VENUS 71 (1–2): 1–11, 2013 ©Ma lacological Society of Japan

Description of Two New Species of Trophoninae s.l. and (: ) from New Caledonia and Comments on Litozamia Iredale, 1929 and Jousseaume, 1880

Roland Houart* Institut royal des Sciences naturelles de Belgique, rue Vautier, 29, B-1000 Bruxelles, Belgium

Abstract: Litozamia acares n. sp. and Siphonochelus (Trubatsa) wolffi n. sp. are described from New Caledonia. The radula and the operculum of Litozamia acares are illustrated and described. The classification of Litozamia in Trophoninae is maintained awaiting molecular data to either confirm or modify this decision. Litozamia longior (Verco, 1909) is reinstated as a valid species. The use of the subgenus Choreotyphis Iredale, 1936 is reinstated in Siphonochelus for a single species from eastern Australia, based on differences in shell morphology.

Keywords: West Pacific, New Caledonia, Trophoninae s.l., Typhinae, new data, new species

Introduction

A large number of oceanographic expeditions conducted by MNHN and ORSTOM (later IRD) were organized starting in the early 1980s to sample the malacological fauna of the West and South Pacific. The richness of the malacofauna of the New Caledonia region, particularly from deep water is without equivalent in other areas of the Indo-Pacific (Bouchet et al., 2008). No less than 83 new species of Muricidae have been described in several papers since 1983, that means about 38% of the total number of named Muricidae (excluding Coralliophilinae) occur in New Caledonia. To date, three major revisions have been published to review the subfamilies Typhinae (Houart, 1991), Ergalataxinae (Houart, 1995a) and Trophoninae (Houart, 1995b). The two species described herein were collected around the island during 11 expeditions between 1978–79 and 1994. All specimens are deposited in the MNHN unless otherwise specified.

Repositories: JW, collection of John Wolff, Lancaster, Pennsylvania, USA; MNHN, Muséum national d’Histoire naturelle, Paris, France; RH, collection of Roland Houart.

Abbreviations: CC, Chalut à crevettes (Shrimp trawl); CP, Chalut à perche (Beam trawl); dd, empty shell(s); DE, Drague Epibenthique (Epibenthic dredge); DR, Drague à roches (rock dredge); DW, Drague Warén (Warén dredge); IRD, Institut de recherche pour le développement; lv, live collected specimen(s); ORSTOM, Office de la recherche scientifique et technique outre- mer.

* Corresponding author: [email protected] 2 R. Houart

Fig. 1. Spiral cords and apertural denticle morphology of Litozamia acares n. sp.

Terminology used to describe the spiral cords (based on Merle, 1999 and 2001) (Fig. 1) (Terminology in parentheses: erratic feature): P, primary cord; IP, infrasutural primary cord (primary cord on subsutural ramp); P1, shoulder cord; P2–P6, primary cords of the convex part of the teleoconch whorl; ADP, adapertural primary cord on the siphonal canal; D1–D5, abapical denticles in aperture.

Systematic Account

Faumily M ricidae Rafinesque, 1815 Subfamily Trophoninae s.l. Cossmann, 1903 Litozamia Iredale, 1929 Type species by original designation: Litozamia rudolphi (Brazier, 1895), Recent, New South Wales, Australia.

Remarks: The subfamily Trophoninae is currently known to be polyphyletic (Barco et al., 2010). To my knowledge, this is the first time that the radula and the operculum of Litozamia are illustrated. Both are typical muricine, but the subfamilial classification of the genus remains doubtful. It is currently classified in Trophoninae in a conservative way (Houart, 2010) as cited by Radwin & D’Attilio (1976: 183), awaiting future study of the molecular characters when more adequate material will become available.

Litozamia acares n. sp. (Figs. 1, 3–5, 29, 31–33)

Type material: Holotype, MNHN 25086; 7 paratypes,N M HN 25088; 1 paratype, RH. New Species of Trophoninae and Typhinae 3

Fig. 2. Distribution of Siphonochelus (Trubatsa) wolffi n. sp.

Type locality: New Caledonia, off Nouméa, reef flat, îlot Maître, 22°20´S, 166°26´E, detritic mounts, tidal [LAGON: stn 1351, Bouchet & Marshall coll., 25 November 1992]. Material examined: New Caledonia. LAGON, stn 1351, off Nouméa, reef flat îlot Maître, 22°20´S, 166°26´E, detritic mounts, tidal. Distribution: Only known from the type material, New Caledonia, off Nouméa, tidal. Etymology: Acares (Greek): short, little, in reference to the tiny shell. Description: Shell small for the genus, reaching 4 mm in height (holotype), height / width ratio 2.4, slender, narrow, nodose. Subsutural ramp strongly sloping, weakly convex. Light tan with slightly darker coloured subsutural ramp and base of siphonal canal; spiral cords P1, P2, P3 and tip of siphonal canal creamy white. Narrow space between cords brown. Aperture creamy white, occasionally with brown lines between. spiral cords seen in transparency. Spire very high with 1.5 protoconch whorls (Fig. 29). Teleoconch of 4 narrow, weakly shouldered, nodose whorls. Suture impressed. Protoconch large, low. Diameter 400 mm. Whorls smooth with single obvious narrow keel adapically. Terminal lip shallow, delicate, thin, weakly curved. Axial sculpture of teleoconch whorls consisting of high, broad, nodose ribs. Other axial sculpture of few weak growth striae. Spiral sculpture of high, strong, broad primary cords and weak lirae. First whorl with visible P1 and P2, second with narrow IP, and broad P1 and P2, third whorl of subadult specimen with narrow IP, P1–P6 and ADP, last whorl of holotype with narrow IP, P1–P6, ADP and weak MP. P1-P3 broadest, of similar size, P4–P6 narrow, also of similar size, ADP narrower. Entire shell covered with narrow spiral striae on and between primary spiral cords. Aperture small, narrow, ovate. Columellar lip narrow, smooth. Rim adherent, very weakly detached for short distance abapically. Anal notch broad, shallow. Outer lip erect with 4 weak denticles within (D1–D2 fused, D3, D4, D5). Siphonal canal relatively short, broad, tapering abapically, narrowly open. Operculum ovate with terminal nucleus on lower right (Fig. 31). Radula (Figs. 32–33) with rachidian bearing broad, long central cusp with weakly smaller and narrower lateral denticle, broad, long lateral cusp on each side and small, narrow, marginal cusp. Lateral teeth sickle-shaped with broad base. 4 R. Houart

Figs. 3–5. Litozamia acares n. sp., New Caledonia, off Nouméa, reef flat îlot Maître, stn 1351, 22°20´S, 166°26´E, detritic mounts, tidal. 3–4. Holotype, MNHN 25086, 4 mm. 5. Paratype, MNHN 25088, 3.4 mm. Figs. 6–10. Litozamia rudolphi (Brazier, 1895). 6–8. Green point, Watsons Bay, Sydney Harbour, NSW, Australia, syntypes, AM C2892; 6, 6.3 mm; 7–8, 5.9 mm. 9–10. North Head, Sydney, NSW, 33 m, 5.9 mm, RH. Figs. 11–16. Siphonochelus (Trubatsa) wolffi n. sp. 11–13. Eastern New Caledonia, 20°57´S, 165°35´E, 205–219 m, holotype, M NHN 25089, 14.3 mm. 14–16. Eastern New Caledonia, 21°52´S, 166°47´E, 120–180 m, paratype, MNHN 25093, 9.7 mm. New Species of Trophoninae and Typhinae 5

Figs. 17–18. Siphonochelus (Trubatsa) wolffi n. sp., Eastern New Caledonia, 21°02´S, 165°38´E, 350–400 m, paratype, MNHN 25094, 19.6 mm. Figs. 19–22. Siphonochelus (Trubatsa) unicornis Houart, 1991. 19–20. New Caledonia, 22°47´ S, 167°22´ E, 390 m, holotype, MNHN 0890, 12 mm. 21–22. South New Caledonia, 22°59´ S, 168°21´ E – 23°S, 168°23´ E, 491–558 m, RH, 11.6 mm. Figs. 23–24. Siphonochelus (Trubatsa) saltantis Houart, 1991, Coral Sea, Nova Seamount, 22°12´ S, 159°23´ E, 332 m, holotype, MNHN 0896, 9.2 mm. Figs. 25–28. Siphonochelus (Choreotyphis) pavlova (Iredale, 1936), Cape Moreton, Queensland, Australia; 25–27, 15.5 mm; 28, 11.2 mm. 6 R. Houart

Figs. 29, 31–33. Litozamia acares n. sp. 29. Protoconch of the specimen illustrated in Text Fig. 1 (scale bar = 250 mm). 31. Operculum (scale bar = 200 mm). 32–33. Radula (32, scale bar = 20 mm; 33, scale bar = 10 mm). Fig. 30. Protoconch of Litozamia rudolphi (Brazier, 1895), RH (scale bar = 250 mm).

Remarks: This is the second known species of Litozamia collected in New Caledonia. Litozamia tropis Houart, 1995 from northeastern New Caledonia, described from empty shells only, differs from L. acares n. sp. in having a double keel on the protoconch, and in having a strongly spirally striate, more strongly shouldered and spiny shell. The protoconch in both species has the same number of whorls, but that of L. tropis is about half the size. Litozamia acares n. sp. differs from the quite similar L. rudolphi (Brazier, 1895) (Figs. 6–10) from New South Wales, Australia in having a relatively smaller and narrower, less shouldered shell with a less sloping and narrower subsutural ramp, broader and more narrowly spaced primary spiral cords, and a relatively narrower and smaller aperture. The protoconch in L. acares n. sp. is more strongly keeled, with a keel situated more abapically (Fig. 29), compared to L. rudolphi, which has a more rounded protoconch with a narrow adapical keel (Fig. 30). The protoconch of L. rudolphi was described as “mammillated” [sic] (Brazier in Henn, 1895: 166). Beechey (2010) also illustrated 3 specimens of L. rudolphi from New South Wales. The species is restricted to New South Wales and Tasmania. The genus Litozamia currently includes 6 species: L. brazieri (Tenison Woods, 1876), L. latior (Verco, 1909), L. rudolphi (Brazier, 1895), L. subtropicalis (Iredale, 1913) and L. tropis Houart, 1995. longior Verco, 1909, until now considered a synonym of T. rudolphi (see Houart, 2010), has a quite different protoconch morphology consisting of a first whorl with a duplicate New Species of Trophoninae and Typhinae 7 keel and a last whorl with a strong single keel (Verco, 1909: fig. 5), and is here reinstated as a valid species.

Subfamily Typhinae Cossmann, 1903 Genus Siphonochelus Jousseaume, 1880 Type species by original designation: arcuatus Hinds, 1843, South Africa (Fig. 41).

Subgenus Trubatsa Dall, 1889 Type species by subsequent designation (Keen, 1944: 57): Typhis longicornis Dall, 1888, Cuba (Fig. 38).

Remarks: Trubatsa differs from Siphonochelus s.s. in having anal tubes with a much broader base than in Siphonochelus s.s. that extend almost over the entire intervarical area, being flattened at their base. The tubes and the axial varices also extend as a broad buttress on preceding whorl. In Siphonochelus s.s. the tubes have a narrower base and they lack the broad buttress observed in Trubatsa. Trubatsa also differs from Siphonochelus s.s. in having the intervarical fold (or ridge) closer to the preceding varix (Figs. 37–39, 41–42). Radwin & D’Attilio (1976: 206) and D’Attilio & Hertz (1988: 62) considered Choreotyphis Iredale, 1936 (type species pavlova Iredale, 1936) a junior synonym of Trubatsa. However, in Siphonochelus (Choreotyphis) pavlova (Figs. 25–28) the axial varices are much broader and rounder than in Trubatsa and the anal tube originates directly from the varix (Fig. 40). In Siphonochelus (Siphonochelus) and S. (Trubatsa) the tubes are connected to the succeeding varices but they originate just behind them. Based on these differences, I now propose the following classification for Trubatsa and Choreotyphis: Eight species are here assigned to Siphonochelus (Trubatsa): Siphonochelus (Trubatsa) erythrostigma Keen & Campbell, 1964 (Queensland, Australia), S. (T.) longicornis (Dall, 1888) (Cuba) (type species), S. (T.) lozoueti Houart, 1991 (new combination) (New Caledonia), S. (T.) saltantis Houart, 1991 (new combination) (New Caledonia), S. (T.) tityrus (Bayer, 1971) (Panama and Venezuela), S. (T.) undulatus Houart, 1991 (new combination) (New Caledonia), S. (T.) unicornis Houart, 1991 (new combination) (New Caledonia) and the new species described herein. One species is referable to Siphonochelus (Choreotyphis): S. (C.) pavlova (Iredale, 1936) (Queensland and New South Wales, Australia).

Siphonochelus (Trubatsa) wolffi n. sp. (Figs. 11–16, 17–18, 34, 37)

Siphonochelus (Siphonochelus) pavlova – Houart, 1991: 229, figs. 4, 35, 45, 56, 63 [not S. (C.) pavlova (Iredale, 1936)]. Type material: Holotype, MNHN 25089; 8 paratypes, MNHN 25090–25094; 2 paratypes, RH; 1 paratype, JW. Type locality: Eastern New Caledonia, 20°57´S, 165°35´E, 205–219 m [BATHUS 1: stn CP 668, Bouchet & Richer coll., 14 March 1993]. Material examined: New Caledonia. VAUBAN 1978–79. stn DR40, 22°30´S, 166°24´E, 250–350m, 10 dd. BIOCAL. Stn DW77, 22°15´S, 167°15´E, 440 m, 1 lv, 1 dd. MUSORSTOM 4. Stn CC246, 22°08´S, 167°11´E, 410–420 m, 1 lv; stn CC247, 22°09´S, 167°13´E, 435–460 m, 1 lv. MUSORSTOM 5. Stn DW301, 22°07´S, 159°25´E, 487–610 m, 1 lv. BATHUS 1. Stn DW639, 21°52´S, 166°47´E, 120–180 m, 1 lv, paratype, MNHN 25093; stn DW640, 21°52´S, 166°48´E, 174 m, 1 dd; stn DW642, 21°52´S, 166°50´E, 302–305 m, 1 dd; stn 8 R. Houart

Figs. 34–36. Protoconchs of Siphonochelus spp. 34. Siphonochelus (Trubatsa) wolffi n. sp., VAUBAN 1978–79, M NHN. 35. Siphonochelus (Trubatsa) saltantis Houart, 1991, M NHN. 36. Siphonochelus (Trubatsa) unicornis Houart, 1991, paratype, MNHN (scale bar = 500 mm). Figs. 37–42. Profile of last teleoconch whorl. 37. Siphonochelus (Trubatsa) wolffi n. sp., New Caledonia, M USORSTOM 4, M NHN. 38. Siphonochelus (Trubatasa) longicornis (Fall, 1888), Cuba, RH. 39. Siphonochelus (Trubatsa) unicornis Houart, 1991, New Caledonia, paratype, MNHN. 40. Siphonochelus (Choreotyphis) pavlova (Iredale, 1936), Queensland, Australia, RH. 41. Siphonochelus (S.) arcuatus (Hinds, 1843), Algoa Bay, South Africa, RH. 42. Siphonochelus (S.) boucheti Houart, 1991, New Caledonia, paratype, MNHN 1045 (scale bar = 1000 mm). New Species of Trophoninae and Typhinae 9

DW653, 21°17´S, 165°57´E, 190–207 m, 1 dd; stn CP656, 21°13´S, 165°54´E, 452–460 m, 4 dd; stn CP668, 20°57´S, 165°35´E, 205–219 m, 1 lv, holotype, M NHN 25089, 2 dd, 1 lv, 2 paratypes, MNHN 25090, 1 RH; stn CP669, 20°57´S, 165°35´E, 255–280 m, 1 dd; stn DW672, 20°48´S, 165°21´E, 347–366 m, 2 dd; stn DW687, 20°35´S, 165°07´E, 408–440 m, 1 dd; stn DE700, 20°57´S, 165°35´E, 160–222 m, 1 dd; stn CP701, 20°58´S, 165°36´E, 302–335 m, 4 dd; stn CP702, 20°35´S, 165°35´E, 591–660 m, 1 lv, 1 dd, paratypes, M NHN 25091; stn DE705, 21°02´S, 165°38´E, 350–400 m, 1 lv, paratype, MNHN 25094; stn DW706, 21°42´S, 166°34´E, 247–252 m, 7 dd, 1 paratype, RH; stn CP707, 21°43´S, 166°36´E, 347–375 m, 4 dd; stn CP711, 21°43´S, 166°36´E, 315–327 m, 1 dd; stn CP712, 21°44´S, 166°35´E, 210 m, 1 lv, 5 dd, 1 paratype, JW; stn CP713, 21°45´S, 166°37´E, 250 m, 2 lv, 1 dd, paratypes, M NHN 25092. BATHUS 2. Stn DW719, 22°48´S, 167°16´E, 444–445 m, 1 dd; stn DW724, 22°48´ S, 167°26´ E, 344–358 m, 1 lv; stn DW730, 23°03´S, 166°58´E, 397–400 m, 1 lv; stn DW739, 22°35´S, 166°27´E, 465–525 m, 2 dd; stn CP760, 22°19´S, 166°11´E, 45 5m, 1 dd. BATHUS 3. Stn DW809, 23°39´S, 167°59´E, 650–730 m, 1 dd; stn DW838, 23°01´S, 166°56´E, 400–402 m, 3 dd. BATHUS 4. Stn DW 885, 22°05´S, 165°58´E, 250–300 m, 1 dd; stn CP947, 20°34´S, 164°58´E 470–490 m, 1 dd (RH). SMIB 8. Stn DW 197–199, 22°51´S, 167°12´E–22°52´S, 168°12´E, 408–436 m, 1 dd. HALIPRO1. Stn CP851, 21°43´S, 166°37´E, 314–364 m, 1 lv; stn CP853 241, 21°45´S, 166°37´E, 241–250 m, 1 lv. Distribution: Chesterfield Islands and New Caledonia, live in 180–591 m, shells in 174–650 m (Fig. 2). Etymology: Named for John Wolff, Lancaster, Pennsylvania, USA, in recognition of his help in improving the quality of the English in many of my papers and for other comments. Description: Shell medium sized for the subgenus, up to 19.6 mm in height at maturity (paratype, MNHN) (Figs. 17–18). Height / width ratio (without tubes) 2.1–2.2, broadly ovate, lightly built. Subsutural ramp narrow, weakly sloping, weakly concave. Creamy white with orange-brown spiral bands at base of anal tubes, at periphery and at base of siphonal canal. Occasionally entirely creamy white. Aperture white. Spire high with 2 protoconch whorls and teleoconch of up to 6 broadly convex, strongly shouldered whorls. Suture impressed. Protoconch large, 800 mm high × 700 mm wide, weakly elongate. Whorls broadly convex, smooth; terminal lip delicate, thin, curved. Axial sculpture of teleoconch whorls consisting of 4 high, broad, shouldered varices from first to last whorl. From second or third whorl, varices more broadly expanded adapically, largely extending above shoulder, forming broad buttress, fused to preceding whorl. Intervarical axial sculpture of single narrow rounded ridge, closer to preceding varix, shell weakly excavated between ridge and succeeding varix. Spiral sculpture of very shallow P1 with anal tube and P2 (shoulder). Other spiral sculpture of numerous striae and lirae. Anal tubes ventrally sealed, rounded, broader at base, forming angle of approximately 45°–80° with axis of shell. Last (apertural) tube occasionally very long (Figs. 14–18). Aperture small, 15% of shell length in holotype, broadly ovate, rim forming a continuous peristome, smooth within. Siphonal canal long, ventrally sealed, weakly abaxially and dorsally bent, broad at base, strongly tapered abapically. Operculum and radula not examined. Remarks: Siphonochelus (Trubatsa) wolffi n. sp. can be compared with S. (T.) saltantis (Figs. 23–24, 35) another New Caledonian species that is quite similar in shell morphology. However, S. (T.) wolffi consistently differs in being comparatively larger, higher and narrower, S. (T.) saltantis reaching a height of 9 mm, with a height / width ratio of 1.9 compared to 2.1–2.2 in S. (T.) wolffi. The new species also has a larger and more elongate protoconch than that of S. (T.) saltantis, which is rounded and quite smaller (600–700 mm high and 500 mm wide). Finally the intervarical 10 R. Houart fold is more obvious in S. (T.) wolffi while very narrow or almost completely covered by the preceding whorl in S. (T.) saltantis and the siphonal canal is comparatively longer and narrower at its base. In S. saltantis the siphonal canal is short, strongly and promptly tapering to become very narrow at its abapical extremity. Siphonochelus (Trubatsa) wolffi n. sp. differs also from S. (T.) unicornis from New Caledonia (Figs. 19–22, 36, 39) in having a markedly lower spire with a smaller protoconch, 800 × 700 mm vs. 1100 × 1000 mm, a comparatively broader shell with broader varices and a narrower, longer siphonal canal. Siphonochelus (Trubatsa) wolffi n. sp. also differs from S. (Choreotyphis) pavlova (Figs. 25–28, 40) with which it was confused by Houart (1991), by the same characters that differentiate Trubatsa and Choreotyphis, i.e., in having the anal tubes connected to the succeeding varices, but originating just behind them, and not directly from them, and in having a narrow, obvious intervarical fold vs. none in. S. pavlova. It also differs in having narrower varices, a more elongate and higher spire and a distinctly smaller protoconch with a height of 800 mm vs. 1000 mm in. S. pavlova and a width of 700 mm vs. 800 mm.

Acknowledgements

I am much indebted to Philippe Bouchet (Muséum national d’Histoire naturelle, Paris) for allowing me to study the Muricidae collected during the expeditions conducted by MNHN and IRD in the Tropical Indo-Pacific. I am also very grateful to Dr. Bouchet and Virginie Héros (MNHN) for the loan of material, comments on this paper and digital images of type material, and to Anders Warén (Natural History Museum, Stockholm) for radula preparation and SEM work on the radula and shell morphology of Litozamia acares n. sp. Thanks are also due to Alison Miller and Mandy Reid (Australian Museum, Sydney) for photographs of the syntypes of Litozamia rudolphi, to John Wolff, Lancaster, Pennsylvania, USA, for checking my English and for other comments, and to two anonymous referees for their helpful comments.

References

Barco, A., Claremont, M., Reid, D. G., Houart, R., Bouchet, P., Williams, S. T., Cruaud, C., Couloux, A. & Oliverio, M. 2010. A molecular phylogenetic framework for the Muricidae, a diverse family of carnivorous gastropods. Molecular Phylogenetics and Evolution 56: 1025–1039. Beechey, D. 2010. The Seashells of New South Wales (Based on the collections of the Australian Museum and of the author). Accessed through: http://seashellsofnsw.org.au/Trophoninae/Pages/litozamia_rudolphi.htm, accessed on 20.03.2012. Bouchet, P., Héros, V., Lozouet, P. & Maestrati, P. 2008. A quarter-century of deep-sea malacological exploration in the South and West Pacific: Where do we stand? How far to go? In: Héros, V., Cowie, R. H. & Bouchet, P. (eds.), Tropical Deep-Sea Benthos 25. Mémoires du Muséum national d’Histoire naturelle 196: 9–40. D’Attilio, A. & Hertz, C. M. 1988. An illustrated catalogue of the family Typhidae Cossmann, 1903. The Festivus 20 (supplement): 1–73. Henn, A. U. 1877. List of found at Green Point, Watson’s Bay, Sydney. With a few remarks upon some of the most interesting species and descriptions of the new species and descriptions of the new species, by John Brazier, F.L.S., C.M.Z.S. Proceedings of the Linnean Society of New South Wales, Second Series 9: 165–182. Houart, R. 1991. Mollusca, Gastropoda: The Typhinae (Muricidae) from the New Caledonian region with description of five new species. Mémoires du Muséum national d’Histoire naturelle, (A), 150. Résultats des Campagnes MUSORSTOM 7: 223–241. Houart, R. 1995a. The Ergalataxinae (Gastropoda, Muricidae) from the New Caledonia region with some comments on the subfamily and the description of thirteen new species from the Indo-West Pacific. Bulletin du Muséum national d’Histoire naturelle, Paris, 4e sér., 16, section A, n° 2–4: 245–197. New Species of Trophoninae and Typhinae 11

Houart, R. 1995b. The Trophoninae (Gastropoda: Muricidae) of the New Caledonia region. Mémoires du Muséum national d’Histoire naturelle, 167. Résultats des Campagnes MUSORSTOM 14: 459–498. Houart, R. 2010. Litozamia rudolphi (Brazier, 1894). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=399198 accessed on 20.03.2012. Keen, A. M. 1944. Catalogue and revision of the gastropod subfamily Typhinae. Journal of Paleontology 18: 50–72. Merle, D. 1999. La radiation des Muricidae (Gastropoda : ) au Paléogène: approche phylogénétique et évolutive. vi + 499 pp. Unpublished thesis, Muséum national d’Histoire naturelle, Paris. Merle, D. 2001. The spiral cords and the internal denticles of the outer lip in the Muricidae: terminology and methodological comments. Novapex 2: 69–91. Radwin, G. & D’Attilio, A. 1976. Murex shells of the world. An illustrated guide to the Muricidae. 284 pp. Stanford University Press, Stanford. Verco, J. 1909. Notes on South Australian marine Mollusca, with descriptions of new species. 12. Transactions of the Royal Society of South Australia 33: 293–342.

(Received March 31, 2012 / Accepted June 25, 2012)

ニューカレドニアからツノオリイレ亜科とパイプヨウラク亜科の 2新種の記載， および Litozamia属と Siphonochelus属についてのコメント

Roland Houart

要 約

ニューカレドニアから Litozamia acares n. sp. と Siphonochelus（Trubatsa） wolffi n. sp.を新種として記載 した。Litozamia acares n. sp.については，この属として初めて歯舌と蓋についても図示し，形態を記載し た。近年，ツノオリイレ亜科は 2つの異なるグループからなる多系統群であることが示されているが， 形態的に区別は困難であり，Litozamia属については分子データによる裏付けがないため，仮にツノオリ イレ亜科に含めた。この属には従来 5種類が知られていたが，T. rudolphi（Brazier, 1895）の異名とされ ていた L. longior（Verco, 1909）は原殻の形態などから有効な種であることを認め，本新種と合わせて 7 種となった。一方， Siphonochelus属については，属内の種の詳しい検討の結果， Trubatsa亜属を有効な タクソンと認め，さらに近年の研究者によってこの亜属の異名とされていた Choreotyphis亜属も Typhina pavlova Iredale, 1936単一種を含む有効な亜属とみなした。