Attendees of the 2014 Northeast Regional DSA Meeting. Photo by Bryan Pfeiffer.
Odonata of Black Moshannon State Park, Centre County, Pennsylvania
Clark N. Shiffer, 301 Farmstead Lane, State College, Pennsylvania, 16803
Abstract Museum (Deans, 2013). Two of the authors (CNS and HBW) attribute their deep interest in Odonata to the Black Moshannon State Park includes a variety of freshwa- Beattys who cultivated that interest early on with frequent ter wetlands that support a diversity of Odonata. It is situ- field trips and sharing of their extensive knowledge. ated 1900 feet (580 m) above sea level in central Pennsylva- nia. Since 1943, 96 species of Odonata have been observed The 2015 DSA meeting will be a great opportunity to within the park. We document the fauna that includes a showcase and explore several rich Odonata habitats such number of species of state and regional conservation con- as Bear Meadows (White, Beatty, and Beatty, 1968), Ten cern. Acre Pond (White, 1963; Shiffer and White, 1995; 2014), and others where almost 100 species have been reported Introduction over the years. One of the lesser-known habitats is Black Moshannon State Park, the focus of this article, which will The 2015 annual Dragonfly Society of the Americas (DSA) hopefully entice many to attend the 2015 DSA meeting. meeting will be held in central Pennsylvania, where the local dragonfly fauna has been sampled frequently for more than Black Moshannon State Park 70 years. The first indications of its rich and diverse fauna emerged from a publication by George H. Beatty (Beatty, Black Moshannon State Park (40.91°N, -78.06°W) in Rush 1946) in which he reported collections at Bear Meadows Township, Centre County, Pennsylvania, is nine miles (15 and a few other locations near State College. In 1955, he km) east of Philipsburg and about 20 miles (32 km) north- moved to the State College area and lived there with his west of State College. It contains a diverse complex of wife, Alice Ferguson Beatty, also an Odonata specialist. wetlands and supports a similarly diverse Odonata fauna. They and coauthors published 14 papers on Pennsylvania Black Moshannon Lake (250 A, 101 ha, elev. 1870 ft, 570 Odonata between 1968 and 1971, of which several relate m), a long-standing impoundment about two miles (3 km) to central Pennsylvania (Beatty, Beatty, and White, 1969a; in greatest length is the park’s centerpiece. It is cradled in a Beatty, Beatty, and Shiffer, 1969b; Beatty and Beatty, 1971). broad valley atop the Allegheny Plateau and is almost sur- While maintaining an interest in Odonata, the Beattys rounded by an extensive bog and heath complex. Numer- never published another paper on Odonata and devoted ous small streams, some with beaver ponds, feed the wet- much of their time to botanical pursuits and photography. lands and lake that below the dam form Black Moshannon Alice passed away in 1987 and George died in 2004. The Creek, a medium-sized rocky stream flowing north in a Beatty’s large Odonata collection is now housed at Penn- narrow forested valley. Close by, the Allegheny Front drops sylvania State University’s Stuart W. Frost Entomological 1300 feet (395 m) into Bald Eagle Valley in the Ridge and
Argia 26(4), 2014 7 Valley physiographic province. Ecologically, the area is perspective, the most thorough surveys have occurred in more typical of boreal habitats in glaciated regions hun- recent years. Thus, recent data provides a more robust indi- dreds of miles to the north than to the habitats fairly close cation of the Odonata fauna. Since 2000, 75 species have to the south and east. A map of the state park is available on been recorded by CNS. Several of these are undoubtedly line at
8 Argia 26(4), 2014 likely a resident species with a small population. It flies over Species that have not been found but could be present spatterdock patches at the lake in June and early July. There are several species that are not common in Pennsyl- Gomphus rogersi (Sable Clubtail) (S2). Although this spe- vania but, based on their distribution in Pennsylvania and cies has been reported only twice, its normal habitat of habitat preference, might be found in Black Moshannon small sandy-bottomed woodland streams with gentle flow State Park. Perhaps participants at the 2015 DSA meeting is often bypassed in surveys. It is likely a rare but resident can document several and thereby top the 100 species mark species in the park. for the park. Ten to look for include:
Libellula flavida (Yellow-sided Skimmer) (S1). Two males Aeshna clepsydra (Mottled Darner) (S2S3). This species of this southern species were seen in late June 2005. They is known from several counties in Pocono Mountains of were likely strays and not part of an established a breeding northeastern Pennsylvania. CNS once observed a drag- population. onfly following the shore of Black Moshannon Lake that appeared to be this species, but its identity was not con- Somatochlora incurvata (Incurvate Emerald) (S1). For many firmed. years, this species was unknown in Pennsylvania. However, once its preferred bog heath habitat and later-season flight Lanthus parvulus (Northern Pygmy Clubtail) (S3). period were recognized, it was looked for and discovered in This species is found on small cold-water streams in the Clinton County (Shiffer, 1969) and subsequently in several Pennsylvania mountains, but it often goes unseen. Black other counties in north-central and northeastern Penn- Moshannon State Park is within its range and has the req- sylvania. It has been seen in three different years at Black uisite habitats. Moshannon since its discovery there in 2002. It is undoubt- edly a resident that flies in places that are hard to access. Epitheca spinigera (Spiny Baskettail) (S1). A few records of this species exist in five northeastern counties of Penn- Gomphus descriptus (Harpoon Clubtail) (S1S2). This sylvania. Black Moshannon State Park is near the southern stream species was recorded in five years before 1970, often range limit of this species. commonly on Black Moshannon Creek in late May and June, but has not been reported since. Whether it is still a Somatochlora elongata (Ski-tipped Emerald) (S2). It is hard resident species is uncertain. to imagine that this species has not been found yet at Black Moshannon because there is plenty of suitable habitat and Calopteryx aequabilis (River Jewelwing) (S2). A single male the species is known from other places in Centre County was collected in 1954 by Stuart W. Frost. The species has not and adjacent counties. been seen since. While there may be populations further downstream on Black Moshannon Creek, finding it again Leucorrhinia glacialis (Crimson-ringed Whiteface) (S3S4). would be a significant discovery. As with the previous species, this species would be expected to be present in Black Moshannon State Park due to its Gomphaeschna furcillata (Harlequin Darner) (S2). This habitat preferences and known geographic range. typical bog species is well known from Tamarack Bog to the north and Bear Meadows (White, Beatty, and Beatty, Leucorrhinia proxima (Red-waisted Whiteface) (S2). 1968) to the south, yet it has been reported only twice from There are relatively few Pennsylvania records for this spe- Black Moshannon State Park where one might expect it to cies, but it has shown up at Ten Acre Pond (Shiffer and be a frequently-seen resident. White, 2014) and Beaver Dam (CNS, unpublished record), and is known from several northern counties . The fact that Somatochlora forcipata (Forcipate Emerald) (S2). Once this it looks a lot like the more common L. frigida (Frosted species was discovered as a resident in 2003, it was found Whiteface) might enable it to escape detection. in low numbers when looked for in late June and early July flying in sunny glades near the Moss-Hanne Trail. Pantala flavescens (Wandering Glider) (S5). This wide- spread migratory species can be expected to appear some- Somatochlora walshii (Brush-tipped Emerald) (S2). This time at any habitat in Pennsylvania. It is surprising that it species is sufficiently common to be found when it is looked has not been reported at least once over a parking lot or field for in its preferred habitat of grassy wet meadows, often in Black Moshannon State Park. near beaver ponds Tramea carolina (Carolina Saddlebags) (S4S5). As with the previous species, T. carolina migrates and could appear
Argia 26(4), 2014 9 in the park. There are breeding populations within Centre Academy of Science Proceedings, 43: 119–126. County. Beatty, G.H., A.F. Beatty, and C.N. Shifffer. 1969b. A sur- vey of the Odonata of central Pennsylvania. Pennsylva- Archilestes grandis (Great Spreadwing) (S4). Twice there nia Academy of Sciences Proceedings, 43: 127–136 have been unconfirmed reports of this late-season damsel- Beatty, G.H. and A.F. Beatty. 1971. The distribution of fly along tributaries near the Mid-State Regional Airport Pennsylvania Odonata. Pennsylvania Academy of Sci- on the west side of the park. ences Proceedings 45: 147–167. Deans, A. 2013. Dragonflies.
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10 Argia 26(4), 2014 Table 1. Decadal distribution of Odonata atDecadal Black Moshannon and Seasonal State Park,Distribution Centre County, of Odonata Pennsylvania at Black Moshannon State Park, Centre Co., Pennsylvania Observation Interval Seasonal Distribution by Monthly Quarters 1943 1950 1960 1970 1980 1990 2000 Tot May June July August September Oct Tot Early - Late Dates Species Scientific Name 1949 1959 1969 1979 1989 1999 2010 Yrs I II III IV I II III IV I II III IV I II III IV I II III IV I II Qtrs Calopteryx aequabilis 0 1 0 0 0 0 0 1 X 1 June 22 Calopteryx amata 0 0 3 2 2 0 1 8 X X X X X X X 7 June 8 - July 29 Calopteryx maculata 0 0 2 2 3 3 9 19 X X X X X X X X X X 10 June 8 - August 12 Lestes congener 0 0 1 0 0 0 0 1 X 1 September 13 Lestes dryas 0 0 1 0 0 1 0 2 X X 2 July 15 - July 31 Lestes eurinus 0 0 0 0 0 1 0 1 X 1 June 25 Lestes forcipatus 0 0 1 0 0 0 1 2 X X 2 Uuly 31 - August 17 Lestes inaequalis 1 1 1 0 1 1 2 7 X X X X X X 6 June 11 - August 4 Lestes rectangularis 1 1 2 1 1 1 6 13 X X X X X X X X X X 10 June 15 - September 10 Lestes unguiculatus 0 0 1 0 0 0 0 1 X 1 July 31 Lestes vigilax 0 0 1 1 0 2 4 8 X X X X X X X 7 June 11 - Sepember 8 Amphiagrion saucium 0 0 0 1 0 0 6 7 X X X X X X X 7 May 30 - August 15 Argia apicalis 0 0 1 2 0 1 0 4 X X X 3 July 11 - August 8 Argia fumipennis violacea 0 0 1 1 1 0 1 4 X X X X 4 June 25 - August 1 Argia moesta 0 0 1 0 0 0 2 3 X X 2 June 11 - June 30 Chromagrion conditum 0 2 5 2 2 1 7 19 X X X X X X X X X 8 May 15 - July 22 Enallagma antennatum 0 0 1 1 0 0 1 3 X 1 June 25 - June 30 Enallagma aspersum 0 0 1 2 2 1 0 6 X X X X 4 June 24 - August 1 Enallagma basidens 0 0 0 0 0 0 1 1 X 1 June 11 Enallagma civile 0 0 1 1 0 0 0 2 X X 2 July 24 - September 8 Enallagma annexum 0 0 1 1 0 0 0 2 X 1 June 24 - June 29 Enallagma divagans 0 0 1 0 0 0 0 1 X 1 June 25 Enallagma ebrium 0 0 1 0 0 0 1 2 X X 2 June 19 - June 27 Enallagma exsulans 0 0 0 1 1 0 0 2 X X X 3 June 29 - August 1 Enallagma geminatum 0 0 0 0 0 1 2 3 X X X X 4 June 27 - August 25 Enallagma hageni 0 2 4 2 2 2 10 22 X X X X X X X X X X 10 June 8 - August 25 Enallagma signatum 0 0 3 1 0 1 1 6 X X X X X 5 June 11 - Sepember 13 Enallagma traviatum 0 0 0 1 1 0 0 2 X X 2 June 24 - July 18 Enallagma vesperum 0 0 1 2 0 1 0 4 X X X X X X 6 June 24 - September 8 Ischnura hastata 0 0 0 0 0 0 2 2 X X X X X 5 June 12 - September 11 Ischnura posita 0 0 1 0 0 0 2 3 X X X 3 June 11 - July 30 Ischnura verticalis 3 3 3 3 3 3 8 26 X X X X X X X X X X X X X X X 14 May 15 - September 13 Nehalennia gracilis 0 1 0 0 1 0 3 5 X X X X X X 6 June 20 - August 15 Nehalennia irene 0 1 3 0 3 1 6 14 X X X X X X X X 8 June 8 - July 30 Aeshna canadensis 2 1 1 3 2 1 4 14 X X X X X X X X X 9 July 29 - September 30 Aeshna tuberculifera 0 0 0 1 1 0 4 6 X X X X X X X 7 August 1 - September 30 Aeshna umbrosa 0 0 1 2 1 2 5 11 X X X X X X X X X 9 June 20 - October 12 Aeshna verticalis 0 0 0 1 0 0 4 5 X X X X X X X X 8 July 10 - October 1 Anax junius 0 0 5 3 1 4 9 22 X X X X X X X X X X X X X X X X X X 17 May 6 - October 1 Basiaeschna janata 0 1 4 0 0 2 3 10 X X X X X 5 May 28 - July 11 Boyeria grafiana 0 0 0 0 2 2 0 4 X X 2 Augiust 1 - August 12 Boyeria vinosa 0 0 0 0 2 1 1 4 X X X X 4 July 29 - September 4 Epiaeschna heros 0 0 1 0 1 0 3 5 X X X X 4 June 9 - August 20 Gomphaeschna furcillata 0 0 0 1 0 0 1 2 X X 2 June 20 - June 24 Rhionaeschna mutata 0 0 0 1 0 1 6 8 X X X X X 5 June 11 -July 26 Arigomphus villosipes 0 1 4 2 0 0 6 13 X X X X X X X X 8 June 11 - August 20 Gomphus borealis 0 1 4 2 1 2 7 17 X X X X X X X 7 May 30 - July 25 Gomphus descriptus 0 2 3 0 0 0 0 5 X X X X 4 May 28 -June 24 Gomphus exilis 0 1 3 2 1 1 4 12 X X X X X 5 June 9 - July 13 Gomphus lividus 0 0 4 1 0 2 2 9 X X X X X X 6 May 20 - June 25 Gomphus rogersi 0 0 1 0 0 0 1 2 x x 2 June 23 - June 30 Gomphus spicatus 0 2 3 4 1 1 5 16 X X X X X X X 7 May 20 - July 17 Hagenius brevistylus 0 1 1 2 0 0 0 4 X X X 3 June 24 - July 27 Lanthus vernalis 0 0 1 0 0 0 0 1 X X X X 4 June 15 - July 14
Argia 26(4), 2014 11 Table 1, continued. Decadal distribution of Odonata at Black Moshannon State Park, Centre County, Penn- sylvania Decadal and Seasonal Distribution of Odonata at Black Moshannon State Park, Centre Co., Pennsylvania ObservationInterval Seasonal Distribution by Monthly Quarters 1943 1950 1960 1970 1980 1990 2000 Tot May June July August September Oct Tot Early - Late Dates Species Scientific Name 1949 1959 1969 1979 1989 1999 2010 Yrs I II III IV I II III IV I II III IV I II III IV I II III IV I II Qtrs Ophiogomphus mainensis 0 0 0 0 2 1 1 4 X X 2 June 10 - August 4 Stylogomphus albistylus 0 0 2 1 0 0 1 4 X X 2 June 11 - June 30 Cordulegaster diastatops 0 0 1 2 2 4 9 18 X X X X X X X X X X X 11 May 20 - August 15 Cordulegaster maculata 0 0 1 0 1 0 4 6 X X X X 4 May 28 - July 6 Cordulegaster obliqua 0 0 0 0 1 2 7 10 X X X X X X 6 June 8 - July 14 Didymops transversa 1 1 1 0 0 0 0 3 X X X 3 June 9 - July 4 Macromia illinoiensis 0 0 0 1 0 0 0 1 X 1 June 29 Cordulia shurtleffi 0 0 3 0 0 0 5 8 X X X X X X 6 June 8 - July 22 Dorocordulia libera 3 3 6 2 2 3 9 28 X X X X X X X X X X 10 May 30 - August 7 Epitheca canis 0 1 2 3 0 2 4 12 X X X X X X X X 6 May 7 - July 3 Epitheca cynosura 1 4 4 2 2 3 9 25 X X X X X X X X X 9 May 20 - July 30 Epitheca princeps 0 0 0 0 0 1 2 3 X X 2 June 27 - Aug 19 Helocordulia uhleri 0 0 4 2 0 0 1 7 X X X 3 May 28 - June 30 Somatochlora forcipata 0 0 0 0 0 0 4 4 X X X X 4 June 19 - July 10 Somatochlora incurvata 0 0 0 0 0 0 3 3 X X X X X X X 7 July 17- September17 Somatochlora tenebrosa 0 0 3 3 2 2 9 19 X X X X X X X X X X X X X X 14 June 15 - September 17 Somatochlora walshii 0 0 0 3 0 1 7 11 X X X X X X X X X X X 11 June 15 - September 8 Celithemis elisa 0 0 0 0 0 0 2 2 X X 2 June 15 - September 4 Celithemis eponina 0 0 0 0 0 0 1 1 X X 2 August 3 - August 15 Erythemis simplicicollis 0 0 0 0 0 0 4 4 X X X X 4 June 23 - August 27 Ladona julia 0 4 5 5 1 4 9 28 X X X X X X X X X X 10 May 28 - August 4 Leucorrhinia frigida 0 2 4 1 2 2 8 19 X X X X X X X X X X 10 June 11 - August 25 Leucorrhinia hudsonica 0 0 2 0 0 0 3 5 X X X X X 5 June 11 - July 13 Leucorrhinia intacta 0 3 1 2 1 3 8 18 X X X X X X X X X 9 June 8 - August 9 Libellula axilena 0 0 0 0 0 0 1 1 X X 2 June 15 - July 21 Libellula cyanea 0 0 0 0 0 0 4 4 X X X X 4 June 15 - August 15 Libellula flavida 0 0 0 0 0 0 1 1 X 1 June 25 Libellula luctuosa 0 0 0 0 0 2 3 5 X X X X X X 6 June 15 - August 4 Libellula pulchella 0 0 1 2 0 1 8 12 X X X X X X X X X X X X 12 May 30 - September 11 Libellula quadrimaculata 0 0 3 2 1 2 9 17 X X X X X X X X X 9 May 20 - August 4 Libellula semifasciata 1 0 2 1 0 1 2 7 X X X X X X X X X X 10 June 5 - August 23 Libellula vibrans 0 0 0 0 0 0 6 6 X 1 June 15 Plathemis lydia 1 0 1 1 0 2 6 11 X X X X X X X X X X X X X 13 May 25 - September 17 Pachydiplax longipennis 0 0 0 0 0 0 1 1 X X X X X X X X X X X X 12 June 11 - September 4 Pantala flavescens 0 0 0 0 0 0 1 1 X 1 September 5 Pantala hymenaea 2 0 1 1 0 3 0 7 X 1 July 13 Perithemis tenera 0 1 3 3 1 2 9 19 X X X X X X 6 June 30 - August 22 Sympetrum janeae 0 0 0 0 0 1 8 9 X X X X X X X X X X X 11 June 15 - September17 Sympetrum obtrusum 1 1 0 0 1 0 0 3 X X X X 4 July 6 - September10 Sympetrum semicinctum 0 0 0 2 1 1 6 10 X X X X X X X X 8 July 29 - September 17 Sympetrum vicinum 2 4 0 0 0 2 5 13 X X X X X X X X 8 July 21- September 17 Tramea lacerata 0 0 0 0 0 0 4 4 X X X X X X X 7 June 30 - October 1 Total Species 12 26 59 50 37 48 75 96 2 3 5 18 19 54 42 67 40 41 42 33 35 24 25 22 23 18 10 4 3 1 96 May 7 - October 12 Number of visits during interval 6 9 23 23 10 28 94 193 2 1 3 3 6 18 21 24 14 12 11 16 13 7 9 8 10 6 4 3 1 1 193
12 Argia 26(4), 2014 Table 2. Seasonal distribution of Odonata at Black Moshannon State Park, Centre County, Pennsylvania. Decadal and Seasonal Distribution of Odonata at BlackDecadal Moshannon and Seasonal State Park, Distribution Centre Co., of Odonata Pennsylvania at Black Moshannon State Park, Centre Co., Pennsylvania Observation Interval Observation Interval Seasonal Distribution by Monthly Quarters Seasonal Distribution by Monthly Quarters 1943 1950 1960 1970 1980 1990 2000 Tot 1943May 1950 1960 June1970 1980 1990July2000 Tot August May SeptemberJune Oct Tot JulyEarly - Late DatesAugust September Oct Tot Early - Late Dates Species Scientific Name 1949 1959 1969 1979 1989Species1999 Scientific2010 Yrs Name I 1949II III1959IV 1969I II1979III 1989IV I 1999II 2010III IVYrs I II I III II IV III I IV II I III II IV III I IV II I QtrsII III IV I II III IV I II III IV I II Qtrs Calopteryx aequabilis 0 1 0 0 0 Calopteryx0 aequabilis0 1 0 1 0 0 X 0 0 0 1 X 1 June 22 1 June 22 Calopteryx amata 0 0 3 2 2 Calopteryx0 amata1 8 0 0 X3 X 2 X X2 X 0 X 1 X 8 X X X X X 7 XJune 8 - JulyX 29 7 June 8 - July 29 Calopteryx maculata 0 0 2 2 3 Calopteryx3 maculata9 19 0 0 X2 X 2 X X3 X 3 X X9 X19 X X X X X X X 10 XJuneX 8 - AugustX X 12 X 10 June 8 - August 12 Lestes congener 0 0 1 0 0 Lestes0 congener0 1 0 0 1 0 0 0 0 1 X 1 September 13 X 1 September 13 Lestes dryas 0 0 1 0 0 Lestes1 dryas 0 2 0 0 1 0 0 1 X 0 X 2 2 XJuly 15 - JulyX 31 2 July 15 - July 31 Lestes eurinus 0 0 0 0 0 Lestes1 eurinus0 1 0 0 0 0 X0 1 0 1 X 1 June 25 1 June 25 Lestes forcipatus 0 0 1 0 0 Lestes0 forcipatus1 2 0 0 1 0 0 0 1 X 2 X 2 Uuly 31 - AugustX 17 X 2 Uuly 31 - August 17 Lestes inaequalis 1 1 1 0 1 Lestes1 inaequalis2 7 1 1 1 X 0 X X1 1 X2 X 7 X X X X 6 JuneX 11 - AugustX X 4 6 June 11 - August 4 Lestes rectangularis 1 1 2 1 1 Lestes1 rectangularis6 13 1 1 2 X 1 X1 1 X X6 X13 X X X X X X X 10 XJuneX 15 - SeptemberX X 10X X X X 10 June 15 - September 10 Lestes unguiculatus 0 0 1 0 0 Lestes0 unguiculatus0 1 0 0 1 0 0 0 0 X 1 1 July 31 X 1 July 31 Lestes vigilax 0 0 1 1 0 Lestes2 vigilax 4 8 0 0 1 X 1 X0 2 X X4 X 8 X X X X 7 XJuneX 11 - SepemberX X 8 X 7 June 11 - Sepember 8 Amphiagrion saucium 0 0 0 1 0 Amphiagrion0 saucium6 7 0 0 X 0 X 1 X X0 X 0 X 6 7 X X X X X X 7 XMay 30 - August 15 X 7 May 30 - August 15 Argia apicalis 0 0 1 2 0 Argia apicalis1 0 4 0 0 1 2 0 1 X X0 4 X 3 XJuly X11 - August X8 3 July 11 - August 8 Argia fumipennis violacea 0 0 1 1 1 Argia fumipennis0 1 violacea4 0 0 1 1 X1 X 0 X1 4 X X X 4 JuneX 25 - AugustX 1 4 June 25 - August 1 Argia moesta 0 0 1 0 0 Argia moesta0 2 3 0 0 1 X 0 X0 0 2 3 X X 2 June 11 - June 30 2 June 11 - June 30 Chromagrion conditum 0 2 5 2 2 Chromagrion1 conditum7 19 X0 2 X X5 X 2 X X2 X 1 X X7 19 X X X X X X X 8 XMayX 15 - July 22 8 May 15 - July 22 Enallagma antennatum 0 0 1 1 0 Enallagma0 antennatum1 3 0 0 1 1 X0 0 1 3 X 1 June 25 - June 30 1 June 25 - June 30 Enallagma aspersum 0 0 1 2 2 Enallagma1 aspersum0 6 0 0 1 2 X2 1 X0 X 6 X X 4 JuneX 24 - AugustX X 1 4 June 24 - August 1 Enallagma basidens 0 0 0 0 0 Enallagma0 basidens1 1 0 0 0 X 0 0 0 1 1 X 1 June 11 1 June 11 Enallagma civile 0 0 1 1 0 Enallagma0 civile0 2 0 0 1 1 0 0 0 X 2 X 2 July 24 - SeptemberX 8 X 2 July 24 - September 8 Enallagma annexum 0 0 1 1 0 Enallagma0 annexum0 2 0 0 1 1 X0 0 0 2 X 1 June 24 - June 29 1 June 24 - June 29 Enallagma divagans 0 0 1 0 0 Enallagma0 divagans0 1 0 0 1 0 X0 0 0 1 X 1 June 25 1 June 25 Enallagma ebrium 0 0 1 0 0 Enallagma0 ebrium1 2 0 0 1 0 X X0 0 1 2 X X 2 June 19 - June 27 2 June 19 - June 27 Enallagma exsulans 0 0 0 1 1 Enallagma0 exsulans0 2 0 0 0 1 X1 0 X 0 2 X X 3 XJune 29 - AugustX 1 3 June 29 - August 1 Enallagma geminatum 0 0 0 0 0 Enallagma1 geminatum2 3 0 0 0 0 X0 X 1 X2 3 X X X 4 JuneX 27 - August 25 X 4 June 27 - August 25 Enallagma hageni 0 2 4 2 2 Enallagma2 hageni10 22 0 2 4 X 2 X X2 X 2 X 10X X22 X X X X X X X 10 XJuneX 8 - AugustX X25 X X 10 June 8 - August 25 Enallagma signatum 0 0 3 1 0 Enallagma1 signatum1 6 0 0 3 X 1 X0 1 X1 6 X X X X 5 JuneX 11 - Sepember 13 X X 5 June 11 - Sepember 13 Enallagma traviatum 0 0 0 1 1 Enallagma0 traviatum0 2 0 0 0 1 X1 0 X0 2 X 2 JuneX 24 - July 18 2 June 24 - July 18 Enallagma vesperum 0 0 1 2 0 Enallagma1 vesperum0 4 0 0 1 2 X0 X 1 X0 X 4 X X X X 6 JuneX 24 - SeptemberX 8 X X 6 June 24 - September 8 Ischnura hastata 0 0 0 0 0 Ischnura0 hastata2 2 0 0 0 X 0 0 0 X 2 2 X X X X 5 XJune 12 - September 11 X X X 5 June 12 - September 11 Ischnura posita 0 0 1 0 0 Ischnura0 posita2 3 0 0 1 X 0 X0 0 2 X 3 X X 3 June 11 - JulyX 30 3 June 11 - July 30 Ischnura verticalis 3 3 3 3 3 Ischnura3 verticalis8 26 X3 3 X 3 X 3 X X3 X 3 X X8 X26 X X X X X X X X X X X X 14 XMayX 15 - SeptemberX X 13X X X X X 14 May 15 - September 13 Nehalennia gracilis 0 1 0 0 1 Nehalennia0 gracilis3 5 0 1 0 0 X X1 0 X X3 5 X X X X 6 XJuneX 20 - AugustX 15 X 6 June 20 - August 15 Nehalennia irene 0 1 3 0 3 Nehalennia1 irene6 14 0 1 X3 X 0 X X3 X 1 X X6 X14 X X X X X 8 XJuneX 8 - JulyX 30 8 June 8 - July 30 Aeshna canadensis 2 1 1 3 2 Aeshna1 canadensis4 14 2 1 1 3 2 1 4 X14 X X X X X X X X 9 July 29 - SeptemberX X 30X X X X X X X 9 July 29 - September 30 Aeshna tuberculifera 0 0 0 1 1 Aeshna0 tuberculifera4 6 0 0 0 1 1 0 4 6 X X X X X X X 7 August 1 - SeptemberX 30 X X X X X X 7 August 1 - September 30 Aeshna umbrosa 0 0 1 2 1 Aeshna2 umbrosa5 11 0 0 1 2 X 1 2 5 11 X X X X X X X X X 9 June 20 - OctoberX 12 X X X X X X X 9 June 20 - October 12 Aeshna verticalis 0 0 0 1 0 Aeshna0 verticalis4 5 0 0 0 1 0 0 X 4 5 X X X X X X X 8 XJuly 10 - October 1 X X X X X X X 8 July 10 - October 1 Anax junius 0 0 5 3 1 Anax junius4 9 22 X 0 0 X X5 X 3 X X1 X 4 X X9 X22 X X X X X X X X X X X X X X X 17 XMayX 6 - OctoberX X 1 X X X X X X X 17 May 6 - October 1 Basiaeschna janata 0 1 4 0 0 Basiaeschna2 janata3 10 0 1 X X4 X 0 X 0 2 X 3 10 X X X X 5 XMay 28 - July 11 5 May 28 - July 11 Boyeria grafiana 0 0 0 0 2 Boyeria2 grafiana0 4 0 0 0 0 2 2 0 4 X X 2 Augiust 1 - AugustX 12X 2 Augiust 1 - August 12 Boyeria vinosa 0 0 0 0 2 Boyeria1 vinosa1 4 0 0 0 0 2 1 1 X 4 X X X 4 July 29 - SeptemberX X 4X X 4 July 29 - September 4 Epiaeschna heros 0 0 1 0 1 Epiaeschna0 heros3 5 0 0 1 X 0 X1 0 3 5 X X X X 4 June 9 - August X20 X 4 June 9 - August 20 Gomphaeschna furcillata 0 0 0 1 0 Gomphaeschna0 1 furcillata2 0 0 0 1 X X0 0 1 2 X X 2 June 20 - June 24 2 June 20 - June 24 Rhionaeschna mutata 0 0 0 1 0 Rhionaeschna1 6 mutata8 0 0 0 X 1 X X0 X 1 X6 8 X X X X 5 JuneX 11 -July 26 5 June 11 -July 26 Arigomphus villosipes 0 1 4 2 0 Arigomphus0 villosipes6 13 0 1 4 X 2 X X0 X 0 X X6 X13 X X X X X 8 XJuneX 11 - AugustX 20 X 8 June 11 - August 20 Gomphus borealis 0 1 4 2 1 Gomphus2 borealis7 17 0 1 X X4 X 2 X X1 X 2 7 X17 X X X X X X 7 May 30 - JulyX 25 7 May 30 - July 25 Gomphus descriptus 0 2 3 0 0 Gomphus0 descriptus0 5 0 2 X 3 X 0 X X0 0 0 5 X X X X 4 May 28 -June 24 4 May 28 -June 24 Gomphus exilis 0 1 3 2 1 Gomphus1 exilis4 12 0 1 X3 X 2 X X1 1 X 4 12 X X X X 5 XJune 9 - July 13 5 June 9 - July 13 Gomphus lividus 0 0 4 1 0 Gomphus2 lividus2 9 0 X 0 X X4 X 1 X X0 2 2 9 X X X X X X 6 May 20 - June 25 6 May 20 - June 25 Gomphus rogersi 0 0 1 0 0 Gomphus0 rogersi1 2 0 0 1 0 x x0 0 1 2 x x 2 June 23 - June 30 2 June 23 - June 30 Gomphus spicatus 0 2 3 4 1 Gomphus1 spicatus5 16 0 2 X 3 X 4 X X1 X 1 X X5 16 X X X X X 7 XMayX 20 - July 17 7 May 20 - July 17 Hagenius brevistylus 0 1 1 2 0 Hagenius0 brevistylus0 4 0 1 1 2 X0 0 X X0 4 X 3 XJuneX 24 - July 27 3 June 24 - July 27 Lanthus vernalis 0 0 1 0 0 Lanthus0 vernalis0 1 0 0 1 X 0 X X0 0 X 0 1 X X X 4 XJune 15 - July 14 4 June 15 - July 14
Argia 26(4), 2014 13 Table 2. Seasonal distribution of Odonata at Black Moshannon State Park, Centre County, Pennsylvania. Decadal and Seasonal Distribution of Odonata at BlackDecadal Moshannon and Seasonal State Park, Distribution Centre Co., of Odonata Pennsylvania at Black Moshannon State Park, Centre Co., Pennsylvania ObservationInterval ObservationInterval Seasonal Distribution by Monthly Quarters Seasonal Distribution by Monthly Quarters 1943 1950 1960 1970 1980 1990 2000 Tot 1943May1950 1960 1970June 1980 1990 July2000 Tot AugustMay SeptemberJune Oct TotJulyEarly - Late DatesAugust September Oct Tot Early - Late Dates Species Scientific Name 1949 1959 1969 1979 1989Species1999Scientific2010 NameYrs I 1949II 1959III IV 1969I 1979II III 1989IV 1999I II 2010III YrsIV I III IIIII IVIII IIV III IIIII IVIII IIV III QtrsII III IV I II III IV I II III IV I II Qtrs Ophiogomphus mainensis 0 0 0 0 2Ophiogomphus1 1mainensis4 0 0 0 X0 2 1 1 4 X X 2 June 10 - AugustX 4 2 June 10 - August 4 Stylogomphus albistylus 0 0 2 1 0Stylogomphus0 albistylus1 4 0 0 2 X1 0X 0 1 4 X X 2 June 11 - June 30 2 June 11 - June 30 Cordulegaster diastatops 0 0 1 2 2Cordulegaster4 diastatops9 18 0 X0 X 1X X2 X 2X X4 X 9X 18X X X X X X X X X 11X MayX 20 -X August 15 X 11 May 20 - August 15 Cordulegaster maculata 0 0 1 0 1Cordulegaster0 maculata4 6 0 0 X 1 X0 1X X0 4 6 X X X X 4 May 28 - July 6 4 May 28 - July 6 Cordulegaster obliqua 0 0 0 0 1Cordulegaster2 obliqua7 10 0 0 0X X0 X 1X X2 X 7 10 X X X X X 6X June 8 - July 14 6 June 8 - July 14 Didymops transversa 1 1 1 0 0Didymops0 transversa0 3 1 1 1 X0 0X X0 0 3 X X X 3 June 9 - July 4 3 June 9 - July 4 Macromia illinoiensis 0 0 0 1 0Macromia0 illinoiensis0 1 0 0 0 1 0 X0 0 1 X 1 June 29 1 June 29 Cordulia shurtleffi 0 0 3 0 0Cordulia0shurtleffi5 8 0 0 3X X0 X 0X 0 X 5X 8 X X X X 6X JuneX 8 - July 22 6 June 8 - July 22 Dorocordulia libera 3 3 6 2 2Dorocordulia3 libera9 28 3 3 X 6X X2 X 2X X3 X 9X 28X X X X X X X X 10X MayX 30 -X AugustX 7 10 May 30 - August 7 Epitheca canis 0 1 2 3 0Epitheca2canis 4 12 X 0X X1 X 2X X3 X 0X 2 4 12 X X X X X X X X 6 May 7 - July 3 6 May 7 - July 3 Epitheca cynosura 1 4 4 2 2Epitheca3cynosura9 25 1 X4 4X X2 X 2X X3 X 9X 25X X X X X X X 9X MayX 20 -X July 30 9 May 20 - July 30 Epitheca princeps 0 0 0 0 0Epitheca1princeps2 3 0 0 0 0 0X 1 2 3 X X 2 June 27 - Aug 19 X 2 June 27 - Aug 19 Helocordulia uhleri 0 0 4 2 0Helocordulia0 uhleri1 7 0 0 X 4 X2 0X 0 1 7 X X X 3 May 28 - June 30 3 May 28 - June 30 Somatochlora forcipata 0 0 0 0 0Somatochlora0 forcipata4 4 0 0 0 0 X 0X X0 X 4 4 X X X 4X June 19 - July 10 4 June 19 - July 10 Somatochlora incurvata 0 0 0 0 0Somatochlora0 incurvata3 3 0 0 0 0 0 0 3X 3 X X X X X X 7 JulyX 17- September17X X X X X X 7 July 17- September17 Somatochlora tenebrosa 0 0 3 3 2Somatochlora2 tenebrosa9 19 0 0 3 X3 X 2X X2 X 9X 19X X X X X X X XX X X X 14X JuneX 15 X- SeptemberX X 17 X X X X X 14 June 15 - September 17 Somatochlora walshii 0 0 0 3 0Somatochlora1 walshii7 11 0 0 0 X3 X 0X X1 X 7X 11X X X X X X X X X 11X JuneX 15 -X SeptemberX 8 X X X 11 June 15 - September 8 Celithemis elisa 0 0 0 0 0Celithemis0 elisa 2 2 0 0 0 X0 0 0 2 2 X X 2 June 15 - September 4 X 2 June 15 - September 4 Celithemis eponina 0 0 0 0 0Celithemis0 eponina1 1 0 0 0 0 0 0 1 1 X X 2 August 3 - AugustX 15X 2 August 3 - August 15 Erythemis simplicicollis 0 0 0 0 0Erythemis0 simplicicollis4 4 0 0 0 0 0 X0 4X X4 X X 4 JuneX 23 -X August 27 X 4 June 23 - August 27 Ladona julia 0 4 5 5 1Ladona 4julia 9 28 0 4 X 5X X5 X 1X X4 X 9X 28X X X X X X X X 10X MayX 28 -X AugustX 4 10 May 28 - August 4 Leucorrhinia frigida 0 2 4 1 2Leucorrhinia2 frigida8 19 0 2 4 X1 X 2X X2 X 8X 19 X X X X X X X X 10X JuneX 11 - AugustX 25 X X X 10 June 11 - August 25 Leucorrhinia hudsonica 0 0 2 0 0Leucorrhinia0 hudsonica3 5 0 0 2 X0 X 0X X0 3X 5 X X X X 5 JuneX 11 - July 13 5 June 11 - July 13 Leucorrhinia intacta 0 3 1 2 1Leucorrhinia3 intacta8 18 0 3 1X X2 X 1X X3 X 8X 18 X X X X X X X 9X JuneX 8 - AugustX 9 X 9 June 8 - August 9 Libellula axilena 0 0 0 0 0Libellula0axilena1 1 0 0 0 X0 0 0 1X 1 X 2 JuneX 15 - July 21 2 June 15 - July 21 Libellula cyanea 0 0 0 0 0Libellula0cyanea4 4 0 0 0 X0 0X 0 X 4 4 X X X 4X June 15 - August 15X 4 June 15 - August 15 Libellula flavida 0 0 0 0 0Libellula0flavida 1 1 0 0 0 0 0X 0 1 1 X 1 June 25 1 June 25 Libellula luctuosa 0 0 0 0 0Libellula2luctuosa3 5 0 0 0 X0 X 0X X2 3X 5 X X X X X 6 JuneX 15 - AugustX 4 6 June 15 - August 4 Libellula pulchella 0 0 1 2 0Libellula1pulchella8 12 0 0 X 1 X2 X 0X X1 X 8X 12 X X X X XX X X X X 12X MayX 30 - SeptemberX X 11 X X X 12 May 30 - September 11 Libellula quadrimaculata 0 0 3 2 1Libellula2quadrimaculata9 17 0 X0 X 3 X2 X 1X X2 X 9X 17 X X X X X X X 9X MayX 20 - AugustX 4 9 May 20 - August 4 Libellula semifasciata 1 0 2 1 0Libellula1semifasciata2 7 1 0 2X X1 X 0X X1 X 2X X7 X X X X X X X 10X JuneX 5 - XAugust 23 X X 10 June 5 - August 23 Libellula vibrans 0 0 0 0 0Libellula0vibrans6 6 0 0 0 X0 0 0 6 6 X 1 June 15 1 June 15 Plathemis lydia 1 0 1 1 0Plathemis2 lydia 6 11 1 0 X 1X X1 X 0X X2 X 6X 11 X X X X X XX X X X X 13X MayX 25 - SeptemberX X 17 X X X 13 May 25 - September 17 Pachydiplax longipennis 0 0 0 0 0Pachydiplax0 longipennis1 1 0 0 0 X0 X 0X X0 X 1X X1 X X X X X X X X X 12X JuneX 11 -X SeptemberX X 4 X X X 12 June 11 - September 4 Pantala flavescens 0 0 0 0 0Pantala 0flavescens1 1 0 0 0 0 0 0 1 1 X 1 September 5 X 1 September 5 Pantala hymenaea 2 0 1 1 0Pantala 3hymenaea0 7 2 0 1 1 0 3 X 0 7 1X July 13 1 July 13 Perithemis tenera 0 1 3 3 1Perithemis2 tenera9 19 0 1 3 3 1X X2 X 9 19X X X X X 6X June 30 -X AugustX 22 X 6 June 30 - August 22 Sympetrum janeae 0 0 0 0 0Sympetrum1 janeae8 9 0 0 0 X0 0X X1 8X X9 X X X X X X X X X 11 JuneX 15 X- September17X X X X X X 11 June 15 - September17 Sympetrum obtrusum 1 1 0 0 1Sympetrum0 obtrusum0 3 1 1 0 0 1 X0 0 X3 X X X 4 July 6 - September10X X X 4 July 6 - September10 Sympetrum semicinctum 0 0 0 2 1Sympetrum1 semicinctum6 10 0 0 0 2 1 1 6 10X X X X X X X X 8 July 29 - XSeptemberX X17 X X X X X 8 July 29 - September 17 Sympetrum vicinum 2 4 0 0 0Sympetrum2 vicinum5 13 2 4 0 0 0 2 5X 13 X X X X X X X 8 JulyX 21- SeptemberX 17 X X X X X X 8 July 21- September 17 Tramea lacerata 0 0 0 0 0Tramea 0lacerata 4 4 0 0 0 0 0X X0 4 4 X X X X XX X 7 June 30 - October 1 X X X X X 7 June 30 - October 1 Total Species 12 26 59 50 37Total Species48 75 96 2 123 526 18 5919 5450 42 3767 4048 41 7542 3396 35 242 253 225 2318 1819 1054 442 367 140 9641 May42 7 -33 October35 1224 25 22 23 18 10 4 3 1 96 May 7 - October 12 Number of visits during interval 6 9 23 23 10Number28 of visits94 during 193interval 2 61 39 3 236 1823 21 1024 1428 12 9411 19316 13 72 91 83 103 66 418 321 124 114 19312 11 16 13 7 9 8 10 6 4 3 1 1 193
Common Names of Species Enallagma antennatum Rainbw Bluet E. aspersum Azure Bluet Calopteryx aequabilis River Jewelwing E. basidens Double-striped Bluet C. amata Superb Jewelwing E. civile Familiar Bluet C. maculata Ebony Jewelwing E. annexum Northern Bluet Lestes congener Spotted Spreadwing E. divagans Turquoise Bluet L. dryas Emerald Spreadwing E. ebrium Marsh Bluet L. eurinus Amber-winged Spreadwing E. exsulans Stream Bluet L. forcipatus Sweetflag Spreadwing E. geminatum Skimming Bluet L. inaequalis Elegant Spreadwing E. hageni Hagen's Bluet L. rectangularis Slender Spreadwing E. signatum Orange Bluet L. unguiculatus Lyre-tipped Spreadwing E. triviatum Slender Bluet L. vigilax Swamp Spreadwing E. vesperum Vesper Bluet Amphiagrion saucium Eastern Red damsel Ischnura hastata Citrine Forktail Argia apicalis Blue-fronted Dancer I. posita Fragile Forktail A. fumipennis violacea Varianble Dancer I. verticalis Eastern Forktail A. moesta Powdered Dancer Nehalennia gracilis SphagnumSprite Chromagrion conditum Aurora Damsel N. irene Sedge Sprite
14 Argia 26(4), 2014 Common Names of Species, continued E. cynosura Common Baskettail E. princeps Prince Baskettail Aeshna canadensis Canada Darner Helocurdulia uhleri Uhler’s Sanddragon A. tuberculifera Black-tipped Darner Somatochlora forcipata Forcipate Emerald A. umbrosa Shadow Darner S. incurvata Incurvate Emerald A. verticalis Green-striped Darner S. tenebrosa Clamp-tipped Emerald Anax junius Common Green Darner S. walshii Brush-tipped Emerald Basiaeschna janata Springtime Darner Celithemis elisa Calico Pennant Boyeria grafiana Ocellated Darner C. eponina Halloween Pennant B. vinosa Fawn Darner Erythemis simplicicollis Eastern Pondhawk Epiaeschna heros Swamp Darner Ladona julia Chalk-fronted Corporal Gomphaeschna furcillata Harlequin Darner Leucorrhinia frigida Frosted Whiteface Rhionaeschna mutata Spatterdock Darner L. hudsonica Hudsonian Whiteface Arigomphus villosipes Unicorn Clubtail L. intacta Dot-tailed Whiteface Gomphus borealis Beaverpond Clubtail Libellula axilena Bar-winged Skimmer G. descriptus Harpoon Clubtail L. cyanea Spangled Skimmer G. exilis Lancet Clubtail L. flavida Yellow-sided Skimmer G. lividus Ashy Clubtail L. luctuosa Widow Skimmer G. rogersi Sable Clubtail L. pulchella Twelve-spotted Skimmer G. spicatus Dusky Clubtail L. quadrimaculata Four-spotted Skimmer Hagenius brevistylus Dragonhunter L. semifasciata Painted Skimmer Lanthus vernalis Southern Pygmy Clubtail L. vibrans Great Blue Skimmer Ophiogomphus mainensis Maine Snaketail Plathemis lydia Common Whitetail Stylogomphus albistylus Eastern Least Clubtail Pachydiplax longipennis Blue Dasher Cordulegaster diastatops Delta-spotted Spiketail Pantala flavescens Wandering Glider C. maculata Twin-spotted Spiketail Pantala hymenaea Spot-winged Glider C. obliqua Arrowhead Spiketail Perithemis tenera Eastern Amberwing Didymops transversa Stream Cruiser Sympetrum janeae (internum) Cherry-faced Meadowhawk Macromia illinoiensis Swift River Cruiser Sympetrum obtrusum White-faced Meadowhawk Cordulia shurtleffii American Emerald S. semicinctum Band-winged Meadowhawk Dorocordulia libera Racket-tailed Emerald S. vicinum Autumn Meadowhawk Epitheca canis Beaverpond Baskettail Tramea lacerata Black Saddlebags
Register Now for the 2015 DSA Central American Regional Meeting! DSA will host its first-ever Central American Regional Meeting in Costa Rica from 31 May–9 June 2015. This is an eight day/nine night meeting in the land of Pseudostigmatidae, resplendent quetzels, and poison dart frogs. The meeting will involve field excursions through the Organization of Tropical Studies (OTS) at two of their field stations in different ecoregions: La Selva, in the Caribbean Lowlands of northern Costa Rica in an area of tropical and premontane wet forest; and Palo Verde, an area in northwestern Costa Rica with deciduous dry forest, and freshwater marshes and wetlands bordering the Tempisque River. Dennis Paulson will be our keynote speaker and main guide, and Ken Tennessen will also present and lead guided field excursions. Bill Haber will also be joining us for the LaSelva portion of the trip. Don’t miss a rare opportunity to learn from an A-team of odonatologists in a gorgeous tropical setting in one of the most biodiverse counties on Earth!
Seats are limited, so don’t delay. Full details regarding the daily itinerary, costs, registration, and more can be found on the meeting web site at
Registration must be completed by 1 February 2015, and deposits must be recived by mid-February. Spaces are filling up fast, so be sure to check out the details soon!
Argia 26(4), 2014 15