FORTHCOMING THIS FALL

Key Updates • Full integration of • Extensively revised • A greatly expanded molecular systematics treatments of 13 families online Photo Gallery Include: into Chapter 2, Methods that have undergone of Vascular Plants. and Principles of significant changes in Biological Systematics. circumscription. SAMPLE PAGES

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For the Student For the Instructor (available to qualified adopters) Photo Gallery of Vascular Plants A Companion to Plant Systematics: A Phylogenetic Instructor’s Resource Library Approach, Fourth Edition The Plant Systematics Instructor’s Resource Library Walter S. Judd, Daniel L. Nickrent, Kenneth R. Robert- includes a collection of visual resources from the textbook son, J. Richard Abbott, Christopher S. Campbell, Barbara for use in preparing lectures and other course materials. S. Carlsward, Tanja M. Schuster, Kurt M. Neubig, Scott The textbook figures have all been sized and formatted Zona, Michael J. Donoghue, and Elizabeth A. Kellogg for optimal legibility when projected. The IRL includes all textbook figures, tables, and floral formulas in JPEG (both The Photo Gallery of Vascular Plants has been greatly high- and low-resolution) and PowerPoint formats. expanded for the Fourth Edition of Plant Systematics, and is now available online. Over 9,700 color photos illustrat- ing the diagnostic characters of (and variability within) For detailed contents the families covered in the text, including or to request an many images showing floral and fruit dissections, are now examination copy, included in the site. The Photo Gallery also includes an visit our website: illustrated glossary of plant terminology. Each new text- book includes an access code for the site, and access may sinauer.com also be purchased separately. Plant Systematics, Fourth Edition Walter S. Judd, Christopher S. Campbell, Elizabeth A. Kellogg, Peter F. Stevens, and Michael J. Donoghue

ABOUT THE BOOK A comprehensive introduction to vascular plant phylogeny, the Fourth Edition of Plant Systematics reflects changes in the circumscription of several families in order to represent monophyletic groups, following the classification of the Angiosperm Phylogeny Group (and recent phyloge- netic analyses). Appendices cover botanical nomenclature as well as field and herbarium methodology. The text is copiously illustrated, using in large part the informative analytical drawings developed as part of the Generic Flora of the Southeastern United States project. Plant Systematics: A Phylogenetic Approach, Fourth Edition is ap- propriate for any course devoted to the systematics of angiosperms or vascular plants and, secondarily, for local flora courses. The text assumes no prerequisites other than introductory botany or biology.

November 2015 • 575 pages (est.) 337 illustrations • 27 plates ISBN 978-1-60535-389-0 • casebound New to the Fourth Edition

• Chapter 2, Methods and • Extensive revisions to 13 fami- Principles of Biological lies that have undergone signifi- AFFORDABLE VALUE OPTIONS Systematics, has been com- cant changes in circumscription pletely rewritten with informa- have been implemented. Revi- FOR EVERY FORMAT! tion on molecular (DNA-based) sions include the breakup of systematics now fully integrat- the non-monophyletic “Wood- Print Edition ed—a change required by the in- siaceae,” “Portulacaceae,” and • Order from our website for a 15% discount creasing importance of molecu- “Clusiaceae,” the segregation from suggested list price.* ($93.46) lar methods in the construction of Cleomaceae and Capparaceae • Free standard ground shipping to U.S. of phylogenetic hypotheses. The from Brassicaceae s.l, Cabomba- addresses on orders $40.00 and up. chapter includes updated and ceae from Nymphaeaceae, Nys- • Orders usually ship in 1–4 days. expanded coverage of maximum saceae from Cornaceae, and Vis- • Offer not available to resellers. likelihood and Bayesian meth- caceae from Santalaceae s.l. In ods, algorithmic approaches to addition, Illiciaceae are included eBooks tree construction, along with in Schisandraceae, Hydnoraceae Suggested list discounted 50% from bound parsimony methods, and new and Lactoridaceae in Aristolo- book list price* for a 180-day subscription. ($54.98) sections on mapping characters chiaceae, Centrolepidaceae and on trees and dating phylogenies. Anarthriaceae in Restionaceae, Suggested list discounted 15% from bound book list price* to own permanently. Turneraceae in Passifloraceae, ($93.46) • Sections on speciation and Zannichelliaceae in Potamoge- Formats include BryteWave, RedShelf, Vital- species concepts have been tonaceae, and Memecylaceae in Source CourseSmart, and YUZU. rewritten to incorporate exten- Melastomataceae. Several familial sive recent research on these clades are for the first time *$109.95 Suggested list price topics. included in the book: Balsamina- $87.96 Net price to resellers ceae, Chrysobalanaceae, Fouqui- eriaceae, Garryaceae, Goodenia- Prices subject to change May 1 and November 1, yearly. ceae, Hypoxidaceae, Musaceae, Phrymaceae, Pittosporaceae, and Thymelaeaceae. CONTENTS Plant Systematics, Fourth Edition Walter S. Judd, Christopher S. Campbell, Elizabeth A. Kellogg, Peter F. Stevens, and Michael J. Donoghue 1. The Science of Plant Systematics What Do We Mean by Plant? THE AUTHORS What Do We Mean by Systematics? The Phylogenetic Approach Walter S. Judd is Distinquished Professor Emeritus in the Department The Practice of Plant Systematics of Biology at the University of Florida. Why Is Systematics Important? Christopher S. Campbell is Professor Emeritus of Botany at the Aims and Organization of This Book University of Maine. 2. Methods and Principles of Elizabeth A. Kellogg is a Member at the Donald Danforth Plant Science Biological Systematics Center in St. Louis, Missouri. Discovering Phylogeny: How Phylogenetic Trees Are Constructed Peter F. Stevens is Professor Emeritus in the Department of Biology • Evolutionary Trees and What They at the University of Missouri, St. Louis, and Curator of the Missouri Depict. Reading the Tree. Botanical Garden. • Rooting Michael J. Donoghue is Yale University’s Sterling Professor of Ecology • Homoplasy • Tree Building: Clustering Methods and Evolutionary Biology and Curator of Botany at the Peabody Museum. and Neighbor-Joining • Models of Evolution • Should You Believe the Tree? Boot- strapping and Comparison of Trees • Methods That Use an Optimality Criterion • Higher-level phylogenetic rela- • A greatly expanded online Photo Parsimony tionships have been updated on the Gallery of Vascular Plants in- ° Maximum Likelihood basis of recently published analyses, cludes over 9,700 photographs ° ° Bayesian Methods and these relationships are reflected illustrating the diagnostic characters • Comparing Trees from Different in the authors’ use of names; e.g., and morphological diversity of each Methods and Sources of Data within the flowering plants we now of the familial clades covered in the Using Phylogenetic Trees recognize Mesangiospermae, Pen- textbook. The photos also include • Constructing a Classification tapetalae, and the Superrosid and taxa of both temperate and tropical ° Grouping: Named Groups Are Superasterid clades. distribution, economically impor- Monophyletic tant species, and examples of impor- ° Naming: Not All Groups Are • Updates to all chapters and to tant anatomical characteristics. Named nearly all cladograms take into ° Ranking: Ranks Are Arbitrary account recent taxonomic meth- ° Comparing Phylogenetic ods and hypotheses. In addition, Classifications with Those several new cladograms and figures Derived Using Other have been added. Since most Taxonomic Methods families have received new phylo- • Describing Evolution: Mapping Characters on Trees genetic study since the previous For detailed contents • Dating Phylogenies edition, the summaries of phyloge- or to request an netic patterns within each of these examination copy, 3. Classification and System in familial clades and their putative visit our website: Flowering Plants: Historical synapomorphies have been up- sinauer.com Background dated, including many changes in Classification, Nature, and Stability generic circumscription. Botany, Classification, and Biology Understanding Relationships Classifications and Memory The Formation of Higher Taxa Plant Groupings over the Years CONTENTS

4. Taxonomic Evidence: Structural 8. Phylogenetic Relationships of • Fabales and Biochemical Characters Angiosperms • Rosales Morphology ANA Grade (Amborellales, • Cucurbitales Pollination Biology Nymphaeales, Austrobaileyales) • Fagales Inflorescences, Fruits, and Seeds • Amborellales ° Malvids (or Euro- Anatomy • Nymphaeales sids II; Malvidae) Embryology • Austrobaileyales • Geraniales Chromosomes Mesangiospermae (including • Myrtales Palynology , Monocots, ) • Brassicales Secondary Metabolites • Chloranthales (of uncertain position, • Malvales Proteins possibly sister to Magnoliids) • Picramniales • Magnoliid Clade (Magnoliidae) • Sapindales 5. The Evolution of Plant Diversity ° Magnoliales • Superasteridae (Caryo- Plant Diversity Is the Result of Evolution ° Laurales phyllales, Santalales, and Variation in Plant Populations and ° Canellales Asterid Clade) Species ° Piperales • Caryophyllales Speciation • Ceratophyllales (placement • Santalales Origins of Reproductive Isolating uncertain) • Asterid Clade (=Sym- Barriers • Monocots (Monocotyledoneae) petalae or Asteridae) Species Concepts ° Acorales ° Cornales ° Alismatales ° Ericales 6. An Overview of Green Plant ° Liliales ° Lamiids (or Phylogeny ° Asparagales Euasterids I, Endosymbiotic Events ° Dioscoreales Lamiidae) “Algae” ° Commelinoid Monocots • Garryales Viridophytes (Green Plants) (Commelinidae) • Solanales (Land Plants) • Arecales • Gentianales Tracheophytes (Vascular Plants) • Commelinales • Boraginales (Seed Plants) • Poales • Lamiales Angiosperms (Flowering Plants) • Zingiberales ° Campanulids (or • Eudicots (Eudicotyledoneae) Euasterids II; 7. , , and ° “Basal Tricolpates” Campanulidae) • Ranunculales • Aquifoliales Lycophytes • Proteales • Apiales • Lycopodiales • Trochodendrales • Dipsacales Monilophytes (Ferns) • Buxales • Asterales • Psilotales ° Core Eudicots (Gunneridae) • Ophioglossales • Gunnerales Appendix 1. Botanical Nomenclature • Equisetales ° Pentapetalae (includes Super- Appendix 2. Specimen Preparation and • Leptosporangiate Ferns rosids and Superasterids) Identification ° Osmundales • Superrosidae (includes Glossary ° Salviniales Saxifragales and Rosid Taxonomic Index °  Clade) Subject Index °  • Saxifragales • Eupolypods I • Rosid Clade (Rosidae) • Eupolypods II ° Vitales Gymnosperms ° Fabids (or Euro- • Cycadales sids I; Fabidae) •  • Zygophyllales • Coniferales • Oxalidales • Gnetales • Celastrales • Malpighiales PRSRT STD Sinauer Associates, Inc., Publishers U.S. Postage Paid 23 Plumtree Road Greenfield, MA PO Box 407 Permit No. 183 Sunderland, MA 01375-0407 sinauer.com

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