World Checklist of the Genus Cardamine L. (Draft 1)
Total Page:16
File Type:pdf, Size:1020Kb
World checklist of the genus Cardamine L. (Draft 1) 1,2Marhold, K., 1Kučera, J., 1Zozomová-Lihová, J. & 3Al-Shehbaz, I. A. 1Institute of Botany, Slovak Academy of Sciences, Dúbravská cesta 9, SK-845 23 Bratislava, Slovak Republic 2Department of Botany, Charles University, Benátská 2, CZ-128 01 Praha 2, Czech Republic 3 Missouri Botanical Garden, P.O. Box 299, St. Louis, Missouri, U.S.A. 63166-0299 Comments on this draft should be sent to: [email protected] Acknowledgements: We acknowledge financial support provided by the Slovak Research and Development Agency (project no. RPEU-0003-06, to Karol Marhold). Distribution data follow: Brummitt, R. K. (2001): World geographical scheme for recording plant distributions. Edition 2. Hunt Institute for Botanical Documentation, Carnegie Mellon University, Pittsburgh. For references to the chromosome number reports see: Marhold, K. & Kučera, J., Karyological database of the genus Cardamine, available at http://www.cardamine.sav.sk/ Cardamine adriatica Jar. Kučera, Lihová & Marhold, Taxon 59: 161 (2010). Ind. loc.: “Dalmatien “Urulia” [near D. Brela village] v. Welden” – Type: unknown – the original material was most likely destroyed in 1849 during the great Zwinger fire (Stafleu & Cowan, 1983); Neotype (Kučera et al. 2010): Croatia, Biokovo Mts., Baška Voda, village of Bast, 43°21´31´´N, 16°59´18´´E, 398 m a. s. l., 24.IV.2003, J. Kučera & M. Kolník s.n. (SAV!). ≡ Pteroneurum bipinnatum Rchb., Fl. Germ. Excurs.: 676. 1830–1832 (non C. bipinnata (C.A. Mey.) O.E. Schulz 1903) ≡ C. maritima f. bipinnata (Rchb.) Sagorski in Österr. Bot. Z. 61: 18. 1911 ? = Pteroneurum microphyllum J. Presl & C. Presl, Delic. Prag. 1: 237. 1822 Ind. loc.: “Hab. in arenosis maritimis Dalmatiae prope Capo Cesto [Primošten]” – Type: unknown. − “C. microphylla” J. Presl & C. Presl, Delic. Prag. 1: 15, 1822 non Adams., nec (Willd.) O.E. Schulz, nom. inval. [Art. 34.1(a); McNeill & al., 2006]. Chromosome number: 2n = 16 (Kučera et al., 2010) Distribution: Europe: 13. YUG-CR Representative herbarium specimens: Europe: Croatia: Velebit Mts.: Velebit, s.d., K. Malý (W); In saxosis ad Karlobag, 3.VI.1890, L. Stipetić (ZA); in litt. Croat. supra Karlobag, 7.VII.1896, Lj. Rossi (ZA); Velebit, ab Carlopago [Karlobag], 10.VI., s. coll. (PRC); Dalmatia: prope Carlopago [Karlobag], s.d., K. Malý (CL); auf dem Velebit bei Carlopago [Karlobag], 10.VI.1895, G. Beck (PRC); Croatia: Karlopago [Karlobag], VII.1836, L.F. Vukotinović (W); Flora Croatica, ad Karlobago, s.d., J.C. Schlosser (WU); Kod Karlobaga, s.d., L.F. Vukotinović & J.C. Schlosser 852 (ZA); Dalmatia: Velebit, Velika Paklenica, 1 Starigrad, 26.VII.1906, A. Degen (PRC); Kroatien, Velebit, Velika Paklenica bei Starigrad, Engstelle der Schlucht, schutt, 120–150 m, 21.IV.1979, E. Vitek (W); Flora Dalmatica: Velebit in rupiorum fissuris Prosena supra Golubič, 13.IX.1913, A. Degen (ZA); Flora von Jugoslawien, Kroatien, Velebit-gebirge, an der Küstenstraße südlich von Jablanac, 22.VI.1968, M. Bässler & I. Quasdorf 1222 (B); ad Vranjak, Jablanac, 6.VII.1896, Lj. Rossi (ZA); Kroatien, Prizna, 20.V.1971, F. Kummert (W); Velebit: in monte Pavelič Kuk, in Šugarska duliba, 1084 m, 14.VII.1909, Lj. Rossi (ZA); Velebit: Stolačka Peč in Šugarska Duliba, 14.VII.1909, Lj. Rossi (ZA); Mosor Mts.: Spalato (Spljet) [Split]: im Mosor am Felsen, 800 m, 29.V.1895, G.A. Poscharsky (CL); Dalmatien, Mt. Mosor, 1869, K. Malý (W); Mosor, s.d., Fr. Petter (PRC); Dalmatien: Gebirgspflanze, an Kalkfelsen auf dem Berge Mossor bei Spalato [Split], V., Fr. Petter (CL, W, ZA); Jugoslavia: Spalato [Split], IV.1903, Fr. Petter (K); Flora Dalmatica exsiccata, an Kalkfelsen auf dem Berge Mossor bei Spalato [Split], V., Fr. Petter (PR); Mosor bei Spalato [Split], V., Fr. Petter (PR); Dalmatien: Spalato [Split], s.d., Fr. Petter (W, WU, BEOU); Dalmatien: Mosor bei Split, Karstregion, Felsheide, Kalk, 1100 m, 13.VII.1964, F. Krendl (W); Spalato [Split] in Dalmatien, s.d., Regel (Z); Ad rup. calcar. montis Mossor pr. Spalato [Split] in Dalmatia, V., Bentych? (Z); Süd Dalmatien, Spalato [Split], s.d., M. Tommasini (B); In montosis Mosor in Dalmatia, s.d., I.I. Schiffner (B); Spalato [Split], s.d., J.C. Equitis-Pittoni & Dannenfeldt (W); Dalmatien: Mosor, auf Felsen am Gipfel des Ljubljan, 22.VI.1927, E. Korb (W); Mt. Ljubljan, V., Fr. Petter (W); auf Felsen, am Weg von Almissa [Omiš] nach den Mühlen an der Cettina, auf dem Berge Ljubljan, und in anderen nördlichen Gegenden des Mossor Gebirges, s.d., Fr. Petter (W); auf Felsen, am Wege von Almissa [Omiš] nach den Mühlen an der Cettina, auf dem Berge Lubljan, und in anderen nördlichen Gegenden des Mossor Gebirges, V., Herbarium Heer (Z); Cettinie, s.d., M. Sardagna (WU); Dalmatien: Mossor planina, nächst dem Sattel kote 1089 m, südöstlich der Ljubirna, 9.VI.1908, E. Janchen (WU); Planina Mosor, Doman staje, 12.VIII.1990, D. Vladović (ZA); Biokovo Mts.: M. Biokovo, s.d., M. Sardagna (WU); Biokovo, 1400 m, VI.1911, A. Teyber (WU); Biokovo, beim Einstieg nach Makarska, 28.VI.1927, E. Korb (W); Flora Croatica, Biokovo, Kuranik – Sv. Ilja, 28.VII.1940, I. Horvat (ZAHO); Flora Croatica, Biokovo, Pod Vošćem, 26.VII.1940, I. Horvat (ZAHO); Kroatien, Dalmatien, Brela, übergang nach Ober-Brela, VI.1980, G. Fitz (W); Dalmatien, Brela, westwärts der Bauernburg (aber Fruchtklappen nicht Zurückrollend), mitte VI.1977, G. Fitz (W); Biokovo, vapenačka mjesta iznad mesta Soline-Brela, 430 m, 16.VI.1956, R. Domac (ZA). Cardamine alpina Willd., Sp. Pl. 3: 481 (1800). Ind. loc.: “Habitat in alpibus Helvetiae, Sabaudiae, Delphinatus, Carinthiae (v.s.).” Type: ≡ Cardamine bellidifolia subsp. alpina (Willd.) B.M.G.Jones, Feddes Repert. 69: 59 (1964). Chromosome number: n=8 (Tischler G., 1950 – without locality) 2n=16 (Manton I., 1932 – without locality) Distribution: Europe: 11. GER; 12. FRA-FR, SPA-AN, SPA-SP; 13. AUT-AU, ITA-IT, SWI. Representative herbarium specimens: Europe: Spain: Huesca, Aneto, 2660-2700 m, 17.8.1984, P. & J.Montserrat (JACA); Spain: Barranco del Portet-Collado Fenás, calcosquitstos y granitos Sallent de Gállego, Huesca, 2400-2660 m, 22.7.1993, L.Villar & J.A.Sesé (JACA); France: da Mandette, Lauret, Hautes Alpes Dauphiné, 8.1904, L.V.Lester- Garland (K); Italy: Longobardia super., Prov. di Sondrio, Bormio, in glareosis montis 2 Leverone, alt. 2600-2900 m, 14.8.1908, M.Longa (K); Switzerland: Col de Grimsel, 17.8.1950, A.Lawalréc (K). Cardamine amariformis Nakai, Bot. Mag. (Tokyo) 26 (311): 324−325 (1912) 'amaraeformis' [Rec. 60G.1a, McNeill et al. 2006]. Ind. loc.: “Hab. secus torrentes montis des diamantes 23. VI. 1906 n. 557 (Faurie) ibidem 18. VIII. 1902 (T. Uchiyama).” Type: LT (proposed): [Korea], secus torrentes montis des diamants, U. Faurie 557, 23.vi.1906 (TI!); isolectotype (P 00279344!). Other original material: Corea, Mt. Kongo-san [in Japanese], T. Uchiyama, 18.viii.1902 (TI!). ≡ C. fauriei Lév., Feddes Repert. 10: 350 (1912) non Franch. (1888), nom. illeg. Ind. loc.: “Corea: secus torrentes montis ’des diamants’, 23 jun. 1906 (Urb. Faurie, 557).” Type: LT (proposed): [Korea], secus torrentes montis des diamants, U. Faurie 557, 23.vi.1906 ( TI!) − C. yezoensis auct. non Maxim.: Y. N. Lee, Fl. Korea: 256 (1996). Chromosome number: Distribution: Asia-Temperate: 31. PRM; 36. CHN, CHM; 38. KOR-NK, KOR-SK. Representative herbarium specimens: Asia-Temperate: Ussuri: Tatrskij Strait, coast of De-kastri bay, 1913, Logousov (LE); Ussuri: Botča valley, 1924, Šiškin 78.301 (LE); Ussuri: Oktrabrskij prisk, 1962, Ogureeva (MHA); Ussuri: Tardoki-Jani Mts., 1963, Šaga (LE, MHA); Ussuri: Privabskoe Lake, 1966, Šipalova (MHA); Ussuri: Upper Bandžal, 1984, Šlothgauer (MHA); Korea: loc. Yungoksu (temple, alt 400 m) – Nogodan (peak alt. 1507 m), Toji myun, Goorae-gun, Junranam-do, 127°35´E, 35°3´N, 1000 m, 2.6.1984, J.Murata 16042 (TI); Korea: Kangwon-do, Yangyang- gun, Seo-myun, Osek-oncheon, Sulak san, 800-1600 m, 4.7.1986, J.Murata & K.Sung-chul 21017 (TI). Cardamine acris Griseb., Spic. Fl. Rumel. 1: 253, 254 (1843). ≡ Cardamine raphanifolia subsp. acris (Griseb.) O.E.Schulz, Bot. Jahrb. Syst. 32: 512 (1903). Ind. loc.: “In regione Macedoniae et Serbiae australis: socialis in herbosis humudis et ad rivulos m. Nidgé alt. 4400´-5500´ (substr. Marmor.), gregariae ad fontem m. Kobelitza juxta Mandram Weizensem alt. 4200´ (substr. Micasch.)!; in rivulo m. Kopaunik 5700´alt (Friedr.).” Type: LT - (Strid 1986:257): Ad fontem gregariae m. Kobelitza juxta Mandramt, s. a., Grisebach no. 943 (GOET). Cardamine acris subsp. acris Chromosome number: 2n=16 (Ančev M., Marhold K. & Goranova M. 1997 – Bulgaria; Perný M., Tribsch A. & Anchev M.E. 2004 - Bulgaria, Montenegro, Greece) Distribution: Europe: 13. ALB, BUL, GRC, YUG-BH, YUG-MA, YUG-MN, YUG-SE. Representative herbarium specimens: Europe: Bosnia and Hercegovina: Ivan Planina, in locis vallis Hlboki potok prope Bradina, ca.1200 m, 24.5.1925, K.Maly (K); Serbia: Kosovo i Metohija, östl. Čakor-Pass, ca. 35 km westl. Peć, ca. 1540 m, 8.8.1976, E.Landolt 971 (ZT); Montenegro: Mt. Bertiscus, in jugo Čakor prope Andrijevica, ca. 1700-1800 m, 29.5.1958, K.H.Rechinger 19601 (K); 3 Montenegro: In rivularibus montis Ljubovički Likan supra Štavnjica gorna pr Peč, ca.1800 m, 8.1914, D.K.Vandaz (PR); Bulgaria: In subalpinis ad rivum Demjanitsa, m. Pirin, 16.7.1929, N.Stojanoff, B.Stefanoff & T.Georgieff (K); Bulgaria: a paludosis ad fontem Catal Cosma, in st. Rila australis sub caucinem Carev vrch, ca. 2100 m, 19.6.1931, N.Stojanoff & T.Georgieff (BRNOU); Bulgaria: Balkan centr.-infra cacumina Baba, 1400 m, 2.5.1902, A.Tošev (SO, SOM); Bulgaria: Zapadna Stara Planina, do Brza reka pod vrh Midzhur, 1900 m, 20.5.1993, V.Vladimirov (SO); Albania: Erammos Range, above Ersekë, by hills on bare upper slopes, ca 5500 ft, 27.6.1933, A.H.G.Alston & N.Y.Sandwith 1938 (BM); Albania: Shemeri, NE of Elbazan, 3000 ft, 22.5.1935, R.V.Pennigton (K); Albania: Distrikt Kalis, Korab, an quellingen Orten der Čafa Korabit, ca.2200 m, 1.7.1918, Dörfler 755 (WU); Macedonia: Debar, montibus Korab, prope karaul Skrtac, ca.