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Maquetación 1
About IUCN IUCN is a membership Union composed of both government and civil society organisations. It harnesses the experience, resources and reach of its 1,300 Member organisations and the input of some 15,000 experts. IUCN is the global authority on the status of the natural world and the measures needed to safeguard it. www.iucn.org https://twitter.com/IUCN/ IUCN – The Species Survival Commission The Species Survival Commission (SSC) is the largest of IUCN’s six volunteer commissions with a global membership of more than 10,000 experts. SSC advises IUCN and its members on the wide range of technical and scientific aspects of species conservation and is dedicated to securing a future for biodiversity. SSC has significant input into the international agreements dealing with biodiversity conservation. http://www.iucn.org/theme/species/about/species-survival-commission-ssc IUCN – Global Species Programme The IUCN Species Programme supports the activities of the IUCN Species Survival Commission and individual Specialist Groups, as well as implementing global species conservation initiatives. It is an integral part of the IUCN Secretariat and is managed from IUCN’s international headquarters in Gland, Switzerland. The Species Programme includes a number of technical units covering Species Trade and Use, the IUCN Red List Unit, Freshwater Biodiversity Unit (all located in Cambridge, UK), the Global Biodiversity Assessment Initiative (located in Washington DC, USA), and the Marine Biodiversity Unit (located in Norfolk, Virginia, USA). www.iucn.org/species IUCN – Centre for Mediterranean Cooperation The Centre was opened in October 2001 with the core support of the Spanish Ministry of Agriculture, Fisheries and Environment, the regional Government of Junta de Andalucía and the Spanish Agency for International Development Cooperation (AECID). -
Spanish Greenish Black-Tip)
EUROPEAN BUTTERFLIES: A PORTRAIT IN PHOTOGRAPHS by B R Watts Euchloe bazae (Spanish Greenish Black-tip) The following pages on Euchloe bazae are an extract from a draft of chapter 4.4 of the publication EUROPEAN BUTTERFLIES: A PORTRAIT IN PHOTOGRAPHS. This extract will be available as a free download at: www.butterflyeurope.co.uk If you have any queries about the publication etc. please contact me personally at [email protected] Because chapter 4.4 contains pages on other Pieridae species too, cross-references in the present extract cannot be followed. EUROPEAN BUTTERFLIES: A PORTRAIT... PIERINAE (THE WHITES) Euchloe bazae Dappled White Group Spanish Greenish Black-tip Taxonomy and Systematics North Africa and may be Two subspecies are recognised in found throughout the year de- Europe: pending on weather and local- ity. E. b. bazae; E. b. iberae. Variation, Identification and The recognition of two subspecies acknowledges geographical Similar Species isolation and different foodplants of the taxa, see below. Broadly, E. bazae is a small, bright yellow species with dark As described on p. 1, the current view is that E. bazae flies in markings like other Euchloe southern and northern Spain and a sibling species, E. charlo- species and both sexes are nia, flies in North Africa. Formerly, the taxa in northern Spain similar. and North Africa were thought to be the same species, E. char- lonia, and the taxon in southern Spain was regarded by some The ups ground-colour is sulphur-yellow and the only dark authors as a different species, E. bazae. markings are on the upfw: a prominent black disco-cellular mark and a black apical patch with a row of yellow (ground-colour) Distribution marks within it. -
Annotated Checklist of Albanian Butterflies (Lepidoptera, Papilionoidea and Hesperioidea)
A peer-reviewed open-access journal ZooKeysAnnotated 323: 75–89 (2013) checklist of Albanian butterflies( Lepidoptera, Papilionoidea and Hesperioidea) 75 doi: 10.3897/zookeys.323.5684 CHECKLIST www.zookeys.org Launched to accelerate biodiversity research Annotated checklist of Albanian butterflies (Lepidoptera, Papilionoidea and Hesperioidea) Rudi Verovnik1, Miloš Popović2 1 University of Ljubljana, Biotechnical Faculty, Department of Biology, Večna pot 111, 1000 Ljubljana, Slovenia 2 HabiProt, Bulevar oslobođenja 106/34, 11040 Belgrade, Serbia Corresponding author: Rudi Verovnik ([email protected]) Academic editor: Carlos Peña | Received 25 May 2013 | Accepted 6 August 2013 | Published 13 August 2013 Citation: Verovnik R, Popović M (2013) Annotated checklist of Albanian butterflies (Lepidoptera, Papilionoidea and Hesperioidea). ZooKeys 323: 75–89. doi: 10.3897/zookeys.323.5684 Abstract The Republic of Albania has a rich diversity of flora and fauna. However, due to its political isolation, it has never been studied in great depth, and consequently, the existing list of butterfly species is outdated and in need of radical amendment. In addition to our personal data, we have studied the available litera- ture, and can report a total of 196 butterfly species recorded from the country. For some of the species in the list we have given explanations for their inclusion and made other annotations. Doubtful records have been removed from the list, and changes in taxonomy have been updated and discussed separately. The purpose of our paper is to remove confusion and conflict regarding published records. However, the revised checklist should not be considered complete: it represents a starting point for further research. -
The Status and Distribution of Mediterranean Butterflies
About IUCN IUCN is a membership Union composed of both government and civil society organisations. It harnesses the experience, resources and reach of its 1,300 Member organisations and the input of some 15,000 experts. IUCN is the global authority on the status of the natural world and the measures needed to safeguard it. www.iucn.org https://twitter.com/IUCN/ IUCN – The Species Survival Commission The Species Survival Commission (SSC) is the largest of IUCN’s six volunteer commissions with a global membership of more than 10,000 experts. SSC advises IUCN and its members on the wide range of technical and scientific aspects of species conservation and is dedicated to securing a future for biodiversity. SSC has significant input into the international agreements dealing with biodiversity conservation. http://www.iucn.org/theme/species/about/species-survival-commission-ssc IUCN – Global Species Programme The IUCN Species Programme supports the activities of the IUCN Species Survival Commission and individual Specialist Groups, as well as implementing global species conservation initiatives. It is an integral part of the IUCN Secretariat and is managed from IUCN’s international headquarters in Gland, Switzerland. The Species Programme includes a number of technical units covering Species Trade and Use, the IUCN Red List Unit, Freshwater Biodiversity Unit (all located in Cambridge, UK), the Global Biodiversity Assessment Initiative (located in Washington DC, USA), and the Marine Biodiversity Unit (located in Norfolk, Virginia, USA). www.iucn.org/species IUCN – Centre for Mediterranean Cooperation The Centre was opened in October 2001 with the core support of the Spanish Ministry of Agriculture, Fisheries and Environment, the regional Government of Junta de Andalucía and the Spanish Agency for International Development Cooperation (AECID). -
Phylogeny of European Butterflies V1.0
bioRxiv preprint doi: https://doi.org/10.1101/844175; this version posted November 16, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY 4.0 International license. A complete time-calibrated multi-gene phylogeny of the European butterflies Martin Wiemers1,2*, Nicolas Chazot3,4,5, Christopher W. Wheat6, Oliver Schweiger2, Niklas Wahlberg3 1Senckenberg Deutsches Entomologisches Institut, Eberswalder Straße 90, 15374 Müncheberg, Germany 2UFZ – Helmholtz Centre for Environmental Research, Department of Community Ecology, Theodor- Lieser-Str. 4, 06120 Halle, Germany 3Department of Biology, Lund University, 22362 Lund, Sweden 4Department of Biological and Environmental Sciences, University of Gothenburg, Box 461, 405 30 Gothenburg, Sweden. 5Gothenburg Global Biodiversity Centre, Box 461, 405 30 Gothenburg, Sweden. 6Department of Zoology, Stockholm University, 10691 Stockholm, Sweden *corresponding author: e-mail: [email protected] Abstract With the aim of supporting ecological analyses in butterflies, the third most species-rich superfamily of Lepidoptera, this paper presents the first time-calibrated phylogeny of all 496 extant butterfly species in Europe, including 18 very localized endemics for which no public DNA sequences had been available previously. It is based on a concatenated alignment of the mitochondrial gene COI and up to 11 nuclear gene fragments, using Bayesian inference of phylogeny. To avoid analytical biases that could result from our region-focus sampling, our European tree was grafted upon a global genus- level backbone butterfly phylogeny for analyses. In addition to a consensus tree, we provide the posterior distribution of trees and the fully-concatenated alignment for future analyses. -
Butterflies of Albania – Fluturat E Shqipërisë. New Surveys, New Species and a New Checklist (Lepidoptera: Papilionoidea)
Butterflies of Albania – Fluturat e Shqipërisë. New surveys, new species and a new checklist (Lepidoptera: Papilionoidea) Sylvain Cuvelier, Laurian Parmentier, Anila Paparisto & Jurgen Couckuyt Abstract. Three new butterfly species were recorded for Albania during two recent surveys: Pyrgus andromedae (Wallengren, 1853), Colias caucasica (Staudinger, 1871) and Leptidea juvernica Williams, 1946. Also Boloria titania (Esper, 1793) is new for the current Albanian territory that had previously been described by Rebel from the historical, larger Albania but situated now in Kosovo. Two data deficient species from the last checklist are confirmed: Carterocephalus palaemon (Pallas, 1771) and Hipparchia semele (Linnaeus, 1758). The status of six species is changed from present to data deficient or potential presence because of lacking recent evidence: Pontia chloridice (Hübner, 1813), Satyrium pruni (Linnaeus, 1758), Neptis rivularis (Scopoli, 1763), Euphydryas maturna (Linnaeus, 1758), Coenonympha glycerion (Borkhausen, 1788) and Kirinia climene (Esper, 1783). The geographical distribution for Pseudochazara tisiphone Brown, 1980 is significantly expanded to central Albania. Dot distribution maps are provided for all Albanian butterfly species. In addition to the new data, the available literature is screened, resulting in a mandatory update of the checklist for the Albanian butterflies: 196 species with recent evidence, 4 are data deficient and at least 9 have the potential to be discovered with future surveys. Përmbledhje. 3 lloje të reja fluturash për Shqipërinë janë përcaktuar nga analiza e materialeve të koleksionuara gjatë dy hetimeve shkencore të realizuara së fundmi: Pyrgus andromedae (Wallengren, 1853), Colias caucasica (Staudinger, 1871) dhe Leptidea juvernica Williams, 1946. Gjithashtu edhe Boloria titania (Esper, 1793) është tashmë një lloj i ri për territorin e Shqipërisë pasi më parë ky lloj është raportuar nga Rebel në zonën që sot ndodhet në Republikën e Kosovës. -
Surveys of Butterfly and Skipper Fauna in the Southwestern Part of the Republic of Macedonia (Lepidoptera: Papilionoidea & Hesperioidea)
NATURA SLOVENIAE 18(2): 23-37 Prejeto / Received: 25.10.2016 ZNANSTVENI ČLANEK Sprejeto / Accepted: 17.11.2016 Surveys of butterfly and skipper fauna in the southwestern part of the Republic of Macedonia (Lepidoptera: Papilionoidea & Hesperioidea) Vid ŠVARA1, Miloš POPOVIĆ2,3, Andrej PETERNEL4, Đorđe RADEVSKI3, Kaja VUKOTIĆ5, Rudi VEROVNIK6 1 Jakčeva ulica 1, 2380 Slovenj Gradec, Slovenia; e-mail: [email protected] 2 University of Niš, Faculty of Sciences and Mathematics, Department of Biology and Ecology, Višegradska 33, 18000 Niš, Serbia; E-mail: [email protected] 3 HabiProt, Bulevar Oslobođenja 106/34, 11040 Beograd, Serbia 4 Javorje 31, 4223 Poljane, Slovenia; E-mail: [email protected] 5 Osek 14b, 5261 Šempas, Slovenia; E-mail: [email protected] 6 University of Ljubljana, Biotechnical Faculty, Department of Biology, Jamnikarjeva 101, 1000 Ljubljana, Slovenia; E-mail: [email protected] Abstract. Data from four surveys within the last six years were combined in order to improve the knowledge of the butterfly distribution in the Republic of Macedonia. These surveys covered a total of 40 localities, with special emphasis on less sampled areas of the south-western part of the country. The study yielded recording of 131 species, including several habitat specialist and potentially threatened butterflies. Interesting records for the following species are discussed in detail: Muschampia tessellum, Anthocharis gruneri, Euchloe penia, Tarucus balkanicus, Cupido alcetas, Pseudophilotes bavius, Polyommatus aroaniensis, Polyommatus escheri, Araschnia levana and Melitaea ornata. Ten species observed are considered threatened at the European level and 18 of them at the country level. This proves high conservation value of the studied region with many butterfly rich habitats still preserved. -
The Phylogenetic Relationships of the Species and Subspecies of The
Atalanta (Dezember 2006) 37 (3/4): 469-482, Würzburg, ISSN 0171-0079 The phylogenetic relationships of the species and subspecies of the subgenus Elphinstonia K l o t s , 1930 (Lepidoptera, Pieridae) by W erner Back , T homas K nebelsberger & M ichael A. M iller received 25.XI.2006 Abstract: This work presents the results of the first molecular investigations of the subgenus Elphinstonia K lots, 1930. A 326 bp fragment of the mitochondrial cytochrome oxidase subunit I (COI) gene was available for the analysis. Monophyly of Elphinstonia is confirmed by the tests, and the further radiation of the subgenus into the „charlonia-group“, comprising the species E. penia (F reyer , 1852), E. transcaspica (Staudinger , 1882), E. lucilla Butler, 1886, E. bazae F abiano , 1993, E. charlonia (D onzel , 1842), respectively, and on the other hand the „tomyris-group“ including E. lessei Bernardi , 1957, E. ziayani Leestmans & Back , 2001, and E. tomyris C hristoph , 1884, become apparent. The respecive species, represented by several samples of different subspecies and populations, show a conspicuous phylogenetic signal and are unequivocally separable from each other. According to the phenotypic characters published earlier by the first author, it is somewhat surprising that E. penia (Freyer ) belongs to the „charlonia-group“, and not to the „tomyris-group“ Aside from that all recently published taxonomic results are affirmed. Especially E. charlonia (D onz .), E. penia (F rey .), E. bazae F abiano , and E. transcaspica (Stgr .), confirmed their status as valid species. This is also the fact for E. lessei Bernardi and E. ziayani L eestmans & Back . Concluding, the re-adjusted taxonomic relationships, the phylogenetic correlation, as well as biogeographic data are presented. -
A Newly Recorded Species, Euchloe Belemia (Lepidoptera: Pieridae) from China
Entomotaxonomia (2016) 38(1): 39–43, DOI: 10.11680/entomotax.2016010, ISSN 2095–8609 39 A newly recorded species, Euchloe belemia (Lepidoptera: Pieridae) from China Changping DING, Yalin ZHANG① Key Laboratory of Plant Protection Resources and Pest Management of the Ministry of Education; Entomological Museum, Northwest A&F University, Yangling, Shaanxi 712100, China Abstract: The species Euchloe belemia (Esper, 1800) of Pieridae is recorded for the first time from China. Characteristics of the genus and the species are revised, and detailed photos/illustrations of male adult are given. A checklist of the species in this genus is provided. Key words: butterflies; Papilionoidea; taxonomy; checklist 中国新纪录种——绿纹荣粉蝶 Euchloe belemia 记述(鳞翅目:粉蝶科) 丁昌萍,张雅林① 植保资源与病虫害综合治理教育部重点实验室,西北农林科技大学昆虫博物馆,陕西 杨凌 712100 摘要:记述中国粉蝶科荣粉蝶属 Euchloe 1 新纪录种:绿纹荣粉蝶 Euchloe belemia (Esper, 1800)。并对 该属、种进行了描述,提供了该属的物种名录。 关键词:蝴蝶;凤蝶总科;分类;名录 Introduction The genus Euchloe was estiblished by Hübner (1819), with the type species Euchloe ausonia var. esperi Kirby, 1871 designated later by the International Commission on Zoological Nomenclature (ICZN) (Francis Hemming, 1954–1955). Euchloe ausonia var. esperi Kirby, 1871 was proposed as a synonymy of Euchloe crameri Butler, 1869 (Back, 1990, 2001) but not accepted. The genus comprises 25 species, which can be divided into two subgenera (Euchloe and Elphinstonia), distributed in Palaearctic and Nearctic regions (Bridges 1988; Back et al. 2006). This genus has been recorded by D'Abrera (1990, 1997), Winhard (2000), Shou (2006) and Wu (2010). In this study, the species Euchloe belemia (Esper, 1800) is recorded for the first time from China and both the genus and species redescribed with photos/illustrations. -
Über Die Biologie Von Euchloe Charlonia Amseli GROSS Und EBERT, 1975 (Lep., Pieridae) 48-60 ©Entomologisches Museum Dr
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Atalanta Jahr/Year: 1982 Band/Volume: 13 Autor(en)/Author(s): Back Werner Artikel/Article: Über die Biologie von Euchloe charlonia amseli GROSS und EBERT, 1975 (Lep., Pieridae) 48-60 ©Entomologisches Museum Dr. Ulf Eitschberger, download unter www.zobodat.at Atalanta, Bd. X III, Heft 1, April 1982, Würzburg, ISSN 0171-0079 Über die Biologie von Euchloe charlonia amseli GROSS und EBERT, 1975 (Lep., Pieridae) von WERNER BACK Die zuletzt 1976 von BACK und EITSCHBERGER erfolgte, weitgehend mit der klassischen Taxonomie übereinstimmende Aufgliederung der Unterarten von Euchloe charlonia muß nach den zuletzt vom Erstautor erzielten Zuchtergebnis sen in wesentlichen Punkten korrigiert werden. So weisen sowohl die Puppen als auch die Raupen der erst vor wenigen Jahren von GROSS und EBERT (1975) beschriebenen Euchloe charlonia amseli derar tig gravierende phänotypische Unterschiede im Vergleich zu Euchloe charlonia penia auf, daß hier nach den bisher üblichen taxonomischen Bewertungsmaßstä ben auf jeden Fall von verschiedenen Arten gesprochen werden muß. Die Bedeutung dieser Zuchtergebnisse für die Taxonomie und die Zuordnung verschiedener weiterer Unterarten von Euchloe charlonia soll nachfolgend eben falls diskutiert werden. Beschreibung der Zuchten von Euchloe charlonia amseli In der Provinz Chuzistan am Persischen Golf (Ramshir-Omidiye, 100 m) wurden am 12.IV .1978 zusammen mit Herrn Dr. E. REISSINGER, Kaufbeuren, zahlrei che Eier und Raupen aller Altersstufen an gelbblühenden Cruciferen gefunden. Die Schoten dieses Kreuzblütlers sind lang mit kurzem Schnabel und zweireihi gen, zahlreichen Samen (vgl. auch Abb.5); die grundständigen, rosettartigen Blät ter sind stark bebuchtet; vermutlich handelt es sich um Diplotaxis sp. -
Türkiye'deki Kelebeklerin Kırmızı Kitabı
RedTürkiye’deki Book of ButterfliesKelebeklerin inKırmızı Turkey Kitabı Evrim Karaçetin ve Hilary J. Welch Türkiye’deki Kelebeklerin Kırmızı Kitabı Karaçetin, E. ve Welch, H.J. 2011 Türkiye’deki Kelebeklerin Kırmızı Kitabı Doğa Koruma Merkezi, Ankara, Türkiye Erişim: [www.dkm.org.tr] © Doğa Koruma Merkezi (DKM), Şubat 2011 Ortadoğu Sitesi, 1589. Sok. No:4, Yüzüncüyıl, Ankara Tel: (312) 287 8144 Faks: (312) 286 6820 www.dkm.org.tr [email protected] 1. Basım Ankara, Şubat 2011-01-19 ISBN: 978-605-89908-1-4 Yazarlar: Evrim Karaçetin ve Hilary J. Welch Haritalar: Mustafa Durmuş, Alper Ertürk Editör: Burcu Meltem Arık Akyüz Grafik tasarım: Güngör Genç Çeviri: Gökhan Şahin, İkarus Tercüme Basım: Dumat Ofset Matbaacılık tel: 0312.278 82 00 Bu kitabın her hakkı saklıdır. Tamamen ya da kısmen çoğaltılması ve metindeki bilgilerin kullanılması Doğa Koruma Merkezi’nin yazılı izni alınmadıkça mümkün değildir. Bilimsel araştırma, tez, makale, kitap ve benzeri eserlerde, kitabın ve Doğa Koruma Merkezi’nin tam adı belirtilerek atıf yapılabilir. Referans gösterme: Karaçetin, E. ve Welch, H.J. (2011). Türkiye’deki Kelebeklerin Kırmızı Kitabı. Ankara: Doğa Koruma Merkezi. Erişim: [www.dkm.org.tr] Ön kapaktaki fotoğraflar: Euphydryas aurinia, Anthocharis cardamines, Polyommatus menalcas ©Onat Başbay : Lycaena dispar ©Bahar Bilgen : probable Polyommatus artvinensis ©Frédéric Bu kitapta kullanılan bütün fotoğrafların kullanım hakkı fotoğrafçılara Carbonell : Polyommatus bollandi ©Dominique aittir. Fotoğraflar, hak sahibinin yazılı izni olmadan çoğaltılamaz ya da Dumont : Polyommatus caeruleus, Polyommatus theresiae başka amaçlarla kullanılamaz. ©Hesselbarth ve ark. (1995) : Polyommatus merhaba ©Evrim Karaçetin : Erebia ottomana ©Matt Rowlings Türkiye’deki Kelebeklerin Kırmızı Kitabı, Hollanda Ekonomi, Tarım ve İnovasyon Bakanlığı tarafından finanse edilen projenin bir etkinliğidir; Arka kapaktaki fotoğraflar: Bakanlık, bu yayının içeriğinden sorumlu değildir. -
DIVERSITY of BUTTERFLIES (Lepidoptera: Papilionoidea & Hesperioidea) in PRESPA NATIONAL PARK (ALBANIA) 1. Introduction
DIVERSITY OF BUTTERFLIES (Lepidoptera: Papilionoidea & Hesperioidea) IN PRESPA NATIONAL PARK (ALBANIA) Dime Melovski Macedonian Ecological Society, Skopje, Macedonia [email protected] 1. Introduction 1.1 Literature review (present knowledge) Due to the fact that Albania was a prohibited country during the communist regime, not much research has been carried out in the period after the Second World War up until the 1990s. The research done in that period was in a form of solemn visits of expeditions and notifications of Moucha (1963a, 1963b), Alberti (1965) and Popescu-Gorj (1971). Before this period there are several publications available that are presenting checklists of butterflies in Albania. The first one published is by Dr. Hans Rebel (Rebel, 1913) in which 89 species of butterflies have been listed for Albania. Later on, Rebel and Zerny (1931) provided much more comprehensive list publishing 167 butterfly species for the Albanian butterfly fauna. The first local entomologist who published a list of butterfly species was Murraj (1972) who compiled an identification key for 93 species (Verovnik & Popović, 2013a). Later on, Misja and Kurrizi (1984) updated the already produced list of species and came up with 180 species. Several new findings for the country were added, including Danaus chrysippus (Linnaeus, 1758) (published also as new for Albania by Luquet and Misja (1989)),Euphydryas maturna (Linnaeus, 1758), Satyrus actaea (Esper, 1781) and Pseudochazara mamurra (Herrich-Schäffer, 1852) (Verovnik & Popović, 2013a). After the 1990s the butterfly research in this country has slightly increased. Beshkov (1995) and Abadijev & Beshkov (1996) published three new species for the Albanian butterfly fauna: Muschampia proto (Ochsenheimer, 1808), Hipparchia senthes (Fruhstorfer, 1908) and Hipparchia volgensis (Mazochin-Porshnjakov, 1952), while Verovnik and Popović (2013b) reported as many during the survey in south-eastern Albania in July 2012: Colias aurorina Herrich-Schäffer, 1850, Pieris balcana Lorković, 1970 and Apatura iris (Linnaeus, 1758).