Eight-Legged Encounters—Arachnids, Volunteers, and Art Help to Bridge the Gap Between Informal and Formal Science Learning
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Curriculum Vitae Bradley Evan Carlson, Ph.D
Curriculum Vitae Bradley Evan Carlson, Ph.D. Byron K. Trippet Assistant Professor of Biology Wabash College Crawfordsville, IN 47933 Email: [email protected] Telephone: (765) 361-6460 Website: carlsonecolab.weebly.com Professional Experience 2014 - present Byron K. Trippet Assistant Professor of Biology, Wabash College Education 2009-2014 PhD in Ecology, minor in Statistics, The Pennsylvania State University, University Park, PA Advisor: Dr. Tracy Langkilde Dissertation: The evolutionary ecology of intraspecific trait variation in larval amphibians 2008 B.S. in Biology, Bethel University, St. Paul, MN Summa cum laude, Honors Graduate Thesis: Temperature and desiccation effects on the antipredator behavior of Centruroides vittatus (Scorpiones: Buthidae) Research Interests Evolutionary ecology – phenotypic diversity, local adaptation, trait integration Behavioral ecology – phenotypic plasticity, predator-prey interactions, personality traits Community ecology – trait-mediated indirect interactions, predation, aquatic ecology Zoology – herpetology, arachnology, comparative morphology Publications (*co-author was undergraduate) Kashon*, EAF, and BE Carlson. 2018. Consistently bolder turtles maintain higher body temperatures in the field but may experience greater predation risk. Behavioral Ecology and Sociobiology 72:9. Carlson, BE, and T Langkilde. 2017. Body size variation in aquatic consumers causes pervasive community effects, independent of mean body size. Ecology and Evolution 7:9978-9990. Lambert, MR, Carlson, BE, Smylie, MS, and L Swierk. 2017. Ontogeny of sexual dichromatism in the explosively breeding Wood Frog. Herpetological Conservation and Biology 12:447-456. Media coverage: InsideEcology.com (https://insideecology.com/2018/02/12/amphibians-that-change-colour/) Carlson, BE, and T Langkilde. 2016. The role of resources in microgeographic variation in Red- spotted Newt (Notophthalmus v. viridescens) morphology. Journal of Herpetology 50:442-448. -
Sexual Selection Research on Spiders: Progress and Biases
Biol. Rev. (2005), 80, pp. 363–385. f Cambridge Philosophical Society 363 doi:10.1017/S1464793104006700 Printed in the United Kingdom Sexual selection research on spiders: progress and biases Bernhard A. Huber* Zoological Research Institute and Museum Alexander Koenig, Adenauerallee 160, 53113 Bonn, Germany (Received 7 June 2004; revised 25 November 2004; accepted 29 November 2004) ABSTRACT The renaissance of interest in sexual selection during the last decades has fuelled an extraordinary increase of scientific papers on the subject in spiders. Research has focused both on the process of sexual selection itself, for example on the signals and various modalities involved, and on the patterns, that is the outcome of mate choice and competition depending on certain parameters. Sexual selection has most clearly been demonstrated in cases involving visual and acoustical signals but most spiders are myopic and mute, relying rather on vibrations, chemical and tactile stimuli. This review argues that research has been biased towards modalities that are relatively easily accessible to the human observer. Circumstantial and comparative evidence indicates that sexual selection working via substrate-borne vibrations and tactile as well as chemical stimuli may be common and widespread in spiders. Pattern-oriented research has focused on several phenomena for which spiders offer excellent model objects, like sexual size dimorphism, nuptial feeding, sexual cannibalism, and sperm competition. The accumulating evidence argues for a highly complex set of explanations for seemingly uniform patterns like size dimorphism and sexual cannibalism. Sexual selection appears involved as well as natural selection and mechanisms that are adaptive in other contexts only. Sperm competition has resulted in a plethora of morpho- logical and behavioural adaptations, and simplistic models like those linking reproductive morphology with behaviour and sperm priority patterns in a straightforward way are being replaced by complex models involving an array of parameters. -
Information to Users
INFORMATION TO USERS The most advanced technology has been used to photograph and reproduce this manuscript from the microfilm master. UMI films the text directly from the original or copy submitted. Thus, some thesis and dissertation copies are in typewriter face, while others may be from any type of computer printer. The quality of this reproduction is dependent upon the quality of the copy submitted. Broken or indistinct print, colored or poor quality illustrations and photographs, print bleedthrough, substandard margins, and improper alignment can adversely affect reproduction. In the unlikely event that the author did not send UMI a complete manuscript and there are missing pages, these will be noted. Also, if unauthorized copyright material had to be removed, a note will indicate the deletion. Oversize materials (e.g., maps, drawings, charts) are reproduced by sectioning the original, beginning at the upper left-hand corner and continuing from left to right in equal sections with small overlaps. Each original is also photographed in one exposure and is included in reduced form at the back of the book. Photographs included in the original manuscript have been reproduced xerographically in this copy. Higher quality 6" x 9" black and white photographic prints are available for any photographs or illustrations appearing in this copy for an additional charge. Contact UMI directly to order. University Microfilms International A Bell & Howell Information Company 300 North Zeeb Road. Ann Arbor, Ml 48106-1346 USA 313/761-4700 800/521-0600 Order Number 9111799 Evolutionary morphology of the locomotor apparatus in Arachnida Shultz, Jeffrey Walden, Ph.D. -
Insect and Arachnid Collection Quest
INSECT AND ARACHNID COLLECTION QUEST This quest encourages you to explore and investigate what you might normally ignore. To complete this quest, find 10 different species of insects, at least one species of arachnids, preserve the insects and arachnids, and then pin your collection. Once you have completed these steps, label the parts of the insects and the arachnids. At the end of this quest, you will receive a “Critters of Texas” pocket guide and points to use toward the trade items at Texas Nature Traders. STEP 1: Identify and collect 10 different insect species. The collection should include five different families of insects. STEP 2: Identify and collect at least one species of arachnids. STEP 3: Perform the proper procurement and preservation methods. STEP 4: Properly pin insects and arachnids. STEP 5: Label parts of insects: head, thorax and abdomen. Label parts of arachnids: cephalothorax and abdomen. STEP 6: Bring in the collection to Texas Nature Traders. FUN FACTS AND ANSWERS TO CURIOUS QUESTIONS What is the difference between insects and arachnids? o Mostly anatomical, such as: arachnids lack antennae; insects have mandibles and arachnids have fangs; insects have six legs and arachnids have eight. Can insects hear? o Yes. Insects use sounds to communicate and navigate their environment. Most insects have a pair of tympanal organs. In many cases, there is also a receptor on the antennae called the Johnston’s organ, which also collects sound vibrations. Are insects/arachnids considered animals? o Yes, they belong to the kingdom Animalia. Where in the world has the most mosquitoes? o Alaska has the most mosquitoes due to the melting snow, which leaves large amounts of standing water – perfect conditions for mosquito breeding. -
Zootaxa,A New Species of Mastigoproctus
Zootaxa 1463: 39–45 (2007) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA Copyright © 2007 · Magnolia Press ISSN 1175-5334 (online edition) A new species of Mastigoproctus Pocock, 1894 (Thelyphonida: Thelyphonidae) from Venezuela CARLOS VIQUEZ1 & LUIS F. DE ARMAS2 1Investigador Asociado. Instituto Nacional de Biodiversidad (INBio), Santo Domingo, Heredia, P. O. Box 22-3100, Costa Rica. E-mail: [email protected] 2P. O. Box 4327, San Antonio de los Baños, La Habana 32500, Cuba. E-mail: [email protected] Abstract A new species of Venezuelan whip scorpions belonging to the genus Mastigoproctus Pocock is herein described from Bolivar state. This new species mainly differs from the Venezuelan Mastigoproctus formidabilis Hirst, and the Colom- bian Mastigoproctus colombianus Mello-Leitão in male pedipalp sculpture and proportions as well as in female genitalia. Now, Mastigoproctus is represented in Venezuela by two apparently very related species. Key words: Thelyphonida, Uropygi, Mastigoproctus, taxonomy, South America, Venezuela Introduction Whip scorpions (Thelyphonida) form one of the smaller arachnid orders (Harvey, 2002, 2003), representing only 0.1% of the arachnids of the world. However, their large size (most are longer than 40 mm length from the pedipalps to the base of the flagellum) and distinctive vinegary odor make them better known for the peo- ple than some of its relatives, e.g., the ricinuleids and schizomids. Whip scorpions are represented in South America by only two genera: Mastigoproctus Pocock and Thely- phonellus Pocock, the first of which shows a higher species richness. A third genus, Amauromastigon Mello- Leitão, was synonymized with Mastigoproctus by Rowland (2002). -
Arachnida: Uropygi: Thelyphonida)
ARTÍCULO: Thelyphonellus venezolanus n. sp., a new species of vinegaroons (Arachnida: Uropygi: Thelyphonida) Joachim Haupt ARTÍCULO: Thelyphonellus venezolanus n. sp., a new species of vinegaroons (Arach- nida: Uropygi: Thelyphonida) Joachim Haupt Abstract: Institut für Ökologie / Biologie, A new species of the genus Thelyphonellus is described. So far, two species of Technische Universität Berlin, Thelyphonellus are known: Th. amazonicus (Butler, 1872) and Th. ruschii Franklinstr. 28/29, FR 1-1, (Weygoldt, 1979). A third species, Th. wetherbeei (Armas), 2002 was transfe- D-10587 Berlin. rred to another genus (Ravilops) (Víquez & Armas, 2005). The new species [email protected] perfectly fits the description of the genus given by Weygoldt (1979). It has more prominent teeth at the coxae of the pedipalps and lacks 'ommatidia' and whip organs entirely. The pedipalpal tibiae are inflated in the male. The male genital atrium is shorter and wider than in the other species. The female remains known. Key words: Arachnida, Uropygi, Venezuela, new species. Taxonomy: Thelyphonellus venezolanus n. sp. Thelyphonellus venezolanus n. sp., nueva especie de vinagrillo (Arach- Revista Ibérica de Aracnología nida: Uropygi: Thelyphonida) ISSN: 1576 - 9518. Dep. Legal: Z-2656-2000. Vol. 17 Sección: Artículos y Notas. Resumen: Pp: 63 − 65. Se describe una nueva especie del género Thelyphonellus. Durante largo tiem- Fecha publicación: 30 Noviembre 2009 po, tan sólo se conocían dos especies de Thelyphonellus: Th. amazonicus (Bu- tler, 1872) y Th. ruschii Weygoldt, 1979. Una tercera especie, Th. Wetherbeei (Armas) fue transferida a otro género (Ravilops) (Víquez & Armas, 2005). La Edita: nueva especie encaja perfectamente en la descripción del género dada por Grupo Ibérico de Aracnología (GIA) Weygoldt (1979). -
Kirill Glebovich Mikhailov: on the Occasion of His 60Th Birthday
Zootaxa 5006 (1): 006–012 ISSN 1175-5326 (print edition) https://www.mapress.com/j/zt/ Biography ZOOTAXA Copyright © 2021 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.5006.1.4 http://zoobank.org/urn:lsid:zoobank.org:pub:B883A8B0-F324-400F-B670-304511C53963 Kirill Glebovich Mikhailov: On the occasion of his 60th Birthday YURI M. MARUSIK1 & VICTOR FET2 1Institute for Biological Problems of the North, Portovaya Street 18, Magadan 685000, Russia Department of Zoology & Entomology, University of the Free State, Bloemfontein 9300, South Africa [email protected]; https://orcid.org/0000-0002-4499-5148 2Department of Biological Sciences, Marshall University, Huntington, West Virginia 25755-2510, USA [email protected]; https://orcid.org/0000-0002-1016-600X Kirill Glebovich Mikhailov was born on 29 July 1961 in Moscow, Russia. Both of his parents, Gleb K. Mikhailov (1929–2021) and Galina R. Mikhailova (1926–2019), were research scientists. Kirill’s father was an expert in the history of mechanics, and mother, a biologist. Since early childhood Kirill was raised mainly by his maternal grandparents, Ro- man P. Nosov and Antonina V. Nosova. Kirill’s grandfather, a CPSU official and a career administrator at the Ministry of Energetics, retired from his post in 1965 to take care of the grandson. Kirill’s grandmother, an obstetrician by profession, received disability at a military plant during the WWII in evacuation, and was a housewife after the war. In 1978, Kirill began his studies at the Division of Biology (Biologicheskii Fakul’tet) of the Moscow State University (below, MSU). Even earlier, as a schoolboy, Kirill used to buy books on zoology, especially separate issues of the Fauna of the USSR and Keys to the Fauna of the USSR, then relatively cheap and available. -
Homeowner Guide to Scorpions and Their Relatives
HOMEOWNER Guide to by Edward John Bechinski, Dennis J. Schotzko, and Craig R. Baird CIS 1168 Scorpions and their relatives “Arachnid” is the scientific classification category for all eight-legged relatives of insects. Spiders are the biggest group of arachnids, with nearly 3800 species known from the U.S and Canada. But the arachnid category includes other types of eight-legged creatures that sometime cause concern. Some of Idaho’s non-spider arachnids – such as scorpions -- pose potential threats to human health. Two related non-spider arachnids – sun scorpions and pseudoscorpions – look fearsome but are entirely harmless. This publication will help you identify these three groups and understand the threats they pose. All three of these groups almost always are seen as lone individuals that do not require any control. Scorpions IDENTIFICATION AND BIOLOGY FLUORESCENT SCORPIONS Scorpions are easily identified by their claw-like pincers at the The bodies of some scorpions – normally pale tan to darker red-brown – front of the head and their thin, many-segmented abdomen that glow yellow-green when exposed to ultraviolet light. Even fossils millions ends in an enlarged bulb with a curved sting at the tip (figure 1). of years old fluoresce under ultraviolet light. Sun spiders similarly glow yel- Five species ranging in size from 2 to 7 inches long occur in low-green under UV light. Idaho. Scorpions primarily occur in the sagebrush desert of the southern half of Idaho, but one species – the northern scorpion (Paruroctonus boreus)– occurs as far north as Lewiston, along the Snake River canyon of north-central Idaho. -
Phylogenomic Resolution of Sea Spider Diversification Through Integration Of
bioRxiv preprint doi: https://doi.org/10.1101/2020.01.31.929612; this version posted February 2, 2020. The copyright holder for this preprint (which was not certified by peer review) is the author/funder. All rights reserved. No reuse allowed without permission. Phylogenomic resolution of sea spider diversification through integration of multiple data classes 1Jesús A. Ballesteros†, 1Emily V.W. Setton†, 1Carlos E. Santibáñez López†, 2Claudia P. Arango, 3Georg Brenneis, 4Saskia Brix, 5Esperanza Cano-Sánchez, 6Merai Dandouch, 6Geoffrey F. Dilly, 7Marc P. Eleaume, 1Guilherme Gainett, 8Cyril Gallut, 6Sean McAtee, 6Lauren McIntyre, 9Amy L. Moran, 6Randy Moran, 5Pablo J. López-González, 10Gerhard Scholtz, 6Clay Williamson, 11H. Arthur Woods, 12Ward C. Wheeler, 1Prashant P. Sharma* 1 Department of Integrative Biology, University of Wisconsin–Madison, Madison, WI, USA 2 Queensland Museum, Biodiversity Program, Brisbane, Australia 3 Zoologisches Institut und Museum, Cytologie und Evolutionsbiologie, Universität Greifswald, Greifswald, Germany 4 Senckenberg am Meer, German Centre for Marine Biodiversity Research (DZMB), c/o Biocenter Grindel (CeNak), Martin-Luther-King-Platz 3, Hamburg, Germany 5 Biodiversidad y Ecología Acuática, Departamento de Zoología, Facultad de Biología, Universidad de Sevilla, Sevilla, Spain 6 Department of Biology, California State University-Channel Islands, Camarillo, CA, USA 7 Départment Milieux et Peuplements Aquatiques, Muséum national d’Histoire naturelle, Paris, France 8 Institut de Systématique, Emvolution, Biodiversité (ISYEB), Sorbonne Université, CNRS, Concarneau, France 9 Department of Biology, University of Hawai’i at Mānoa, Honolulu, HI, USA Page 1 of 31 bioRxiv preprint doi: https://doi.org/10.1101/2020.01.31.929612; this version posted February 2, 2020. The copyright holder for this preprint (which was not certified by peer review) is the author/funder. -
Giant Whip Scorpion Mastigoproctus Giganteus Giganteus (Lucas, 1835) (Arachnida: Thelyphonida (=Uropygi): Thelyphonidae) 1 William H
EENY493 Giant Whip Scorpion Mastigoproctus giganteus giganteus (Lucas, 1835) (Arachnida: Thelyphonida (=Uropygi): Thelyphonidae) 1 William H. Kern and Ralph E. Mitchell2 Introduction shrimp can deliver to an unsuspecting finger during sorting of the shrimp from the by-catch. The only whip scorpion found in the United States is the giant whip scorpion, Mastigoproctus giganteus giganteus (Lucas). The giant whip scorpion is also known as the ‘vinegaroon’ or ‘grampus’ in some local regions where they occur. To encounter a giant whip scorpion for the first time can be an alarming experience! What seems like a miniature monster from a horror movie is really a fairly benign creature. While called a scorpion, this arachnid has neither the venom-filled stinger found in scorpions nor the venomous bite found in some spiders. One very distinct and curious feature of whip scorpions is its long thin caudal appendage, which is directly related to their common name “whip-scorpion.” The common name ‘vinegaroon’ is related to their ability to give off a spray of concentrated (85%) acetic acid from the base of the whip-like tail. This produces that tell-tale vinegar-like scent. The common name ‘grampus’ may be related to the mantis shrimp, also called the grampus. The mantis shrimp Figure 1. The giant whip scorpion or ‘vingaroon’, Mastigoproctus is a marine crustacean that can deliver a painful wound giganteus giganteus (Lucas). Credits: R. Mitchell, UF/IFAS with its mantis-like, raptorial front legs. Often captured with shrimp during coastal trawling, shrimpers dislike this creature because of the lightning fast slashing cut mantis 1. -
Arachnid Or Insect?
Suggested levels for Guided Reading, DRA,™ Lexile,® and Reading Recovery™ are provided Life Science in the Pearson Scott Foresman Leveling Guide. ArachnidArachnid oror Insect?Insect? Comprehension Genre Text Features Skills and Strategy Expository • Compare and • Captions nonfiction Contrast • Labels • Main Idea • Callouts • Summarize Scott Foresman Reading Street 2.3.3 ISBN-13: 978-0-328-50838-9 ISBN-10: 0-328-50838-1 9 0 0 0 0 by Kristin Cashore 9 780328 508389 50838_CVR.indd Page 1-2 3/19/13 2:34 AM gg-041 /110/SF00327_R4/sf00327_1of1/work%250/indd%250/SF_RE_TX:NL_Leveled_G2/On/2.3.3O%2 ... Vocabulary been Arachnid believe caught or finally Insect? today tomorrow whatever Word count: 284 by Kristin Cashore Note: The total word count includes words in the running text and headings only. Numerals and words in chapter titles, captions, labels, diagrams, charts, graphs, sidebars, and extra features are not included. Glenview, Illinois • Boston, Massachusetts • Chandler, Arizona Upper Saddle River, New Jersey 50838_001-012.indd Page 1 3/19/13 2:22 AM gg-041 /110/SF00327_R4/sf00327_1of1/work%250/indd%250/SF_RE_TX:NL_Leveled_G2/On/2.3.3O%2 ... Many people believe that all bugs and creepy crawly things are insects. This is not true! A spider is not an insect. It is an arachnid. What is the difference? Photographs Every effort has been made to secure permission and provide appropriate credit for photographic material. The publisher deeply regrets any omission and pledges to correct errors called to its attention in subsequent editions. Unless otherwise acknowledged, all photographs are the property of Pearson Education, Inc. -
Giant Spiders Cast Webs Over River Using Super Biomaterial 17 September 2010
Giant spiders cast webs over river using super biomaterial 17 September 2010 access journal for the communication of all peer- reviewed scientific and medical research published by the Public Library of Science. Caerostris darwini, a giant orb spider and namesake of Charles Darwin, weaves a web of tremendous strength and size never before seen. Photo courtesy of Matjaz Gregoric. (PhysOrg.com) -- The antithesis of the itsy-bitsy spider, Caerostris darwini, a giant orb spider and namesake of Charles Darwin, weaves a web of super strength never before seen, says Dr. Todd Blackledge, Leuchtag Endowed Chair at The University of Akron. This close-up view of a giant orb spider web spanning a Blackledge, an associate professor of biology; Ingi river demonstrates the tremendous strength and elasticity of this type of silk. Photo courtesy of Matjaz Kuntner. Agnarsson, Blackledge’s former postdoctoral researcher at UA and current assistant professor and director of Museum of Zoology at the University of Puerto Rico; and Matjaž Kuntner, In their PLoS ONE paper, Agnarsson, Kuntner and chair of the Institute of Biology at the Scientific Blackledge report on the testing of material Research Centre, Slovenian Academy of Sciences properties of Darwin’s bark spider. The authors and Arts, reveal this new spider species from predicted that the expansive webs would be spun Madagascar and its incredibly expansive web. using extraordinary silk and they proved their prediction correct. First, the researchers considered “Darwin’s bark spider or C. darwini,” as it is that spider silks already combine high strength with referenced, makes one of the largest known webs elasticity, demonstrate exceptional toughness and and suspends the giant webs across rivers and are able to absorb three times more energy than lakes, according to the researchers.