Pre Cambrian – Devonian
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Morphology and Developmental Traits of the Trilobite Changaspis Elongata from the Cambrian Series 2 of Guizhou, South China
Morphology and developmental traits of the trilobite Changaspis elongata from the Cambrian Series 2 of Guizhou, South China GUANG-YING DU, JIN PENG, DE-ZHI WANG, QIU-JUN WANG, YI-FAN WANG, and HUI ZHANG Du, G.-Y., Peng, J., Wang, D.-Z., Wang, Q.-J., Wang, Y.-F., and Zhang, H. 2019. Morphology and developmental traits of the trilobite Changaspis elongata from the Cambrian Series 2 of Guizhou, South China. Acta Palaeontologica Polonica 64 (4): 797–813. The morphology and ontogeny of the trilobite Changaspis elongata based on 216 specimens collected from the Lazizhai section of the Balang Formation (Stage 4, Series 2 of the Cambrian) in Guizhou Province, South China are described. The relatively continuous ontogenetic series reveals morphological changes, and shows that the species has seventeen thoracic segments in the holaspid period, instead of the sixteen as previously suggested. The development of the pygid- ial segments shows that their number gradually decreases during ontogeny. A new dataset of well-preserved specimens offers a unique opportunity to investigate developmental traits after segment addition is completed. The ontogenetic size progressions for the lengths of cephalon and trunk show overall compliance with Dyar’s rule. As a result of different average growth rates for the lengths of cephalon, trunk and pygidium, the length of the thorax relative to the body shows a gradually increasing trend; however, the cephalon and pygidium follow the opposite trend. Morphometric analysis across fourteen post-embryonic stages reveals growth gradients with increasing values for each thoracic segment from anterior to posterior. The reconstruction of the development traits shows visualization of the changes in relative growth and segmentation for the different body parts. -
Download Abstract Booklet Session 4
Abstract Volume 17th Swiss Geoscience Meeting Fribourg, 22nd + 23rd November 2019 4 Palaeontology 106 4. Palaeontology Torsten Scheyer, Christian Klug, Lionel Cavin Schweizerische Paläontologische Gesellschaft Kommission des Schweizerischen Paläontologischen Abhandlungen (KSPA) Symposium 4: Palaeontology TALKS: 4.1 Alleon J., Bernard S., Olivier N., Thomazo C., Marin-Carbonne J.: Molecular characteristics of organic microfossils in Paleoarchean cherts 4.2 Antcliffe J.B., Jessop W., Daley A.C.: Prey fractionation in the Archaeocyatha and its implication for the ecology of the first animal reef systems 4.3 Bastiaans D., Kroll J.F., Jagt J.W.M., Schulp A.S.: Cranial pathologies in a Late Cretaceous mosasaur from the Netherlands: behavioral and immunological implications. 4.4 Daley A.C., Antcliffe J.B., Lheritier M.: Understanding the fossil record of arthropod moulting using experimental taphonomic approaches 4.5 Dziomber L., Foth C., Joyce W.G.: A geometric morphometric study of turtle shells 4.6 Evers S.W.: A new hypothesis of turtle relationships provides insights into the evolution of marine adaptation, and turtle diversification 4.7 Fau M., Villier L., Ewin T.: Diversity of early Forcipulatacea (Asteroidea) 4.8 Ferrante C., Cavin L.: Weird coelacanths from the Triassic of Switzerland 4.9 Frey L., Coates M.I., Rücklin M., Klug C.: A new early symmoriid with an unusual jaw articulation from the Late Devonian of Morocco 4.10 Friesenbichler E., Hautmann M., Bucher H.: Palaeoecology of benthic macroinvertebrates from three Middle Triassic -
Sequence of Post-Moult Exoskeleton Hardening Preserved in a Trilobite Mass Moult Assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco
Editors' choice Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco HARRIET B. DRAGE, THIJS R.A. VANDENBROUCKE, PETER VAN ROY, and ALLISON C. DALEY Drage, H.B., Vandenbroucke, T.R.A., Van Roy, P., and Daley, A.C. 2019. Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco. Acta Palaeontologica Polonica 64 (2): 261–273. Euarthropods have a tough exoskeleton that provides crucial protection from predation and parasitism. However, this is restrictive to growth and must be periodically moulted. The moulting sequence is well-known from extant arthropods, consisting of: (i) the long inter-moult stage, in which no changes occur to the hardened exoskeleton; (ii) the pre-moult stage where the old exoskeleton is detached and the new one secreted; (iii) exuviation, when the old exoskeleton is moulted; and (iv) the post-moult stage during which the new exoskeleton starts as soft, thin, and partially compressed and gradually hardens to the robust exoskeleton of the inter-moult stage. Trilobite fossils typically consist of inter-moult carcasses or moulted exuviae, but specimens preserving the post-moult stage are rare. Here we describe nine specimens assigned to Symphysurus ebbestadi representing the first group of contemporaneous fossils collected that preserve all key stages of the moulting process in one taxon, including the post-moult stage. They were collected from a single lens in the Tremadocian part of the Fezouata Shale Formation, Morocco. Based on cephalic displacement and comparison to other trilobite moults, one specimen appears to represent a moulted exoskeleton. -
1414 Hughes.Vp
The depositional environment and taphonomy of the Homerian Aulacopleura shales fossil assemblage near Lodìnice, Czech Republic (Prague Basin, Perunican microcontinent) NIGEL C. HUGHES, JIØÍ KØÍ, JOSEPH H.S. MACQUAKER & WARREN D. HUFF Excavation of Joachim Barrande’s classic fossil locality of the “Aulacopleura shales” exposed on Na Černidlech Hill, near Loděnice reveals that most specimens were recovered from a 1.4 m interval exposed in “Barrande’s pits”. These are located at the eastern end of a 0.4 km trench dug in the mid 1800’s to expose the interval along strike. Over an hundred bedding planes occur within the 1.4 m interval, and thousands of articulated trilobites have been collected at the site. In- dividual bed surfaces vary in the density, size, and taxonomic composition of the fossils contained. Some preserve a di- verse benthic shelly fauna, others are almost exclusively dominated by the trilobite Aulacopleura koninckii, and a third variety is apparently barren of all shelly fossils. Isolated sclerites of A. koninckii are rare, and on almost all bedding sur- faces exoskeletons are predominantly partially articulated and lack both alignment and sclerite fragmentation. The oc- currence of A. koninckii conforms in many ways to the characteristics of a Type I trilobite lagerstätte of Brett et al. (2012). The presence of enrolled A. koninckii suggests that final burial may have resulted from relatively rapid obrution, although the condition of partial articulation indicates that many carcasses or exuviae partially disaggregated before burial. The mean size and density of A. koninckii specimens varies markedly among bedding planes, with some assem- blages entirely comprised of juveniles, suggesting that notably dense trilobite clustering was not restricted only to repro- ductively mature individuals. -
Plumulitid Machaeridian Remains from the Silurian (Telychian) of Severnaya Zemlya, Arctic Russia
NORWEGIAN JOURNAL OF GEOLOGY Plumulitid mochoeridion remains from the Silurion, Arctic Russio 53 Plumulitid machaeridian remains from the Silurian (Telychian) of Severnaya Zemlya, Arctic Russia Anette E.S. Hogstrom, Olga K. Bogolepova & Alexander P. Gubanov Hogstrom, A.E.S., Bogolepova, O.K. and Gubanov, A.P.: Plumulitid machaeridian remains from the Silurian (Telychian) of Sevemaya Zemlya, Arc tic Russia. Norsk Geologisk Tidsskrift, Vol. 82, pp. 53-55. Trondheim 2002, ISSN 029-196X. The machaeridian genus Plumulites is reported for the first time from the Severnaya ZemlyaArchipelago of Arctic Russia, where it occurs in limes tone concretions within the Sredninskaya Formation. Graptolites fromthe same concretions indicate the late crispus- griestoniensisBiozones of the mid Telychian (Uandovery). Similarities to plumulitid sclerites from the Upper Ordovician of the Tairnyr Peninsula promotes further interest in machaeridian faunas fromthis region. A.E.S. Hogstromi, O.K. Bogolepova and A.P. Gubanov, Historical Geology & Palaeontology, Dept. of Earth Sciences, Uppsala University, Norbyviigen 22, SE-752 36, Uppsala, Sweden. I Temporaryaddress: Dept. of Earth Sciences, Wills Memorial Building, Queen's Road, Bristol BSB l RJ, UK. lntroduction Stratigraphy and locality The global record of Silurian machaeridians is limited, Machaeridians discussed herein originate from the but includes rare articulated specimens, and more com Lower Silurian Sredninskaya Formation (Matukhin et monly isolated sclerites that have been found in Britain al. 1999). To avoid nomenclatural questions, it should (de Koninck 1857; Woodward 1865; Withers 1926 and be noted that these rocks were previously referred to as Adrain et al. 1991), the Baltic Region (Aurivillius 1892; the Golomaynnaya Formation (Menner et al. -
A Case Study of Eyeless, Dimorphic Devonian Trilobites from Poland
[Palaeontology, Vol. 59, Part 5, 2016, pp. 743–751] ANCIENT ANIMAL MIGRATION: A CASE STUDY OF EYELESS, DIMORPHIC DEVONIAN TRILOBITES FROM POLAND † by BŁAZEJ_ BŁAZEJOWSKI_ 1, CARLTON E. BRETT2,ADRIANKIN3, , † † ANDRZEJ RADWANSKI 4, and MICHAŁ GRUSZCZYNSKI 3, 1Institute of Palaeobiology, Polish Academy of Sciences, Twarda 51/55, 00-818, Warszawa, Poland; [email protected] 2Department of Geology, University of Cincinnati, Cincinnati, OH 45221-0013, USA; [email protected] 3‘Phacops’ – Association of Friends of Geosciences, Grajewska 13/40, Warszawa, 02-766, Poland 4Institute of Geology, University of Warsaw, Zwirki_ i Wigury 93, 02-089, Warszawa, Poland Typescript received 15 February 2016; accepted in revised form 8 July 2016 Abstract: We report evidence of one of the oldest known have migrated periodically to shallow marine areas for mass animal migratory episodes in the form of queues of the eye- mating and spawning. The sudden death of the trilobites in less trilobite Trimerocephalus chopini Kin & Błazejowski,_ the queues may have been caused by excess carbon dioxide from the Late Devonian (Famennian) of central Poland. In and hydrogen sulphide introduced into the bottom water by addition, there is evidence for two morphs in this popula- distal storm disturbance of anoxic sediments. This study tion, one with nine segments and the other with ten. We demonstrates the potential for further research on the evolu- infer that these queues represent mass migratory chains tion and ecology of aggregative behaviour in marine arthro- coordinated by chemotaxis, comparable to those observed in pods. modern crustaceans such as spiny lobsters, and further sug- gest that the two forms, which occur in an approximately Key words: animal migration, eyeless trilobite, Phacopinae, 1:1 ratio, may be dimorphs. -
PROGRAMME ABSTRACTS AGM Papers
The Palaeontological Association 63rd Annual Meeting 15th–21st December 2019 University of Valencia, Spain PROGRAMME ABSTRACTS AGM papers Palaeontological Association 6 ANNUAL MEETING ANNUAL MEETING Palaeontological Association 1 The Palaeontological Association 63rd Annual Meeting 15th–21st December 2019 University of Valencia The programme and abstracts for the 63rd Annual Meeting of the Palaeontological Association are provided after the following information and summary of the meeting. An easy-to-navigate pocket guide to the Meeting is also available to delegates. Venue The Annual Meeting will take place in the faculties of Philosophy and Philology on the Blasco Ibañez Campus of the University of Valencia. The Symposium will take place in the Salon Actos Manuel Sanchis Guarner in the Faculty of Philology. The main meeting will take place in this and a nearby lecture theatre (Salon Actos, Faculty of Philosophy). There is a Metro stop just a few metres from the campus that connects with the centre of the city in 5-10 minutes (Line 3-Facultats). Alternatively, the campus is a 20-25 minute walk from the ‘old town’. Registration Registration will be possible before and during the Symposium at the entrance to the Salon Actos in the Faculty of Philosophy. During the main meeting the registration desk will continue to be available in the Faculty of Philosophy. Oral Presentations All speakers (apart from the symposium speakers) have been allocated 15 minutes. It is therefore expected that you prepare to speak for no more than 12 minutes to allow time for questions and switching between presenters. We have a number of parallel sessions in nearby lecture theatres so timing will be especially important. -
Species of the Devonian Aulacopleurid Trilobite Cyphaspides from Southeastern Morocco
Journal of Paleontology, 94(1), 2020, p. 99–114 Copyright © 2019, The Paleontological Society. This is an Open Access article, distributed under the terms of the Creative Commons Attribution licence (http://creativecommons.org/ licenses/by/4.0/), which permits unrestricted re-use, distribution, and reproduction in any medium, provided the original work is properly cited. 0022-3360/20/1937-2337 doi: 10.1017/jpa.2019.71 Species of the Devonian aulacopleurid trilobite Cyphaspides from southeastern Morocco Brian D.E. Chatterton,1 Stacey Gibb,1 and Ryan C. McKellar2 1Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, AB T6G 0R2, Canada <[email protected]>, <[email protected]> 2Royal Saskatchewan Museum, 2445 Albert Street, Regina, Saskatchewan, S4P 4W7, Canada <[email protected]> Abstract.—Three new species of Cyphaspides are proposed: C. ammari, C. nicoleae, and C. pankowskiorum. These species are based on specimens obtained from Middle Devonian (Eifelian) strata of the Bou Tchrafine Group, near Erfoud, in the Province of Errachidia, southeastern Morocco. The present contribution enhances our knowledge of Cyphaspides by providing details of three new species that are based on well-preserved, complete, and articulated types. The genus Cyphaspides is discussed, and an emended diagnosis is provided. The paleobiogeography, ontogeny, and relationships of the genus are discussed. UUID: http://zoobank.org/4a7aab8f-8c8e-4498-9cc2-6f8c69b85213 Introduction Republic (Barrande, 1846, 1872;Růžička, 1939; Prantl and Přibyl, 1950), the Armorican Massif (Massif Armoricain) of Moroccan Lower and Middle Devonian trilobite faunas are northwestern France (Pillet, 1972), Morocco (Alberti, 1969; known for their abundance and diversity, frequent occurrence Crônier et al., 2018; and herein), China (Yi and Hsiang, 1975; of articulated specimens, and exceptional quality of preserva- Luo and Jiang, 1985), and Uzbekistan (Kim et al., 1978b). -
The Evolution of Trilobite Body Patterning
ANRV309-EA35-14 ARI 20 March 2007 15:54 The Evolution of Trilobite Body Patterning Nigel C. Hughes Department of Earth Sciences, University of California, Riverside, California 92521; email: [email protected] Annu. Rev. Earth Planet. Sci. 2007. 35:401–34 Key Words First published online as a Review in Advance on Trilobita, trilobitomorph, segmentation, Cambrian, Ordovician, January 29, 2007 diversification, body plan The Annual Review of Earth and Planetary Sciences is online at earth.annualreviews.org Abstract This article’s doi: The good fossil record of trilobite exoskeletal anatomy and on- 10.1146/annurev.earth.35.031306.140258 togeny, coupled with information on their nonbiomineralized tis- Copyright c 2007 by Annual Reviews. sues, permits analysis of how the trilobite body was organized and All rights reserved developed, and the various evolutionary modifications of such pat- 0084-6597/07/0530-0401$20.00 terning within the group. In several respects trilobite development and form appears comparable with that which may have charac- terized the ancestor of most or all euarthropods, giving studies of trilobite body organization special relevance in the light of recent advances in the understanding of arthropod evolution and devel- opment. The Cambrian diversification of trilobites displayed mod- Annu. Rev. Earth Planet. Sci. 2007.35:401-434. Downloaded from arjournals.annualreviews.org ifications in the patterning of the trunk region comparable with by UNIVERSITY OF CALIFORNIA - RIVERSIDE LIBRARY on 05/02/07. For personal use only. those seen among the closest relatives of Trilobita. In contrast, the Ordovician diversification of trilobites, although contributing greatly to the overall diversity within the clade, did so within a nar- rower range of trunk conditions. -
Th TRILO the Back to the Past Museum Guide to TRILO BITES
With regard to human interest in fossils, trilobites may rank second only to dinosaurs. Having studied trilobites most of my life, the English version of The Back to the Past Museum Guide to TRILOBITES by Enrico Bonino and Carlo Kier is a pleasant treat. I am captivated by the abundant color images of more than 600 diverse species of trilobites, mostly from the authors’ own collections. Carlo Kier The Back to the Past Museum Guide to Specimens amply represent famous trilobite localities around the world and typify forms from most of the Enrico Bonino Enrico 250-million-year history of trilobites. Numerous specimens are masterpieces of modern professional preparation. Richard A. Robison Professor Emeritus University of Kansas TRILOBITES Enrico Bonino was born in the Province of Bergamo in 1966 and received his degree in Geology from the Depart- ment of Earth Sciences at the University of Genoa. He currently lives in Belgium where he works as a cartographer specialized in the use of satellite imaging and geographic information systems (GIS). His proficiency in the use of digital-image processing, a healthy dose of artistic talent, and a good knowledge of desktop publishing software have provided him with the skills he needed to create graphics, including dozens of posters and illustrations, for all of the displays at the Back to the Past Museum in Cancún. In addition to his passion for trilobites, Enrico is particularly inter- TRILOBITES ested in the life forms that developed during the Precambrian. Carlo Kier was born in Milan in 1961. He holds a degree in law and is currently the director of the Azul Hotel chain. -
Olenid Trilobites: the Oldest Known Chemoautotrophic Symbionts?
Olenid trilobites: The oldest known chemoautotrophic symbionts? Richard Fortey* Department of Paleontology, Natural History Museum, Cromwell Road, London SW7 5BD, United Kingdom Communicated by Lynn Margulis, University of Massachusetts, Amherst, MA, April 3, 2000 (received for review March 1, 2000) Late Cambrian to early Ordovician trilobites, the family Olenidae, Given the widespread occurrence and taxonomic spread of were tolerant of oxygen-poor, sulfur-rich sea floor conditions, and chemoautotrophic symbiosis, it is likely to have been an ancient a case is made that they were chemoautotrophic symbionts. adaptation. Because direct evidence of the fossil bacteria seldom Olenids were uniquely adapted to this habitat in the Lower preserves, evidence of such habits tends to be inferred from Paleozoic, which was widespread in the Late Cambrian over Scan- taxonomic and͞or morphological data, which is not always dinavia. This life habit explains distinctive aspects of olenid mor- reliable (11). The long-ranging solemyid and lucinid bivalves (10) phology: wide thoraces and large numbers of thoracic segments, indicate that by the Silurian (ca. 420 million years), this life mode thin cuticle and, in some species, degenerate hypostome, and the already had been adopted by some groups with living descen- occasional development of brood pouches. Geochemical and field dants. Ancient vent associations have been recognized from at evidence is consistent with this interpretation. Olenids occupied least the Devonian (12). their specialized habitat for 60 million years until their extinction I show in this paper that even by the late Cambrian period (505 at the end of the Ordovician. million years ago) certain extinct arthropods, trilobites belonging to the family Olenidae, evolved features best understood as evidence olorless sulfur bacteria, a heterogeneous category of bacte- of sulfur chemoautotrophic mode of metabolism. -
For Peer Review
Page 1 of 29 Geological Magazine 1 2 3 1 A new nektaspid euarthropod from the Lower Ordovician of Morocco 4 5 6 7 2 8 9 1* 1,2,3 1 1 1 10 3 Francesc Pérez-Peris , Lukáš Laibl , Lorenzo Lustri , Pierre Gueriau , Jonathan B. Antcliffe , 11 1 1 12 4 Orla G. Bath Enright and Allison C. Daley 13 14 15 5 16 17 6 1Institute of Earth Sciences, University of Lausanne, Géopolis, CH-1015 Lausanne, Switzerland 18 19 For Peer Review 20 7 2The Czech Academy of Sciences, Institute of Geology, Rozvojová 269, 165 00 Prague 6, Czech 21 22 23 8 Republic 24 25 3 26 9 Institute of Geology and Palaeontology, Faculty of Science, Charles University, Albertov 6, 27 28 10 Prague, 12843, Czech Republic 29 30 31 11 *[email protected] 32 33 34 12 35 36 37 13 Abstract 38 39 40 14 Nektaspids are Paleozoic non-biomineralised euarthropods that were at the peak of their diversity 41 42 15 in the Cambrian period. Post-Cambrian nektaspids are a low diversity group with only a few 43 44 16 species described so far. Here we describe Tariccoia tazagurtensis, a new species of small 45 46 47 17 bodied nektaspid from the Lower Ordovician Fezouata Shale of Morocco. The new species 48 49 18 differs from the type (and only other known) species from the Ordovician of Sardinia (Italy), 50 51 19 Tariccoia arrusensis, in possessing more pointed genal angles, a cephalon with marginal rim, a 52 53 20 pygidium with anterior margin curved forwards, a rounded posterior margin and longer and more 54 55 56 21 curved thoracic tergites.