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Y Y A A P P OFFICERS AND EDITORS OF THE PALEONTOLOGICAL SOCIETY Grand Patrons ANDREW H. KNOLL RICHARD K. BAMBACH CHARLES LANE ARTHUR J. BOUCOT ALLAN R. LARSON President NADIA FRÖBISCH, MARY SILCOX, Education/Outreach Coordinator RODNEY M. FELDMAN AND CARRIE SCHWEITZER CECILIA LENK AND PAUL K. STROTHER ARNOLD I. MILLER, Museum für Naturkunde Univ. of Toronto Scarborough ROWAN LOCKWOOD, ROBERT AND ELVIRA GASTALDO JERE H. LIPPS Univ. of Cincinnati College of William & Mary PETER J. HARRIES, EDUARDO LEORRI SORIANO, THE FAMILY OF N. GARY LANE ALAN R. LORD [email protected] [email protected] North Carolina State East Carolina University, Greenville A. R. PETE PALMER CHRIS MAPLES AND SARA MARCUS J. WILLIAM SCHOPF President-Elect JANET WOOD MCGREGOR JOACHIM HAUG, DAVID VARRICCHIO, Councilors RAMAN J. SINGH BRUCE MacFADDEN, CHERYL L. METZ Ludwig Maximilian Montana State Univ. LAUREL COLLINS, STEVEN M. STANLEY Florida Museum of Natural History MOLLY F. AND CALVIN MILLER University of Munich Florida International Univ. bmacfadd@fl mnh.ufl .edu STEVE WESTROP, RONALD W. MORIN collinsl@fi u.edu Patrons OSBORNE B. NYE, JR. MICHAEL HAUTMANN, Oklahoma Museum EDWARD J. ANDREW, JR. Past President Univ. of Zurich TERESA O’NEILL DAVID M. WORK, KATE BULINSKI, WILLIAM I. AUSICH STEVEN M. HOLLAND, Bellarmine University RONALD L. PARSLEY Univ. of Georgia MARTIN J. HEAD, Maine State Museum BRUCE M. BELL MARK E. PATZKOWSKY [email protected] [email protected] Brock Univ. STIG M. BERGSTROM JOHN POJETA, JR. PATRICK WYSE JACKSON, DAVID J. BOTTJER MARY LOU POJETA THOMAS HEGNA, Univer. of Dublin Trinity College Student Representative DONALD W. BOYD Secretary JOHN K. POPE Western Illinois University SHARON McMULLEN, DEREK E. G. BRIGGS MARGARET M. YACOBUCCI, MARGARET M. YACOBUCCI, ANNE RAYMOND Bowling Green State Univ. Univ. of Wisconisin–Madison JAMES C. BROWER SAMANTHA HOPKINS, Bowling Green State Univ. MARGARET N. REES [email protected] [email protected] JOHN L. CARTER Univ. of Oregon SAMUEL ZAMORA, PETER R. CRANE FRANK H. T. RHODES EMILY ORZECHOWSKI, RICHARD A. ROBISON Treasurer NIGEL HUGHES, Geological Survey of Spain THE CUSHMAN FOUNDATION PETER J. HARRIES, Univ. of Calif.-Berkeley J. ROBERT DODD JUNE R. P. AND CHARLES A. ROSS UC Riverside RENBIN ZHAN, North Carolina State Univ. [email protected] MARY L. DROSER CHRISTOPHER A. SHAW Nanjing Institute of Geology and [email protected] DANIEL KSEPKA, LUCY E. EDWARDS FREDERICK C. SHAW Palaeontology, Bruce Museum Section Chairs ROBERT J. ELIAS LAWRENCE M. SMALL Chinese Academy of Sciences JP Editors Cordilleran: J. MARK ERICKSON GEORGE D. STANLEY, JR. JAMES LAMSDELL, BRENDA R. HUNDA, SHUNXIN ZHANG, NICOLE BONUSCO, DOUGLAS H. ERWIN ROBERT J. STANTON, JR. West Virginia University Cincinnati Museum Center Canada-Nunavut Geoscience Offi ce CA State Univ. Fullerton HOWARD R. FELDMANW CARL W. STOCK [email protected] ROGER D. K. THOMAS MAX LANGER, [email protected] ROBERT M. FINKS Paleobiology Senior Editor KARL W. FLESSA BRUCE H. TIFFNEY JISUO JIN, Universidade de São Paulo MARK PATZKOWSKY, North Central: TERRENCE J. FREST SALLY E. WALKER Univ. of Western Ontario STEPHEN A. LESLIE, Penn State Univ. JAMES THOMKA, PHILIP D. GINGERICH RON F. WASZCZAK [email protected] James Madison Univ. Paleobiology Co-Editors Univ. of Akron BRIAN F. GLENISTER JOHNNY A. WATERS [email protected] F. D. BUD HOLLAND, JR. GARY D. WEBSTER HANS-DIETER SUES, BRUCE LIEBERMAN, CATHERINE BADGLEY, J. STEWART HOLLINGSWORTH MARK A. WILSON Smithsonian Institution Univ. of Kansas Univ. of Michigan Northeastern: NIGEL C. HUGHES SCOTT L. WING [email protected] JUDITH NAGEL-MYERS, STEVEN LoDUCA, CARLOS A. JARAMILLO, THOMAS W. KAMMER ABBY YOCHELSON ELLEN CURRANO, Eastern Michigan Univ. Smithsonian Tropical Research St. Lawrence Univ. ERLE G. KAUFFMAN CHARLES YOCHELSON University of Wyoming Institute [email protected] PATRICIA H. KELLEY JEFFERY B. YOCHELSON RICK LUPIA, [email protected] WOLFGANG KIESSLING, SUSAN M. KIDWELL LOUIS G. ZACHOS Univer. of Oklahoma Rocky Mountain: Universitat Erlangen JP Associate Editors LEIF TAPANILA, NICHOLAS BUTTERFIELD, GABRIELA MÁNGANO, SHUHAI XIAO, Idaho State Univ. Th e Journal of Paleontology publishes manuscripts on all aspects of paleontology, including systematics, phylogeny, paleoecology, paleobiogeography, biostratigraphy, Cambridge Univ. Univ. Saskatchewan Virginia Polytechnic Institute [email protected] taphonomy, and evolution. It emphasizes specimen-based research and features illustrations of the highest quality possible. Manuscript treating all fossil organism Southeastern: STYLIANOS CHATZIMANOLIS, GREG McDONALD, groups are welcome, including invertebrates, vertebrates, plants, algae, and microorganisms of all sorts, as well as ichnofossils (trace fossils). Peer Review Coordinator MICHELLE CASEY, Univ. Tennessee–Chattanooga Bureau of Land Management JESSICA KASTIGAR, Murray State Univ. [email protected] Th e Journal of Paleontology accepts only online manuscript submissions through the website: TOM CRONIN, MALGORZATA [email protected] [email protected] USGS MOCZYDLOWSKA-VIDAL, https://mc.manuscriptcentral.com/paleo Editor – Priscum Uppsala Univ. Communications Offi cer WILLIAM A. DiMICHELE, MATT POWELL, ______TONY MARTIN, Smithsonian Institution ALISON MURRAY, Juniata College Univ. of Alberta Emory Univ. Author guidelines can be downloaded at: JAN OVE EBBESTAD [email protected] [email protected] Museum of Evolution, PAULA NOBLE, https://www.cambridge.org/core/journals/journal-of-paleontology/information/instructions-contributors Editor – Social Media Uppsala University Univ. of Nevada Program Coordinator MATTHEW CLAPHAM, PHOEBE COHEN, ______BRIAN PRATT, Univ. of California, Santa Cruz Williams College ROBERT J. ELIAS, For questions regarding online submissions, please contact the editorial offi ces at: Univ. of Manitoba University of Saskatchewan [email protected] [email protected] [email protected] ______Cover illustration: A representative selection of some Cambrian fossils presented in this Cambrian Explosion Special Issue. Central image: New Membership Information radiodontan oral cone (NIGPAS 162529) from the Wulongqing Formation, Gaoloufang, in Kunming, eastern Yunnan Province, South China. Members of the Paleontological Society automatically receive electronic subscriptions to Paleobiology and Zeng et al., 2017, Fig. 2.1. Specimen 7 cm in diameter. Bottom left : Discinella micans (NFM F-2475) internal view from Middle Shale member, the Journal of Paleontology as a benefi t of membership. To apply for individual membership, please go to Forteau Formation, Fox Cove, southern Labrador. Skovsted and Topper 2017, Fig. 3.9. Specimen is 0.9 mm in diameter. Bottom right: Kutorgina the Paleontological Society’s homepage at http://paleosoc.org/ where links to an electronic application, a printable paper application form, chengjiangensis (ELI BK-064A) from the Chengjiang fauna of China. Holmer et al., 2017, Fig. 2.1. Specimen is 12 mm in diameter. and a complete list of membership benefi ts are provided under the Membership heading. ISSN 0022-3360 ______

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Cambrian Explosion VOL 92 | NO 1 | 2018 Zhifei Zhang and Glenn Brock, Guest Editors

1 Zhifei Zhang and Glenn A. Brock New evolutionary and ecological advances in deciphering the Cambrian explosion of animal life 3 Jian Han, Guoxiang Li, Xing Wang, Xiaoguang Yang, Junfeng Guo, Osamu Sasaki, and Tsuyoshi Komiya Olivooides-like tube aperture in early Cambrian carinachitids (Medusozoa, Cnidaria) 14 Huaqiao Zhang, Andreas Maas, and Dieter Waloszek New material of scalidophoran worms in Orsten-type preservation from the Cambrian Fortunian Stage of South China

26 Shixue Hu, Bernd-D. Erdtmann, Michael Steiner, Yuandong Zhang, Fangchen Zhao, VOLUME 92 Zhiliang Zhang, and Jian Han Malongitubus: a possible pterobranch hemichordate from the early Cambrian of South China 33 Lars E. Holmer, Zhifei Zhang, Timothy P. Topper, Leonid Popov, and Thomas M. Claybourn The attachment strategies of Cambrian kutorginate brachiopods: the curious case of two pedicle openings and their phylogenetic signiﬁ cance 40 Han Zeng, Fangchen Zhao, Zongjun Yin, and Maoyan Zhu Ⅵ

A new radiodontan oral cone with a unique combination of anatomical features from the early NO 1 Cambrian Guanshan Lagerstätte, eastern Yunnan, South China 49 Yuning Yang, Xingliang Zhang, Yuanlong Zhao, Yiru Qi, and Linhao Cui New paleoscolecid worms from the early Cambrian north margin of the Yangtze Platform,

South China Ⅵ 59 Bing Pan, Timothy P. Topper, Christian B. Skovsted, Lanyun Miao, and Guoxiang Li PAGES 1-114 Occurrence of Microdictyon from the lower Cambrian Xinji Formation along the southern margin of the North China Platform 71 Christian B. Skovsted and Timothy P. Topper Mobergellans from the early Cambrian of Greenland and Labrador: new morphological details and implications for the functional morphology of mobergellans 80 Natalie I. Schroeder, John R. Paterson, and Glenn A. Brock Eldonioids with associated trace fossils from the lower Cambrian Emu Bay Shale Ⅵ Konservat-Lagerstätte of South Australia 2018 87 Yunhuan Liu, Qi Wang, Tiequan Shao, Huaqiao Zhang, Jiachen Qin, Li Chen, Yongchun Liang, Cheng Chen, Jiaqi Xue, and Xiaowen Liu New material of three-dimensionally phosphatized and microscopic cycloneuralians from the Cambrian Paibian Stage of South China 99 Stephen Pates, Allison C. Daley, and Bruce S. Lieberman Hurdiid radiodontans from the middle Cambrian (Series 3) of Utah 114 Retraction

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