);NB (1); PE (1);LB (2);NF (2) Subfamily CEPHALOINAE Genus CEPHALOON Newman Drachj,lis Casey Ichnodes Dejean Sponia',ium Casey Typitium Casey Key to species: Arnett (1953) C. bicolor Horn BC simulans (Casey) C. lepturoid.es Newman oN PQ uarians Haldeman C. pacifrcum Van Dyke -BC C. tenuicome LeConte _BCAB piceuri }Iorn ornatum Casey uersicolor Casey C. ungulare LeConte ON PQ PE LB NF 262 Meloidae Subfamily STENOTRA.CHELINAE Genus STENOTRACHELUS Latreille S. aeneus (Fabricius) AK YK NT BC AB MB ON PQ LB NF Genus ANELPISTUS Horn A. americanus Horn PQ A. canadensis Mank NT BC AB Subfamily NEMATOPLINAE Genus NEMATOPLUS LeConte N. collaris LeConte ON PQ NB Family MELOIDAE blister beetles J.M. Campbell Meloidae comprises over 3000 species, which cantharidin (Spanish fly), which causes blisters on are found in most regions of the world. However, contact with the skin and, if present in dried hay, the majority are restricted to warmer, drier areas. has been known to be fatal to horses. Cantharidin Only 49 species and subspecies are known to occur is also used for medical purposes. in Canada and Alaska, and most of these are found Most genera of North American Meloidae are in the semiarid areas of the Prairie Provinces and well known taxonomically. A few genera such as British Columbia. Larvae of Epicautini feed on Tricrania have not been revised, and the large, grasshopper eggs, and those of aII other Canadian important genus Epicauta is in need of additional blister beetles feed on eggs, larvae, and food study. The Canadian fauna is well known, and reserves of ground-nesting bees. Larvae of Meloini therefore few additional species are expected to be and Zonitinae are phoretic. The eggs are laid on added to the checklist in the future. Van Dyke flowers, the larvae hatch, and the frrst-instar larvae (1928a) and Arnett (1968) provide the most recent (called triungulin) attach themselves to bees that treatments of the family at the generic level. visit flowers and are transported to the bee's nest, where they drop off. Adults of Canadian meloids are phytophagous or feed on nectar; many are consid- aK (3); vK (1); NT (3); sC (25); AB (26); SK (23); ered economic pests. Adults of this family produce MB (20); ON (17);PQ (12);NB (3);NS (3) Subfamily MELOINAE Tribe Meloini Genus EURYMELOE Reitter E. aleuticus (Borchmann) AX Genus MELOE Linn6 Key to North American species: Pinto and Selander (1970) (Subgenus TREIODOUS Dugbs) M. barbarus LeConte BC (Subgenus MELOE s. str.) Melittophagus Kirby M. americanus Leach ON tnoerens LeConte temnus Dillon M. angusticollis Say BC AB SK MB ON PQ NS montanus LeConte rugipennis LeConte M. carbonaceus LeConte M. dianellus Pinto & Selander ---ncffisx-oNPe -NS M. impressus Kirby AK YK NT BC AB SK MB ON PQ NB occidentalis Van Dyke 263 Meloidae M. niger Kirby NTBCABSKMBONPQ_NS hirbyi Drllon opacus LeConte perplee:us LeConte M. occultus Pinto & Selander _BC M. strigulos,us Mannerheim AK BC M. vandykei Pinto & Selander - BC Tlibe Lyttini Subtribe Lyttina Genus LYTTA Fabricius Key to species: Selander (1960) (Subgenus PARALYTTA Selander) L. vulnerata tl,eContet - BC anguicollis (Dugbs) cooperi LeConte (Subgenus I'OREOSPASTA Horn) L. cyanipen:ris (Leconte) - BC AB solicis LeConte L. nuttalli Say AB SK MB fulgifera (LeConte) L. stygica (LeConte) - BC arbore<:.,. Wellman difficihs (FalI) dolosa LeConte neuadezsis Van Dyke purpurzscens (FaII) smarag'dula (LeConte) stolida (Fa11) L. viridana -eConte - BC AB SK MB (Subgenus I'OMPHOPOEA LeConte) L. sayi LeCr,nte - ON pe pyriuorz (Fitch) Subtribe Epicautina Genus EPICAUTA Dejean Key to North American species: Werner (1945) (see also Ballmer 1g79; pinto 1g80) (Subgenus EPICAUTA s.str.) (Anomelony c hus Saylor) (Gnathospasta }Jorn) (Henous Haldeman) (Maculicauta Dillon) (Nomaspis LeConte) (P le u rop om p ha LeOonte) E. atrata (Frrbricius) MB coracin,z (Illiger) trichrus (Pallas) E. cinerea (I'orster) - SK MB ON pe fissilabris (LeConte) E. fernrgine,r (Sayt AB SK MB ON E. immerita Walker - BC E. maculata (Say) - SK MB ON conspersa (LeConte) media I)ugds nogales Werner punctuata (Dugds) E. normalis Werner - BC AB SK E. oregona Florn - BC AB SK MB E. pennsylvcLnica (DeGeer) AB SK MB ON PQ NB morio (l,eConte) zigro (Woodhouse) E. piceiventris Maydell - BC AB SK MB elong att>calcarafo Maydell E. pruinosa LeConte - BC AB SK MB 264 Meloidae E. puncticollis Mannerheim - BC AB barberi Werner oblita LeConte E. sericans Leconte AB sK E. ventralis werner AB sK E. vittata (Fabriciust - ON PQ chapmani (Woodhouse) I e rnni scata (Fabricius) (Subgenus MACROBASIS LeConte) (Ap teros pasta LeConte) E. fabricii (LeContet AB SK MB oN PQ debilis (LeConte) E. Ilavocinerea (Blatchleyr MB ON E. murina (LeConte) - ON PQ NB unicolor Kirby E. subglabra rFall I - BC AB SK MB Genus LINSLEYA MacSwain Key to species: Selander (1955) (Subgenus LINSLEYINA Selander) L. sphaericollis {Say) - BC AB SK MB chalybea (LeConte) c haly b e ata ( Gemrninger) infidelis (Fall) Subfamily ZONITINAE T?ibe Zonitini Subtribe Nemognathina Genus NEMOGNATIIA Iiliger Key to North American species: Enns (1956) (Subgenus MEGANEMOGNATHA Enns) N. lutea dichroa LeConte - BC AB SK MB N. lutea lutea LeConte AB N. soror l\lacSwain - BC (Subgenus PAURONEMOGNATIIA Enns) N. nemorensis Henlz - ON b irnac ulata Melsheimer (Subgenus NEMOGNATHA s. str.) N. piazata bicolor LeOonte AB SK MB discolor LeConte stellaris Beauregard texana LeConte Genus TRICRANIA LeConte Tricraniodes Wellman T. mur"rayi LeConte - BC T. sanguinipennis {Say) - ON PQ T. stansburl (Haldeman) NT BC Genus HORNIA Riley H. minutipennis Riley AB Subtribe Zonitina Genus ZONITIS Fabricius Key to North American species: Enns (1956) (Subgenus NEOZONITIS Enns) Z. bilineata Say - ON PQ lineata Mels}reilner m an d ib u laris Mel sheimer Z. vermiculata Schaeffer - BC (Subgenus PARAZONITIS Enns) 265 Oedemeridae Z. sayi Wickham AB SK MB imrnactilata (Say) Genus GNAIHIUM Kirby Key to species: MacSwain (1952) G. minimurrL (Say) SKAB Family OEDEMERIDAE false blister beetles J.M. Campbell The family Oedemeridae comprises at least 100 species have been reported to cause blister:ing simi- genera arrd 1500 species, distributed in all faunal lar to that produced by some blister beetles,. regions. Thirteen species are known to occur in Arnett (1951) revised the North American Canada and Alaska. The biology of most species is species of oedemerids; little information has since unknown. Larvae of some species live in rotten been added to our knowledge of the Clanadian wood, and adults of most species are found on flow- fauna. ers, where they feed on pollen and nectar. One species, Nacerdes melanura (wharf borer), is an eco- AK(2); BC (10);AB (4); MB (3); ON (5); PQ (5);NB nomic pest of pilings and ship timber. A few exotic (1);NS (2);NF (1) ' I (1) Subfamily CALOPODINAE Genus CAJ-OPUS Fabricius C. ansustus LeConte BC AB oN PQ Subfamily NACERDINAE Genus XANTHOCHROA Schmidt X. testacea llorn BC Genus NACIIRDES Dejean Anogcotles Dejean N. melanura (Linn6)t BC MB ON PQ NB NS NF apicalis (Say) Subfamily OEDEMERINAE Tlibe Ditylini Genus DITYLUS Fischer von Waldheim D. caeruleus (Randall) _MB ONPQ_--__ D. gracilis L,:Conte AK BC uestitus LeConte D. quadricol)is LeConte AK BC AB consors LeConte Tribe Asclerini Genus XANIIHROCHROINA Ganglbauer X. bicolor (LerConte) BC Genus ASCL,ERA Stephens A. discolor L,eConte _BC A. excavata]-eConte -BCAB A. nigra LeConte A. puncticollis (Say) ---:t--^ras oN pq A. ruficollis (Say) ON PQ NS carinatct (Newman) Genus COPIDITA LeConte C. quadrimaculata LeConte BC 266 Mycteridae Family PROSTOMIDAE prostomid beetles J.M. Campbell Prostomidae is a small family that contains only In most North American literature Prostomis two genera and about 20 species. Most faunal arnericanus has been referred to as P. mandibularis regions, except South America, have a few species of (Fabricius), a European species. the family; only one species occurs in Canada. Nothing is known about the habitat of this species. BC (1) Genus PROSTOMIS Latreille P. americanus Crotch BC FamilySYNCHROIDAE synchroidbeetles L. LeSage As currently defined (Crowson 1966, Lawrence There is no recent treatment of the North 1982a), the family Synchroidae is reduced to two American species of this family. genera and 10 species, which were previously placed in the Melandryidae. Only one species of the genus Synchroa occurs in Canada. Both adults and Iarvae live under the bark ofold dead trees. uB (1);oN (1);PQ (rl Genus SYNCHROA Newman S. punctata Newman MB ON PQ nzurina Haldeman umbrina Melsheimer Family MYCTERIDAE mycterid beetles J.M. Campbell Mycteridae includes about 160 species around (Hatch 1965). I have followed Lawrence (1982o) in the world, of which four are known to occur in considering mycterids to be a distinct family. The Canada. Larvae live under bark, and adults are species are rather poorly known taxonomically, and often found on flowers. a modern revision is needed badly. Species of this family have been included in a variety of families, including Melandryidae (Leng L920), Salpingidae (Arnett 1968), and Pythidae sC (2); ON (2);PQ (2) Genus LACCONOTUS LeConte L. pinicola Horn BC L. punctatus LeConte oN PQ Genus MYCTERUS Clairville Key to North American species: Hopping (1935) M. concolor LeConte BC flauipennis Horn M. scaber Haldeman ON PQ 267 Pythidae Family BORIDAE J.M. Campbell Boridae contains three genera and six species; (Arnett 1968). Young (19856) transferred Lecontia only two species occur in Canada. Larvae of Boros from the family Pythidae to the Boridae. I have fol- occur under the bark of coniferous trees and nroba- lowed Lawrence (1982a) in recognizing Boridae as a bly feed on decayed cambium (Lawrence L9824). separate family. The species of Boridae have been placed in a variety of families such as Tenebrionidae (Leng NT (1); sC (1); AB (2); SK (2); MB (2); O.N (2); PQ 1920), P;'thidae (Hatch 1965), and Salpingidae (2);NB (1) Genus BOR()S Herbst B. unicolor Siay SK MB ON PQ Genus LECONTIA Champion Crymoctes LeConte L. discicollis (LeConte) NT BC AB SK MB ON PQ NB Family PYTHIDAE pythid beetles J.M. Campbell The family P1'thidae forms a small group com- (Arnett 1968) or, conversely, Pythidae has included prising orrly about 50 species around the world, of a number of taxa now transferred to other families which six occur in Canada and Alaska. Members of such as Salpingidae, Cononotidae, and Mycteridae. the subfermily Pythinae occur primarily in the The arrangement followed here is based on Holarctic region, whereas members of the other two Lawrence (1982a) with the exception of Lecontia, subfamilies are found in Central and South which has been transferred to Boridae as Young America, .New Zealand, and Australia. Both larvae suggests (1985b). Pollock is currently revising the and adults are found under bark ofconiferous trees. genus Pytho, so that most species of the family will Larvae fe':d on decaying inner bark (Pytho) or sap- soon be well known taxonomically. wood (Prirgnathus). Meml,ers of this family have previously been AK (3); YK (3); NT (3); BC (5); AB (4); SK (3); MB assigned to other families such as Salpingidae (a);ON (a);PQ (a);NB (3);NS (a); LB (1);NF (2) Subfamily PYTHINAE Genus PYTIIO Latreille P. americamrs Kirby AK YK NT BC AB SK MB ON PQ NB NS LB deplanatus Mannerheim P. niger Kirtry NT BC AB SK MB ON PQ NS NF P. seidlitzi Blair 5v MB ON PQ NS P. strictus L,-.Conte ItBCy NB Genus PRIOGNATHUS LeConte P. monilicornis (Randall) AK YK BC SK MB ON PQ NB NS NF sahl b e rg i (Mannerheim) Genus SPHI'LMA Horn S. quadricollis Horn BC 268 Pyrochroidae Family PYROCHROIDAE fire-colored beetles J.M. Campbell Twelve species of the family Pyrochroidae are Neopyrochroo and Schizotus ceruicalis are attracted known to occur in Canada and Alaska. Adult to cantharidin. pyrochroids are rarely collected, although adults of The family is well known taxonomically, thanks Dendroides and Neopyrochroo occasionally are to the revision and catalog of the North American attracted to light, and those of most species can be species by Young (1975 and 1983, respectively). collected in flight intercept traps or Malaise traps. However, in recent papers, Young has suggested Larvae of many species are common and are among that Ischalia be transferred to the Anthicidae the most frequently collected and distinctive species (1985o) and that the genus Pedilus be transferred of Coleoptera found under bark and in rotting wood. from Pedilidae to Pyrochroidae (1984). I prefer to Young (1975) summarized the feeding habits and postpone making these changes until his more biology of pyrochroids. Larvae live in dead wood detailed analysis of Pedilus is published. and are apparently fungivorous rather than preda- ceous or xylophagous, as reported by earlier authors. Adults rarely feed, but may be attracted at AK (2); NT (1); BC (5); AB (a); SK (1); MB (3); ON night to fermenting fruit or bait. Adults of (7);PQ (7); NB (3); NS (a);NF (1) Subfamily ISCIIALIINAE Genus ISCHALIA Pascoe (Subgenus EUPLEURIDA LeConte) L costata (LeConte) I. vancouverensis Harrington ecAB--9*PQ-TS--- Subfamily PYROCHROINAE Genus SCHIZOTUS Newman S. cervicalis Newman NT BC AB MB ON PQ NB NS S. fuscicollis (Dejean) AK Genus NEOPYROCHROA Blair N. femoralis (LeConte) -oNPQ N. flabellata (Fabricius) -ONPQ Genus DENDROIDES Latreille D. canadensis Latreille MB ON PQ NB NS blcolor (Newman) cyanipennis Latreilie D. concolor (Newman) _ONPQ NB NS NF D. ephemeroides (Mannerheim) BC AB D. marginatus Van Dyke T BC pacificus Barrett D. picipes Horn BC D. testaceus LeConte SKAB MB ON pe 269 Salpingidae Family SALPINGIDAE narrow-waistedbarkbeetles J.M. Campbell Salpingidae comprises 10 species in Canada and pingid genera were previously included in families Alaska and about 350 species around the world. such as the Pythidae and Colydiidae (Agleruus). The The adults are most often found under the bark of family is poorly known taxonomically, and a com- conifers, on flowers, or in detritus. Their feeding prehensive revision of most North American genera habits are unknown. is needed. Salpingidae, as currently constituted (Lawrence Ig82a), includes groups previously treated as dis- tinct families, such as the aegialitids, dacoderids, AK (3); YK (1); NT (1); BC (7); AB (1); SK (1); MB elacatidsr, eurystethids, inopeplids, othniids, and (1); oN (3); PQ (2); NB (1); NS (1); LB (1); NF (2) tretothoracids. On the other hand. some of the sal- . I(1) Subfamily SALPINGINAE Genus SPHAERIESTES Stephens Salpingus atct. S. alternatus (LeConte) BC S. virescens (LeCon!e) YK NT BC AB SX nrS ON pe NS - is Nr Genus VINCENZELLUS Reitter V. elongatus (Mannerheim) AI( Genus RHINOSIMUS Latreille Key to Ncrth American species: Blair (1932) R. viridiaerLeus Randall AK BC oN PQ NS a e n e ir nt r is Mannerheim nitens LeConte R. iecontei Blair -BC R. pallipes Boheman _BC Subfamily AE GIALITINAE Key to species: Spilman (1967) Genus AEC|IALITES Dejean Eurystethus Seidlitz A. californi,:us Motschulsky AK BC Subfamily OTHNIINAE Genus ELA.CATIS Pascoe Othnius LeOonte E. lugubris (Horn) BC Subfamily AGLENINAE Genus AGLENUS Erichson A. brunneus (Gyllenhal)t ON obsoler)us (Shuckard) 270 Anthicidae Family ANTHICIDAE antlike flower beetles Y. Bousquet Anthicidae comprises 55 species in Canada and and Anthicinae. Thanks to the work of Werner and Alaska. The adults are usually found under debris of Chandler, the taxonomy of most of the groups and in leaf litter on the ground or on flowers and that occur in North America has been worked out. foliage; the larvae live in the soil and in decayed vegetation. Apparently, members of many species are scavengers; those of some species are known to be egg predators. AK (2); YK (2); NT (5); BC (29); AB (25); SK (24); The North American species of this family are MB (19); ON (25); PQ (21); NB (9); NS (5); PE (1); usually grouped into two subfamilies, Notoxinae NF(3) .I(3) Subfamily NOTOXINAE TYibe Notoxini Genus MECYNOTARSUS LaFert6-S6nectbre Key to New World species: Chandler (1977) M. candidus LeConte AB Genus NOTOXUS Fabricius Ceratoderus Blanchard Monocerus Faldermann Key to North Arnerican species: Chandler (1982) N. anchora Hentz _NT_ABSK MB ON PQ PE N. bifasciatus (LeConte) MB-_ Iu_ N. cavicornis LeConte BC breuiusculus Fall obesulus Blaisdell N. desertus Casey ON constrictus Casey N. intermedius Fall AB SK paruidens Fa\l N. manitoba Chandler MB_ N. marginatus LeConte -AB N. montanus Casey _AB microcerus Casey similis Fall N. pictus Casey BC AB N. robustus Casey BC humboldti Casey N. schwarzi Horn BC N. serratus (LeConte) BC AB ix digitalis LeConte uandykei Blaisdell N. subtilis LeConte AB SK MB ON Subfamily ANTHICINAE TYibe Anthicini Genus AMBLYDERUS LaFert6-S6nectdre A. granularis LeConte ON PQ A. obesus Casey BC A. pallens LeConte BC AB SK MB ON PQ NB Genus ANTHICUS Paykull Nathicus Casey Key to North American species: Werner (1964) A. ancil)a Casey AB SK caseyi Pic parallelus Casey A. bellulus LeConte BC 277 Anthicidaer heluin.us Casey minusculus Champion A. biguttulus LeConte BC ouicol,'.is Casey protectus Casey A. cervinusi LaFert6-S6nectbre NT BC AB SK MB ON PQ bifasc;iatus Say b izonc'.t u s LaFert6-S6nectbre gilenst.s Casey seminotatus Casey termir:alis LeConte A. coracinu.s LeConte AK YK NT BC AB yu PQ NB NS A. cribratus LeConte BC :" 9* hecate Casey A. ephippium LaFert6-S6nectbre BC AB SK MB ON PQ confusus LeConte diffici;.is LeConte luteolus LeConte pinguescens Casey simiolus Casey A. flavicans LeConte _NTBCAB SK MB ON PQ NB NS A. haldemr.ni LeConte -NT_AB SK_ON PQ NE decorellus Casey q uadr iguttatus Haldeman sozcias Casey A. hastatus Casey BC AB SK MB solidus Casey subcati.uus Casey A. heroicus Casey PQ NB A. lecontei Champion --ec AB SK _ONPQ auriger Casey junctus Casey stellat,^ts Casey uulneratus Casey A. lutuient.rs Casey BC SKAB ON agilis Casey herifu.,lus Casey mund.ts Casey A. melanch olicus LaFert6-S6nectbre ON PQ facilis Casey latebntns LeConte spretus LeConte A. nanus L.:Conte BC amoenus Casey innocens Casey inscitt',s Casey nympltn Casey obliquus Casey peninsularis Casey uagan:; Casey A. nigritus Mannerheim AK YK NT A. praeceps Casey BC A. punctule.tus LeConte BC AB S^ l,fn - cephalotes Casey decrepitus Casey mercurialis Casey monticola Casey A. scabriceps LeConte AB SK MB ON PQ NB NS NF compositus Casey Genus SAPINTUS Casey Key to Ncrth American species: Werner (1962) S. fulvipes l.LaFert6-56nectdre) BC _MBONPQ colonus Casey rustict:s Casey S. lemniscatus Werner ON S. pubescer.Ls (LaFert6-S6nectdre) oN PQ S. pusillus (LaFert6-S6nectbre) _MBONPQ_NS binomi.natus Pic festinans Casey mollis Casey 272 Anthicidae Genus STRICTICOMUS Pic S. tobias (Marseul)i PQ Genus OMONADUS Mulsant & Rey Hemantus Casey Key to North American species: Werner (1964) O. floralis (Linn6)t - BC AB SK MB ON PQ NB - NF (?) basillaris (Say) O. formicarius (Goeze)f - BC AB SK MB ON PQ NB enodis (Casey) rixator (Casey scenicus (Casey) Genus MALPORUS Casey M. cinctus tsay) - SK - ON M. formicarius (LaFert6-S6nectbre) - SK MB ON PQ NB NS Genus THICANUS Casey T. mimus Casey - BC AB SK MB Genus EIJVACUSUS Casey E. coloradanus Casey - BC AB SK MB Genus VACUSUS Casey Key to species: Werner (1961) V. formicetorum (Wasmann) AB SK desertorum Casey prominens Casey suspectus Casey V. nigritulus (LeConte) - BC arcanus Casey Genus ISCHYROPAIPUS LaFert6-S6nectbre Lappus Casey Key to North American species: Werner (1973) L cochisei Werner - BC L nitidulus (LeConte) - BC - ON PQ alacer (Casey) asperulus (Casey) cursor (Casey) nubilatus (Casey soliuagans (Casey) uigilans (Casey) I. obscurus (LaFert6-s6nectdre) - ON PQ I. sturmi (LaFert6-S6nectbre) - ON e le g ans (LaFert6-S6nectbre) I. turgidicollis (Casey) - BC animatus (Casey) Genus TANARTHRUS LeOonte Key to species: Chandler (1975) (Subgenus TANARTHROPSIS Casey) T. salicola LeConte - SK densus Casey mormon Wickham nubifer Casey tricolor Casey uirginalis Wickham 273 Pedilidae Family PEDILIDAE pedilid beetles Y. Bousquet Pedilidae currently includes 19 species in Eurygeniinae. Young (1984) suggestecl that the Canada; none has been reported from Alaska. The genus Pedilus be transferred from Pedilidae to adults usually occur on flowers or foliage and feed Pyrochroidae and placed in its own subfamily on flowers, pollen, and nectar; they are attracted to (Pedilinae) and that other North American pedilids light at night. The larvae live in decaying plant be tranferred to Anthicidae. I prefer to wait until matter, uLsually in wet places. Young's revision of the genus Pedilus is published The group is probably closely related to before incorporating these changes. Anthicidae, and some authors, including Crowson (1955) and Lawrence (1982o), have included the pedilids in Anthicidae. Members of Pedilidae are BC (8); AB (4); SK (1); MB (2); ON (6); PQ (e); NB usually classified in two subfamilies, Pedilinae and (1); NS (1) Subfamily PEDILINAE Genus PEDILUS Fischer von Waldheim Corphyra Say P. abnormis (Horn) _?AB-_ P. canaliculatus (LeConte) pe----- P. cyanipen.nis Bland PQ P. elegans (Hentz) MB ON PQ fuluipe's Newman h a e m a t ho i dal i s Zi e gler lugubris Newman newmani LeConte ruficollis Ziegler P. flabellat:rs (Horn) BC P. impressrLs (Say) ON PQ P. inconspicuus (Horn) BC P. longilobrs Fall P. lugubris (Say) _ _ _ f _ _ ON pe NB NS imzs llewman inornatus (Randall) infumtr.tus (Hentz) nigricctns Ziegler rufithorax Newman P. oregonus Fall _BC P. punctulatus LeConte _BC funebris (Hotn) P. terminalis (Say) ON PQ guttulu Newman Subfarnily EURYGENIINAE Tlibe Eurygeniini Genus PEBGETUS Casey P. campanulatus (LeOonte) BC Genus STEREOPAJ-PUS LaFert6-S6nectdre Key to species: Abdullah (1965) S. columbianus Hopping BC S. guttatus LeConte BC AB impressicollis Casey subalbicans Casey S. hirtus Hatch S. rufipes C:asey __BC_-_-_ po S. vestitus Say) SK MB ON PQ bad,iipennis LeOonte 274 Scraptiidae Tribe Macratriini Genus MACRATRIA Newman M. confusa LeConte ON PQ Family ADERIDAE antlike leaf beetles Y. Bousquet Aderidae is a small family that includes nine Except for Werner's (1990) recent review of the species within the borders of Canada. Very little is eastern North American species, members of this known about these beetles. The adults are usually family has been little studied taxonomically. found on foliage and flowers; the larvae of some species have been found in rotten wood and leaf litter. gC (1); ON (8); PQ (8); NB (1); NS (1) ' I (1) Genus ELONUS Casey E. basalis (Leconte) - ON PQ princeps Casey E. nebulosus (LeConter - ON PQ Genus ADERUS Westwood Phomalus Casey A. populneus (Panzertt - BC - ON PQ Genus VANONUS Casey Tanilotes Casey V. calvescens Casey PQ V. piceus (LeConte) - ON congener Casey fusciceps Casey t ub er c ulifer ( Hamilton) V. vigilans Casey - ON PQ V. wickhami Casey - ON PQ NB NS lacustris (Casey) Genus ZONANTES Casey Z. fasciatus (Melsheimert - ON PQ tricuspis Casey Z. pallidus Werner - ON PQ Family SCRAPTIIDAE scraptiid beetles L. LeSage The family Scraptiidae, which comprises about There is no recent treatment of the family in 400 species around the world and 21 in Canada and North America; the Canadian fauna is poorly Alaska, represents a group of beetles with uncertain known except in the provinces of Quebec and affinities. Previously placed in Mordellidae, or British Columbia. Young (1976) transferred the Melandryidae, or Anthicidae, they are now consid- genus Pectotom.a, previously included in ered to be a separate farnily closely related to those Tetratomidae, to Scraptiini. families (Crowson 1955; Franciscolo 1963; Lawrence I982a). Adults are often found in large numbers on flowers, and larvae occur in decaying wood or in leaf AK (2); yK (1); gC (16); AB (5); SK (1); MB (1); ON litter. (8);PQ (10); NB (3);NS (2); NF (3) 275 Scraptiidac Subfamily ANASPIDINAE Tribe Anaspidini Key to North American species: Liljeblad (1945) Genus ANIiSPIS Geo{Iroy A. atrata Champion YK BC AB atra LzConte A. duryi Liljeblad BC A. flavipennis Haldeman BC ON PQ NB A. nigrina Csiki BC ON PQ NF zlgro (Haldeman) A. rayi Hat,ch BC A. rufa Say AK BC AB ON PQ NB lls - NF dimidiata Melsheimer filiforrnis LeConte nigrice:ps LeConte pallescens Mannerherm uenlro'is Melsheimer A. seposita Liljeblad _BC A. sericea ( \{annerheim) AK BC luteipennis LeConte Genus SILARIA Mulsant S. revelstokei Liljeblad BC Genus LARISIA Emery L. nigricolo r Liljeblad BC AB ON Genus NASSIPA Emery N. hopping Liljeblad BC Subfamily SCRAPTIINAE Tlibe Allopodini Genus ALLOPODA LeOonte A. californir:a Schaeffer BC A. lutea (Haldeman) fuscos ut ur alis (Blatchley) Tlibe Scraptiini Genus SCRAPTIA Latreille S. sericea t lylelsheimer) PQ Genus CAI IFA LeConte C. pallipennis LeConte PQ C. pailipes' Melsheimert BC MB ON PQ NS NF biimpressa (Haldeman) C. plagiata ( Melsheimert ON PQ americana (Haldeman) C. pusilla (I{aldeman) ON PQ NB Genus PECTOTOMA Hatch P. hoppingi Hatch BC AB T?ibe Pentariini Genus PENTARIA Mulsant Anthobates LeOonte Anthobatula Sftalrd. Key to North American species: Liljeblad (1945) P. fuscula (l,eConte) BC PQ P. trifasciata Melsheimer BC AB SK -ONPQ nubila (LeConte) 276 Cerambycidae Superfamily Chrysomeloidea Family CERAMBYCIDAE longhorned beetles J. McNamara Cerambycidae is represented in Canada and in forest ecolory because they help in the decompo- Alaska by 354 species and subspecies; it is one of sition ofdead and dying trees. the largest groups of wood-boring insects. These Thanks to the works of Linsley (1962a, 1962b, beetles have always been popular with collectors 1963, 1964) and Linsley and Chemsak (1972, 1976, because of their bright colors and the large size of 1985), the North American fauna is taxonomically adults of many species. The adults generally feed well known; only the second part of the subfamily on wood, roots, leaves, and pollen. Flower-feeding Lamiinae remains to be done. Chemsak and Linsley adults may visit many different blossoms, but some (1982) also published a checklist of the North are host-specific; they are important plant-pollina- American cerambycids. tion agents. The larvae bore into wood and roots; the host wood may be living or thoroughly decayed. AK (31); YK(27); NT (32); BC (145); AB (88); SK Some may even totally destroy timber for commer- (56); MB (100); ON (20e); PQ (178); NB (68); NS cial purposes. Cerambycids play an important role (51);PE (11);LB (13);NF (24) . H (4);I (1) Subfamily PARANDRINAE Tribe Parandrini Genus PARANDRA Latreille (Subgenus NEANDRA Lameere) P. brunnea brunnea (Fabricius) oN PQ ampliceps Casey conformis J. Thomson dentata J. Thomson grauidula Casey minuta J. Thomson purpureus (Herbst) q uad.r icoll is J. Thomson sayi J. Thomson P. brunnea quebecensis Abdullah PQ Subfamily PRIONINAE Tribe Ergatini Genus ERGATES Audinet-Serville Trichocnemis LeConte E. spiculatus spiculatus (LeConte) - BC californicus (A. White) spiculigerus (A. White) Tlibe Prionini Genus ORTHOSOMA Audinet-Serville O. brunneum (Forster) oN PQ NS amplians Casey cy lindr icum (Fabricius ) cylindroides (Gmelin) pennsyluanicum (DeGeer) spadix Casey unicolor (Drury) Genus PRIONUS Geoffroy (Subgenus PRIONUS s. str.) (Prionellus Casey) P. californicus Motschulsky AK BC acomanus Casey alutaceus Casey 277 Cerambycidee ambiguus Casey angustulus Casey coloradensis Casey compar Casey consors Casey crassict>rnis LeConte curuatr^.s LeConte fissifrons Casey horni l.ameere humerc:lis Casey ineptus Casey nanus i)asey orbiceps Casey ouipenr:is Casey paruicollis Casey proxim,zns Casey punctui',atus Casey scutellaris Casey seniger Casey solidus Casey spaldingi Casey spiculosus Casey stultus Casey suspectus Casey terminulis Casey texanus: Casey uintanus Casey P. laticollis Druryt oN PQ b reu ico." nis Fabricius densus Casey frosti C'.asey hempi Casey nigrescans Casey oblongu.s Casey paruus Casey P. lecontei I,ameere BC P. pocularis Dalman oll pe - bicolor Casey curticornis LeConte laeuigatus Hanis obliqui:ornis LeConte prolixus Casey ualidiceps Casey (Subgenus I{EOPOLYARTHRON Semenov) (Riporuts Casey) P. imbricorrris (Linn6) ON brunneus Casey cuneatus Casey diuersus Casey mimus Casey robustu,s Casey (Subgenus III.{TENNAJ,IA Casey) P. fissicorni s Haldeman ON paruiceps Casey thoracicus Casey transuersus Casey Tribe Tragosomini Genus TRATIOSOMA Audinet-Serville T. depsariurn (Linn6)* NT BC SK MB ON PQ NB NS NF harisii LeConte paruicc'lle Casey repens Casey sodale Casey spiculu.tn Casey 278 Cerambycidae Subfamily SPONDYLINAE Genus SPONDYLIS Fabricius S. upiformis Mannerheim AK BC AB ON PQ basalis Casey collaris Casey laticeps LeConte parua Casey robustula Casey subpubescens Casey Subfamily ASEMINAE TYibe Asemini Genus ARHOPAJ-US Audinet-Serville Criocephalum Dejean Criocephalus Mulsant A. asperatus (LeConte) NT BC AB SK filitarsis (Casey) A. foveicollis (Haldeman) AK YK NT AB SK MB ON PQ NB NS LB NF agrestis (Kirby) ceruinus (Casey) lacustrinus (Casey) A. productus (LeConte) BC AB elongatus (Casey) propinquus (Casey) A. rusticus obsoletus (Randall) ON mexicanus (J. Thomson) obscurus (LeConte) Genus MEGASEMUM Kraatz M. asperum (LeConte) BC AB impressum (Casey) rectipennis (Casey) uteanum (Casey) Genus ASEMUM Eschscholtz Liasetnum Casey A. australe LeConte PQ c ham p lainianum (C asey) A. caseyi Linsley BC A. nitidum LeConte BC mokelumne Casey A. striatum (Linn6)8 NTYK BC AB SK MB ON PQ NB NF amputatum Casey atrum Eschscholtz breuicorne Casey brunneum Haldeman carolinum Casey costulatum Casey curtipenne Casey ebenutn Casey fuluipenne Casey fuscutn Haldeman gracilicorne Casey juuencum Haldeman moestum Haldeman obsoletum Haldeman paruicorne Casey pugetanum Casey stoctonense Casey substriatum Haldeman Genus TETROPIUM Kirby T. abietis FaIl -BC T. cinnamopterum Kirby AKYKNTBCAB_MB ON PQ NS-- NF T. parvulum Casey AKYKNTBCAB_MB ON PQ LB alaskanumFall 279 Cerambycii.ae T. schwarzilnum Casey T. velutinurn LeConte ---ncAB--oNPQ----y boreak Casey columlianum Casey hexagonum Casey Tlibe Atimiini Genus ATItr{IA Haldeman A. confusa confusa (Say) - ON frislls l{aldeman A. confusa corsalis LeConte - BC A. hoppingi Linsley - BC Subfamily CERAMBYCINAE Tribe Opsimini Genus OPSIMUS Mannerheim O. quadrilineatus Mannerheim AK BC biplectralis Casey Genus DICI)NTRUS LeConte Paraopsimus Champlain & Knull D. bluthner LeConte - BC TYibe Smodicini Genus SMODICIIM Haldeman S. cucujiforrre (Say) - ON conuergens Casey cy lindr;;d.e s (Newrnan) Tlibe Hesperophanini Genus TYLONOTUS Haldeman T. bimacular,us Haldeman MB ON PQ Genus HESIIEROPHANES Mulsant AnoplitLm Haldeman H. pubescens {Haldeman) - ON PQ Genus EBU:RIA Audinet-Serville E. quadrigerninata (Say) - ON Tribe Elaphidiini Genus PSYI|,ASSA Pascoe PseudiL,idion Casey P. unicolor (.Randallt - ON PQ - NS p loch ionoc era (Haldeman) Genus STEIIOSPHENUS Haldeman S. notatus (Olivierr - ON pQ deflend,zs (Newman) discicol,:!.is (Dej ean) discoiderus (Sturm) pinorunz Casey Genus ENAIIIIALODES Haldeman Hy pe rm a ll us Lacordaire Romaleum A. White Thersalus Pascoe 280 Cerambycidae E. atomarius (Drury) ON bispinosus (Pascoe) mary landicus (Fabricius) multicolor (Gmelin) p ulu er ule ntus (DeGeer) thoracicurn (Casey) uariegatus (Goeze) E. cortiphagus (Craigheac, ON E. rufulus (Haldeman) ON PQ crinicornis (Casey) mancus (Casey) Genus PARELAPHIDION Skiles Key to species: Skiles (1985) P. incertum (Newman) ON breuicorne (Casey) neglectum (LeConte) obsolescens (Casey) Genus ANELAPIIUS Linsley E I ap hidionoides Linsley A. parallelus (Newman) MB ON PQ orclzs (Newman) canadensis (Casey) defectus (Casey) densicollis (Casey) medialis (Casey) molliculus (Casey) oblitus (LeConte) p ulu erulentus (Olivier) A. villosus (Fabricius) MB ON PQ abruptus (Casey) breuiusculus (Casey) lacustris (Casey) militaris (Casey) p r uinos u s (Gu6rin-M6neville) putator (Peck) scuticularis (Casey) Genus ELAPHIDION Audinet-Serville Cycliopleurus Hope E. mucronatum (Say) ON muriaticum (Haldeman) muricatum Haldeman neb ulosum Gu6rin-M6neville uestitum Haldeman Tlibe Ibidiini Genus HETERACHTHES Newman Heteracthon J. Thomson Key to species: Martins (1970) H. quadrimaculatus Haldeman - ON PQ euanescens (J. Thomson) fluuiatilis Casey pallidus Haldeman Tlibe Obriini Genus OBRIUM Dejean Diozodes Haldeman Phyton Newman O. maculatum (Olivier) howdeniRnrTl limum (Newman) pallidum (Say) O. rufulum Gahan MB ON PQ 28r Cerambycide.e Tlibe Hyboderini Genus HYBODERA LeOonte H. tuberculaua LeConte _BC Genus LAMPROPTERUS Mulsant Callitne.llum Sftand Callitnus Mulsant Pilema.LeConte L. ruficollis (LeConte) _BC longicol'.is (Casey) opacipennis (Casey) Tfibe Eumichthinr Genus EUMI.CHTHUS LeConte E. oedipus LcConte - BC ater Hardy & Preece funebrus (Van Dyke) malkini (Fisher) ruber H,*dy & Preece Genus POECIILOBRIUM Horn P. chalybeunr (LeConter - BC gibsoni [Iopping minuturn Casey rugosipt:nne Linell Tlibe Molorchini Genus MOLORCIIUS Fabricius Glaphyra Newman Laphyrt,. Newman M. bimaculal;us bimaculatus Say AB SK MB ON PQ affinis l.eCorrte obscuru:; (LeConte) M. longicollis LeConte - BC Tribe Stenopterini Genus CALLIMOXYS Kraatz C. sanguinicollis luscipennis - BC (LeCcnte) C. sanguinic,rllis sanguinicollis - ON PQ (Olivier) pinorurr;, Casey Tlibe Rosaliini Genus ROSALIA Audinet-Serville R. funebris l\lotschulsky AK BC acorrrano, Casey mandibitlaris Casey Tlibe Callidiini Genus PRONOCERA Motschulskv Gonocallus LeConte P. coilaris coilaris (Kirby) AK YK NT BC AB SK MB ON PQ NB - NF lepida (LeConte) Genus HYLOTRUPES Audinet-Serville 282 Cerambycidae H. bajulus (Linn6)t PQ bullatus Haldeman inaequalis Casey incertus Casey latus Casey minutus Casey nematocerus Casey pedicellatus Casey puncticollis Casey Genus SEMANOTUS Mulsant Anocomis Casey Hemicallidium Casey S. amethystinus (Leconte) _BC S. ligneus amplus (Casey) AK BC angustus (Casey) obliuiosus (Casey) terracensis (Casey) S. ligneus basalis (Casey) BC thujaeYanDyke S. ligneus ligneus (Fabricius) MB ON PQ NS nicolas (A. White) russicus (Stephens) S. litigiosus (Casey) NT BC AB SK MB ON PQ NB canadensis (Casey) terminatus (Casey) Genus MERIELLUM Linsley Merium LeConte M. proteus (Kirby) AK YK NT BC AB SK MB ON PQ bifossulatum (Casey) simile (I{trby) Genus CALLIDIUM Mulsant Meridion des Gozis C. antennatum hesperum Casey -BC C. cicatricosum Mannerheim AKYK_BCAB subopacum Swaine udncouuerense Van Dyke C. frigidum Casey oN PQ lacustre Casey C. hoppingi Linsley C. schotti Schaeffer C. violaceum (Linn6)t NN NS NF janthinum LeConte Genus PHYMATODES Mulsant Microcallidium Casey Phymatod,ina Casey Poecilium Fairmaire P. aeneus LeConte BC P. aereus (Newman) oN pe pallipes (Haldeman) P. amoenus (Say) oN PQ coccineico llis (Haldeman) hudsonicus Casey P. ater LeConte _NS P. decussatus decussatus (LeConte) BC collaris Casey harfordi Casey latifasciatus Hardy & Preece obliquus Casey P. dimidiatus (Kirby) AK YK BC AB SK MB ON PQ NB haltnii (Schon]nen) manner h eimii (LeConte) palliatus (Haldeman) P. fulgidus Hopping _BCPQ P. hirtellus densipennis Casey _BCAB P. lecontei Linsley _BC 283 Cerambycidae grandi; Casey obscur,ts (LeConte) P. maculico.lis LeConte - YK - BC PQ NB elonga;:us Hopping hard,yi (YanDyke) P. nitidus f.eConte - BC crucialis Casey elegant Casey exilis Qasey P. testaceus tl-innerf - ON PQ uariabilis (Linn6) uentralis Haldeman P. varjus tFabricius) - ON PQ albofasciatus (Bland) diffidens Casey latipennis Casey s emic ir c ular is (Blano.r subfast:iellus (A. White) uersicoJor (Gmelin) P. vulneratrrs (LeConte) - BC concinnus Casey nigrescens Hardy & Preece Genus PHYSOCNEMUM Haldeman Dulari,ts J. Thomson P. andreae tHaldeman) - ON luscum (J. Thomson) P. breviline,rm t Sayt MB ON PQ angulatum Casey compressipes Casey densun't Casey longitarse Casey pictum Casey P. violaceip'rnne HamilLon Genus ROP AJ-OPUS Mulsant Euryotr tera Horn R. sanguini.:ollis (Horn r MB ON PQ NB Genus ELA'IOTRYPES Fisher E. hoferi Fir,her AB Genus XYLTJCRIUS LeOonte X. agassizi {Leconter - BC Tribe Clytini Genus MEC'.ACYLLENE Casey Cyllene Newman M. angulifera (Casey) AB SK MB M. caryae (Gahan) -oN M. decora (Olivier) AB_MB cas teln,zui (Aurivillius) chara (Say) genitiuo (Casey) horni (.A,urivillius) infausta (LeConte) hansana (Casey) mediana (Aurivillius) solida (Casey) M. robiniae (Forster) _ONPQ angulata (Fabricius) flexuos'z (Fabricius) picta (Drtry) Genus GLYOOBIUS LeConte G. speciosus (Say) _ONPQ haiji (Gtay) 284 Cerambycidae Genus CAI-LOIDES LeConte C. nobilis nobilis (Harris) MB ON PQ Genus SAROSESTHES J. Thomson S. fulminans (Fabricius) ON PQ Genus XYLOTRECHUS Chevrolat X. aceris Fisher _ONPQ X. annosus annosus (Say) NT AB SK MB ON PQ NB X. annosus emotus W.J. Brown -BC X. colonus (Fabricius) MB ON PQ NB agrestis (LeConte) campestris (OIivier) te rrnina n s (Fabricius ) upsilon (Stephens) uillicus (Olivier) X. integer (Haldeman) ON PQ NB NS PE frosli Van Dyke X. lengi Schaeffer _BC X. longitarsis Casey NT BC AB columbianus Casey inflaticollis Casey montanicus Casey X. mormonus (LeConte) YK BC AB SK MB X. quadrimaculatus (Haldeman) MB ON PQ NB NS--- X. sagittatus sagittatus (Germar) MB ON PQ cylindrus Casey pubescens (Haldeman) salebrosus Casey X. schaefferi Schott X. undulatus (Say) AK Yr( Nr ec AB sx MB 3N FB - Ns pn is Nr' fuscus (Kirby) interruptus (Castelnau & Go.y) lunulatus (Kirby) soyi (Castelnau & Gory) undatus (Kirby) Genus NEOCLYIUS J. Thomson R ho p alop achys Chevrolat R ho p alome ru s Chevrolat N. acuminatus acuminatus (Fabricius) MB ON PQ NB americanus (Gmelin) aspericollis (Germar) erythrocep halus (Fabricius) N. balteatus LeOonte BC N. caprea (Say) ON PQ eleuatus (Castelnau & Go.y) gibbicollis (Castelnau & Gory) N. conjunctus (LeConte) BC occiduus Casey tularensis Casey N. fulguratus Casey confusus Van Dyke N. mucronatus mucronatus ON (Fabricius) humeralis (Newman) luscus (Fabricius) tnaculosus (Gmelin) strictulus Casey N. muricatulus muricatulus (Kirby) AK YK NT BC AB SK MB ON PQ NB PENS NFLB &lr6yi Aurivillius leucozonus (Castelnau & Gory) longipes (Kirby) N. provoanus Casey BC AB basalis Van Dyke basillaris Van Dyke Genus CLYTUS Laicharting C. canadensis Hopping BC 285 Cerambycirlae C. margini<:ollis Castelnau & Gory PQ NB NS carolinensis Casey uespoides Haldeman C. pacificus {Van Dyke) - BC C. planifrons (LeConte.t - BC AB C. ruricola (Olivier) AB SK MB ON PQ NB catulus (Casey) hammatus Say Genus CLYTOLEPTUS Casey C. albofasciatus (Castelnau & Gory) - ON PQ gracilicornis Casey frisfis Casev Tribe Tillomorphini Genus EUDERCES LeConte Apelocera Chevrolat Eplophorus Chevrolat E. picipes picipes (Fabricius) -oNPQ Tlibe Anaglyptini Genus TILI,OCLYTUS Bates T. geminat rs (Haldeman) - ON duplicztus (LeConte) Genus CYI';TOPHORUS LeConte C. verrucosus (Olivier) AB SK MB ON PQ - NS Genus MICROCLYTUS LeConte M. compre-.sicollis (Castelnau - ON PQ & Gory) frosti Oasey gibbulus (LeConte) insinuans (Casey) niger (LeConte) M. gazellula (Haldeman) - ON Tlibe Purpuricenini Genus PUI|PURICENUS Dejean P. humeralis lFabricius.t MB ON PQ Genus KNIILLIANA Linslev Chion Newman K. cincta cincta tDrury) - ON PQ baltea;la (DeGeer) filicontis (Casey) garganica (Fabricius) q uad r ii.sp inosa (Haldeman) ruricoia (Gmelin) rzsficcr (Fabricius) Genus TRAGIDION Audinet-Serville Trichascelis Bates T. coquus ( Linn6t apicat.um Casey filiconte Casey fuluipenne (Say) leuipes Casey lynceum (Fabricius) melsheimeri (Germar) nubiferum Casey 286 Cerambycidae Genus BATYLE J. Thomson Batyleoma Casey B. suturaiis suturalis (Say) SKAB ON PQ fontinalis (Casey) miniata (Germar) rubro (Newman) B. suturalis pearsalli (Bland) _ABSKMB B. ignicollis ignicollis (Say) _ ^AB acotnana Casey punctipennis Casey sang uinicoll is ( Germar) seclusa Casey Genus CROSSIDIUS LeConte Oxoplus LeConte C. carolinus (LeConte) SK C. discoideus discoideus (Say) AB pulchrior Bland C. discoideus blandi Casey -{B C. pulchellus LeConte AB debilis Casey lanaticollis Casey nitidicollis Casey palmeri Bates retractus Casey sinuosus Casey uegasensis Casey C. punctatus LeConte BC crossipes Casey hirsuticollis Casey plagiatus Casey uinosus Casey Genus ELYTROLEPTUS Dug6s E. floridanus (LeOonte) ON Tribe Holopleurini Genus HOLOPLEURA LeConte H. marginata LeConte BC helena LeConte Subfamily LEPTURINAE Tlibe Desmocerini Genus DESMOCERUS Audinet-Serville D. auripennis auripennis Chevrolat D. auripennis cribripennis Horn D. auripennis piperi Webb D. palliatus (Forster) ON pe NB \TIT blandus (Fabricius) cyaneus (Fabricius) elongatus Bland Mbe Necydalini Genus NECYDAI-IS Linn6 N. cavipennis LeConte BC N. diversicollis diversicolhs BC Schaeffer N. laevicollis LeConte BC seminiger Linsley N. mellita (Say) ON annericanus Haldeman 287 CerambyciJae Genus ULOCIIAETES LeOonte U. leoninur; LeConte BC Tlibe Lepturini Genus CEI{TRODERA LeConte Parapachyta Casey C. decolorata (Harris) oN PQ NB NS lacustris Casey rubidct (Haldeman) C. spurca (LeOonte) BC ceruin.z. (Walker) Genus STEiNOCORUS Fabricius Anisorus Mulsant Eutoxotus Casey Toxotc,psis Casey ToxottLs Dejean S. cylindricollis (Say) atratus (Haldeman) dentip ennis (Haldeman) diues (Newman) S. fl avolineatus (LeConte) _BC S. nubifer (LeConte) _BCAB apiciu,:ntris Casey hesperus Casey lateratis (Casey) margi:tellus Casey pacifit us Casey paruiceps Casey plagiatus Casey rufipe,znis Casey spinosus (Hopping) tenellt..s Casey truncctulus Casey S. obtusus .LeConte) BC SKAB breuicollis Casey S. schaumii(LeConte) MB ON PQ croceus (Leng) S. trivittatus (Say) _ABSK MB S. vestitus (Haldeman) BC AB ater (L,eng) aureatus Casey flaccia'.us Casey giluicarnis Casey oregonensis Casey sericai.us Casey subpir;guis Casey uirgat,rc (LeConte) S. vittiger (Randall) oN PQ nigrip,zs (Haldeman) Genus ENCYCLOPS Newman E. caerulea (Say) ON PQ pallipes Newman Genus PYR,OTRICHUS LeConte P. vitticollis LeConte BC cribripennis Casey Genus PACHYTA Dejean Argaleus LeConte Linsle;lana Podany P. armata LeConte _BC P. lamed lit,urata Kirby AK YK NT BC AB meoNpe: LB NF amerit:ana Podany confl ag1 r ata Motschul sky nitens (LeConte) 288 Cerambycidae Genus EVODINUS LeConte E. monticola monticola (Randall) MB ON PQ NB NS LB NF carolinensis Casey E. monticola vancouveri Casey AK NT BC Genus ANTHOPHYLAX LeOonte A. attenuatus (Haldeman) oN PQ NB NS A. cyaneus tHaldeman) oN PQ NB malac hitic u s (Haldeman) A. viridis LeConte -PQ NS uiridipennis Casey Genus NEANTHOPHYLAX Linsley & Chemsak N. mirificus (Bland) BC AB Denustus (Bland) N. tenebrosus nigrolineatus BC (Van Dyke) Genus RHAGIUM Fabricius Allorhagium Kolbe Hargium Samouelle Stenocorus Geoffroy R. inquisitor (Linndtx YK NT BC AB SK MB ON PQ NB NF americanurn Podany boreale Casey californicum Casey canadense Podany cariniuentre Casey crosslpes Casey i nuesliga lor Mannerheim lineatus (Olivier) mexicanurn Casey montanum Casey paruicorne Casey quadricostatum Podany thoracicum Casey Genus SACHALINOBIA Jacobson Pseudopachyta Swaine & Hopping S. rugipennis rugipennis (Newman) MB ON PQ NB Genus GAUROTES LeConte G. cyanipennis tSayt ON PQ ioze (Newman) lapor te i ( Gu6rin-M6neville) leonardi (Haldeman) ser u illei (Audinet-Serville) Genus ACMAEOPS LeOonte A. proteus proteus (Kirby) AK YK NT BC AB SK MB ON PQ NB LB aurora Casey cauicollis Casey coloradensis Casey gibbula LeConte lacustrina Casey obsoleta Casey parkeri Casey pinicola Schaeffer puncticeps Casey scutellata Casey sublineata (Haldeman) Genus LEPTALIA LeConte L. macilenta (Mannerheim) AK BC AB frankenhaeuseri (Mannerheim) fuscicollis (LeConte) 289 Cerambycidae Genus CO IITODERA Mulsant Acma,zopsilla Casey Lepta.:maeops Casey C. coniferae Hopping & Hopping - BC C. longicornis {Kirbyt - BC AB SK alticoia (Casey) 6oher,' Hopping & Hopping dorsa,',is (LeConte) harneyensis Hopping & Hopping ligata (LeConte) marginalis (LeConte) oregonensis Hopping & Hopping punctiuentris (Casey) suturalis (Casey) uincta (LeConte) C. militari'r militaris (LeConr,eJ - BC C. subpilosa (Leconte) - BC AB intermedia (Casey) lupino (LeConte) pugetuna (Casey) quadriceps (Casey) Genus PSEIIDOGAUROTINA Plavilstshikov P. abdomirralis (Bland I - ON PQ P. cressoni cressoni (Bland) - BC AB Genus BR.I\CHYSOMIDA Casey B. atra (Leconte) - BC AB aterriina Casey morata Casey B. bivittata (Say) AB SK MB ON PQ NB NS fuscicttps (LeConte) incert r (Bland)' nigrip,ennis (LeConte) oblonga (Casey) uariarts (LeConte) Genus GN.{.THACMAEOPS Linsley & Chemsak G. pratens s (Laicharting)* AK YK NT BC AB SK MB ON PQ NB LB NF fu lu i p ennis (Mannerheim) longiceps (Kirby) s em i - marg inara (Randall) s t rig il ata (Fabricius) Genus ME'IACMAEOPS Linsley & Chemsak M. vittata Swederus) PQ direcfrz (Newman) pallidc (Haldeman) q uadr iu ittata (Schonherr) Genus STIIANGALIA Audinet-Serville Ophistomis J. Thomson S tr anil alina Auriviilius S. acuminz.ta (Olivier) - ON emaciata Newman unicolor Haldeman S. bicolor (:Swederus) - ON simulons (Casey) S. famelica solitaria Haldeman - ON S. luLeicorrris (Fabriciust - ON PQ carolinae (Casey) euersa (Casey) Genus BELLAMIRA LeConte B. scalaris (Say) - SK MB ON PQ NB NS coarcto.ta (Haldeman) 290 Cerambycidae Genus ANALEPTURA Linsley & Chemsak A. lineola (Say) - ON PQ NB cincta (Haldeman) indirecta (Newman) lateralis (HaIdeman) obsoleta (Haldeman) Genus CHARISALIA Casey C. americana (Haldeman) - ON Genus GRAMMOPTERA Audinet-Serville Parallelina Casey G. exigua (Newman) - ON PQ saucia (LeConte) G. haematites (Newman) - SK MB ON PQ NB zono (Newman) ruficeps (LeConte) G. molybdica {Leconte) - BC G. subargentata (Kirby) AK YK NT BC AB SK MB ON PQ NB filicornis (Casey) quebecensis (Couper) rufi.basis (LeConte) similis (I\trby) Genus JUDOLIA Mulsant J. gaurotoides gaurotoides (Casey) AB SK hnulli (Swaine & Hopping) uiuaria Casey J. instabilis (Haldeman) - YK - BC AB SK MB conuexa (LeOonte) fl au iu entris (Schaeffer) pacifica Casey trajecta Casey J. montivagans montivagans (Couper) AK YK NT BC AB SK MB ON PQ NB NS - LB seminigra Casey J. quadrata (LeConte) - YK NT BC - SK MB ON PQ Genus PYGOLEPTURA Linsley & Chemsak P. carbonata (LeConte) - BC P. nigrella nigrella (Say) AK - NT BC AB SK MB ON PQ NB LB NF lacustris (Casey) nigrita (HaLch) praestans (Casey) serricornis (Casey) Genus DORCASINA Casey D. matthewsi (LeConte) - BC macrocera (Casey) Genus IDIOPIDONIA Swaine & Hopping l. pedalis (LeConte) - ON PQ NB NS Genus TYPOCERUS LeConte T. acuticauda acuticauda Casey - ON PQ NB NS thoracicus Casey T. balteatus Horn NT - AB T. confluens Casey MB caligans Casey T. lugubris tSayt - ON T. octonotatus (Haldeman) - ON arapahoe Casey sinuatus LeConte T. sparsus LeConte MB ON PQ T. velutinus velutinus (Olivier) MB ON PQ NB NS fugo* (Fabricius) manitobensis Swaine & Hopping tenuior (Kirby) 297 Cerambycidae T. zebra (Olivier) ON aurige" (Newman) carolir,La (Weber) zebratus (Fabricius) Genus NEOALOSTERNA Podany N. capitata {Newman) PQ NB NS N. rubida (l,eConte) BC _9* heeni (Casey) Genus PSEUDOSTRANGAIIA Swaine & Hopping P. cruentat a (Haldemant ON PQ maneii'. (Casey) Genus LEPTURA Linn6 (Subgenus LEPTURA s. str.) L. obliterata obliterata (Haldeman) BC idahoensis (Casey) perductor (Walker) uitiosa (LeConte) L. plagifera LeConLe BC L. plebeja Flandall yuoNpeNe_ L. propinquaBland AK BC minuscula (Casey) regularis (Casey) L. subhami.ta Randall ON PQ NB NS annatfl (Haldeman) elegan; (Haldeman) interrt..pta Newman (Subgenus iTENURA Dejean) (Mega,i.eptura Casey) L. emargin,rta Fabricius ON Genus STENOSTROPHIA Casey S. amabilis {Leconte) BC S. tribalteaba serpentina (Casey) BC Genus TRACIfYSIDA Casey T. aspera aspera (LeConte) AK YK NT BC AB parkeri (Hippisly) T. aspera b:revifrons (H. Howden) T. mutabili; (Newman) scABixreBNiSNENS- - NF I ae tifi<.: ( Provancher) I ur idi 1., enni s (Haldeman) prou at;.cher i (Aurivillius) quadncollis (LeConte) Genus PIDIfNIA Mulsant (Subgenus .PIDONIA s. str.) (Hapai,osalia Casey) P. ruficollis {Sayr ON PQ NB NS alleclo (Newman) fragilis (Casey) laeuiceps (Casey) Iineicornis (Casey) nitidicollis (Horn) paupercula (Newman) sphaericollis (Say) straus:;i (Webb) uiber (Newman) P. scripta (l,eConte) AK BC (Subgenus'IHESAJ-IA Casey) P. gnathoides (LeConte t BC lisa (Leng) rubriceps (Casey) 292 Cerambvcidae (Subgenus CERATOPIDONIA Linsley & Chemsak) P. quadrata Hopping - BC Genus ORTHOLEPTURA Casey O. valida (Leconte) - BC oculea Casey Genus STRANGAI-EPTA Casey S. abbreviata (Germarr - ON PQ NB NS gulosa (Kirby) nitidipe nnis ( Provancher) saratogensis (Rat) semiuittata (Kirby) S. pubera tSay) MB ON PQ Genus STICTOLEPTURA Casey S. canadensis canadensis (Olivier) MB ON PQ NB NS PE - NF c innamop te r a (Haldeman) erythroptera (Kirby) te nuic or n is (Haldeman) S. canadensis cribripennis (LeConte) - BC AB SK MB coccinea (LeConte) diuisa (Casey) ebena (Leng) Genus ANASTRANGAJ-IA Casey A. laetifica (LeConte) - BC lugens (LeConte) A. sanguinea (LeOonte) AK YK - BC AB SK MB ON PQ NB apicata (Casey) bould.erensis (Casey) Genus XESTOLEFTURA Casey X. behrensi (LeConte) - BC X. crassipes (LeContet - BC fas ciu en tr is ( LeConte) muliebris Casey shastana Casey uancouueri Casey tanthogaster (LeConte) X. octonotata (Sayt - ON q uadr ip unctato ( Haldeman) stictica (Newman) X. tibialis (LeConte) - \'K - BC AB ON PQ - NS - LB NF columbica Casey hirtella (LeConte) miquelonensis (Pic) pictipennis Casey Genus LEPTUROPSIS Linsley & Chemsak L. biforis (Newman) - ON PQ NB laurentica (Casey) L. dolorosa (Leconte) - BC Genus TRIGONARTHRIS Haldeman Cercolia Casey T. minnesotana (Casey) MB ON PQ NB NS T. proxima (Sayt MB ON PQ - NS hempiana (Casey) subpubescens (Kirby) Genus ACMAEOPSOIDES Linsley & Chemsak A. rufulus (Haldeman) - BC AB SK MB ON PQ - NS Genus BRACHYLEPTURA Casey B. champlaini Casey - ON PQ - NS PE 293 Cerambycidae B. dehiscens (LeConte) BC B. rubrica ([iay) ON PQ NB erythraptera (Germar) B. vexatrix, Mannerheim) AK BC conuoluens (Casey) quadrii,lum (LeConte) Genus STROPHIONA Casey S. laeta (Le(lonte) BC ostenta Casey S. nitens (Forster) MB ON PQ NS bellina Casey quagga (Germar) zebra (t)livier) Genus COS.MOSAJ-IA Casey C. chrysocorna (Kirby) YK NT BC AB SK MB ON PQ NB NS PE NF aureolc,. (Casey) auripilis (LeConte) densep,'.losa Casey Subfamily LAMIINAE Tlibe Parmenini Genus PLEOTRURA Mannerheim P. spinicauc.a Mannerheim AK BC produc.la LeConte Tlibe Moneileminr Genus MO]TIEILEMA Say Collapt:eryr Newman M ono p,'.e sa Motschulsky Omosc"ilon J. Thomson M. annulatrLm Say AB SK angulatumPsota demissutn Casey hybridum Psota montanum Psota nubeculum Casey puncticolle Psota Ttibe Monochamini Genus HEBESTOLA Haldeman Cacoplt.a LeConte H. nebulosa Haldeman PQ albata (Knu]l) pullato (Haldeman) pruinosa (LeConte) Genus MONOCHAMUS Dejean M. carolinensis (Olivier) oN PQ NB dentator (Fabricius) minor (LeConte) M. clamator' latus Casey BC M. marmorirtor Kirby MB ON PQ NB NS acutus (Lacordaire) fautor (.LeConte) M. mutator LeConte NT _SKMB oN PQ NB M. notatus lDrury) NT BC AB SK MB oN PQ NB NS PE confusor l{trby morgani Hopping M. obtusus rbtusus Casey _BC 294 Cerambycidae M. scutellatus scutellatus (Say) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF resutor Kirby M. scutellatus oregonensis (LeConte) BC AB monticola Casey M. titillator (Fabricius) -oN angusticollis Casey obesus Casey Genus MICROGOES Casey M. oculatus (LeConte) -ONPQ tenuicornis Casey Genus GOES LeConte G. debilis LeConte ON PQ G. pulcher (Haldeman) oN PQ fuscatus IKntTl G. pulverulentus (Haldeman) ON PQ laurenticus Casey Tribe Mesosini Genus SYNAPHAXTA J. Thomson S. guexi (LeConte) - BC annulata Casey breuicollis Casey hutneralis Casey Tlibe Dorcaschematini Genus DORCASCHEMA Haldeman Hetoemis Haldeman D. nigrum (Say) oN PQ TYibe Apodasyini Genus EUPOGONIUS LeConte Eriopsilus Bates Phidola J. Thomson E. pauper LeConte _ONPQ frafiniKnull uestitus (Say) E. subarmatus (LeConte) _ONPQ E. tomentosus (Haldeman) _ONPQ piniuora Filc}r Genus PSENOCERUS LeConte Paralarica Breuning P. supernotatus (Say) MB ON PQ NB funebris Casey picfzs (Breuning) pini LeConte frislis Casey Tlibe Pogonocherini Genus POLIAENUS Bates P. oregonus (Leconte) - BC Genus ECYRUS LeOonte Oebaceres J. Thomson E. dasycerus dasycerus (Say) - ON PQ exiguus (Haldeman) obscurus (Haldeman) 295 Cerambycirlae Genus LOI'HOPOGONIUS Linsley L. crinitus LeConter - BC Genus POCiONOCHERUS Dejean (Subgenus .POGONOCHERUS s. str.) P. penicillalus Leconte AK - NT BC AB - MB oN pe NB NS pE NF alashanus Schaeffer - carinatus Casey (Subgenus .OUPOGONOCHERUS Linsley) P. mixtus l:laldeman - yK NT BC AB - MB ON pe NB simple.x LeConte P. parvulus LeConte AB SK MB ON pe salicol'r Casey P. pictus Fi,ll NT BC AB emarginatus Casey fastigiutus Casey P. propinquus Fall - BC Tribe Onciderini Genus ONCTIDERES Lepeletier & Audinet-Serville O. cingulatrrs cingulatus rsayr - ON pallesczns Casey praecit:.tens Casey Tribe Hippopsini Genus HIPI)OPSIS Lepeletier & Audinet-serville H. lemniscata tFabriciusr - ON lineata Gt6rin-M6neville lineola la Audinet-Serville meinerli Aurivillius Tribe Acanthoderini Genus AXGIMORPHUS Haldeman Aethioltoctines J. Thomson A. modestus {Gyllenhal } - - MB ON pe decipie:zs Haldeman A. morrisi (lJhler) leucogenus (J. Thomson) A. quadrigit,bus tSayr ON lucidus (Knull) - Genus OPLOSIA Mulsant O. nubila (LeContet - ON pe Tribe Acanthocinini Key to North American species: Dillon (1956) Genus CANIfNURA Casey C. princeps Walkerr - BC uincta (Casey) Genus NEACANTHOCINUS Dilion N. obliquus,rbliquus (LeConter AB acornanLls (Casey) angulosus (Casey) sedulus (Casey) N. obliquus racificus (Casey) - BC N. obsoletus (Olivier) - ON pe floridar.:us (Casey) N. pusillus (Kirby) AK - AB SK MB ON pe NB NS 296 Cerambycidae Genus CERATOGRAPHIS Gahan C. biguttata (LeConte) PQ Genus GRAPHISURUS Kirbv Urographis Horn G. despectus (LeConte) - ON PQ hebes (Casey) G. fasciatus (DeGeer) - ON PQ mirlus (Fabricius) p enn sy lu anicus ( Gmelin) reductus (Casey) Genus AMNISCUS Haldeman Astylopsis Casey A. collaris Haldeman - ON PQ intenuptus Haldeman A. macula (Say) - ON PQ NB NS sticticus Haldeman A. sexguttatus (Say) MB ON PQ commixtus Haldeman guttatus (Say) Genus LEPTOSTYLUS LeConte L. transversus transversus - ON PQ (Gyllenhal) aculiferus (Say) rnarg inell us (Haldeman) t ub erculatu s (Frolich) Genus STERNIDIUS LeConte Key to North American species: Lewis (1986) S. alpha(Say) - ON PQ cinereus (LeConte) coloradensis DilIon diuergens (Haldeman) lateralis (Haldeman) nelsonicus (Casey) suturalis Dillon uicinus (Haldeman) S. fascicularis (Harris) - ON PQ - NS dentatus (Casey) nigricans Dillon obscurellus (Casey) pleuralis (Casey) scapalis (Casey) testaceus (Casey) timid,us (Casey) xanthoxyli (Shimer) s. misellus (Leconte) MB oN PQ S. rusticus (LeConte) MB Genus ASTYLEIOPUS Dillon A. variegatus (Haldeman) AB SK MB ON PQ NB obscurus (Haldeman) tr ifa sciatu s ( Haldeman) Genus DECTES LeConte D. sayi Dillon & Dillon AB - MB ON spizosus (Say) Genus LEPTURGES Bates (Subgenus LEPTURGES s. str.) L. confluens (Haldeman) - ON PQ L. symmetricus (Haldeman) - ON PQ 297 Cerambycidae Genus URGLEPTES Dillon U. facetus (liay) -ONPQ U. querci (Fitch) _ONPQ /risfis (Casey) U. signatus (LeConte) -oNPQ tenebrosus (Casey) Genus HYPERPLATYS Haldeman H. aspersa (Say) BC AB SK MB ON PQ NB NS NF uigilans Casey H. maculata Haldeman _MBONPQNBNS amnicola Casey binocul.aris Casey cryptiat Casey delicatr:.t Casey frigida Casey lentigit;.osa Casey nigrellcr Haldeman robustula Casey uariolata Casey H. montana Casey BC AB SK MB Tlibe Cyrtinini Genus CYR|CINUS LeConte Myrmo,bmia Bates C. pygmaeu,r (Haldeman) ON Tlibe Saperdini Genus SAPI)RDA Fabricius Eutetropha Bates Key to species: Abdullah and Abdullah ( 1966) S. calcarata Say BC AB SK MB ON PQ PE adspersa LeConte S. candida brpuncLata Hopping SK MB ON S. candida candida Fabricius -oNPQ NS biuittato Say S. concolor unicolor Felt & Joutel SK MB ON PQ NB NS NF mecasotdes Casey S. cretata Nr)wman MB S. discoidea Fabricius _ONPQ dorsalit Haldeman fuscipes Say S. fayi Blan< oN PQ S. horni Jou;el BC AB _MB S. imitans F.:lt & Joutel oN PQ S. lateralis F'abricius -MBONPQ NB NS abbreuiata Fitch connecte. Felt & Joutel disconolata Pic suturalis Fitch transeuns Breuning S. mutica Say MB ON PQ S. obliqua Say MB ON PQ NB NS PE S. populnea :moesta LeConte YK NT SC AB SK MB ON PQ LB S. populnea'populnea Linn6 BC AB S. puncticoll s Say BC -oNPQ trigeminata Randall S. tridentata Olivier SK MB ON PQ dubiosa Haldeman interme':lia Fitch rubronotata Fitc}r trifasciota Casey S. vestita Say MB ON PQ atkinsoni Crrtis 298 Cerambycidae Tribe Phytoecini Genus OBEREA Mulsant Amaurostoma J. Miilier Isosceles Newman Key to species: Breuning (1960) O. appaiachiana Casey _ON-NB O. bimaculata (Olivier) MB ON PQ affinis Haris basalis LeConte basicollis Breuning bicallosicollis Breuning bipunctata (Creutzer) broo&si Wallis delicatula Casey dolosa Casey filum Casey flauocephala Blatchley insignior Breuning insignis Casey iowensis Casey rnontana Casey per s picill ata Haldeman tr ip unctata ( Fabricius ) ulmicola Chittenden umbra Casey O. canadensis Fisher _SKMBONPQ O. cylindricollis Casey -oN O. deficiens Casey -ONPQ O. delongi Knull -oNPQ anter ufi collis Breuning O. exilis Casey -oN O. ocellata Haldeman -oN plagiata Casey O. oculaticollis (Say) MB ON O. pallida Casey MB ON PQ O. praelonga Casey _ONPQ O. pruinosa Casey AB_ -oN O. quadricallosa LeConte _BC O. schaumii LeOonte -BC AB SK MB ON PQ NB caseyi Plavilstshikov ferruginea Casey infrarufa Breuning su bcy I ind r i collis Breuni ng uittipennis Breuning wapleri Chevrolat O. tripunctata (Swederus) oN PQ NB amabilis Haldeman intermedia Casey mandarina (Fabricius) rnyops Haldeman subdeficiens Breuning subexilis Breuning submondarina Breuning texana Horn Genus MECAS LeConte (Subgenus MECAS s. str.) M. bicallosa Martin BC Tribe Tetraopini Genus TETRAOPES Schiinherr Key to species: Chemsak (1963) T. annulatus LeConte AB-MB canescens LeConte fontinalis Casey uteanus Casey uestitus Casey 299 Bruchidae T. femoratus LeConte BC AB SK MB ON amnicc'la Casey atrisetosus Casey breuisetosus Casey coseyj ,\urivillius collaris Horn fortis Casey monticola Casey nigripes Casey oregonetnsis LeConte Punctiltennis Qasst robustt,s Casey ruber (.Asey spissicornls Casey uegaser,,sis Casey uelutin ^ts Casey T. quinquemaculatus Haldeman ON T. tetrophthalmus (Forster) -MBONPQ humerc:Iis Casey iowensis Casey torndtor (Fabricius) t r e dec im punc tatu s (Dr apiez) Family BRUCHIDAE seed beetles J. McNamara The t ruchids comprise 19 species in Canada Borowiec (1987) published a key to the genera of and Alasra. Their habits vary considerably, but the world, and Udayagiri and Raj Wadhi (1989) they depend almost entirely on seeds for their published a catalog of the world species. development. The larvae live in seeds of legumes, notably l)eas and beans, and are often pests of stored leguminous seeds. The adults are commonly found in flowers and on foliage. The pea weevil (Bruchus pisorum) and the bean weevil aK (1);YK (1); NT (2); BC (10); AB (8); SK (7); MB (Acanthoscelides obtectus) are major pests of peas (e); ON (11); PQ (5); NB (2); NS (2); PE (1); NF (1) and beans, and have spread around the world. . I(6) Subfamily AMBLYCERINAE Genus AMB LYCERUS Thunbere A. robiniae ( Fabricius) AB gleditsiae (Castiglioni) Subfamily KYTORHININAE Genus KYTORHINUS Fischer von Waldheim K. prolixus (Fall) AK YK NT BC AB SK MB Subfamily BRUCHINAE Genus MEG.A.CERUS Fahraeus Pachybruchus Pic (Subgenus NTEGACERUS s. str.) M. discoidus (Say) BC AB SK MB ON PQ NB NS Genus CALI,OSOBRUCHUS Pic C. maculatus (Fabricius) SK ON ambiguzs (Gyllenhal) quadrinzaculatus (Fabricius) sinuatus (Fahraeus) 300 Chrysomelidae Genus GIBBOBRUCHUS Pic Key to North American species: Whitehead and Kingsolver (1975) G. mimus (Sayt ON borealis (Schdnherr) crataegi (Fahraeus) Genus MEIBOMEUS Bridwell Key to North American species: Kingsolver and Whitehead (1976) M. musculus tSay) ON PQ a lb og ut tis (Motschulsky) erythrocerus (Riley) Genus ACANTHOSCELIDES Schilskv A. aureolus (Horn) _BC AB SK MB A. calvus (Horn) -oN A. fraterculus (Horn) NT BC AB SK MB ON A. obtectus (Say)t _BC MB ON PQ fabae (Pi'iley) u ar icor ni s (Motschulsky) A. pauperculus (LeConte) BC p ulicar iu s (Motschulsky) rulzs (Motschulsky) simplex (Motschulsky) A. perforatus (Horn) AB _MB A. pullus (FalI) BC A. seminuium (Horn) AB SK MB Genus BRUCHUS Linn6 B. brachialis Fahraeust _BC -oN B. pisorum (Linn6)t _BCAB MB ON PQ NU NS PE NF pisi Linn6 B. rufimanus (Boheman)t SK MB ON PQ Genus BRUCHIDIUS Schilsky Key to North American species: Bottimer (1968) B. cisti (Fabricius)t BC unicolor (Olivier) B. villosus (Fabricius)f ON oler (Marsham) Family CHRYSOMELIDAE leaf beetles L. LeSage The family Chrysomelidae is one of the largest in remaining subfamilies are either root feeders, root group of beetles in the world, with over 50 000 miners, stem miners, or leaf miners. known species and many additional thousands to be Chrysomelidae is usually classifred into 19 sub- discovered, especially from Asia and the Neotropical families (Seeno and Wilcox 1982) of which 14 occur region; 569 species and subspecies are listed here for in Canada. The fauna of the leaf beetles is relatively Canada and Alaska. All known adults are phy- well known in eastern Canada (except for some tophagous. Primitive plants and ferns are used alticine, galerucine, and cryptocephaline genera) but infrequently, and conifers are barely touched; decid- is poorly known in western Canada. The present list uous trees, bushes, and weeds are preferred host- follows essentially the arrangement proposed by plants. Some species are pollen eaters when in the Seeno and Wilcox (1982), except where the generic adult stage. Larvae show a broader ecological diver- concepts have been recently modifred. sity: Donaciinae are aquatic; Clytrinae live in ant nests, where they feed on plant detritus and on ant larvae or pupae; most Cryptocephalinae feed on decaying matter in leaf litter; Criocerinae, AJ{ (42); YK (41); NT (72); BC (181); AB (217); SK Chlamisinae, Chrysomelinae, and Cassidinae and (174); MB (25a); ON (363); PQ (27a); NB (116); NS some Galerucinae and Alticinae feed on leaves: those (103);PE (19); LB (1a); NF (3a) o H (9); I (38) 301 Chrysome lidae Subfamily DONACIINAE Key to New World genera: Askevold (1990) Tlibe Plateumarini Genus PL,A.TEUMARIS C.G. Thomson P. aurifera (LeConte) AK YK NT AB_MB PQ NF P. balli Askevold PQ NB P. chalcea (Lacordaire) _BC AB SK MB ON PQ NB NS PE lodingi .Schaeffer) s hoema,ize ri ( Schaeffer) P. diversa (Schaeffer) P. dubia (Schaeffer) AK--nCAB-yuoN- idola (Hatch) P. flavipes (Kirby) AK YK NT BC AB SK MB ON PQ NB LB NF wallil;i (Schaeffer') P. frosti (Schaeffer) NT BC AB SK MB ON PQ NB NS P. fulvipes (Lacordaire) AK_NTBCABSKMBONPQNBNSNF P. germari (Mannerheim) AK YK NT BC AB SK MB ON PQ NB NS _ LB NF diues (LeConte) fl au ip ennis ( Mannerheim) serricauda (Schaeffer) P. metallir,a (Ahrens) ON PQ NB NS catar'rctae (Newman) femor alis (Kirby) gentitis (Leconte) indicu (Melsheimer) zaza (Melsheimer) parua: (Lacord.afte) P. nitida (tlermar) BC AB SK MB ON PQ NB NS PE LB NF emarSlinata (Kirby) juncina (Couper) paciftca (Schaeffer) P. neomex icana (Schaeffer) BC long it:' ol I i s ( Schaeffer) u er mti. c ulat a ( Schaeffer) P. pusilla Say) YKAX NT BC ff SK MB ON PQ NB NS NFLB pyritt: sa (LeConte) rugiftons (Newman) P. robusta (Schaeffer) oN PQ P. rufa (Say) _ y:'TSKyu oN PQ NB NS affinis (I\trby) jucunda (LeConte) hirbyi (Lacordaite) s ulcic ollis (Lacordaire) Tribe Donaciini Genus DO NACIELLA Reitter D. pubicollis ( Suffrian) AB SK MB ON Genus DONACIA Fabricius (Subgenus DONACIA s. str.) Key to l\lorth American species: Marx (1957) D. cinctico:rnis Newman MB ON PQ NS antillarum Suffrian lucidl:, Lacordaite pulch,zlla LeConte rufipennis Lacordaire tenuis Schaeffer tryphera Schaeffer D. palmata Olivier ON PQ NB NS angustipes Marx claud;icans Germar D. parvide.ns Schaeffer -oN D. piscatrir Lacordaire MB ON PQ NB alutacea LeOonte carolina Lacordaire conge;'Ler LeConte cuprea Kirby 302 Chrysomelidae D. proxima Kirby NT BC AB SK MB ON PQ NB NS NF californica LeConte e pisco palis Lacordaire D. rufescens Lacordaire ON PQ D. texana Crotch ON minor Schaeffer (Subgenus DONACIOMIMA Medvedev) D. biimpressa Melsheimer MB ON PQ a ur ichalcea Melsheimer torosa Le0onte D. caerulea Olivier MB ON PQ NB aequalis Say confusa Lacordaire confusa LeConte D. canadensis Askevold AB SK MB ON PQ D. cazieri Marx NT BC AB _ONPQ LB D. confluenta Say _BC AB SK MB ON PQ D. cuprea Kirby -SK -oNPQ curticollis Knab D. distincta LeConte AK NTYK BC AB SK MB ON PQ LB NF occidentalis Mead D. fulgens LeConte _SK MB ON PQ NB NS PE D. hirticollis Kirby AK YK NT BC AB SK MB ON PQ NB NS_LBNF rudicollis Lacordaire D. limonia Schaeffer _ONPQ D. magnifrca LeConte BC AB SK MB ON PQ NB D. megacornis Blatchley -oN megaloceraWeise D. porosicollis Lacordaire NT BC AB SK MB ON D. pubescens LeConte BCTSK MB ON PQ D. rugosa LeConte -oN D. subtilis Kunze BC AB SK MB ON PQ NS aenea Ahtens aerea Lacord.aire magistrigata Mead quadricollis Say D. tuberculifrons Schaeffer BC MB ON PQ NB Tribe Haemoniini Genus NEOFIAEMONIA Sz6kessy Key to North American species: Askevold (1988) N. flagellata Askevold MB ON N. melsheimeri (Lacordaire) YSK MB ON PQ N. minnesotensis Askevold MB ON N. nigricornis tKirby) BC MB ON PQ americana (Gu6rin-M6neville) Subfamily ORSODACNINAE Genus ORSODACNE Latreille Orsodacna Dejean O. atra (Ahrens) YK NT BC AB SK MB ON PQ NB NS anneniacae Germar childreniKirby hepatica Say inconstans Newman luctuosa Lacordaire ruficollis Newman tibialis Iltrby tricolor Melsheimer triuittata Lacordaire uittata Say 303 Chrysomelidae Subfamily ZEUGOPHORINAE Genus ZEIIGOPHORA Kunze (Subgenus ZEUGOPHORA s. str.) (Auchenia Thunberg) (Tararis LeConte) Z. abnormis (LeConte) AK YK NT BC SK MB ON PQ hirbyi Baly reineckei Gtote Z. atraFall YK NT BCABSKMB_PQ Z. consanguinea Crotch BC AB SK Z. multnomah Hatch YK BC_ON Z. puberula Crotch -Nr ON Z. scutellaris Suffriant BC AB SK MB ON PQ Z. varians Crotch BC AB SK MB ON PQ Subfamily CRIOCERINAE Tlibe Criocerini Genus LIL IOCERIS Reitter (Subgenus LILIOCERIS s. str.) L. lilii (Scopoli)t oN PQ lilioru,m (C.G. Thomson) melanocephala (Say) m e r di,;1 e r a (Fabricius ) Genus CRIOCERIS O.F. Mriller Pleurcphora Chewolat C. asparagr (Linn6)t _BCABSKMBONPQ NB C. duodecirnpunctata (Linn6)f _BCABSKMBONPQ NB Tribe Lemiini Genus LEIIA Fabricius (Subgenus LEMA s. str.) L. cyanella Linn6f L. gaspensis W.J. Brown PQ L. palustris Blatchley _ONPQ L. trilinea trilinea R.E. White MB ON PQ bilineo.ta Omer-Couper & Miles medionata Schaeffer medionota Schaeffer trilineuta atct. L. trivittatr,L trivittata Say ON PQ imma< ulicollis Chevrolat Iecontti Clatk nigrou.ittata Schaeffer triuirgzta LeConte Genus OUI.,EMA des Gozis (Subgenus I)ULEMA s. str.) O. melanopus (Linn6)t ON PQ Subfamily CLYTRINAE Key to New World genera: Moldenke (1981) Key to North American species: Moldenke (1970) TYibe Clytrini Genus ANC,TMOEA Lacordaire Anomoia Dejean Euthyna Gistel A. laticlavia laticlavia (Forster) MB ON PQ ephipptum (Germar) lloridana Schaeffer 304 Chrysomeiidae hansana Schaeffer nigripes (Fabricius) o6silo (Fabricius) ruficauda (Foersberg) Tribe Megalostomini Genus COLEOTHORPA Moldenke C. dominicana dominicana NT_AB_MBON (Fabricius) C. vittigera vittigera (LeConte) _BCABSKMB Tribe Babiini Genus BABIA Chevrolat Harpasta Gistel (Subgenus BABIA s. str.) B. quadriguttata quadriguttata ON (Olivier) biguttata Crotch quadriforis (Newman) Genus SAXINIS Lacordaire (Subgenus BOREOSAXINIS Moldenke) S. omogera omogera Lacordaire ON S. saucia saucia LeOonte ec Subfamily CRYPTOC EPHALINAE Tlibe Pachybrachini Genus PACHYBRACHIS Chevrolat (Subgenus PACH\aBRACHIS s. str.) (Pachystylus Rey) P. abdominalis (Say) AB SK MB P. alticola Fall AB P. atomarius (Melsheimer) AB-MBON atomus Bowditch infaustus Haldeman P. autolycus FaIl -BCABSKPQ P. bivittatus (Say) -BCABSK-ONPQ albescens Suffrian uiduatus Suffrian P. calcaratus Fall P. coloradensis Bowditch ---AB--oN-- seuier Bowditch P. discoideus Bowditch _AB P. donneri Crotch BC AB P. hepaticus (Melsheimer) -ABSKMBONPQ punctdtus Haldeman P. luridus (Fabricius) ON PQ oesculi (Melsheimer) femoratus (Say) festiuus Fall moerens Stil, nigrinus Blatchley P. melanostictus Suffrian _YKNTBCABSKMB P. minor Bowditch MB- P. mnigrum (Melsheimer) - - - :c - - pe - - - - - intricatus Suffrian P. nigricornis carbonarius AB_MBON (Haldeman) P. nigricornis diffrcilis Fall _BCABSKMBONPQ wahsatchensis Fall P. nigricornis nigricornis (Say) AB SK MB janus Fall 305 Chrysomelidae P. obsoletr.,s Suffrian BC AB _MBONPQ P. othonous othonous (Say) _MBONPQ rna rg r. naticoll is Randall P. othonous pallidipennis Suffrian SK striatus LeConte P. othonous sioux Balsbaugh AB SK MB P. peccans Suffrian AB MB ON PQ NB NS--- P. pectoralis (Melsheimer) _ONPQ NB NS oculatus Suffrian sobrin.us Haideman stricticus Blatchley P. praeclarus Weise MB ON elegar,.s Blatchley P. relictus Fall -oN P. spumar:us Suffrian --sx MB ON PQ robori:s Fall P. subfasci atus LeConte MB ON bigutiatus Suffrian P. tridens tMelsheimer) P. trinotatrrs (Melsheimer) P. vau Fal] -__AB P. vacillatus Fall BC Tribe Monachini Genus LEIIIPHANES Gistel Mona,:hulus Leng Mona:hus Chevrolat Key to l\'orth American species: Balsbaugh (1g66) L. saponat rs ( Fabricius t MB ON PQ NB oler (IIaldeman) uiridanus (Falll Tlibe Cryptocephalini Genus CRIIFTOCEPHAIUS O.F. Mtiller (Subgenus CRYIvIOCEPHAJ-US s. str. ) (C antiiostethus Haldeman) Key to North American species: White (1968) C. calidus Suffrian _MBON C. confluer:.tus confluentus Say BCY:"-o*pe C. gibbicollis gibbicollis Haldeman NB NS C. guttulatus Olivier oN PQ loufur Newman C. incertus Olivier -oN PQ lineoltfius Haldeman C. insertus Haldeman AB SK MB ON PQ ellipscidalis Casey C. leucomelas leucomelas Suffrian ON C. mutabil: s Melsheimer _MBON PQ discoideus Suffrian dispersus Haldeman C. notatus notatus Fabricius ON PQ NB NS C. notatus quadrimaculatus Say Bcy SK MB ON PQ C. notatus sellatus Schaeffer _ _ _MB C. quadruplex Newman _MBONPQ NS q uadr tg uttulus Suffrian C. sanguin collis Suffrian BC nigerrimus Crotch C. schreibersii Suffrian ON rugicallis Haldeman C. spurcus vandykei R.E. White BC C. striatulrLs LeConte _PQ C. tinctus LeConte _PQ C. trivittatrs Olivier _PQ lirus Newman uilolus Haldeman C. venustus cinctipennis Randall SK MB ON 306 Chrvsomelidae C. venustus hamatus Melsheimer MB C. venustus ornatulus Clavareau MB 9" ornatus Fabricius C. venustus simplex Haldeman ON C. venustus venustus Fabricius AB SX nlS ON PQ flaccidus Suffrian Genus DIACHUS LeConte Fabricianus Weise BC MB D. aeruginosus LeConte NT pe D. auratus (Fabricius) -BC MB ON NS aeneus (OIiier) chalconotu s (Mannerheim) urrldis (Melsheimer) D. catarius (Suffrian) BC ON PQ NBI'- D. pallidicornis (Suffrian) l.BsKMB ON Genus BASSAREUS Haldeman B. formosus (Melsheimer) -oN PQ NS egenus Suffrian guttatus Haldeman B. lituratus (Fabricius) -oN B. Iuteipennis (Melsheimer) -oN PQ_ B. mammifer (Newman) -oN B. pretiosus (Melsheimer) MB B. sellatus Suffrian MB ON PQ (?) notatus (Kirby) (?) q uadris ig nolus Suffrian B. sulphuripennis (Melsheimer) ON Genus TRIACHUS LeConte T. atomus (Suffrian) -oN T. vacuus LeConte -oN Ne- Subfamily CHLAMISINAE Genus NEOCHLAMISUS Karren Key to North American species: Karren (1972) N. alni (W.J. Brown) -oN N. bebbianae (W.J. Brown) MB ON PQ lls N. chamaedaphnes (W.J. Brown) -ONPQ N. comptoniae (W.J. Brown) NB NS N. coryli (W.J. Brown) -oN N. cribripennis (LeConte) _ONPQ NS N. eubati (W.J. Brown) _ONPQ N. fragariae (W.J. Brown) NB N. tectus (W.J. Brown) -oN Genus EXEMA Lacordaire Key to North American species: Karren (1966) E. canadensis Pierce MB ON PQ NB E. dispar Lacordaire -oN pennsyluanica Pierce Subfamily SYNETINAE Genus SYNETA Dejean Key to species: Edwards (1953) Key to eastern North American species: Brown (1961) S. albida LeConte BC suturalis LeConte S. carinata Mannerheim AK BCT S. extorris borealis W.J. Brown --oNpeNnNs-- NF S. ferruginea (Germar) _MBONPQNBNS NF cosfolo (Newman) rubicunda Lacordaire tripla (Say) 307 Chrysomel:idae S. hamata Horn - BC AB SK S. pilosaW.J. Brown - YK NT BC AB SK MB ON PQ NB NS NF S. simplex simplex LeConte - BC S. simplex subalpina Edwards - BC AB Subfamilv ELIMOLPINAE Tri6e Nodinini Genus METACHROMA Chevrolat Atracli.ela Gistel Key to syrecies: Blake (1970) M. angustr:lum Crotch AB SK MB ON parall,zlum}J.orn M. longulun Horn AB SK no u e m s triat um Schaeffer opacipenne Fall Genus PAF:IA LeOonte Key to North American species: Wilcox (1957); Balsbaugh (1970) P. aterrima tOliviert PQ P. fragaria,r fragariae Wilcox - ON PQ P. pratensi s Balsbaugh PQ P. quadrigr.ttata LeConte AB SK MB salicetLWllcox P. quadrin< tata (Say) - ON PQ giluipts Crotch infusafta (LeConte) melan.t ra (Melsheimer) P. sexnoLat r tSayt MB P. scutellaris (Notmanr - ON P. thoraci*L (Melsheimert MB ON PQ Tribe Eumolpini Genus SPIITITHEROPTryTA Dei ean Chalc<'paria Crotch Chrysc,dina BaIy Key to Nrrth American species: Schultz (1976) S. globosa (Oliviel - ON Genus NOL)ONOTA Le{bwe BrachS,pnoea Gislel Nodo (lhevrolat Key to Nrrth American species: Schultz (1980) N. clypealis Horn AB N. margaretae Schultz MB ON N. puncticollis tSayt AB SK MB ON PQ oenea ('Melsheimer) humeralis (LeConte) s t r ig icollis (Lelbvre ) N. tristis (Crlivier) - ON ouata (.Say) pilula (Getrnar) Genus COI.ASPIS Fabricius Key to N,rrth American species (part): Blake (1974) C. brunnea (Fabricius) - ON PQ flauidtt Say flauicans LeGvre C. costipennis CroLch - ON crotchi Lelevre C. favosa Say - ON C. suilla bo: ealis Blake MB ON Genus RIIABDOPTERUS LeGvre Rhabdophorus Lelbvre Key to North American species: Schultz (1977) 308 Chrysomelidae R. deceptor Barber AB MB ON R. praetextus (Say) -oNPQ PE spiculatus Barber ueisei (Schaeffer) Genus CHRYSOCHUS Chevrolat Atymius Gistel Key to western North American species: Hatch (1971) C. auratus (Fabricius) _BCABSK MB ON PQ C. cobaltinus LeConte _BC californicus Marshall castaneus Marshall tenebricosus Marshall Tribe Adoxini Genus GRAPHOPS LeConte Heteraspis LeConte Key to species: Blake (1955) G. cupraeus ( Provancher) PQ G. curtipennis curtipennis -oN (Melsheimer) G. marcassitus (Crotch) _AB MB ON PQ G. nebulosus (LeConte) G. negrellus Blake . - _ TSK MB G. nigellus Blake MB G. pubescens (Melsheimer) MB ON PQ NS G. varians LeConte yu_ G. viridis Blake SK _ Genus XANTHONIA Baly X. decemnotata (Sayt SKMBONPQ-NS X. vil losula (Melsheimer) _MBONPQ_NS plagiata (Melsheimer) steuensi Blake Genus FIDIA Walsh F. viticida Walsh -oN lurida Lefevre Genus BROMIUS Chevrolat Adoxus Kirby Eumolpus Redtenbacher B. obscurus (Linn6)* AK YK NT BC AB SK MB ON PQ NB NS PE NF cochlearius Say concinnus Weise leaisi Weise Genus MYOCHROUS Erichson Key to North American species: Blake (1950) M. denticollis tSayt PQ M. squamosus LeConte AB SK Genus GLYPTOSCELIS Chevrolat Key to species: Blake (1967) G. albida albida LeConte - BC G. artemisiae Blake BC G. longior LeConte G. pubescens (Fabriciust _-Bc_- MB ON PQ pini (Say) hirsuta (Gmelin) hirta (Olivier) G. septentrionalis Blake BC 309 Chrysomelirlae Subfamily CHRYSOMELINAE Key to North American species: Wilcox (1972) T?ibe Timarchini Genus TIMARCHA Latreille (Subgenus r\MERICANOTIMARCHA Jolivet) Key to New World species: Jotivet (1948) T. cerdo StAl _BC aenea Jtolivet intento Jolivet nigra Jolivet T. intricata Haldeman _BC adusta Jolivet uiolacet Jolivet uiridis Jollet Tribe Chrysomelini Genus ZYGDGRAMMA Chewolat Z. conjuncta conjuncta (Rogers) AB SK MB Z. continua LeConter AB Z. disrupta'Rogersr - ON Z. exclamationis (Fabricius) AB SK MB ON pe Z. suturalis casta ( Rogers) MB fesliuo .Fabricius) Z. suturalis suturalis tFabriciusr - ON pe pzlcro ,Fabricius) Genus CALI,IGRAPHA Chevrolat B oliog r tp ha Motschulsky M eta llc; g r ap ha Motschul sky Phyllis Gistel Polyspita Hope Key to North American species: Brown (1g45) (Subgenus C|.RAPHICAI-LO Monros) C. lunata hlbrida tSayr AB SK MB lateuittuta (Achard) C. lunata lunata (Fabricius) - ON brunnea, (Thunberg) bowditchi (Achard) mediori:tpta (Achard) (Subgenus I IDENSOMELA Monros) C. bidenticola bidenticola - ON PQ W.J. Brown hilaris rAchard) intermedia (Achard) similis (Rogers) (Subgenus COREOPSOMELA Monros) C. californica californica Linell - BC C. californica coreopsivora NT BC AB SK MB ON PQ NB - pE W.J. Brown elegans (Olivier) (Subgenus ACALLIGRAPHA Monros) C. praecelsis (Rogers) MB (Subgenus CAILIGRAPHA s. str.) C. alni Schaeffer - - AB SK MB ON Pe NB NS confinis (Kirby) C. alnicola V/.J. Brown - ON PQ NB NS C. amator S'.J. Brown - ON C. apicalis Notman - ON PQ C. confluens Schaeffer - ON PQ NB NS C. dolosa W.J. Brown - ON PQ 310 Chrysomelidae C. igrrota W.J. Brown MB ON PQ NB NS PE C. incisa (Rogers) MB C. knabi W.J. Brown MB ON PQ C. multipunctata bigsbyana (Kirby) nc AB sx y" PQ NE NS C. multipunctata multipunctata _AB 9* (SaY) C. ostryae W.J. Brown -oNPQ C. philadelphica (Linn6) BC_SK MB ON PQ NB NS decipiens (Weber) ruficornis (Olivier) C. pnirsa (StAl) ON PQ labyrinthica Horn C. pruni W.J. Brown -oN C. rhoda Knab MB ON C. rowena Knab MB ON PQ NB C. scalaris (LeConte) MB ON PQ multiguttis Stitl C. sigmoidea (LeConte) BC C. spiraeae {Say) T -oN C. tiliae W.J. Brown _ON-NB C. verrucosa (Suffriant NT BC AB SK MB ON C. vicina Schaeffer -oNPQNB C. virginea W.J. Brown -ONPQNB Genus LEPTINOTARSA SIAI Myocoryna Sthl Polygratnma Chevrolat Key to North American species: Jacques (1988) L. decemlineata decemlineata (Say) BC AB SK MB ON PQ NB NS PE albida Tower intermedia Towet melanica Tower minuta Towet multitaeniata (StA\:l oblongata "Iower pallidaTower rubicundaTower rubriuittata 'Iower tortuosa Tower Genus I"ABIDOMERA Chevrolat P arop simena Motschulsky L. trimaculata (Fabriciust MB ON PQ cliuicollis (Kirby) rogersi LeConte Genus CHRYSOLINA Motschulsky Ap te r osoma Motschulsky Chrysomela auct. OreinaMonros & Bechyn6 Key to species: Brown (I962a) (Subgenus CHRYSOLINA s. str.) (S tic hosoma Motschulsky) C. staphylea tLinn6)t PQ NS NF (Subgenus SPHAEROMELA Bedel) C. varians (Schaller)t BC (Subgenus HYPERICIA Bedel) C. hyperici (Forster)* BC_ON NB NS C. quadrigemina (Suffrian)t BC_ON NB NS gemellata atct. gemtnata anct. (Subgenus CHALCOIDEA Motchulsky) C. frnitima W.J. Brown AK YK C. fl avomarginata fl avomarginata AB (Say) C. flavomarginata vidua (Rogers) BC AB 311 Chrysomelirlae C. hudsonica W.J. Brown NT MB ON PQ NF (Subgenus I'LEUROSTICA Motschulskv) C. cavigera iJ.R. Sahlberg)x AK yK lolli Jar:obson (Subgenus I,RCTOLINA Kontkanen) C. caurina Vy'.J. Brown AK C. magniceps (J. Sahlberg) AK C. subsuicat.a (Mannerheim) AK yK (Subgenus I'EZOCROSITA Jacobson) C. auripenn:s (Say) AK MB inornalz (Rogers) C. basilaris rSay) - YK BC AB Genus GASI ]ROPHYSA Chevrolat Key to North American species: Jolivet (1951) (Subgenus CASTROPIrySA s. str.) (Gastro'zidea Hope) G. cyanea caesia Rogers NT BC AB SK G. cyanea c;anea Melsheimer ON G. dissimilis (Sayr N'T_ABSK_ G. lormosa t iay r AB SK MB G. polygoni tLinner+ _BCABSKMB ON PQ NB NS PE caeruli1tennis (.Say) r uftcoll, s (Fabricius) Genus PHAINDON Megerle von Miihlfeld Key to Nodh American species: Balsbaugh (1983) (Subgenus PFIAEDON s. str.) P. armoraciae armoraciae (Linn6)* AK YK NT BC AB SK MB ON PQ NB NS NF americcnus Schaeffer P. armoraciae niger Hatch _BC planus Jatch P. laevigatus (Duftschmid)* AK NT BC AB SK MB ON PQ NB NS carri Hr*ch cochleariae auct. amer. uniform:is Fall uancout'erensis Hatch P. oviformis lleConte) AK YK NT BC AB sK oN PQ:tuNS--- P. prasinellus (LeConter _BCAB yu punctat.us Hatch uandyke'iHatch P. viridis Melsheimer SK MB ON PQ NB aeruginosus Suffrian dietrichi Hatch micr o re i: ic ula tzs Hatch oklahortensis }{ahch Genus HYDIIOTHASSA C.G. Thomson (Subgenus HYDROTIIASSA s. str.) (Erernosis des Gozis) H. boreela Schaeffer - YK AB SK MB H. obliquata (Leconte) v:' ON PQ H. ovalis tBlatchleyr ON H. vittata tOlivierJ* NT AB SK MB ON PQ NS triuittatt (Say) uaripes r.LeConte) Genus PRASOCURIS Latreille Helodes Paykull P. phellandrii (Linn6)* AK NT BC AB SK MB ON PQ Genus PLAGIODERA Chevrolat (Subgenus PI-AGIODERA s. str.) (Me laso tn ida Schaeffer) (Pseudoiina Schaeffer) 312 Chrysomelidae P. versicolora (Laicharting)t MB ON PQ NB ar tnor aciae ( Fabricius) Genus CHRYSOMELA Linne Key to New World species: Brown (1956) (Subgenus CHRYSOMELA s. str.) (Lina Dejean) (Melasoma Stephens) C. aeneicollis (Schaeffer) BC AB quadriguttoides Beller & Hatch C. blaisdelli (Van Dyke) AK NT BC C. confluens (Rogers) _BC maculicollis Schaeffer C. crotchi W.J. Brown AK NT_ABSKMB ON PQ NB NS C. engelhardti (Hatch) AK YK NT C. falsa W.J. Brown AK YK NT BC AB SK MB ON PQ _LB C. invicta W.J. Brown AB v C. knabi hesperia W.J. Brown AB SK MB C. knabi knabi W.J. Brown oN-- C. laurentia W.J. Brown NT_AB oN PQ C. lineatopunctata (Forster) NT BC AB SK MB oN PQ nouaboracensis Gmelin obsoleta (.Say) s cr ip toide s ( Schaeffer) uariegata Olivier C. mainensis interna W.J. Brown BC AB C. mainensis littorea W.J. Brown AK BC C. mainensis mainensis Bechyn6 YK NT _AB MB PQ NB NS _LBNF C. schaefferi W.J. Brown AK YK BC AB MB 9* immaculata (Schaeffer) C. scripta Fabricius BC AB SK MB ON C. semota W.J. Brown YK BC AB C. walshi W.J. Brown )' ON PQ Genus GONIOCTENA Chewolat Key to North American species: Brown (1942o) (Subgenus C.ONIOCTENA s. str.) (Phytodecta Kirby) G. americana (Schaeffer) AK YK NT BC AB SK MB ON PQ NB LB arctica Bechyn6 guttifera Bechyn6 G. nivosa alberta W.J. Brown -NT_AB G. nivosa arctica (Mannerheim) AK YK NT BC AB MB G. nivosa nivosa (Suffrian) G. notmani (Schaeffer) YKNT_ABSK MB ON PQ LB G. occidentalis (W.J. Brown) YK-BCAB Genus PHRATORA Chevrolat Key to North American species: Brown (1951) (Subgenus PHRATORA s. str.) (Phyllodecta l\rby) P. americana canadensis W.J. Brown ON PQ NB NS P. frosti frosti W.J. Brown NS__ P. frosti remissa W.J. Brown AK EC AB SK MB ON PQ P. hudsonia W.J. Brown AK YK NT BC AB sK yu oN PQ P. kenaiensis W.J. Brown AK P. interstitialis Mannerheim AK YK NT BC aklauiki (Can) P. purpurea novaeterrae W.J. Brown NF P. purpurea purpurea W.J. Brown YK NT BC AB Sf ff,fS ON Pe NN NS Genus ENTOMOSCELIS Chewolat C hry sornelop silo Strand Chrysomelopsis Achard Key to North American species: Brown (1942b) E. americana W.J. Brown AK YK NT BC AB SK MB 313 Chrysomelic;.ae Subfamily GALERUCINAE Tribe Galerucini Key to North American species: Wilcox (1965) Genus DEROSPIDEA Blake Key to North American species: Blake (1931) D. brevicollis (LeContet ON Genus TRIRHABDA LeConte Key to North American species: Blake (1931) T. adela Bla- Genus GAI-IDRUCELLA Crotch (Subgenus C:ALERUCELLA s. str.) (Hydrollaleruco Laboissidre) G. nymphaerre (Linn6)x AK YK NT BC AB SK MB ON PQ NB PENS LB NF fernoralis (Meisheimer) Iuctuosu (Mannerheim) margin,zlla (Kirby) sagittariae (Kirby) stilleri ?app (Subgenus I\'EOGAIERUCELLA Chujo) G. quebecen;is W.J. Brown PQ G. stefansso:ri W.J. Brown AK YK NT PQ_NS Genus TRICHOLOCHMAIA Laboissibre T. alni (Fall. -oN PQ NB NS T. decora calbo (LeConte) - BC AB SK MB T. decora de,:ora (Say) -oN PQ NB solicis (Randall) T. cavicollis il,eConte) AB SK MB ON PQ haematlca (LeConte) T. kalmiae (Fallt -oN PQ NB NS T. perplexa tFallt PQ NB T. punctipennis pallida - BC _9* (Beller & Hatch) T. punctipennis punctipenms - BC (Mannerheim) T. ribicola cc'nfusa (W.J. Brown) ON T. ribicola ribicola (W.J. Brown) AB SK MB T. rufosangu.inea (Sayt _PQ T. sablensis (W.J. Brown) NS- T. spiraeae t Fall.) oll pe - T. spiraeophila (Hatch & Beller) - BC T. tuberculaba (Say) ON PQ NB ols - T. vaccinii (Fallt ON PQ NB Genus PYRRHALTA Joanni s P. viburni (Paykullt* ON PQ Genus XANI|HOGALERUCA Laboissibre X. luteola (O.F. Miiller)t - BC oN PQ 3L4 Chrysomelidae Genus ERYNEPHAI-A Blake Key to species: Blake (1936) E. rnaritima (LeConte) NS texana Schaeffer E. puncticollis (Say) AB SK MB Genus MONOXIA LeConte Key to species: Blake (1939) M. angularis (LeConte) M. consputa LeConte M. debilis LeConte M. grisea Blake M. inornata Blake M. puberula Blake M. sordida LeConte Genus OPHRAELLA Wilct.rx Key to North American species: LeSage (1986) O. americana (Fabricius) AB SK MB PQ O. arctica LeSage NT MB 9* O. bilineata (Kirby) -BC AB SK MB O. communa LeSage _BC AB SK ON notulata anct. O. conferta (LeConte) AB SK MB oN PQ NB NS O. cribrata (LeConte) AB SK yu ON PQ O. notata (Fabricius) oN PQ O. nuda LeSage AB O. pilosa LeSage _BC AB SK MB ON PQ Genus GA.LERUCA O.F. Miiller Key to North American species: Blake (1945) (Subgenus GALERUCA s. str.) (Adimonia Laicharting) G. browni Blake G. externa (Say) G. rudis (LeConte) Tribe Luperini Genus DIABROTICA Chewolat D. barberi Smith & Lawrence MB ON PQ NB longicornis atct. D. cristata (Harris) - SK MB ON atripennis (Say) fossata LeConte D. undecimpunctata howardi Barber oN PQ d uodec im p unclofa (Fabricius) D. virgifera virgifera LeConte ON filicornis Hotn Genus ACAI-YMMA Barber Key to Norbh American species: Munroe and Smith (1980) A. gouldi Barber _ONPQ A. vittatum (Fabricius) MB ON PQ NE NS atnericanutn (Gmelin) melanocep halurz ( Fabrici us) pallipes (OIivier) stolatum (Gmelin) Genus CEROTOMA Chevrolat Andrector Horn Cerotana Bowditch Ceratorna Doziet C. trifurcata (Forster) MB ON PQ 315 ChrysomeliCae Genus PSELIDOLUPERUS Beller & Hatch P. lecontii (tlrotch) _BC P. longulus (LeConte) -BC baheri Beller & Hatch Genus SCELOLYPERUS Crotch S. cyanellusr (LeConte) ON NS S. nigrovirescens (FalI) -BCAB S. schwarzi Horn _BC S. torquatus (LeConte) _BC S. varipes (i-eConte) _BCAB concauus Beller & Hatch Genus PHI'LLOBROTICA Chevrolat P. decorata (Say) BC AB SK MB PQ NS P. discoidea (Fabricius) -oN circumdata (Say) P. limbata (Fabricius) -oN PQ Subfamilv ALTICINAE Genus BLEPHARIDA Chevrolat Key to North American species: Mignot (1971); Balsbaugh and Hays (1972) (Subgenus ULEPHARIDA s. srr.t (BlephuridulaWeise) B. rhois (Forster) AB SK MB ON atripennis }Iotn dorothea Mignot meticu'.osa (Olivier) roi (Ol vier) s/olido (Fabricius) Genus PHYLLOTRETA Chewolat Tanygctster Blatchley Key to North American species (part): Smith (1985) P. alberta Chittenden AB P. albionica (LeconteJ BC AB SK P. aerea Allardt ON P. armoraciae (Koch)t _SK ms oN PQ NB PE biplagiata Chittenden P. bipustule.ta t Fabricius t oN PQ P. brevipen'ris Chittenden MB P. chalybeil'ennis tCrotch t -NS P. conjunctrr Gentner AB SX MB ON PQ P. constrict r E.H. Smith nc AB SK pe P. crucifera : I Goeze tt AB SK MB oN NE NS columl: iana Chittenden P. decipiens Horn BC AB ordina;ia Chittenden P. denticon is Horn BC a m pl ic cr ni s Chittenden aequal.is Hatch P. inconspicua Chittenden BC-SKMB P. lewisii (Crotcht BC P. liebecki Schaeffer _MBON P. oblonga Chil,tenden _ABSKMB P. pusilla Horn -ABSKMBON P. ramosoides E.H. Smith NT BC P. robusta l,eConte _ABSKMBONPQ P. striolata (Fabricius)f - YK NT BC AB SK MB ON PQ NB NS artiu ittata Chittenden lineolata Chittenden uerniccsa Chittenden uillolo (Fabricius) P. utanula li.H. Smith BC P. zimmerrnanni (Crotch)* AK - NT BC AB SK MB ON PQ NB NS PE NF sinuatu Horn 316 Chrysomelidae Genus APHTHONA Chevroiat Cerataltica Ctotch A. cyparissiae (Koch)t AB?_ON-? A. flava Guillebeaut AEr I ',' _ ') A. nigriscutis Foudrast AB_MBON Genus LONGITARSUS Berthold (Subgenus LONGITARSUS s. str.) (Apterius Blatchley) (T einodacty la Chevrolat) (Thyamis Stephens) L. erro Horn AB-MBONPQ NS L. fermgineus (Foudras)t YBC -oNPQ PE NF L. ganglbaueri Heikertingert MB L. insolens Horn AB ON PQ L. jacobaeae Waterhouset -nc L. luridus (Scopoli)f -oNPQ NU NS L. melanurus (Melsheimer) MB ON PQ v L. pallescens Blatchley -oN L. pellucidus (Foudras)t _ONPQ L. pratensis (Panzer)t _ONPQ PE L. rubiginosus (Foudras)f _ONPQ L. succineus Foudrasf NT L. suspectus Blatchley -oN L. testaceus (Melsheimer) _ONPQ NB Genus GLYFTINA LeConte Balophila Ctotch G. abbreviata Gentner ON G. atriventris Horn BC sx lle ON G. brunnea Horn T _MB ON PQ G. cyanipennis (Crotch) ON G. spuria LeConte _MB ON PQ lissotorques LeConte Genus SYSTENA Chevrolat S. bitaeniata (LeConte) AB S. blanda (Melsheimer) AB_MBONPQ ligata (.LeConte) ochracea (LeOonte) taeniata (.Say) S. carri Blake _AB S. corni Schaeffer oN- S. dimorpha Blake _AB s^- S. elongata (Fabricius) _MB floridana Blatchley subdenea (LeConte) S. frontalis (Fabricius) _MBONPQ S. hudsonias (Forster) -MBONPQ S. marginalis (Illiger) ON PQ oblonga (LeConte) Genus STRABAJ-A Chevrolat (Subgenus STRABAI-A s. str.) Key to New World species: Blake (1953) S. rufa (Illiger) oN PQ Genus AJ-TICA O.F. Miiller (Subgenus ALTICA s. str.) (.Gr a p toder a Chevrolat) (Megaltica Hatc}l':' A. aeneola Blatchley -oN PQ aenescens Blatchley A. ambiens LeConte NT BC MB ON PQ NS alni Hatris latiplicata Blake A. betulae Schaeffer PQ 317 ChrysomeLidae A. bimargirrata Say AK YK NT BC AB SK MB ON PQ LB p licipe nnis ( Mannerheim) A. browni }4ohamedsaid MB ON PQ NB NS potentillae W.J. Brown A. canadensis Gentner AB SK MB A. chalybar,ra Illiger -oN PQ uitiuota Thomas A. carinata Germar MB ON PQ NB e*apta Say A. caurina Blake BC AB _MB A. corni Woods BC -MBONPQ A. elongatula Csiki _AB _MBON elongata Gentner A. foliaceae LeConte MB ON puncti.oennis LeOonte A. heucherae FaiI MB A. ignita Illiger -oN A. inaerata LeConte NT MB A. kalmiae Melsheimer PQ NB NS uirida:za Schaeffer A. litigah lrall -oN A. populi S'.J. Brown SK yu oN PQ NB A. prasina LeOonte BC AB SK A. probata Fall A. ribis W.,1. Brown :-BC__ -oN A. rosae Woods MB ON PQ A. subplica;a LeConte _ONPQ A. sylvia Malloch -oNPQ NU NS cuprascens FalI torquo.ta LeConte A. tincta LeConte -BC A. tombacina (Mannerheim) AK YK NT BC AB SK l,fB ON Pe NU WS LB euicta LeConte shoemtkeri Schaeffer A. ulmi Wocds MB ON PQ A. vicaria Ilorn AB A. woodsi Iiely -oN Genus HOIINAJ-TICA Barber H. atriventris (Meisheimer) ON PQ Genus ORT'HALTICA Crotch Leptot,brix Heikertinger & Csiki Le ptot. icha I ti co Heikertinger Leptot,ix Horn M ic re p itr ix Laboissidre Serraticollis R.E. White Key to North American species: Scherer (1974) O. copalina (Fabricius) oN PQ fortico.nis (Illiger) Genus DEFiOCREPIS Weise D. erythroJ'us (Melsheimer) PQ rufipes (Ctotch) D. oregoni .{atch BC Genus ORIISTIA Germar Orestirtides Hatch PyladiaWeise O. robusta LeContet BC SK MB ON PQ NF Genus HIPPURIPHILA Foudras Key to North American species: Brown (19426) H. canadensis W.J. Brown AK -NT_ABSK y" oN PQ H. equiseti Beller & Hatch BC AB v H. manculzL (LeConte) AK BC AB SK modeeri anct. 318 Chrysomelidae Genus CREPIDODERA Chevrolat Key to North American species: Parry (1986) C. browni Parry -oN C. decora Parry -oNPQ C. digra Parry AK _ECAB s^ MB ON PQ r.rs lls C. heikertingeri (Lazorko) _BCAB SK MB ON PQ NB NS NF T\TF C. luminosa Parry _ONPQ NB NS C. nana (Say) NT BC AB SK MB ON PQ NB c hittende ni Heikertinger C. populivora Parry NT BC AB SK MB ON PQ NB NS C. sculpturata (Lazorko) _BC MB ON C. solita Parry _BCAB SK MB ON PQ C. spenceri (Lazorko) C. vaga Parry -oN C. violacea Melsheimer _ _ _:'_ _ONPQ Genus EPITRIX Foudras E. brevis Schwarz -oN E. cucumeris (Harris) _SKMBONPQ NS NS (?) pubescens llliget seminulum LeConte E. hirtipennis (Melsheimer) E. humeralis Dury E. subcrinita (LeConte) E. tuberis Gentner Genus MANTURA Stephens M. chrysanthemi (Koch)f M. floridana Crotch AB SK lrs on peNeNs--Nil Genus CHAXTOCNEMA Stephens (Subgenus CHAETOCNEMA s. str.) C. concinna (Marsham)'l _PE C. confrnis Crotch AB SK lrs om pe Ne NS flauicornis LeConte C. cribrata LeConte NT BC AB SK MB ON PQ NB NS NF rudis LeConte C. cribrifrons LeConte _ONPQ C. decipiens LeConte _SK C. denticulata (Illiger) -oN americana Motschulsky s emichalchea Melshermer texana Ctotc}r C. ectypa Horn _SKMB C. minuta Melsheimer MB ON PQ NB NS C. opulenta Horn AB SK MB C. perturbata Horn _SKMB C. protensa LeConte -nc AB SK MB ON PQ C. pulicaria Melsheimer v _BC AB ON (?) aeneola LeConte C. subcylindrica LeConte NT BC AB SK MB ON PQ cylindrica LeConte C. subviridis LeConte AB SK MB Genus DISONYCHA Chevrolat Key to North American species: Blake (1933) D. alternata (Illiger) BC AB SK MB ON PQ NB NS punctigera Schaeffer (? ) q uinq ueu ittaro ( Say) D. arizonae Casey MB ON borealis Blake douisi Schaeffer glabrata Jacoby D. latifrons Schaeffer BC AB SK MB ON PQ NB NS osferis Schaeffer laticollis Schaeffer D. latiovittata Hatch _BC D. pensylvanica (Illiger) ON PQ 319 Chrysomel,dae parua Blatch\ey pensyiuania Stwn (?) s e x,t ineata (Olivier) D. procera Casey NT BC A3 SK MB ON PQ n igriutntris Schaeffer (?) pallipes Crctch (?) uici.na l\trby D. punctigera LeConte NT AB SK MB 'neglecta Schaeffer punctipennis Schaeffer D. schaefferi Blake D. triangularis (Sayr - scABSf MB ON PQ montanensis Blake - puncticollis (Kirby) D. uniguttr.ta { Say t MB ON PQ uicina \{ttby D. xanthonrelas ( Dalman) NT AB SK MB ON PQ PE atrella Blake ceruicctlis LeConte collaris (Illiger) merdiuora Melsheimer xanthomelaeno Gemminger & Harold Genus KUSICHELINA Bechyn6 Key to North American species: Blake (1927) K. barberi (Blake) _AB (Forster) K. frmbriat.r - -4,8 circun cincta (Crotch) suture'Ia (Say) K. gibbitanra (Say) ON K. lugens (li,eConte) NT BC AB SK MB ON lamprt)cydnea (B1ake) K. perplexzi (Blake) ON K. thoracica. (Fabricius) ON flaua (Gmelin) K. vians (Il iger) NT BC SK MB ON PQ NS abdon',.inalis ( Olivier) badia Blake) scripti.:ollis (Say) Genus CAPRAITA Bechyn6 ChloephagaWeise Key to Nrrrth American species: Blake (1927) C. circumditta (Randall) ON limbalis (Melsheimer) (?) que"cata (Crotch) C. scalaris t,Melsheimer) oN PQ lobata (LeConte) C. sexmacuLata (Illiger) ON cretica (Jacoby) palliatt (RandalI) C. subvittata (Horn) AB SK MB oN PQ NB NS C. thyamoides (Crotch) ON Genus DISTIGMOPTERA Blake Key to North American species: Blake (1943) D. apicalis l3lake -oN D. borealis Blake BC AB SK MB ON PQ NB D. pilosa (Illiger) -oNPQ Genus DIBOLIA Latreille Key to N'rrth American species: Parry (1974) (Subgenus IDIBOLIA s. str.) D. borealis tlhevrolat AB SK MB _PQNBNS D. chelones Parr5r oNPQ-NS D. Iibonoti l{orn sc D. melampyri Parry T ONPQNBNS___ D. penstemrrnis Parry D. reyheria Mignot AB _MB 9*PQIUNS_-_ 320 Chrysomelidae Genus PSYLLIODES Berthold P. affinis (Paykull)t -oN P. chrysocephala (Linn6)t --Ne P. convexior LeConte -oN P. cucullata (Illiger)f PQ NB P. elegans Horn AB P. napi (Fabricius)f _ONPQ NB pn-*" P. picina (Marsham)-l _ONPQ P. punctulata Melsheimer AB MB ON PQ _NS I\TE] aenescens Casey extricata Casey paruicollis LeConte Subfamily HISPINAE Tribe Chalepini Key to North American genera: Butte (1968o) Genus ANISOSTENAWeise (Subgenus ANISOSTENA s. str.) A. californica Van Dyke _BC A. nigrita tOlivier) -BCMBONPQ NB Genus SUMITROSIS Butte Anoplitis Chapuis Key to North American species: Butte (1969) S. ancoroides (Schaeffer) AB SK MB S. inaequalis (Weber) - BC AB SK MB oN pe N" NS bazcis (Newman) flauipes (Germar) neruosa (Hotn') obsoleta (Say) pallida (Say) suturalis (Fabricius) S. rosea (Weber) SK MB ON PQ NB NS PE philemon (Newman) Genus ODONTOTA Chevrolat Key to North American species: Butte (1968c) O. dorsalis tThunberg) oN PQ hanisi Crotch scutellaris (Olivier) suturalis (Harris) O. mundulus (Sanderson) MB O. scapularis t Oliviert -oN lateralis (Say) Genus CHALEPUS Thunberg Anoplitis Kirby Key to North American species: Butte (1968b) C. bicolor (Olivier) ON rufiuentris (Suffrian) rufogaster Thunberg Genus BALIOSUS Weise B. nervosus (Panzer) ON PQ NB NS marginatus (Say) marmoratus (Baly) obsoletus (Say) pallipes (Germar) quadratus (Fabricius) quercifoliae (Harris) rubellus (Schonherr) ruber (Weber) 32L ChrysomelLdae Tlibe Uroplatini Genus STEiNOPODIUS Horn Key to North American species: Blaisdell (19386) S. lateralis lSchaeffer) AB SK uand.uzeei Blaisdell S. texanus (Schaeffer.t AB SK MB Genus BRACHYCORYNA Gu6rin-M6neville Key to North American species: Staines (1986) B. hardyi ((lrotcht - BC AB confusa Hatch horni'Weise melsheimeri Horn B. montana (Horn) AB SK MB ON Genus GL)?HUROPLATA llhmann Key to North American species: Riley (1985b) G. pluto (Newman) AB - MB ON PQ porcatd (Melsheimer) Genus MIC.IRORHOPALA Chevrolat Key to North American species: Clark (1983) M. excavat.r cyanea { Say } - BC AB SK MB M. excavatrr excavata (Olivier) MB ON PQ NB NS M. vittata tFabricius) - BC AB SK MB ON PQ NB laetuluLeConte M. xerene (Newman) AB SK MB ON PQ NB interrt.tpta Couper Subfamily CASSIDINAE Tribe Stolaini Key to genera: Hincks (1952) Genus CHINLYMORPHA Chevrolat C. cassidea (Fabricius) _SKMBONPQ orgus iLichtenstein) cribraria (Olivier) geniculata Boheman lewisi Crotch s e p t e rrtde ci m p u nc t at a (S ay) Tribe Physonotini Genus PHI'SONOTA Boheman Key to North American species: Sanderson (1948) (Subgenus PIfYSONOTA s. str.) P. helianthi tRandallt MB ON PQ q uinquepu nct ato (Walsh) sep ten,tr io nalis Boheman P. unipunctata (Say) - SK MB ON PQ T?ibe Cassidini Key to North American genera: Riley (1986a) Genus CASSIDA Linn6 Key to North American species: Riley (1986b) (Subgenus CASSIDA s. str.) C. azurea F'abriciusf - SK C. flaveola Thunbergt - YK NT BC AB SK MB ON PQ C. rubiginosa O.F. Miiller+ AB SK MB ON PQ NB Genus JON'THONOTA Spaeth J. nigripes .Olivier) AB SK MB ellipsis (LeConte) 322 Nemonychidae Genus AGROICONOTA Soaeth A. bivittata (Say) MB ON PQ striolata (Boheman) uittula (Boheman\ Genus PLAGIOMETRIONA Spaeth (Subgenus PARAMETRIONA Spaeth) P. clavata (Fabricius) ON PQ NB diu er s icoll is ( Schaeffer) Genus STRONGYLOCASSIS Hincks Strongylaspis Spaeth Key to species: Riley (1985o) S. atripes (LeConte) MB ON creolo (Sanderson & King) loedingi (Schaeffer) Genus DELOYALA Chewolat Chirida Chapuis D. guttata (Olivier) SK MB ON PQ NB PENS bohemani (Spaeth) columbica (Spaeth) cruciata (Fabricius) fuliginosa (OIivier) hamulata (Boheman) immunita (Boheman) immunda (Boheman) lucidula (Boheman) lugubrina (Boheman) p ennsy lu anic a ( Spaeth) signifera (Herbst) trabeata (Boheman) Genus CIIARIDOTELLA Weise (Subgenus CHAEROCASSIS Spaeth) C. purpurata (Boheman) SK MB ON PQ NS C. sexpunctata bicolor (Fabricius) -oN PQ NB NS auric halcea (Fabricius) auris plendens (Mannerheim) b istrip ustulata (Herbst) floridana (Schaeffer) (? ) mary landico ( Herbst) pallida (Herbst) Superfamily Curculionoidea Family NEMOI{YCHIDAE J. McNamara Nemonychidae comprises eight species in Anthribidae. Kuschel (1989) recently published a Canada. The adults feed on the staminate flowers revision of this family for North America The list ofvarious conifers. below follows O'Brien and Wibmer (1982). The nemonychids are often included in Curculionidae, but their larvae show an aflinity to BC (6); AB (3);MB (1); ON (2);PQ (3); NB (1) 323 Anthribidae Subfamily DOYDIRFIYNCHINAE Tribe Cimberidini Genus CIMBERIS des Gozis C. bihirsuta Hatch BC C. compta (LeConte) BC paruulus lJatch C. decipiens Kuschel BC C. eiongata (LeConte) BC MB ON PQ C. pallipennis (Blatchley) AB PQ C. pilosa (LeConte) -oN PQ NB Genus PITYOMACER Kuschel P. pix Kuschel BC AB Genus ACFI OMACER Kuschel A. bombifrr: ns (LeConte) BC AB Family ANTHRIBIDAE fungus weevils J. McNamara Anthribidae includes 18 species in Canada and lar habitats. Adults feed on fungr, pollen, or even Alaska. One of these, the coffee bean weevil bark; larvae feed in plant stems, fungr, orvvoods. (Araecerus fasciculatus), is introduced and may be Valentine (1960) has revised the North establishred in warehouses; this species damages American genera of Anthribidae. many stored plant materials, including seeds. Adults of anthribids are often collected by beating AJ{ (2); YK (2); NT (2); BC @); AB (8); S.K (6); MB dead or diseased vegetation or by sweeping in (7); ON (12); PQ (12); NB (1);NS (1);NF (1) o fl weedy areas. The larvae are usually found in simi- (1); I (1) Subfamily CHORAGINAE T?ibe Araecerini Genus ARAECERUS Schrinher"r A. fasciculatus (DeGeer)t BC ON PQ Tlibe Choragini Genus EIIXENUS LeConte E. punctatus LeConte PQ Subfamily ANTHRIBINAE TYibe Eupariini Genus EUPARIUS Schdnherr Key to Norlh American species: Valentine (1960) E. marmoreus (Olivier) MB ON PQ NB albifrons (Boheman) lunatus (Fabricius) Tlibe Anthribini Genus TRICIONORHINUS Wollaston Anthrihulus LeConte Brachy tarsoides Pierce Key to wrstern North American species: Vaientine (1971) T. alternatus (Say) AB MB ON T. annulatus (Carr) _BC AB 324 Ithyceridae T. griseus (LeConte) T. limbatus (Say) - - - - ffi3f reoNpe - - - - - T. sticticus (Boheman) NT BC AB SK MB ON PQ obsoletus (Fahreus) uariegatus (Say) T. tomentosus tomentosus (Say) ON 6reuis (Fahreus) paululus (Casey) Tribe Ormiscini Genus ORMISCUS Waterhouse Eusphyrus LeOonte Gonops LeConte Toxotropis LeConte O. saitator LeConte oN PQ angulatus Pietce O. walshii (LeConte) ON Tribe Tropiderini Genus TROPIDERES Schbnherr Eurymycter LeConte Gonotropis LeConte 'Key to North American species: Valentine (1960) T. dorsalis (Thunberg)* AK YK NT AB SK MB PQ NF gibbosus (LeConte) T. fasciatus (Olivier) _BC SK MB ON PQ bicarinatus (Pierce) T. latifascia (Pierce) NS T. tricarinatus (Pierce) _ _ _ oNiS - Tribe Allandrini Genus AI,LANDRUS LeConte Key to North American species: Valentine (1960) A. bifasciatus LeConte A. brevicornis Frost ----AB-nreoNPQ----- A. populi Pierce AK \'K AB SK _ ON PQ Family ITIIYCERIDAE New York weevil J. McNamara Ithyceridae includes only one species. The Sanborne (1981) studied the morphology of both adults feed on the tender shoots and buds ofvarious larvae and adults, as well as the biology of the hardwoods and occasionally on some cultivated fruit species. trees. The larvae feed on the roots of the host plants. ON (1); PQ (1) Genus ITHYCERUS Schonherr Pochyrhynchus Kirby L noveboracensis (Forster) oN PQ cur culionoides (Herbst) p unc ta t uI us ( Fabricius) schoenherri (Kirby) 325 Rhynchitidrre FamilyATTELABIDAE leaf-rollingweevils J. McNamara Four species of Attelabidae occur in Canada. The classification used here follows that of The females form rolls from leaves and lay a single Robert W. Hamilton (unpublished Ph.D. thesis, egg in each roll. The larvae feed on the inner part 1969), who revised the North American species. of the roll. SK (1); MB (2); ON (3); PQ (a); NB (1); NS (2) T?ibe Attelabini Genus HIMATOLABUS Jekel H. pubescerrs (Say) MB ON PQ NB NS colora<:loensis Yoss d,is p ar.'.p e s (Chittenden) maculutus (Provancher) rhois (:3oheman) Genus HOI\{OEOLABUS Jekel H. analis (Illiger) SK MB ON PQ similis (Kirby) Genus ATTELABUS Linn6 (Subgenus IiYNOLABUS Jekel) A. bipustulatus Fabricius -ONPQ_NS A. nigripes,J.E. LeConte PQ genalis LeConte Family RHYNCHITIDAE leaf and bud weevils J. McNqmara Rhynchitidae comprises 13 species and sub- family, but only parts of his work have been pub- species in Canada. Some lay their eggs in the lished to date; he also wrote a key to the North young fruit or buds of trees that are then destroyed American genera ( 1971). by the larvae; others feed on blossoms or foliage. Robert W. Hamilton (unpublished Ph.D. thesis, NT (3); BC (5); AB (6); SK (5); MB (6); ON (7); PQ 1969) revised the North American species of the (5);NB (3);NS (3); PE (2);NF (2) Subfamily PTEROCOLINAE Genus PTEROCOLUS Say ApotonLus Kirby P. ovatus (Fabricius) ON Subfamily RIIYNCHITINAE Tribe Rhynchitini Genus EUGNAMPTUS Schttnherr Key to Norih American species: Hamilton (1990) E. angustatus (Herbst) ON abdominalis Yoss collaris (Fabricius) fuscipes Pierce Nigripennis ( Fabricius) nigripes Pierce rubricollis (Say) ruficeps Pierce 326 Brentidae ruficollis (Germar) sulcifrons Gyllenhal Genus PSELAPHORHYNCHITES Schilsky Key to North American species: Hamilton (1971) p. cyanellus (LeConte) NT - AB sK MB ON PQ NB NS PE - NF P. perplexus (Blatchley) - ON PQ NB NS PE Genus MERHYNCHITES Sharp Key to North American species: Hamilton (1985) M. bicolor bicolor (Fabricius) NT BC AB SK MB oN PQ antennalis (Yoss) intermedius (Yoss) M. bicolor cerdonis (Voss) - BC AB M. bicolor cockerelli (Pierce) AB erythrosoma (Cockerell & Harris) M. bicolor nigricephalus Hamilton - BC M. wickhami wickhami (Cockerell) NT BC AB SK MB homocidae (Yoss) mandibularis (Yoss Genus HAPLORHYNCHITES Voss Key to North American species: Hamilton (1974) H. aeneus (Boheman) - SK MB consobrinus (Yoss) TYibe Auletini Genus AULETOBIUS Desbrochers Key to North American subgenera: Hamilton (1983) (Subgenus ALETINUS Desbrochers) A. ater (Leconte) MB oN PQ (Subgenus MESAULETES Voss) A. cassandrae (LeConte) - ON PQ NB NS NF albou e stitus (Blatchley) blatchleyiYoss minor (Blatchley) (Subgenus AULETOBIUS s. str.) Metopon Waterhouse A. congruus (Walker) - BC AB SK MB subcoeruleus LeOonte Family BRENTIDAE brentid beetles J. McNamara Only one species of Brentidae, the oak timber- into the heartwood, often causing serious injury to worm, occurs in Canada. The adults are found oak stock. under bark or in decayed wood. The larvae bore ON (1); PQ (1) T?ibe Arrhenodini Genus ARRHENODES Schdnherr A. minutus (Drury) - ON PQ brunneus (Panzer) lecontei (Power) maxillosus (Olivier) p aup er c ula (Blatchley) sallei (Power) s epte ntrionis (Herbst) 327 Apionidae Family APIONIDAE pear-shaped weevils J. McNamara Thirty-five species of Apionidae are recorded in Canada and Alaska. The larvae feed on leaves and stems, carrsing galls, or develop within the seeds of various plants. The adults usually feed on leaves. Than}s to the work of Kissinger (1968), the tax- aK (3); vK (3); NT (3); BC (12); AB (20); SK (13); onomy of t;he North and Central American species is MB (15); ON (17); PQ (13); NB (2); NS (2); NF (1) well known. . I(1) Tlibe Nanophyini Genus NAN{)PHYES Schonherr Nanoda cty lus Blatchley Nanodes Schtinherr P se udo ty c hius Blatchley (Subgenus NANOPHYES s. str.) N. canadens s W.J. Brown AB SK MB T?ibe Apionini Genus APION Herbst (Subgenus ALOCENTRON Schilsky) A. attenuatum Smith BC A,E MB ON PQ (Subgenus C ERATAPION Schilsky) A. contusum Smith PQ notabile, Buchanan spinipes Fall A. extensum Smith SK (Subgenus C OELOCEPHALAPION Wagner) A. carinatun. Smith ON concoloratum Srlrilh A. emaciipes Fall ON A. segnipes Siay ON (Subgenus E UTRICHAPION Reitter) A. alaskanun Fall AK YK NT BC AB A. cavifrons .-eConte _BCAB -pe A. cyanitinctum Fall AK YK NTBCAB_MB A. huron Falt AB_MB ON (Subgenus F,{ILAPION Kissinger) A. finitimum FaIi BCT SK MB ON PQ NB A. impeditum Fall _ _ PQ A. melanarium Gerstaecker -oN PQ A. occidentale Fall _AB A. pennsylvanicum Boheman BC AB SK mn oN PQ erythrocerum Srlrith A. robustum Smith AB SK obesum Srrlith A. speculifenrm Fall AB SK MB (Subgenus L-APION Roudier & Tempere) (ftios Deville) A. frosti Kissinger -oNPQ NS A. idiastes K.,ssinger -ONPQ (Subgenus PERAPION Wagrrer) A. punctinasrm Smith BC AB SK MB ON A. wickhami Kissinger _ABSK 328 Curculionidae (Subgenus PSEUDAPION Schilsky) A. amaurum Kissinger -AB_MB A. capitone Kissinger _ABSKMB A. disparatum Sharp _AB_MB nasutum Fall (Subgenus RHOPAIAPION Schilsky) A. longirostre Oliviert BC ON PQ (Subgenus TRICHAPION Wagner) A. centrale Fall AB SK MB ON A. commodum Fall _:" A. porcatum Boheman TSKMB ON A. proclive LeConte -BC crassinasum LeConte nunenmacheri Wagner A. reconditum Gyllenhal -oN PQ turbulentum Smith A. simile Kirby BC AB SK MB ON PQ NB NS NF lanuginosum. Walsh uicinum Srrrith walshii Smit}r A. tenuirostrum Smith AB SK MB $ A. antennatum Smith AK YK BC Tlibe Aplemonini Genus PODAPION Riley P. gallicola Riley ON PQ alemnderi Sleeper Family CURCULIONIDAE snout beetles or weevils J. McNamara The family Curculionidae includes 609 species the North American species with supplements in and subspecies in Canada and Alaska. AII of them 1984 and 1989, the latter under Wibmer and are phytophagous. Most of the larvae live inside O'Brien. The classification used here follows that the tissues of the host plant, but some feed exter- checklist. nally on the leaves. The adults feed on leaves, pollen, flowers, developing fruit, and fungi, and some burrow in wood; they do not necessarily feed on plants preferred by the larvae. Many weevils are host-specifrc and some are economic pests. AK (61); YK (51); NT (55); BC (261); AB (206); SK Kissinger (1964) wrote a key to the genera. (I24); MB (179); ON (312); PQ (280); NB (72); NS O'Brien and Wibmer (1982) published a checklist of (91); PE (17); LB (11); NF (68) ' H (14); I (69) Subfamily BRACITYDERINAE Tribe Naupactini Genus ASYNONYCHUS Crotch A. cervinus (Boheman)t ON fulleri (Il.orn) 329 Curculionidae Tribe Barynotini Genus TRI(IONOSCUTA Motschuisky Key to North American species: Pierce (1975) (Subgenus I)RIGONOSCUTA s. str.) T. pilosa iritlescens Pierce - BC Genus STAMODERES Casey S. lanei {Van Dyke) - BC Genus BAR'INOTUS Germar Key to Nc'rth American species: Brown (1950) B. moerens Fabricius)t NS B. obscurus (Fabricius)t - BC PQ NB NS PE - NF B. schoenherri (Zetterstedt)t PQ NB NS NF Tribe Polydrosini Genus POLYDRUSUS Germar Cyphontimus Horn Poly drt ts u s Schcinherr Key to Ncrth American species: Sleeper (1957a) P. americanrs (Gyllenhalt - ON PQ d.orsali:; (Horn) P. cervinus,Linn6tt PQ P. impressif'rons (Gyllenhal)+ - - MB ON PQ - NS P. sericeus ( Schallertf - ON PQ NB Genus PACI{YRHINUS Schrinherr S cy t hrt:. p u s Schdnherr P. californic -rs (Horn) - BC P. elegans(Oouper) - BC AB ON PQ - NS metall i c us (Desbrochers) P. ferrugine:s (Casey) - BC Tlibe Brachyderini Genus BARYPEITHES Jacquelin du VaI & Fairmaire B. pellucidus (Boheman)t - BC - ON PQ - NS PE - NF Genus SCIAPHILUS Schtinherr S. asperatus (Bonsdorff)t - BC - ON PQ NB NS NF Genus BRAOHYSOMUS Schonherr Thricol zpoide s O'Brien B. echinatus (Bonsdorfl)* PQ NF clauisetis (O'Brien) Tribe Strophosomini Genus STROPHOSOMA Billberg S. melanogrammum (Forster)f - BC - ON PQ - NS NF Tlibe Cneorhinini Genus PHILOPEDON Schtinherr P. plagiatunL (Schaller)t _NBNSPE-NF 330 Curculionidae Tribe Sitonini Genus SITONA Germar S. californicus (Fahraeus) BC angustulus (Casey) apacheanus (Casey) cockerelli Blaisdell explicatus (Casey) ertrusus (Casey) hispidiceps (Casey) oregonensis Tanner prominens (Casey) S. cylindricollis (Fahraeus)t _BCABSK MB ON PQ NB NS alpinensis Tanner S. hispidulus (Fabricius)t -BC oN PQ NB NS S. lepidus Gyllenhal f_ _BC AB MB ON PQ NB NS NF S. Iineatus (Linn6)t -BC S. lineellus (Bonsdorfi)t NT BC AB sr MB ON NB NS -YK _BC S. lupinus Sleeper pe S. scissifrons Say AK_ NT BC AB SK ln" oN lln NS NT uittatus (Couper) S. tibialis (Herbst)t AK _BC AB _ONPQ NF T[ibe Tanymecini Genus MINYOMERUS Horn Elissa Casey Pseudelissa Casey M. innocuus Horn AB Genus TANYMECUS Germar T. confusus Say NT-ABSK MB confertus Gyllenhal uariabilis Fahraeus Genus PANDELETEIUS Schdnherr Key to North American species: Howden (1959) P. hilaris (Herbst) _ONPQ pauperculus Gyllenhal Subfamily OTIORFIYNCHINAE Tlibe Otiorhynchini Genus OTIORIIYNCHUS Germar B r ac hy r hinu s Latreille Key to North American species: Warner & Negley (1976) O. desertus Rosenhauerf NF O. Iigneus (Olivier)t NSNS- NF O. ligustici (Linn6)t -Olf- O. ovatus (Linn6)t AK YK NT SC AB SK MB ON PQ NB NS LB NF O. porcatus (Herbst)t PQ NF O. raucus (Fabricius)t -oN ___ (Gyllenhal)f PQ NB NS NF O. rugifrons _NS O. rugosostriatus (Goeze)f -sc- -ONPQ O. scaber (Linn6)t _NS O. singularis (Linn6)l _BC -ONPQ NB NS NT picipes (Fabricius) O. sulcatus (Fabricius)t AK _BC _MBON PQ NB NS PE NF Genus SCIOPITHES Horrr S. obscurus Horn BC angustulus Casey brumalis Casey significans Casey 331 Curculionidae Genus AGRONUS Horn A. carri Bu<.hanan - BC AB Tribe Peritelini Genus OMIAS Germar O. erectus Flatch - BC O. minor Hatch - BC O. saccatus {LeconteJ - BC Genus PERITELINUS Casey P. oregonus Van Dyke - BC P. variegatus Casey - BC Genus GEODERCODES Casey G. latipennis Casey - BC Genus NEI!-OCESTES Van Dyke Key to North American species: Van Dyke (1936) N. horniVanDyke - BC - ON PQ NB NS N. incomptr.s tHornt - BC N. montanus Van Dyke - BC N. puncticollis tCaseyr - BC sordidt:s Van Dyke N. tuberculi,tus Van Dyke - BC Genus PAR,\PTOCHUS Seidlitz P. sellatus ( BohemanJ BC califorr'.icus Seidlitz Tribe Trachyphloeini Genus TRAT,IHYPHLOEUS Germar Chaete':.:hus }lorrl Key to Ncrth American species: Brown (1965) T. aristatus tGyllenhal r* - ON T. asperatur Bohemant - ON NS setiger:.Hotn) T. bifoveolatus (Beck)f - BC AB ON PQ NB NS PE - NF doujsi l3latchley Genus CER{IOPEDIUS Sleeper C. artemisiae (Pierce) - BC paruultls (Van Dyke) Tlibe Hormorini Genus HORMORUS Horn H. undulatus (Uhler) AB - MB ON PQ NB NS Genus AGAIIPIIAEROPS Horn A. niger Hor n - BC sulcirostris (Pierce) Tlibe Phyllobiini Genus PHY,LOBIUS Germar P. glaucus tiicopoli t+ P. oblongus r.Linn6)f - BC - ON PQ NB - PE Genus EVOIIUS LeConte E. naso (Le(ronte) NT BC AB SK 332 Curculionidae Genus APHRASTUS Say M icrony c h us Provancher A. taeniatus Say ON PQ sulcatus (Provancher) Tribe Leptopiini Genus PHYXELIS Schdnherr Geoderces }Iotn P. rigidus (Say) oN PQ NB NS PE NF glomerosus Boheman melanothrix (Kirby) setiferus Boheman Genus LEPIDOPHORUS Kirby Lophalophus LeConte Key to North American species: Buchanan (1936b) L. inquinatus (Mannerheim) AI{ BC L. lineaticollis Kirby AK YK NT BC L. pumilus Buchanan -BCAB Genus CIMBOCERA Horn C. pauper Horn AB Genus PLINTHODES LeConte P. taeniatus (LeConte) BC Genus TRICHALOPHUS LeConte T. alternatus (Say) _YK-BC AB MB ON T. brunneus Van Dyke AK BC T. didymus (LeConte) AK BC AB SK MB constrictus (LeConte) T. seriatus (Mannerheim) AK T. simplex LeConte NT AB SA MB T. stefanssoni Leng AK_NT Genus LEPESOMA Motschulsky Amnesia Horn Dyslobus LeConte Melamomphus l{orn Thricomigus Horn Key to North American species: Van Dyke (1933) L. alternatum (Horn) BC L. bakeri (Van Dyke) BC YSK L. decoratum (LeOonte) BC L. granicolle granicolle (LeConte) BC debilis (Casey) discors (Casey) sculptilis (Casey) L. lecontei (Casey) BC L. luteum (Horn) BC SK L. raucum (Horn) BC ciliatum (Pierce) L. tanneri (Van Dyke) BC L. ursinum (Horn) BC L. verruciferum (Casey) BC b ituberculatum (Pierce) Genus PANSCOPUS Schdnherr Key to North American species: Buchanan (1936o) (Subgenus NEOPANSCOPUS Pierce) P. aequalis (Horn) BC AB SK MB uestitus (Casey) (Subgenus NOCHELES LeConte) (Panscopidius Pierce) P. torpidus (LeContet BC 333 Curculionidae (Subgenus IIIOMIDUS Casey) P. abruptus (Casey) - BC P. ovalis Pi,rrce - BC AB SK P. pallidus Suchanan - BC AB P. rugicollis Buchanan - BC (Subgenus I)ANSCOPUS s. str.) P. erinaceu,r (Say) - ON PQ carinat:us Pierce (Subgenus PARAPANSCOPUS Buchanan) P. maculosu s Blatchley - ON P. ovatipennis Buchanan - ON (Subgenus PlIfMAfl NUS LeConte) P. gemmatu.s (LeConte) - BC (Subgenus PSEUDOPANSCOPUS Buchanan) P. costatus Buchanan - BC Genus CON NATICHELA Anderson C. artemisir,Le Anderson AK YK NT Genus ANAMETIS Horn A. granulatrr (Say) - BC AB SK MB ON PQ NF grisea':Jorn Genus TRO PIPHORUS Schrinherr T. elevatus Herbst)t - NF carinatus (O.F. Mtiller) T. obtusus (Bonsdorff)t NS NF T. terricola.Newman)l PQ - NS PE - NF Tribe Ophryastini Key to North American species: Kissinger (1970) Genus OPIIRYASTES Say AmydrDgmus Pierce Eupagoderes Horn ?ososfes Sharp O. cinerascens (Pierce) - BC columbianus (Van Dyke) O. sulcirost:ts (Say) NT - AB SK MB ligatus LeConte O. tuberosu; LeConte AB collaris Champion Subfamily EREMNINAE Tribe Cyphicerini Genus CYR'IEPISTOMUS Marshall C. castaneu; (Roelofs.tt - ON Genus CAI-(JMYCTERUS Roelofs C. setarius lloelofsf - ON Subfamily RIIYTIRRHININAE Genus LISTRONOTUS Jekel Hyperodes Jekel MacroP'9l{n"rY Key to Nc,rth American species: Henderson (1940); O'Brien (1981) L. alternatus (Dietz) PQ NB L. anthracir.us {Dietz) - ON PQ 334 Curculionidae L. appendiculatus (Boheman) BC SK MB ON PQ floridensis Blatchley impressus Van Dyke leucozonatus Chittenden L. californicus (Dietz) -AB prl L. caudatus (Say) BC ON L. delumbis (Gyllenhal) BC AB SK MB ON pa:) indistinctus (Dietz) schwarzi (LeConte) solutus (Boheman) spurcus (Boheman) L. echinodori O'Brien _MB L. frontalis LeConte EC AB SK_ON PQ L. humilis (Gyllenhal) A"- BC PQ NF subcribratus (Dietz) L. maculicollis (Kirby) AK YK BC AB PQ mirabilis (Dietz) myasellus (Dietz) L. montanus (Dietz) AB SK MB L. oregonensis oregonensis (LeConte) _MBON PQ impressifrons LeConte rudipennis Blatchley L. oregonensis tessellatus Casey AB L. porcellus (Say) BC PQ minitnus (Blatchley) L. punctiger LeConte BC AB gracilis LeConte leechi Sleeper L. sordidus (GyIlenhal) ON PQ disting uendu s (Gyllenhal) obliquus LeConte L. sparsus (Say) NT_AB _MBONPQ inbellis (Dietz) itnmundus (Boheman) latiusculus (Boheman) lineatulus (Say) longulus (Dietz) squalidus (Gyllenhal) L. squamiger (Say) BC AB SK MB ON PQ inaeq uali pennis (Boheman) L. truncatus (Hatch) BC L. tuberosus LeConte _MBON Pc) L. vitticollis (Kirby) AB SK MB interp unctatulus (Dietz ) inter s tit ialis (Dietz) tenebrosus (Dietz) ulkei (Dietz) Genus LIXELLUS LeConr,e Anchodemus LeConte L. filiformis LeConte L. Iutulentus (Boheman) - Tf'BCfsKMBoNiS- - - - - angustus (LeConte) Genus BYRSOPAGE S Schdnherr B. carinatus Motschuisky AK Subfamily THECESTERNINAE Genus THECESTERNUS Say Lithodus Germar T. affinis (LeConte) AB eroszs (LeConte) morbillosus (LeConte) rectus (LeConle) rudis (LeConte) T. humeralis (Say) AB 335 Curculioni lae Subfamily FIIPERINAE Tribe Hyperini Genus HYPERA Germar Phytonomus Sch6nherr H. castor (LeConte) -sKMB-PQ H. compta (Say) -BC-ONPQ diuersz (Gyilenhal) H. diversipunctata (Schrankrx AK - AB SK - ON H. eximia (LeConte) H. meles (['abricius)t - - - - Y- ItuSNpeN"Ns- - - H. nigriros iris (Fabricius)i -BC-ONPQNBNSPE-NF uiridi.. (Provancher) H. postica iGyllenhal)f ABONPQ-NS H. pubicoli.s (LeConte) _BC H. punctatir (Fabricius)t -BC-ONPQ-NSNF opimus (LeOonte) H. quadricollis (LeConte) AB-MB H. rumicis (Linn6)+ AB SK MB ON H. seriata (Mannerheim) AK YK H. trivittata (Say) NTBCAB_MB setigera (LeConte) Subfamily CLEONINAE TYibe Cleonini Key to New World species: Anderson (1988) Genus CLtr.ONIS Dejean Cleont..s Sch Genus CLEONIDIUS Casev Lirestt.:s Reitter C. calandrordes tRandall) pe NB C. erysimi tFallr - BC AB - MB C. frontalis (LeConter AB SK MB sparsus (LeConte) sfralus (Csiki) C. longinasus R.S. Anderson - BC mixtus (LeConte\ C. poricollis (Mannerheim) - BC AB SK hirbyi (Casey) lobigerinus (Casey) uirgatu.s (LeConte) uittatus (Kirby) C. puber"ulus (Lecontet AB - MB C. trivittatr:s (Sayt - BC AB SK MB bicarinatus (Casey) inornai'.us (LeConte) praepotlens (Say) 336 Curculionidae Tribe Lixini Genus LIXUS Fabricius L. caudifer LeConte L. concavus Say -:"--MBoNpe- coloradensis Chittenden crassulus Notman rectus LeCorte soror Casey L. laramiensis Casey AB bischoffi Fall L. macer LeOonte -oN blatchleyi Csiki capitatus Chittenden cauicollis Blatchley nouellus Blatchley L. mucidus LeConte ON PQ morulus Blatchley L. parcus LeConte AB ON pygmaeus Casey L. punctinasus LeOonte albisetiger Chittenden ele phantu lus Chittenden planicol I is Chittenden L. rubellus Randall NT BC AB SK MB ON PQ auctus LeConte L. terminalis LeConte AB_MBON blakeae Chittenden Tlibe Rhinocyllini Genus RHINOCYLLUS Germar R. conicus (Frolich)f _SK_ONPQ Subfamily MOLYTINAE Tlibe Emphyastini Genus EMPHYASTES Mannerheim E. fucicola Mannerheirn AK BC Genus THALASSELEPHAS Egorov & Korotyaev Neophycocoetes O'Brien & Wibmer Phycocoetes LeConte T. testaceus (LeContet - BC Tlibe Hylobiini Genus LEPYRUS Germar Key to North American species: Van Dyke (1928b) L. capucinus (Schaller) NT L. gemellus gemellus Kirby AK YK NT BC L. Iabradorensis Blair NT PQ - - is- L. nordenskioeldi alternans Casey LB L. nordenskioeldi caesius Csiki AK YK cinereus Van Dyke L. nordenskioeldi canadensis Casey AI{ NT BC AB SK L. oregonus oregonus Casey AK _BC perforatus Casey L. oregonus tessellatus Van Dyke _BCAB L. palustris (Scopoli)x NTBCAB_MBONPQ is- colon (Lint6) geminatus Say Genus IfYLOBIUS Germar Hypomolyx LeConte Key to North American species: Millers, Benjamin, and Warner (1963) 337 Curculioni*1ae H. congener Dalla Torre, AK YK NT BC AB SK MB ON PQ NB NS LB NF Schenkling & Marshall confusus Ilttby H. pales (Ilerbst) AB oN PQ NS macelous (Germar) H. pinicola (Couper) _YKNTBC AB SK MB ON PQ NB NS NF H. radicis Iluchanan MB ON PQ :" H. warreni Wood -BC AB MB ON PQ NB NS NF Genus PACIIfYLOBIUS LeConte P. picivoru:; (Germar) ON LB stupitius (Boheman) Tribe Plinthini Genus STEREMNIUS Schcinherr Parap,t,inthus Faust S. carinatus (Boheman) AK BC S. tuberosus Gyllenhal AK BC s crob ic ulatus (Mannerheim) Genus STFI EREUS Motschulsky Lobosuma Buchanan P hilo s lrat u s Zimmerman Key to Nrrth American species: Zimmerman (1g64) S. horridus (Mannerheim) AK _BC S. multitutrerculatus Buchanan AK _BC S. ptinoides; (Germar)x AK _BCNS I\TIT boreaL,s Motschulsky fas cic u.Iatu s Motschulsky S. quadritu.berculatus Motschulsky AK BC SK Genus GASTROTAPHRUS Buchanan G. barberi ]luchanan BC Subfamily ACICNEMIDINAE Genus TRACHODES Germar T. hispidus (Linn6)t NF Subfamily PISSODINAE Genus PISIIODES Germar P. affinis Rrrndall _NTBCAB ON PQ NSrtlB NF czrriel Hopkins P. costatus Mannerheim AK BC P. fasciatus LeConte BC P. fiskei Hopkins rasoNFeNs---- P. nemorensis Germar 5_:cysK MB ON PQ NB NS approrimatus Hopkins canadensis Hopkins deodarae Hopkins P. radiatae Hopkins -BC P. rotundatus LeConte AK YK _BCAB_MBONPQ olu Ns - alascensis Hopkins nigrae Hopktns P. schwarzi Hopkins BC AB yosemite Hopkins P. similis Hopkins BC -PQ _NS NF utahensis Hopkins P. striatulus (Fabricius) BC ON PQ NB NS NF dubius Ratdall froserl Hopkins prperi Hopkins P. strobi (Peck) BC AB SK MB ON PQ NB NS NF 338 Curculionidae engelmanni Hopkins sitchensis Hopkins P. terminalis Hopping BC Subfamily ERIRHININAE TYibe Erirhinini Genus DORYIOMUS Germar Alycod,es Dietz Key to North American species: O'Brien (19706) D. brevicollis LeConte ON PQ dubius (Dietz) fuscipes Casey indifferens Casey D. frostii Blatchley YK NT BC AB SK MB ON PQ D. hirtus LeConte -BCAB columbianus Sleeper hispidus LeConte nubeculinus Casey D. imbecillus Faust AK YK NT AB oN PQ alaskanus Casey subsimilis Blatchley D. inaequalis Casey -BCAB SK MB D. laticollis LeConte AK BC AB sK oN PQ D. lecontei O'Brien AK Yu D. leucophyllus (Motschulsky) AK YK NT BC adustus Motschulsky bajulus Farst longulus LeConte s ub sig natus (Mannerheim) te s sel latu s Motschulskv D, luridus (Mannerheim) AK YK NT BC AB SK _PQ _NS cuneatulus Casey D. mannerheimi (Gemminger) AK NT BC AB -PQ uestitus (Mannerheim) D. marmoreus Casey AB ON PQ _NS D. mucidus (Say) v_ AB _PQ D. parvicollis Casey -ec AB SK mn oN PQ NB_NF amplus Casey D. rufulus (Mannerheim) AK YK AB oN PQ NS D. squamosus LeConte _1 YBC AB SK MB angustatus (Dietz) tesselatus (WaIsh) D. vagenotatus Casey -YK NT BC AB SK MB ON PQ NF Genus ERIRHINUS Schdnherr E. aethiops (Fabricius)* AK YK NT BC AB SK MB ON PQ NS morio Mannerheim Genus GRYPUS Germar Grypidius Schiinherr Key to species: Cawthra (1957) G. brunnirostris (Fabricius) ON PQ G. equiseti (Fabricius)* AK YK NT BC AB SK MB ON PQ G. Ieechi (Cawthra) -BCAB SK MB ON Genus NOTARIS Germar Key to North American species: Buchanan (1927) N. bimaculatus (Fabricius) _YKNTBCABSKMBON PQ NB NS NF wyoming ensis Chittenden N. flavipilosus Chittenden pe N. puncticollis (LeConte) T- NT BC AB SK MB ON NS NT Genus PROCAS Stephens P. lecontei Bedel YK NT MB ON PQ 339 Curculioniclae Tlibe Smicronychini Genus SMICRONYX Schonhen Key to North American species: Arderson (1962) (Subgenus DESMORIS LeConte) S. fulvus LrrConte AB SK - ON S. humilis rDietzr - SK MB S. incertus tDietzt MB rustict',s Dietz uitiostt:.s Dietz S. pinguis Jllatchley - ON S. sordidus LeConte MB montazus (Dietz) (Subgenus .PACFfYPHANES Dietz) (P ac h5, ty c hius LeConte) S. amoenus (Say) - SK MB S. centralis (Dietz t AB lateralis (Dietz) S. discoideus {LeConte) - SK MB ON carus lDietz) (Subgenus PSEUDOSMICRONYX Dietz) (Syner:ha Dietz) S. commixt rs Dietz AB SK MB coseyi Blatchley S. corniculatus (Fahraeusl - ON PQ columl:'ianus Dietz conniuens Casey fallacir>sus l{]ima fallar Dietz laungi;zosus Dietz spurcus Casey squam'.tlatus LeConte S. flavicans LeConte - ON dietzil1ima macuh:tus Dietz nebulosus Dietz ornatil:tennis Dietz S. squalidur Casey - ON cinerascens Dietz morio l)ietz S. tessellatrLs Dietz MB ON im p r essiros t ris Dietz S. utilis Bu<'hanan AB SK (Subgenus TIMICRONYX s. str.) S. cinereus 'Motschulsky) - BC perplexus Dietz S. congesturi Casey MB S. sculpticollis Casey - ON gibbirostris Casey sagittatus Casey S. tychoides LeConte - ON S. vestitus l.,eConte AB - MB Genus PRO.MECOTARSUS Casey P. densus Cnsey - BC P. lumatus tlasey - SK Tlibe Tanysphyrini Genus TANYSPIIYRUS Germar T. ater Blatchley - BC - SK - ON makolskii Smreczynski T. lemnae (Fabriciustf - BC AB SK MB ON PQ 340 Curculionidae Tribe Bagoini Genus BAGOUS Germar Key to North American species: Tanner (1943) B. americanus LeConte B. aLratus Blatchley 3N;:_ B. bituberosus LeConte ______B. blanchardi Blatchley ON B. magister LeConte ON B. mamillatus Say _ABONPQ B. nebulosus LeConte BCABSK_NB ochraceus Blatchley B. obliquus LeConte _ONPQ B. planatus LeConte _ONPQ NB B. restrictus LeConte --ec-- -ONPQ B. tanneri O'Brien -oN longirostrus Tanner B. transversus LeConte BC AB SK MB ON PQ carinatus Blatchley Genus LISSORHOPTRUS LeConte Key to species: Kuschel (1951) L. oryzophilus Kuschel _AB L. simplex tSayt _ABONPQ egezus (Gyllenhal) Genus BRACHYBAMUS Germar B. electus Germar PQ inceratus Boheman Genus NOTIODES Schtinherr Endalus Castelnau Notionomus Erichsorr N otiop hilus Schtinherr Key to North American species: Tanner (1943) N. aeratus (LeConte) _ .AB N. limatulus tGyllenhal t -ABSKMBONPQ N. ovalis iLeConte) _ABONPQ N. punctatus (LeConte) BCAB_MB Genus ONYCHYLIS LeConte Key to North American species: Burke (1961) O. nigri rostris (Boheman t _ONPQ Tribe Derelomini Genus PHYLLOTROX Schrinherr Euclyptus Dietz P. ferrugineus LeConte -oN testaceus (Dietz) P. nubifer LeConte _BCAB_MB fuluipennis Sleeper P. rutilus (FaIl) _BC Genus ACAI-YPTUS Sch Subfamily OTIDOCEPHALINAE Tribe Otidocephalini Genus MYRMEX Sturm O tidoce p halus Chewolat M. chevrolatii (Horn) _ONPQ rny r mecodes (Chewolat) M. laevicollis (Horn) -oN 341 Curculionitiae M. myrmex (Herbst) PQ americ tnus (Chevrolat) herbstti Stunn myrmecodes (Say) Subfamily MAGDALINAE Tribe Magdalini Genus MA(I.DALIS Germar M. aenescerLs LeConte AK BC AB M. alutacea LeConte AK YK NT BC AB PQ M. armicoll s (Say) SK PQ pallida (Say) -oN M. austera austera Fall -oN M. austera substriga Fall PQ M. barbita (Say) MB ON PQ M. cuneiforrnis Horn BC M. gentilis LeConte YK BC AB SK PQ M. gracilis (LeConte) BC M. hispoides LeConte YK BC AB -oN pe NF M. imbellis (LeConte) BC AB M. inconspir:ua Hor-rr -oN PQ M. lecontei ,lecepta Sleeper BC M. olyra (Horbst) pe brunnices (Gyllenhal) M. pandura (Say) MB ON PQ M. perforatr,r Horn -oN PQ M. piceae B rchanan -oN PQ M. salicis Hcrn -oN PQ M. subtincta LeConte -BCAB sx Tribe Laemosaccinr Genus LAE.MOSACCUS Schainherr L. nephele (Flerbst) ON plagiatus (Fabricius) Subfamily CURCTILIONINAE Genus CURCULIO Linn6 Balani:tus Germar Key to Nc,rth American species: Gibson (1969) C. aurivestis Chittenden BC b reu inti. s us Chittenden C. caryae (lliorn) _ONPQ C. caryatrypes (Boheman) -oN cylindricollis (Casey) hariolus (Casey) picus (Boheman) C. confusor,.Hamilton) ON C. iowensis lOasey) SKMBONPQ_NS e*ilis Chittenden funicul aris Chittenden i6ls Chittenden nume n;',u s Chittenderr C. nasicus (fjay) ON PQ NB NS PE auctus (Casey) ordinatus (Casey) C. obtusus (Bianchard) MB ON PQ neocorylus Glbson C. pardalis (Chittenden) MB ON appalachius (Casey m ul t ifa sciatu s Chittenden paruicollis (Casey) C. proboscideus Fabricius ON PQ cuneatus (Casey) nasutus (Say) 342 Curculionidae quercus (I{orrr) r ectiro str is ( Gyllenhal ) rectus (Say) rostrdtus (Gyllenhal) sparsellus (Casey) C. sayi (Gyllenhal) _ONPQ acuminatus (Casey) algonquinus (Casey) auriger (Casey) macilentus (Casey) mollis (Casey) perexilis (Casey) setosicornis (Casey) sparsus (Gyllenhai) strigosus (Casey) C. strictus (Casey) MB ON longipes (Casey) tubulatus (Casey) utensis (Casey) uirginicus (Casey) C. sulcatulus (Casey) ON PQ NS baculi (Chittenden) curtus (Chittenden) striatus Chittenden C. uniformis (LeConte) BC occidentis (Casey) Subfamily ANTHONOMINAE Genus MACRORHOPTUS LeConte Key to North American species: Sleeper (1957b) M. hispidus Dietz SK M. sidalceae Sleeper nc v Tlibe Anthonomini Genus ANTHONOMOPSIS Dietz A. mixta (LeConte) _ONPQ Genus CHELONYCHUS Dietz C. Iongipes Dietz _BCABSKMB Genus ANTHONOMUS Germar Anthomorphus Weise Cnemocyllus Dietz Paranthonomus Dietz Serarthrus Blatchley Tachypterellus Fall & Cockerell Tachypterus Dietz A. confusus Dietz _BCAB A. consimilis Dietz PQ A. consors (Dietz) -BC A. convictus Gates A. corvulus LeConte - - - ncAB- me3N;3Ns NS NF A. deceptus Sleeper -BC A. elongatus LeConte -oNPQ A. haematopus Boheman _BCABSKMBONPQ NS sycophantaWalsh A. hirtus Leconte -BC A. interstitialis Dietz -ONPQNB A. lecontei Burke -BCABSKMBONPQ uariegatus LeConte A. melancholicus Dietz -BC murinus Dietz A. molochinus Dietz _ONPQ NS A. musculus Say _ONPQ sulcifrons LeConte A, nebulosus LeConte -BC-ONPQ 343 Curculionidae A. nigrinus.Boheman NT PQ A. ochreopiltsus Dietz _BC A. pictus Blatchley PQ A. profundus LeConte ON PQ NB NS uulpinz,:s Dietz A. quadrigil:bus (Say) BC MBSKAB ON PQ cerasi (List) magna (List) A. quesnelensis Sleeper BC AB A. robustuh:.s LeConte _ABSK A. rubidus L,eConte BC -oN rqDr\ A. rubricus lJchenkling & Marshall BC MB rubellus Dietz A. rufrpes LeConte AB A. rutilus (Boheman) PQ nubiloides FalI A. signatus lJay BC AB SK MB ON PQ NS PE NF bisignalus Gyllenhal pallidus Dietz scut eI lo. t u s Gyllenhal A. simiolus Illatchley PQ NB A. sphaeralciae Fall ---AB-l4e A. squamosr.s LeConte BCTSKMB A. squamulatus Dietz _ _ A. subfascia;us LeConte ON PQ latiusct;.Ius Dietz rantus Blatchley A. suturalis LeConte BC ON erythropterus Say flauicornis Boheman hicoriat: Pierce uespertinus Dietz A. tectus Le(lonte SK MB A. xanthocnr'mus Dietz ON Genus COC()OTORUS LeConte Key to No:th American species: Brown (1966) C. pumilae r:rajor (W.J. Brown) _MB C. pumilae pumilae (W.J. Brown) ON PQ C. scutellari,; ( LeconteJ _MB prunicida (Walsh) Genus DIETZIANUS Sleeper Xanthus Dietz D. pygmaeus. tDietzt MB Tribe Bradybatini Genus PSEI.IDANTHONOMUS Dietz Key to No;th American species: Clark (1987) P. crataegi (Walsh) AB SK MB ON PQ NB NS brunneus Dietz longulus Dietz paruulus Dietz p un ct icol I is Blatchley P. helvolus (13oheman) ON hamamelidis Pie'rce incipiens Dietz relictus Dietz P. rufulus Dietz _MB-PQ P. seriesetosus Dietz oNPQ-NS P. validus Dietz YT BCAB-MBONPQNBNS facetus Dietz 344 Curculionidae Subfamily RFIYNCIIAENINAE Key to North American species: Anderson (1989) Genus RHYNCHAENUS Clairville Orchestes llTiger R. griseus Sleeper _BC R. mixtus (Blatchley) _BCABSKMBONPQNBNSPE R. pallicornis (Say) _BCAB_MBONPQNBNSNF puberulus (Boheman) uniformis (W.J. Brown) R. testaceus (O.F. Mriller)x AK BC AB SK MB ON PQ NB NS NF canus (Horn) pallidior (Leng) Genus TACHYERGES Schtinherr T. ephippiatus (Say) -BCABSKMBONPQNB T. niger (Horn) AK YK NT BC AB SK MB ON PQ NB NS _ LB illinoisensis (Fall) paruicollis (LeConte) T. salicis (Linn6)x NTBCASSKMBONPQ-NSNF subhirtus (Horn) Genus ISOCHNUS C.G. Thomson L arcticus (Korotyaev)x AK_NT L flagellum (Erichson)* AK YK NT I. rufrpes (LeConte) -BCAB-MBONPQ-NSNF minutus (Horr') I. populicola (Silfverberg)J _ONPQNBNS Subfamily PRIONOMERINAE Tlibe Prionomerini Genus ODONTOPUS Say O. calceatus tSay) - ON PQ c ar bonar iu s ( Gyllenhal) Tribe Piazorhinini Genus PIAZORHINUS Schdnherr P. pictus LeOonte MB ON PQ P. scutellaris (Say) MBONPQ_NS Subfamilv TYCHIINAE Tribe-nlleschini Genus ELLESCHUS Dejean Alyca LeConte E. bipunctatus (Linn6) MB ON PQ E. borealis Carr AK YK AB - MB E. ephippiatus (Say) - YK - BC AB SK MB ON PQ NF rufus (Say) E. scanicus (Paykullt AB SK - ON PQ Genus PROCTORUS LeConte Encalus LeConte P. armatus LeConte AK - NT BC AB SK MB ON PQ NB P. decipiens (LeConte) AK YK NT BC AB SK MB ON PQ NB NS Tribe Lignyodini Key to genera: Clark (1982) Genus LIGNYODES Dejean Key to North American species: Ciark (1980) (Subgenus LIGNYODES s. str.) (T hy sanocnemis LeConte) 345 Curculionid:re L. bischoffi tBlatchley) - ON PQ L. fraxini tl,eContet - SK - ON PQ L. helvolus tl.eConte) MB ON PO b alaninoide s ( Schaeffer) breuis (Casey) caseyi (IAima) L. horridulus (Casey) - ON PQ punctatus (Casey) squamtger (Casey) L. ocularis ('laseyt - ON Genus PLO(IETES LeConte Hamaba Casey Rosella Whitehead Key to spercies: Clark (1982) P. ulmi LeConte - ON PQ Tribe Tychiini Genus SIBINIA Germar Key to New World species: Clark (1978) S. maculata (LeConte) AB maculilbr (Hatch) Genus TYCIIIUS Germar Miccotrcgus Schdnherr Paraty<:hius Casey Key to North American species: Clark (1971) T. aratus Say - SK arator ()yllen}:al T. liljebladi .3latchley AB SK MB T. lineellus l,eConte - BC AB SK dilectus Casey hesperis Casey probus Jasey radians Casey tacitus'Jasey T. montanus Clark AB T. picirostrit (Fabricius)t - BC AB SK MB ON PQ - NS PE - NF posticu:,: Gyllenhal T. semisqua:nosus LeConte - BC intermi.ttus }Iatc}l lamello.;us Casey squamosus Hatch T. soltaui Casey - SK MB hirsutus Clark hirtellu,s LeConte T. stephensi Schrinherrt - BC AB ON PQ NB NS griseus Schaeffer tontentosus (Herbst) T. tectus LeOonte AK YK - BC AB SK MB languidus Casey mixtus Hatch Subfamily GYMNETRINAE Tlibe Gymnetrini Genus GYM.NETRON Schdnherr G. antirrhinr (Paykull)t _BCABONPQ-NSNF G. netum (Germar)t _BC G. pascuorurn (Gyllenhal)t -BC_ONPQ G. tetrum (Fabricius)t -BC-ONPQ 346 Curculionidae TYibe Mecinini Genus MIARUS Schtinherr Cleopomiarus Pierce M. hispidulus LeConte MB ON PQ consuetus Casey illini Casey nanus Casey puritanus Casey Genus MECINUS Germar M. pyraster (Herbst)t - BC Subfamily CRYPTORITYNCHINAE Tribe Ithyporini Genus RHYSSOMATUS Schdnherr R. aequalis Horn ON R. Iineaticollis (Say) oN PQ Genus CONOTRACHELUS Dejean Loceptes Casey Key to North American species: Schoof ( 1942) C. anaglypticus (Say) ON PQ C. aratus (Germar) _PQ C. corni W.J. Brown -MBONPQ C. crataegi Walsh ON PQ C. elegans (Say) ON PQ C. geminatus LeConte _PQ C. juglandis LeConte oN PQ C. nenuphar (Herbst) BC -MBONPQ NS -NF orgula (Fabricius) cerasi (Peck) C. nivosus LeOonte -AB plagiatus LeConte C. posticatus Boheman -oNPQ C. recessus (Casey) -oN atokanus Fall C. schoofr Papp -oN tibialis Schoof C. seniculus LeConte -ONPQ Genus PHELOCONUS Roelofs P. cribricollis (Say) -oN puncticollis (Walsh) Genus MICROIIYUS LeConte Echinaspis Blatchley M. setiger LeConte ON wolcotti (Blatchley) Tlibe Cryptorhynchini Genus GERSTAECKERIA Champion Opuntiaphila Pietce Philopuntia Pietce Key to North American species: O'Brien (1970a) G. basahs (LeConte) AB Genus ACALLES Schdnherr A. carinatus LeConte ON Genus TYLODERMA Say Analcis Schdnherr Key to North American species: Wibmer (1981) T. aereum (Say) ON PQ 347 Curculionidae T. capitale \Vibmer -oN T. foveolaLum tSayt -oN T. minimun Blatchley AB T. nigrum Casey -oNPQ NB asclepiusae Sleeper T. oenotherare Wibmer AB SK MB Genus CRYTTORHYNCHUS Illiger C ry p to r hy nc hidiu s Pierce C. fuscatus .-eConte ON PQ C. lapathi tl,inn6tx BC ru _MB ONPQNBNSPE_NF Genus APTINROMECHUS Faust Key to Nc,rth American species: Whitehead (19Tg) A. ferratus (Say) ON Genus COPIIES Champion Key to North American species: Sleeper ( 1955) C. fallax (LeConte) ON C. obtentus,'Herbst) ON ypsilon (Boheman) Genus EUBULUS Kirsch Key to species: Fiedler (1939) E. bisignatu; rSayr ON PQ o b liq ue,hs ciatzs (Boheman) E. parochus (Herbstr oN PQ (.?) luctt:osus (Boheman) Subfamily ZYGOPINAE Key to North American genera: Sleeper (1963) Tribe Zygopini Genus CYLI NDROCOPTURUS Heller Copturt'des Casey Crjtrotus Casey C. deleoni Brrchanan BC AB C. longulus (LeConte) ON disperst:.ts (Casey) mucidu; (Casey) nubilat:.ts (Casey) obscure,'.lus (Casey) subcupreus (Casey) C. medicatusr Carr Tlibe Lechriopini Genus LECHRIOPS Schdnherr Gelus Casey Key to North American species Sleeper (1963) L. californice. (LeConte) L. oculata (Say) - :"- - oll PQ NS--- ^ne Genus ACOf'TUS LeConte Homogaster Provancher A. suturalis i,eConte ON PQ NB NS q ue be cens is (Provancher) Genus PSOMUS Casey P. armatus (.r)ietz) ON PQ politus ()asey 348 Curcul.ionidae Subfamily CEUTORIIYNCHINAE Tlibe Mononychini Genus MONONYCHUS Germar M. vulpeculus (Fabricius) ON PQ Tlibe Cnemogonini Key to genera: Colonnelli (1979) Genus CRAPONIUS LeConte C. inaequalis (Say) ON Genus CNEMOGONUS LeConte C. iecontei Dietz AK YK NT BC AB MB ON PQ NB Genus ACANTHOSCELIDIUS Dalla Torre & Hustache Acanthoscelis Dietz A. acephalus (Say) - BC AB SK MB ON PQ PE leprosus (Boheman) s ub uliro str i s (Gyllenhal) A. curtus tSay) PQ Genus AULEUTES Dietz A. epilobii (Paykull)* - YK NT BC AB SK MB ON PQ NB NS cruralis (LeConte) A. nebulosus (LeConte) - BC - SK MB ON PQ NB A. tenuipes (Leconte) -oNPQ Genus PERIGASTER Dietz Key to North American species: Buchanan (1931) P. cretura (Herbst) oN PQ quadrispinosa (Say) P. liturata (Dietz) oN PQ NF longirostris Buchanan Genus DIETZELLA Champion Coelogaster Schdnherr D. zimmermanni (Gyllenhai) - BC ON PQ NF Tribe Ceutorhynchini Genus CEUTORIIYNCHUS Germar C euthorhy nchidius J acqrelin du Val C. americanus Buchanan AK BC PQ- NS C. assimilis (Paykull)f )' BC 18:"MB 9* C. bolteri Dietz BC hirticollis Dietz C. carteri W.J. Brown AB C. convexipennis Fall MB C. cupreus Hatch sc C. dubitans W.J. Brown _SK C. echinatus Fall MB C. erysimi (Fabricius)f BC ON PQ NS C. floralis (Paykull)f T_ _ ON PQ NB NS I\'D puberulus LeConte sep tent r ionalis Gyllenhal C. fulvotertius FaII NT BC SK oN- C. handlordi W.J. Brown TY f MB C. hearnei W.J. Brown MB C. invisus Fall s^ y" C. lecontei Colonnelli -sc AB conuexicollis LeConte C. munki W.J. Brown MB C. neglectus Blatchley sc AB sx MB ON PQ C. omissus Fall T_ _ABSK MB oN PQ C. opertus W.J. Brown BC 349 Curculionidae C. oregonersis Dietz _YK-BCAB_MB C. ovipenni; Dietz C. pauxillur; Dietz - - - l" AB sx nre------C. pervestit,us FaIl AB C. punctige r Gyllenhalt _YK_BCABSKMBONPQNBNSNF C. querceti lGyllenhal) AK _ NT BC AB _ MB - PQ NB NS PE _ NF hamiltoniDietz pzslo Mannerheim C. rapae Gyllenhalt AKYK_BCABSKMBONPQ affiuentus Dietz C. squamatus LeConte _ONPQ C. squamosulus Sleeper _BC sleeperi Colonnelli C. subpubescens LeConte _YKNT_MB Genus POOPHAGUS Schttnherr P. sisymbri:i (Fabricius)t PQ Genus AMAIUS Schrinherr A. scortillurn (Herbst)t -BCABSKMBONPQ_NSNF robinsoni (Blatchley) Genus CAL'CSIRUS Thomson C. medialis (LeConte) _ONPQ C. semirufu.s (LeOonte) AB ON PQ NF C. zimmern rannii (Gyllenhal) MB ON Genus ALLI)SIROCAIUS Colonnelli A. angulatu.s (LeConte) _BCAB A. consangu.ineus (Dietz) AB SK MB A. disturbatus (Dietz) _BC Genus HAT; ROPLONTUS Thomson H. litura (F;rbricius)f -oN Genus SIR()CALODES Voss S. sericans (LeOonte) MB Genus NEDYUS Schdnherr Cidnorhinus Thomson N. fl avicaudis (Boheman) AB SK MB ON PQ apicalis (Dietz) Tribe Scleropterini Genus HONIIOROSOMA Frivaldszky H. sulcipenrre (LeConte) - YK - SK MB ON PQ Genus RUTIDOSOMA Stephens R. decipiens (LeConte) AK YK - BC AB SK MB ON PQ Genus ACA.LLODES LeConte A. saltoides Dietz PQ A. ventricosus LeConte MB ON Tlibe Phytobiini Key to New World genera: Colonnelli (1980) Key to genera: Colonnelli (1986) Genus PIIYIOBIUS Schdnherr Hydaticus Schdnherr Li todacty I us Redtenbacher Parenthis Dietz 350 Curculionidae P. leucogaster (Marsham) YK NT BC AB SK MB ON PQ griseomicans Schwarz my r iop hy I li (Gyllenhal) P. vestitus (Dietz) ON Genus EUBRYCHIOPSIS Dietz E. albertana (W.J. Brown) NT BC AB SK Genus EUBRYCHIUS Thomson E. velatus (Beck) BC aquaticus Thomson Genus RHINONCUS Schtinherr C ampy lir hy nch zs Dej ean Key to North American species: Hoebeke & Whitehead (1980) R. castor (Fabricius)t _BC_ONPQ NB NS NF R. longulus LeConte -BCABSKMBONPQ R. pericarpius (Linn6)t _BCABONPQ NF umbellae (Fabricius) R. pyrhopus Boheman NTBC_SKMBONPQ PE R. triangularis (Say) AB SK MB ON PQ occidentalis Dietz Genus PELENOMUS Thomson Mecopeltus Dietz P. asperulus Dietz \tK AB ON aeneosquamoszs (Dietz) P. congenialis (Dietz) AB P. fuliginosus (Dietz) BC AB ON PQ NB P. gracilipes Dietz -PQ griseus Blatchley P. pusillus Dietz _PQ scandens (.Dietz) P. scoliasus (Dietz) AB MB P. squamipennis (Sleeper) -BC P. squamosus LeConte YK _BC AB ON pe- im pressi uen tris Blatchley P. sulcicollis (Fahraeus) AB ONMB PQ P. ventralis (Sleeper) YK BC AB Genus NEOPIIYTOBIUS Wagner N. cavifrons (LeConte) BC AB toruipes (Dietz) Subfamily BARIDINAE Tribe Baridini Genus BARIS Germar Baridius Schiinhen Cy p hi r h inu s Schainherr Orthobaris Casey B. canadensis Blatchley -oN B. interstitialis (Say) -oNPQ agnita Casey anaplata Casey appalachia Casey apposita Casey aprica Casey apricoides Fall attonsa Casey austiniana Casey bispeculata Casey brachyrhina Casey breuipennis Casey carinulata (LeConte) caseyi HusLache cribrifera Casey cuneipennis Casey 351 Curculionidae demissa Casey dolosa Casey fultoni Casey immur,.is casey irrita Casey lauaca,za Casey lubrica Casey nephiana Casey oblata Casey oblong,tla Casey penins,ilae Horn quadr(:..ta (LeConte) retrusa Casey subaenea (LeConte) surrufcL Casey tumescens (LeConte) uagans Casey zuniana Casey B. lengi Hustache ON carbon,zria Blatchley B. stacesmit;hi Sleeper BC B. strenua (LeConte) MB c o I o r a c:;'.e n s i s C as ey formali',s Casey longulicollis Casey montanica Casey satelles Casey separara Casey subpar'.lis Casey uinta (.lasey B. striata (Say) _MBON B. subsimilis Casey ON Genus COS.MOBARIS Casey C. americarra Casey SK oN PQ sionilli Hayes squamLger Hayes Genus AUL OBARIS LeConte A. pusilla (l,eConte) PQ Genus AMF'ELOGLYPIER LeConte A. ampeiopsis (Riley) ON ater LeConte Genus PSEIJDOBARIS LeConte P. angusta (LeConte) angustula LeConte P. nigrina (Siay) oN PQ Genus TRICIHOBARIS LeConte T. trinotata (Say) ON impotens Casey jejuniosa Casey plumbea (LeConte) ueslila (Boheman) Genus ORTFIORIS LeConte O. crotchii LeConte O. tenuirostris Casey _ _ _:"ffi______ Tlibe Madarini Genus ONYCHOBARIS LeConte O. subtonsa LeConte - i{B O. pectorosa. LeConte PQ 352 Curculionidae Genus MADARELLUS Casey Willinkia Bondar M. undulatus (Say) - ON PQ Tribe Centrinini Genus GERAEUS Pascoe Centrinaspis Casey G. picumnus (Herbst) - ON albotectus (Casey) compactus (Casey) floridensis (Casey) lasciuus (Casey) oliuaceus (Gyllenhal) paululus (Casey) per scillus (Gyllenhal) sulor (Harris) Genus ODONTOCORYNUS Schdnherr O. denticornis (Casey) - ON O. scutellumalbum t$sy) - ON PQ Genus CENTRINOGYNA Casey C. canadensis Casey AB - MB Genus CALANDRINUS LeConte C. grandico)lis LeConte AB angustulus Casey C. obsoletus Casey AB insignus Casey Genus CATAPASTUS Casey C. conspersus tl,eConte) - ON PQ Genus CYLINDRIDIA Casey C. prolixa tl-eContet MB ON PQ Genus DIRABIUS Casey (Subgenus DIRABIUS s. str.t D. rectirostris (LeConte) ON PQ tenua (Blatchley) Genus PROSALDIUS Oglobin Anacentrinus Buchanan P. subaequalis (Casey) MB Genus STETHOBARIS LeConte S. commixta Blatchley - ON PQ S. laevimargo {Championt - ON S. ovata (LeConte) - ON PQ Genus IDIOSTETHUS Casey l. tubulatus tSayt - SK PQ s t rig apu nc t us (Harnilton) Genus HAPLOSTETHOPS Casey H. ellipsoideus (Casey) - ON Genus PLOCAMUS LeConte Epeuchaetes Lyman Euchaetes LeConte Eunyssobia Casey P. echidna (LeConle) - ON P. hispidulus LeConte - NB 353 Curcuiionii.ae Subfamily RIfYNCHOPHORINAE Tribe Sphenophorini Genus SPHENOPHORUS Schrtnherr Calandra Clairville Trichi:;chius LeConte Key to North American species: Vaurie (1951) S. aequalis aequalis Gyllenhal - BC AB SK MB ON PQ scirpi {Jhittenden uniuitia Chittenden S. australis australis Chittenden - ON PQ typhae' Chittenden S. callosus Oliviert - ON PQ jugosus Chittenden S. cariosus ,Oliviert NS cicatricosus (Say') flexuosus Gyllenhal larualis (Germar) sculptilis Uhler S. cicatristriatus Fahraeus AB SK MB cic at r i.,cennis Fahraeus ulkei llorn uariolc'sus LeConte S. costipennis Horn - BC AB SK MB ON PQ NB laeu ig t:.Lt u s Chittenden me do r aensis ( Satterthwait) S. incongruus Chittenden - ON elephantulus Chittenden S. melanoccphalus {Fabricius) - ON PQ nubilus Gyllenhal S. minimus Hart - ON S. mormon Chittenden - BC AB SK MB distich lidis Chittenden S. parvulus Gyllenhal - ON PQ pumilus Gyllenhal S. pertinax pertinax (Olivier) - ON - NB NS canaliculatus Boheman inter u c: I I at us Chittenden nasutu.s Chittenden penins ularis Chittenden sefiger Chittenden truncatus (Say) S. robustior Chittenden MB costifer Chittenden S. robustus Horn - BC MB ON m uI t ih, neat us ( Satterthwait) r ect i s t r iat u s Chittenden S. sayi Gyllenhal - BC s ubca r inat u s Mannerheim S. serratipes Chittenden AB S. striatipe:rnis Chittenden AB SK MB ON NS S. venatus ;enatus (Say) - ON PQ NB confusus Gyllenhal fallax Boheman immunis (Say) placid;ts (Say) S. villosiventris Chittenden PQ S. vomerinus LeConte AB baridioides llotrr S. zeae Walsh - ON PQ blatchieyi Chittenden Genus RHODOBAENUS LeConte H otnaiosty I us Chelrolat Key to North American species: Vaurie (1981) R. quinquepunctatus (Say) - ON PQ R. tredecimpunctatus tllliger) - ON PQ cribrarius (Fabricius) leptocerus (Panzer) q uatuor decirn punc tatus (Panzer) 354 Curculionidae Tribe Sitophilini Genus SITOPHILUS Schonherr NF S. granarius (Linn6)f - BC MB ON PQ r emotep unctatus Gyllenhal S. oryzae (Linn6)t BC - ON PQ NF S. zeamais Motschulsky* MB ON Genus DIOCALANDRA Faust D. elongata (Roelofs)t - BC Subfamily COSSONINAE Tribe Acamptini Genus ACAMPTUS LeConte A. echinus Casey _ A. rigidus LeConte ______BNpe _ _ _ _ Tfibe Cossoninr Genus HIMATIUM Wollaston Himatinum Cockerell H. errans LeConte PQ - NS nigritulum Casey paruula Blatchley Genus COSSONUS Clairville Borophloeus Wollaston C. americanus Buchanan PQ - NS NF C. crenatus Horn - BC C. impressifrons Boheman - ON PQ concinnus Boheman C. pacificus Van Dyke - BC AB C. piniphilus Boheman - BC californicu s Motschulsky scrobiculatus LeConte C. platalea Say - ON PQ bohetnanniHotn subareatus Boheman C. quadricollis Van Dyke - BC AB Tlibe Rhyncolini Genus PENTARTHRUM Wollaston P. huttoni Wollastont PQ NF Genus EUOPHRYUM Broun E. confine (Broun)f - NF Genus HEXARTHRUM Wollaston H. thujae W.J. Brown - BC H' ulkei Horn - oN PQ Genus ELASSOPTES Horn E. marinus Horn - BC Genus STENOSCELIS Wollaston S. brevis (Boheman) - ON PQ Genus PHLOEOPHAGUS Schonherr P. apionides Horn - ON PQ P. cinadensis Van Dyke - BC AB - MB - PQ P. minor Horn PQ carinatus (Blatchley) 355 Scolytidae Genus RHIINCOLUS Germar Xylocc.mesus Thatcher R. brunneu.s Mannerheim AK YK NT BC AB MB ON PQ NB NF c alifor ni c u s Wollaston relictus Casey R. cylindricollis Wollaston -BC R. knowltoni (Thatcher) MB R. macrops Buchanan _BC ON pr) NS R. oregonensis Horn _BC R. pallens Oasey -BC R. stacesm:thi Sleeper _BC Genus CA-I|,PHONOTUS Casey C. testaceus Casey AK BC AB ON PQ NB NF Subfamily DRYOPHTHORINAE Tribe Dryophthorini Genus DRYOPHTHORUS Germar D. america:rus Bedel MB ON PQ NB NS corticalis Say Family SCOLYTIDAE bark beetles J. McNamara In Canada and Alaska 204 species of Scolytidae The North American scolytids are well known are recorded. Most of the Canadian species attack taxonomically; Wood (1982) revised all t;he North forest trees, especially conifers, and may cause con- American species, and Bright (1976) covered the siderabler damage to timber. Both the larvae and Canadian and Alaskan ones. adults form galleries under the bark. Some of the galleries may be so distinctive that the genus and, AK (52); YK (33); NT (31); BC (13a); AB (80); SK in some ,iases, the species that made them can be (40); MB (a7); ON (89); PQ (81); NB (53); NS (a6); identifieci. PE (8);LB (6);NF (27) . I (5) Subfamily FIYLE SININAE Tribe Hylastini Genus SCIItrRUS LeOonte S. annectans LeConte AK_NTBCAB ONPQ-NS NF S. pubescens Swaine AKYK-BCAB Genus HYLURGOPS LeConte H. porosus (LeConte) NT BC AB SK lecontei Swaine H. reticulatus Wood NT BC H. rugipennis pinifex (Fitch) BC AB SK MB ON PQ NB NS H. rugipenrris rugipennis AK YK BC AB (Mannerheim) H. subcostulatus subcostulatus AK BC (Mannerheim) c r i s tat u s ( Mannerheim) Genus IfYLASTES Erichson H. gracilis LeOonte BC asper Swaine Iongus LeConte nitidus Swaine H. longicollis Swaine _BCAB H. macer LeConte _BC 356 Scolytidae H. nigrinus (Mannerheim) AK YK BC AB yukonis Fall H. porculus Erichson MB ON PQ NB NS canadensis Blackman carbonarius Fihch cau ernos u s ( Zimmermann) granosus Chapuis scober Swaine swainei Eggets ruebbi Swaine H. ruber Swaine NT BC AB H. tenuis Eichhoff _BC criticus Eichhoff minutus Blackman pd.ruus Blackman pusillus Blackman Tribe Hylesinini Genus HYLASTINUS Bedel H. obscurus (Marsham)f BC _ONPQ NS Genus HYLESINUS Fabricius Leperisinus Reitter H. aculeatus Say SK MB ON PQ NS cinereus (Swaine) imperialis Eichhoff H. californicus (Swaine) MB hoferi (B\ackman) H. criddiei (Swaine) MB ON PQ H. fasciatus LeConte -oN H. pruinosus Eichhoff -oN Genus AI-NIPI{AGUS Swaine H y la s tinoides Spessivetsev A. aspericollis (LeConte) AK _BC A. hirsutus Schedl _BC Tribe Tomicini Genus PSEUDOIIYLESINUS Swaine P. dispar dispar Blackman _BC P. dispar pullatus Blackman -BC P. granulatus (LeConte) _BC P. nebulosus nebulosus (LeConte) T -BC AB P. nobilis Swaine _BC furnissi Blackman P. pini Wood _BC P. sericeus (Mannerheim) AK _BC grandis Swaine P. sitchensis Swaine AK _BC P. tsugae Swaine AK _BC heeni Blackman obeszs Swaine similis Blackman Genus HYLURGOPINUS Swaine H. rufrpes (Eichhoffl SK MB ON PQ NB NS opaculus (LeConte) Genus XYLECHINUS Chapuis X. americanus Blackman X. montanus Blackman AK ecABsr-9*PQ-Tt--- Genus DENDROCTONUS Erichson D. bre.ricomis LeConte BC AB barberi Hopkins 357 Scolytidae D. murrayar:.ae Hopkins _BCAB D. ponderosrLe Hopkins _BC _ y"9*______monticolae Hopkins D. pseudotsugae Hopkins _BCAB D. punctatus, LeConte AK YK NT BC AB -oN PQ NB NF johanse.ci Swaine D. rufipennis (Kirby) AK YK NT BC AB SK MB ON PQ NB NS NF borealis Hopkins engelmc:.nni Hopkins o6eszs (Mannerheim) piceaperda Hopkins similis .i,eConte D. simplex LeConte AX _ NT BC AB SK MB ON PQ NB NS PE LB NF D. valens Lelonte _NTBCAB_MBONPQNBNSNF Tribe Phloeotribinr Genus PHLOEOTRIBUS Latreille P. dentifrons (Blackman) P. lecontei Schedl ---ecAB- _9" puberulus LeConte P. liminaris tHarris) SK MB ON PQ NB mississt pp iensis (Blackman) P. piceae Sweine AK YK NT BC AB MB ON PQ NB NS Tribe Phloeosinini Genus PHL()EOSINUS Chapuis P. canadensis Swaine -oNPQNB P. cupressi l{opkins AK BC nitidus Swaine P. dentatus ( Say) ON enieus l:llackrr.an graniger (EichhofD haagi (Eichhoff) P. hoferi Blar:kman BC P. hoppingi Swaine BC woodiBrig}rt P. keeni Blackman BC P. pini Swaine AK YK NT BC AB MB PQ alaskanus Blackman piceae Swaine P. punctatus LeConte BC buckhorni Blackman chamberlini Blackman haniksu Blackman rubicun,lulus Swaine rzsli Blackman P. scopulorum scopulorum Swaine _BCAB P. sequoiae llopkins AK- _BC blackmcLni Schedl squamo.sus Blackman Genus CHRAMESUS LeConte C. hicoriae LeConte oN PQ lecontei (Chapuis) Tribe Hypoborini Genus CHA.ETOPHLOEUS LeConte Pse ud oc' ryp h alus Swaine Renocis Casey C. heterodoxus (Casey) BC AB SK MB brittaini:. (Swaine) brunneu.s (Blackman) commix,ius (Blackman) criddlei (Swaine) /asczs (Blackman) 358 Scoly'tidae Tbibe Polygraphini Genus CARPHOBORUS Eichhoff C. andersoni Swaine AKYKNT_AB C. carri Swaine AKYKNT_AB MB -NS-- C. dunni Swaine _NB C. intermedius Wood AK BC C. ponderosae Swaine C. sansoni Swaine - - - :"AB- engelmanniWood. C. vandykei Bruck -BC Genus POLYGRAPHUS Erichson Lepisotnus IKtrby P. convexifrons Wood AKYK_BCAB_PQ P. rufrpennis (Kirby) AK YK NT BC AB SK MB ON PQ NB NS PE LB NF breuicornis (Kirby) nigriceps (Kirby) saginatus Mannerheim Subfamily SCOLYTINAE Tribe Scolytini Genus SCOLYTUS Geoffroy S. fagi Walsh S. laricis Blackman _ S. mali (Bechstein)t _ _ _:"_ _ _:x;,_ sulcatus LeConte S. monticolae Swaine -BCAB S. multistriatus (Marsham)f S. muticus Say S. opacus Blackman _ - _:'AB_ I"3NTl"- abietis Blackman S. piceae (Swaine) AK YK NT BC AB SK MB ON PQ NB S. praeceps LeConte _:"f" S. quadrispinosus Say ___:"___oNpe__ carya Riley coryoe Walsh S. rugulosus (P.W.J. Mtiller)t -BC-oNPQ NS_-_ S. subscaber LeConte _BC S. tsugae (Swaine) _BCAB S. unispinosus LeOonte -BCAB fis&ei Blackman sobrinus Blackman S. ventralis LeConte BC Ttibe Micracini Genus PSEUDOTHYSANOES Blackman Chalcohyus Blackman C ry ptocle ptes Blackman Cryptulocleptus Wood P. rigidus (LeConte) _PQ drakeiBlackrr'an Genus MICRACIS LeConte M. swainei Blackman ON populi Swaine Genus HYLOCURUS Eichhoff M icr acisoides Blackman H. hirtellus LeConte _BC crinitus Blackman 359 Scolytidae Tribe Ipini Genus PITYOGENES Bedel P. carinulatus (LeConte) BC hannatus (LeConte) P. fossifrons (LeConte) BC P. hopkinsi Swaine - SX l,fg ON pq NB NS NF P. plagiatus knechteli Swaine _YK BC AB SK P. plagiatus plagiatus (LeConte) AB SK MB ON PQ lecontei. Swaine Genus PITYOKTEINES Fuchs Orthotomides Wood, P. elegans Swaine BC AB P. Iasiocarp'. (Swaine) BC AB P. minutus rSwaine) BC AB jasperi Swaine P. ornatus ( lwaine) BC P. sparsus (.LeConte) _ABSK MB ON PQ NB NS PE LB NF balsameus (LeConte) Genus ORTHOTOMICUS Ferrari O. caelatus i.EichhofD AK YK NT BC AB SK MB ON PQ NB NS NF decretus (Eichhoff) Puncti7'ennis (LeConte) uicinus (.LeConte) Genus IPS DeGeer I. borealis Swaine AI{ YK NT BC AB SK MB ON PQ NB NS NF swaine. Hopping thomasi Hopping I. calligraphus (Germar) SK oN PQ NS chloroticus Dejean conforntis Dejean eresus rlsay) ponderosae Swaine praetnorsus (Eichhoffl I. concinnus (Mannerheim) AK BC chamberlini (Swaine) hirsutus (Eichhoff) I. emarginatus (LeConte) BC I. grandicollis (Eichhofl) lae oN pe cacographus (LeConte) chagno;zi Swaine cloudcrcfti Swaine c r i b r icct llis ( Eichhofl) I. integer (E ichhofl) BC I. latidens (l.,eConte) YK_ BC AB SK ON PQ guildilllackrnan longidens Swaine I. mexicanuri (Hopkins) AK BC AB radiatae Hopkins I. montanus (Eichhof0 BC uancoutseri Swaine L perroti Swaine _BCABSKMBONPQNB I. perturbatus (EichhofD AK YK NT BC AB SK MB ON PQ NB LB hudsonicus (LeConte) interp u.nctus (EichhofD I. pini (Say) AK YK NT BC AB SK MB ON PQ NB NS NF dentatus (Sturm) laticolli.s Swaine oregoni:s (Eichhoff) pallipes (Sturm) praefrictus (Eichhoff) rectus (LeConte') I. plastographus plastographus BC (LeConte) L tridens (Mannerheim) YK NT BC AB am isk iuL iensis Hopping 360 Scolytidae dubius Swaine engelmanni Swaine inter r u ptu s (Mannerheim) semirostris Hopping yohoensis Swaine I. woodi Thatcher AB Tribe Dryocoetini Genus LYMANTOR Lovendal L. alaskanus Wood AK L. decipiens (LeConte) oN pe Genus DRYOCOETES Eichhoff D. affaber (Mannerheim) AK YK NT BC SK MB ON PQ NB NS NF piceoe Hopkins pubescens Swaine D. autographus (Ratzeburg) YKAK NT BC AB SK MB ON PQ NB NS LB NF americanus Hopkins pseudotsugae Swaine setn ica steneus (Mannerheim) sep te nt r ionis ( Mannerheim) D. betulae Hopkins BC ON PQ NB NS NF eichhoffi Hopkins I iq uidatnb ar is Hopkins D. caryi Hopkins AK BC AB D. confusus Swaine BC AB -T abiells Hopkins D. granicollis (LeConte) -PQ NB D. sechelti Swaine BC Tlibe Crypturgini Genus CRYPTURGUS Erichson C. borealis Swaine AK-NTBCABSKMBONPQNBNSNF corrugatus Swaine C. pusillus (Gyllenhal)t _SK_ONPQNBNSPE-NF dtotnus LeConte Genus DOLURGUS Eichhoff D. pumilus (Mannerheim) AK BC Tlibe Xyloterini Genus XYLOTERINUS Swaine X. politus (Say) MB ON PQ NB NS unicolor (Eichhofft Genus TRYPODENDRON Stephens T. betulae Swaine NT BC AB -MBONPQNBNSPE T. lineatum (Olivier) AKYK_BCAB SK MB ON PQ NB NS PE NF biuittatum (Kirby) borealis Swaine c au ifron s (Mannerheim) uittigerum Eichhoff T. retusum (LeConte) AKYK-BCABSKMBONPQNB T. rufrtarsis (Kirby) AKYK-BCAB-MBONPQNBNS ponderosae Swaine Tlibe Xyleborini Genus XYLOSANDRUS Reitter X. germanus (Blandford) ON 361 Scolytidae Genus XYLI.BORUS Eichhoff Ambros;odmus Hopkins Anisantl.rus Ferrari X. affinis Eichhoff - ON X. dispar (Fabricius) - BC - ON PQ - NS NF pyri (Peck) swainei (Drake) X. intrusus Blandford - BC futcfri Hopkins howardt Hopkins scopulorum Hopkins X. obesus Letlonte - ON PQ populi (Swarne) serratus Swaine X. pubescens Zimmermann - ON propinguus Eichhoff X. sal (Hopl:inst - ON PQ minor Swaine neardus Schedl X. xylographrs (Sayt - ON PQ canadensis Swaine inermis Eichhoff Genus XYLEBORINUS Reitter X. saxeseni { latzeburg) - BC - ON PQ orbufi (llopkins) floridem;is (Hopkins) Iibocedr,:. Swaine pecanus (Hopkins) quercus (Hopkins) lszgoe Swaine Tlibe Cryphalini Genus TRYPOPHLOEUS Fairmaire T. populi Hopkins - BC AB SK MB NB T. striatulus (Mannerheim) AK YK AB - MB - PQ - NS NF nitidus fiwaine punctipennis Hopkins T. thatcheri (Woodt - BC Genus PROCRYPHAIUS Hopkins P. mucronatrLs (LeContet AX BC AB idahoensis Hopkins populi L{opkins P. utahensis Hopkins AK BC AB - PQ oceris Hopkins solicis F.opkins Genus CRYPHALUS Erichson Taeniogi),yptes Bedel C. pubescens Hopkins - BC subconcentralis Hopkins C. ruficollis f-aseri Hopkins - - PQ balsameus Hopkins C. rufrcollis ruficollis Hopkins AK YK - BC AB - MB ON PQ NB NS PE - NF amabilis Chamberiin approximatus Hopkins canadensis Chamberlin coloradensis (Wood) grandis Chamberlin mainensis Blackman Genus HYPGTHENEMUS Westwood Stephanoderes Eichhoff H. dissimilis (Zimmermann) - ON PQ chapuisti (Eichhoff) 362 Scolytidae H. eruditus Westwood - ON asiminae Hopkins citri Ebling euonymi Hopkins hamamelidis Hopkins juglandis Blackman nigripennis Hopkins przni Hopkins punctifrons Hopkins rumseyi Hopkins schwarzi (Hopkins) Tlibe Corthylini Genus PSEUDOPITYOPHTHORUS Swaine P. minutissimus (Zimmermann) PQ P. pubipennis ( LeConte) sc _9* Genus CONOPHTHORUS Hopkins C. banksianae McPherson -oN C. coniperda (Schwarz) -oN PQ NB NS clunicus Hopkins taedae Hopkins C. ponderosae Hopkins BC contortae Hopkins flerilis Hopkins tn lambertianae Hopkins monticolae Hopkins scopulorum Hopkins C. resinosae Hopkins oN PQ NB NS uirginianae Hopkins Genus PITYOPHTHORUS Eichhoff Key to North American species: Brieht (1981) (Subgenus PITYOPHTHORUS s. str.) (Myeleborus Blackman) P. absonus Blackman _BC AB demissus Blackman inyoensis Big}rt P. alpinensis Hopping AB P. angustus Blackman __ON_NENS P. aquilus Blackman ___sc AB SK aristatae Bright caelator Blackman P. balsaneus Blackman -NT ON PQ NB NS patchiBlackman P. bassetti Blackman AK YK NT P. biovalis Blackman -ONNS P. boycei Swaine \tK BC AB catulus (Blackman) iniquus (Blackman) siouxensis Bight P. briscoei Blackman -oN PQ NB mundus Blackman P. carinatus carinatus Bright PQ NB P. carinatus monticolae Bright sc AB_ P. cariniceps LeConte ? AB SK ff,fe O* PQ NB NS canad,ensis Swaine cognatus Blackman P. carinulatus Swaine AK opimus Blackman P. cascoensis Blackman NT -AB ON NF pilifer Schedl P. cavatus Bright SK MB P. concavus Blackman oN PQ NB NS P. confertus Swaine BC agndtus Blackman burkeiBlackman com.Dtus Blackman 363 Scolytidae P. confinis LeConte - BC P. consimilis LeConte - BC AB SK MB ON PQ - NS granulatus Swaine nudus Siwaine P. dentifrons Blackman AB ON PQ NB NS PE - NF P. digestus ( LeContet - BC AB aplanatus Schedl hophinsi Blackman idoneus Blackman ponderosae Blackman P. fuscus Bla.ckman - BC smithi Schedl P. grandis B'ackman - BC P. hesperius Bright - BC P. intextus Swaine - BC AB SK MB ON PQ NB NS NF shepardi Blackman lonsus lJlackman P. lautus Eichhoff - ON PQ natalis Blackman rhois Svraine P. murrayanae Biackman AK - NT BC AB - MB ON - NB acceptut: Bight aurulentus Bight cutleri S,waine depygis Blackman elongatt.!.s Swaine e:rilis Swaine gracilis Swaine tenuis Swaine watsoni Schedl P. nitidulus tMannerheim) AK BC AB atratulus (LeConte) punctict. llis (LeConte) P. nitidus Svvaine AK YK NT BC AB ON PQ NB NS NF anceps Illackman aquilon,:us Bight borealis Swaine uorions Schedl P. opaculus L,eConte AK YK NT BC AB SK MB ON PQ NB NS NF abietis Eilacknan albertensis Blackman exiguus Blackman pygmaetts Schedl P. orarius Bright - BC P. pseudotsugae Swaine - BC thatcheriBight P. pubemlus (LeConte) - ON PQ NB NS NF infans Eichhoff P. pulchellus Eichhoff - YK NT BC AB SK MB ON PQ NB hirticept, LeConte pusio LeConte P. pulicarius (Zimmermann) - SK MB ON PQ NB NS P. ramiperda Swaine - ON PQ fiuazi (Blackman) P. recens Bri.;ht - YK - BC P. scalptor Blackman - BC P. scalptus Bright - BC P. serratus Swaine - BC P. sierrensis Bright - BC P. toralis Wo,rd - BC AB collinus Bng}rt confusus (Bright) P. tuberculatus Eichhoff AK YK - BC AB rugicollis Swaine P. venustus Rlackman - BC artifexBlackman Genus GNAT'HOTRICHUS Eichhoff Ancyloderes Blackman Gnathot,"ic hoides Blackman 364 Platypodidae G. materiarius (Fitch) - SK MB ON PQ NB NS corthyloides Eichhoff G. retusus (LeConte) - BC alni Blackman G. sulcatus (LeContet - BC aciculatus Blackman Genus MONARTHRUM Kirsch M. fasciatum (Say) - ON PQ gracile (Eichhoff) simile (Eichhofl\ M. mali (Firch) - oN PQ NB longulum Eichhoff M. scutellare (LeConte) - BC cauum (LeConte) obl iq uecaudatum ( Schedl) Genus CORTF{YLUS Erichson C. punctatissimus (Zimmermann) - ON PQ FamilyPLAIYPODIDAE pinholeborers J. McNamara One species of Platypodidae occurs in Canada. Wood (1979) has published a catalog of the These beetles infest the wood of weakened trees, North American species. where they mine galleries. The larvae feed on the fungus that grows on the gallery walls. BC (1) Subfamily PLATYPODINAE TYibe Platypodini Genus PLATYPUS Herbst P. wilsoni Swaine - BC 365 Halictophagidae Order Strepsiptera twisted-winged parasites S.B. Peclt In Canada 11 species of Strepsiptera are now Adults may be reared from parasitized hosts. known in three of the nine families recognized Males may be attracted to ultraviolet black light around the world. These species are distributed in traps or may be lured by placing virgin females (in Southern Canada. Another 10 species occur in the their hosts) in frnely screened cages. northern tier of the United States and may eventu- The systematic position of this group is becom- ally be found in Canada. Many taxa are known ing clear. It has been grouped near the similarly only throrrgh adult females. endoparasitic Rhipiphoridae in general works on The biology of this endoparasitic group is highly beetles (Arnett 1968, Crowson 1981, White 1983). unusual. Studies on species similar to those in Extensive morphological studies by Kinzelbach Canada suggest the following. The frrst-instar tri- (1971) and Kathirithamby (1989) suggest that ungulin-like larvae are free-living at birth but strepsipterans are the sister group to the beetles as immedial;ely seek and parasitize immature speci- a whole. They are then ranked as a separate order mens of lvgaeid bugs, various leafhoppers and their most clearly related to the beetles (Kinzelbach 1971, relatives, and Andreno solitary bees. The hetero- Lawrence and Newton 1982, Kathirithamby 1989). morphic larvae mature in the host insect and Overviews of the group in North Arrrerica are pupate there. Adult female strepsipterans remain found in Bohart (1941), Kinzelbach (1$71), and larva-like and in the host. Adult males are winged Luna de Carvalho and Kogan (1990). The classifica- and live r)nly a day and an evening or two, during tion used here follows Kathirithamby (1980). which tinre they actively search for females. After the female has been inseminated, eggs develop and hatch inside her. A thousand or more larvae may then leave the female, to seek new hosts and begin the cycle again. BC (2);AB (1); SK (3); ON (2);PQ (2);NS (1) Suborder Stylopidia Family CORIOXENIDAE Subfamily CORIOXENINAE Genus LOANIA Kinzelbach L. canadensis Kinzelbach ON Family HALICTOPHAGIDAE Subfamily IIALICTOPHAGINAE Genus HALICTOPHAGUS Bohart Agalliaphagus Pterce Anthericomma Pierce Bruesia Perkins Cyrtocaraxenos Pierce Dacyrtocara Pietce Dioxocera Pietce Diozocera Pierce Megalechthrus Perkins Membracixenos Pierce NeocholaxPierce Pentac{.adocera Pierce Pe n log'qm mqPft i1a Pierce Pentaxocera Pierce Pentazocera Pietce Pentozoe Pierce Pseudopatella Bohaft Pyrilloxenos Pierce 366 Stylopidae Key to North American species: Bohart (1943) H. americanus Perkins PQ H. bidentatus Bohart _SK H. mackal Bohart _SK H. omani Bohart AB Genus STENOCRANOPHILUS Pierce Delphacircnos Pierce Muirixenos Pierce S. canadensis Kinzelbach SK Family STYLOPIDAE Subfamily STYLOPINAE Genus STYLOPS Kirby Katastylops Pierce Neostylops Pierce Prostylops Pierce S. advarians Pierce -BC S. childreni Gray NS S. leechi Bohart _BC prf S. vicinae Pierce Subfamily XENINAE Genus PSEUDOXENOS Saunders P. tigridis Pierce ON 367 REFERENCES Abdullah, M. 1965. 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Abacidus LeConte 33 Acupalpus Latreille 47 Abax Bonelli 35 Acylamus Sharp 226 Abdera Stephens 248 Acylophorus Nordmann 122 Abletobium Casey 115 Adalia Mulsant 235 Abraeinae 135 Adelina Dejean 255 Abraeini 135 Adephaga 7 Abraeus Leach 135 Aderidae 275 Abstrulia Casey 243 Aderocharis Sharp 113 Acalles Schiinherr 347 Aderus Westwood 275 Acalligrapha Monros 310 Adimonia Laicharting 315 Acallodes LeConte 350 Adistemia Fall 240 Acalymma Barber 315 Adota Casey 101 Acalyptus Schtjnherr 341 Adoxini 309 Acamptini 355 Adoxus Kirby 309 Acamptus LeConte 355 Adranes LeConte 129 Acanthocinini 296 Adranini 129 Acanthoderini 296 Aegialia Latreille 147 Acanthoscelides Schilsky 301 Aegialiini 146 Acanthoscelidius Dalla Torre & Hustache 349 Aegialites Dejean 270 Acanthoscelis Dietz 349 Aegialitinae 270 Achenomorphus Motschulsky 113 Aegomorphus Haldeman 296 Acholerops Casey 83 Aeletes Horn 136 Acicnemidinae 338 Aeolus Eschscholtz 176 Acidota Stephens 89 Aethiopoctines Thomson 296 Aciliini 71 Agabetes Crotch 63 Acilius Leach 71 Agabetinae 63 Acletus LeConte 212 Agabetini 63 Aclypea Reitter 85 Agabini 67 Acmaeodera Eschscholtz l.6l- Agabinus Crotch 67 Acmaeoderinae 161 Agabus Leach 67 Acmaeoderini 161 Agalliaphagus Pierce 366 Acmaeops LeConte 289 Agaporus Zimmermann 67 Acmaeopsilla Casey 290 Agasphaerops Horn 332 Acmaeopsoides Linsley & Chemsak 293 Agathengis des Gozis 223 Acoptus LeConte 348 Agathidini 80 Acrepis LeConte 200 Agathidium Panzer 80 Acritini 136 Agleninae 270 Acritus LeConte 136 Aglenus Erichson 270 Acrolochct Thomson 88 Aglyptinus Cockerell 79 Acromacer Kuschel 324 Agonoderus Dejean 47 Acropteroxys Gorham 224 Agonodromius Reitter 52 Acroschatia Wilke 254 Agonum Bonelli 52,53 Acrotona Thomson 100 Agostenus MotschulskY 50 Acrotrichiinae 76 Agrabia Casey 236 Acrotrichis Motschulsky 76 Agrilaxia Kerremans 165 Acrulia Thomson 88 Agrilinae 167 Actedium Motschulsky 21 Agrilini 167 Actiastes Casey 127 Agrilus Curtis 167 Actidium Matthews 75 Agriotella Brown 182 Actium Casey 127 Agriotes Eschscholtz 182 Actizona Chandler 127 Agriotini 182 Actobius Fauvel 117 Agroiconota Spaeth 323 397 Agronus Horn 332 Ampeloglypter LeConte 352 Agyrtecanus Reitter 77 Amphasia Newman 45 Agyrtes Itrdlich 77 Amphicerus LeConte 200 Agyrtidae 77 Amphichroum Kraatz 90 Ahasverus des Gozis 220 Amphicrossus Erichson 216 Airora Reitter 207 Amphidora Eschscholtz 257 Akephoms LeConte 17 Amphimallon Berthold 153 Alamona Casey 155 Amphivectura Blaisdell 212 Alaobia Thomson 102 Amphizoa LeConte 61 Alaus Eschscholtz 175 Amphizoidae 61 Aleocharil Gravenhorst I04 Amydetinae 191 Aleocharinae 99 Amydrogmus Pierce 334 Aleocharini 104 Anabolus Gistel 12 Aleodoru,s Say 106 Anacaena Thomson 132 Aletinus Desbrochers 327 Anacentrinus Buchanan 353 Alitargus Casey 242 Anadaptus Casey 43 Allandrini 325 Anaedus Blanchard 262 Allandrus LeConte 325 Anaglyptini 286 Alleculinne 259 Anagrylius Casey 156 Alleculinii 259 Analcis Schdnherr 347 Alloeocnemis LeConte 199 Analeptura Linsley & Chemsak 291 Allonychus Zaitzev 67 Anambodera Barr 161 AllopodaLeConte 276 Anametis Horn 334 Allopodini 276 Anaquedius Casey 122 AllorhagtumKolbe 289 Anaspidinae 276 Allosirocrilus Colonnelli 350 Anaspidini 276 Alniphagus Swaine 357 Anaspis Geoffroy 276 Alobates Motschulsky 26L Anqstictodera Casey l2I Alobus LeConte 151 Anastrangalia Casey 293 Alocentron Schilsky 328 Anatis Mulsant 235 Alphitobius Stephens 258 Anatrinodia Casey 157 Alphitophagus Stephens 255 Anchastus LeConte 184 Altica Mtiller 317 Anchicera Thomson 223 Alticinae 316 Anchodemus LeConte 335 Alyca Le(lonte 345 Anchomenus Bonelli 52 Alycodes Dietz 339 Aruchus LeConte 52 Amacylophorus Smetana 122 Anchytarsus Gu6rin-M6neville 169 Amalus Schdnherr 350 Ancistronycha Markel 193 Amara Bonelli 35, 38 Ancylochira Eschscholtz 163 Amarygrrrini 259 Ancyloderes Blackman 364 Amqurosromo Mtiller 299 Ancylonycha Blanchard 152 Amblopusa Casey 107 Ancyronychini I72 Amblycerinae 300 Ancyronyx Erichson L72 Amblycerus Thunberg 300 Ancy rop horus I{r aatz 94 Amblyctis LeConte 247 Andrector Horn 315 Amblyderus LaFert6-S lnecldre 27 | Androchirus LeConte 260 Ambrosiodmus Hopkins 362 Anelaphus Linsley 281 Americanotimarcha Jolivet 310 Anelastes Kirby 188 Americonr,asezs Csiki 34 Anelpistus Horn 263 Amerinusr Casey 46 Anepsiini 253 Amerizus Chaudoir 28 Anepsius LeConte 253 Amecocerus Solier 213 Anepsiota Casey 101 Ametor Semenov 132 Anisa.ndrus Ferrari 362 Amischa Thomson 101 Anisocaluio Crotch 235 Arnnesia.Florn 333 Anisodactylus Dejean 43, 44 Amniscus, Haldeman 297 Anisomera LeConte 67 Ampedini 183 Anisorus Mulsant 288 Ampedus Dejean 183 Anisostena Weise 321 398 Anisosticta Dejean 233 Aphanotus LeConte 258 Anisotarsus Chaudoir 45 Aphelogenio Chaudoir 56 Anisotoma Panzer 80 Apheloplastus Brown 79 Anisoxya Mulsant 248 Aphenolia Reitter 215 Anobiidae 201 Aphodiinae 146 Anobiinae 202 Aphodiini 147 Anobiini 203 Aphodius Illiger 147 Anobium Fabricius 203 Aphonus LeConte 156 Anocomis Casey 283 Aphorista Gorham 238 Anogcodes Dejean 266 Aphrastus Say 333 Anogdus LeConte 78 Aphthona Chevrolat 317 Anomala Samouelle 154 Apion Herbst 328 Anomalaegialia Brown 147 Apionidae 328 Anomalepta Casey 154 Apionini 328 Anomalini 154 Aplemonini 329 Anomalonychus Saylor 264 Aplocentrus LeConte 44 Anomalopus Casey 154 Aploderus Stephens 93 Anomoea Lacordaire 304 Apocellus Erichson 95 Anomoglossus Chaudoir 49 Apodasyini 295 Anomoia Dejean 304 Apothinus Sharp 126 Anomophagus Reitter 218 Apotomus Kirby 326 Anoplis Kirby 163 Apristus Chaudoir 58 Anoplitis Chapuis 321 Apteralium Casey 115 Anoplitis Kirby 321 Apterius Blatchley 317 AnopliurnHaldeman 280 Apteroloma Hatch 77 Anoplotrupes Jekel 150 Apteromechus Faust 348 Anotylus Thomson 95 Apterosoma Motschulsky 311 Antennalia Casey 278 Apterospasta LeConte 265 Anthaxia Eschscholtz 165 Araecerini 324 Anthaxiini 165 Araecerus Schtinherr 324 Anthericomma Pierce 366 Araeopidius Cockerell 143 Antherophagus Latreille 221 Araeopus LeConte 143 Anthicidae 271 Archostemata 6 Anthicinae 271 Arctelaphrus Semenov 16 Anthicini 271 ArctobiaThomson 15 Anthicus Paykull 271 Arctoclinidium Bell 7 Anthobates LeConte 276 Arctodytes Thomson 67 Anthobatulo Strand 276 Arctolina Kontkanen 312 Anthobium Leach 91 Areus Casey 109 Anthobium auct. 87 Argaleus LeConte 288 Anthocornus Erichson 211 Argante Gistel 162 Anthomarp,hus Weise 343 Argutor Dejean 34 Anthonominae 343 Arhopalus Audinet-Serville 279 Anthonomini 343 Aridius Motschulsky 240 Anthonomopsis Dietz 343 Arpedium Erichson 89 Anthonomus Germar 343 Arrhenodes Schiinherr 327 Anthophagini 89 Arrhenodini 327 Anthophylax LeConte 289 Arrhenopeplus Kocb. 84 Anthrenini 198 Arrhenoplita l{trby 255 Anthrenus Schaeffer 198 Artematopidae 174 Anthribidae 324 Arthrolips Wollaston 228 Anthribinae 324 Arthromacral{trby 262 Anthribini 324 Artochia Casey 90 Anthribulus LeConte 324 AsaphesKirby 181 Apator Semenov 67 Asaphidion des Gozis 20 Apatura Castelnau & Gory 165 Asclera Stephens 266 Apelocera Chevrolat 286 Asclerini 266 Apenes LeConte 59 Ascoliocerus M6quignon 178 399 Aseminae 279 Baeckmanniolus Reichardt 137 Asemini 279 Baeocera Erichson 125 Asemobius Horn 95 Bagoini 341 Asemum Eschscholtz 279 Bagous Germar 341 Asidini Il54 Balaninus Germar 342 Asidopsis Casey 254 Baliosus Weise 321 Asiobates Thomson 74 BalophilaCrotch 317 Aspidius Mulsant & Rey 257 Baridinae 351 Astenina 117 Baridini 351 Astenus Dejean 117 Baridius Schtinherr 351 Asteria Fauvel 113 Baris Germar 351 Astyleiopus Dillon 297 Barynotini 330 Astylopsis Casey 297 Barynotus Germar 330 Asynonyt:hus Crotch 329 Barypeithes Jacquelin du Val & Fairmaire 330 Ataenius Harold 149 Barytachys Chaudoir 28 Atheta Thomson 101 Bassareus Haldeman 307 Athetalia Casey 101 Batrisinae 127 Athetini 100 Batrisini 127 Athetota Casey 101 Batrisodes Reitter 127 Athoini .176 Batyle Thomson 287 Atholus llhomson 139 Batyleoma Casey 287 Athous Eschscholtz 177 Baudia Ragusa 49 Atimia Haldeman 280 Beeria Hatch 120 Atimiini 280 Bellamira LeConte 290 Atomaria Stephens 223 Belonuchus Nordmann 119 Atomariirae 223 Bembidiini 20 Atomarii:ri 223 Bembidion Latreille 20, 27 Atrachela. Gistel 308 Bembidionetolitzkya Strand 23 Atractopterus LeConte 182 Berninelsonius Leseigneur 178 Atranus l,eConte 52 Berosinae 131 Atrecus Jacquelin du Val 123 Berosus Leach 131 Attagenirri 197 Bessobia Thomson 102 Attagenus Latreille 197 Bibloplectus Reitter 127 Attalus Erichson 212 Bidensomela Monros 310 Attelabidae 326 Bidessini 63 Attelabini 326 Bidessonotus R6gimbart 63 Attelabus;Linn6 326 Biphyllidae 241 Atymius (]istel 309 Biphyllini 241 Auchenia Thunberg 304 Bitoma Herbst 251 Augyliini 169 Bius Mulsant 258 AulaconArnett 175 Blapstinus Dejean 257 Auletini 327 Blapylis Horn 257 Auletobiu.s Desbrochers 327 Blechrus Motschulskv 58 Auleutes Dietz 349 Bledius Leach 93 Aulobaris LeConte 352 Blemus Dejean 19 Aulonium Erichson 251 Blepharida Chewolat 316 Aulonothr:oscus Horn 186 Blepharidulo Weise 316 Autalia Leach 107 Blepharoplataphus Netolitzky 23 Autaliini 107 Blethisa Bonelli 15 Axinopalpus LeConte 58 Blitophaga Reitter 85 Bolboceras Kirby 150 Babia Chevrolat 305 Bolboceratini 150 Babiini 305 Bolbocerosoma Schaeffer 150 Babnormrrdes Park 127 Boliographa Motschulsky 310 Bacaniini 138 Bolitobiini 98 Bacanius LeConte 138 Bolitobius Leach 98 Bactridium LeConte 218 Bolitobus Tottenham 98 Badister Clairville 49 Bolitochara Mannerheim 107 400 Bolitocharina I07 Brennus Motschulsky 13 Bolitophaginl 255 Brenthidae 327 Bolitophagus Illiger 255 Bromius Chevrolat 309 Bolitotherus Candbze 255 Brontes Fabricius 220 Bomius LeConte 58 Broscini 19 Boreades Parsons 214 Broscodera Lindroth 19 Boreaphilus Sahlberg 89 Broscus Panzer 19 BoreobiaTschitsch6rine 31 Bruchidae 300 Boreocanthon Halffter 146 Bruchidius Schilsky 301 Boreoglymmius Bell & BeIl 7 Bruchinae 300 Boreophilia Benick 102 Bruchus Linn6 301 Boreosaxinis Moldenke 305 Bruesia Perkins 366 Boreostiba Lohse 101 Brumoides Chapin 233 Boridae 268 Bryaxis Leach 128 Borophloeus Wollaston 355 Brychius Thomson 60 Boros Herbst 268 Bryonomus Casey 119 Boschella Barr 210 Bryoporus lkaatz 98 Bostrichidae 199 Buprestidae 160 Bostrichinae 200 Buprestinae 162 Bostrichini 200 Buprestini 163 Bostrichoidea 199 Buprestis Linn6 163 Bothrideres Erichson 252 Buprestoidea 160 Bothriderinae 252 Byrrhidae 158 Bothriderini 252 Byrrhinae 158 Bothriopterus Chaudoir 32 Byrrhini 158 Brachiacantha Dejean 232 Byrrhodes LeConte 205 Brachinini 18 Byrrhoidea 158 Brachinus Weber 18 Byrrhus Linn6 158 Brachybamus Germar 341 Byrsopages Schijmherr 335 Brachycepsis Brendel 83 Bythinini 128 Brachycis Casey 245 Bythinopsis Raffray 128 Brachycoryna Gu6rin-M6neville 322 Byturidae 241 Brachycrepis LeConte 184 Byturini 241 Brachyderinae 329 Byturus Latreille 241 Brachyderini 330 Brachygluta Thomson 128 CacopliaLeConte 294 Brachyglutini 128 Cacoternnus LeConte 203 Brachyini 168 Cq.docera Eschscholtz 149 Brachyleptura Casey 293 Caenia Newman 189 Brachylobus Chaudoir 50 Caeniella Cockerell 189 Brachymorphzs Chewolat 210 Caenocara Thomson 204 Brachynotus Kirby 192 Caenocyrta Brown 79 Brachypnoeo Gistel 308 Caenoscelini 222 Brachypterolus Grouvelle 214 Caenoscelis Thomson 222 Brachypterus Kugelann 214 Caerosternus LeConte 138 Brachyrhinus Latreille 331 Cafius Stephens 119 Brachys Solier 168 Calandra Clairville 354 Brachysomida Casey 290 Calandrinus LeConte 353 Brachysomus Schijnherr 330 Calathus Bonelli 51 Brachystilus Chaudoir 31 Calitys Thomson 207 Brachytarsoides Pierce 324 Calleida Dejean 58 Brachyusa Mulsant & Rey 100 Callidiini 282 Bracteomimus Lindroth 21 Callidium Mulsant 283 Bracteon Bedel 20 Calligrapha Chewolat 310 Bradybatini 344 Callimellum Strand 282 Bradycellus Erichson 46 Callimoxys lkaatz 282 Bradytus Stephens 36 Callimus Mulsant 282 Brathinus LeConte 89 Callipara Motschulsky 10 401 Callisten,',a Vacher de Lapouge 11 Catharellus Casey 46 Callisthenes Fischer von Waldheim 11 Cathartosilvanus Grouvelle 220 Callisthiri 49 Catogenus Westwood 219 Callitrop;r Motschulsky 10 Catopocerinae 80 Calloides LeConte 285 Catopocerus Motschulsky 80 Callosobruchus Pic 300 Catops Paykull 81 Calochara Casey 104 Catoptrichus Murray 81 Calochromus Gu6rin-M6neville 188 Catorama Gu6rin-M6neville 205 Caloderma Casey 113 Cedius LeConte 129 Calodrepa Motschulsky 10 Celetes Newman 189 Calomycterus Roelofs 334 CeliaZimmermann 36 Calopodinae 266 Celiamorphus Casey 43 Calopterini 189 Celina Aub6 63 Calopteron Gu6rin-M6neville 189 Centrinapsis Casey 353 Calopus Irabricius 266 Centrinini 353 Calosirus Thomson 350 Centrinogyna Casey 353 Calosoma Weber 10 Centrodera LeConte 288 Calvia Mulsant 235 Centronopus Solier 258 Calyrnmaderini 204 Centuriatus Pacheco 169 Calymmaderus Solier 204 Ceophyllini 129 Calyptomerus Redtenbacher 141 Ceophyllus LeConte 129 Camptorl;.ino Kirby 151 Cephalogyna Casey 44 Campylir.hynchus Dejean 351 Cephaloidae 262 Canastottr Casey 100 Cephaloinae 262 Canifa LeConte 276 Cephaloon Newman 262 Canonura Casey 296 Ceracis Melli6 245 Cantharidae 192 Ceralimnius Hinton 173 Cantharinae 192 Cerambycidae 277 Cantharirri 193 Cerambycinae 280 Cantharis Linn6 193 CerandriaLucas 255 Cantharoidea 188 Cerataltica Crotch 317 Canthon .lloffmansegg L46 Ceratapion Schilsky 328 Canthosttthus Haldeman 306 Ceratoderus Blanchard 271 Caphalistes Motschulsky 212 Ceratographis Gahan 297 Capnochroa LeConte 260 CeratomaDozier 315 Capraita Bechyn6 320 Ceratomegilla Crotch 233 Carabidae 8 Ceratopidonia Linsley & Chemsak 293 Carabini 10 Cercolia Casey 293 Carabus Linn6 11 Cercopedius Sleeper 332 Carcinops Marseul 138 Cercyon Leach 134 Carcinus.Marseul 138 Cercyonini 134 Cardiophorinae 185 Cerophorus Castelnau 216 Cardiophorus Eschscholtz L85 CerotaruaBowditch 315 Carebq,ra LeConte 247 Cerotoma Chewolat 315 Carpelimus Leach 94 Ceruchini 144 Carphacis des Gozis 98 Ceruchus MacLeay 144 Carphoborus Eichhoff 359 Cerylon Latreille 227 Carphonobus Casey 356 Cerylonidae 227 Carpophilinae 2L4 Ceryloninae 227 Carpophilus Stephens 214 Cetoniinae 157 Carrhydms Fall 69 Cetoniini 157 Cartodere Thomson 240 Ceutorhynchidius Jacquelin du Val 349 Cassida Linn6 322 Ceutorhynchinae 349 Cassidinae 322 Ceutorhynchini 349 Cassidini 322 Ceutorhynchus Germar 349 Catapastus Casey 353 Chaerocassis Spaeth 323 Cateretes Herbst 214 Chaetarthria Stephens 131 Cateretinae 214 Chaetarthriinae 131 402 Chq.etechus Horn 332 Cicindelidia Rivalier 10 Chaetocnema Stephens 319 Cicindelini 8 Chaetophloeus LeConte 358 Cidnorhinus Thomson 350 Chaetophora Stephens 158 Clidae 244 Chalcoidea Motschulsky 311 Ciinae 244 Chalcoparia Crotch 308 Ciini 244 Chalcophora Solier 161 Cilea Jacquelin du Val 97 Chalcophorinae 161 Cimberidini 324 Chalcophorini 161 Cimberis des Gozis 324 Chalcohyus Blackman 359 Cimbocera Horn 333 Chalepini 321 Ciocotalpa Saylor 155 Chalepus Thunberg 321 Circinalia Casey 54 Chapmania Bernhauer 95 CircinalidiaCasey 54 Charaphloeus Casey 219 Circinoporus Guignot 65 Charhyphus Sharp 95 Circomus Fleischer 258 Charidotella Weise 323 Cis Latreille 244 Chariessa Perty 210 Cistelct. Fabricius 259 Charisalia Casey 291 Clambidae 141 Chauliognathinae 195 Clambus Fischer von Waldheim 141 Chauliognathini 195 Clamoris des Gozis 253 Chauliognathus Hentz 195 Clavigerinae I29 Chelonychus Dietz 343 Clavilispinus Bernhauer 92 Chelymorpha Chewolat 322 Cleonidius Casey 336 Chilocorinae 233 Cleoninae 336 Chilocorini 233 Cleonini 336 Chilocorus Leach 233 Cleonis Dejean 336 ChionNewman 286 Cleonus Schdnherr 336 Chionotyphlus Smetana 113 Cleopomiarus Pierce 347 Chirida Chapuis 323 Cleptelmis Sanderson 172 Chitalia Sharp 106 Cleridae 208 Chlaeniellus Reitter 50 Clerinae 208 Chlaenius Bonelli 49, 50 Clerini 209 Chlamisinae 307 Cleroidea 206 ChloephagaWeise 320 Clinidium Kirby 7 Cholevinae 81 Clivina Latreille 18 Cholevini 81 Clivinini 17 Choraginae 324 Clusiota Casey 102 Choragini 324 Clytini 284 Chramesus LeConte 358 Clytoleptus Casey 286 Chromatia LeConte 259 Clytrinae 304 Chrysobothrinae 166 Clytrini 304 Chrysobothrini 166 Clytus Laicharting 285 Chrysobothris Eschscholtz 166 Cnernidotus Erichson 61 Chrysobracteon Netolitzky 20 Cnemocyllus Dietz 343 Chrysochus Chevrolat 309 Cnemogonini 349 Chrysodina Baly 308 Cnemogonus LeConte 349 Chrysolina Motschulsky 311 Cneoglossini 143 Chrysomela auct. 311 Cneorhinini 330 Chrysomela Linn6 313 Coccidophilus Brethes 229 Chrysomelidae 301 Coccidula Kugelann 233 Chrysomelinae 310 Coccidulinae 233 Chrysomelini 310 Coccidulini 233 Chrysomeloidea 277 Coccinella Linn6 235 Chrysomelopsilo Strand 313 Coccinellidae 229 Chrysomelopsis Achard 313 Coccinellinae 233 Chrysophana LeConte 161 Coccinellini 233 Chrysostigma Kirby 10 Coccotorus LeConte 344 Cicindela Linn6 8 Coelambus Thomson 64 403 Coelini 254 Coptotomini 70 Coelocephalapion Wagner 328 Coptotomus Say 70 Coelocnernis Mannerheim 261 Copturodes Casey 348 Coelogaster Schiinherr 349 Coraebini 167 Coelometopinae 260 Cordalia Jacobs 105 Coelometopini 260 Cordoharpalus Hatch 41 Coelomorpha Casey 254 Coreopsomela Monros 310 Coelostethus LeConte 203 Corioxenidae 366 Coelotaxis Horn 253 Corioxeninae 366 Coelus Eschscholtz 254 CorphyraSay 274 Colaspis Fabricius 308 Corthylini 363 Colastus Erichson 214 Corthylus Erichson 365 Colenis Erichson 79 Corticaria Marsham 240 Coleomegilla Timberlake 233 Corticariinae 240 Coleothorpa Moldenke 305 Corticarina Reitter 241 Collapterl,'x Newman 294 Corticeus Piller & Mitterpacher 256 Colliuris DeGeer 56 Corticotomus Sharp 207 Collops Erichson 211 Cortilena Motschulsky 241 Colon Her:bst 80 Cortinicara Johnson 241 Coloninaer 80 Cortodera Mulsant 290 Colopterus Erichson 214 Corylophidae 228 Colposura', Casey 101 Corylophinae 228 Colydiida,: 251 Corymbites Latreille 179 Colydiinae 251 Coryphiini 89 Colydiini 251 Coryphium Stephens 89 Colydium Fabricius 252 Cosmobaris Casey 352 Colymbet,:s Clairville 69 Cosmosalia Casey 294 Colymbet.nae 67 Cosnania Dejean 56 Colymbet:ni 69 Cossoninae 355 Comstocle,'.a Van Dyke 52 Cossonini 355 Conalia lVlulsant &Rey 249 Cossonus Clairville 355 Coninomu'.s Thomson 240 Cotalpa Burmeister 155 Coniocleonus Motschulsky 336 Coxelus Horn 251 ConiontelLus Casey 253 Craponius LeConte 349 Coniontides Casey 253 Cratacanthus Dejean 39 Coniontini 253 Cratidus LeConte 257 Coniontis Eschscholtz 253 Cratomorphini 190 Coniophagus Mink 213 Cratonychus Dejean 184 Con iop hth:.alma Kangas 221 Crerya LeConte 210 Conithassa Thomson 239 Cremastocheilini 157 Connatichela Anderson 334 Cremastocheilus Knoch 157 Connophron Casey 82 Creniphilus Horn 132 Conoderus Eschscholtz 176 Crenitis Bedel 132 Conophthorus Hopkins 363 CrenitulusWinters 132 Conoplectus Brendel 126 Creophilus Samouelle 120 Conotelus Erichson 214 Crepidodera Chevrolat 319 Conotrachelus Dejean 347 Crepidophorus Mulsant & Guillebean 177 Copelatinae 62 Crepidotritus LeConte 184 Copelatini 62 Crine Pascoe 218 Copelatus Erichson 62 Criocephalum Dejean 27 I Cophes Champion 348 Criocephalus Mulsant 279 Copidita LeConte 266 Criocerinae 304 Coprini 1,46 Criocerini 304 Copris Mtiller 146 Crioceris Mi.iller 304 Coprochara Mulsant & Rey 104 CrioscaphaHorn 246 Coprophibrni 92 Crosimus Casey 221 Coprophilus Latreille 93 Crossidius LeConte 287 Coproporus Kraatz gG Cryrnodes LeConte 268 404 Cryobius Chaudoir 34 Cychrini 12 Cryocarabus Vacher de Lapouge 11 Cychrus Fabricius 13 Cryphalini 362 Cyclinus I{trby 72 Cryphalus Erichson 362 Cycliopleurus Hope 281 Cryptarcha Shuckard 217 Cyclocephala Latreille 155 Cryptarchinae 217 Cyclocephalini 155 Cryptobiina 116 Cycloneda Crotch 236 Cryptobiurn Mannerheim 116 Cyclotrachelus Chaudoir 35 Cryptocephalinae 305 Cylindera Westwood 10 Cryptocephalini 306 Cylindridia Casey 353 Cryptocephalus Miiller 306 Cylindrocharis Casey 31 Cryptocleptes Blackman 359 Cylindrocopturus Heller 348 Cryptohypnus Eschscholtz 17 8 Cylister Cooman 140 Cryptolestes Ganglbauer 220 Cylistix Marseul 140 Cryptophagidae 221 Cyllene Newman 284 Cryptophaginae 221 Cyllodes Erichson 217 Cryptophagini 221 Cyllopodes Murray 2\4 Cry ptophagistes Crotch 221 Cymatodera Gray 208 Cryptophagus Herbst 221 Cymatopterus Dejean 69 Cryptophilinae 224 Cymbiodyta Bedel 133 Cryptophilini 224 Cymindis Latreille 59 Cryptophilus Reitter 224 Cynaeus LeConte 255 Cryptopleurum Mulsant 135 Cypha Leach 108 Cryptorhopalum Gu6rin-M6neville 198 Cyphea Fauvel 107 Cryptorhynchidius Pierce 348 Cyphelophorus Kuwert 131 Cryptorhynchinae 347 Cyphicerini 334 Cryptorhynchini 347 Cyphina 107 Cryptorhynchus Illiger 348 Cyphirhinus Schiinherr 351 Cryptoweisea Gordon 229 Cyphomimus Horn 330 Cryptulocleplus Wood 359 CyphonPaykull 142 Crypturgini 361 Cyrtepistomus Marshall 334 Crypturgus Erichson 361 Cyrtinini 298 Ctenicera Latreille 179 Cyrtinus LeConte 298 Ctenistini 129 Cyrtocaraxenos Pierce 366 Ctenobium LeConte 202 Cyrtomoptera Motschulsky 193 Ctenodactylini 56 Cyrtomoptila Motschulsky 193 Ctenonychus Stephens 184 Cyrtophorus LeConte 286 Cucujidae 219 Cyrtoscydmus Motschulsky 83 Cucujinae 219 Cyrtotriplax Crotch 225 Cucujoidea 213 Cyrtusa Erichson 79 Cucujus Fabricius 219 Cytilus Erichson 160 Cupedidae 6 Cupedinae 6 Dacne Latreille 225 Cupedini 6 Dacninae 225 Cupes Fabricius 6 Dacyrtocara Pierce 366 Cupila Casey 127 Dalopius Eschscholtz 182 Curculio Linn6 342 Dalotia Casey 100 Curculionidae 329 Dampfius Pacheco 170 Curculionoidea 323 Danae Reiche 238 Curculioninae 342 Daniela Netolitzky 23 Curimopsis Ganglbauer 158 Danosoma Thompson 175 Curtonotus Stephens 35 Dascillidae 143 Curvimanon Fleischer 80 Dascillinae 143 Cybister Curtis 72 Dascilloidea 143 Cybistrini 72 Dasydera LeConte 150 Cybocephalinae 2I7 Dasyglossal{raatz 99 Cybocephalus Erichson 217 DasypogonLeConte 175 Cychramus Kugelann 216 Dasvrhadus Fall 213 405 Dasy'bes Faykull 212 Diaperini 255 Dasytinae 212 Diaperis Miiller 255 Datomicra Mulsant & Rey 100 Diaphoetes Waterhouse 120 Dearthrus LeConte 198 Diaphoromerus Chaudoir 45 Decarthron Brendel 128 Diaulota Casey 107 Dectes LeConte 297 Diazus LeConte 151 Deinopsini 108 Dibolia Latreille 320 Deinopsis Matthews 108 Dibolocelus Bedel 133 Deinopteroloma Jansson 91 Dicaelus Bonelli 48 Deleaster Erichson 92 Dicentrus LeConte 280 Deloyala Chewolat 323 Dicerca Eschscholtz 162 Delphacixenos Pierce 367 Dicercini 162 Delphastrrs Casey 229 Dicheirus Mannerheim 45 Delphota Jasey 103 Dichelonycho Kirby 153 Deltometopus Bonvouloir 187 Dichelonyx Harris 153 Deltostethus Sharp 134 Dichrornino Casey 155 Dendroctonus Erichson 357 Dictyopterus Mulsant 188 Dendroiders Latreille 269 Dicyrtocercyon Ganglbauer 134 Dendrophagus Schrinherc 220 Didetus LeConte 50 Dendrophilinae 138 Didion Casey 229 Dendrophilini 138 Dienerella Reitter 239 Dendrophilus Leach 138 Dietzella Champion 349 Denticollis Piller & Mitternacher t77 Dietzianus Sleeper 344 Deratanclrus Casey 55 Dimetrota Mulsant & Rey 100 Deratopeus Casey 115 Dimetrotina Casey 100 DerelomirLi 341 Dinaraea Thomson 102 Deretaphrini 252 Dineutus MacLeay 72 Deretaphrus Newman 252 Dinoderinae 200 Dermestes Linn6 196. 197 Dinodromius Casey 58 Dermestic{ae 196 Diocalandra Faust 355 Dermestirri 196 Diocarabus Reitter 12 Dermestinus Zhantiev 196 Diochini 123 Dermestoidea 195 Diochus Erichson 123 Derocrepis Weise 318 Diomus Mulsant 231 Derodontidae 195 DioxoceraPierce 366 Derodontinae 196 Diozocera Pierce 366 DerodontrLs LeConte 196 DiozodesHaldeman 281 Deronectes auct. amer. 66 DiphyaulonArnett 175 Derospidea Blake 314 Diplocampa Bedel 27 Derulus Tschitch6rine 31 Diplocheila Brull6 48 Derus Motschulsky 31 Diplocoelus Gu6rin-M6neville 241 Descarpentriesina Leraut 163 Diplotaxis Kirby 151 Desmatogaster Knutson 203 Diplous Motschulsky 30 Desmocerini 287 Dirabius Casey 353 Desmocerrrs Audinet-Serville 287 Dircaea Fabricius 248 Desmopachria Babington 63 Dirhaginae 187 Desmoris LeConte 340 Dirhagus Latreille 187 Devia Blar:kwelder 99 Disamara Lindroth 35 DiabroticeL Chewolat 315 Discoderus LeConte 43 Diacheila Motschulsky 15 Disonycha Chevrolat 319 Diachus LeConte 307 Distemmus LeConte 88 Diagraphct Newman 189 Distichalius Casey 122 Dialytellus Brown 149 Distigmoptera Blake 320 Dialytes Harold 149 Ditemnus LeConte 194 Dianchomeno Chaudoir 57 Ditropalia Casey 107 Dianous Samouelle 109 Ditylini 266 Diqnusa Casey 107 Ditylus Fischer von Waldheim 266 Diaperinae 254 Dolerosomus Motschulskv 182 406 Dolichocis Dury 244 Ecyrus LeConte 295 Dolichosoma Stephens 212 Elacatis Pascoe 270 Doliema Pascoe 255 Elaphidiini 280 Dolopiosomus Motschulsky 184 Elaphidion Audinet-Serville 281 Dolurgus Eichhoff 361 El ap hidionoide s Linsley 281 Donacia Fabricius 302 Elaphrini 15 Donaciella Reitter 302 Elaphropus Motschulsky 28 Donaciinae 302 Elaphrotatus Semenov 16 Donaciini 302 Elaphroterus Semenov 16 Donaciomima Medvedev 303 Elaphrus Fabricius 16 DonesiaCasey 103 Elasmocerus LeConte 208 Dorcaschema Haldeman 295 Elassoptes Horn 355 Dorcaschematini 295 Elater Linn6 181 Dorcasina Casey 291 Elateridae 175 Dorcatoma Herbst 204 Elaterinae 181 Dorcatominae 204 Elaterini 181 Dorcatomini 204 Elateroidea 174 Dorcini 144 Elateroides Schaeffer 206 Dorcus Macleay 144 Elathous Reitter 177 Dorthesia Say 250 Elatotrypes Fisher 284 Dorytomus Germar 339 Elatrinus Horn 184 Doydirhynchinae 324 Eleates Casey 255 Drachylis Casey 262 Eleodes Eschscholtz 256 Drapetes Dejean 186 Eleodini 256 Drasterius Eschscholtz 176 Eleodopsis Blaisdell 257 Drastophus Casey 82 Eleusinini 92 Dromaeolus Kiesenwetter 187 Elissa. Casey 331 Dromius Bonelli 58 Elleschini 345 Drusilla Leach 106 Elleschus Dejean 345 Dryocoetes Eichhoff 361 Ellipsoptera Dohktourow 10 Dryocoetini 361 Ellychnia LeConte 191 Dryophilinae 2Ol Elmidae 172 Dryophthorinae 356 Elminae 172 Dryophthorini 356 Elmini 172 Dryophthorus Germar 356 Elodes Latreille 142 Dryopidae 171 Elonium Leach 88 Dryopoidea 169 Elonus Casey 275 Dryops Olivier 171 Elytroleptus Dug6s 287 Dubiraphia Sanderson 172 ElytrusaCasey 101 Dularius Thomson 284 Embaphion Say 257 Dynastinae 155 Emmesa Newman 246 Dyschirius Bonelli 17 Empelinae 87 Dysidius Chaudoir 32 Empelus LeConte 87 Dyslobus LeConte 333 Emphanes Motschulsky 27 Dysmathes Mannerheim 61 Emphyastes Mannerheim 337 Dytiscidae 62 Emphyastini 337 Dytiscinae 71 Empinodes Park I27 Dytiscini 71 Emplenota Casey 105 Dytiscus Linn6 71 Enaphalodes Haldeman 280 Encalus LeConte 345 Eanus LeConte 181 Enchodes LeConte 247 Eburia Audinet-Serville 280 Encyclops Newman 288 Ecarinosphaerula Hatch 79 Endalus Castelnau 341 Echinaspis Blatchley 347 Endecatominae 200 Echocerus Horn 255 Endecatomus Melli6 200 Echochara Casey 105 Endectus LeConte 252 Echyra Erichson 154 Endeitoma Sharp 251 Ectopria LeConte 174 Endeodes LeConte 211 407 Endomychidae 237 Euaesthetus Gravenhorst 112 Endomychinae 239 EubaeoceraCornell 125 Endomychini 239 Eubriinae 174 Endomychus Panzer 239 Eubrychiopsis Dietz 351 Endrosq.LeConte 152 Eubrychius Thomson 351 Engamota Casey 100 Eubulus Kirsch 348 EngisPayxull 225 Eucanthus Westwood 150 Enhydrini 72 Euceratocerini 202 Enicmus Thomson 239 Euchaetes LeConte 353 Enicoceru-. Stephens 74 Euchariruq Casey 104 EnnearthronMelli6 244 Eucicones Sharp 251 Enochrus'lhomson 132 Eucinetidae 141 Enoclerus Gahan 209 Eucinetoidea l4l Enopliini 210 Eucinetus Germar 141 Enoplurus Hope 131 Euclyptus Dietz 341 Entomophthalmus Bonvouloir 187 Eucnecosum Reitter 90 Entomosce:Iis Chevrolat 313 Eucnemidae 186 Epauloecus Mulsant & Rey 205 Eucneminae 187 Epeuchaet,zs Lyman 353 Euconnini 82 Ephalus LeConte 257 Euconnus Thomson 82 Ephistemtrs Stephens 223 Eucrada LeConte 201 Epicauta Dejean 264 Eucradini 201 Epicautina 264 Eucryptusa Casey 107 Epierus Edchson 138 Euderces LeConte 286 Epilachna Dejean 237 Eudromus l{trby 21 Epilachninae 237 Euferonia Casey 33 Epilachnirri 237 Eugnamptus Schijnherr 326 Epiphanis Eschscholtz 188 Euharpalops Casey 41 EpiphloeirLi 210 Eulathrobium Casey 116 Episcopell,ts Casey 43 Euliusa Casey 100 Episernus Thomson 202 Eumalus Sharp 92 Epismilia C ocker ell 229 Eumichthini 282 Epitrix Foudras 319 Eumichthus LeConte 282 Eplophorus Chevrolat 286 Eumicrota Casey 107 Epuraea Erichson 215 Eumitocerus Casey 99 Epuraeanella Crotch 215 Eumolpinae 308 Erchomus Motschulsky 96 Eumolpini 308 Eremninar: 334 Eumolpus Redtenbacher 309 Eremosis cles Gozis 312 Eumorphinae 238 Ergates Audinet-Servllle 27 7 Eunonia Casey 93 Ergatini 2',77 EunyssobiaCasey 353 Erichsonius Fauvel 117 EuophryumBroun 355 Eriopsilus Bates 295 Eupactus LeConte 204 Erirhininae 339 Eupagoderes Horn 334 Erirhinini 339 Eupariini 149 Erirhinus Schiinherr 339 Eupariini 324 Erirhipidi,z Casey 157 Euparius Schiinherr 324 Erirhipis Ilurmeister 157 Eupetedromus Netolitzky 25 Ernobiinae 202 Euphorhipis Casey 157 Ernobiini 202 EuphoriaBurmeister 157 Ernobius illhomson 202 Euphoriaspls Casey 157 Eronyxa Reitter 207 Eupisenus Casey 243 Eropterus Green 189 Euplectinae 126 Eros Newrnan 189 Euplectini 126 Erotides $Iaterhouse 189 Euplectus Leach 126 Erotylidae 225 Eupleurida LeConte 269 Erynephala Blake 315 Eupogonius LeConte 295 Euaesthetinae Il2 Eupogonocherus Linsley 296 408 Eupristocerus Deyrolle 167 Fallapion Kissinger 328 EurealeConte 174 Faroninae 126 Eurelymis Casey 212 Faronini 126 Euremus Bierig 119 Feronia Latreille 30 Eurhizophagus M6quignon 218 Fidia Walsh 309 Europhilus Chaudoir 52 Flavohelodes Klausnitzer 142 Eurydactylus LaFert6-S6nectbre 49 Florilinus Mulsant & Rey 199 Euryderus LeConte 40 FluuicolaDeKay 174 Eurygeniinae 274 Fornax Castelnau 187 Eurygeniini 274 Foveoscapha Park & Wagner 126 Eurylister Bickhardt 140 Funda Blackwelder 104 Eurymeloe Reitter 263 Furcacampa Netolitzky 26 Eurymetopini 253 Fuscocalathzs Ndgre 51 Eurymycter LeConte 325 Euryoptera Horn 284 Gabrius Stephens 119 Eurypalpus LeConte 174 Gabronthus Tottenham 119 Eurypogon Motschulsky 175 Galerita Fabricius 59 Eurysphindinae 213 Galeritino Jeannel 59 Eurysphindus LeConte 213 Galeritini 59 Eurystethus Seidlitz 270 Goleritula Strand 59 Eurytrachelus Motschulsky 21 Galeruca Miiller 315 Eurytrichus LeConte 45 Galerucella Crotch 314 Eusomalia Casey 160 Galerucinae 314 Eusphalerini 87 Galerucini 314 Eusphalerum Kraatz 87 Galeruclerus Gahan 210 Eusphyrus LeConte 325 Gastrallini 202 Euspilotus Lewis 137 Gastrallus Jacquelin du Val 202 Eustilbus Sharp 226 Gastrellarius Casey 32 Eustrophinae 246 Gastroeideq.Hope 312 Eustrophinus Seidlitz 246 Gastrolobium Casey 116 Eustrophus Illiger 246 Gastrophysa Chewolat 312 Eutetrapha Bates 298 Gastrotaphrus Buchanan 338 Eutheia Stephens 83 Gaurambe Thomson 207 Eutheiini 83 Gaurodytes Thomson 67 Euthyna Gistel 304 Gauropterus Thomson 123 Eutoxotus Casey 288 Gaurotes LeConte 289 Eutrichapion Reitter 328 Gehringia Darlington 19 Eutrichites LeConte 128 Gehringiini 19 Euvacusus Casey 273 Gelus Casey 348 Euvrilletta Fall 204 Genattus Sharp 197 Euxenus LeConte 324 Gennadota Casey 99 Euxestinae 227 Genyon Smetana 134 Evarthrus LeConte 35 Geoderces Horn 333 Evodinus LeConte 289 Geodercodes Casey 332 Euoplus LeConte 255 Geodromicus Redtenbacher 91 Evotus LeConte 332 Geomorus Schiinherr 336 Excavodes Park 127 Geomysaprinus Ross 137 Exema Lacordaire 307 Geopatrobus Darlington 29 Exochomus Redtenbacher 233 Geopinus LeConte 43 Exomella Casey 159 Georyssidae 135 Exopioide s Gu6rin-M6n6ville 199 Georyssus Latreille 135 Exosternini 140 Geostiba Thomson 101 Explorator Pacheco 169 Geotrupes Latreille 150 Geotrupinae 150 Fabricianus Weise 307 Geotrupini 150 Falacer Castelnau 259 Gephelophorus Sharp 131 Falagria Leach 105 Geraeus Pascoe 353 Falagriini 105 Gerstaeckeria Champion 347 409 Gibbiinae 205 C4tmnodus Kirby 157 Gibbium Scopoli 205 Qtmnota Gistel 163 Gibbobruchus Pic 301 Gymnusa Gravenhorst 108 Glaphyra Newman 282 Gymnusini 108 Glaphyrinae 150 Gynandropus Dejean 43 Glaphyrocanthon Martinez 146 Gynandrotarsus LaFert6-S6nectbre 44 Glaresis Erichson 150 Gynopterus Mulsant & Rey 206 Glipa LeConte 249 Gyrinidae 72 Glipostenoda Ermisch 250 Gyrininae 72 Glischrochilus Reitter 217 Gyrinini 73 Glycerius Casey 46 Gyrinulus Zaitzev 73 Glycobius LeConte 284 Gyrinus Mi.iller 73 Glyphony>: Candbze 182 Gyrohypnus Samouelle 124 Glyphuroplata Uhmann 322 Gyrophaena Mannerheim 106 Glyptasida Casey 254 Gyrophaenina 106 Glyptina l,eConte 317 Gyrophaenini 106 Glyptoderus LaFert6-S6nectdre 49 Gyrotus Casey 348 Glyptoma Erichson 92 Glyptophorus Park 221 Habrocerinae 99 Glyptoscelis Chevrolat 309 Habrocerus Erichson 99 Gnathacmaeops Linsley & Chemsak 290 Hadraule Thomson 245 Gnathium Kirby 266 Hadrobregmini 203 Gnathoncus Jacquelin du Val 138 Hadrobregmus Thomson 203 Gnathospa'.sta Horn 264 Hadroplontus Thomson 350 Gnathotri<'hoide s Blackman 3 64 Hadrotes Miiklin 120 Gnathotrit:hus Eichhoff 364 Haemoniini 303 Gnathusa Fenyes 99 Haida Keen 91 Gnatocerus Thunberg 255 Halictophagidae 366 Gnorimellrr Casey 158 Halictophaginae 366 Gnypeta Thomson 100 Halictophagus Bohart 366 Goes LeConte 295 Haliplidae 60 Golgia Mulsant 238 Haliplus Latreille 60 Goniacerirrae 128 Hallomeninae 246 Goniaderinae 262 HallomenusPanzer 246 Gonioctena Chewolat 313 Halobrecthina Bernhauer 103 Goniolophus Casey 47 Hamq.ba Casey 346 Gonocallus LeConte 282 Hapalaraea Thomson 88 Gonocephalum Chevrolat 257 Hapalorhinus LeConte 211 Gonoderini 259 Hapalosalia Casey 292 Gonoderus Motschulsky 31 Haplandrus LeConte 260 Gonops LeConte 325 Haplanthaxia Reitter 165 Gonotropi:t LeConte 325 Haplochile LeConte 29 Grammoptr;era Audinet-Serville 291 Haplocoelus Chaudoir 31 Graphicallo Monros 310 Haplophoria Casey 157 Graphisurus Kirby 297 Haplorhynchites Voss 327 Graphoderus Dejean 71 Haplostethops Casey 353 Graphops LeConte 309 Hargium Samouelle 289 Graptodera Chewolat 317 Harminius Fairmaire 177 Gratusus (Jasey 242 Harmoniaspls Casey 236 Gronevus Casey 228 Harpalellus Lindroth 42 Grynocharis Thomson 207 Harpalini 39 Grypidius Schiinherr 339 Harpalobrachys Tschitsch6rine 40 Grypus Germar 339 Harpalomerus Casey 41 Gymnetrirrae 346 Harpalus Latreille 40, 4l Gymnetrini 346 Harpasta Gistel 305 Gymnetron Schtinherr 346 Hebestola Haldeman 294 Gymnis LeConte 152 Helichus Erichson 171 Gymnocht.hebius d'Orchymont 74 Helobium Leach 15 410 Helocerus Mulsant & Rey 199 Himatium Wollaston 355 Helocharini 132 Himatolabus Jekel 326 Helocombus Horn 133 HippodamiaDejean 234 HelodesPaykull 312 Hippopsini 296 Helophorinae 130 Hippopsis Lepeletier & Audinet-Serville 296 Helophorus Fabricius 130 Hippuriphila Foudras 318 Helopini 258 Hirmoplataphus Netolitzky 23 Helops Fabricius 258 Hispinae 321 Hemantus Casey 273 Hister Linn6 139 Hemicallidium Casey 283 Histeridae 135 Hemicarabus G6hin 12 Histerinae 139 Hemicoelus LeConte 203 Histerini 139 Hemicrepidius Germar 181 Histeropsis Chevrolat 255 HemicyphonLeConte 142 Holciophorus LeConte 31 Hemimedon Casey 113 Holoboreaphilus Campbell 89 Hemiptychus LeConte 205 Hololepta Paykull 140 Hemiquedius Horn 122 Hololeptini 140 Hemisopalus Casey 43 Holopleura LeConte 287 Hemispilota Casey 154 Holopleurini 287 Hemistenus Motschulsky 109 Holostrophus Horn 246 Henoticoides Hatch 221 Holotrochus Erichson 92 Henoticus Thomson 221 Homaeotarsus Hochhuth 116 Henotiderus Reitter 221 Homalochthebius Kttwefi 7 4 Henous Haldeman 264 Homalostylus Chevrolat 354 Hesperobaenus LeConte 2I8 Homalota Mannerheim 107 Hesperobium Casey 116 Homalotina 107 Hesperolinus Casey 124 Homalotusa Casey 102 Hesperophanes Mulsant 280 Homoeocarabus Reitter 11 Hesperophanini 280 Homoeolabus Jekel 326 Hesperosaprinus Wenzel 137 Hornoeosomo Austin 80 Hesperus Fauvel 119 HomogasterProvancher 348 Hetaeriinae 141 Homorosoma Frivaldszky 350 Hetaeriini 141 Hoplandria Kraatz 105 Hetaerius Erichson 141 Hoplandriini 105 Heterachthes Newman 281 Hoplia Illiger 154 Heteracthon Thomson 281 Hopliini 154 Heterarthron Dejean 199 Hoplocephalo Castelnau & Brull6 255 Heteraspis LeConte 309 Hoppingiana Blaisdell 212 Heterhelus Jacquelin du Val 214 Horistonotus Canddze 185 Heterlimnius Hinton 173 Hormorini 332 Heteroceridae 169 Hormorus Horn 332 Heterocerini 170 Hornaltica Barber 318 Heterocerus Fabricius 170 Hornia Riley 265 Heterodromla Haldeman 220 Horticola Lindemann 241 Heteronychus Seidlitz 67 Hybodera LeConte 282 Heterops Mannerheim 90 Hyboderini 282 Heteropus Castelnau 257 Hydaticini 71 HeterosilphaPortevin 85 Hydaticus Leach 71 Heterosternus Zimmermann 65 Hydaticus Sch6nherr 350 Heterosternuta Strand 65 Hydnobiini 78 Heterostethus Falkenstrom 65 Hydnobius Schmidt 78 Heterothops Stephens 121 Hydnocera Newman 208 Hetoernis Haldeman 295 Hydraena Kugelann 74 Hexarthrum Wollaston 355 Hydraenidae 74 Hexirhexius Grigarick & Schuster 126 Hydraeninae 74 Hiloro Mulsant & Rey 102 Hydraenini 74 Hilarina Casey 100 Hydriomicrus Casey 23 Himatinum Cockerell 355 Hydrium LeConte 21 tr Hydrobiinae 131 Hypoborini 358 Hydrobiini 131 Hypocaccus Thomson 137 Hydrobiusr Leach 131 HypocoelusDejean 188 Hydrocan-:hus Say 62 Hypocoprinae 223 HydrocharaBerthold 133 Hypocoprus Motschulsky 223 Hydrochinae 130 Hypocyphtus Gyllenhal 108 Hydrochu,s Leach 130 Hypodacne LeConte 227 Hydrogale,ruco Laboissidre 314 Hypolithus Eschscholtz 178 Hydrophilidae 130 Hypomedon Mulsant & Rey 113 Hydrophilinae 133 HypomolyxleConte 337 Hydrophiloidea 130 Hypophlaeus Fabricius 256 Hydrophilus Miiller 133 Hypophloeinae 256 Hydroporinae 63 Hypophloeini 256 Hydroporini 64 Hypostenus Rey 109 Hydroponrs Clairville 65 Hypothenemus Westwood 362 HydrosmectaThomson 100 Hypulini 247 Hydrothassa Thomson 312 Hypulus Paykull 247 Hydrovatini 63 Ibidiini 281 Hydrovatrrs Motschulsky 63 262 Hygrotus Stephens 64 Ichnodes Dejean 195 Hylastes Ilrichson 356 Ichthyurini 134 Hylastini 356 Ictinosternun Smetana 259 Hylastinotdes Spessivetsev 357 Idiobates Casey Swaine & Hopping 291 Hylastinus Bedel 357 Idiopidonia Hylecoetirrae 206 Idiostethus Casey 353 Hylecoetus Latreille 206 Ilendus Casey 242 Hylesininae 356 Ilybiosoma Crotch 67 Hylesinin; 357 Ilybius Erichson 69 99 Hylesinus Fabricius 357 Ilyobates Kraatz 77 Hylis des,]ozis 188 Ipelates Reitter 261 Hylobiini 337 Iphthiminus Spilman Hylobius (]ermar 337 Ipini 360 360 Hyloceotu,:,; Melsheimer 206 Ips DeGeer 12 Hylocurus Eichhoff 359 Irichroa Newman 187 Hylota Casey 99 Isarthrus LeConte Hylotrupes Audinet-Servllle 282 Ischalia Pascoe 269 269 Hylurgopinus Swaine 357 Ischaliinae Hylurgops LeOonte 356 Ischnosoma Stephens 98 273 Hymenochara Campbell 260 Ischyropalpus LaFert6-S6nectdre 226 Hy menoph.orus Mulsant 259 Ischyrus Lacordafte 345 Hymenortrs Mulsant 259 Isochnus Thomson Hypebaeus Kiesenwett er 212 Isoglossa Casey 99 209 Hypera Gr:rmar 336 Isohydnocera Chapin 259 Hyperaspidius Crotch 231 Isomira Mulsant Hyperaspini 231 Isoplastus Horn 79 Hyperaspis Redtenbacher 23I Isopleurus Kirby 36 Isorembus 48 Hypericia Bedel 311 Jeannel Hyperinae 336 Isorhipina Casey 157 Boisduval & Lacordaire 187 Hyperini 336 Isorhipis Hyperisus Mulsant &Rey 202 Isosceles Newman 299 Isostenia Vacher de Lapouge 11 Hypermal,t u.s Lacordaire 280 Isotachys Casey 29 Hyperodes Jekel 334 Hyperplatys Haldeman 298 Ithyceridae 325 Sch6nherr 325 Hypherpes Chaudoir 31 Ithycerus 347 Hyphydrini 63 Ithyporini Tempere 328 Hypnoryra Casey 124 Ixapion Roudier & .Irios Deville 328 Hypnoidus Dillwyn 178 4t2 Jonthonota Spaeth 322 Latheticus Waterhouse 258 Judolia Mulsant 291 Lathridiidae 239 Lathridiinae 239 Kalissus LeConte 84 Lathridius Herbst 239 Katastylops Pierce 367 Lathrimaeunz Erichson 91 Knowltonia Fisher 166 Lathrium LeConte 89 Knulliana Linsley 286 Lathrobiella Casey 116 Korynetinae 210 Lathrobiina 115 Korynetini 211 Lathrobioma Casey 115 Kuschelina Bechyn6 320 Lathrobium Gravenhorst 115 Kytorhininae 300 Lathrolepta Casey 115 Kytorhinus Fischer von Waldheim 300 Lathrotaxis Casey 116 Lathrotropis Casey 116 Labidomera Chevrolat 311 Lebia Latreille 56 Laccobius Erichson 131 Lebiini 56 Lacconotus LeConte 267 Lechriopini 348 Laccophilinae 63 Lechriops Schcinherr 348 Laccophilini 63 Lecontea Olivier 190 Laccophilus Leach 63 Leconteus Lutshnik 30 Laccornis des Gozis 67 Lecontia Champion 268 Lachnocrepis LeConte 49 Lederia Reitter 246 LachnosternaHope 152 Leiestini 238 Lacon Castelnau 175 Leiochiton Curtis 19 Laemophloeinae 219 Leiodes Latreille 79 Laemophloeus Dejean 219 Leiodidae 77 Laemosaccini 342 Leiodinae 78 Laemosaccus Schdnherr 342 Leiodini 78 Laemostenus Bonelli 51 Leiolinus Casey 124 Laestris Melsheimer 109 Leionotus Kirby 71 Laetulonthus Moore & Legner 119 Leirus Zimmermann 35 Lagarus Chaudoir 33 Leistus Frrilich 13 Lagriidae 261 Lema Fabricius 304 Lagrlinae 262 Lemiini 304 Lamiinae 294 Lerua Casey 7I3 Lamiota Casey 101 Leperisinus Reitter 357 Lamperos Allard 258 Lepesoma Motschulsky 333 Lamprias Bonelli 56 Lepiarcha Sharp 217 Lampropterus Mulsant 282 Lepidophorus Kirby 333 Lampyridae 190 Lepidotus Stephens 175 Lampyrinae 190 Lepisomus Kirby 359 Languria Latreille 224 Leptacinodes Casey 123 Languriidae 223 Leptacinus Erichson 123 Languriinae 224 Leptacmaeops Casey 290 Langurlini 224 Leptaegialia Brown 147 Lanternarius Pacheco 170 Leptalia LeConte 289 Lapethus Casey 227 Leptinillus Horn 82 Laphyra Newman 282 Lentininae 82 Lappus Casey 273 Leptinotarsa StAl 311 Lapsus Pacheco 170 Leptinus Mtiller 82 Lara LeConte 172 Leptoferonia Casey 32 Laricobius Rosenhauer 196 Leptomus Casey 8 Lafinae I72 Leptophloeus Casey 220 Larini 172 Leptopiini 333 Larisia Err,ery 276 Leptorus Casey 114 Lasconotus LeConte 251 Leptoscydmini 83 Lasioderma Stephens 204 Leptoscydmus Casey 83 Lasiodermini 204 Leptostylus LeConte 297 Lasiotrechus Ganglbauer 19 Leptothrix Heikertinger & Csiki 318 413 Leptotrachelus Latreille 56 Liparocephalus Miiklin 108 Leptotrichaltica Helkertinger 3 18 Lispinini 92 LeptotrixHorn 318 Lissominae 186 Leptotyphlinae 113 Lissorhoptrus LeConte 341 Leptura Linr'6 292 Listemus Casey 160 Lepturges Bates 297 Listronotus Jekel 334 Lepturinae 287 Listrus Motschulsky 213 Lepturini 288 Litargus Erichsor' 242 Lepturopsis Linsley & Chemsak 293 Litheleodes Blaisdell 256 Leptusa I{raatz 107 Lithocharis Dejean 113 Lepyrus Germar 337 Lithocharodes Sharp 123 LestqBlackwelder 90 Lithodus Germar 335 Lesteva Latreille 90 Litochropus Erichson 226 Leuchydrium Casey 25 Litodactylus Redtenbacher 350 Leucopaed,erus Casey 116 Litolethra Casey 115 Lexiphaners Gistel 306 Lixellus LeConte 335 Liaphlus ()uignot 61 Lixestus Reitter 336 Liq,semum, Casey 279 Lixini 337 Librodor Reitter 217 Lixus Fabricius 337 LichenophLanes Lesne 200 Loania Kinzelbach 366 LichnantL,e Burmeister 150 Loberini 224 Lichnocarabus Reitter 11 Loberus LeConte 224 Licinini 48 Lobiopa Erichson 216 Ligmargus Stibick 178 Lobosoma Buchanan 338 Lignyodes Dejean 345 Lobrathium Mulsant & Rey 116 Lignyodini 345 Loceptes Casey 347 Ligyrellus Casey 156 Lodinus Mulsant &Rey 257 Liryrodes Casey 156 Longitarsus Berthold 317 Ligyrus Burmeister 156 Lopha Stephens 27 Lilioceris Reitter 304 Lophalophus LeConte 333 Limnebiui; Leach 74 Lopheros LeConte 189 Limnichiciae 171 Lophioderini 82 Limnichir.ae 171 Lophioderus Casey 82 Limnichir.i 171 Lophoglossus LeConte 30 Limnichites Casey 171 Lophomucter Notman 105 Limnocharis Horn 74 Lophopogonius Linsley 296 Limonius Eschscholtz 176 Lordithon Thomson 98 Limulodes Matthews 76 Loricera Latreille 15 Limulodidae 76 Loricerini 15 LinaDejean 313 Lorinota Casey 105 Lindorus'Casey 233 Loxandrus LeConte 30 Liniolis Grigarick & Schuster 126 Loxopeza Chaudoir 56 Linohespe'rus Smetana 123 Lucanidae 143 Linolathro.Casey 116 Lucaninae 144 Linsleya MacSwain 265 Lucanini 144 Linsleyana, Podany 288 Lucanus Scopoli 144 Linsleyina Selander 265 Lucidota Castelnau 191 Liocellus Motschulsky 46 Lucifotychus Park & Wagner 129 Liocellus Tschitch6rine 46 Ludius Berthold 181 Liocosmius Casey 25 Lumetus Zaitzev L32 Liocyrtusa Daffner 79 Luperini 315 Liodessus Guignot 63 Lutrochus Erichson 171 Liodicaelus Casey 49 Lycidae 188 Liogluta Thomson 102 Lycinae 188 Lioligus Casey 159 Lycoperdina Latreille 238 Lionepha Casey 21 Lycoperdinini 238 Lionothus Brown 78 Lyctidae 201 Lioon Casey 159 Lvctini 201 4L4 Lyctus Fabricius 201 Mathrilaeurn Moore 91 Lygerus Kiesenwetter 195 Matini 71 Lygistopterini 188 Matus Aub6 71 Lymantor Lovendal 361 Mecas LeConte 299 Lymexylidae 206 Mecinini 347 Lymexyloidea 206 Mecinus Germar 347 Lyperopherus Motschulsky 35 Mecopeltus Dietz 351 Lyperosomus Motschulsky 33 Mecynotarsus LaFert6-S6nectdre 271 Lyperostenia Vacher de Lapouge 10 Medon Stephens 113 Lyperus Chaudoir 33 MedonellaCasey 113 Lypoglossa Fenyes 103 Medonina 113 Lyrosoma Mannerheim 77 Medonodonta Casey 114 Lytta Fabricius 264 Megacerus Fahraeus 300 Lyttina 264 Megacrotono Scheerpeltz 103 Lyttim 264 Megacyllene Casey 284 Megalechthrus Perkins 366 Macdonaldium Abdullah & Abdullah 76 Megaleptura Casey 292 Machaerodes Brendel 128 Megaliridia Casey 12 Macratria Newman 275 Megalodacne Crotch 225 Macratriini 275 Megalostomini 305 Macreurops Casey 218 MegalticaHatch 317 Macrobasis LeConte 265 Meganemognatha Enns 265 Macrodactylini 153 Megapangus Casey 40 Macrodactylus Latreille L54 Megapenthes Kiesenwetter 184 Macrodea Casey 222 Megaquedius Casey 121 MacrodytesThomson 71 Megarafonus Casey 126 Macronaemia Casey 233 Megarthrus Curtis 87 Macronoxia Crotch 152 Megasemum l{raatz 279 Macronychini 173 Megasternini 134 Macronychus Miiller 173 Megasternum Mulsant 134 Macropogon Motschulsky 174 Megastilicus Casey 114 Macropogoninae 174 Megatoma Herbst 198 Macrops Kirby 334 Megatomini 197 Macrorhoptus LeConte 343 Megilla Mulsant 233 Macrosiagon Hentz 250 Megodontus Solier 12 Macrotelus Klug 208 Megomus Casey 8 Maculicq.uta Dillon 264 Meibomeus Bridwell 301 Madarellus Casey 353 Melalgus Dejean 199 Madarini 352 Melamomphus Horn 333 Magdalinae 342 Melanagonum Casey 53 Magdalini 342 Melandrya Fabricius 246 Magdalis Germar 342 Melandryidae 245 Malachiinae 2l]- Melandryiinae 246 Malachius Fabricius 211 Melandryini 246 Malacocis Gorham 245 Melaneleodes Blaisdell 256 Mallodrya Horn 243 Melanius Bonelli 33 Malporus Casey 273 Melanophila Eschscholtz L65 Malthacus Kirby 192 Melanophilini 165 Malthinini 194 Melanophthalma Motschulsky 240 Malthinus Latreille 194 Melanotini 184 Malthodes Kiesenwetter 194 Melanotrupes Blanchard 150 Mantura Stephens 319 Melanotus Eschscholtz 184 Maphoca Casey 245 Melanthaxia Richter 165 Margaiostus Stibick 178 Melasinae 187 Margarinotus Marseul 139 Melasis Olivier 187 Marginus LeConte 241 Melq.soma Stephens 313 Margus Dejean 258 Melasomida Schaeffer 312 Masoreini 56 Melba Casey 127 4r5 Meligethes Stephens 217 Microhyus LeConte 347 Meligethinae 217 MicrolathraCasey 116 Melittophcgus Kirby 263 Microlestes Schmidt-Goebel 58 Meloe Lir.n6 263 Microlipus LeConte 211 Meloidae 263 Micromalthidae 6 Meloinae 263 Micromalthus LeConte 6 Meloini 263 Micromaseus Casey 34 MelolontkLinae 151 MicromedonCasey 113 Melolonthini 151 Micromelomalus Casey 22 Melomalus Casey 22 Micromota Casey 100 Melyridaer 211 Micronychus Provancher 333 Membrqctxenos Pierce 366 Micropeplidae 84 Menechid'zs Motschulsky 258 Micropeplini 84 Meracantha Kirby 259 Micropeplus Latreille 84 Merhynchites Sharp 327 Microrhagus Dejean 187 Meridion des Gozis 283 Microrhopala Chewolat 322 Meriellum Linsley 283 Microsaurus Stephens 121 Merinus LeConte 260 Microscaph a LeConte 246 Merium LeConte 283 Microschatia Solier 254 Meronera Sharp 100 Microscydmus Saulcy & Croissandeau 83 Mesauletes Voss 327 Microtachys Casey 29 Mesosini 295 Microtonus LeConte 247 Metablap";'lis Blaisdell 257 Microweise a Cockerell 229 Metqbletus Schmidt-Goebel 58 Microweisini 229 Metabola Chaudoir 57 Millidium Motschulsky 75 Metachro:na Chewolat 308 Minyomerus Horn 331 Metaclisa Jacquelin du Val 258 Mioptachys Bates 29 Metacmaeops Linsley & Chemsak 290 Miscodera Eschscholtz 19 Metallina Motschulsky 21 Mitosynum Campbell 93 Metallogra,pho Motschulsky 310 Mocyta Mulsant & Rey 102 Metamelanlus Tschitsch6rine 33 Molamba Casey 228 Metaxyodonla Casey 113 Molorchini 282 Methlini 63 Molorchus Fabricius 282 Methydrus Rey 132 Moluciba Casey 99 MetoponWaterhouse 327 Molytinae 337 Metrius Eschscholtz 19 Monachini 306 Mezium Clurtis 205 Monachulus Leng 306 Miarus Schiinherr 347 Monachus Chevrolat 306 Miccotrogus Schajnherr 346 Monarthrum Kirsch 365 Micracini 359 Moneilema Say 294 Micracis l,eConte 359 Moneilemini 294 Micracisoides Blackman 359 M onillop atrob us }Iatcli' 29 Micraegialia Brown 146 Monocerus Faldermann 27 | Micragonum Casey 54 Monochamini 294 Micralymma Westwood 88 Monochamus Dejean 294 Micrathous Lane I77 MonoferoniaCasey 32 Micrepitrix Laboissidre 318 Mononychini 349 Microaug;vles Pacheco 170 Mononychus Germar 349 Microbledius Herman 93 Monophylla Spinola 208 Microbregma Seidlitz 203 Monoplesa Motschulsky 294 Microcallidium Casey 283 Monotoma Herbst 218 Microcara Thomson 142 Monotominae 218 Microclytus LeConte 286 Monoxia LeConte 315 Microcylloepus Hinton 173 Mordella Linn6 248 Microdota Mulsant & Rey 102 Mordellidae 248 Microedus LeConte 91 Mordellini 248 Microgoes Casey 295 Mordellistena Costa 249 Micrograrnrne W alkJey 239 Mordellistenini 249 416 Morphnosoma Lutshnik 33 Neacanthocinus Dillon 296 Morychus Erichson 160 Neandra Lameerc 277 Motschulskium Matthews 75 Neanthophylax Linsley & Chemsak 289 Muirixenos Pierce 367 Neatus LeConte 259 Mulsantina Weise 237 Nebria Latreille 14 Murmidiinae 227 Nebriini 13 Murmidius Leach' 227 Necrobia Olivier 211 Mutinus Casey 112 Necrodes Leach 85 Myas Dejean 31 Necrophila Kirby & Spence 85 Mycetaea Stephens 238 Necrophilus Latreille 77 Mycetaeinae 238 Necydalini 287 Mycetaeini 238 Necydalis Llnn6 287 Mycetina Mulsant 239 Nedyus Sch 4r7 Neophytobius Wagner 351 Obriini 281 Neopolya.rthron Semenov 278 Obrium Dejean 281 Neoporus Guignot 65 Ocalea Erichson 99 Neopyrochroa Blair 269 Occlephelinus Hatch 89 Neorthopleura Barr 210 Ochodaeinae 149 Neosaprinus Bickhardt 137 Ochodaeus Audinet-Serville 149 Neoscutopterus Balfour-Browne 70 Ochrosidia Casey 155 Neostylops Pierce 367 Ochthebiinae 74 Neotrochus Blackwelder 92 Ochthebius Leach 74 Neozonitis Enns 265 Ochthedromus LeConte 21 Nephanes Thomson 76 Ochthephilum Stephens 116 Nephus Mulsant 231 Ochthephilus Mulsant & Rey 94 Nicaginae 144 Ochthexenus Motschulsky 88 Nicagini 144 Octotemnus Melli6 245 Nicagus l,eConte 144 Ocyusa l{raatz 99 Nicobiini 202 OdacanthellaLiebke 56 Nicrophorinae 85 Odacanthini 56 Nicrophorus Fabricius 85 Odontaeus Fischer von Waldheim 150 Niptus Boieldieu 205 Odontium LeConte 20 Nlsa Casey 128 Odontocorynus Schijnherr 353 Nisaxis Casey 128 Odontomus Kirby 166 Nitidula Fabricius 216 Odontonyx Stephens 51 Nitidulidae 214 Odontopus Say 345 Nitidulinae 215 Odontosphindini 213 Nitionini 254 Odontosphindus LeConte 213 Nocheles LeConte 333 Odontota Chewolat 321 Noda Chevrolat 308 Odontotaenius Kuwert 145 Nodinini 308 OebaceresThomson 295 Nodonota Lefbvre 308 Oedemeridae 266 Nomaretus LeConte 13 Oedemerinae 266 Nomaspis LeConte 264 Oedostethus LeConte 178 Nomidus Casey 334 Oestodes LeConte 181 Nomiini 29 Oestodini 181 Nominus Motschulsky 59 Oiceoptoma Leach 85 Nomius Castelnau 29 Olibrini 226 Nordenskioeldia Sahlberg Il2 Olibrus Erichson 226 Nosodendridae 196 Oliganomalo Casey 154 Nosodenclron Latreille 196 Oligolinus Casey 123 Nosodes LeConte 207 Oligomerodes Fall 204 Notaphusr Stephens 25 Oligomerus Redtenbacher 202 Notaris Ciermar 339 Oligomia Casey 100 Noteridaer 61 Oligopterus Casey 113 Nothopus LeConte 40 Oligotini 108 Nothus Olivier 248 Olisthaerinae 92 Notiodes Schiinherr 341 Olisthaerus Dejean 92 Notionomus Erichson 341 Olisthopus Dejean 51 Notiophilini 15 Olla Casey 236 Notiophilus Schtinherr 341 Olophrum Erichson 89 Notiophilus Dum6ril 15 Omq.lq. Motschulsky 27 Notobia Perty 45 Omaliinae 87 Notoxinae 271 Omaliini 88 Notoxini 271 Omalium Gravenhorst 88 Notoxus Fabricius 271 Omq.seidius Jeannel 33 Novelsis Casey 197 Omaseulus Lutshnik 34 Nudatoconnus Franz 83 Omuseus Dejean 33 Nudobius Thomson l-23 Omias Germar 332 Omoglymmius Ganglbauer 7 Oberea Mulsant 299 Omonadus Mulsant & Rev 273 418 Omophron Latreille 16 OrsonyxLeConte 151 Ornophronini 16 Orthaltica Crotch 318 Omorgus Erichson 150 Orthoboris Casey 351 OmoscylonThomson 294 Orthocis Casey 245 Omosita Erichson 216 Ortholeptura Casey 293 Omus Eschscholtz 8 Orthoperini 228 Oncideres Lepeletier & Audinet-Serville 296 Orthoperus Stephens 228 Onciderini 296 Orthoris LeConte 352 Onibathum Tottenham 87 Orthosoma Audinet-Sewllle 27 7 OnichodonNewman 187 Orthotomicus Ferrari 360 Ontholestes Ganglbauer 120 Orthotomides Wood 360 Onthophagini 146 Orus Casey 114 Onthophagus Latreille 146 Oryctini 156 Onthophilus Leach 138 Oryzaephilus Ganglbauer 220 Onychobaris LeConte 352 Osmoderma LePeletier & Audinet-Serville 157 Onychotrupes Jekel 150 Osmodermini 157 Onychylis LeConte 341 Osoriinae 92 Oodes Bonelli 49 Osoriini 92 Oodini 49 Osphya Illiger 248 Opatrinae 256 Osphyinae 248 Opatrini 257 Osphyini 248 Ophistomis Thomson 290 Ostoma Laicharting 207 Ophonus Dejean 40 Ostomodes Reitter 207 Ophraella Wilcox 315 Othiini 123 Ophryastes Say 334 Othniinae 270 Ophryastini 334 Othnius LeConte 270 Opilo Latreille 209 Otidocephalinae 341 Opisthiini 15 Otidocephalini 341 Opisthius Kirby 15 Otidocephalzs Chevrolat 341 Oplosia Mulsant 296 Otiorhynchinae 331 Opresus Casey 83 Otiorhynchini 331 Opsimini 280 Otiorhynchus Germar 331 Opsimus Mannerheim 280 Oulema des Gozis 304 Optioservus Sanderson 173 Oulimnius des Gozis 173 Opuntiaphila Pierce 347 Oxoplus LeConte 287 Orchesia Latreille 246 Oxybleptes Smetana 124 Orchesiini 246 Oxygonus LeConte 180 Orchestes Illiger 345 Oxylaemus Erichson 252 Ordobrevia Sanderson 173 Oxymedon Casey 113 Oreina Monros & Bechyn6 311 Oxynychus LeConte 231 Oreocharq Casey 104 Oxyomus Stephens 148 Oreodytes Seidlitz 67 Oxypoda Mannerheim 99 Oreoxenus Tschitch6rine 46 Oxypodini 99 Orestia Germar 318 Oxyporinae 109 Orestioides Hatch 318 Oxyporus Fabricius 109 Ormiscini 325 Oxypselaphus Chaudoir 52 Ormiscus Waterhouse 325 Oxypteris Kirby 165 Orobanus LeConte 90 Oxytelinae 92 Orochares Kraatz 91 Oxytelini 94 Orophiinae 245 Oxytelus Gravenhorst 95 Orophiini 245 Oropus Casey 126 Pachybrachini 305 Orphelophorus d'Orchymont 131 Pachybrachys Chevrolat 305 Orphilini 199 Pachybruchus Pic 300 Orphilus Erichson 199 Pachydela Rivalier 8 Orsodacna Dejean 303 Pachydeniini 153 Orsodacne Latreille 303 Pachylobius LeConte 338 Orsodacninae 303 Pachyphanes Dietz 340 4t9 Pachyrhjnus Schcinherr SB0 Paraplinthus Faust 338 Pachy rhSt nchus l{trby 325 Paraptochus Seidlitz 332 Pachysctrelini 168 Paraquedius Casey 122 Pachyschelus Solier 168 Parargutor Casey 32 Pachystilicus Casey 114 Parascydmus Casey 83 Pachystylus Rey 305 Parastenus Heyden 110 Pachyba Dejean 288 Paratachys Casey 29 Pachytychius LeConte 340 Paratenetus Spinola 262 Pactopus LeConte 186 Paratrogophloeus Hatch 94 Poederillus Casey 116 Paratropa Vacher de Lapouge 10 Paederina 116 Paratychius Casey 346 Paederinae 113 Parazonitis Enns 265 Paederini 113 Parelaphidion Skiles 281 Paederus Fabricius 116 Parenthis Dietz 350 Palaminus Erichson 113 Paria LeConte 308 Pallodes Erichson 216 Parilendus Casey 242 Palorus Mulsant 258 Parmenini 294 Palporus Campbell 97 Parmulus Harold 228 Panagaeini 49 ParocaleaBernhauer 105 Panagaeus Latreille 49 Paromalini 138 PanalotaCasey 101 Paromalus Erichson 138 Pandeletr:ius Schtinherr 331 Paropsimeno Motschulsky 311 Panscopi,l,ius Pierce 333 Parothius Casey 123 Panscopu.s Schiinherr 333, 334 Pasimachus Bonelli 17 Parabracteon Notman 20 Passalidae 145 Paracercrron Seidlitz 134 Passalotaenlus Kuwert 145 Paraclivi:ra Kult 18 Passandrinae 2I9 Parac5rmus Thomson 132 Patrobini 29 Paradelitrthrum Hatch gt Patroboidea Van Dyke 30 Paradicaelus Ball 48 Patrobus Dejean 29 Paradilacra Bernhauer 102 Pauronemognatha Enns 265 Parafilum,is Casey 207 Paussini 19 Paragonu.m Casey 53 Pectotoma Hatch 276 Paralarica Breuning 295 Pedetes Kirby 182 ParalathraCasey 116 Pediacus Shuckard 219 Paralesteua Casey g0 Pedilldae 274 Paraliaphlus Wallis 60 Pedilinae 274 Paralister Bickhardt 139 Pedilophorini 160 Paralitargus Casey 242 Pedilus Fischer von Waldheim 274 Parollelin.a Casey 291 Peditatus Pacheco 170 Parallelostethus Schwarz 181 Pelates }lorn 77 Paralopha Casey 27 Pelatines Cockerell 77 Paralytta Selander 264 Pelecomalium Casey 90 ParqmedonCasey 114 Pelecotoma Fischer von Waldheim 250 Parametriona Spaeth 323 Pelecotominae 250 ParamysiaReitter 235 Pelec;ryhorus Solier 254 Paranaemia Casey 233 Pelenomus Thomson 351 Paranchus Lindroth 52 Pelidnota Macleay 155 Parandra Latreille 277 Pelidnotidia Casey 155 Parandrinae 277 Pelophila Dejean 13 Parandrini 277 Peltastica Mannerheim 196 Paranomctla Casey 154 Peltasticinae 196 Pq,ranomus Kiesenwetter 181 Peltideo Motschulsky 207 Paranove].sis Casey 197 Peltinae 207 Paranthonomus Dietz 343 Peltini 207 Paraopsiraus Champlain & Knull 280 Peltodytes R6gimbart 61 Parapach;yto Casey 288 Pemphus Motschulsky 13 Parapanscopus Buchanan 334 Penetini 253 420 Pentacladocera Pierce 366 Philodes LeConte 4? Pentagonica Schmidt-Goebel 50 Philonthini 117 Pentagonicini 50 Philonthus Curtis 117 Pentagrammaphila Pierce 366 Philopedon Schiinherr 330 Pentanota Bernhauer 100 Philophuga Motschulsky 58 Pentaphyllus Dejean 256 Philopuntia Pierce 347 Pentaria Mulsant 276 Philostrq.tus Zimmerman 338 Pentariini 276 Philothermus Attb6 227 Pentarthrum Wollaston 355 PhiloxylonLeConte 202 Pentaxocerq. Pierce 366 Phlaeopterus Motschulsky 90 Pentazocera Pierce 366 Phloeocharinae 95 Penthe Newman 243 Phloeonaeus Erichson 93 PenthelispaPascoe 252 Phloeonomus Heer 88 Penthinae 243 Phloeophagus Schtinherr 355 Pentodontini 156 Phloeopora Erichson 105 Pentozoe Pierce 366 Phloeopterus LeConte 90 Peranus Lewis 139 Phloeosinini 358 Perapion Wagner 328 Phloeosinus Chapuis 358 Percosia Zimmermann 36 Phloeostiba Thomson 88 Pergetus Casey 274 Phloeotribini 358 Perigaster Dietz 349 Phloeotribus Latreille 358 Perigona Castelnau 51 Phloeotrya Stephens 248 Perigonini 51 Phlogistosternus Wolcott 210 Perimecus Stephens 184 Phobetus LeConte 153 Perimegatoma Horn 198 Phomalus Casey 275 Peristethus LeConte 33 Phosphaenus Castelnau 191 Peritelini 332 Photinini 190 Peritelinus Casey 332 Photinus Castelnau 190 PerthalycraHorn 216 Photurinae 191 Peryphanes Jeannel 24 Photuris Dejean 191 Peryphodes Casey 25 Phratora Chevrolat 313 Peryphus Stephens 24 Phrenapatinae 253 Petaliini 204 Phryganophilus Sahlberg 247 Petalium LeConte 204 Phrypeus Casey 28 Pezocrosita Jacobson 312 Phthora Mulsant 253 Phacophallus Coiffait 123 Phycocoetes LeConte 337 Phaedon Megerle von Miihlfeld 312 Phyla Motschulsky 21 Phaenocerus Bonvouloir 187 Phfi.etes Redtenbacher 255 Phaenog'yra Seevers 107 Phylethus Redtenbacher 255 Phaenonotum Sharp 133 PhyIIis Gistel 310 Phaenops Dejean 165 Phyllobaenini 208 Phalacridae 226 Phyllobaenus Dejean 208 Phalacrini 226 Phyllobaenus Spinola 210 Phalacrus Paykull 226 Phyllobiini 332 Phaleriini 254 Phyllobius Germar 332 Phaleromela Reitter 254 Phyllobrotica Chewolat 316 Pharalus Casey 40 Phyllodecta Kirby 313 Phausis LeConte 191 Phyllodrepa Thomson 88 Phelister Marseul 140 Phyllophaga Harris 152 Phellidius LeConte 255 Phyllotreta Chevrolat 316 Phellopsis LeConte 252 Phyllotrox Schtinherr 341 Pheloconus Roelofs 347 Phymaphora Newman 238 Phengodes Illiger 190 Phymatinus LeConte 334 Phengodidae 190 Phymatodes Mulsant 283 Phenolia Erichson 216 Phymatodina Casey 283 Phidola Thomson 295 Physocnemum Haldeman 284 Philq.s Portevin 85 Physonota Boheman 322 Philhygra Mulsant & Rey 102 Physonotini 322 427 Physorhini 184 Platypodidae 365 Phytobiini 350 Platypodinae 365 Phytobius Schonherr 350 Platypodini 365 PhytodectaKirby 313 Platypsyllus Ritsema 82 Phytoecini 299 Platypsyllus Westwood 82 PhytonNewman 281 Platypus Herbst 365 Phytonomu,s Schiinherr 336 Platysoma Leach 140 Phytosini 107 Platysomatini 140 Phyxelis Schtinherr 333 Platystethus Mannerheim 94 Piazorhinini 345 Platytrachelus Motschulsky 21 Piazorhinus Schiinherr 345 Platyusa Casey 106 Pidonia Mulsant 292 Plectographo Rivalier 10 Piestinae 92 Plectralidus Casey 40 Piestini 92 Plectrura Mannerheim 294 PilemaLeConte 282 Plegaderini 136 Pilopius Casey 129 Plegaderus Erichson 136 Pinacodera Schaum 59 Plesiocis Casey 245 Pinodytes Florn 80 Pleurophoro Chewolat 304 Pinophilini 113 Pleurophorus Mulsant 149 Piosoma LeConte 40 Pleuropompha LeConte 264 Pisenus Casey 243 Pleurostica Motschulsky 312 Pissodes G'ermar 338 Pleurotobia Casey 107 Pissodinae 338 Plinthini 338 Pityobini 176 Plinthodes LeConte 333 Pityobius l,eConte 176 Plocamus LeConte 353 Pityogenes,Bedel 360 Plocetes LeConte 346 Pityokteines Fuchs 360 Pocadius Erichson 216 Pityomacer Kuschel 324 Podabrini 192 Pityophagrrs Shuckard 217 Podabrus Westwood 192 Pityophthorus Eichhoff 363 Podapion Riley 329 Placonotusr Macleay 219 Poecilium Fairmaire 283 Placopterus Wolcott 209 Poecilobrium Horn 282 Placusa Erichson 107 Poecilochroo Chevrolat 209 Plagiodera Chevrolat 312 Poecilus Bonelli 30 Plagiometriona Spaeth 323 Pogonocherini 295 Planesus Motschulsky 59 Pogonocherus Dejean 296 Plataphodes Ganglbauer 22 Polemius LeOonte 194 Plataphus Motschulsky 22 Poliaenus Bates 295 Platerodini 188 Polistoma Casey 105 Plateros Bourgeois 189 Pollaclasis Newman 191 Plateumarini 302 Polycaon Castelnau 199 Plateumaris Thomson 302 Polycaonini 199 Platidiolus Chaudoir 30 Polycestinae 161 Platidius Chaudoir 30 Polycestini 161 Platybregmus Fisher 203 Polyclasis LeConte 191 Platycerini 144 Polyderis Motschulsky 29 Platyceropsis Benesh 144 Polydrosus Schdnherr 330 Platycerus Geoffroy 144 Polydrosini 330 Platycis Thomson 189 Polydrusus Germar 330 Platydema Castelnau & Brull6 255 Polygrammo Chewolat 311 Platydracus Thomson 120 Polygraphini 359 Platylomaius Cooman 138 Polygraphus Erichson 359 Platymedon Casey 114 Polyphaga 74 Platynidius Casey 56 Polyphylla Harris 152 Platynini 51 PolyspilaHope 310 Platynomicrus Casey 53 Polystoma Stephens 105 Platynus Bonelli 55 PomphopoeaLeConte 264 Platypatrobus Darlington 29 Pontomalota Casey 103 422 Poophagus Schiinherr 350 Psammoporus Thomson 147 Popillia Audinet-Serville 155 Pselaphidae L25 Porcinolus Mulsant l3z Rey 159 Pselaphinae t29 Poreospasta Horn 2134 Pselaphini 129 Porotachys Netolitzky 29 Pselaphorhynchites Schilsky 327 Porrhodites Kraatz',91 Pselaphus Herbst 129 Potamodytes Zimmermann 66 Psenocerus LeConte 295 Potamonectes Zimmermann 66 Psephenidae 174 Prasocuris Latreille 312 Psepheninae 174 Priacma LeOonte 6 Psephenus Haldeman 174 Priacmini 6 Psephus Kirby 149 Priobium Motschulsl 423 Pseudota Casey 101 Pycnoplectus Casey 126 Pseudothysanoes Blackman 359 Pycnorus Casey 114 Pseudotrtaeno Minsk & Hatch 39 Pycnotomina Casey 218 Pseudotrox Robinson 150 Pygoleptura Linsley & Chemsak 291 Pseudotychlus Blatchley 328 PyladiaWeise 318 Pseudousipalia Lohse 102 Pyractomena Melsheimer 190 Pseudow eisea Schwarz 229 Pyrectosomo Motschulsky 191 Pseudoxenos Saunders 367 Pyres Champion 258 Pseudoxyletinus Pic 203 Pyrilloxenos Pierce 366 Pseudoxyporus Nakane & Sawada 109 Pyrochroidae 269 Psiloscelis Marseul 140 Pyrochroinae 269 Psoa Herbst 200 Pyrophorinae 175 Psoinae 199 Pyrophorini 175 Psoini 2(t0 Pyropyga Motschulsky 191 Psomophrrs Casey 83 Pytotrichus LeConte 288 Psomus Clasey 348 Pyrrhalta Joannis 314 Psydrus LeConte 29 Pyrrocis Castelneau 243 Psylliodes Berthold 321 Py.thidae 268 Psyllobora Dejean 237 Pybhinae 268 Psylloborini 237 Py'tho Latreille 268 Psyrassa Pascoe 280 Pytna Casey 129 Ptenidium Erichson 75 Pterna Seidlitz 260 Quadrifrons Blatchley 216 Pterocolirrae 326 Quediellus Casey L22 Pterocolus Say 326 Quediini 120 Pteroloma Gyllenhal 77 Quediochrus Casey 121 Pteropalus Casey 43 Quedionuchus Sharp 121 Pterostichini 30 Quedius Stephens 121 Pterostichus Bonelli 31 Pteryngium Reitter 221 Rafonus Casey 126 Ptiliidae 75 Ramecia Casey 126 Ptiliinae 75 RamonaCasey 113 Ptilininae 203 Rantogiton des Gozis 70 Ptilinini 203 Raphirus Stephens 122 Ptilinus I\fiiller 203 Redistes Casey 252 Ptiliola Haldeman 75 Reesa Beal 198 Ptiliolum Flach 75 Refonia Casey 32 Ptiliopycra Casey 76 Reichenbachia Leach 128 Ptilium Gyllenhal 75 Rembus Dejean 48 Ptilodact5'la Illiger 169 Reminius Casey 267 Ptilodacty'lidae 169 Remus Holme 119 Ptinidae 205 Renardia Motschulsky 92 Ptininae 205 Renocis Casey 358 Ptinodes LeConte 202 Reuelstohea Hatch 90 Ptinus Linn6 206 Rhabdophorus Leftvre 308 Ptomaphagini 81 Rhabdopterus Lefbvre 308 Ptomaphagus Illiger 81 Rhadalinae 212 Ptomister Houlbert & Monnot 139 Rhadine LeConte 52 Pubimodes Park I27 Rhaeboscelis LeConte 167 Pullus Mulsant 230 Rhagium Fabricius 289 PunctagonumCasey 53 Rhagomicrus Fleutiaux 187 Purpuricenini 286 RhagonychaEschscholtz 193 Purpuricenus Dejean 286 Rhanidea Strohecker 238 Pycnoglyprta Thomson 88 Rhanis LeConte 238 Pycnomerini 252 Rhantus Dejean 70 Pycnomerus Erichson 252 Rhaphiona Fischer von Waldheim 15 Pycnoruotida Casey 254 Rhaxonycha Motschulsky 193 Pycnophus Casey 83 Rhinocyllini 337 424 Rhinocyllus Germar 337 Salpinginae 270 Rhinoncus Schiinherr 351 Salpingus auct. 270 Rhinosimus Latreille 270 Sandalus Knoch 143 Rhipiceridae 143 Sanfilippody'tes Franciscolo 65 Rhipidandrini 254 Saperda Fabricius 298 Rhipidandrus LeConte 254 Saperdini 298 Rhipiphoridae 250 Sapintus Casey 272 Rhipiphorino Casey 157 Saprininae 136 Rhipiphorinae 250 Saprinus Erichson 136 Rhipiphorus Bosc dAntic 250 Saprobia Ganglbauer 216 Rhizophagidae 218 Saprophilus Streubel 120 Rhizophaginae 218 Sarosesthes Thomson 285 Rhizophagus Herbst 2l-8 Sarpedon Bonvouloir 187 Rhodobaenus LeConte 354 Saxinis Lacordaire 305 Rhombodera Reiche 50 Scalopterus Motschulsky 2\2 Rhombonalia Casey 154 Scaphidema Redtenbacher 254 Rhopalapion Schilsky 329 Scaphidiidae I24 Rhopalelophorus Kuwert 130 Scaphidiini 125 Rhopalomerus Chewol at 285 Scaphidium Olivier 125 Rhopalopaclzys Chevrolat 285 Scaphinotus Dejean 12 Rhynchaeninae 345 Scaphisoma Leach 125 Rhynchaenus Clairville 345 Scaphisomini 125 Rhynchitidae 326 Scaphium l{trby I25 Rhynchitinae 326 Scarabaeidae 145 Rhynchitini 326 Scarabaeinae 146 Rhynchophorinae 354 Scarabaeini 146 Rhyncolini 355 Scarabaeoidea 143 Rhyncolus Germar 356 Scarites Fabricius 17 Rhysodidae 7 Scaritini 17 Rhysothorax Bedel 147 Scelolyperus Crotch 316 Rhyssemus Mulsant 149 Schedarosus Reitter 255 Rhyssomatus Schijnheru 347 Schistogloss a I{r aatz lO2 Rhytirrhininae 334 Schizogenius Putzeys 18 Rhyzobius Stephens 233 Schizophilus Bonvouloir 188 Rhyzopertha Stephens 200 Schizotus Newman 269 RileyaOlivier 191 Sciaphilus Schiinherr 330 Riponus Casey 278 Scierus LeConte 356 RomaleumWhite 280 SciodrepaThomson 81 Ropalopus Mulsant 284 Sciodrepoides Hatch 81 Rosalia Audinet-Serville 282 Sciopithes Horn 331 Rosaliini 282 Scirtes Illiger 142 Rosella Whitehead 346 Scirtidae 142 Roualida Casey 103 Scleropterini 350 Rugilus Leach 114 Scobicia Lesne 200 Rushia Forel 247 Scoliocerus Motschulsky 178 Rutelini 155 Scolytidae 356 Rutelinae 154 Scolytinae 359 Rutidosoma Stephens 350 Scolytini 359 Rybaxis Saulcy 128 Scolytus Geoffroy 359 Rypobiini 228 Scopaeina 114 Scopaeoma Casey 114 Sachalinobia Jacobson 289 Scopaeopsis Casey 114 Saciinae 228 Scopaeus Erichson 114 Saciini 228 Scopophus Casey 83 Sacium LeConte 228 Scotias Czenpinski 205 Sa.codes LeConte 142 Scotobates Rye 258 Salebius Casey 22I Scotochroa LeConte 247 Salpingidae 270 Scotochroides Mank 247 425 Scotocryptini 79 Sitophilini 355 Scraptia Latreille 276 Sitophilus Schiinherr 355 Scraptiidae 275 Smicronychini 340 Scraptiinae 276 Smicronyx Schdnherr 340 Scraptiini 276 SmiliaWeise 229 Sculptotheca Schilsky 204 Smodicini 280 Scydmaenidae 82 Smodicum Haldeman 280 Scydmaeninae 82 Sonoma Casey 126 Scydmaenrni 83 Sonornota Casey 102 Scydmaenr-rs Latreille 83 Soronia Erichson 216 Scymninaer 229 Spectralia Casey 162 Scymnini 229 Sperchopsis LeConte 132 Scymnobius Casey 231 Sphaeracra Say 56 Scymnus Kugelann 230 Sphaericus Wollaston 205 Scythropus Schtlnherr 330 Sphaeridiinae 133 Selenalius Casey 43 Sphaeridiini 133 Selenophorus Dejean 43 Sphaeridium Fabricius 133 Semanotus Mulsant 283 Sphaeriestes Stephens 270 Semicampa Netolitzky 27 Sphaerites Duftschmid 135 Semiclivina Kult 18 Sphaeritidae 135 Semijulistus Schilsky 212 Sphaeroderus Dejean 12 Sepedophilus Gistel 97 Sphaeromela Bedel 311 Serangiini 229 Sphalera LeConte 248 Serica Ma<:Leay 151 Sphalma Horn 268 Sericini lti1 Sphenophorini 354 Sericoda Kirby 52 Sphenophorus Schiinherr 354 Sericoderinae 228 Sphindidae 213 Sericoderini 228 Sphindinae 213 Sericoderus Stephens 228 Sphindini 213 Sericosomus Dejean 182 Sphindus Chewolat 213 Sericus Eschscholtz 182 Spilodiscus Lewis 140 Serraticoll;i.s White 318 Spilosota Casey 155 Serropalpini 247 SpilotaBurmeister 154 Serropalpus Hellenius 248 Spilotus LeConte 247 Sexarthrus Blatchley 343 Spintherophyta Dejean 308 Siagonium Kirby & Spence 92 Spondylinae 279 Siberia Blzrckwelder 95 Spondylis Fabricius 279 Sibinia Germar 346 Spongoloba Chaudoir 58 SibiotaCasey 101 Spongopus LeConte 45 Sicinus Champion 255 Sponidium Casey 262 Silaria Mulsant 276 Stagetus Wollaston 204 Silini 194 Stamoderes Casey 330 Silis Chartrrentier 194 Staphygraphis Horn 90 Silphidae 84' Staphylinidae 86 Silphinae 85 Staphylininae l1-7 Silusida Casey 107 Staphylinini 120 Silvaninae 220 Staphylinoidea 74 Silvanini 220 Staphylinus Linn6 120 Siluanophloeus Sharp 219 Statira Audinet-Serille 262 Silvanus Latreille 220 Statirinae 262 Simplocaria Stephens 159 SteatoderusDejean 181 Simplocariini 159 Stegobiini 202 Sinistrocedius Park 129 Stegobium Motschulsky 202 Sinodendrini 144 Stelidota Erichson 215 Sinodendron Hellwig 144 StenagriaSharp 105 Sirocalodes Voss 350 Stenelmis Dufour 173 Sitona Germar 331 Stenichnini 83 Sitonini 3ll1 Stenichnus Thomson 83 426 Steninae 109 Stomis Clairville 30 Stenister Reichardt 139 Stornphax Fischer von Waldheim 149 Stenistoderus Jacquelin du Val 123 Storualota Mannerheim 101 Stenocantharus Gistel 13 Storthephora Miiklin 262 Stenocellus Casey 46 Strabala Chewolat 317 StenochiaKirby 261 Strangalepta Casey 293 Stenocorus Geoffroy 289 Strangalia Audinet-Serville 290 Stenocorus Fabricius 288 Strangalina Aurivillius 290 Stenocranophilus Pierce 367 Strepsiptera 366 Stenocrepis Chaudoir 49 Stricticomus Pic 273 Stenolophus Stephens 47 Strigocis Dury 245 Stenopodius Horn 322 Strigoderma Burmeister 155 Stenopterini 282 Strigodermella Casey L55 Stenoscelis Wollaston 355 Strongylaspis Spaeth 323 Stenosphenus Haldeman 280 Stronryliini 261 Stenostrophia Casey 292 Strongylium Kirby 261 Stenotarsinae 238 Strongylocassis Hincks 323 Stenotarsini 238 Strophiona Casey 294 StenothoraxHarris 154 Strophogastra Fenyes 103 Stenotrachelinae 263 Strophosoma Billberg 330 Stenotrachelus Latreille 263 Strophosomini 330 Stenous Chaudoir 49 Stylopidae 367 Stenura Dejean 292 Stylopidia 366 Stenuris Kirby 162 Stylopinae 367 Stenus Latreille 109, 110 Stylops Kirby 367 Stephanocleonus Motschulsky 336 Subhaida Hatch 89 Stephanoderes Eichhoff 362 Sulcacis Dury 245 Stephanopachys Waterhouse 200 Sumitrosis Butte 321 Stephostethus LeConte 240 Sunius Curtis 113 Stereagonum Casey 55 Suphisellus Crotch 62 Steremnius Schtinherr 338 Symbiotes Redtenbacher 238 Stereocerus Kirby 31 Symphora LeConte 247 Stereopalpus LaFert6-S inectdre 27 4 Synaphaeta Thomson 295 Steriphanus Casey 253 Syncalosomo Breuning 10 Sternidius LeConte 297 Syncalyptinae 158 Sternuchus LeConte 141 Synchita Hellwig 251 Steroderus Motschulsky 32 Synchitini 251 Steropus Dejean 32 Synchroa Newman 267 Stethobaris LeConte 353 Synchroidae 267 Stethon LeConte 187 Synchytodes Crotch 251 Stethorhanis Blaisdell 238 Syndesinae 144 Stethorini 229 SynerthaDietz 340 Stethorus Weise 229 Syneta Dejean 307 Sthereus Motschulsky 338 Synetinae 307 Sticholotidinae 229 Synolabus Jekel 326 Stichosoma Motschulsky 311 Synstrophus Seidlitz 246 Stictalia Casey 107 Syntomium Curtis 93 Stictanchus Casey 55 Syntomus Hope 58 Stictocranius LeConte 112 Synuchus Gyllenhal 51 Stictoleptura Casey 293 Systena Chewolat 317 Stictolinus Casey 124 Stigrnatoma LeConte 260 Tachinoderus Motschulsky 96 Stilbolemrna Harris 151 Tachinus Gravenhorst 96 Stilbolidus Casey 45 Tachistodes Casey 48 Stilbus Seidlitz 226 Tachyerges Schiinherr 345 Stilicina 114 Tachymenis Motschulsky 29 Stilicosoma Casey 114 Tachyporinae 96 Stolaini 322 Tachyporini 96 427 Tachyporu.s Gravenhorst 97 Tetrascapha Schuster & Marsh 126 Tachypterellus FaII & Cockerell 343 Tetratoma Fabricius 243 Tachypteru.s Dietz 343 Tetratomidae 243 Tachypus Dejean 20 Tetratominae 243 Tachys Stephens 29 Tetropium Kirby 279 Tachyta Kirby 28 Teuales Casey 90 Tachyura Motschulsky 28 Texania Casey 161 Taenioglyptes Bedel 362 Thalasselephas Egorov & Karotyaev 337 Taenobqtes Motschulsky 260 Thalycra Erichson 216 Tanaocarabus Reitter 12 Thanasimus Latreille 209 Tanaops LeConte 212 Thanatophilus Leach 85 Tanarthropsis Casey 273 Thaneroclerini 209 Tanarthrus LeConte 273 Thecesterninae 335 Tanilotes Casey 275 Thecesternus Say 335 Tanygaster Blatchley 316 Thermonectus Dejean 72 Tanymecirri 331 Thersalus Pascoe 280 Tanymecus Germar 331 Thes Semenov 239 Tanyrhinus Mannerheim 91 Thesalia Casey 292 Tanysphyrini 340 Thesium Casey 126 Tanysphyrus Germar 340 Thicanus Casey 213 Taphranch.us Casey 53 Thinobius Kiesenwetter 94 Taphrocerus Solier 168 Thinodromus Kraatz 94 Taphroscydmus Casey 83 Thinopinus LeConte 120 Tapinosthenes Kolbe 10 Thinusa Casey 108 Taraxis LeConte 304 Thoracophorus Motschulsky 92 Tarpela Bates 258 Thricolepoides O'Brien 330 Tarphiota Casey 103 Thricomigus Horn 333 Tarsosteninae 2ll Thrimolus Casey 243 Tarsostenrrs Spinola 211 Throscidae 186 Taxicerellc,. Casey 100 Throscinae 186 Tecnophilus Chaudoir 59 Throscus Latreille 186 Tectosternum Balfour-Browne 134 Thyamis Stephens 317 Tedinus Crasey 259 Thylodrias Motschulsky 199 Teinodactylo Chevrolat 317 Thylodriini 199 Telacis Poey 259 Thymalus Latreille 207 Telephanus Erichson 220 Thy sanocnemis LeConte 345 Telephorus Schaeffer 193 Tilargus Casey 243 Telmatophilinae 22L TileaFauvel 9O Telmatophilus Heer 221 Tillini 208 TemnochilaWestwood 207 Tilloclytus Bates 286 Tenebrio L,inn6 259 Tillomorphini 286 Tenebrioni.dae 253 Timarcha Latreille 310 Tenebrioninae 258 Timarchini 310 Tenebrionini 258 Tipnus Thomson 205 Tenebrionoidea 242 Tmesiphorini 129 Tenebroides Piller & Mitterpacher 208 Tmesiphorus LeConte 129 Tenomergil Neboiss 6 Tomq.rus LeConte 224 Tentyriinae 253 Tomicini 357 Teredini 252 Tomoxia Costa 248 Teretriini 136 Toraminae 224 Teretrius Erichson 136 Toramim 224 Tesnus Rey 112 Toramus Grouvelle 224 Tetartoperrs Czwalina 115 Torrentomus Bierig 94 Tetragonorlerus Dejean 56 Tosastes Sharp 334 Tetraleucus Casey 52 Toxidium LeConte 125 Tetralina (lasey 100 Toxotopsis Casey 288 Tetraopes Schtjnherr 299 Toxotropis LeConte 325 Tetraopini 299 Toxotus Dejean 288 428 Trachelonepha Casey 22 Trigonodemus LeConte 91 Trachodes Germar 338 Trigonogenius Solier 205 Trachyinae 168 Trigonorhinus Wollaston 324 Trachykele Marseul 163 Trigonoscuta Motschulsky 330 Trachypachidae 7 Trigonurini 92 Trachypachus Motschulsky 7 Trigonurus Mulsant 92 Trachyphloeini 332 Triliarthrus Casey 46 Trachyphloeus Germar 332 Trimioplectus Brendel 127 Trachysectus Casey 113 Triplacinae 225 Trachysida Casey 292 Triplax Herbst 225 Tragidion Audinet-Serville 286 Triplectrus LeConte 44 Tragosoma Audinet-Serville 278 Trirhabda LeConte 314 Tragosomini 278 Tritoma Fabricius 225 Trechicus LeConte 51 Trixagus Kugelann 186 Trechini 19 Trogastrini 126 Trechonepha Casey 21 Troginae 150 Trechus Clairville 19 Trogoderma Dejean 197 Treiodous Dugds 263 Trogophloeus Nannerheim 94 Trepanedoris Netolitzky 27 Trogossita Olivier 208 Triachus LeConte 307 Trogossitidae 206 Triaena LeConte 39 Trogossitinae 207 Triarthron Markel 78 Trogossitini 207 Tribalinae 138 Trogoxylon LeConte 201 Triboliini 258 Tropicini 170 Tribolium Macleay 258 Tropicus Pacheco 170 Tribonia Rivalier 8 Tropideres Sch0nherr 325 Tribrachys LeConte 2I4 Tropiderini 325 Trichalophus LeConte 333 Tropiphorus Schiinherr 334 Trichapion Wagner 329 Tropisternus Solier 133 Trichelaphrus Semenov 16 TrotomrnaLeConte 247 Trichesthes Erichson 152 Trox Fabricius 150 Trichiinae 157 Trymochthebius Kuweft 7 4 Trichiini 158 Trypherus LeConte 195 Trichiotinus Casey 158 Trypodendron Stephens 361 Trichischius LeConte 354 Trypophloeus Fairmaire 362 Trichiusa Casey 103 Turboscymnus Gordon 231 Trichobaris LeConte 352 Tychiinae 345 Trichocanthus Motschulsky 120 Tychiini 346 Trichocellus Ganglbauer 46 Tychini 129 Trichochrous Motschulsky 2I3 Tychius Germar 346 Trichocnemis LeConte 27 7 Tychobythinus Ganglbauer l-28 Trichodes Herbst 210 Tylicini 160 Trichodesma LeConte 202 Tylicus Casey 160 Tricholochmaea Laboissidre 314 Tylistus LeConte 204 Trichophya Mannerheim 99 Tyloderma Say 347 Trichophyinae 99 Tylonotus Haldeman 280 Trichoplataphus Netolitzky 23 Tympanophorus Nordmann 120 Trichopteryx Kirby & Spence 76 Typhaea Stephens 243 Trichoscelis Bates 286 Typhloleiodes Hatch 80 Trichotichnus Morawitz 43 Typitium Casey 262 Tricoryninae 205 Typocerus LeConte 291 Tricorynini 205 Tyrini 129 Tricorynus Waterhouse 205 TyrusAub6 129 Tricrania LeConte 265 Tricraniodes Wellman 265 Uleiota Latreille 220 TrigaFauvel 95 Uleiotini 220 Trigites Handlirsch 95 Ulitusa Casey 107 Trigonarthris Haldeman 293 Ulochaetes LeConte 288 429 Uloma Ltrtreille 257 Xyleborinus Reitter 362 Ulomini 257 Xyleborus Eichhoff 362 Unamis Casey 90 Xylechinus Chapuis 357 Upis Fabricius 261 Xyletininae 203 Urgleptes Dillon 298 Xyletinini 203 Urograph',is Horn 297 Xyletinus Latreille 204 Uroplatini 322 Xylita Paykull 247 Utobium FalI 202 Xylobiops Casey 200 LJvarus Guignot 64 Xylobius Latreille 188 Xylocomesus Thatcher 356 Vacusus Casey 273 Xylocrius LeConte 284 Vanonus Casey 275 Xylodromus Heer 91 Variopalpis Solier 58 Xyloecus Dejean 188 Vellica Casey 90 Xyloperthini 200 Venusa Casey 107 Xylophilus Mannerheim 188 Veraphis Casey 83 Xylopinus LeOonte 260 Vicelva l\:[oore & Legner 95 Xyloryctes Hope 156 Vincenze.llus Reitter 270 Xylosandrus Reitter 361 Vrilletta LeConte 203 Xyloterini 361 Xyloterinus Swaine 361 WillinkiaBondar 353 Xylotrechus Chevrolat 285 Xanthochroa Schmidt 266 Yrdops Steffan 171 Xanthochroina Ganglbauer 266 Xanthoga.leruca Laboissidre 314 Zabrini 35 Xantholirrinae 123 Zacotus LeConte 19 Xantholinini 123 ZaitzeviaChampion 173 Xantholirrus Dejean 124 ZalepiaArnett 175 Xanthonia Baly 309 Zalobius LeConte 95 Xanthoph,ius Motschulsky 123 Zeadolopus Broun 79 Xanthopl'gini 120 Zenodosus Wolcott 209 Xanthus Dietz 344 Zeugophora Kunze 304 Xeninae 367 Zeugophorinae 304 Xenochara Mulsant & Rey 104 ZezeaCsiki 39 Xenocharis Bierig 113 ZiloraMulsant 247 Xenodacne Boyle 225 ZonantesCasey 275 Xenodusa Wasmann 106 Zonitina 265 Xenoscelinae 224 Zonltinae 265 Xenota Mulsant & Rey 103 Zonitini 265 Xerosaprinus Wenzel 137 Zonitis Fabricius 265 XestipygerMarseul 138 Zonoptilus Solsky 93 Xestobiini 202 Zopheridae 252 Xestobiurn Motschulsky 202 Zophefinae 252 Xestocis Casey 244 Zuphiini 59 Xestoleptura Casey 293 Zuphium Latreille 59 Xestolinus Casey 124 Zygogramma Chewolat 3 10 Xestonotrrs LeConte 45 Zygopinae 348 Xestophu;s Casey 83 Zygopini 348 Xyleborini 361 ZyrasStephens 106 Recycled ,*r1. faper ,pn *""" GP 430 Canad'd