Further records and descriptions of

Bactra species (, ),

chiefly in Dr. H.G. Amsel collection

by

A. Diakonoff

(Rijksmuseum van Natuurlijke Historie, Leiden)

The short forms of and records 15 present paper a continuation my species, or notes on some already

of of the known with five partial revision certain species genus species, new synonyms. the (Zool. Verhand., no. 29, 1956). It is chiefly based Continued study revealed to be gratifying, as

revision the entire material of in the Bactra to on a of Bactra genus more and more appears represent a collection of the well-known of field for taxonomie studies, due specialist Microlepi- very attractive chiefly the doptera, Dr. H. G. AMSEL, Karlsruhe, Germany; to the clear-cut genital characters of two sexes.

with the material of the from the males the and the details of the along new genus Irak, In shape minor and that from of the German armature of the valva But the Afghanistan, Afghanis- may vary slightly. spe- tan Expedition 1956, collected by Dr. AMSEL himself cific rigidity of the genital characters in the females and entrusted for identification. These characters to me Besides, inter- is simply astonishing! are extremely esting material was received from Mr. TOSHIIO OKU, useful for the descrimination of the species. There

Sapporo, Japan, and from some other sources. The seems to be no end to the specific variety within author the kind of the the The division of four gratefully acknowledges help genus. Bactra in subgenera senders, and their permission to retain certain dupli- (1956) proved to be natural and useful, at least with cates for the collection of the Leiden Museum. the three of the four, the New Zealand subgenus

Noteraula remaining monobasic and therefore little

Since the publication of my above mentioned paper known. a few errors were detected and some alterations be- The author intends a world-wide revision of the came necessary. They will be found below, together genus Bactra. Any material for re-examination or with descriptions of five new species and one sub- identification will be received with gratitude.

Bactra (Bactra) lanceolana (Hübner 1796)

Tortrix lanceolana Hübner Samml. 970 25.IV.1956 G. 1796, europ. Schmett., Afghanistan, Herat, m, (H. vol. 7, Tortr., t. 13, fig. 80. AMSEL), 7 <3, gen. nos. 2515, 2729, 2730, 2732, 2741, Bactra (Aphelia) fumosana Kennel 1900, Iris, vol. 13, 2742, 2744; 8(3, nos. 2794-2801; 9(3, nos. 2806- p. 263. Syn. nov. Palaearkt. 2814; 17(3, nos. 2822-2838. 41 <3. Bactra fumosana, Kennel 1910, Tortr., p. 472-

473, t. 18, fig. 73a. — Rebel, in Staudinger & Rebel 1901,

Catal. Lep. Pal. Faun., vol. 2, p. 113. Furthermore, identification of the following material Bactra (Bactra) fumosana, Diakonoff 1956, Zool. Ver- was confirmed. Sardinia, Isili, 20.VII.1936 (H. G. hand., no. 29, p. 16, fig. 16 (gen. $ type fig.). AMSEL), 1 <3, gen. no. 2720; Porto Santoru, 18.VI.1936

G. 2 nos. G.U. 90 and 95; 1 (H. AMSEL), (3 , gen. Ç, gen. Distribution. Palaearctic region. no. G.U. 101 (Ams.); Anitro, 6.VI.1934, 1 (3, gen. no. G.U. G.U. 98 91; Tempio, 18.IX.1934, 1(3, gen. no. (PBEDOTA Iran, Shiraz, 15.VII. 1940, 1 Ç, gen. no. 2856; no leg., SCHAWERDA Collection). Syria, Damascus, 27.VI.1950 also 2853. date, probably Shiraz, 2(5, gen. nos. 2849, G. 2722. (H. AMSEL), 1(3, gen. no. Iraq, Haj Omran, Barfkhanen, 9—12.000 feet, IV. 1940, 1(3, gen. no. Rayat, 5000-6000 feet, 2-13.VI. 1956, e coll. WILTSHIRE,

2854 P. WILTSHIRE AMSEL 3$. 1 G.U. 3372 (E. leg., Coll.). $• 1(3, gen. no. (Ams.).

Bactra (Bactra) lanceolana (Hübner 1796) forma ￿ fumosana Kennel status nov. (Pl., fig. 3)

and available seemed be rather distinct In my 1956 revision I was dubious about the status men type to of “Bactra fumosana”. At the time the unique speci- from lanceolana and I preferred to keep it separate,

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additional material. that with pending further evidence of e.g., gen. no. 2775, illustrated on Pl., fig. 3.

than could have In this the colliculum is in This evidence turned up sooner I specimen dilatedabove, as

in hoped, in the form of three specimens from Iran lanceolana (cf. my Figs. 12—13, 1956) and its ventral

has small also the AMSEL Collection, all females again. They were rim a spout, as in that species; the collected side by side with typical lanceolana, males characteristic punctulate field above the ostium (la-

female. of the mella has also intermediate and a After critical re-investigation postvaginalis) an shape; for the folds of genitalia I was satisfied that there are no reasons deceiving ƒ fumosana, giving it a peculiar

as a also but weak, maintaining fumosana separate species. In my aspect, are present here, are etc. but female form sub- It worth while for the opinion it is a (not even a seems to retain present species) of Bactra (B.) lanceolana. KENNEL'S name for this form.

The slight differences in the shape and in minor Iran, Resht, Tabergourabe, 20.V.1955 (FH. of the somewhat 1 2775. Fars, Pireh Zan, particulars sterigma appear exagger- SCHAUFFELE), 9, gen. no. and ated in the 16, 1000 feet, 4.X. 1950, 1 no. 2851. Shiraz, type specimen (cf. my Fig. 1956) Ç, gen.

variation. At 1 represent the extreme of a moderate 15.VII. 1940, $, gen. no. 2855 (E. P. WILTSHIRE).

have intermediate 3 present I specimens at hand, as, ?.

Bactra (Bactra) honesta Meyrick 1909 (Pl., fig. 1)

Bactra honesta Meyrick 1909, J. Bombay Nat. Hist. Soc., valvae turned flat as much as possible. From this vol. 19, 585 ( Ç, Assam). — Diakonoff 1950, Bull. Brit. p. (5 figure it is evident that the dorsal portion of the Mus., Ent., vol. 1, p. 287, pi. 6, fig. 23, pi. 7, fig. 33 (lecto- genitalia (uncus and socii) is exactly similar to that type select., gen. <3Ç fig.). lanceolana but the much less robust in tegumen is

and similar that the Distribution. , Bengal. more to in furfurana. Especially

The is close to lanceolana but decided- valva is it is shorter, broad at base and dis- species very different,

In sacculus is ly distinct. the British Museum (Natural History) tinctly pointed at the apex, while the

allowed rather and sacculus where the type is preserved, I was to re- acutely prominent triangular (left mount the genitalia and to re-examine them closely; is broken in two).

with the they are figured once more on Pl., fig. 1,

Bactra (Bactra) furfurana (Haworth 1811)

Haworth 1957 1 1 Tortrix furfurana 1811, Lep. Brit.,p. 466. (G. EBERT), <3, gen. no. 2717; Ç, gen. no.

Distribution. Palaearctic 2746. East 1100 28.VI. region. Afghanistan, Sarobi, m,

1956 (H. G. AMSEL), 1 <5, gen. no. 2727; 1 Ç, no. Iran, Fars, Sineh-Safid, ca. 6500 ft., 2.IX.1950 (E. 2739. P. WILTSHIRE), 1 S no. 2852. . gen. Spanish Morocco, Buhasen, 1000 m marsh, South 5.V. West Afghanistan, Herat, 910 m, 9.VI.1954 (E. DE BROS), 1 <3, gen. no. 2723. G. 1 <5, 2731; Hilmend 1956 (H. AMSEL), gen. no. All specimens in the AMSEL Collection. 500 18.V. River, Darweshan, Registan Desert, m,

Bactra (Bactra) longinqua spec. nov. (Fig. 1)

17 With and olive and dark fuscous. Basal fourth of with Ç mm. a general appearance markings costa a of but with and of dark marks and dot a lanceolana male, narrower wings series minute marginal a larger distinct genitalia. on extreme base; posterior part of costa with a series Head, palpus, and thorax ochreous-whitish, termin- of dark fuscous longitudinal larger blotches, alter- al of dark rather with few dark scales scattered segment palpus grey, slender, long, nating minute points; a ochreous-whitish. drooping. Abdomen Fore wing over olive markings and along veins in disc; olive

sublanceolate, costa in the usual a discal narrow, gently curved through- markings arranged pattern;

connected with dorsum faint out, apex tolerably pointed, termen oblique, gently patch at 2/5, by a concave. Ochreous-whitish, markings pale ochreous- marbling; the same marbling indicating a basal patch;

Downloaded from Brill.com09/23/2021 08:15:01PM via free access EXPLANATION OF PLATE

Fig. 1. Male genitalia of Bactra (Bactra) honesta Meyr. (holotype). Fig. 2. Male genitalia of

B. straminea 9545 (Chiloides) Butl. forma ￿ iomolybda Meyr., holotype (gen. no. B.M., phot. no. lanceolana forma ￿ 11509). Fig. 3. Female genitalia of B. (B.) fumosana Kennel, Iran, no. 2775.

Fig. 4. Male genitalia of B. (C.) truculenta Meyr., Pakistan, no. 2129. (Fig. 2, by the courtesy

of the Trustees of the British Museum. Figs. 1, 3 and 4, author phot.)

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Figs. 1-4. Female genitalia of Bactra (Bactra). 1. longinqua spec. nov. (Afgha-

nistan). 2. longinqua iranica subspec. nov. (Iran). 3. robustana Chr. (Sardinia).

4. amseli spec. nov. (Afghanistan)

from middle of end of several lines with less an oblique streak costa to cell, costa; a more or interrupted

there with an crescentic two terminal fascia, from below to tornus. ending elegant mark; apex Cilia erect blotches on dorsum, before and beyond pre- ochreous-whitish, with a fine ochreous-olive subbasal

and cilia mixed with dark fuscous. ceding, outwardly inwardly oblique, respectively; line, opposite apex

blotch in connected Hind and irrorated a longitudinal apex faintly by wing evenly finely with fuse-

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with distinct Cilia ochreous- not aciculate but ous, a purple gloss. nal, slightly sclerotized folds, and

ochreous towards above these each side whitish, becoming deeper apex; a on a small dark crescentic

fuscous subbasal line. structure. Ostium bursae resembling that in furfurana

Female sclerotized. genitalia (Text-fig. 1). Anapophyses strong- but distinctly A small signum present.

ly bent and sclerotized below base. Sterigma, or (Slide no. 2726, holotype).

the entire lamella as a East 1100 28.VI.1956 rather, postvaginalis shaped Afghanistan, Sarobi, m,

sclerotized in middle of trapezoid, slightly plate, (H. G. AMSEL), 1 Ç, holotype, gen. no. 2726. Unique.

less sclerotized related to Col- which, a T-shaped portion, edged by Apparently furfurana. Type in AMSEL

slight folds; above this there is a series of lortgitudi- lection.

Bactra (Bactra) longinqua iranica subsp. nov. (Fig. 2)

and Ç 11—17.5 mm (holotype 17.5 mm). Head thorax Paratype (imperfect) much smaller, with ground

whitish. Antenna dark median colour duller but grey. Palpus whitish, brighter white, markings deeper, faint- and rather well- segment strongly expanded by roughish scales, more olive-tawny tinged, forming

suffused with middle terminal defined of these ly grey in externally, transverse bands, posterior angulate

dark in segment slender, rather long, exposed, grey. middle; posterior third of wing entirely suffused Abdomen ochreous-whitish. with deeper olive-tawny, with anterior edge well-

rather streaks of Fore wing elongate-sublanceolate, narrow, defined, triangularly incised; 3—4 oblique

pointed, termen faintly sinuate, very oblique. Ochre- glossy whitish ground colour, originating from pos-

marbled with vivid ochre- terior of ous-white, irregularly light part costa, running across upper part of wing

colour form small basal breadth. Otherwise similar ous, this tending to a patch, to holotype.

a submedian, and a median transverse and some Female 2). simi- band, genitalia (Text-fig. Sterigma very lar indefinite marbling posteriorly; rather distinct costal to that of the nominate form but somewhat more

blotches variable size formed this sclerotized. ventral rim ostium of by marbling; The of the cup of

extreme edge of costa with minute dark marks formed bursae (lamella antevaginalis proper) stronger sclero-

few dark fuscous ill-defined tized and encircled a series of fine by a fine scales; cloudy ventrally by

dark faint suffused the lateral folds inside the dotting along dorsum; a longitud- hyaline folds; hyaline

inal streak of extend- the lamella brighter tawny-ochreous colour, ostium, as in nominate form, absent, post-

of ing from base to apex and occupying about 1/4 vaginalis similar, being a corrugated transverse-oval basal wing breadth; this streak interrupted between sclerite, but with stronger outward edges; small folds

patch and the two transverse fasciae, and becoming on each side and above this lamella, more distinct.

darker, rather well-defined, tinged orange, and con- The trapezoidal sclerite uniting the two lamellae, less fascia tinuous; this portion of longitudinal including distinct in the subspecies, rounded, with only the end cell and indicated short folds. a whitish ocellus on of strewn through- extreme angles by (Slides nos.

but with dark brown scales. out, more so posteriorly, 2779, holotype, 2777, paratype).

Cilia whitish-ochreous, with a white basal line, fol- South East Iran, Djiroft, Andar-Abad, 1—18.V.

lowed a faint darker ochreous subbasal line, cilia 2779; by 1956 (W. RICHTER), 1 Ç, holotype, gen. no.

mixed with dark fus- Resht, 1954 (FR. SCHAUFFE- opposite apex tawny-ochreous, Tabergourabe, VII—VIII.

scales. 1 2777. 2 in cous LE), Ç, paratype, gen. no. Ç. Holotype

Hind wing glossy light fuscous-grey, darker to- AMSEL Collection.

Cilia wards apex. whitish-grey.

Bactra (Bactra) robustana (Christoph 1872) (Fig. 3)

Horae Soc. Ent. Checklist Brit. 124. — Vâri Entom. Aphelia robustana Christoph 1872, Ross., Ins., p. 1951, Ber., vol.

198. — vol. 9, p. 13, t. 1, f. 10. 13, p. Bentinck 1951, Tijdschr. v. Ent., vol. 94,

Bactra robustana, Rebel, in Staudinger & Rebel 1901, p. 334. — Toll 1958, Annales Zool. Polon., vol. 17, p. 65—

— 11. Catal. Lep. Pal. Faun., vol. 2, p. 113. Kennel 1910, Pa- 72, figs. 5-6, t. 2, f. 4, t. 3, f.

t. 72. — Bactra Pierce laearkt. Tortr., p. 472, 18, f. Kloet & Hincks 1945, scirpicolana 1935, Entomologist, vol. 68, p.

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vol. South 970 25.IV. 148—149. Bentinck 1936, Tijdschr. v. Ent., 79, p. West Afghanistan, Herat, m,

XXVII. - L. 201. - Toll 1958, Annales Zool. Polon., 1 c., p. 1956 (H. G. AMSEL), Ç, gen. no. 2716. Iran, 65. vol. 17, p. Resht, Tabergourabe, 20.V.1955 (FR. SCHAUFFELE), Grapholitha (Bactra) scirpicolana, Bentinck 1936, Tijd- 1 2772. schr. 209. Ç, gen. no. v. Ent., vol. 79, p.

Bactra griseana Djakonov 19£9, Rev. russe Entom., vol.

23, 164, 21-22. confirmed. p. figs. Identification of the following material was Ver- Bactra grisea Diakonoff (nec Djakonov) 1956, Zool. Central Hungary, Dömsöd, 922.V.1923 (sic) (SCHMIDT),

hand. no. 29, 13 (lapsus). 910.VII.1918 p. 1(5, gen. no. 2802, 1,5, no. 2804; Izsâk,

(SCHMIDT), 1(5, no. 2803. Germany, Borkum Island, e 1. Coll. 22.VI.1931, Scirp. marit., 1(5, Gen. no. G.U. 89. Distribution. Palaearctic region. BRINKMANN. Sardinia, Teulada, 10.11.1934, 1 $, gen. no. G.U.

103 (Ams) (H. G. AMSEL, in his collection).

Bactra amseli (Bactra) spec. nov. (Fig. 4)

Head and thorax Cilia sordid with fuscous Ç 15—18 mm (holotype 18 mm). posteriorly. whitish, a pale

pale ochreous; palpus strongly dilated by roughish subbasal line.

scales above and terminal Female beneath, concealing seg- genitalia (Text-fig. 4). Eighth segment

in rather sclerotized. ment (paratype); holotype palpus worn, strongly Sterigma well-developed, shaped

with terminal Ab- a robust rounded os- slender, segment short, exposed. as subtriangular plate, below; with and thick- domen whitish-ochreous. tium, a triangular opening, a plicate

ened lower Lamella Fore wing narrow, elongate-lanceolate, moderately edge. postvaginalis, a sinuate

dilated, broadest at 3/4; costa little curved anteriorly, ridge, adorned with minute transverse strigulations.

small straight posteriorly, gently impressed along its third A signum present. (Slides nos. 2734, holotype; fourth, rather 2778, apex pointed, termen oblique, gently paratype).

convex. Whitish-ochreous, finely dotted with pale The paratype specimen has identical genitalia, but

different the fore tawny-ochreous, more than dorsal half of wing from markings: wing is whitish, finely

base suffused with this colour; streaked with a series of 4—5 to apex evenly upper pale greyish-olive; pale edge of this suffusion rather well-defined, posteriorly fuscous oblique strigulae originating from costal dots

fourth of costal and dorsal becoming concave, the suffusion running to apex and along posterior costa;

lower suffusion with of dark occupying its half; a deeper tawny edges throughout a continuous series

in series lower half of apex and along termen, becoming fuscous points; a of points before termen,

clouded with faint fulvous suffusion. narrower and ill-defined downward; a couple of min- distinct,

ute dark brown points scattered before termen (ab- South West Afghanistan, Hilmend River, 500

in Darweshan, sent left wing). Cilia whitish-ochreous, with a pale m, Registan Desert, 18.V. 1957 (G. basal suffused 1 2734. line, beyond this with pale ochreous- EBERT), $, holotype, gen. no. Iran, Resht,

small brownish dot lower 20.V. 1955 1 tawny, a opposite apex, on Tabergourabe, (FR. SCHAUFFELE), 5.

in Col- half of cilia. paratype, gen. no. 2778. Holotype AMSEL

Hind wing glossy grey-fuscous, becoming paler lection, paratype in the Leiden Museum. towards with of Related robustana. base, a suggestion a purplish gloss to

Bactra (Bactra) festa spec. nov. (Figs. 9, 13, 14)

15—16 15 Head and irrorated and dotted between (5 mm, Ç mm. palpus pale coarsely markings, es-

basal dark fuscous. ochreous; palpus moderate, median segment rough- pecially on half, with Markings ish towards below, terminal moderate, blackish-fuscous, with Some apex segment suffusedly edged tawny.

Antenna Thorax marks and of slender, exposed. pale grey. light eight transverse on costa, oblique vari- Abdomen with blackish ochreous-fuscous. glossy pale silvery-grey. ous sizes, alternating minute transverse

Fore wing narrowly elongate, dilated, broadest at jots; 011 basal half of costa these marks short and uni- bent colorous half the marks 4/5, costa moderately bent at base, stronger at blackish; on posterior larger

termen suffused with 4/5, apex rounded-pointed, gently concave, longitudinal, more tawny; penultimate undulate oblique. Pale ochreous, suffused with ashy-grey, mark giving rise to a tawny, slightly line,

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Figs. 5-9. Female genitalia of Bactra. 5. B. (Chiloides) telaviviana Amsel,

holotype (Israel). 6. B. (B.) graminivora Meyrick, no. 2585 (E. Turkestan).

7. The same, ostium bursae stronger magnified, no. 2718. 8. B. (Nannobactra)

leonina spec, nov., holotype (Pakistan). 9. B. (B.) festa spec, nov., allotype (Japan)

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of before the smaller. obliquely crossing to upper part termen, strong uncus longer, socii Especially

is termen abruptly bent downward; costal mark pre- characteristic the large sacculus of the valva, being

this but shorter rather than robustana but smaller ceding prolonged into a similar line, erect-oval, larger in

not reaching beyond vein 9; ultimate costal mark than in loeligeri; the usual sheaf of spines Spci is dark situated less and forming a small longitudinal blotch; a fuscous more or superficially not in a

rounded in rounded in most of this spot apex; a large slightly oblique V- cavity, as species subgenus;

shaped mark in disc, from middle of costa to lower cucullus rather slender and slightly more sinuate

towards not than in the two above mentioned As usual, angle of cell, thence rising apex but species.

of rather the of the valva much affected its reaching it; a series semicrescentic, irregular shape is very by

marks fold: a in the cf. 13 lateral along group together, beyond base, position mount, Fig. (completely

of basal inwards- of the and 14 not indicating edge patch, a larger aspect valva) Fig. (valva in a quite

blotch before and smaller lateral oblique 1/3, a crescentic aspect).

mark and before the of the Female and below slightly top V- genitalia (Text-fig. 9) peculiar highly

and interconnected marks lamella cordi- shaped mark, two slender, characteristic, antevaginalis being a

between crescentic mark brown with a forming a narrow zigzag form-V-shaped, vividly sclerite, corru-

and small additional mark surface and and towards termen; a longitudinal gated edges, two concavities

4 on termen, between veins and 5, connected with middle; the ostium bursae proper invert-triangular,

small with fine lateral folds and dark of character- posterior arm of the V-shaped mark; fuscous edges a

blotches extended Cilia illustrated small irregularly along termen. istic shape, as (Fig. 9). Signum, a

with brownish and substellate sclerite of usual and pale ochreous, a subbasal, a grey shape structure.

fuscous suffused blotch subapical, narrow lines; a (Slide no. 2841, allotype).

Hokkaido, Nukabira, 5.VII.1958, 1(5, holo- opposite apex. Japan, allo- Hind wing rather dark grey, with a faint brown type, gen. no. 2839; Jozankei, 19.VII.1956, 1 5, Cilia the and gloss. pale ochreous-greyish, with a faint sub- type, gen. no. 2841; same locality date, 2 S

basal fuscous line. paratypes, gen. nos. 2846 and 2847; Teine, 9.VIII. The female allotype (rather worn) is paler, more 1956, 1 (5, paratype, gen. no. 2845 (TOSHIIO OKU). ochreous tinged, with dark markings ochreous-tawny 4 (5, 1 $. Holo- and allotype in the Leiden Museum,

instead of dark additional in the collection of Mr. T. OKU. fuscous; an transverse paratypes

strigula originating from the ultimate costal mark and An interesting novel member of the subgenus Bac-

around to of nearest to robustana running apex upper part termen. Other- tra, perhaps Christoph or loeligeri

wise apparently similar to the holotype. Diakonoff, from Amur, but with the appearance of a Male genitalia (Text-figs. 13, 14) resemble those of gaudy coloured lanceolana. the Palaearctic robustana but are quite distinct, with

Bactra (Bactra) graminivora Meyrick 1922 (Figs. 6, 7)

vol. 2, no. 1899 recorded Bactra graminivora Meyrick 1922, Exot. Microl., p. (originally by me as furfurana,

521 (<3Ç, Bengal). — Fletcher 1932, Life-histories Ind. 1956, p. 12). East Turkestan, Charchui, 10—15

Microl., ser. 2, 24, pi. 14, figs, a-d (biol., food-plants). p. and 23.VI (no year) (J. VON RENNENKAMPF), 2(5, gen.

— Bull. Brit. vol. 287, Diakonoff 1950, Mus., Ent., 1, p. nos. 2579 and 2580, and 7 Ç, nos. 2580—2585. pi. 6, fig. 24, pi. 7, figs. 34, 36 (lectot. select., gen.$ Ç il- gen. lustr.). — Amsel 1958, Beitr. nat. Forsch. S.W. Deutschi., (Leiden Museum). vol. 17, 76 (N. Arabia). S. W. p. Afghanistan, Hilmend River, 500 m, Dar- Bactra (Bactra) graminivora, Diakonoff 1956, Zool. Ver- weshan, Registan Desert, 18.V. 1957 (G. EBERT), 3 $, 10. hand., no. 29, p. 18, fig. gen, nos. 2718, 2719, Herat, 970 m, 25.IV. Bactra Amsel Bull. Soc. Sei. 2733; cyperana 1951, nat. Maroc, 1956 nov. (H. G. AMSEL), 2 <3, nos. 2743, 2745. N. W. vol. 31, p. 68, fig. 4. ((5, Morocco). Syn.

Bactra Faunula Al- mediterraneana Agenjo 1952, Lep. Afghanistan, Korees Duobi, 600 m, near Bala pi. figs. pi. figs. 7 ( $ meriense, p. 99, 4, 32, 33, 12, 6, Ç, 19. V. 1956 G. 1 Murghab, (H. AMSEL), Ç, gen. no. Spain, Almeria). Syn. nov. 2721. N. 700 Afghanistan, Polichomri, m, 28. V.

Distribution: ? 1956 G. 1 1 India, Bengal; Kashmir, Shrinagar; (H. AMSEL), $, 9, gen. nos. 2862 and

Northern 2863. Pakistan, Karachi; Arabia; Spain; Morocco. E. Afghanistan, Sarobi, 1100 m, 3.VII. 1956

Central 1 G. 5 Asia, Margelan, Hbh, 1880, Ç, gen. (H. AMSEL), 1(5, gen. no. 2713; 9, 28. VI. 1956,

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2740. Col- described as follows. Ostium bursae gen. nos. 2714, 2724, 2725, (AMSEL shaped as a fine,

lection). hardly traceable ring, only ventral portion of which 4.IX.1940 P. 1 visible this Iran, Shiraz, (E. WILTSHIRE), 9» usually is (Fig. 7); portion is distinctly

2850. colliculum rather narrowed gen. no. (AMSEL Collection). sclerotized; is abruptly below Irak, Bagdad, 10.V.1937 (E. P. WILTSHIRE), 1 $, the sclerotized portion, hyaline, but apparent-

under this gen. no. 2865. ly rigid; rigid portion the membraneous

The nice, well-preserved series from Afghanistan, ductus bursae proper begins. Lamella post-vaginalis

well hitherto as as from Turkestan, reveal that this represented by a series of slightly sclerotized, but not little known variable. The males folds species is strongly hairy or aciculate, straight (in furfurana, a single usually are fuscous, and resemble the European fur- curved lip-like fold). (Slides figured, 2585 and 2718).

with furana most. The females are pale ochreous Although I was not able to study the type material light greyish hind wings and variably marked fore of Bactra mediterraneana Agenjo, the nice figure of

of these the dark trans- the male of this distinct wings; markings mostly only genitalia species is so (I.e., and but verse strigulae on costa dorsum are visible, PL 12, fig. 6), that I do not hesitate to identify

the brownish shadow from mediterraneana with the sometimes longitudinal present species. base also above the middle of Dr. H. G. AMSEL male to apex is present just kindly sent me a paratype

Turkestan dark fuscous B. Palestine, wing; two specimens possess specimen of his cyperana from express- discal smaller fold of his that this also be the spots: a one in beyond 1/4 ing surmise species might

and end had indeed confirm this. wing a larger, irregularly elongate one, on same as graminivora. I to of cell.

The male identical to those of the to genitalia are Furthermore, I was able confirm the following identi-

fications AMSEL Indian examples. The large oval field of the sacculus of the present species in the Collection.

Iraq, Bagdad, 18.V.1956, ex coll. Wiltshire, 1(5, size and female gen. may slightly vary as to shape. The no. 3371 Ams.; North East Arabia, Dhahran, 11.III. and of which genitalia (Figs. 6 7) a photograph was 1957, e coll. Talhouk, 1(5, gen. no. 3340 Ams.; Ras Ta-

earlier PI. 7, are illustrated 3346 published (1950, fig. 34) nura, 18.IV.1957, the same, 1(5, gen. no. Ams. here at a larger scale (Figs. 6 and 7) and may be

Bactra (Bactra) telaviviana Amstel 1935 (Fig. 5)

Bactra telaviviana Amsel 1935, Zoogeographica, vol. 2, p. opportunity to describe and illustrate, with Dr. AM-

46, 123 (nom. nud.) — 1935, Mitt. Zool. Mus. Berlin, vol. SEL'S consent, the female genitalia for the first time.

20, 291, pi. 11, fig. 89 (Tel Aviv, Palestine). — Veröff. p. Female genitalia (Text-fig. 5). Eighth segment Deutsch. Kol. Uebers. Mus., vol. 1, p. 261, no. 230. formed sclerotized, apophyses strong. Sterigma by a

U-shaped lamella antevaginalis; lamella postvaginalis

Distribution. Palestine. indicated only by small semicircular folds. Colliculum

sent for closer the Dr. AMSEL kindly me study narrow, long, funnel-shaped, hyaline. Signum ap- female holotype of this interesting species. I use this parently absent. (Slide no. 2770).

Bactra (Chiloides) simpliciana Chrétien 1915 (Figs. 12, 18)

Chrétien Head with Bactra simpliciana 1915, Ann. Soc. ent. France, 5 18 mm. smoothly appressed scales-, vol. 301 (adult), 302 (larva and pupa; on Cyperus 84, p. pale ochreous. Thorax pale ochreous. Palpus moder- conglomeratus Rottb.). — Amsel 1951 Bull. Soc. Sei. Nat. ate, triangularly dilated, rather smooth, except a tuft Maroc, vol. 31, p. 68. on at the ochreous, median Bactra telaviviana, Amsel 1951 (nec Amsel 1935!), Beitr. upper edge top; pale

suffused with on basal nat. Forsch. S.W. Deutschl., vol. 17, p. 76, fig. 18 ($, segment grey externally except N.E. Arabia; attrib. subgen. to Chiloides). short, third, terminal segment very exposed. Ab- domen whitish-ochreous.

Owing to the great kindness of Mr. J. BOURGOGNE, Fore wing elongate-semioval, almost sublanceolate, almost Muséum National d'Histoire Naturelle, Paris, I was dilated, broadest beyond 2/3; costa straight, able the the to study unique type specimen of present apex distinctly rounded, termen rounded, oblique.

be redescribed as follows. Unicolorous ochreous, with a species. It may very light slight gloss

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Figs. 10-13. Genitalia of Bactra. 10. B. (Chiloides) diakonoffi Amsel, female,

allotype. 11. B. (C.) grethae spec, nov., female, holotype (Pakistan). 12. B. Chrét. (C.) simpliciana (no. 3334, N. Arabia). 13. B. (B.) festa spec, nov.,

paratype (no. 2845), left valva in lateral aspect

and with faintest terminal and with suffused with to- a pale tawny tinge; very pale ochreous, greyish apical edges narrowly suffused with tawny-brownish. wards apex, with a golden gloss. Cilia white with a

Cilia faint submedian faintest and half ochreous-whitish, a tawny yellowish tinge opposite apex upper shade, base pale. of termen.

Hind wing whitish, apical fourth faintly suffused Male genitalia (Figs. 12, 18). Socius rather small.

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Figs. 14-17. Male genitalia of Bactra. 14. B. (B.) festa spec, nov., holotype.

15-16. Left valva of the types of B. (C.) diakonoffi, Amsel, monochorda Diak., and excelsa Diak., respectively (16 and 17 after DIAKONOFF 1956)

with those side of the sacculus shown the left uncus resembling parts in B. furfurana. (as in valva; in

Valva submembraneous, rather weak, cucullus cloth- the right the spines are fully drawn for the sake of ed, except along costa, with fine hairs, not with clarity). bristles. Valvula short, the apical bristles ("corona") Holotype (monotype), male, gen. no. 2876, with

characteristic black moderately long; two bristles on the labels: "Cyp. conglom., 26.4.09" in ink; red inner side of the stalk. Sacculus large but rather label "TYPE"; "Bactra simpliciana" in violet ink. In

with a covered the condition but left half of thorax and the amorphous, weak, big area with good with characteristic round and with fine short basal half of the left fore with brown impressions wing a stain. hairs. The stout spines are attached on the external The two male specimens, recorded by Dr. AMSEL

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(1958) from Northern Arabia, one of which I was solution of this problem for a long time, though.

kindly allowed to study, and attributed by him to Material studied: 1 <3, North Arabia, Hofuf, 15.11.

telaviviana 12. B. (described after a single female) ap- 1957, gen. no. 3334 Ams., figured on Text-fig.

to be identical with the above re-described This with unicolorous with pear type large species long wings

of Whether and tel- rounded of for simpliciana. simpliciana (<3) a apex, is especial interest, it seems to

of and the form of the aviviana (9) are only sex partners one same represent an early subgenus Chiloides,

species, it is not possible at present to decide. The with small sacculus and valvula as compared with the

two names should be kept separate, pending further large cucullus. evidence of bred material. of this material The of of described Obtaining types two species Bactra, by

of simpliciana seems to be not very difficult, since COUNT E. TURATI from Cyrenaica, viz., lactosana and

CHRÉTIEN indicated the food and the of available plant timp pauperrima, were not to me. Apparently, of the larval The these lost. occurrence stages. present political types are It is probable that they represent

unrest the of North Africa the the and of in countries may delay same species, are synonyms simpliciana.

Bactra (Chiloides) straminea(Butler 1881) (Pl., fig. 2)

Chiloides Butler of additional moderate straminea 1881, Ann. Mag. Nat. Hist., transverse series spines across

ser. 5, vol. 7, 393, no. 23 (). — Meyrick 1885, p. the inner side of the disc of the saucculus, running Trans. N. Zeal. Inst., vol. 17, p. 142 (part.). — Walsingham beside and parallel to the base of the median coronate Fauna 5. - 1907, Haw., vol. 1, p. 687-688, pi. 11, fig. of the valva This series of Zool. no. appendage (valvula). spines Diakonoff 1956, Verhand., 29, p. 26, figs. 4, 24-25 (gen. 3$). is indicated in the photograph by an arrow (Pl., fig. Bactra (Chiloides) straminea, Diakonoff 1956, Zool. Ver- the of the of the British 2). By courtesy Trustees hand., no. 29, p. 26, figs. 4, 24-25 (gen.

A in comparison of the genitalia of the type specimen however, is very slight and occurs several male and close of further material the British study at specimens from Hawaii studied by me. Tentatively

Museum revealed that Bactra is this iomolybda Meyrick I propose to retain name as Bactra (Chiloides)

straminea straminea only a synonym of Butler. The only slight Butler forma $ iomolybda Meyrick (status difference between the which traceable types is is a nov.).

18. Bactra Museum Fig. Male genitalia of simpliciana Chrét., holotype, gen. no. 2876, Paris

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Bactra (Chiloides) venosana(Zeller 1847)

the the median Phoxopteris venosanaZeller 1847, Isis, p. 738. spines on external portion of the Aphelia venosana, Herrich-Schäffer 1849, Syst. Bearb., sacculus: these much are longer and more robust, but 244. vol. 4, p. less truculenta. The numerous in females, however, Bactra venosana, Rebel in Staudinger & Rebel 1901, vol. are different (cf. Figs. 28—32, 1956). — my 2, p. 113. Kennel in Spuler 1910, Schmett. Eur., vol. 2, clearly

273. - Kennel Pal 472, 18, 73. p. 1910, Tortr., p. pi. fig.

Bactra (Chiloides) venosana, Diakonoff 1956, Zool. Ver- Identifications confirmed. Palestine, Jericho, Licht-

31—33 hand., no. 29, p. 33, figs. (gen. <3Ç fig.). fang, 31.V.1930, 1(5, gen. no. 2816; Tabgha, Tiberias,

10.V. 1 2815 G. 1930, (5, gen. no. (H. AMSEL). Jordan Val- to The more material of this below level, species comes my ley, Zerqa R. Colony, c. 100 m sea at light,

25.VI. 2 the closer to be its to Bactra 1954 and 22.IX.1954 (TREVOR THOUGHT), (5, gen. notice, appears affinity 2819 and 2821. Porto 12.VI.1936 truculenta nos. Sardinia, Santoru, (Chiloides) Meyrick, a widely distributed, G. 2817 2805. Tem- (H. AMSEL), 1(3, 1 Ç, gen. nos. and tropical from Southern Asia. In the male the species Amsel G. pis, 6.V.1932, 1 (5, gen. no. G.U. 92, (H. AMSEL). reliable difference is the and the number of only size (AMSEL Collection).

Bactra (Chiloides) truculenta Meyrick 1909 (Pl., fig. 4)

Bactra truculentaMeyrick 1909, J. Bombay Nat. Hist. Soc., Caroline Islands.

— Diakonoff Bull. Brit. vol. 19, p. 586 ((5, India). 1950, One male specimen, from Mauripur, clearly shows 30 Mus., Ent., vol. 1, p. 289, pi. 5, fig. 16, pi. 7, fig. of the which al- a peculiarity genitalia apparently is and (lectotype select., syn. scythropa geraropa). but not visible in Bactra scythropa Meyrick 1911, Prot'. Linn. Soc. N.S ways present always as clearly as Dilli). this therefore I of these Wales, vol. 36, p. 284 (Ç Timor, case; present a photograph Bactra Meyrick 1932, Exot. Microl., vol. 4, geraropa genitalia (Pl., fig. 4). This photograph shows a kind 147 (Ç Formosa). p. of sclerotized rounded the of the cap over top sac- Bactra ( Chiloides) truculenta, Diakonoff 1956, Zool. culus (between arrows). This structure is omitted Verhand., no. 29, p. 27, figs. 28-30 (gen. (JÇ re-descr., from is fig-)- my previous figures (1956, figs. 29, 30). It

equally present in venosana.

Distribution. India, Andaman Islands, Pakistan, Mauripur, 29.VII. 1956, at light, 2 c5 Ceylon, >

South , Formosa, Timor, Hawaii, ?, gen. nos. 2129 and 2131. Thailand, Bangkok, 21—

India, 27.XI.1957, 1 1 no. ?. Known to me previously from Java, Ç, gen. no. 2439, $, gen. 2440;

Phen 8. 2443 Borneo, Kisar Island, Philippine Islands, Hawaii, Keng Dao, XII. 1957, 1 Ç, gen. no. (L.

D. BRONGERSMA). 3(3, 2 $.

Bactra (Chiloides) diakonoffi Amsel 1958 (Figs. 10, 15)

Bactra (Chiloides) diakonoffi Amsel 1958, Beitr. nat. only, gently decreasing in size proximad. Further-

Forsch. S.W. Deutschl., vol. 17, 76, f. 19 N. p. (<5$, more the cucullus is slightly broader, and the valvula

Arabia, <3 gen. illustr., attrib. to subg. Chiloides). is without the bristles along the stalk. (That the

the kindness of AMSEL I had an of the valvula absent, as stated in the de- Owing to Dr. op- corona is

the and the portunity to examine holotype allotype scription, is a delusion). (Slide no. 3339, Ams.). be related specimens of this recently described species, and also The allotype proves to equally to ex- but also be distinct. have doubt that was kindly granted permission to figure the genitalia celsa, to I no

somewhat scale than the the indeed. The differences at larger in original figure. types are sex partners

The close the Asiatic from the female of excelsa as follows. species is very to tropical genitalia are instead of al- excelsa Diak. and monochorda Diak., as Text-figs. Eighth tergite (Text-fig. 10) erect-oval,

15—17 distinct from both. The lamella less show, but sufficiently most circular, as in excelsa; antevaginalis

of the males the and the with folded and with lower differences are shape arma- broad, upper angles over,

and without ture of the sacculus. In diakonoffi it is broader (distal) portion differently shaped, a split;

colliculum shorter. larger, armed with some six large marginal spines distinctly (Slide no. 2771, Diak.).

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Bactra (Chiloides) grethae spec. nov. (Fig. 11)

$ 16 mm. Head, palpus and thorax (rather rubbed) sternite with two elongate-oval, sublateral, heavily

pale ochreous, palpus moderate, rather slender, termi- sclerotized plates, narrowed and united mesially rather nal segment infuscated. Abdomen pale ochreous. Colliculum well-developed, straight and long irrorated with rather substellate sclerite of usual Fore wing (rubbed) pale ochreous, Signum, a large

fuscous, brown-fuscous suffusion in disc before and shape and structure. (Slide no. 2130, holotype).

beyond cell, connected with a more or less distinct, Pakistan, Mauripur, 29.III.1956, at light, 1 $,

of brownish-fuscous marks 2130 D. interrupted row cloudy on holotype, gen. no. (L. BRONGERSMA). Unique.

terminations of veins. (Cilia lacking). Dedicated to the wife of the collector, Dr. M. BRON-

Hind wing ochreous-whitish, faintly irrorated with GERSMA-SANDERS.

fuscous towards The rather but the pale or greyish, more distinctly so specimen is rubbed, genitalia

Cilia whitish. are so distinct that I do not hesitate describing this apex. with far male Female genitalia (Text-fig. 11) sterigma com- new species. I expect the, so unknown,

sclerotized in plicated and characteristic. Eighth sternite mem- to possess a long and aedoeagus, ac-

with broad middle which fits cordance with the colliculum of the female. At braneous, a split in in long

able indicate the of this a broad, funnel-shaped and sclerotized, enlarged present I am not to affinity

ostium bursae, plicate at the sides; below this the interesting species, pending the discovery of the male.

Bactra (Chiloides) leucogama Meyrick 1909

Bactra leucogama Meyrick 1909, J. Bombay Nat. Hist. Distribution. Ceylon, Celebes.

vol. 584 — Soc., 19, p. (Ceylon, (5$). Diakonoff 1950, 1 Thailand, Keng Phen Dao, 8.XII.1957, $, gen Bull. Brit. Mus., Ent., vol. 1, p. 287, pi. 6, fig. 22, pi. 7, no. 2442 (L. D. BRONGERSMA). 1 specimen. fig. 33.

Bactra (Chiloides) leucogama, Diakonoff 1956, Zool,

Verhand., no. 29, p. 45, figs. 39-41.

leonina Bactra (Nannobactra) spec. nov. (Fig. 8)

ochreous-whitish. tri- wards base, towards dark and Ç 14 mm. Head, thorax Palpus apex becoming grey

dilated, of median Cilia ochreous-whitish, with a subbasal angularly apex segment produced opaque. grey

terminal concealed; shade. into an acute point, segment whitish, basal and lower edge of apical half, suffused Female genitalia (Text-fig. 8) resembling those of

aciculate with black. Abdomen whitish-ochreous. minima Meyrick. Sterigma, a minutely cup,

costa ab- much than in that also Fore wing sublanceolate, rather narrow, larger species, upper angles lower ruptly curved at base, slightly curved throughout pos- slightly produced, lateral walls much thicker;

rim Ninth sclerotized teriorly, apex pointed, termen oblique, gently convex. or lip, similar. segment more

Pale ochreous-yellowish (rather rubbed), faintly suf- throughout, anapophyses straight, sclerotized and fused in disc, and in as minima, not sclerotized, along termen, apex upward longer (in curved). Signum far with a continuous series of of usual and a as vein 8, grey; very shape structure: moderate, cup-shaped short, blackish-brown strigulae alternating with points minutely aciculate sclerite, stronger sclerotized than

and of short blackish- in minima. along costa, a series marginal (Slide no. 2441).

1 brown strigulae along termen. Cilia dull sordid ochre- Thailand, Bangkok, 24—27.XI.1957, at light,

D. ous-whitish, base narrowly white; faintly barred with $, holotype, gen. no. 2441 (LEO BRONGERSMA).

dark fuscous, bar opposite apex distinct, grey. Unique. Nearest to Bactra (N.) minima Meyrick,

Hind wing glossy fuscous-grey, semipellucent to- differing by the genitalia.

Bactra (Nannobactra) phaulopa Meyrick 1911

Bactra phaulopa Meyrick 1911, Proc. Linn. Soc. N.S. Bull. Brit. Mus., Ent., vol. 1, p. 288, pi. 8, fig. 42 (gen.

— Diakonoff 1950, Wales, vol. 36, p. 253 (Ç, Kei Islands). descr., fig.).

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Bactra (Nannobactra) phaulopa, Diakonoff 1956, Zool. cited paper (1956), references in the text to the

no. 51 (gen. redescr., Verhand., 29, p. 52, figs. 50, <3Ç of the of this figures genitalia species on the opposite fig-)- 50 of to the page got interchanged: fig. refers, course,

Correction. Unfortunately, on p. 55 of the above female, fig. 51, to the male.

Bactra (Nannobactra) minima Meyrick 1909

1909, J. Nat. Hist. Soc., close Bactra minima Meyrick Bombay A comparison of the types of minima and vol. — Diakonoff 19, p. 586 (<5, Barberyn Island, Ceylon). microtripta in the British Museum (Natural History) 1950, Bull. Brit. Mus., Ent., vol. 1, p. 288, pi. 6, fig. 25 me that the proved to they are same species. (lectot. select., gen. <3 fig.). It should be noted that the of the valva of Bactra (Nannobactra) minima, Diakonoff 1956, Zool. edge 52—53 this armed with but with Verhand., no. 29, p. 55, figs. (syn. phaeopis, gen. species is not a single series

(5 $ redescr., fig.). series are several of stout spines which arranged Bactra phaeopis Meyrick 1911, Proc. Linn. Soc. N.S. another and make the of closely to one impression Sudest — Wales, vol. 36, p. 254 ( Ç, Island, N. Guinea). dense of 53 of being a single row spines, as in Fig. Diakonoff 1950, Bull. Brit. Mus., Ent., vol. 1, p. 288, pi. revision 6, fig. 28 (gen. (J), pi. 8, fig. 41 (gen. Ç). (Lectot. select., my above cited (1956).

— Zool. 55. gen. (5 $ descr., fig.). 1956, Verhand., no. 29, p. Bactra microtripta Meyrick 1927, Ins. Samoa, vol. 3,

fase. 75 ( Samoan nov. Lep., 2, p. <5, Islands). Syn.

“Bactra” 1900 roseana Walsingham

Hist., but to the tribe Olethreutini Bactra roseana Walsingham 1900, Ann. Mag. Nat. genus Bactra, belongs ser. 7, vol. 6, p. 334 ((J, Japan). and might be placed somewhere near the genus

This large, elegantly coloured and marked species Olethreutes. I studied the male type specimen at the from Japan has nothing whatever to do with the British Museum (Natural History).

POSTSCRIPT

before this sent to the 2—5, 16 1958). In this Shortly paper was printers', pl. text-figs., Warszawa, paper

Dr. S. me new Palaearctic Bactra B. is TOLL, Katowice, Poland, kindly sent a species, gozmanyana,

described. do venture make comments his just appeared interesting paper on some new I not to any

Tortricidae der Familie this before close of TOLL'S (Drei weitere neue Arten upon species, a study Dr.

Tortricidae vol. material which he send aus Polen, Ann. zool., 17, pp. 65—80, very kindly promised to me.

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