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Journal of Natural History Publication details, including instructions for authors and subscription information: http://www.informaworld.com/smpp/title~content=t713192031 Revision of the Australian wolf spider genus Dingosa Roewer, 1955 (Araneae, Lycosidae)

Online Publication Date: 01 January 2007 To cite this Article: Framenau, Volker W. and Baehr, Barbara C. (2007) 'Revision of the Australian wolf spider genus Dingosa Roewer, 1955 (Araneae, Lycosidae)', Journal of Natural History, 41:25, 1603 - 1629 To link to this article: DOI: 10.1080/00222930701475717 URL: http://dx.doi.org/10.1080/00222930701475717

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The publisher shall not be liable for any loss, actions, claims, proceedings, demand or costs or damages whatsoever or howsoever caused arising directly or indirectly in connection with or arising out of the use of this material. © Taylor and Francis 2007 Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 l pce auei aesme owne;fmlswt gsccnuulyb on nJl and July in considered found here be is usually species can This possible. eggsac not with is females entrance. identification Australian winter; burrow of species the their accurate to of patterns around summer holotype turret late colour The characteristic August. in a distinct mature construct they and species where are All region genus vegetation mark) the cardiac of cephalic within species. (serrated cover Species opisthosoma some sparse Lycosinae. raised and in Australian female eyes) anteriorly the a between within the enlarged bent band unique as part of an longitudinal transverse has septum light such the (narrow which median of prosoma characters, pedipalp, The corners male somatic apophysis. the the with tegular Additional of T-shaped elongated structure inverted unusually is the an epigyne by and spiders division lycosine embolic other all from differs 95)adfrel ecie ytesm uhri usqetpbiainbsdon based publication subsequent a in of author same holotype the female by the described formerly and 1955a) urnl nlddi hsgnsd o ofr otedanssof Australia diagnosis outside the from to species conform Other not do morphology. genus lycosine this derived in much included with currently genus Australian an is serrata D. ( utai,and Australia, rma mauefml ooyewihi ot hsseisi eeconsidered here is species this lost; is which holotype female immature an from History Natural of Journal O:10.1080/00222930701475717 49, online DOI: Bag 1464-5262 Locked print/ISSN Museum, 0022-2933 [email protected] Australian ISSN Email: Western Australia. Invertebrates, 6986, Terrestrial W.A. of DC, Department Welshpool Framenau, W. Volker Correspondence: genus spider wolf The Introduction Keywords: descriptions: original (Angola), the of comb. evaluation n. critical a 1959), on (Roewer, based placements generic new genus spider wolf Australian The Abstract 1 FRAMENAU W. VOLKER Lycosidae) genus (Araneae, spider 1955 wolf Roewer, Australian the of Revision ads hartmanni Pardosa simsoni Australia, ob Cia.W lospotMzfainadMrsks(01 rvossgetdcombination suggested previous (2001) Marusik’s and Mozaffarian persica support Trochosa also We (China). comb. aooy turret-building taxonomy, cetd2 a 2007) May 25 Accepted eateto ersra netbae,WsenAsrla uem et,WsenAustralia, Western Perth, Museum, Australian Western Invertebrates, Terrestrial of Department Smn 88 tp species); (type 1898) (Simon, L oh 87,n comb. n. 1877), Koch, (L. 2 colo nmlBooy nvriyo etr utai,Caly etr Australia, Western Crawley, Australia, Western of University Biology, Animal of School rcnd,Aaee Australia, Araneae, Arachnida, 3 uesadMsu,SuhBibn,Qenln,Australia Queensland, Brisbane, South Museum, Queensland Ree,1955). (Roewer, Ree,15) .cm.(azna,and (Tanzania), comb. n. 1959), (Roewer, Dingosa yoasimsoni Lycosa 07 12–8:1603–1629 41(25–28): 2007; , igs traghardi Dingosa Dingosa owr 95wsiiilylse as listed initially was 1955 Roewer, 1,2 Dingosa .humphreysi D. # ads completa Pardosa ABR .BAEHR C. BARBARA & 07Tyo Francis & Taylor 2007 igs topaziopsis Dingosa owr 95i eie oicuefu species: four include to revised is 1955 Roewer, io,19 olce nLucso,Tasmania Launceston, in collected 1898 Simon, eog otesbaiyLcsneSneal 83and 1833 Sundevall, Lycosinae subfamily the to belongs igs,Lycosa Dingosa, Lwec,14)(ot fia a described was Africa) (South 1947) (Lawrence, MKy 95,n comb.; n. 1985), (McKay, Ree,15) .cm.(Mozambique), comb. n. 1959), (Roewer, Hg,19)i nimtr pdrand spider immature an is 1896) (Hogg, rcoaursina Trochosa e species, new , Dingosa Dingosa 3 rfrsnyhbtt iha with habitats sandy prefer epooetefollowing the propose We . oe nudum nomen oe dubium nomen Dingosa Pardosa .murata D. Shne,13) n. 1936), (Schenkel, oe dubium nomen ads angolensis Pardosa systematics, , .s. and sp.; n. (Roewer . Dingosa Dingosa . Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 1604 pce is,floe yalohrseisi lhbtclodr h morphological The order. Species alphabetical type (mm). in the millimetres list species publication other in this all of are by part measurements taxonomic followed the All first, in species h. at 1–3 re-descriptions acid lactic and for in either descriptions submersion by was temperature examination for overseas room prepared were lodged genitalia institutions. Female material respective Germany). the Type all visited author from country. senior spiders the wolf this whilst examined of personally in records or 20,000 borrowed collections about of museum examination comprehensive major a on based is of study Our species. these of Methods descriptions original the on based mainly placements generic alternative (e.g. 2007 genus Framenau, to The belong Lycosinae clades. Artoriinae Australian distinct that suggested and two terminals analysis spider phylogenetic molecular wolf 1979, Hippa, recent world-wide A 70 and 2006). Hippa, including Lehtinen Framenau and Zoicinae Yoo and the 2007; 2006c, to Framenau Lehtinen addition in Venoniinae subfamilies, speciose spider 1979, most wolf the Australian are Lycosinae the The of 1986). (Dondale 1833 Sundevall, been Lycosinae subfamily pedipalp, has male the behaviour of apophysis Similar their around genus 1985a). Holarctic rocks 1979, the or of litter McKay species of Australian some 1976; turret the or Main in palisade reported 1965; a constructing (Lane of burrow habit unusual their for transferred he Subsequently, placed. pdrgenus American spider South the with clade monotypic a all gallopavo whereas formed taxa, Hemisphere Lycosinae Northern to Australian relationships closest other have to appear 2006 Yoo, and ecie iiaiyo h aepdpl.H lodsrbdascn pce ihsimilar with both species morphology, second genital a and described somatic peculiar also He pedipalp. in male morphology the of similarity perceived rtclyrve l te o-utainseiscretylse in listed currently species non-Australian other all review critically ca 17)ddraieta h nsa opooyo h aepdpl xlddthis excluded pedipalp male the of morphology unusual from the species that realize did (1979) McKay oh 87 i o eonz htti pce scneei with congeneric is species this that recognize not did 1877, Koch, 2003). remote Museum only Australian 1970; shows Mascord that name (e.g. this scientific Spider’’, Spider of the Wolf and material wolf ‘‘Grey literature trapdoor-building type the as true enigmatic, the the known an with examined commonly described similarity never Australia and he in illustrated that erroneously spider, 2007). he appears 1998, as it Platnick be species (e.g. to however, catalogues believed 1804), later he in Latreille, what accepted re-described not (1967) was Hickman (1966) Guy by decisions Subsequently, Allocosa 1955b). (Roewer (Australia) pdr eeeaie n7%EO ihaWl troirsoe(Heerburg, stereomicroscope Wild a with EtOH 70% in examined were Spiders tegular extended an and division embolic enlarged an to due unusual somewhat Although h i forsuyi orvs aooial h orAsrla pce ftewolf the of species Australian four the taxonomically revise to is study our of aim The ti hrfr o upiigta ca 17) hnh re-described he when (1979), McKay that surprising not therefore is It ak,10 Gy16)btti yoyyi diint ayohrtaxonomic other many to addition in synonymy this but 1966) (Guy 1900 Banks, (Mello-Leita .W rmnu&B .Baehr C. B. & Framenau W. V. Dingosa Dingosa Lycosa Pardosa or o 91 ugsigacaeo odaa rgn(upye l 2006). al. et (Murphy origin Gondwanan of clade a suggesting 1941) ˜o, including , om ato ytmtcrvso ftewl pdr fAsrlaand Australia of spiders wolf the of revision systematic a of part forms Geolycosa , .simsoni D. .humphreysi P. ans longipes Mainosa Venatrix Dingosa .murata D. otoey 94 where 1904, Montgomery, notntl,Hcmns(97 itk esse in persisted mistake (1967) Hickman’s Unfortunately, . Geolycosa .simsoni D. .serrata L. owr 90adterelated the and 1960 Roewer, lal ofrst h igoi ftewl spider wolf the of diagnosis the to conforms clearly ca,18 MKy18a.I diint a to addition In 1985a). (McKay 1985 McKay, .s. hc snwt cec.I diin we addition, In science. to new is which sp., n. Dingosa L oh 88 Faea 06)adin and 2006a) (Framenau 1878) Koch, (L. Wlae 1942). (Wallace, .serrata P. to scmol trbtdt h ryWolf Grey the to attributed commonly is Pardosa a osdrdajno yoy of synonym junior a considered was igs simsoni Dingosa and .L oh 87bsdo a on based 1847 Koch, L. C. .humphreysi P. yoaserrata Lycosa Tuberculosa .simsoni D. Dingosa ndti (in detail in yoaserrata Lycosa r elknown well are a previously was n suggest and However, . Framenau Pavocosa Lycosa L. Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 Taxonomy i ftglraohsso aepdpl oeo essrih Fgr B;lateral 5B); (Figure straight less or more pedipalp male of apophysis tegular of Tip – .Tpo eua ppyi fml eiapdsicl uvdvnrly(iue3B); (Figure ventrally curved distinctly pedipalp male of apophysis tegular of Tip 3. i ftglraohssbut itntykee nvnrlve Fgr A;female 6A); (Figure view ventral in keeled distinctly blunt, apophysis tegular of Tip – .Tpo eua ppyi fml eiapcre etal Fgr D;epigyne 2D); (Figure ventrally curled pedipalp male of apophysis tegular of Tip 2. h pia ueo troirsoe(Z;LiaMcoytm,Wtlr emn)with Germany) (MaxView Wetzlar, set Microsystems, adapter Leica (MZ6; optical stereomicroscope an a of tube optical which the for (2000). genitalia Sierwald female of of terminology exception the the applied with we (2006b) Framenau follows nomenclature rsae A,SuhAsrla uem dlie M,SnknegMuseum, Senckenberg fu Museum, Museum SMF, Queensland ZMB, Perth; Adelaide; QM, Humboldt-Universita Museum, der Museum, of Australian Zentralinstitut Melbourne; Western Australian Museum WAM, Victoria, South Frankfurt; MCZ, Museum d’Histoire SAM, London; National NMV, Museum Brisbane; Museum, Australian MHNP, Paris; MA; History AM, Naturelle, Cambridge, Collections: Natural Harvard, width. Zoology, BMNH, opisthosoma Comparative prosoma OW, Sydney; PW, length; length; prosoma Museum, PL, opisthosoma length; total OL, TL, stated): width; otherwise not software Measurements if lateral. posterior spiders, the PLE, (adult median; with posterior PME, combined posterior; PE, lateral; and anterior 2006). images) al. 10–15 et (Khmelik (ca 4.07 planes Focus focal Helicon different package in taken were akcricmr noitooawal ertd ihtofa,tohlk shapes tooth-like flat, two with serrated, weakly opisthosoma on mark cardiac Dark – a forms serration anterior first serrated, strongly opisthosoma on mark cardiac Dark 1. ca 95,p1115 ca 95,p85. p 1985b, McKay 101–105; p 1985a, McKay Pardosa Lycosa htgah eetknwt iia aea(6 ao,Jpn htwscnetdto connected was that Japan) Canon, (G6; camera digital a with taken were Photographs h olwn brvain r sd ys E neir M,atro ein ALE, median; anterior AME, anterior; AE, Eyes: used. are abbreviations following The de ftases ato einspu nfml pgn truncated epigyne female in . septum median . of . part transverse 5C) (Figure of edges aea de ftases ato einspu nfml pgn ecigunder reaching . epigyne female 3C) in (Figure septum pockets median anterior of the part transverse of edges lateral pgn ogrta ieadatro agnntcniuu Fgr C . . 6C) (Figure continuous not margin anterior and wide than longer epigyne ie hnln n ihcniuu neirmri Fgr E . . 2E) (Figure margin anterior continuous with and long than wider Fgrs3,5) aea de ftases ato einspu extend septum median of . part transverse . tooth of basal 2 edges with 5C) pedipalp 3C, (Figures lateral with male anteriorly of . 5A); epigyne apophysis tegular 3A, female 1C–F); (Figures (Figure 6C) (Figure side 6A); wide either than on 2C, longer (Figures much or tooth 2E) (Figure basal margin anterior without continuous pedipalp 45 male ca of angle distinct aril,10 i at:L oh17,p9092 ika 97 80. p 1967, Hickman 930–932; p 1877, Koch L. part): (in 1804 Latreille, .L oh 87(npr) io 99 9–9;MKy17,p225–229; p 1979, McKay 191–192; p 1909, Simon part): (in 1847 Koch, L. C. ufml LYCOSINAE Subfamily aiyLYCOSIDAE Family u e oteseisof species the to Key ...... ihbd xs(iue1,B ,H;tglraohssof apophysis tegular H); G, B, 1A, (Figure axis body with re ARANEAE Order Dingosa 2 ls cptoi,Cp oa,F,UA.Photographs USA). FL, Coral, Cape Scopetronix, Plus; eiino h utainwl spider wolf Australian the of Revision t Berlin. ¨t, ...... 3 owr 1955 Roewer, lrk 1757 Clerck, udvl,1833 Sundevall, Dingosa udvl,1833 Sundevall, igs 1605 Dingosa .murata D. rNaturkunde, ¨r .humphreysi D. .simsoni D. .serrata D. .sp. n. Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 p .fo ot ai okNtr eev,WsenAsrla(A 112) G aeof male (G) 71/1428); (WAM Australia Western Reserve, Nature Rock Tarin North from n. sp. udc nvriycmu,Prh etr utai WMT62470). (WAM Australia Western Perth, campus, University Murdoch oh rmBl ak et,WsenAsrla(A 532;()fml of female (H) T58362); (WAM Australia Western Perth, Park, Bold from Koch) of holotype of of female of male (D) female (C) 16745); T53725); (B) (WAM T62462); Australia (WAM Australia Western pce ftegenus the of Species Diagnosis feminine). simsoni Lycosa species. Type obnto fsmtcadgntlccaatr.Tecpai rao h rsm is prosoma the of area cephalic The characters. genitalic and somatic of combination 1606 iue1 htgah dra iw fseisof species of view) (dorsal Photographs 1. Figure Dingosa Geolycosa Allocosa .murata D. .W rmnu&B .Baehr C. B. & Framenau W. V. owr15a 4 ( 240 p 1955a, Roewer ak,10 i at:Gy16,p3,4,63. 47, 33, p 1966, Guy part): (in 1900 Banks, otoey 94(npr) owr15a 4;MKy17,p380. p 1973, McKay 243; p 1955a, Roewer part): (in 1904 Montgomery, p .fo utop etr utai WMT10) F eaeprtp of paratype female (F) T51303); (WAM Australia Western Zuytdorp, from n. sp. .humphreysi D. Dingosa io,19,b rgnldsgain(owr15b (gender 1955b) (Roewer designation original by 1898, Simon, MKy rmGogri,WsenAsrla(A 469;()male (E) 84/649); (WAM Australia Western Goongarrie, from (McKay) ifrfo l te pce ihnteLcsdeb a by Lycosidae the within species other all from differ .humphreysi D. oe nudum nomen Dingosa .simsoni D. MKy rmHmig,SuhAsrla(A NN (SAM Australia South Hambidge, from (McKay) A aeof Male (A) . ;Ree 95,p762–763. p 1955b, Roewer ); Smn rmCrei oeta,Western Homestead, Carnegie from (Simon) .simsoni D. Smn rmGasPatch, Grass from (Simon) .serrata D. L oh from Koch) (L. .serrata D. .murata D. (L. Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 ai e eght rsm it:lg1 leg width: prosoma (average the males in to between particular 1A–H). band in length (Figure Lycosinae, light mark other leg median cardiac many in ratio serrated narrow than unique a longer a comparatively is longitudinal has are there and Legs opisthosoma and short The two PLE, 1A–H). has (Figure and prosoma eyes fovea the between of band lines median dark light the 2A), (Figure F). (E, elevated Left mm D) 0.10 Scale (C, D); respectively. (C, view. views, dorsal mm frontal 0.50 and and ventral B); epigyne, lateral (A, female mm carapace, F) 1.00 (E, Male bars: respectively; views, B) lateral (A, and ventral T53723). pedipalp, (WAM Australia Western Numbudding, 2. Figure pce) ncmaio,ti e ai smc oe o h pce fteAsrla genus Australian the of species the for lower much is ratio Venatrix leg this comparison, In species). igs simsoni Dingosa lg1 (leg 5 .,lg2 leg 3.6, Smn,ml rmGasPth etr utai WMT26)adfml from female and T62462) (WAM Australia Western Patch, Grass from male (Simon), 5 .,lg3 leg 3.2, eiino h utainwl spider wolf Australian the of Revision 5 .,lg4 leg 3.0, 5 .,lg2 leg 4.7, 5 .;dt o 7seisdrvdfrom derived species 17 for data 4.2; 5 .,lg3 leg 4.5, 5 .,lg4 leg 4.4, igs 1607 Dingosa 5 5.9; n 5 4 Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 ih ryt akoiege ihdsic akrsrae ada aki neirhalf, anterior in ( mark pattern IV cardiac formula humphreysi dark serrated Leg darker 1A–H). longitudinal dorsally (Figure generally distinct Opisthosoma bands with but females. with lateral wide, in light olive-grey long as by than dark long accentuated slightly wider as to little row about and grey largest AE males Labium median light in row; size. wide the PME equal than with longer of than teeth, little teeth shorter promarginal retromarginal row three three AE with medially and setae Chelicerae ALE, lines white 2B). dark of than (Figure line short larger procurved white two Narrow AME dark with V. to eyes. a band of forming brown Median between sometimes prosoma Flanks bands. fovea, of 2A). and submarginal (Figure shield PLE and Dorsal between region median 2B). (Figure cephalic light view with elevated frontal brown distinctly in steep with longer region Prosoma profile cephalic females. dorsal than smaller wide, Males than 6.50–15.00). (TL spiders wolf Medium-sized is apophysis Description points it tegular which in The Lycosinae most 6A). to curved contrast 5A, in and the 3A, apically, long less of with retrolaterally. 2C, or (Figures cavity, members more cymbium points for embolus and the large elongated of and unusually length apophysis the is half division terminal least embolic at The occupying 2001). Lycosinae Vink and Framenau mm 0.50 bars: Scale ventral respectively. views, pedipalp, dorsal Left D). and B) (C, ventral (A, mm epigyne, 0.10 84/649). female B); (WAM D) (A, (C, Australia respectively; Western views, Goongarrie, lateral from and female and T48586) (WAM 3. Figure 1608 w erltrl( retrolateral two igs humphreysi Dingosa .W rmnu&B .Baehr C. B. & Framenau W. V. , .murata D. .humphreysi D. .Siaino es ae:fmr he osl w apicoprolateral, two dorsal, three femur: males: legs: of Spination ). MKy,ml rmQenVcoi pig aueRsre etr Australia Western Reserve, Nature Springs Victoria Queen from male (McKay), , .serrata D. .serrata D. , ny,oeaioerltrl(isn in (missing apicoretrolateral one only), .simsoni D. ridsic ih ace ( patches light indistinct or ) . I . II . I.Lg brown Legs III. D. D. Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 prahclhassmwa lnae n oeie ihdsic neirtp( tip anterior distinct with sometimes and elongated humphreysi somewhat heads Spermathecal hsgnsmyb on ntemntpcAsrla genus Australian sinuous monotypic of the relative a closest in The with 2006). found al. apically) be et Murphy may elongated 1986; genus (Dondale somewhat surface this dorsal (although the on apophysis channel tegular transverse genus The Remarks in Females: (missing apicoretrolateral. retrolateral one apicoprolateral, two one dorsal, apicoventral, prolateral, two three one pairs, ventral three retrolateral, metatarsus: retrolateral; one two prolateral, two pairs, ventral murata tistp mou rgntn rltrlyo moi iiinadsmcrua,ln and long spines. semicircular, or and macrosetae division without embolic tip on Cymbium and prolaterally broad thin. narrow and originating semicircular relatively Subtegulum embolus apophysis ventrally. tip, terminal curved large, its or division at straight Embolic bulb. tip on its basally apically, situated two less pairs, or more ventral pointing three apicoventral. one tibia: pairs, retrolateral; ventral one three metatarsus: prolateral, prolateral; one patella: apicoretrolateral; rnvrebr na tews lcihoitooa svr different. morphology. very genital with and is genitalia spiders, opisthosoma, female blackish of and otherwise coloration male an However, the on males. bars and in transverse particular differences in considerable legs, long show very has and burrow ooye eae anetn(41 Launceston female, Holotype: examined Material sensu serrata Pardosa to Similar 2006a). (Framenau (misidentification). 550. p 1998, Platnick 83. p 1985b, McKay 379; p 1973, not but anteriorly, curve edges epigyne lateral female the the and in species, septum two median other in the the so of in part absent transverse is the that of pedipalp male the in apophysis in flat comparatively are bands these on shapes humphreysi tooth-like the whereas opisthosoma, the opooy ae fboth of Males morphology. eaeeiyewt netdTsae einspu idsic in (indistinct septum median T-shaped inverted with epigyne Female elongated, apophysis Tegular tegulum. narrow relatively undivided, with pedipalp Male h orseisof species four The ads serrata Pardosa not simsoni Dingosa simsoni Lycosa .simsoni D. yoasimsoni Lycosa ;ptla n rltrl n erltrl ii:oedra naia af three half, apical in dorsal one tibia: retrolateral; one prolateral, one patella: ); , and .murata D. Dingosa and .murata D. io 88 9 ano 91 7;Bne 97 62 McKay 2622; p 1957, Bonnet 273; p 1911, Rainbow 29; p 1898, Simon L oh:MClog 00 53,Fgrs31–.0 .5(not 3.25 3.17–3.20, Figures 35–36, p 2000, McCullough Koch): (L. Smn:Ree 95,p20 owr15,p30 iue199; Figure 360, p 1959, Roewer 240; p 1955a, Roewer (Simon): Simon .serrata. D. .Kc,17,misidentification). 1877, Koch, L. eog otesbaiyLcsnea h aepdpl a a has pedipalp male the as Lycosinae subfamily the to belongs .Semtea tlsshort. stalks Spermathecal ). Dingosa igs simsoni Dingosa igs simsoni Dingosa hs w rusaeas vdn hncmaiggenital comparing when evident also are groups two These . .humphreysi D. sensu Dingosa a esprtdit w itntgop ae nsomatic on based groups distinct two into separated be can Fgrs1,B AF 4) 2A–F, B, 1A, (Figures ika 97 0 iue 4,13 lt 3 iue3 Figure 13, Plate 143, 142, Figures 80, p 1967, Hickman u 26 eiino h utainwl spider wolf Australian the of Revision , 9 ,147 S, Mainosa and and .serrata D. Smn 1898) (Simon, u .murata D. 08 .humphreysi D. ulsatre rudteetac ftheir of entrance the around turret a builds 9 ,Tsai,Asrla MN 4640). (MHNP Australia) Tasmania, E, aevr itntsrae ad on bands serrated distinct very have aeatohbslyo h tegular the on basally tooth a have ,toaiorltrl one apicoprolateral, two ), Mainosa Mainosa igs 1609 Dingosa rmnu 2006 Framenau, aiglight having .serrata D. D. D. ). Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 7mlsS 32 S, miles 17 of Records 4. Figure 1610 te aeilexamined. material Other MKy lgtsurs 1records). 41 squares; (light (McKay) elhHl tto,29 Station, Hill wealth 34 Station, Calperum Lake, Clover 145 135 S42) w ae,FwesGp 31 Gap, Fowlers males, two KS84122); gsc on il ubln,33 Euabalong, Hill, Round eggsac, SMN170;oeml,tofmls u. onla mW 31 W, km 6 Koonalda, juv., 1 females, two male, one NN16770); (SAM n ae lno am,3k ,FneNtoa ak 24 Park, National Finke S, km 3 Palms, of Glen male, one A S05) n eae aeClaon,29 Callabonna, Lake female, one KS30556); (AM he eae,Lk arnr 32 Gairdner, Lake females, three N67) n eae ahrn itit 14 District, Katherine female, one NN16779); ihega,Lk ils 32 Gilles, Lake eggsac, with eae,RprRvrrgo,14 region, River Roper females, eae rofedCnevto ak 34 Park, Conservation Brookfield female, uyroCek B ag,31 Range, ABC Creek, Bunyeroo aaob,No nwe ae 31 Cave, Knowles of N Catacombs, u u 57 50 9 9 SMN170;oeml,HnigKol mW 26 W, km 2 Knoll, Hanging male, one NN16750); (SAM E A S42) n eae oa,3 ie ,31 E, miles 30 Cobar, female, one KS84121); (AM E .W rmnu&B .Baehr C. B. & Framenau W. V. u 12 Dingosa 9 ,141 S, u 57 nAustralia. in utai:NwSuhWales South New Australia: 9 u ,134 S, 27 u 47 9 Q 698;oefml ihega,Cbr 31 Cobar, eggsac, with female one S64938); (QM E 9 ,136 S, u u u 49 10 18 u u 00 u 00 igs simsoni Dingosa 9 9 24 9 ,134 S, SMN176;oefml,Hmig,33 Hambidge, female, one NN16726); (SAM E 00 u 9 9 u 48 ,140 S, ,146 S, 9 07 ,138 S, 0 u ,135 S, 04 9 u 9 SMN173;tofmls onreGap, Moonarie females, two NN16753); (SAM E ,130 S, 21 9 u ,141 S, 08 u 9 u ,139 S, 47 24 u u 9 32 50 NVK8165). (NMV E 9 9 u SMN170;oeml,Common- male, one NN16700); (SAM E u Smn fl ice;12records); 112 circles; (full (Simon) A KS84120). (AM E 52 9 37 u 9 u SMN1723;trefemales, three NN16732–3); (SAM E 20 28 43 u 9 9 28 00 SMN1646;oefemale, one NN16694–6); (SAM E 0 9 9 SMN1783) n female one NN16728–30); (SAM E ,132 S, A S39) n eaewith female one KS43591); (AM E 0 9 ,140 S, SMN169;tofemales, two NN16699); (SAM E n eae u. rknHill, Broken juv., 5 female, one : u 05 u 16 u 08 9 50 9 NVK15;two K8125); (NMV E 9 02 0 u ot Australia South u ,132 S, 29 19 otenTerritory Northern 0 SMNN16693); (SAM E 9 9 ,146 S, 23 u 0 u 27 ,130 S, 50 9 9 ,129 S, 00 u .humphreysi D. 20 0 u (SAM E 9 23 (AM E u u u 9 29 23 :one 36 51 9 9 0 9 S, S, E E : Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 696;trefmls on een 38 Nepean, Point females, three S64936); 34 31 Pound, Wilpena (A, mm 0.50 and bars: Scale ventral respectively. pedipalp, views, Left dorsal B) and D). (A, ventral (C, 71/1428). epigyne, mm (WAM 0.10 female B); D) Australia (C, Western respectively; Reserve, views, Nature lateral Rock Tarin North from 5. Figure 75k N,26 NNE, km 17.5 utai n xc oain et ein .J aecleto;seLn 95 (WAM 1965) Lane see collection; Lane J. W. region, Perth ? location, exact (no Australia Australia 75k N,26 NNE, km 17.5 SMN177;oefml,oefml ihega,Trwe 33 Terowie, eggsac, with female one female, one NN16727); (SAM N,26 NNW, SMN1712;oefml,Wyla 33 Whyalla, female, one NN16721–2); (SAM 26 n eaewt gsc hal an ak 33 Park, Fauna Whyalla eggsac, with female one N32) w eae,Wpn on ag,Fa fMoai a,31 Gap, Moonarie of E Flat Range, Pound Wipena females, two NN23325); 33 WM7/3–) w eae,WdnaHl,32 Hill, Wudinna females, two 71/637–8); (WAM 8). 138 S64937). 36 254;tofmls abtW na) ron re,Gmo ags 30 Ranges, Gammon Creek, Arcoona (near), WH Sambot females, two S24514); 139 u u u u 06 29 19 04 u u Tasmania 39 02 9 9 9 9 ,139 S, 57 50 ,141 S, 9 9 SMN171;fu ae,Rnak 9k N,33 NNW, km 29 Renmark, males, four NN16711); (SAM E SMN1767;oefml ihsielns ct re,na Morgan, near Creek, Scott spiderlings, with female one NN16736–7); (SAM E igs murata Dingosa 0 0 ,130 S, ,137 S, u Victoria w eae,tofmlswt gsc n eaewt pdrig,Western spiderlings, with female one eggsac, with females two females, two : 29 u 9 u 40 56 55 n eae ugaeCecn,Lucso,41 Launceston, Crescent, Mulgrave female, one : 9 0 u u 9 SMN174;oeml,SrbyPa,4k ,32 S, km 4 Peak, Scrubby male, one NN16704); (SAM E ,130 S, u 14 NVK24;oefml,Ltl eet 36 Desert, Little female, one K9264); (NMV E 26 u 19 19 n eae aeAbcta Wcre,c 5k S fYaapeet, of WSW km 15 ca corner, NW Albacutja, Lake female, one : 9 9 u 00 9 20 p . aehltp rmZydr,WsenAsrla(A 533 n female and T51303) (WAM Australia Western Zuytdorp, from holotype male n., sp. 9 36 20 20 0 0 u 9 SMN174;oefml,Pr il,Aead,34 Adelaide, Hills, Para female, one NN16754); (SAM E 0 SMN178;oeml,Mnao osrainPark, Conservation Munyaroo male, one NN16778); (SAM E 27 0 ,138 S, ,130 S, ,130 S, 9 22 0 u SMN1747;oeml,M itr,84k NW, km 8.4 Kintore, Mt male, one NN16774–7); (SAM E u u 07 37 07 9 9 56 9 SMN1745;oeml,M Cooperinna, Mt male, one NN16734–5); (SAM E 56 eiino h utainwl spider wolf Australian the of Revision 0 SMN1712;fu ae,M itr,7km 7 Kintore, Mt males, four NN16771–2); (SAM E 0 SMN173;tremls tCooperinna, Mt males, three NN16773); (SAM E u u 18 02 9 9 ,144 S, u ,137 S, u 59 02 9 9 44 ,137 S, u 0 u 34 ,135 S, 39 9 9 SMN236;oemale, one NN23326); (SAM E u A KS84119). (AM E 34 u 9 32 SMN176 SAM NN16766, (SAM E 9 52 u u u 26 u 40 33 31 0 SMNN16767– (SAM E 9 9 9 ,147 S, 48 9 ,141 S, igs 1611 Dingosa ,140 S, u 0 u ,138 S, 31 36 9 9 ,135 S, ,138 S, u u u 08 38 24 Western u 9 9 57 9 (QM E (QM E (QM E u u u u 9 48 26 19 44 37 9 9 9 0 9 S, S, E E E Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 529;fu ae,Boahn tto,24 24 Station, Station, Boolathana Boolathana males, males, four 24 Station, T51289); three Boolathana T51281); male, one (WAM T53900); T53884–6, T53880, 6. Figure 1612 iemls oltaaSain 24 Station, Boolathana males, five iw,rsetvl;(,D eaeeiye eta n oslves epciey cl as .0m A B); (A, mm 0.50 bars: Scale respectively. views, dorsal and lateral and ventral ventral D). epigyne, pedipalp, (C, Left female mm B) 0.10 D) (A, (C, T62470). (WAM respectively; Australia views, Western Perth, campus, University Murdoch orbn 31 Boorabin, 121 31 eae ann tc ot,Wl 01 24 10–1, Well Route, Stock Canning female, angeHmsed . mW angecm,25 camp, Carnegie W, km 0.5 Homestead, Carnegie WMT54,T55) n eaewt gsc ors,30 Forrest, eggsac, with female one T45857); T45843, (WAM nwneu/odnRas 34 Roads, Gnowangerup/Borden ae odaie a5k W 31 NW, km 5 ca Doodlakine, male, 561;oefml,Goaln od 31 Road, Goomalling female, one T53661); 31 onare 29 Goongarrie, tto,28 Station, 32 1 33 41, iemls xlmto ae 32 Lake, Exclamation males, five WMT19) iemls uhBy 25 Bay, Bush males, five T51290); (WAM xlmto ae 32 Lake, Exclamation ae 32 Lake, u u u 12 57 46 u 26 9 9 9 u ,120 S, 51 26 14 9 WMT54) n ae n eae xlmto aeae,32 area, Lake Exclamation female, one male, one T45849); (WAM E igs serrata Dingosa u 0 0 9 .W rmnu&B .Baehr C. B. & Framenau W. V. 49 ,117 S, ,121 S, ,121 S, u 47 9 18 u u 9 13 17 ,117 S, u 0 55 u ,121 S, u u 9 9 26 43 ,120 S, WMT14) n eae odn f tjnto Kojonup/ junction at of, N Borden, female, one T51249); (WAM E 56 9 ,121 S, 9 9 49 9 WMT26) n eae ra otenHgwy 1 iepeg mile 214 Highway, Northern Great female, one T62462); (WAM E 05 L oh,ml rmBlii,WsenAsrla(A 604 n eaefrom female and T60074) (WAM Australia Western Baldivis, from male Koch), (L. u 22 0 u u WMT59,T59,T59,T59–,T12,T51230); T51228, T45898–9, T45896, T45894, T45892, (WAM E 0 51 24 WMT14) 1mls w eae,EcaainLake, Exclamation females, two males, 11 T51248); (WAM E u 9 19 WM6/1) n ae uhBy 25 Bay, Bush male, one 69/819); (WAM E u 9 15 9 43 45 9 WMT15,T15) w eae,Boai Rock, Boorabin females, two T51253); T51251, (WAM E 9 0 WMT35) n ae rspth izeadLocality Fitzgerald Grasspatch, male, one T53850); (WAM E 0 ,121 S, WMT13) n ae xlmto ae 32 Lake, Exclamation male, one T51232); (WAM E u 00 u 36 u 9 u 24 ,118 S, 50 u 9 24 ,117 S, 9 9 49 28 9 04 0 0 u ,113 S, ,121 S, 16 0 u u u WMT13) n ae Exclamation male, one T51231); (WAM E u 04 48 33 51 9 WM6/1) n eae Burnabinmah female, one 69/915); (WAM E 9 u 9 9 9 40 WMT56) n ae rknRoad, Erikin male, one T55264); (WAM E ,116 S, 24 ,121 S, u 42 u u 0 9 23 48 ,113 S, 49 9 24 9 9 17 0 ,122 S, u u ,113 S, 0 29 39 WMT18) w females, two T51286); (WAM E 0 WMT12) i males, six T51229); (WAM E u u 9 9 24 36 WM7/9) n female, one 70/194); (WAM E NVK10;tofemales, two K8170); (NMV E u 9 9 58 u 49 37 40 9 0 0 E(WAMT53725–6);one u 9 113 S, E(WAMT51285);one 30 44 u u 24 07 0 9 ,127 S, WMT51288); (WAM E 9 9 49 34 u 39 0 0 u ,113 S, ,113 S, 47 9 u 47 41 9 ,121 S, 0 9 (WAM E E(WAM u u 40 49 u u 9 9 46 30 24 22 9 0 9 0 S, E E E Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 n xc oain WM6/5;oefml,Hrimt at 32 East, Harrismith female, one 69/85); (WAM location) exact (no 011;tofmls u. ye,TeHms 32 Humps, The Hyden, juv., 1 females, two 70/191); oain WM6/5) n eae ebn,Wo,31 of, W Yelbeni, female, one 68/856); (WAM location) 1) n eaewt gsc ot fWlbn,30 Walebing, of South eggsac, with female one 514); n ae oedeNtr eev,30 Reserve, Nature Jouerdie male, one 119 574;oeml,Ynann ae,32 Lakes, Yenyanning male, one T53724); n ae n eae amn 31 Tammin, female, one male, one ai okRsre 33 Reserve, Rock Tarin 561;egtmls ag tto,26 Station, Nanga males, eight T53621); aj aj,S f ln oeln,29 powerline, along N of, SE Kadji, Kadji ae orkn ok 31 Rock, Yokrakine male, 31 117 he eae,NrmenCm ie 32 Site, Camp Narembeen females, three argnRsre 32 Reserve, Narrogin elrernae,31 area, Kellerberrin h neirpcesaesprtdin separated are pockets anterior the hre hnta of in that (keeled than pedipalp shorter male the of apophysis simsoni Dingosa Diagnosis 118 aueRsre 32 Reserve, Nature ainlPr,24 Park, National rw,bocymria ad;wiestei einadsbagnlbns otherwise bands; dark submarginal bands; and submarginal median brown in diagonal light setae and distinct clypeus white PLE; to bands; and V; PLE marginal open PME behind blotchy an towards between from brown, form eyes setae fovea that between white from lines setae of longitudinal widening white line brown band of dark median line and central brown short narrow two light including setae; here white and PLE of lines radial white Male. Description 116 No ri rc) 31 track), train of (N et 30 West, 26 on tto,7 mNo aln–hrutnRa,GetVcoi eet 29 Desert, Victoria Great Road, Rawlina–Wharburton of N km 74 Station, Downs 125 30 WM7/9–) n eae ognHls AArclueDprmn eerhStation, Research Department 30 Agriculture WA Hills, Wongan female, one 70/192–3); (WAM 28 114 rsm,dra hed(iue A A:dr rw;idsic lc ailpattern; radial black indistinct brown; dark 2A): 1A, (Figures shield dorsal Prosoma, u u u u u 22 51 57 54 56 u u u u u u 39 37 31 12 43 30 9 9 9 9 9 ,117 S, 14 09 ,116 S, ,117 S, ae nWMT62462. WAM on Based 9 9 9 9 9 9 00 00 WM7/9) ormls eto alai,RsrorRa,32 Road, Reservoir Karlgarin, of West males, four 70/190); (WAM E 09 36 SMN171;oefml,ot uaigHl,32 Hill, Cunaring female,South one NN16781); (SAM E u 0 0 31 ,120 S, ,116 S, 0 0 0 0 WMT18) n ae tEvr,25k 160 km 2.5 Elvire, Mt male, one T51283); (WAM E WMT34) i ae,M are tto,26 Station, Narryer Mt males, six T53745); (WAM E WMT15) he ae,Ws eejr aueRsre 29 Reserve, Nature Perenjori West males, three T51257); (WAM E WMT16) n eaewt gsc iua mS elri turnoff, Yeelirrie S, km 3 Wiluna, eggsac, with female one T51268); (WAM E 9 41 u u u 58 35 43 0 ,117 S, u scoeyrltdto related closely is 9 9 u 9 WMT35) n eaewt gsc agoRa,8mls(oexact (no miles 8 Road, Yalgoo eggsac, with female one T53750); (WAM E 17 WM69/911). (WAM E u 39 WM7/8) n ae ygeHmsed 7k,125 km, 17 Homestead, Wydgee male, one 70/189); (WAM E 33 u 9 41 9 38 .serrata D. 29 9 05 u u 9 u 56 0 20 27 33 0 SMN172;tofmls ognHls 30 Hills, Wongan females, two NN16782); (SAM E u WMT15) n ae adtuaSain 24 Station, Mardathuna male, one T51254); (WAM E 0 u 06 ,114 S, 9 11 9 0 9 ,117 S, ,117 S, ,118 S, 54 9 9 u ,118 S, 49 25 0 WMT16) n ae aeNnnSieRsre East, Reserve, Shire Ninan Lake male, one T51260); (WAM E 0 n a itntadcniuu neirmri whereas margin anterior continuous and distinct a has and 9 u ,117 S, ,117 S, 57 u u 58 u 11 45 u 9 32 13 9 u 9 eiino h utainwl spider wolf Australian the of Revision 30 9 WM6/01;oeml,NrhDao Rocks Dragon North male, one 69/1011); (WAM E .serrata D. 38 WMT20) he ae,KneyRange Kennedy males, three T62304); (WAM E .serrata D. 0 9 WMT18) n ae u. aeMollerin, Lake juv., 4 male, one T51282); (WAM E WM7/42;oefml,Toii,32 Toolibin, female, one 71/1432); (WAM E u u 0 u 9 WMT16) n eae ubdig of W Numbudding, female, one T51263); (WAM E 32 38 31 ,117 S, u u 10 u 28 04 9 9 9 51 11 WM7/42;oefml,Yorkrakine, female, one 71/1422); (WAM E .serrata D. 9 9 u 18 40 9 13 ,118 S, 0 0 WMT32) n eae Premier female, one T53723); (WAM E ,118 S, u ifr ntecre i ftetegular the of tip curled the in differs but , u 0 0 . 29 9 ,116 S, ,114 S, 38 9 WM6/1,6/1) n female, one 69/914); 69/912, (WAM E 0 ,117 S, u u u 24 .Teeiyeof epigyne The ). 42 u 25 u u 27 23 04 9 9 WM6/0–0;oefemale, one 69/908–10); (WAM E 9 ,116 S, 39 9 9 ,118 S, 9 16 u 34 u 13 0 10 WMT16) n male, one T51264); (WAM E 0 0 WMT15) n male, one T51259); (WAM E 9 WMT18,T51287); T51284, (WAM E 08 9 u 24 u 13 25 u 0 54 WMT16) one T51267); (WAM E 0 u 9 u ,117 S, 9 WM6/3) WA, 69/833); (WAM E 35 56 ,116 S, 9 WM71/1425–7); (WAM E 9 u 9 ,115 S, ,117 S, .simsoni D. rm 29 from, igs 1613 Dingosa u u 39 u 48 54 9 u 9 u 32 55 E(WAM68/ 9 51 ,116 S, 0 9 9 E(WAM E(WAM u u u u E(WAM 16 32 27 25 smuch is u u u 9 9 9 9 u from, 40 57 00 52 49 43 43 0 9 0 0 9 0 9 S, S, S, S, S, S, E Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 1614 .073,5.32 4.20–7.35, .5 5.10 5.85, Variation. oee ihwieste u e rw ea aeal n napthcnrlyi rn of front in centrally patch a in and centrally; laterally brown. darker setae brown Spinnerets somewhat few furrow. brown, a epigastric light but brown Venter setae, dark white bands. carry with yellow-brown they covered as in darker setae somewhat olive-brown; white corners dark serrated setae; opisthosoma setae, of silvery sides with lateral band band; serrated serrated of laterally 2E). (Figure lines yellow-brown ventrally curled apophysis tegular of tip tip; pointed bristle long one AE, below bristles six eyes; around macrosetae AME. brown between few setae; black 4.88 two pairs, one ventral apicoretrolateral. apicoprolateral, Female. three one half, apicoprolateral, two metatarsus: one apical dorsal, retrolateral; apicoventral, in three two one dorsal retrolateral, femur: prolateral, one one two I: tibia: prolateral, pairs, leg retrolateral; ventral ventrally one of setae prolateral, three Spination scopulate one legs; I. other patella: leg than apicoretrolateral; darker of I leg metatarsi of on tarsi and metatarsi tibiae, banding; .8(.3,O .8(.5.Lg:lntso emns(femur Sternum segments (2.03). of OL 1.79 lengths Legs: PLE (0.82/0.76). (5.25). (1.61), 0.64/0.73 2.88 1.39 OW (length/width) PME (7.13), PLE Labium (1.33), 4.38 (0.45), 1.12 (2.13/1.88). 0.45 AE PME 2.13/1.88 (0.18), eyes: 0.15 (length/width) of ALE Row (0.24), 0.21 (0.41). AME 0.42 Eyes: (4.38). 4.00 PW (5.38), two pairs, ventral Measurements. one three apicoventral. apicoprolateral, one tibia: two pairs, retrolateral; retrolateral, ventral one two three metatarsus: dorsal, prolateral, prolateral; three one femur: patella: I: apicoretrolateral; leg of Spination darker. short. spermathecae continuous. margin anterior widening; brown. Spinnerets: tarsus I4.75 II 3.00 es e oml VI formula leg Legs: with light band; strong median serrated embolus laterally with tip, black to broad brown dark with 1A): (Figure Opisthosoma apophysis terminal D): white. 2C, setae. and white (Figure dense truncated with covered Pedipalps basally end discolorations; darker front few with brown; brown margins. light dark Chelicerae: towards wide; longer than are longer which Labium: setae white brown; Sternum: es e oml IV of formula stalks triangular; leg elongated Legs: spermathecae of heads 2F): end (Figure anterior view but dorsal setae. T-shaped, Epigyne, inverted white septum median with 2E): (Figure covered view yellow-brown, ventral Epigyne, Venter white and 1B). (Figure truncated end male front as brown; dark Opisthosoma: setae. long; white than long wider very male. Labium: with as covered chelicerae: yellow-brown; and Sternum: 1B), (Figure shield dorsal Prosoma, + 5 3.25 5 27 pdpl 1.56 (pedipalp 22.75 + oa egh:pdpl 1.56 pedipalp length): total ae nWMT73725. WAM on Based 5.63 .W rmnu&B .Baehr C. B. & Framenau W. V. + ¡ ae fmls rne mean (range, (females) Males 2.00 .2 W31–.5 3.80 3.15–4.35, PW 0.52; + 4.75 aeWMT26 fml A 775:T 02 1.3,P 5.13 PL (12.63), 10.25 TL T73725): WAM (female T62462 WAM Male 5 ¡ 41,II3.75 III 14.13, .4 W30–.5 3.86 3.00–5.25, PW 0.84; + 2.38 . . 5 I I 75,II4.50 III 17.50, . . + 2.38 II II . . + I;clrto sml u itlsget flgInot I leg of segments distal but male as coloration III; I;lgtbon eoawt oebonlongitudinal brown some with femora brown, light III; 4.38 + – + 1.88 + + 1.75 3.38 ¡ + ¡ 5 + 0.43; + – 5.13 2.00 D:T .01.5 9.68 7.80–12.75, TL SD): .9 4.38 I 5.69, ¡ + 1.88 0.58; + 5 n 4.63 5 35,I 5.00 IV 13.50, 5 T .01.0 11.59 8.40–15.00, (TL 9 .1 5.00 I 5.31, n + 5 2.50 23). + 5.25 5 67,I 5.88 IV 16.75, + 3.13 + + + patella/tibia 6.13 6.25 + 1.75 + + 5.25 4.63 ¡ 5 .5 L4.20– PL 1.55; 45,I 4.00 II 14.50, + + 2.50 + 2.25 + metatarsus ¡ 6.88 .5 PL 1.75; 5 5 18.88). + 18.13, 7.00 + + + Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 hti omni ot-atr utai nldn amna tapasta Hickman that appears of It female holotype Tasmania. the including examine Australia not did south-eastern (1967) in common is that oee,ti omnnm sbsdo h iietfcto fti pce yHickman by species this of misidentification the on re-described based who is (1967) name common this However, simsoni patchy Dingosa of consisted vegetation The Remarks bedrock. sandstone grasses. over introduced and shale native of of reefs remnants on soil loam McCullough soils. dry eroded or sandy on illustrated habitats (2000) vegetated sparsely open, for of preference shrubland ‘‘open bushland’’, ‘‘remnant erBra ot utai.Here, Australia. South Burra, near eu nTsai.Wlkne 13,p35 tesdi atclra lvtdcephalic elevated an particular in stressed 325) p (1837, (‘‘ region Walckenaer Tasmania. in genus acear 87 ae naml olce nTsai.Teseisi currently is housed species be this species to that a expected suggests be description was The original material could Walckenaer’s The Tasmania. communication). where personal MNHP, in Leguin, the (E.-A. in collected found be male not could a on as catalogued based 1837, spiders Walckenaer, wolf by Australian of illustrated genus and unnamed Spider an Wolf books of which as species Grey to spider undescribed Australia an The is popular south-eastern (1967) misidentification. from Hickman Some (1967) the Spider identify (1959). Hickman’s to Wolf continued perpetuating Roewer 2003) Grey Museum in trapdoor-building Australian 108; common epigyne p. 1970, Mascord the (e.g. webpages of illustration poor September. and August in as common found most been be have to eggsacs appear with Females but October. July, and as females, early for search in appear males simsoni Dingosa preferences habitat and history Life anadAsrlasuho 24 of south Australia Mainland Distribution detailed not is description support the However, undoubtedly spiders. to wolf enough Australian in uncommon very is which ads humphreysi Pardosa ooye ae aeCoi (32 Cronin Lake male, Holotype: examined Material 378. p 0Fbur 91 .F upry ta. A odilsSre ieLR7 mallee/ 7, LCR site Survey Goldfields WAM al., et Humphreys F. Triodia W. 1981, February 10 acear(87 ecie h is ofsie rmAustralia, from spider wolf first the described (1837) Walckenaer for descriptions Habitat uet,ptalta WM8/0) aaye:egtmls aaa ooye(WAM holotype as data males, eight Paratypes: 84/602). (WAM trap pitfall lunette, e uteyu e exlge poste lignes deux des yeux quatre Les yoaleucophaeoides Lycosa Dingosa oe dubium nomen swne-aue dl pdr a efudbtenArl hntefirst the when April, between found be can spiders Adult winter-mature. is scretyascae ihtecmo ae‘Ge ofSpider’’. Wolf ‘‘Grey name common the with associated currently is .simsoni D. igs humphreysi Dingosa otlikely most , ca 95,p1113 iue1–,e ,h lt ;Panc 1989, Platnick 1; Plate h, g, e, 1a–c, Figure 101–103, p 1985a, McKay yoasimsoni Lycosa .simsoni D. (as Panc 07 ae nRee 95) h yematerial type The 1955a). Roewer on based 2007; (Platnick L oh 87 lobelongs. also 1877) Koch, (L. Fgrs1,D AD 4) 3A–D, D, 1C, (Figures u yoairrorata Lycosa .simsoni D. ads serrata Pardosa aiueadTsai Fgr 4). (Figure Tasmania and latitude S u 22 eiino h utainwl spider wolf Australian the of Revision 9 .simsoni D. 15 salre rporbidn,broigwl spider wolf burrowing trapdoor-building, large, a as nld ‘addns’o ‘aeo addunes’’, sand of ‘‘base or dunes’’ ‘‘sand include 0 rerssn porte sont ´rieures ,119 S, MKy 95,n comb. n. 1985), (McKay, Acacia sti pce steol ersnaieo this of representative only the is species this as sasno yoy of synonym senior a as n eotdi omna i td site, study his at common it reported and ) u .simsoni D. 49 cuidsoy adpcigclay–sandy hard-packing stony, occupied ’ad‘mle’ r‘mla’sgetn a suggesting ‘‘mulga’’ or ‘‘mallee’’ and ’’ 9 30 0 ,WsenAsrla utai) 4– Australia), Australia, Western E, ssru le un sur ´s u ae i dniiaino a on identification his based but grrnlmn ucorselet du renflement ´ger .simsoni D. igs 1615 Dingosa yoairrorata Lycosa . Dingosa ’’), , Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 4637 MS0)eaie;1 ae,Lk rnn(32 Cronin Lake males, 15 examined; S409) QM 84/603–7, te aeilexamined. 84/649). (WAM material turret litter Other slack, dune 3, GGR site Survey Goldfieds WAM Humphreys, 135 S64935). utai) –0Fbur 91 .F upry ta. A odilsSre ieLCR site Survey Goldfields WAM al., mallee/ et Humphreys 8, F. W. 1981, February 4–10 Australia), 3k ,36 N, km 13 119 462;fu ae,PitSlain – mWW 28 WNW, km 7–8 Salvation, Point males, four 84/602); ak 35 Park, 1616 rw einbn ieigfo oe oad L n eeicuigaV-shaped a including here and PLE towards fovea from widening band median brown Male. in in evident as septum Description part median transverse The the epigyne. on the edges of truncated shape have in the not apically does in differ curved clearly is Females which apophysis, tegular 30 Reserve, Nature Springs Victoria Queen females, two males, odilsSre ieLR5(A 4632) n eae orbn(31 Boorabin female, one 84/623–26); (WAM 5 LCR site Survey Goldfields osrainPr,Pnao–odronRa,35 Road, Pinnaroo–Bordertown Park, Conservation ada ako h pshsm sls ertdta in than serrated less is opisthosoma the on mark place cardiac characters genitalic and Somatic Diagnosis T53114). T53111, T48802, T48734, T48629, T48586, T48559, T48555, T48546, 120 pce n ae ar ml ot ttebs ftetglraohsso h male the of apophysis tegular the of base the at tooth small of a Males carry pedipalp. males and species odilsSre ieBR2 et ihocsoa ale(A 467;oefemale, one 84/627); (WAM mallee occasional with heath (31 2, Boorabin BNR site Survey Goldfields N69) n ae gra osrainPr,Pnao–odronRa,35 Road, Pinnaroo–Bordertown Park, Conservation Ngarkat male, one NN16692); 140 l,WMGlfed uvyBR1, BNR Survey Goldfields WAM al., od mNo onay 35 boundary, S of N km 5 Road, ae,Gogri (29 Goongarrie males, osrainPr,bre rc,6k ir,35 Kirra, E km 6 track, border Park, Conservation twomales,fivefemales,PinkawillinieConservationPark,33 o,WMGlfed uvyst G ,dn ie ifl rp(A 4693) eight 84/629–31); (WAM trap pitfall site, dune (29 2, Goongarrie GGR males, site Survey Goldfields WAM How, ,N 66,NN16764). 16762, NN 9, o,WMGlfed uvyst G ,dn ae ifl rp(A 4629;nine 84/632–9); (WAM trap pitfall base, dune 3, (29 GGR Goongarrie males, site Survey Goldfields WAM How, o,WMGlfed uvyst G ,dn,ptalta WM8/2–8;one 84/629–48); (WAM trap pitfall dune, 1, (29 GGR site Goongarrie Survey female, Goldfields WAM How, rsm,dra hed(iue1) rw;idsic akbonrda atr;light pattern; radial brown dark indistinct brown; 1C): (Figure shield dorsal Prosoma, u u u u 57 48 45 13 ae nWMT48586. WAM on Based 9 9 9 9 SMN173 N64–,N179 N65) n eae elr Station, Keilira female, one NN16751); NN16749, NN16745–6, NN16743, (SAM E 00 00 ,WsenAsrla,2–9Mrh18,W .Hmhese l,WAM al., et Humphreys F. W. 1981, March 24–29 Australia), Western E, u etr Australia Western 47 .W rmnu&B .Baehr C. B. & Framenau W. V. Triodia 0 0 SMN162;oeml,NaktCnevto ak Pinnaroo–Bordertown Park, Conservation Ngarkat male, one NN16682); (SAM E ,WsenAsrla,41 eray18,W .Hmhese l,WAM al., et Humphreys F. W. 1981, February 4–10 Australia), Western E, u 9 ,140 S, 35 u 15 9 ,140 S, 9 ifl rp(A 4682) ormls aeCoi (32 Cronin Lake males, four 84/608–22); (WAM trap pitfall , ,120 S, .humphreysi D. u 36 u u u u 55 55 55 9 10 u SMN165 N68) n ae n eae Ngarkat female, one male, one NN16687); NN16685, (SAM E u 04 55 9 9 9 utai:SuhAustralia South Australia: 9 20 20 20 SMN168;oeml,oefml,NaktConservation Ngarkat female, one male, one NN16678); (SAM E 9 ,WsenAsrla,2–9Mrh18,W .Hmheset Humphreys F. W. 1981, March 24–29 Australia), Western E, 9 Victoria ,121 S, 0 0 0 n ae A aeCoi,32 Cronin, Lake WA, male, one : ,121 S, ,121 S, ,121 S, u 43 and 9 31 u u u u 08 07 07 07 n eae iteDsr,36 Desert, Little female, one : 0 ,140 S, .murata D. 9 9 9 9 .humphreysi D. Callitris ,WsenAsrla,1–1Jl 91 .F. W. 1981, July 15–21 Australia), Western E, 55 55 55 0 0 0 ,WsenAsrla,1–6Mrh17,R A. R. 1979, March 10–16 Australia), Western E, ,WsenAsrla,1–6Mrh17,R A. R. 1979, March 10–16 Australia), Western E, ,WsenAsrla,1–6Mrh17,R A. R. 1979, March 10–16 Australia), Western E, u 47 etioae ers(A 468;three 84/628); (WAM debris heat/isolate, 9 .humphreysi D. 48 u a edsigihdb h hp fthe of shape the by distinguished be can u 46 48 0 SMN168;oeml,Ngarkat male, one NN16688); (SAM E 9 16 9 ls to close u 08 12 iemls abde 33 Hambidge, males, five : 0 u ,140 S, 0 9 .simsoni D. 07 ,140 S, ,123 S, u 22 9 u S,136 22 9 .murata D. 30 u u 9 57 u o oin so not but u 48 15 0 36 u ,119 S, 9 14 u 9 0 40 9 00 .murata D. 00 ,119 S, u WMT27) 10 T62372); (WAM E and 33 9 0 ,123 S, 9 0 SMNN23324) (SAM E E(SAMNN16755– SMNN16683, (SAM E 9 u ,141 S, h longitudinal The . .serrata D. 49 u 49 9 35 .humphreysi D. u 9 41 . 30 0 u .murata D. ,Western E, 38 9 0 E(WAM u u E(WAM 23 42 9 E(QM nboth in u u 9 9 15 13 23 14 0 9 9 0 S, S, S, S, . Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 eaass he eta ar,topoaea,oertoaea,oeaioeta,one apicoventral, one Female. retrolateral, retrolateral; one two prolateral, prolateral, two two pairs, apicoretrolateral. pairs, ventral one retro- apicoprolateral, three ventral one half, apical three prolateral, in one metatarsus: dorsal furrow. patella: one epigastric apicoretrolateral; tibia: of one lateral; front retrolateral, in two centrally apicoprolateral, patch a in yellow; Venter setae tip setae. brown dark brown. of setae, Spinnerets curled edges dark white by ventrally with accentuated half covered and anterior in mark tooth cardiac brown basal with apophysis 3B). tegular (Figure sickle-shaped; embolus four brown eyes; AME. light around between macrosetae bristle median distinct long brown in eyes; one few setae AE, setae; between white below brown bands; setae bristles otherwise marginal bands; white brown submarginal dark of and narrow line indistinct bands; median submarginal narrow pattern; darker tibia short. spermathecae pockets. anterior covered half; the yellow-brown, under posterior reaching Venter in mark. brown. and cardiac light on mark Spinnerets: setae setae. cardiac brown white of few with lateral and setae patches silvery-white yellow-brown irregular serrated; .0.O .8(.5,O .3(.3.Lg:lntso emns(femur segments of lengths (2.19). Legs: 1.71 (5.63). PLE (1.71), 3.13 1.38 OW (0.90/ (0.58), PME 0.67/0.67 (6.75), 0.52 (1.44), (length/width) PME 5.38 1.17 (0.25), Labium OL AE 0.17 (2.38/2.13). eyes: 0.90). ALE 2.25/1.88 of (0.35), (length/width) Row 0.29 Sternum AME (0.52). Eyes: 0.44 (4.50). PLE one 4.00 PW prolateral, (6.25), two pairs, Measurements. ventral three apicoventral. apicoretrolateral; one one tibia: pairs, apicoprolateral, ventral retrolateral; two three dorsal, one metatarsus: three retrolateral; prolateral, femur: I: one leg patella: of Spination II. and 2.25 6.13 6.50 V6.88 IV es e oml VI formula setae; leg light Legs: of cover dense a through grey brownish light 1C): (Figure indented; and Opisthosoma broad tip its semicircular, apophysis basally. terminal setae B): white. white and 3A, few (Figure truncated frontally; Pedipalps end brown front light brown; brown, wide; Chelicerae: macrosetae. as brown long fewer as and Labium: setae white yellow-brown; Sternum: pgn,dra iw(iue3) ed fsemtea ihatro is tlsof stalks tips; anterior IV with formula leg spermathecae Legs: of heads ends 3D): lateral (Figure its view T-shaped, dorsal inverted Epigyne, septum median 3C): (Figure view ventral Epigyne, weakly mark cardiac longitudinal setae. brown brown few dark basally, olive-brown; setae 1D): white (Figure few Opisthosoma brown; distinct. reddish less dark male. bands Chelicerae: as submarginal labium: male, and as Sternum 1D): (Figure shield dorsal Prosoma, + 5 + + metatarsus 3.13 5.75 60,I 4.25 II 16.00, + ae nWM84/649. WAM on Based 8.13 5 + 2.88 20,I 5.25 II 22.00, + 7.63 aeWMT88 fml A 469:T 12 1.5,P 5.75 PL (13.25), 11.25 TL 84/649): WAM (female T48586 WAM Male 5 + tarsus 20.75). + 3.63 + 5.00 . 5 . 5 I oa egh:pdpl 2.50 pedipalp length): total I . + . 62 pdpl 2.38 (pedipalp 26.25 3.50 II + II 6.50 . . I;bon cplt ea nmttriadtrio e I leg of tarsi and metatarsi on setae scopulate brown; III; + I;bon pnto flgI eu:tredra,two dorsal, three femur: I: leg of Spination brown. III; 2.25 + 5.75 eiino h utainwl spider wolf Australian the of Revision 5 50,II4.25 III 15.00, + 3.00 5 05,II5.00 III 20.50, + 2.75 + + + 2.25 – 4.75 + 2.25 + + – 3.75 5 + + 2.13 .8 4.50 I 7.38, 6.25 + 2.13 5 + .8 5.63 I 6.88, 6.13 5 igs 1617 Dingosa 48,I 5.63 IV 14.88, + 3.00 + 5.63 + 5 patella/ + + 20.38, 3.63 7.13 + + + Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 etr utai,SuhAsrla nowsenVcoi Fgr 4). (Figure Victoria western into Australia, South Australia, Western Distribution simsoni D. ‘‘ also caring but and bases’’, Females collected. April been autumn. in not and peaking far summer July, so late and have in February brood activity for between reproductive found suggests been which March, mainly have spiders Adult preferences habitat and history Life ooye ae utop(27 Zuytdorp Male, Holotype: examined Material ta.(A 533.Prtps he eae,NrhTrnRc aueReserve Nature Rock Tarin North Harvey females, S. three M. ZU2, Paratypes: Survey T51303). Carnarvon (33 (WAM CALM/WAM trap, al. pitfall et wet 1995, May 16–21 4.87 4.35–5.70, te aeilexamined. material Other 1428–30). .5 4.88 5.85, etr Australia Western M 83) itra iteDsr ainlPr,c 36 ca Park, National Desert Little Victoria: K8132). (MV Variation. N15/2 WM6/1) n ae xlmto ae 32 Lake, Exclamation male, one 68/515); (WAM 1959/A24 FN ieS 2,2–9Arl20,R el,G aod .Snes .Hgs r ifl traps 29 pitfall Eneabba, dry Higgs, female, P. Sanders, one A. Harold, T51246); G. Teale, (WAM R. 2002, 1–6 April 23–29 02A, SG site ciln iest,RCmnrlsns(A 418;oefml,Gni,31 Gingin, female, one T42138); (WAM sands mineral RGC site, mine McMillan, f( fFed od,33 Road), Fields of (E of 115 coe 99t oebr20,P a erke l,wtptaltas(A T51258); (WAM traps pitfall wet 31 al., et Airport, Heurck Perth van male, P. 2000, one November 1 to 1999 October 4Jn o2 uy19,J .Wloke l,wtptaltas(A 600;tomales, two T68030); (WAM traps pitfall wet al., et Waldock M. 27 J. 1993, Zuytdorp, July 28 to June 24 o1 a 95 .S avye l,wtptaltas A,T10;fu males, four T51302; WAM, traps, pitfall wet al., et Harvey S. M. 27 1995, Zuytdorp, May 18 to o1 a 95 .S avye l,wtptaltas(A 534;fu ae,same males, 27 four Zuytdorp, T51304); male, (WAM one T51309); traps (WAM pitfall 1995 wet August al., 16 et to May Harvey 18 S. except M. data 1995, May 18 to 1618 avye l,wtptaltas(A 537;tomls aedt xet1–1May 16–21 except data same males, two T51307); (WAM T51308). traps (WAM pitfall 1995 wet al., et Harvey 114 aia ecitosfrti pce nld ‘uesrbad’ ‘ue’ n ‘‘dune and ‘‘dunes’’ shrubland’’, ‘‘dune include species this for descriptions Habitat u u u 06 55 11 9 9 9 ,118 S, ,Jl 94 .Lnge,sn li WM7/43;tomls aeKn,SE King, Lake males, two 71/1433); (WAM plain sand Lindgren, E. 1954, July E, 16 .W rmnu&B .Baehr C. B. & Framenau W. V. . 0 ¡ ae fmls rne mean (range, (females) Males ,CL/A anro uvy U,1 aur o1 a 95 .S. M. 1995, May 17 to January 10 ZU5, Survey, Carnarvon CALM/WAM E, .6 W28–.5 3.45 2.85–4.05, PW 0.56; u u u 15 15 13 ¡ 9 9 41 28 Trioida 9 .9 W30–.0 3.46 3.00–4.20, PW 0.49; ,WsenAsrla utai) 3My17,A ans(A 71/ (WAM Baynes A. 1971, May 23 Australia), Australia, Western E, 0 0 n eae taae 32 Attadale, female, one : ,114 S, ,114 S, u rsln’ n ‘ale’sgetn iia aia rfrne as preferences habitat similar suggesting ‘‘mallee’’ and grassland’’ 06 Australia: u u 9 u 01 09 46 58 igs murata Dingosa 9 9 0 9 Fgrs1,F AD 7) 5A–D, F, 1E, (Figures 48 02 ,121 S, 34 u 16 0 0 0 ,CL/A anro uvy U,1 January 11 ZU2, Survey, Carnarvon CALM/WAM E, ,CL/A anro uvy U,1 January 10 ZU4, Survey, Carnarvon CALM/WAM E, ,115 S, 9 41 e ot ae:Dnlqi,22 Deniliquin, Wales: South New u 11 0 ¡ ,114 S, 9 ¡ u 0.36; 35 58 D:T .51.0 8.94 7.35–10.50, TL SD): ¡ 0 9 ,CL aiiyAto ln ieG ,15 6, GP site Plan, Action Salinity CALM E, 25 u 0.43; 01 u n 01 0 .sp. n. 5 ,WMUbnBsln uvy PA7, Survey, Bushland Urban WAM E, 9 u ,115 S, 56 9 2(L84–20,10.15 8.40–12.00, (TL 12 48 n 9 5 ,115 S, 0 ,WsenAsrla Australia), Australia, Western E, 9). u 48 u 33 9 u ,5My15,A .Main, R. A. 1959, May 5 E, 17 0 ,141 S, 9 ,1 a 91 .P. R. 1991, May 15 E, u 47 u 9 18 50 ¡ 0 0 u ,121 S, M K8113). (MV E .6 L4.05– PL 0.96; 22 0 ,121 S, ¡ u u .6 PL 1.26; 24 15 9 u 9 u 34 21 25 18 0 9 0 0 E, S, S, E Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 af akbonaon oe;nro eta ieo ht ea ewe ysfo behind from eyes between setae white anterior of in line bands central short narrow irregular fovea; two around with brown and dark anteriorly half; widening band median brown light Male. Description in epigyne the in of truncated pockets are which anterior septum, median T-shaped inverted in h eua ppyi,wihi uvdvnrlyin ventrally curved is which apophysis, tegular the h pcfcnm sa detv napsto eie from derived apposition in adjective an is name specific The Etymology of Records 7. Figure igs murata Dingosa species other all and Diagnosis species this of behaviour turret-building the genus to the refers within and wall) a by lgtsurs 7rcrs.Tesnyesre of series syntype The records). 17 squares; (light rsm,dra hed(iue1) akbon nitntdr rw ailpattern; radial brown dark indistinct brown; dark 1E): (Figure shield dorsal Prosoma, .murata D. ae nWMT51303. WAM on Based eae fbt pce r itnuse ytesaeo h aea iso the of tips lateral the of shape the by distinguished are species both of Females . sms iia to similar most is Dingosa Dingosa nAustralia. in . .humphreysi. D. .humphreysi D. igs serrata Dingosa eiino h utainwl spider wolf Australian the of Revision .serrata D. L oh fl ice;14records); 184 circles; (full Koch) (L. rmSde sntmpe ic h aeili lost. is material the since mapped not is Sydney from ae ifri h hp fteaia i of tip apical the of shape the in differ Males . .humphreysi D. .murata D. muratus n oeo esstraight less or more and u ec ne the under reach but , (Latin—surrounded igs 1619 Dingosa .murata D. p n. sp. Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 1620 .857,4.89 3.88–5.75, akste elwbonbnsltrlo ada ak elwbonirglrsosin brown. spots light Spinnerets irregular straight setae. yellow-brown brown less mark; with by or cardiac covered laterally brown; of accentuated more Venter serrations, lateral half. lateral tip bands posterior minor yellow-brown its two setae; with and mark dark cardiac tooth brown dark basal setae; with apophysis tegular 5B). tip; (Figure bent with setae. black few otherwise long one AE, below bristles four PLE; bands; marginal behind submarginal brown and and dark AME. median eyes in between narrow around bristle setae bands; macrosetae white setae, submarginal brown white few brown interspersed light and black distinct bands; clypeus; to PLE e) w erltrl eaass he eta ar,topoaea,oertoaea,one Female. retrolateral, one prolateral, apicoretrolateral. two one right one pairs, on prolateral, apicoprolateral, (three ventral one prolateral one three two apicoventral, patella: metatarsus: pairs, ventral retrolateral; retrolateral; three two half, one apical leg), in apicoprolateral, dorsal two one tibia: dorsal, retrolateral; three femur: I: tlso prahceS-shaped. spermathecae of stalks male. as Spinnerets lighter. slightly male, .0 4.34 6.00, 3.38 .8(.0,O .8(.3.Lg:lntso emns(femur Sternum segments (1.67). of OL 1.44 lengths Legs: PLE (0.58/0.69). (3.63). (1.25), 0.60/0.52 2.38 1.11 OW (length/width) PME (5.00), PLE Labium (1.06), 3.38 (0.42), 0.90 (1.63/1.50). 0.42 AE PME 1.63/1.50 (0.15), eyes: 0.13 (length/width) of ALE two Row (0.25), 0.21 (0.40). dorsal, AME 0.35 Eyes: (3.19). three 3.00 PW femur: (4.25), I: leg one Measurements. prolateral; of one apicoventral; Spination one pairs, I. ventral ventral three leg three apicoprolateral. tibia: metatarsus: retrolateral; of prolateral; one two prolateral, tarsi pairs, one patella: and retrolateral; two metatarsi apicoprolateral, on setae Variation. tarsus I3.38 II 2.38 es e oml VI formula leg Legs: white and silvery of cover dense a through grey brownish light 1E): (Figure Opisthosoma thin, embolus tips, two with broad, apically apophysis terminal basally, B): setae 5A, (Figure white truncated. Pedipalps few end frontally; front band brown; longitudinal brown otherwise light brown, brown, dark dark marginally. basally Chelicerae: denser wide; are than which longer setae Labium: white brown; shiny Sternum: pgn,dra iw(iue5) ed fsemtea ihdsic neirtips; anterior distinct with IV spermathecae formula leg of Legs: anteriorly. heads truncated 5D): septum (Figure median view of as edges dorsal Venter lateral contrast. Epigyne, 5C): less (Figure view with ventral and Epigyne, lighter overall but male, as 1F): male. (Figure distinct. as Opisthosoma more labium: and wider and bands dark chelicerae, marginal Sternum, male, as 1F): (Figure shield dorsal Prosoma, + 5 2.31 5 63 pdpl 1.61 (pedipalp 16.30 + oa egh:pdpl 1.54 pedipalp length): total ae nWM71/1428. WAM on Based 3.92 .W rmnu&B .Baehr C. B. & Framenau W. V. + ¡ ae fmls rne mean (range, (females) Males 1.54 .2 W26–.5 3.16 2.63–4.25, PW 0.72; + 3.15 aeWMT10 fml A 112) L70 90) L3.88 PL (9.00), 7.00 TL 71/1428): WAM (female T51303 WAM Male 5 ¡ 02,II2.61 III 10.23, .6 W28–.3 3.51 2.88–4.13, PW 0.66; + 1.92 . . I 5 I . . 23,II3.23 III 12.38, II + II 1.69 . . I;dr rw ihirglrlgtpths pnto fleg of Spination patches. light irregular with brown dark III; I;dr rw ihirglrlgtpths ekscopulate weak patches; light irregular with brown dark III; + + 3.15 – + + 1.38 + 1.38 2.54 ¡ + ¡ 0.51; 5 + + – 3.61 D:T .81.5 8.41 6.88–12.75, TL SD): .9 3.23 I 4.69, ¡ 1.46 + 1.54 0.45; + n 5 5 3.46 5 .7 V3.69 IV 9.77, 6(L76–27,10.68 7.63–12.75, (TL 16 n .6 3.61 I 4.46, + 5 1.85 10). + 3.54 5 21,I 4.31 IV 12.15, + 2.46 + + + patella/tibia 5.54 4.23 + 1.54 + + 3.92 3.23 ¡ 5 .1 L3.63– PL 1.61; 07,I 3.00 II 10.77, + + 1.92 + 1.92 + metatarsus 4.69 ¡ .6 PL 1.56; 5 5 14.07). + 13.00, 4.92 + + + Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 (33 of Syntypes examined Material 500. p 1993, Platnick 383; p 1989, Platnick 85; p in established (synonymy 322 p 1992, Moritz 378; p 1973, 1979). McKay McKay 3407; p 1958, 142. Bonnet p 1976, Main 2664; p 1957, Bonnet 272; p South New southern and Victoria 7). western (Figure and Wales Australia, Western from records Disjunctive Distribution other been of has preferences eggsac habitat with with T53903). agreeing May, (WAM female in date one peak and Only a data with locality autumn. November exact in and without activity April collected, reproductive between found suggests been which mainly have spiders Adult preferences habitat and history Life 8k Wo arnea 29 Narrandera, of SW km 88 examined. material Other female, 11118); (ZMB 1905 24360). June (MHNP of 1 (32 114, data Station Syntypes same Fremantle Hartmeyer, R. near 2005). Michaelson, W. Hill, Framenau Australia), Buckland lost; female, considered Simon: Collection (Bradley akRne tto,2k ,34 W, km 2 Station, Ranger Park otn tto,Coog 36 Coorong, Station, Cortina ihega,BamnRa ertroft hreil nCnmlaBlo Road, Cunamulla–Bollon on Charleville to turnoff near Road 26 Boatman eggsac, with coadCnevto ak 35 Park, Conservation McDonald 151 gsc rn ae 31 Lake, Frans eggsac, ak 34 Park, 37 138 SMN174 N64–) n eae u. imsWl,M eceConservation Rescue Mt Well, 35 Jimmys juv., 1 Park, female, one NN16747–8); NN16744, (SAM osrainPr,35 Park, Conservation gsc elr tto,1 mN 36 N, km 13 Station, Keilira eggsac, ads serrata Pardosa serrata Geolycosa praevelox Pardosa serrata Lycosa 71/1433), (WAM plain’’ ‘‘sand is species this for available description habitat only The u u u 39 24 u u 53 07 46 9 9 ,140 S, ,146 S, 9 9 9 ,151 S, Q S64941). (QM E SMN1761) w ae,oefml,Hmig,33 Hambidge, female, one males, two NN16706–10); (SAM E u u 59 51 9 9 29 u yoaserrata Lycosa ,140 S, u 13 14 u 0 13 ,140 S, 9 .Kc 87 3–3,Pae8,Fgrs5 a ,6;Ribw1911, Rainbow 6a; 6, 5a, 5, Figures 80, Plate 930–932, p 1877, Koch L. 9 SMN173;tomls w eae,3jv,Cfi a National Bay Coffin juv., 3 females, two males, two NN16723); (SAM E L oh:MKy17,p2529 iue af ad ca 1985b, McKay 2a–d; 1a–f, Figures 225–229, p 1979, McKay Koch): (L. Q 699;tofmls aeBodae erDly 27 Dalby, near Broadwater Lake females, two S64939); (QM E 9 ,NwSuhWls utai) rde olcin o examined Not Collection. Bradley Australia), Wales, South New E, L oh:Ree 95,p23 ca 93 380. p 1973, McKay 243; p 1955a, Roewer Koch): (L. io 99 9–9;Ribw11,p26 owr15a 185; p 1955a, Roewer 276; p 1911, Rainbow 191–192; p 1909, Simon u 18 u u u 51 28 50 9 SMN163;tremls imsWl,M Rescue Mt Well, Jimmys males, three NN16673); (SAM E igs serrata Dingosa utai:NwSuhWales South New Australia: 9 9 9 ot Australia South ,140 S, 22 ,134 S, .Kc:ukonnme fmlsadfmls Sydney females, and males of number unknown Koch: L. 0 u SMN1678;oeml,CnnaNtoa Park, National Canunda male, one NN16697–8); (SAM E 08 Fgrs1,H AD 7) 6A–D, H, 1G, (Figures u 17 u 9 u ,150 S, 18 50 9 u u u ,139 S, 13 eiino h utainwl spider wolf Australian the of Revision 35 9 31 9 SMN1646;tremls n eaewith female one males, three NN16674–6); (SAM E SMN172;fv eae,Gow,35 Goolwa, females, five NN16742); (SAM E 9 9 9 ,139 S, ,140 S, ,135 S, u 02 Dingosa u L oh,n comb. n. Koch), (L. 50 n ae n eae ilatConservation Billiatt female, one male, one : 9 NVK7614). (NMV E u u 9 10 u 08 SMN162;oeml,Ferries male, one NN16672); (SAM E 19 9 9 SMN167 N67–1;one NN16679–81); NN16677, (SAM E 9 SMN171;oefml with female one NN16701); (SAM E species. SMN1784) n female, one NN16738–41); (SAM E u 01 9 ,115 S, n eae eelHighway, Newell female, one : u Queensland 45 9 ,WsenAustralia, Western E, igs 1621 Dingosa ads praevelox Pardosa u 23 9 n female one : ,135 S, u u u 21 30 57 9 9 9 S, S, E Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 34 089;oeml,Blii,77Blii od 32 Road, Baldivis 707 Baldivis, male, one 1038–9); 31 Ranges, Gawler Sands, Kokatha female, ole 33 Collie, ainlPr,25 Park, National WM7/9–) n ae aknRa,32 Road, Darkin male, one 70/195–6); (WAM ae ag,3 rso a,31 Way, Preston 31 Balga, male, ainlPr,25 Park, National 139 aiog 35 Malinong, 31 35 35 ae,Dri od f(rvt ad,32 land), (private of E Road, Darkin males, 522;oeml,Betod ihRa,32 Road, High Brentwood, male, one T51252); e n xc oain WM6/6;oefml,Genuh uyBah 28 Beach, Lucy Greenough, female, one 69/86); (WAM location) exact (no peg 115 N fYaet 35 Yaapeet, of WNW 133 eaewt gsc ulbok 31 Bullsbrook, eggsac, with female eae alntn 31 Darlington, female, od 35 Road, 140 26 aet eev) 33 Reserve), Magenta 114 onr a1 mWWYaet 36 Yaapeet, WSW km 15 ca Corner, ak inroBretw od mNo onay 35 boundary, S of N km 5 Road, Pinnaroo–Bordertown Park, uir ubaSain 24 Station, Quobba Cuvier, 1622 eet 36 Desert, N69–) n eae eoaNtv oetRsre 37 Reserve, Forest Native Penola female, one NN16690–1); 36 N62) n eae eoaNtv oetRsre erWeg,37 edge, W near Reserve, Forest Native Penola female, one NN16724); 113 WMT14) n eae as nrne 26 Entrance, False female, one T51245); (WAM 24 Station, Quobba Cuvier, WMT37,T38–,T38–4 589;fu ae,2 eae,2juv., 2 females, 24 males, 1965) four Lane see T53899); collection; Lane T53887–94, J. 32 W. T53881–3, Attadale, region, T53879, Perth locality, (WAM exact (no Australia Western Australia Western SMN175;oefml,Pn ilae,So rnBrn 33 Baron, Iron of S area, Hill Pine female, one NN16725); (SAM eae izeadRvrae n xc oain WM7/0) n ae Forrestfield, male, one 70/207); (WAM location) 32 exact Park, (no Hartfield area River Fitzgerald female, 0–,7/1) w eae,Bdeu wm,Wneo,31 Wanneroo, Swamp, Badgerup females, two 71/811); 401–2, SMN172;trefmls u. ikwlii osrainPr,33 Park, Conservation Pinkawillinie juv., 1 females, three NN16752); (SAM uir ubaSain 24 Station, Quobba Cuvier, 136 ec,Abn,a lf re,34 Creek, Bluff at Albany, Beach, u u u u u u 45 57 05 24 31 33 u u u u u u u 43 38 28 36 42 29 00 9 9 9 9 9 9 ,135 S, ,115 S, ,117 S, ,138 S, 39 ,141 S, 9 9 9 9 9 9 9 SMN1701 N66) n ae otLnonCfi a area, Bay Lincoln/Coffin Port male, one NN16763); NN16760–1, (SAM E SMN175;oeml,Ymar okoe mW 31 W, km 1 Rockhole, Yumbarra male, one NN16705); (SAM E WMT33) n eaewt gsc ra otenHgwy 1 mile 214 Highway, Northern Great eggsac, with female one T53436); (WAM E 20 SMN166;oeml,NaktCnevto ak Pinnaroo–Bordertown Park, Conservation Ngarkat male, one NN16686); (SAM E WMT33) n ae resilsOk 7mlsEo igu (Lake Pingrup of E miles 17 Oak, Greenshields male, one T53637); (WAM E 57 0 .W rmnu&B .Baehr C. B. & Framenau W. V. u ,113 S, u u 0 0 42 22 SMNN16769). (SAM E WMT19–) n ae xlmto ae 32 Lake, Exclamation male, one T51295–7); (WAM E 33 u u 01 9 u 9 u u 9 14 38 u u ,116 S, 58 ,141 S, 46 29 40 56 9 ,115 S, 0 9 9 9 9 ,140 S, Q 690;oefml,Mle,36 Mallee, female, one S64940); (QM E 9 SMN173;oefml,NaktCnevto ak 35 Park, Conservation Ngarkat female, one NN16703); (SAM E 9 u WMT86) n ae orbn 31 Boorabin, male, one T58362); (WAM E ,139 S, SMN175;oefml ihega,Wnra 35 Wynarka, eggsac, with female one NN16765); (SAM E WMT50) n eae w gsc,Glrp 33 Gelorup, eggsacs, two female, one T55309); (WAM E 31 u u u 58 00 52 1fmls orfmlswt gsc n eaewt spiderlings, with female one eggsac, with females four females, 11 : u u 9 36 09 50 9 u 9 9 34 00 40 u 31 u 48 u 30 9 0 9 u u WMT19–) 7mls dlLn,26 Land, Edel males, 27 T51298–9); (WAM E WM7/41;tofmls otso,31 Cottesloe, females, two 71/1431); (WAM E 48 NVK14;oefml,Ltl eet ooaRa,Nihil, Road, Goroka Desert, Little female, one K8134); (NMV E 0 0 9 30 0 55 ,113 S, ,115 S, ,141 S, 9 ,113 S, 9 WM6/011,6/015,6/9 936–8B 69/ 69/376B–384B, 69/99, 68/1051–56, 68/1001–11, (WAM E ,118 S, 9 u 9 00 9 SMN172;oefml ihega,Mln,near, Milang, eggsac, with female one NN16702); (SAM E 11 ,116 S, u u 13 13 0 9 SMN164;tomls gra Conservation Ngarkat males, two NN16684); (SAM E 35 u u 9 9 u u u 55 22 27 u 59 34 55 Victoria 33 52 u 0 ,113 S, 53 9 0 0 9 9 9 9 u NVK45;tofmls aeAbcta NW Albacutja, Lake females, two K7435); (NMV E ,113 S, ,113 S, u 43 9 16 ,118 S, u 01 04 40 ,115 S, 9 04 WM7/41;oeml,GniiNature Gunyidi male, one 71/1421); (WAM E 0 0 0 WMT55) he ae,Faci Peron Francois males, three T55554); (WAM E 9 9 WMT19) n ae rnha Bay, Frenchman male, one T51294); (WAM E WMT19) 2mls rnosPeron Francois males, 12 T51293); (WAM E 9 WMT38) i ae,Ee Land, Edel males, six T53785); (WAM E ,115 S, ,141 S, u u 27 ee eae,Lk lauj ak 5km 15 Park, Albacutja Lake females, seven : 25 u u u u u 25 30 27 08 51 9 u 9 20 07 ,135 S, 9 9 9 9 WM6/1) n eaewt eggsac, with female one 68/516); (WAM E 9 12 41 u 26 u WM7/2) n ae odPark, Bold male, one 73/221); (WAM E 0 59 55 9 WMT19) n eae Cheyne female, one T51292); (WAM E u 40 23 u 0 0 0 u WMT19) n ae Cape male, one T51291); (WAM E WMT10) i ae,Cape males, six T51300); (WAM E 03 9 ,116 S, 9 18 WM6/1) n ae Cape male, one 69/916); (WAM E NVK63;oefml,Little female, one K7613); (NMV E 0 9 u ,116 S, ,113 S, 9 25 9 ,115 S, ,115 S, 9 u SMN171;oefemale, one NN16731); (SAM E 10 u 31 u u u 30 9 19 45 ,146 S, u 9 u u 45 51 50 47 9 9 9 19 SMN170;one NN16780); (SAM E 31 u 9 9 u 9 14 9 WMT14) one T51247); (WAM E WM6/7) one 69/672); (WAM E WMT07) one T60074); (WAM E 28 ,115 S, 0 u 0 WMT86) six T58462); (WAM E 47 ,140 S, u u 9 9 54 ,120 S, 21 ,140 S, 9 18 9 9 NVK8127). (NMV E 25 u u u 0 50 u 59 28 ,121 S, 0 47 ,137 S, u 9 u 9 19 9 WM69/ (WAM E ,115 S, 51 ,140 S, 9 48 9 9 (WAM E u u (SAM E 0 u 46 31 (SAM E 24 u 4 u u u u u 9 9 u u 9 9 07 19 23 47 54 44 07 45 34 51 11 9 9 0 9 9 0 9 9 0 9 0 S, S, S, S, S, S, S, E E E E Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 eev,Mdad od 30 Road, Midlands Reserve, 34 eev,Nma etRa,30 Road, West Namban Reserve, 27 eae u. adkt 32 Jandakot, juv., 1 female, u. ot fFtgrl n uet iesjt,34 jct., Rivers Susetta and Fitzgerald of North juv., ignlln fwiestebtenPEadPE itntlgtbonsubmarginal brown light distinct PLE; and PME and between eyes between setae setae white white of dark of line and median short line narrow two anteriorly; diagonal PLE; widening behind band lines median longitudinal brown light brown setae; white of lines radial Male. Description within species other all fteml eiapwihi eldaothlwyadhsabodtp(one i curled tip (pointed tip broad a has and halfway about keeled in is ventrally which pedipalp male the of serrata Dingosa Diagnosis 32 ae,1jv,Nrhmtn f i od 28 Road, Tip of, S Northampton, juv., 1 males, 31 ae anr,32 Warnbro, male, 31 scrub, Wanneroo males, two 71/1423–4); (WAM ihod 32 Richmond, 32 Parmelia, male, ae n eae n eaewt gsc ace,31 Yanchep, eggsac, with female one female, one male, WM6/8) n eae ognHlsrgo,30 region, Hills Wongan female, one 69/882); (WAM aueRsre 30 Reserve, Nature 487 483 483 489 486 496 495 592 T53057, T52912, Hill, T48817, T48985, Reabold male, T48810, T48956, one T53115); 31 T48741, T53112, T48807, T48896, Pert, T53093, T53089, T48801, T48709,T48713, T48889, T53085, T53081, T48792, T48883, T48705, T53061, T48785, T48873, T48696, T48594, T48780, T48867, T48583, T48693, T48777, T48577, T48485, T48689, T48554, T48747, T48482, T48549, T48619, T48478, T48545, T48474, T48611, T48536, T48470, T48532, T48467, T48501–2, T48464, T48461, T48395–6, WMT15,T34) w ae,Mnee–ulw od 28 Road, Mingenew–Mullewa males, two T53541); T51559, (WAM 531;oefml,Yr,7mlsWo,o okPrhRa,31 Road, York–Perth on of, W miles 7 York, female, one T58321); 32 888;svnmls n eae utop 27 Zuytdorp, female, one males, seven 68/858); WMT16) n ae oly 31 Morley, male, one T51262); (WAM sad 32 Island, trigRa,31 Road, Sterling n ae utop 27 Zuytdorp, male, one 30 aps 32 campus, rsm,dra hed(iue1) akbon nitntbakrda atr;white pattern; radial black indistinct brown; dark 1G): (Figure shield dorsal Prosoma, u u u u u u 18 15 06 21 02 rm 31 from, 9 9 9 9 9 55 25 ,115 S, 21 22 ae nWMT62462. WAM on Based u 56 0 0 0 0 ,118 S, ,114 S, ,116 S, ,115 S, u 9 00 .simsoni D. u ,115 S, u 04 u 47 9 52 ,115 S, sms iia to similar most is u 9 9 17 ,115 S, u u ,115 S, 9 u u 45 11 WMT75) w ae,1jv,JlmrCnevto ak North, Park, Conservation Julimar juv., 1 males, two T47250); (WAM E 11 u 13 9 59 u u ,115 S, 20 46 9 9 u 9 9 39 16 15 14 04 u 9 u .Tefml pgn scnieal ogrta iei otatto contrast in wide than longer considerably is epigyne female The ). 27 14 9 9 30 u ,115 S, WM7/5) ormls n eaewt gsc Rossmoyne, eggsac, with female one males, four 73/151); (WAM E 0 0 9 0 u 0 44 WM6/5,6/34 981 114) n eae Rottnest female, one 71/1847); 69/881, 69/83–4, 68/857, (WAM E WMT51306). (WAM E Dingosa ,115 S, WMT69) n ae u. rg,31 Trigg, juv., 1 male, one T56398); (WAM E WMT16) he eae,Lea,29 Leeman, females, three T51265); (WAM E 49 0 9 ,117 S, 9 ,123 S, u WM6/5) w ae,Siln ag aaa Park, Caravan Range Stirling males, two 68/859); (WAM E 9 15 u WMT36) n ae he eae,MrohUniversity Murdoch females, three male, one T53464); (WAM E 9 43 WMT15,T27) n eae abnNature Namban female, one T62470); T51250, (WAM E 9 u 42 u u 9 43 11 WM7/9) 2 ae,QenVcoi Springs Victoria Queen males, 127 70/197); (WAM E 07 u u 0 47 nwihi sgnrlywdrta long. than wider generally is it which in u 9 24 .simsoni D. ,114 S, 9 u WMT14) n ae isn 32 Wilson, male, one T51244); (WAM E 9 41 10 22 ,115 S, 9 9 WMT52) n eae on eo,Lake Peron, Point female, one T55429); (WAM E 19 9 0 eiino h utainwl spider wolf Australian the of Revision 9 WMT88,T88,T88–,T48393, T48388–9, T48386, T48384, (WAM E ,116 S, 17 0 u WMT16) he ae,Mlao,4k at km 4 Mullaloo, males, three T51261); (WAM E u 53 0 09 ,115 S, u 50 9 u ifr ntesaeo h eua apophysis tegular the of shape the in differs but , ,115 S, 9 09 u 9 01 WMT25) n eae Jandakot, female, one T62452); (WAM E u 0 31 u WMT10) n ae Zuytdorp, male, one T51301); (WAM E 9 59 51 u u 9 15 54 57 9 0 u u 12 WMT15) he ae,one males, three T51255); (WAM E 01 45 9 u 9 0 28 53 ,114 S, WMT36) n ae Mount male, one T53660); (WAM E 0 WMT15) he eae,3 females, three T51256); (WAM E 9 9 u ,119 S, ,115 S, 0 33 9 ,114 S, ,116 S, 9 ,115 S, u 44 u u 48 27 u u 9 40 09 18 9 9 u WM6/3–) one 69/831–2); (WAM E WM7/–0;four 71/5–10); (WAM E 41 9 9 0 u WM9/90;one 94/1910); (WAM E 02 WMT16) one T51266); (WAM E 53 u 55 9 WMT42157–8, (WAM E 0 9 WMT51305); (WAM E ,116 S, 9 06 igs 1623 Dingosa u u u 0 56 53 02 ,115 S, u 9 9 9 ,114 S, 39 ,115 S, ,115 S, 9 (WAM E u 25 u u u 9 45 55 58 49 9 9 9 0 E E E E Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 1624 .073,6.12 4.80–7.35, ht ea nylo-rw ad.Vne elwbon oee ihwieste few brown. setae, setae; light white Spinnerets brown with furrow. dark covered epigastric carry of yellow-brown; they front Venter as in bands. darker setae yellow-brown brown corners in serrated setae setae, olive-brown; dark white silvery opisthosoma of with sides band lateral band; serrated serrated of laterally lines yellow-brown 6B). (Figure broad apophysis tegular of tip tip; long one AE, below bristles four bands; eyes; AME. submarginal around between and macrosetae bristle median brown in few setae setae; white black bands; otherwise marginal blotchy brown, dark bands; w rltrl w erltrl eaass he eta ar,topoaea,two prolateral, two pairs, ventral Female. apicoretrolateral. three pairs, one ventral metatarsus: apicoprolateral, three one half, retrolateral; femur: apicoventral, apicoretrolateral; apical one I: in one two retrolateral, leg dorsal retrolateral, of one prolateral, tibia: Spination two retrolateral; II. two apicoprolateral, one and two prolateral, I one prolateral; leg patella: of one metatarsi dorsal, on three ventrally setae scopulate banding; prahceshort. spermathecae indistinct. septum brown. light .8 5.56 6.88, 4.13 .5(.0,O .3(.5.Lg:lntso emns(femur Sternum segments (2.18). of OL 1.64 lengths Legs: PLE (0.91/0.94). (6.75). (1.70), 0.94/0.91 4.13 1.52 OW (length/width) PME (8.50), PLE Labium (1.48), 6.25 (0.61), 1.36 (2.50/2.38). 0.48 AE PME 2.75/2.38 (0.21), eyes: 0.24 (length/width) of ALE Row (0.27), 0.30 (0.52). AME 0.45 Eyes: (5.00). 5.25 PW (6.88), three Measurements. metatarsus: prolateral; two pairs, patella: apicoventral. ventral apicoretrolateral; one three one pairs, tibia: retrolateral, ventral retrolateral; two one apicoprolateral, prolateral, two one dorsal, three femur: I: tarsus Variation. I6.00 II 02 pdpl 2.88 (pedipalp 30.25 es e oml VI formula leg Legs: light band; median serrated laterally with black to brown dark 1G): (Figure pointed Opisthosoma with sickle-shaped embolus broad, apophysis setae. terminal white B): white. dense 6A, (Figure and with Pedipalps truncated covered end brown; macrosetae. front light brown brown; Chelicerae: long wide; 10 than ca longer and Labium: setae white yellow-brown; Sternum: pgn,dra iw(iue6) ed fsemtea lnae;sak of stalks elongated; spermathecae of heads IV formula leg 6D): (Figure Legs: view dorsal median separated, pockets Epigyne, anterior wide, than longer Spinnerets 6C): setae. (Figure view white ventral with Epigyne, covered yellow, white. Venter and 1H). truncated (Figure end male front as male. brown; Opisthosoma as wide; setae than pigmentation; longer greyish Labium: with centrally male. yellow-brown, as Sternum: 1H): (Figure shield dorsal Prosoma, + 2.38 5 + 7.63 oa egh:pdpl 3.00 pedipalp length): total ae nWMT73725. WAM on Based .W rmnu&B .Baehr C. B. & Framenau W. V. 5 ¡ ae fmls rne mean (range, (females) Males 68,II4.50 III 16.88, + .8 W30–.5 4.04 3.00–5.25, PW 0.68; 6.13 aeWMT86 fml A 640:T 37 1.0,P 6.88 PL (15.00), 13.75 TL T62470): WAM (female T58362 WAM Male ¡ + .8 W34–.0 4.43 3.45–5.40, PW 0.68; 3.50 + 2.50 . 5 . I 32,II5.63 III 23.25, I . . + + 5.25 II II – . . + I;clrto n cplt ea sml.Siaino leg of Spination male. as setae scopulate and coloration III; I;bon eoawt aea akrbonlongitudinal brown darker lateral with femora brown, III; + 1.88 4.38 + 5 2.88 + .5 5.88 I 7.25, 2.38 + ¡ 7.13 + ¡ 0.60; 5 – D:T .51.5 10.72 8.55–13.75, TL SD): + ¡ 65,I 6.13 IV 16.50, + 6.88 2.50 0.51; n 5 + + 5 7(L94–68,13.21 9.45–16.80, (TL 17 3.50 6.50 .8 6.13 I 8.38, n 5 5 25). + 4.25 31,I 7.38 IV 23.13, + 7.63 + 2.38 + + + patella/tibia 6.75 8.38 5 90,I 4.75 II 19.00, + + 2.63 + 6.38 ¡ 9.63 .4 L4.80– PL 1.44; 5 + + + 3.50 23.13). metatarsus 9.00 ¡ .3 PL 2.13; 5 + + 24.38, 4.25 5.63 5 + + Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 rps oeaporaegnrcpaeet ae anyo h rgnldescriptions. original these the of on none mainly that based placements reveals generic description appropriate species more to original propose attributed the be in or can material species listed type were of evaluation East Middle the to addition In Dingosa to misattributed Species and Victoria, Wales, Discussion South New in found occasionally 7). Australia, (Figure Queensland South species-level and to Western respect in cautiously Distribution treated be and (1965) must Lane above by reliably listed provided traits. be information ecological as cannot the that (1979) Consequently, material level. McKay numbers, juvenile specific smaller and to in misidentifications included, identified of also number study material considerable his His a that species. revealed one WAM the than simsoni of more collection on the based in was material lost reference his species. considered of this examination are of the identity types of the illustration the about and doubt therefore description no accurate and (1877) leave Koch’s unknown epigyne L. is However, 2005). collection (Framenau this of whereabouts serrata Dingosa observation). Remarks the personal increase Framenau, to W. seemingly turret V. 1965). the 1965; on (Lane (Lane stands short vision slope spider and of The no to leaves distance prey. restricted or for use usually waiting is little to are whilst turret trend Burrows is barricade 1965). the a there (Lane summer, is widening where there During and areas renovating winter, 1979). for towards (McKay wood but of materials pieces grass, of of variety constructed a usually of up and made spinifex’’ palisade of edge ‘‘outer swamp’’, with dry lake’’. sand ‘‘reedy of ‘‘on ‘‘edge complex’’, and ‘‘samphire—lithic plains’’ also ‘‘sand but hills’’, ‘‘sand include descriptions led nFbur n slt sJn.Mtr eae eefudfo a to (McKay females May season Some from following 1979). the found McKay copulate were and 1965; were months eggsac Lane summer females with also the 1979). Mature Females through (see persist recorded August. November June. to were and until appear as some May July but between late from April, numbers recorded as and highest March and with in November, February found were in males already mature of majority The preferences habitat and history Life ae(95 eiae ag oto fhsPDsuisto studies PhD his of portion large a dedicated (1965) Lane a by surrounded is It deep. cm 10–20 ca and vertical open, is species this of burrow The iia oalother all to Similar and .murata D. .simsoni D. a rgnlydsrbdfo aeilo h rde olcin The Collection. Bradley the of material from described originally was iial,rcrslse nMKy(99 as (1979) McKay in listed records Similarly, . Dingosa ieseisfo utai,Arc,SuhAeia hn,and China, America, South Africa, Australia, from species nine , Dingosa , .serrata D. Dingosa ofcltt uuervsoso hs pce ehere we species these of revisions future facilitate To . eiino h utainwl spider wolf Australian the of Revision ro oti td Panc 07.Critical 2007). (Platnick study this to prior sms omno ad ol.Habitat soils. sandy on common most is igs serrata Dingosa igs serrata Dingosa ads serrata Pardosa Scirpa sstetre sa as turret the uses igs 1625 Dingosa and however, , Callitris include D. ’’, Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 20)i lcn hsseisin species this placing in in (2001) placement a the of inverted supports an examination of shows the representatives (1955b) on in Roewer that based Dingosa by like not epigyne much was female septum median the it T-shaped of as illustration (2007) The Platnick material. by type in accepted not publication was same genus which the the in in 10467/1)) described combination (RII formally 9910467 he (SMF which Iran Lahidschan, in collected hartmanni u oisimtr odto tcno eatiue oayautwl pdrfrom spider wolf adult any to attributed be cannot area. considered it cephalic here elevated condition is an and immature have Australia not its does to and coloration Due opisthosoma and prosoma typical usqetyrpre rmcnrlAsrlabsdo eodjvnl (Rainbow and juvenile examined) second a Australia, on South a based not River, clearly Australia is Stevenson holotype central The from 1915). from reported K0943, subsequently (NMV holotype oitrrt u lal hwta hs pce r rnl lcdin placed wrongly are species difficult these are that illustrations show clearly stylized but (1959) interpret, Roewer’s to (RII/11723)). 9911723 (RII/10469/1)), SMF and Tanzania; (RII/10015/1)), 9910015 9910469 SMF Mozambique, SMF Angola; the in described Ree,15) ob nov., comb. 1959), three (Roewer, remaining the transfer we Consequently, 1992). MH22–75 ontcnomt h ufml yoia,btaecnitn with consistent are Brazil; but Sul, (E Lycosinae, do 1986 Dondale, subfamily Grande Allocosinae Rio the subfamily Grande, to the Rio in conform species from not female do and 2724–2725) male BMNH material (type Brazil from lutain faltreseisapa oso iiaiiswith similarities show to appear species three all of illustrations only: females single on based described genus the genus of the member a of to not reminiscent transfer an clearly epigyne and A is pattern female species colour The the a 1885. in shows Simon, septum description original median The T-shaped 282). inverted (MCZ Peru Ana, Santa from communication). personal 1626 eei hrcesaebsdo eia opooy si per ob nieyta the that unlikely be consider to we appears elucidated, it be can As many species morphology. as this doubtful genital considered of on communica- be identity based personal must are Nordander, spiders wolf (T. characters is juvenile lost generic it of considered where placement be Sweden, generic must Gothenburg, Any and Museum tion). subsequently History housed was Natural be the to It in expected present Africa. not South is species Umbonumbi, the in collected to holotype transferred female juvenile a rnfrto transfer A igs persica Dingosa igs topaziopsis Dingosa igs tragardhi Dingosa igs venefica Dingosa he pce eeoiial lcdin placed originally were species Three igs liopus Dingosa nebulosa nadto,teclrto fti pce sdsrbdb owr(1955b) Roewer by described as species this of coloration the addition, In . .W rmnu&B .Baehr C. B. & Framenau W. V. Ree,15) ob o. edn eiino African of revision a pending nov., comb. 1959), (Roewer, gopwithin -group Dingosa Hogna Hogna Allocosa Cabri,11) hsseiswsdsrbdbsdo igefemale single a on based described was species this 1916): (Chamberlin, owr 95 owr(95)dsrbdti pce rmafemale a from species this described (1955b) Roewer 1955: Roewer, Kyeln,19) h eiai hrceso hsseisdescribed species this of characters genitalic the 1891): (Keyserling, Lwec,14) hsseiswsdsrbdi h genus the in described was species this 1947): (Lawrence, spnig(E pending is yRee 15)wr usqetysnnmzdadtaserdto transferred and synonymized subsequently were (1959) Roewer by Ree,15b n ial to finally and 1955b) (Roewer, Hg,19) hsseiswsdsrbdfo neryimmature early an from described was species this 1896): (Hogg, h igegnswti hssbaiy spnig(E pending is subfamily, this within genus single the , Trochosa Pardosa Trochosa Dingosa ads completa Pardosa Trochosa ´ oe dubium nomen , lae,proa communication). personal Alvares, . osqety eflo oafra n Marusik and Mozaffarian follow we Consequently, . .lusingana P. .L oh 87bsdo h rgnlillustration, original the on based 1847 Koch, L. C. oafra n aui 20)pooe new a proposed (2001) Marusik and Mozaffarian . Dingosa : igs angolensis Dingosa rcoapersica Trochosa Dingosa igs completa Dingosa sdansdhr,a tde o hwthe show not does it as here, diagnosed as . Ree,15) ob o. and nov., comb. 1959), (Roewer, owr 99(lewied n Jocque and (Alderweireldt 1959 Roewer, .hartmanni D. yRee 15) n l he were three all and (1959), Roewer by ´ Dingosa Dingosa lae,proa communication). personal Alvares, . .traghardi D. Ree,1955). (Roewer, owr 99(Mossamedes, 1959 Roewer, Trochosa Ree,15) h yeof type The 1959). (Roewer, to Pardosa Dingosa Pardosa owr 99(Aruscha, 1959 Roewer, owr 99(Tete, 1959 Roewer, a Dingosa oe dubium nomen n eydfeetto different very and Pardosa w te species other Two . : sdansdhere. diagnosed as ads angolensis Pardosa h epigyne The . . Lycosa ´ Alvares, . Pardosa . Hogna from ´ Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 ecito fauiomydr rw pdrceryd o ofr to conform not do clearly spider brown dark uniformly a of description rnsetadTrtnNradrasse nteufruaeyuscesu erhfrthe for search of unsuccessful to unfortunately material misattributed the in type were assisted that Nortander Torsten America and South Kronestedt from species on information in species this listed (1955b) h upr fteeisiuin n hi nhsatcpronl E personnel. enthusiastic without their impossible and been have institutions would these study and of This (WAM), support (ZMB). Waldock Peter Nawai the Julianne (AM), Shahin and for Smith and Harvey respective Helen Dunlop Mark (MHNP); staff and their Jason (SAM), Rollard Milledge museum Hirst to Graham Christine David visits and (QM), (NMV), and during Seeman Lillywhite Leguin managers, Owen or Elise-Anne and collection loan order): Raven as Robert curators, particular either no of collections (in variety their institutions a accessing in to assistance indebted are We Acknowledgements comb. n. 1936), (Schenkel, Dingosa einbn ftepooa osqety etase hsseisto species this transfer we Consequently, prosoma. the of band median either or in Trochosa 2005), placement a Stratton 2000), Logunov support e.g. and not Zyuzin genus; does Mediterranean description putatively original (1936) Schenkel’s References paper. Harvey this Mark of drafts to earlier on (ABRS) and comments Dondale, Study helpful Charles Thomas, provided Resource Melissa Yoo Adelaide). Biological of Jung-Sun (University Australian Austin Andy the and (WAM) by funded spiders genus rmnuV,Vn J 01 eiino h genus the of Revision 2001. CJ. genus Vink Australian VW, new genus Framenau the spider of wolf Revision Australian 2007. VW. the Framenau of Revision 2006c. VW. Framenau 2006b. VW. Framenau Jocque M, Alderweireldt odl D 96 h ufmle fwl pdr Aaee yoia) ca ogeoItrainlde Internacional genus Congreso spider X wolf Actas The Lycosidae). 2005. (Araneae: VW. spiders 3027–4230. Framenau Douladoure. wolf Toulouse: of N–S. subfamilies 1926–3026. 4: The Douladoure. part 1986. Toulouse: 2, CD. G–M. Volume 3: Araneorum. Dondale part Bibliographia 1958. 2, Volume P. Araneorum. Bonnet Bibliographia 1957. P. Bonnet Lyco family spiders, wolf sheets: Fact 2003. Museum Australian rmnuV.2006a. VW. Framenau igs ursina Dingosa aooy15:927–970. Taxonomy roine(rna:Lcsde.Zoaa1391:1–34. Zootaxa Lycosidae). (Araneae: Artoriinae 1304:1–20. Zootaxa Lycosidae). 1281:55–67. 34:206–213. Arachnology of Journal Aracnologı 5:73–113. wolf_spiders. Zoology Tropical Lycosidae). (Araneae species variable highly some Aaee yoia) eod fteWsenAsrla uem22:265–292. Museum Australian Western the of Records Lycosidae). (Araneae, Lycosa Ree 99.Sn ta.(99 regarded (1999) al. et Song 1959). (Roewer nbt h hp fteeiyeadtecaatrsi ogtdnlbnsi the in bands longitudinal characteristic the and epigyne the of shape the both in a aa Espan Jaca, ´a, Shne 96.Teoiia lutaino h eaeeiyeadthe and epigyne female the of illustration original The 1936). (Schenkel yoatraghardi Lycosa Shne,13) hsseisfo hn a rgnlydsrbdi the in described originally was China from species this 1936): (Schenkel, .19.Arve fthe of review A 1992. R. ´ Knoelle Mainosa a1:327–332. ˜a e ooyi ofsie eu rmAsrla(rna:Lcsde.Zootaxa Lycosidae). (Araneae: Australia from genus spider wolf monotypic new a , Dingosa e eu o h utain‘suteokwl pdr’(rna Lycosidae). (Araneae spider’’ wolf ‘‘shuttlecock Australian the for genus new a , Schizocosa hssuyfrspr farvso fAsrla wolf Australian of revision a of part forms study This . sdansdhr,btspot ls iiaiywith similarity close a supports but here, diagnosed as eiino h utainwl spider wolf Australian the of Revision Artoria nebulosa hmeln 94bth ae rnfre tto it transferred later he but 1904 Chamberlin, hrl nAsrla e yoyisadgnrctransfers generic and synonymies new Australia: in Thorell gopof -group Venatrix ia oln] http://www.am [online]. sidae Artoriopsis Pardosa owr(rna,Lcsde.Invertebrate Lycosidae). (Araneae, Roewer .ursina D. Anomalosa nanwsbaiyo ofspiders, wolf of subfamily new a in oh14 nArc,acmlxwith complex a Africa, in 1847 Koch Schizocosa ´ as e lae provided Alvares der owr 90(Araneae: 1960 Roewer, online.net.au/factSheets/ Trochosa Dingosa aNaci genus; Nearctic (a igs 1627 Dingosa pce inquirenda species Dingosa Torbjo . : Lycosa Roewer . .ursina T. ¨rn (a . Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 aeW.16.Tebooyo,adeooia set fsome of aspects ecological and and of, Bauer biology Nuremberg: The abgebildet. 1965. und WJ. beschrieben Lane Natur der nach http:// Australiens, Co. Arachniden Helicon p. Die (Ukraine): 112 1877. Kharkov Gallery. L. [online]. Art 4.07 Koch and Focus Helicon Museum 2006. Tasmanian A. Hobart: Glazunov Tasmania. D, of Kozub spiders VV, common Khmelik Some 1967. VV. Hickman u .16.Cnrbto a Contribution 1966. Y. Guy 1628 io .19.Dsrpin ’rcndsnueu e ailsdsAeeia,Psuia,Lcsdeet Lycosidae Pisauridae, Agelenidae, des familles des nouveaux d’arachnides Descriptions 1898. E. Simon of male the of Description Provinzen Schluss). 2000. nordwestlichen P. und Sierwald den (Fortsetzung nach (Lycosidae) Expedition II wissenschaftliche Lycosiformae Schwedisch-chinesische 1936. Araneae E. 1959]. Schenkel de date National [imprint Parc du 1960 Exploration (Lycosidae). CF. II Roewer Lycosiformae Araneae 1958]. date [imprint 1959 CF. Roewer culuhT.20.Eooyo h biaebroigsiesascae ihteAead ym bluetongue pygmy Adelaide the with associated spiders p. burrowing 112 obligate Reed. the of Sydney: Ecology colour. 2000. in TP. spiders McCullough p. Australian 296 1970. Collins. R. Sydney: Mascord Spiders. 1976. BY. Main owrC.15b i rne e O der Araneen Institut Brussels: Die 2a, Volume 1940. 1955b. bis 1758 CF. von Araneae 39:483–489. Roewer der Australia Katalog South 1954]. 9:107–319. of date Museum Society [imprint Australian Royal History, 1955a the the Natural of CF. of of Records Transactions Roewer Museum Araneidae. Arachnida. Australian 1915. American of WJ. York: census Rainbow New A 7.5. 1911. WJ. Version Rainbow catalog. spider world 1940– The redescriptions 2007. With NI. 1992–1995. Platnick taxonomy, spider in 1940–1980. Advances transfers 1997]. date and [imprint synonymies 1998 With NI. Platnick 1988–1991. taxonomy, p. spider 673 Press. in University Advances Manchester Manchester: 1993. 1981–1987. taxonomy, NI. spider Platnick in Advances 1989. NI. Platnick reconstruction Phylogenetic Araneae: 2006. 10:67–74. AD. X. Selecta Austin Arthropoda Y-C, Berlin. Park (Aranei). MS, spiders Harvey Museums Iranian SC, Donellan of Zoologischen VW, checklist Framenau A des NP, 2001. Murphy Arachniden-Sammlung YM. Marusik der F, Mozaffarian Typen Die 1992. Arachnida: M. 3, Volume Moritz Australia. of catalogue genus Zoological the editor. of DW, species Walton new In: A Lycosidae. 14. 1985b. Lycosidae): RJ. (Araneae: McKay Australia of spiders wolf The 1985a. RJ. McKay 9. The Lycosidae): (Araneae: 1. Australia of Lycosidae): spiders (Araneae: wolf The Australia 1979. RJ. of McKay spiders wolf The 1973. RJ. McKay ap.p889–968. p Raspe. www.helicon.com.ua. 33:1–172. vce avec xoia.Anlsd aSocie la de Annales Oxyopidae. 28:133–140. Arachnology of Journal fo Lycosidae). Arkiv 1927–1930. Hummel David Su Dr Prof. Expedition und der Hedin Arzt Sven Dr von Leitung 55:519–1040. unter Witte Chinas, de GF l’Upemba—Mission de National Parc du Exploration 55:1–518. Witte de GF l’Upemba—Mission 164:751–782. iad( lizard p. 186 Science. of Department Australia, Western O p. 923 Belgique. de Naturelles Sciences des Royal Natural of Museum American the http://research.amnh.org/entomology/spiders/catalog/INTRO1.html. with association in Press p. 976 University History. History. Manchester Natural Manchester: of 1980. Museum American the with association Molecular in p. 846 Press behavior. University building Manchester Manchester: web of evolution the genes: NADH1 and and 38:583–602. rRNA Evolution biogeography 28 and rRNA, 12S Phylogenetics classification, the from for sequences using Lycosidae) implications (Araneae: spiders wolf the of Canberra: 68:309–329. Berlin Museums Palpigradi. Zoologischen des Amblypygida, Mitteilungen Lycosidae. Pseudoscorpionida, 73–88. p part, Service. Publishing in Government Australian Araneomorphae Mygalomorphae, 22:101–104. Museum Queensland the of Memoirs 19:225–229. Museum Queensland the of uesadMsu 16:375–398. Museum Queensland ¨ trecice kdmedrWseshfe,Mteaic-auwseshflceKas,AtiugI Abteilung Klasse, Mathematisch-Naturwissenschaftliche Wissenschaften, der Akademie sterreichischen td spe ´tude .W rmnu&B .Baehr C. B. & Framenau W. V. iiu adelaidensis Tiliqua caedsespe des ´ciale l’e ` hnustei] dlie dlieUiest.15p. 115 University. Adelaide Adelaide: thesis]. [honours ) td e araigne des ´tude csd ao.Taaxd ’ntttSiniiu hrfe,Se Cherifien, Scientifique l’Institut de Travaux Maroc. du `ces ´te noooiu eBliu 42:5–34. Belgique de Entomologique ´ ¨ oipsplacidus Sosippus trecice rnEpdto 995.Stugbrct der Sitzungsberichte 1949/50. Iran–Expedition sterreichischen e el ail e yoia td asu-ail e Lycosinae des sous-famille la de et Lycosidae des famille la de ´es igCag rna eaml o schwedischen vom gesammelt Araneae Ping-Chang. ¨ ihntso h ufml oipne(Araneae, Sosippinae subfamily the on notes with , Lycosa rZooi29:1–314. Zoologi ¨r pce PDtei] et:Uiest of University Perth: thesis]. [PhD species ads serrata Pardosa bicolor L oh17) Memoirs 1877). Koch (L. ru.Mmiso the of Memoirs group. reZoologie ´rie Pardosa . Downloaded By: [Danish Veterinary and Agricultural Library] At: 19:32 2 September 2007 o -,Faea W 06 ytmtc n igorpyo h he-e uligwl pdrgenus spider wolf building sheet-web the of biogeography and genus Systematics the 2006. VW. of Framenau spiders J-S, Yoo burrowing the of revision A 1942. HK. Wallace Apte insectes. des naturelle Histoire 1837. in Publishing CA. behaviour Technology Walckenaer courtship and and Science ornamentation Hebei of Shijiazhuang: Evolution China. 2005. of G. spiders Stratton The 1999. Su J. Fauna Chen Die M, editors. R, Zhu Hartmeyer D, W, Song Michaelsen In: partie. 2me Araneae, 1909. E. Simon yznA,LgnvD.20.Nwadltl-nw pce fteLcsdefo zrajn h Caucasus the Azerbaijan, from Lycosidae the of species little-known and New 2000. DV. Logunov AA, Zyuzin iln auait27:1–62. Naturalist Midland p. 33:347–376. 682 Arachnology of Journal Lycosidae). (Araneae, morphology on based p. 640 House. su Hamburger der Ergebnisse Aaee yoia) ultno h rts rcnlgclScey11:305–319. Society Arachnological British the of Bulletin Lycosidae). (Araneae, 20:675–712. Systematics Invertebrate Lycosidae). (Araneae: detasrlshnFrcugrie10.Jn:Gsa ice.p155–212. p Fischer. Gustav Jena: 1905. Forschungsreise ¨dwest-australischen eiino h utainwl spider wolf Australian the of Revision rs oue1 ai:LbareEncyclope Librairie Paris: 1. Volume `res. Geolycosa Schizocosa Aaee yoia) American Lycosidae). (Araneae, nihsfo phylogeny a from insights : igs 1629 Dingosa ¨dwest-Australiens. dqeRoret. ´dique Venonia